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  • Angiosperms
  • Springer  (236)
  • 1995-1999  (15)
  • 1990-1994  (221)
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  • Springer  (236)
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Year
  • 1
    Electronic Resource
    Electronic Resource
    DNA probes for the Y-chromosome ofSilene latifolia, a dioecious angiosperm (1992)
    Springer
    Sexual plant reproduction 5 (1992), S. 86-88 
    Details
    ISSN: 1432-2145
    Keywords: Sex chromosome markers ; Y-chromosome ; Angiosperms ; Silene latifolia ; Melandrium album
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary In order to obtain markers for the Y chromosome ofSilene latifolia, we pooled equal weights of leaf tissue from 18 female siblings into one sample and repeated the process with 18 male siblings. Pooling was intended to provide a common genetic background for each sample, leaving the absence or presence of the Y chromosome as the primary difference between the two samples. DNA was extracted from each sample and subjected to polymerase chain reaction (PCR) amplification with arbitrary 10 bp primers. Four of 60 primers used gave an amplification with the male DNA not found among those from the female DNA. Each of these was subsequently shown to provide a reliable marker for the Y chromosome.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00714562
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  • 2
    Electronic Resource
    Electronic Resource
    Fingerprinting plant genomes with oligonucleotide probes specific for simple repetitive DNA sequences (1992)
    Springer
    Theoretical and applied genetics 83 (1992), S. 691-694 
    Details
    ISSN: 1432-2242
    Keywords: DNA fingerprinting ; Repetitive DNA ; Genotype identification ; Angiosperms
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Oligonucleotides hybridizing to simple repetitive DNA patterns are highly informative as probes for DNA fingerprinting in all investigated animal species, including man. Here we demonstrate the applicability of this technique in higher plants. The oligonucleotide probes (GTG)5 and (GATA)4 were used to investigate the differences in DNA fingerprint patterns of the following angiosperm species: Triticum aestivum, Secale cereale, Hordeum vulgare, Beta vulgaris, Petunia hybrida, Brassica oleracea, and Nicotiana tabacum. Two species, Hordeum vulgare as a monocot and Beta vulgaris as a dicot, were analyzed in more detail. Their genomes differ considerably in both amount and organization of the simple repetitive sequences (GATA)n, (GACA)n, (GTG)n, and (CT)n due to the evolutionary distance of these two species. Furthermore, several lines and cultivars of Beta vulgaris and Hordeum vulgare can clearly be distinguished on the basis of their highly polymorphic patterns of these repetitive sequences.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00226686
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  • 3
    Electronic Resource
    Electronic Resource
    Angiosperm origin and early stages of seed plant evolution deduced from rRNA sequence comparisons (1991)
    Springer
    Journal of molecular evolution 32 (1991), S. 253-261 
    Details
    ISSN: 1432-1432
    Keywords: Chloroplast 4.5S rRNA ; Cytosolic and chloroplast 5S rRNAs ; 5.8S rRNA ; 18S rRNA ; Nucleotide sequences ; Phylogenetic trees ; Angiosperms ; Gymnosperms ; Monocotyledons ; Dicotyledons
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Complete or partial nucleotide sequences of five different rRNA species, coded by nuclear (18S, 5.8S, and 5S) or chloroplast genomes (5S, 4.5S) from a number of seed plants were determined. Based on the sequence data, the phylogenetic dendrograms were built by two methods, maximum parsimony and compatibility. The topologies of the trees for different rRNA species are not fully congruent, but they share some common features. It may be concluded that both gymnosperms and angiosperms are monophyletic groups. The data obtained suggest that the divergence of all the main groups of extant gymnosperms occurred after the branching off of the angiosperm lineage. As the time of divergence of at least some of these gymnosperm taxa is traceable back to the early Carboniferous, it may be concluded that the genealogical splitting of gymnosperm and angiosperm lineages occurred before this event, at least 360 million years ago, i.e., much earlier than the first angiosperm fossils were dated. Ancestral forms of angiosperms ought to be searched for among Progymnospermopsida. Genealogical relationships among gymnosperm taxa cannot be deduced unambiguously on the basis of rRNA data. The only inference may be that the taxon Gnetopsida is an artificial one, andGnetum andEphedra belong to quite different lineages of gymnosperms. As to the phylogenetic position of the two Angiospermae classes, extant monocotyledons seem to be a paraphyletic group located near the root of the angiosperm branch; it emerged at the earliest stages of angiosperm evolution. We may conclude that either monocotyledonous characters arose independently more than once in different groups of ancient Magnoliales or that monocotyledonous forms rather than dicotyledonous Magnoliales were the earliest angiosperms. Judging by the rRNA trees, Magnoliales are the most ancient group among dicotyledons. The most ancient lineage among monocotyledons leads to modern Liliaceae.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF02342748
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  • 4
    Electronic Resource
    Electronic Resource
    Nucleotide distribution in gymnosperm nuclear sequences suggests a model for GC-content change in land-plant nuclear genomes (1994)
    Springer
    Journal of molecular evolution 39 (1994), S. 34-46 
    Details
    ISSN: 1432-1432
    Keywords: CpG suppression ; GC content ; Angiosperms ; Isochores ; GC bias ; Mutational pressure ; Error-prone repair ; Transcriptionally coupled repair
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Nuclear protein coding sequences from gymnosperms are currently scarce. We have determined 4 kb of nuclear protein coding sequences from gymnosperms and have collected and analyzed 〉60 kb of nuclear sequences from gymnosperms and nonspermatophytes in order to better understand processes influencing genome evolution in plants. We show that conifers possess both biased and nonbiased genes with respect to GC content, as found in monocots, suggesting that the common ancestor of conifers and monocots may have possessed both biased and nonbiased genes. The lack of biased genes in dicots is suggested to be a derived character for this lineage. We present a simple but speculative model of land-plant genome evolution which considers changes in GC bias and CpG frequency, respectively, as independent processes and which can account for several puzzling aspects of observed nucleotide frequencies in plant genes.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00178247
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  • 5
    Electronic Resource
    Electronic Resource
    The evolution of angiosperm seed proteins: A methionine-rich legumin subfamily present in lower angiosperm Clades (1996)
    Springer
    Journal of molecular evolution 43 (1996), S. 399-404 
    Details
    ISSN: 1432-1432
    Keywords: Asarum ; Dioscorea ; Angiosperms ; Evolution ; Legumins ; Seed proteins
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Analysis of legumin-encoding cDNAs fromDioscorea caucasica Lipsky (Dioscoreaceae) and fromAsarum europaeum L. (Aristolochiaceae) shows that there is an especially methionine-rich legumin subfamily present in the lower angiosperm clades including the Monocotyledoneae. It is characterized by a methionine content of 3–4 mol% which is roughly triple the methionine proportion of most other legumins. These “MetR” legumins, if present, still have to be detected in the higher angiosperms including the important seed crops. Evolutionary analysis suggests that the MetR legumins are the result of a gene duplication allowing the differentiation of legumin genes according to their sulfur content. The duplication event must have taken place before the split into mono- and dicotyledonous plants but probably after the separation of angiosperms and gymnosperms.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF02339013
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  • 6
    Electronic Resource
    Electronic Resource
    Isolation of the male germ unit: organization and function in tobacco (Nicotiana tabacum L.) (1998)
    Springer
    Plant cell reports 18 (1998), S. 143-147 
    Details
    ISSN: 1432-203X
    Keywords: Key wordsNicotiana tabacum ; Male germ unit ; Scanning electron microscopy ; Sperm isolation ; Angiosperms
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Sperm cells are released from pollen tubes of tobacco as linked cells, associated with the vegetative nucleus in an assemblage known as the male germ unit (MGU). Using light microscopy, the MGU assemblage appears to be ensheathed by cytoplasmic material of the pollen tube, which may stabilize their association. Following their release, the shape of the sperm cells and vegetative nucleus changes from an ellipsoidal to a more spheroidal morphology. When most of the cytoplasmic material is dispersed, a boundary remains around the two sperm cells. Using scanning electron microscopy, the cytoplasmic material surrounding the MGU appears filamentous, sometimes twisted and rope-like. Based on these observations, the function of the MGU of tobacco is discussed.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/s002990050547
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  • 7
    Electronic Resource
    Electronic Resource
    Seed size, pollination costs and angiosperm success (1991)
    Springer
    Evolutionary ecology 5 (1991), S. 231-247 
    Details
    ISSN: 1573-8477
    Keywords: Angiosperms ; pollination ; seed size
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Seed plants capture pollen before seeds are dispersed and abort unpollinated ovules. As a result, each seed is associated with an accessory cost that represents the costs of pollen capture and the costs of aborted ovules. Accessory costs may explain the minimum seed size among species, because these costs are likely to comprise a greater proportion of total reproductive allocation in species with smaller seeds. For very small propagules, the costs of pollination may not be worth the benefits, perhaps explaining the persistence of pteridophytic reproduction at small propagule sizes. The smallest angiosperm seeds are much smaller than the smallest gymnosperm seeds, both in the fossil record and in the modern flora. This suggests that angiosperms can produce pollinated ovules more cheaply than gymnosperms. Pollination becomes less efficient as a species decreases in abundance, and this loss of efficiency is greater for species with a higher accessory cost per seed. We propose that the greater reproductive efficiency of angiosperms when rare can explain why angiosperm-dominated floras were more speciose than the gymnosperm-dominated floras they replaced.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF02214230
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  • 8
    Electronic Resource
    Electronic Resource
    Double fertilization inEphedra trifurca, a non-flowering seed plant: The relationship between fertilization events and the cell cycle (1991)
    Springer
    Protoplasma 165 (1991), S. 106-120 
    Details
    ISSN: 1615-6102
    Keywords: Angiosperms ; Cell cycle ; Double fertilization ; Endosperm ; Ephedra
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Fertilization inEphedra trifurca was examined with a combination of light and fluorescence microscopy. Developmental analysis clearly indicates that double fertilization events, similar to those described inE. nevadensis, regularly occur during the process of sexual reproduction inE. trifurca. In addition to the typical fusion of a sperm nucleus and egg nucleus, a second fertilization event occurs between the second sperm nucleus from an individual pollen tube and the ventral canal nucleus. Both of the fertilization events take place within the confines of an individual egg cell of the female gametophyte. Microspectrofluorometric data demonstrate that each nucleus involved in a sexual fusion event proceeds through the synthesis phase of the cell cycle and increases its DNA content from 1C to 2C before the process of nuclear fusion is completed. Photometric data also confirm that the product of the second fertilization event is equal in DNA content (4C) to the zygotic nucleus derived from the first fertilization event, and is prepared to enter into mitosis as a fully functional diploid nucleus.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF01322281
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  • 9
    Electronic Resource
    Electronic Resource
    Distribution of naturally occurring quinones (1991)
    Springer
    Pharmacy world & science 13 (1991), S. 70-73 
    Details
    ISSN: 1573-739X
    Keywords: Algae ; Angiosperms ; Arthropods ; Bacteria ; Fungi ; Lichens ; Marine animals ; Quinones, naturally occurring
    Source: Springer Online Journal Archives 1860-2000
    Topics: Chemistry and Pharmacology
    Notes: Abstract Angiosperms, fungi (including lichens), and bacteria are the main sources of natural quinones. Small numbers are present in algae, ferns, conifers, sponges, echinoderms, other marine animals, and arthropods. In angiosperms quinones have some chemotaxonomic value at the genus and family level but more surveys are required.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF01974983
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  • 10
    Electronic Resource
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    Unidirectionality of floral colour changes (1996)
    Springer
    Plant systematics and evolution 200 (1996), S. 125-140 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Colour change ; floral colour phase ; innate flower detection ; colour preference ; pollination
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Many angiosperms have arranged their flowers in inflorescences forming a distinct signalling unit to flower visitors. In some species, the flowers of inflorescences undergo a temporal colour change corresponding exactly to a change in the reward status. Based on information obtained from the spectral reflection curves of pre-change and postchage colours of flower corollas and/or floral guides, it was possible to demonstrate that the colour phase associated with reward closely corresponds to the visual stimuli which trigger behavioural responses of inexperienced flower visitors, and that the colour phase associated with less reward corresponds to visual stimuli less attractive to naïve flower visitors. Reciprocal colour changes were not observed. It is to be assumed that the unidirectionality of floral colour changes is an adaptation of angiosperms aimed at the guidance of first-time flower visitors. Signalling reward to inexperienced flower visitors is an additional function of floral colour changes. The main function of floral colour changes, however, is to provide cues with which the flower visitors can learn to associate one colour phase with reward.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00984753
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  • 11
    Electronic Resource
    Electronic Resource
    Foliar anatomy ofPolygonum (Polygonaceae): Survey of epidermal and selected internal structures (1992)
    Springer
    Plant systematics and evolution 182 (1992), S. 71-106 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; dicotyledons ; Polygonaceae ; Polygonum ; Crystals ; extrafloral nectaries ; leaf anatomy ; secretory structures ; stomata ; subepidermal fibers ; trichomes
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Several anatomical characters in leaves were described, and their distribution determined, for 153 species ofPolygonum, mostly from herbarium specimens. Structures surveyed were epidermal (glandular and nonglandular trichomes, nodules, specialized parenchyma, stomatal apparatus) and internal (cavities, crystals, laticifer-like cells, nodules, subepidermal fibers). Cleared leaves were supplemented by resin-embedded sections and SEM preparations of selected species. No feature defines any taxonomic section, but some features occur only within one section. Laticifer-like cells, epidermal and internal nodules, resin cups, and unique epidermal and subepidermal cavities seem to be unknown elsewhere; other features (invaginated epidermal cells; enlarged crystal cells confined to paraveinal layer) are rare among angiosperms.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00941415
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  • 12
    Electronic Resource
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    Evolution ofPeperomia (Piperaceae) in the Juan Fernandez Islands, Chile (1992)
    Springer
    Plant systematics and evolution 182 (1992), S. 107-119 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Piperaceae ; Peperomia ; Flavonoids ; cytology ; phenetics ; cladistics ; Flora of the Juan Fernandez Islands ; Chile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Four species ofPeperomia (Piperaceae) occur in the Juan Fernandez Islands, Chile:P. berteroana, P. margaritifera, P. skottsbergii, andP. fernandeziana. The last species is found also in continental Chile, whereas the other three are endemic to the archipelago.Peperomia margaritifera is found only on the older island of Masatierra, whereasP. skottsbergii is confined to the younger island of Masafuera, andP. berteroana occurs on both islands. Phenetic analyses of mainland taxa suggest thatP. fernandeziana belongs to subg.Sphaerocarpidium whereas the endemic taxa form their own subg.Tildenidium connecting to subg.Tildenia. Cladistic analyses indicate thatP. margaritifera is the most primitive species in the archipelago and thatP. berteroana is the most derived, especially patristically. Chromosomally, the four species are all n = 22, which may be tetraploid on a base of x = 11. Sulfated flavones occur only inP. berteroana andP. skottsbergii, which are otherwise unknown for the family. Dispersal of propagules to the islands from the continent and between islands is believed to have been accomplished by birds.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00941416
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  • 13
    Electronic Resource
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    Asphodelus tenuifolius andA. fistulosus (Liliaceae) are morphologically, genetically, and biologically different species (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 1-12 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Liliaceae ; Asphodelus tenuifolius ; A. fistulosus ; Cytogenetics ; electrophoretics ; morphology ; duplication genes ; speciation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The biological analysis of six populations ofAsphodelus tenuifolius and 12 populations ofA. fistulosus has confirmed that they are separate species. Both their floral structures (length of the tepals, stamens, anthers and style) and also their pollen size are clearly different.A. tenuifolius has only the 2n = 28 chromosome race, whileA. fistulosus has 2n = 28 and 2n = 56.A. tenuifolius is genetically less variable thanA. fistulosus and they have different electrophoretic mobilities. Gene duplication phenomena exist in the 2n = 28 level of both species.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00935979
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  • 14
    Electronic Resource
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    Allozyme and life history variation in five northwardly colonizing North American weed species (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 41-54 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Abutilon theophrasti ; Datura stramonium ; Panicum miliaceum ; Sorghum halepense ; Setaria faberi ; Weeds ; allozymes ; life history ; variation ; Flora of N. America
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The relationships of allozyme and life history variation in a particularly narrow ecological setting are studied. Levels of genetic variation are compared in five introduced, predominantly selfing weedy species that are undergoing rapid range expansion northward in eastern N. America, mostly in monocultures of soybean and maize. In all of these species, a low level of allozyme variation contrasts sharply with the substantial inter- and intrapopulational variation in morphological and phenological life history features. Evolutionary and historical factors, determining variability of the species examined are reviewed, including founder effects, breeding system, environmental homogeneity, polyploidy, domestication, and crop-weed interactions.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00935983
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  • 15
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    A natural hybrid ofMonophyllaea (Gesneriaceae) in the tropical rain forests of West Sumatra (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 55-63 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Gesneriaceae ; Monophyllaea ; M. hirtella ; M. horsfieldii ; Chromosome number ; hybridization ; fertility ; tropical rain forest ; habitat segregation ; Flora of Indonesia ; Sumatra
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A natural hybrid (2n = 21) between the parapatric rain forest speciesMonophyllaea hirtella (2n = 20) andM. horsfieldii (2n = 22) (Gesneriaceae) has been observed at Sg. Lubuk Paraku, Padang, W. Sumatra. The hybrids showed intermediary characters between the parental species in the inflorescence structure, flower size and colour, indumentum, chromosome numbers, and habitat. The 29% pollen fertility of a single hybrid specimen suggests that the genetical isolation between the parental species may break down and reproduction over subsequent generations may occur.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00935984
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  • 16
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    Cytogenetic investigations onOenothera nutans (Onagraceae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 69-80 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Onagraceae ; Oenothera sect.Oenothera subsect.Oenothera ; O. nutans ; Chromosomal analysis ; complex analysis ; structural heterozygosity ; complex heterozygosity ; taxonomy ; numerical taxonomy ; factor analysis ; reciprocal translocations ; Sifactors ; lethal factors ; sublethal factors
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Oenothera nutans, common to the Appalachian Mts between 650 and 1 700 m altitude, was investigated cytogenetically and taxonomically. The species is permanently structurally heterozygous. It consists of two genomes of the B-type which are more or less indistinguishable phenotypically. Nearly all of the strains investigated possess a self-incompatibility factor in one of the two complexes. Both complexes show a close relationship to the predominantly homozygousO. grandiflora, a native of the southern lowlands.O. nutans andO. grandiflora possess the same plastid type, plastome III. Probably,O. nutans evolved by an accumulation of reciprocal translocations within an originally structurally homozygous population, which must be regarded ancestral to the present forms ofO. grandiflora.
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    URL: http://dx.doi.org/10.1007/BF00935986
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  • 17
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    Pollen amino acids—an additional diet for a nectar feeding butterfly? (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 111-121 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Aesculus californica ; Amsinckia lunaris ; Brodiaea pulchella ; Carduus pycnocephalus ; butterflies ; Battus philenor ; Heliconius ; Pollen amino acids ; diffusion ; nectar ; butterfly-feeding
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The increase of the amino acid concentration over different time intervals in artificial nectar (i.e., a sucrose solution) due to pollen contamination was investigated in four Californian plant species (Aesculus californica, Amsinckia lunaris, Brodiaea pulchella, Carduus pycnocephalus), which are important nectar resources for a Californian colony of the butterflyBattus philenor as well as for other insects. The increase of the amino acid concentration in the medium is different in all four species and seems to be determined by a variety of factors including permeability of the pollen grain wall and presence or absence of pores. The results suggest a passive diffusion process of the free pollen amino acids into the medium rather than an active release. Implications from the experiments forBattus philenor and for other nectar feeding pollinators are discussed. A possible complementary effect of free pollen and nectar amino acids is proposed for plant species in which pollen is likely to be knocked into nectar by their flower visitors. A possible evolutionary pathway from nectar feeding butterflies such asBattus philenor to the complex derived pollen feeding habit in theHeliconius butterflies is proposed.
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    URL: http://dx.doi.org/10.1007/BF00935989
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  • 18
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    Ant nests as primary habitats ofSilybum marianum (Compositae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 209-217 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Compositae ; Silybum marianum ; Formicidae ; Messor semirufus ; Myrmecochory ; preadaptations ; synanthropic plant ; grasslands
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The common Mediterranean ruderal thistleSilybum marianum is associated with nutrient-rich sites. Its wind-dispersed achenes possess an oily food body, that is attractive to harvester-ants. Following removal of the oily body, the achenes are deposited in the refuse zone together with rich organic material and soil removed from the nest; while in the nest the achenes are partly protected from fires. The thistle grows successfully in the nutrient-enriched refuse zone and thus dominates patches in the grassland. Preadaptations to live in association with harvester-ants enableS. marianum to occur also on marking stations of male gazelle, on cattle dung deposits, and in synanthropic ruderal habitats.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00937675
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  • 19
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    Phylogenetic relationships and gene flow between sympatric diploid and tetraploid plants ofDactylis glomerata (Gramineae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 81-96 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Gramineae ; Festuceae ; Dactylis glomerata ; Enzymatic markers ; phenology ; interploid exchanges ; autopolyploidy ; evolution
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Phylogenetic relationships between sympatric, morphologically indistinguishable diploid and tetraploid plants ofDactylis glomerata L. (Gramineae) in Galicia (Spain) were assessed using allozyme markers for 6 distinct systems. The study exploited recent introduction in Galicia and subsequent hybridization of an alien 4xDactylis subspecies possessing distinct allozymes from those of all the native plants. Opportunities for gene exchanges between the ploidies were estimated from in situ observations of flowering, examination of progenies in 2x/4x natural and experimental crosses, and enzyme analyses. Results show a high genetic similarity between the Galician diploids and tetraploids, which possess peculiar alleles in common. Although the ploidy levels usually have distinct flowering periods, interploidal crosses do occasionally occur. Gene flow is likely much more important from the diploid to the tetraploid level. A good genetic intermixing occurs between the Galician and the alien tetraploid entities which have simultaneous flowering. Autopolyploidization of the diploids followed by various rates of hybridization is proposed as one very probable origin of natural tetraploids inDactylis.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00935987
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  • 20
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    Invasive tapetum and tricelled pollen inAmbrosia trifida (Asteraceae, tribeHeliantheae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 237-243 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Heliantheae ; Ambrosia ; Anther ; crystals ; pollen ; sperm cells ; tapetum
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Staminate flowers of giant ragweed,Ambrosia trifida L. (Asteraceae, tribeHeliantheae, subtribeAmbrosiinae) were processed into resin and sectioned 1–2 µm thick. The invasive (amoeboid) anther tapetum remains parietal until microspores are released from tetrads, then it swells and invades the locule, merging gradually into a single protoplast that flows among the microspores. After the tapetal membrane ruptures at late microspore stage, tapetal debris fills the locule, then disappears as pollen matures. Pollen becomes tricelled before anthesis. The two sperm cell nuclei are slender and wormlike. The present report supports the two generalizations that invasive tapetum and tricelled pollen are attributes of theAsteraceae.
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    Distribution of diploid sexual plants ofTaraxacum sect.Ruderalia in east-Central Europe, with special reference to Czechoslovakia (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 71-84 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Taraxacum sect.Ruderalia ; Sexual diploids ; mixed populations ; distribution pattern ; ecogeographical relations ; taxonomical problems
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Herbarium specimens (c. 1 000) and population samples (76) have been screened for the occurrence of diploid representatives ofTaraxacum sect.Ruderalia (T. sect.Vulgaria, nom. illeg.). The area studied comprises German Democratic Republic, Poland, Czechoslovakia, Hungary and parts of Austria, Yugoslavia, and Rumania. Diploids are common from the Valley of the Moravia river on southwards. Isolated outpost localities are found in S. Poland and extend far to the north: neighbourhood of Berlin. Based on the pattern of distribution in Czechoslovakia, the Pannonian (thermophilous) phytogeographical character of the diploids is discussed. Special attention has been paid to morphological differences between di- and triploids in Czechoslovakia. The study provides a basis for further biosystematical and taxonomic studies in the relationships between diploids and polyploids as well as in the stability of agamospecies.
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    Transfer cells and lipid droplets of someValerianaceae (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 37-51 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Valerianaceae ; Transfer cells ; lignification ; lipid droplets ; oil cells ; tannins ; Development ; structure ; axial distribution ; transmission electron microscopy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Development, structure and the axial distribution of transfer cells and their lignification were investigated inValerianella locusta, Valeriana officinalis, andV. tuberosa (Valerianaceae). Fundamental new results are: (1) Transfer cells often contain numerous lipid droplets. Within the stem the distribution of cells containing lipid droplets correlates to that of transfer cells. (2) InValeriana officinalis persisting protuberances are frequently found on pit membranes of xylem transfer cells. Lignified transfer cells can undergo a second modification: a layer covering the secondary wall forms wall ingrowths similar to those of transfer cells. (3) Peripheral pith cells, abuting transfer cells, are able to modify into transfer cells. Cambial derivatives are only temporarily developed as transfer cells. (4) Phloem transfer cells are found in vascular bundles of the whole axis. (5) In roots, xylem transfer cells are poorly developed or absent. (6) Oil cells with oil bodies are present in the rape ofValeriana tuberosa. They are absent however in the stem of the species investigated. (7) Tannins occur in elements of the primary cortex, phloem and secondary xylem ofValeriana officinalis.
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    Hair morphology and systematics ofPhysalis (Solanaceae) (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 193-204 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Solanaceae ; Physalis ; Hair morphology ; systematics ; hair development
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Hair morphology was exammed on developing seedlings and mature plants of ten species from five of seven series inPhysalis. The taxonomic importance of hair development and the distribution of hair types then was evaluated with respect to other comparative data. Two classes of hair types were observed, as found in earlier investigations ofSolanum andRhododendron. The first hair class comprises a sequence from unbranched hairs to dendroid-stelliform hairs. Hairs of this class show considerable variability among taxa in size, presence or absence of glandular tips, and presence and degree of branching, and so were useful in making taxonomic comparisons. The second hair class consists of short hairs, each tipped with a multicellular gland. This second hair class was not useful taxonomically because of its lack of variability in morphology and its often rare occurrence on a plant. The taxa studied exhibit distinctive trichome features which serve to distinguish series and, in some cases together with other data, provided new insight into previously enigmatic species relationships.
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    The cytotaxonomy ofEmilia spp. (Asteraceae: Senecioneae) occurring in Brazil (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 229-236 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Emilia ; Chromosome numbers ; C-banding ; cytogeography ; evolution
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Analysis of several populations in a large part of the distribution area of the genusEmilia in Brazil has revealed only two species: the diploidE. sonchifolia and the tetraploidE. fosbergii. The more widely reportedE. coccinea was not found. They show a karyotype constancy in morphology and chromosome number (2n = 10 and 2n = 20, respectively), C-banding pattern and number of secondary constrictions. Some indications were found thatE. fosbergii may be an allopolyploid and that its ancestors had different genome sizes.
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    Reproductive biology ofPolyalthia littoralis (Annonaceae) (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 237-245 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Annonaceae ; Polyalthia littoralis ; Extended spiral thickenings ; flowering phenology ; reproductive biology ; self-pollination ; Malesian tropics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Flowering phenology of some annonaceous trees and reproductive biology ofPolyalthia littoralis (Annonaceae) were studied. The trees showed various types of flowering phenology within the family. Among them,P. littoralis had hermaphroditic and protogynous flowers, and exhibited continuous flowering throughout the year. Bagged flowers set fruits and seeds comparable to the control. The observations of meiotic stages and the results of castrated tests indicated no possibility of apomictic reproduction. The abscised anthers attached to torus by the extended spiral thickenings, and accomplished self-pollination. Outbreeding possibly occurs, but the following self-pollination guarantees seed set.
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    Pattern analysis in pseudanthia (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 57-88 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Inflorescence ; pseudanthium ; pseudocorolla ; flower symmetry
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Pseudanthia occur in more than 40 angiosperm families. With regard to the underlying inflorescence structure they can be classified into the following groups: (a) floral and (b) hyperfloral pseudanthia, each with (c) or without (d) pseudocorollas. Pseudanthia have developed along independent evolutionary lines and are not bound to a particular inflorescence structure. They are the result of (a) the specific morphological predisposition of the taxon concerned, (b) aggregation and diminution of the flowers, giving rise to the formation of an attraction unit (for animal pollination), (c) variation, and (d) selection. Ontogenetical abbreviation is regarded to play an essential role in the origin and elaboration of pseudanthia.
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    The systematic significance of surface features of theBalanophora tuber (Balanophoraceae) (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 129-134 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Balanophoraceae ; Balanophora ; Tuber surface morphology ; systematics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The surface ofBalanophora tubers consists of a nonepidermal layer made up of two distinctive types of cells, armature cells and stellate wart cells. Both cell types are provided with a heavy wall, and are dead at maturity. Stellate warts in the three species investigated,B. elongata, B. fungosa, andB. hansenii, séem to be uniform in appearance, but armature cells are extremely distinctive for each species. They are present in large, agglomerate masses in the first, singly or in very small clusters in the second, and as completely free, individual, acicular cells in the third species. Such differences are believed to be significant systematically, and the separation ofB. hansenii is thus probably justified. Notwithstanding superficial similarities, stellate warts do not seem to be comparable to the lenticels of other plants.
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    Ribosomal DNA spacer-length variation inSecale spp. (Poaceae) (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 205-220 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Secale ; Ribosomal DNA ; spacer subrepeats ; restriction fragment length polymorphism (RFLP) ; phenetic relationships
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Variation in ribosomal DNA spacer length was analysed in 23 populations of 12Secale spp. by restriction enzyme analysis. Digestion of rDNA with Taq I endonuclease enzyme yields spacer fragments that include the subrepeat array and the non-repetitive region downstream of the array. Extensive spacer length variation existed in most species with Taq I fragment lengths ranging from 0.9–3.1 kb. These length variants have been attributed to the differences in number of 134 bp spacer subrepeats within rDNA arrays.S. silvestre was the only species to exhibit a unique spacer length variant of 0.9 kb and this was shown to result from the presence of an extra Taq I site in the spacer. rDNA spacer length frequencies were determined for the species. These frequencies were used to derive phenetic relationships between the species by numerical taxonomic methods. In plots constructed fromGower's distance matrices,S. silvestre appeared well separated from the major cluster consisting of the other species. On the basis of morphological and cytogenetic criteria,S. silvestre is considered the most ancient species. The rDNA data is consistent with this interpretation as it shows a clear differentiation ofS. silvestre from all the other species based on length and nucleotide sequence composition of the spacer region.
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    Identification and partial characterization of two species-specific repeat families in the great millet (Sorghum vulgare, Poaceae) genome (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 249-257 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Sorghum vulgare ; great millet ; DNA hybridization ; repetitive DNA
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The 1.4 kbp Xba I and the 1.3 kbp EcoRI repeat families in great millet were partially characterized with respect to their genomic distribution and their homology with the EcoRI and Xba I families of five other millet DNAs. The digestions of great millet DNA using increasing amounts of the two enzymes show that these two families are disperse in nature. The hybridization of these two families to the genomic digests of great millet indicates that they are arranged in a clustered and scrambled manner. Similarly, the hybridization with the EcoRI and Xba I digests of five other millet DNAs reveals the speciesspecific nature of these two repeat families. The latter also hybridize to the total repetitive fraction of great millet isolated at a highly stringent temperature of 75°C suggesting that the members of these two families are probably largely homogeneous.
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    A study of genetic diversity among host-specific populations of the witchweedStriga hermonthica (Scrophulariaceae) in Africa (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 1-12 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Scrophulariaceae ; Striga hermonthica ; Hemiparasitism ; hostspecificity ; enzyme electrophoresis ; genetic diversity ; Flora of Burkina Faso ; Sudan
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Striga hermonthica is a root hemiparasite that attacks onlyGramineae, includingSorghum and millet for which it is a principal cause of lowered yield. Enzyme electrophoresis was used to investigate genetic diversity inStriga hermonthica and to determine the level of differentiation between host-specialized populations. Nine genetic loci coding eight enzymes were interpreted and data obtained from three populations: oneSorghum-adapted population from Sudan and two populations from Burkina Faso, oneSorghum-adapted and the other millet-adapted. Levels of polymorphism were similarly high in all three populations (P=0.625, A=2.6−2.8, H=0.293−0.401). Genotypic frequencies at most loci conformed to Hardy-Weinberg expectations in each population, consistent with outcrossing as predicted from previous studies of floral biology. Occasional heterozygote deficiencies were probably the result of Wahlund effect. The mean value of FST over the three populations was 0.068, indicating a slight to moderate level of genetic differentiation among the populations. The two Burkina Faso populations were more closely related (S=0.940, D=0.006) than either was to the Sudan population, suggesting that geographic separation is more important than host specialization in contributing to population differentiation. TheSorghum-adapted population was slightly closer to the Burkina FasoSorghum-adapted population (S=0.873, D=0.047) than to Burkina Faso millet-adapted population (S=0.851, D=0.074). The absence of substantial genetic divergence between host-specific populations ofStriga could result either from recent evolution of host-specialized strains or from strong selection for physiological specialization in the face of substantial gene flow between the populations.
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    Hair morphology inPlantago sect.Coronopus (Plantaginaceae) (1992)
    Springer
    Plant systematics and evolution 179 (1992), S. 107-113 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Plantago sect.Coronopus ; Hair morphology ; bottle-like hairs
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The presence of bottle-like hairs has been described in four hitherto uninvestigated taxa ofPlantago sect.Coronopus sensuDietrich. This hair type is a feature characteristic of sect.Coronopus.
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    Embryology ofMalaxis saprophyta, with comments on the systematic position ofMalaxis (Orchidaceae) (1992)
    Springer
    Plant systematics and evolution 179 (1992), S. 95-105 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Orchidaceae ; Epidendreae ; Malaxidinae ; Malaxis saprophyta ; Embryology ; systematics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract InMalaxis saprophyta, anther wall development corresponds to the Monocotyledonous type. The uninucleate tapetum is of secretory type and the endothecium develops U- and V-shaped thickenings on the inner tangential and radial walls. Cytokinesis is simultaneous; tetrahedral, isobilateral and T-shaped tetrads are formed which are compactly aggregated in pollinia. At anthesis the microspore tetrads are 2-celled. The ovule is anatropous, bitegmic and both integuments are dermal in origin. A single hypodermal cell develops directly into a megaspore mother cell. Embryo sac development is predominantly monosporic and less often bisporic. Irrespective of the type of development, the mature embryo sac is 6-nucleate. Although double fertilization occurs, the primary endosperm nucleus degenerates. Embryogeny is of the Onagrad type. The mature embryo lacks differentiation into cotyledon, plumule and radicle. The reticulate seed coat is formed entirely by the outer layer of outer integument. There are three sterile and three fertile valves in the ovary. Although initially parenchymatous, the entire three sterile valves in the ovary and the upper half of the three fertile valves become sclerified after fertilization. The embryological characters support the disputed systematic position ofMalaxis within subtribeMalaxidinae ofEpidendreae.
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    Allozyme divergence among species ofWolffia (Lemnaceae) (1995)
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    Plant systematics and evolution 197 (1995), S. 59-69 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Lemnaceae ; Wolffia ; Allozymes ; genetic divergence ; electrophoresis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The genusWolffia was surveyed electrophoretically at 14 allozyme loci. A total of 133 clones representing 10 of the 11 recognized species was examined. Genetic identities among most pairs of species are zero, with non-zero values ranging from 0.14 to 0.40.Wolffia angusta and the newly describedW. neglecta show the highest similarity, and the former species has an identity of 0.14 withW. australiana. The next highest similarity (0.34) occurs betweenW. globosa of Southeast Asia andW. cylindracea of southern Africa, which until recently, had generally been viewed as members of the same species. Other species showing some common alleles are members of a complex involvingW. arrhiza, W. columbiana, W. cylindracea, andW. globosa. WithinW. arrhiza, plants from South Africa and Europe are easily distinguished electrophoretically because each contains unique alleles at two loci. Strains from other parts of Africa vary at these loci and are not totally distinct from either the plants from South Africa or from Europe. Species ofWolffia are much more divergent at allozyme loci than the majority of congeners of flowering plants. This suggests that the species are quite old and that the difficulties in distinguishing taxa morphologically are the result of reduction rather than lack of divergence due to recent speciation. Because of the lack of shared alleles between the majority of species pairs inWolffia, enzyme electrophoresis provides limited resolution of species relationships in the genus.
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    Notes on the colour of pollen (1995)
    Springer
    Plant systematics and evolution 198 (1995), S. 235-252 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Pollen ; pollen colour ; pollen signal ; spectral reflection
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The spectral reflection of pollen in 67 plant species out of 28 families was measured by means of mass recording of pollen grains. Various types of spectral reflection curves were found, but 75% belonged to two categories: 1. Human-yellow pollen with strong reflection in the green and red, and low reflection in the ultraviolet and blue range of wavelengths. 2. Human-whitish pollen with strong reflection in the green and red and additional reflection of shorter wavelengths. It is shown that it is important to have information about the mode of the visual pollen display — crypsis or colour contrast against the corolla, pollen advertisement, or concealment — and the visual capabilities of the presumed pollinators in order to be able to discuss the signalling function of pollen colours.
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    Chemistry and micromorphology of compound epicuticular wax crystalloids (Strelitzia type) (1994)
    Springer
    Plant systematics and evolution 193 (1994), S. 115-123 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Strelitzia ; Heliconia ; Typha ; Benincasa ; Epicuticular wax ; wax ultrastructure ; wax chemistry ; systematics ; taxonomy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Certain non-liliiflorous taxa within the monocotyledons (e.g.,Strelitzia, Heliconia, Typha) are characterized by compound epicuticular wax rodlets (Strelitzia type). Similar rodlets are also encountered on the surface of the dicotyledonous plantBenincasa hispida (Thunb.)Cogn. Chemical analysis of the surface wax from both sources showed that the rodlets are chemically distinct. The rodlets of the monocotyledons consist exclusively of aliphatic wax lipids, mainly wax esters. In contrast, the rodlets ofBenincasa are cheifly composed of triterpenol acetates and triterpenols. Formation of rodlets is therefore interpreted as ultrastructural convergency. It is concluded that taxonomical studies on wax crystalloids can be misleading when interpreted in terms of micromorphology of crystalloids only.
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    Chemo- and karyotaxonomic studies on some rhizomatous species of the genusSymphytum (Boraginaceae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 31-39 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Boraginaceae ; Symphytum tuberosum complex ; S. grandiflorum agg ; Chemotaxonomy ; pyrrolizidine alkaloids ; triterpenes
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract InS. tuberosum subspp.tuberosum andnodosum, S. grandiflorum andS. ibericum the presence of the pyrrolizidine alkaloids lycopsamine, echimidine and symphytine could be demonstrated. The taxonS. tuberosum contains an unknown compound that seems to be specific for this taxon. This compound is not the pyrrolizidine alkaloid anadoline which has previously been reported for this species. It is possibly represented by a peak on GC/MS with a molecular ion peak at m/z 623 (as TMS derivative) and can be used as a chemotaxonomic marker for the speciesS. tuberosum. The pyrrolizidine alkaloid pattern of the two subspecies ofS. tuberosum reinforces the close relationship. Fresh material ofS. tuberosum contained the triterpene isobauerenol, but in herbarium material isobauerenol was lacking. InS. grandiflorum, neither fresh nor dried material contains isobauerenol. In herbarium material ofS. ibericum also no isobauerenol could be found. More extensive chemotaxonomical research is necessary to support the view thatS. abchasicum is more closely related toS. ibericum than toS. grandiflorum.
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    Investigations into the evolutionary history of theAntennaria rosea (Asteraceae: Inuleae) polyploid complex (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 97-110 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Compositae ; Inuleae ; Antennaria ; A. rosea ; Agamic complex ; agamospermy ; asexual reproduction ; clonal organism ; dioecy ; morphology ; numerical taxonomy ; phenetics ; polyploidy ; systematics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract TheAntennaria rosea polyploid agamic complex is one of the most morphologically diverse and widespread complexes of N. AmericanAntennaria. The group is taxonomically confusing because of numerous agamospermous microspecies, having been recognized as distinct species. Morphometric analyses have demonstrated that the primary source of morphological variability in the complex derives from six sexually reproducing progenitors,A. aromatica, A. corymbosa, A. media, A. microphylla, A. racemona, andA. umbrinella. Additionally, two other sexually reproducing species,A. marginata andA. rosulata, may have contributed to the genetic complexity of theA. rosea complex. Cluster analysis indicates that four discrete morphological groups exist within theA. rosea complex. Each group could be the result of predominance of genes from different groups of sexual progenitors. AsA. rosea is of multiple hybrid origin, from among several sexual progenitors, it is advisable to recognizeA. rosea as a distinct species from its sexual progenitors.
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    Patterns of intraspecific variation inAntennaria alborosea, A. corymbosa, A. marginata, A. microphylla, A. parvifolia, andA. umbrinella (Asteraceae:Inuleae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 123-150 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Inuleae ; Antennaria ; Geographic variation ; clinal variation ; principal component analysis ; correlation coefficients
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Patterns of intraspecific variation were examined inAntennaria alborosea A. E. Porsild,A. corymbosa E. Nels,A. marginata Greene,A. microphylla Rydb.,A. parvifolia Nutt., andA. umbrinella Rydb. AlthoughA. alborosea was initially considered arctic in distribution, it became apparent that a southern montane element also exists. Our results suggest that morphological differences between arctic and southern montane specimens represent clinal variation. The additional morphological data for specimens that occur more than 1,500 km south of the species' range as it was initially described result in a better understanding of this once presumed arctic taxon. Morphological variation in the dioecious speciesA. corymbosa, A. marginata, A. microphylla, A. parvifolia, andA. umbrinella was greater between the genders than was geographic variation within each gender. These results demonstrate that both pistillate and staminate specimens must be examined in dioecious species ofAntennaria if morphological variation in the respective species is to be fully understood. Character size or number of broadly distributed species (A. microphylla andA. parvifolia) generally decreased with increasing longitude, whereas characters of species with more restricted distributions (A. alborosea, A. corymbosa, andA. marginata) generally increased in size or number with increasing latitude or longitude.Antennaria umbrinella was an exception in this respect.
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    A redescription ofPotamogeton distinctus including remarks on the taxonomy of thePotamogeton nodosus group (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 245-259 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Potamogetonaceae ; Potamogeton distinctus ; P. indicus ; P. nodosus ; P. tepperi ; Taxonomy ; distribution ; description
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Potamogeton distinctus has an E. Asian distribution and is closely related toP. nodosus. P. tepperi seems to be a synonym ofP. tricarinatus, even though the name has frequently been applied to other species likeP. distinctus. P. indicus is a synonym ofP. nodosus notwithstanding the fact that not all Indian broad-leaved pondweeds belong to that taxon.
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    Systematics ofJusticia sect.Pentaloba (Acanthaceae) (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 219-235 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Acanthaceae ; Justicia ; Siphonoglossa ; Cytology ; flavonoids ; systematics ; taxonomy ; generic relationships
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The transfer of the four taxa ofSiphonoglossa sect.Pentaloba toJusticia is proposed. It is shown that the taxa of this section were placed inSiphonoglossa primarily because of a single-character phenetic relationship and that they correctly belong inJusticia. In addition to morphology, data from cytology and flavonoid chemistry are also presented that support this intergeneric transfer. A key to the taxa and a detailed taxonomic treatment of the section are provided.
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    The genusAntennaria (Asteraceae: Inuleae) in western North America: Morphometric analysis ofAntennaria alborosea, A. corymbosa, A. marginata, A. microphylla, A. parvifolia, A. rosea, andA. umbrinella (1990)
    Springer
    Plant systematics and evolution 169 (1990), S. 151-175 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Inuleae ; Antennaria alborosea ; A. corymbosa ; A. marginata ; A. microphylla ; A. parvifolia ; A. rosea ; A. umbrinella ; Taxonomy ; multivariate morphometrics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Multivariate analysis of vegetative and reproductive characters was used to examine morphological relatedness amongAntennaria alborosea A. E. Porsild,A. corymbosa E. Nels.,A. marginata Greene,A. microphylla Rydb.,A. parvifolia Nutt.,A. rosea Greene, andA. umbrinella Rydb. Both pistillate and staminate plants were examined. Some of the characters examined were variable in one species, but stable in another (i.e., presence or absence of papillae on the achenes). Our analyses indicate that the seven species are morphologically distinct. It is hypothesized that theA. rosea agamic complex arose through hybridization amongA. corymbosa, A. microphylla, A. umbrinella, and possiblyA. dioica (L.)Gaertn. However, hybridization between the three former species and others, as well as their subsequent morphological responses to different environmental conditions causes confusion in recognizing the taxa.Antennaria angustifolia Rydb.,A. arida E. Nels.,A. confinis Greene,A. scariosa E. Nels.,A. foliacea humilis Rydb.,A. concinna E. Nels., andA. viscidula E. Nels. are considered to represent F 1 hybrids.
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    Surface features of the leaves ofBalanophoraceae — A family without stomata? (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 29-35 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Balanophoraceae ; Balanophora ; Hachettea ; Langsdorffia ; Lophophytum ; Scybalium ; Thonningia ; Stomata ; epidermal surface
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Both adaxial and abaxial leaf surfaces were searched for stomata inBalanophora elongata, B. fungosa, Hachettea austro-caledonica, Langsdorffia hypogaea, Lophophytum mirabile subsp.mirabile, Scybalium jamaicense, andThonningia sanguinea (Balanophoraceae). Neither stomata nor guard cells were observed. The epidermal surfaces of these species are extremely diverse with respect to cell shape, cell size, and surface ornamentation, these features providing valuable systematic criteria.
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    Numerical analyses of karyotypic diversity in the genusEupatorium (Compositae, Eupatorieae) (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 215-228 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Compositae ; Eupatorium ; Chromosomal distance ; chromosomal identity ; karyotype analysis ; statistics phenetics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Somatic metaphase karyotypes were analyzed for 22 diploid species ofEupatorium. The karyotypic comparisons were made using two indices: minimal chromosomal distance (MCD), measuring overall dissimilarities, and chromosomal identity (CI), measuring number of morphologically identical chromosomes between species. The resulting phenograms from these indices are largely compatible. The 22 species cluster into four groups in the phenogram using MCD, and the grouping corresponds well with morphology or geographic distribution into the three N. American groupsEutrochium, Uncasia, Traganthes, and the E. Asian group. These results suggest that karyotypes in perennialEupatorium have been considerably conservative and changed not through large chromosomal mutations but through small chromosomal mutations gradually fixed.
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    Systematics and evolution within theCompositae, seen with the eyes of a chemist (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 1-14 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Compositae ; Chemistry ; systematics ; evolution
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The broad knowledge of the chemistry of theCompositae allows the discussion of its relevance for the systematics and evolution within the family. Furthermore a separation into subfamilies can be supported by the observed differences in the distribution of the main constituents in the tribes.
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    Anthecology ofSchrankia nuttallii (Mimosaceae) on the tallgrass prairie (1990)
    Springer
    Plant systematics and evolution 170 (1990), S. 247-255 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Mimosaceae ; Schrankia nuttallii ; Halictidae ; Bee-pollination ; anthecology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Schrankia nuttalii flowers through late spring on the tallgrass prairie. Although each stem produces an average of 26 capitate inflorescences only 12% of those inflorescences will open each day to disperse and receive polyads. Each inflorescence may live up to 48 hours but anthers abscise by late afternoon on the first day and the filaments change color and lose their scent. The 78–93 florets comprising each inflorescence open synchronously before dawn or during early morning hours. First day inflorescences ofS. nuttallii are herkogamous and fragrant. They are nectarless. Bombyliid flies and male bees are infrequent floral foragers so the major pollinators include female bees representing five families;Anthophoridae, Apidae, Colletidae, Halictidae, andMegachilidae. All foraging insects ignore second day inflorescences although stigmas are still receptive. Although 97% of all bees collected onS. nuttallii carrySchrankia polyads in their scopae or corbiculae 59% also carry the pollen/pollinaria of one or more coblooming angiosperms. At least 98% of all bees carrying mixed pollen loads incorporate the pollen/pollinaria of one or more nectariferous taxa (e.g.Asclepias spp.,Asteraceae, Convolvulaceae, Delphinium spec., etc.). Species of halictid bees are more likely to carry pure loads ofS. nuttallii polyads (70%) than bees of the four remaining families. Due to the nectarless florets and high degree of polylectic foraging bee-pollination inS. nuttallii converges more closely with the pollination systems of some AustralianAcacia spp. than with most other xeric/tropical genera of mimosoids studied in the western hemisphere.
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    The morphology and relationships ofMedusagyne (Medusagynaceae) (1990)
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    Plant systematics and evolution 171 (1990), S. 27-55 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Theales ; Medusagynaceae ; Medusagyne ; Systematics ; anatomy ; pollen ; morphology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The comparative vegetative and reproductive morphology and anatomy of the endangered, monotypic, dicotyledonous genusMedusagyne was studied, and detailed descriptions of leaf, axis, nodal, wood, floral, pollen, fruit, and seed structure are presented. Overall, the genus has many specialized features, including the possession of extreme, habitat-related specializations. Flowers are either bisexual or staminate, and are interpreted as retaining some primitive aspects, such as many free parts spirally arranged on an elongate floral axis. One of the most salient structural features of the plant is the massive development of ensheathing fibrous elements around the vascular system of both vegetative and reproductive tissues. Diffuse foliar sclereids are absent. Particular attention is paid to the unusual multicarpellate, synovarial gynoecium and comparisons are made with theCaryocaraceae. The totality of morphological and anatomical evidence confirms the view thatMedusagyne is a very distinct and evolutionarily isolated genus, best treated as forming the monotypic familyMedusagynaceae. Observations are presented, including the occurrence of stamen fascicle traces, that link the family to the dillenialean and thealean assemblage. Like other isolated thealean taxa,Medusagyne shows affinities to several different families, without having especially close relationships with any particular extant taxon.
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    Life cycle and adaptive strategies of three non-annual, ruderalCardueae: Onopordum nervosum, Carthamus arborescens, andCirsium scabrum (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 117-128 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Asteraceae ; Cardueae ; Onopordum nervosum ; Carthamus arborescens ; Cirsium scabrum ; Life cycle ; adaptive strategies ; autoecology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The life cycle and autoecology of three Mediterranean thorny species,Onopordum nervosum (Iberian endemic),Cirsium scabrum (Western Mediterranean) andCarthamus arborescens (Iberian-North African), of potential use in agriculture as bioenergetic, forage or oil producing plants were studied. These threeCardueae spp. are good examples of ways in which Mediterranean plants adapt their typical annual cycle to avoid summer droughts.O. nervosum is a perennial, monocarpic, heliophilous species which flowers in early summer and grows even under the extreme xeric conditions of the Mediterranean summers. It is a basiphilous plant usually found in sandy-loamy and sandy-clayey-loamy soils with a marked geographical variability in its seed germination.C. scabrum is a perennial, monocarpic species which has a long life cycle, flowering in summer and growing only during the wet months. It is not well-adapted to droughts and is usually restricted to acid soils.C. arborescens is a perennial, polycarpic species which flowers in spring and dries up in summer. It is usually found in basic loamy-sandy or sandy-loamy soils poor in organic matter.
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    Pollination ecology ofOxalis violacea (Oxalidaceae) following a controlled grass fire (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 147-155 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Oxalidaceae ; Oxalis violacea ; Andrenidae ; Bombyliidae ; Halictidae ; Megachilidae ; Pollination ecology ; heterostyly
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    Topics: Biology
    Notes: Abstract Vernal grass fires may encourage profuse flowering in clonal, colonies ofOxalis violacea. Long-styled colonies appear to be more floriferous than short-styled colonies and set a greater number of capsules. Individual flowers of both morphs live one or two days, change position on their respective pedicels and advertise nectar concealed at the base of the floral throat. AlthoughDiptera, Hymenoptera, andLepidoptera forage for nectar, bees (Andrenidae,Anthophoridae, Halictidae, andMegachilidae) probably make the only effective pollen transfers between the two morphs. Both male and female bees may transport pollen of both morphs and short-tongued bees (e.g.,Augochlorella spp.,Dialictus spp.) may be more common but as effective as pollinators as long-tongued bees (e.g.,Calliopsis andreniformis andHoplitis spp.). The conversion rate of flowers into capsules is only 13–17%. The spreading style in the short-styled morph is interpreted as an adaptation restricting insect-mediated, self-pollination but encouraging bee-stigma contact during nectar foraging.
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    Sympodial and monopodial branching inAcer (Aceraceae): Evolutionary trend and ecological implications (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 187-197 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Aceraceae ; Acer ; Sympodial and monopodial branching ; evolution ; adaptive strategy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The evolutionary trend and its ecological implications in sympodial and monopodial branching patterns has been investigated in 20 JapaneseAcer spp. through comparison of shoot tip abortion and terminal bud formation. The genus is divided into two species groups according to its branching pattern, one (6 species) predominantly exhibiting sympodial branching with frequent monopodial branching in short shoots (sympodial species), and the other (14 species) exhibiting only monopodial branching (monopodial species). The early ontogeny of leaf and bud scales is described. Despite the difference in branching patterns, the bud scales of terminal buds are essentially the same in having a leaf base developed to function as a protecting organ. In all the sympodial species, during the abortion of a sympodium shoot tip, one or two pairs of primordia were found to occur on the apex, and later wither. These primordia resemble bud scales of terminal buds in their ontogeny and morphology, and appear to be rudimentary. It is suggested that a rudimentary terminal bud develops together with the establishment of sympodial branching, and that sympodial branching has originated from monopodial branching. Based on this proposed evolutionary trend, it is suggested thatAcer has moved from less shady habitats into shady habitats with monopodial branching (advantageous for vertical growth) changing into sympodial branching (advantageous for lateral spread).
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    Crossing experiments of lettuce cultivars and species (Lactuca sect.Lactuca, Compositae) (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 233-248 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Compositae ; Lactuca sativa ; L. serriola ; L. saligna ; L. virosa ; Hybridization
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The degree of relationships withinLactuca sativa and three wild relativesL. serriola, L. saligna, andL. virosa was studied by observing the performance, vigour and fertility of the F 1 hybrids obtained from crosses made in and between the four species. The crosses ofL. saligna ×L. virosa and the reciprocal crosses produced no hybrids.L. saligna andL. virosa are the least related of the four species.L. sativa ×L. serriola and the reciprocal crosses were successful and produced fertile hybrids These two species are genetically very closely related.L. saligna is known to produce, as a female parent, hybrids withL. sativa andL. serriola. Now the reciprocal cross was successful for the first time, so the unability to obtain hybrids in the past was based on the choice of accessions and not caused by unilateral incompatibility.L. virosa ×L. sativa and the reciprocal combination produced hybrids. The combinationL. serriola ×L. virosa produced hybrids with very limited fertility. In contrast to earlier reports (sterile hybrids) one combination of the reciprocal cross too produced hybrids with very limited fertility.—Some of theL. saligna ×L. sativa (and reciprocal) hybrids were found to look strikingly likeL. serriola. This adds evidence for the descent ofL. serriola andL. sativa:L. saligna also made part of the ancestral complex of the cultivated lettuce.
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    Molecular divergence in the Galapagos Islands—Baja California species pair,Gossypium klotzschianum andG. davidsonii (Malvaceae) (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 99-115 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Malvaceae ; Gossypium ; Chloroplast DNA ; allozymes ; isozymes ; molecular evolution ; long-distance dispersal
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Molecular divergence betweenGossypium klotzschianum andG. davidsonii was studied. The former is endemic to five of the larger islands of the Galapagos, whileG. davidsonii is restricted to the southern half of Baja California, approximately 2 500 km distant. A substantial body of genetic and taxonomic data suggests that these two species are related as progenitor and derivative. Interspecific hybrids are fully fertile, with no evidence of F2 breakdown and normal segregation of genetic markers. Allozyme analysis of 33 populations for 41 loci indicated that the allelic composition ofG. klotzschianum represents a subset ofG. davidsonii. Although genetic diversity is relatively restricted in both species, calculated measures demonstrate higher levels of genetic variability and greater population structuring inG. davidsonii than inG. klotzschianum. The interspecific genetic identity of 0.87 is typical for progenitor-derivative species pairs. Chloroplast DNAs were surveyed for variation with 25 restriction enzymes using hybridization probes that cover the entire chloroplast genome. No intraspecific and little interspecific variation was detected among 560 cpDNA restriction sites, representing sequence information for approximately 3200 nucleotides. Only 3 mutational differences distinguished the two species, resulting in a sequence divergence estimate of 0.09%. Divergence times were estimated from both the isozyme data and the cpDNA restriction site data. Although these estimates have several sources of error, both molecular data sets were congruent in suggesting that the two lineages diverged between 250000 and 700000 years ago. Accumulated evidence suggests that dispersal was from Baja California to the Galapagos Islands rather than the reverse, and most likely was mediated by trans-oceanic drift.G. klotzschianum may be the only species of the endemic Galapagos flora to have arisen from a northern Mexican progenitor.
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    The systematic position of theHolarrheninae (Apocynaceae) (1990)
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    Plant systematics and evolution 171 (1990), S. 157-185 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Apocynaceae ; Plumerioideae ; Apocynoideae ; Holarrheninae ; Holarrhena ; Carruthersia ; Spirolobium ; Floral structure ; pollen morphology ; indole alkaloids ; steroidal alkaloids
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The genusHolarrhena, described byRobert Brown in 1811, has had a problematic taxonomic history, in part due to a suite of characters that does not conform with accepted concepts within theApocynaceae. In a number of important taxonomic charactersHolarrhena is typical of subfam.Apocynoideae. But due to the relatively unspecialized structure of the anthers most recent authors have placedHolarrheng, together withCarruthersia andSpirolobium, as the subtribeHolarrheninae in subfam.Plumerioideae. For the present investigation the floral structure and pollen morphology ofHolarrhena, Carruthersia andSpirolobium were analyzed. From the chemical literature reports of the occurrence of steroidal alkaloids in thePlumerioideae were evaluated. Our results indicate that the three genera belong to subfam.Apocynoideae in the tribeNerieae, but that the “Holarrheninae” is an unnatural group, and that the three genera should be accommodated individually within the tribe.
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    Ultrastructural evidence of endoplasmic reticulum changes during the maturation of the olive pollen grain (Olea europaea L., Oleaceae) (1990)
    Springer
    Plant systematics and evolution 171 (1990), S. 221-231 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Oleaceae ; Olea europaea ; Endoplasmic reticulum ; stacks cisternae ; pollen grain ultrastructure
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Characteristic features of rough endoplasmic reticulum (rER) distribution and proliferation were noted during olive pollen (Olea europaea L.) development, suggesting the physiological significance of this organelle. Initially scarce in the young microspore, ER increases as cytoplasmic vacuoles form. At the vacuolated microspore stage the cytoplasm contains numberous polysomes and elongated rER cisternae arranged preferentially in stacks, with an average intracisternal width of 0.07 µm. Stacks persist in the bicellular pollen grain but consist of fewer, shorter, dilated cisternae (mean intracisternal width 0.1 µm) containing a considerable electron-dense matrix. Cisternae in the mature grain are fragmented, leaving behind an ER of swollen pockets. Pockets of ER containing a material of greater electron density are evenly deposited along the plasmalemma, in close relation with it. A dense material is seen in the tubules of the apertural region, which was lacking in earlier stages. Our results show that ER may be involved in protein transport to the intine.
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    Pollination biology of two species ofKielmeyera (Guttiferae) from Brazilian cerrado vegetation (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 35-49 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Guttiferae ; Kielmeyera coriacea ; K. speciosa ; “Buzz pollination” ; carpenter bees ; breeding system ; andromonoecy ; self-mimicry ; neotropical savanna
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The pollination biology and breeding systems ofKielmeyera coriacea andK. speciosa, two sympatric woody species common in the cerrado vegetation of C. Brazil, were studied. Both species have similar nectarless, polystemonous “Papaver-type” flowers which are visited by a similar spectrum of insects, though they bloom in different seasons and are thus phenologically isolated. Large carpenter bees seem to be the most important pollinators and these and other bees effect “buzz pollen” retrieval despite the fact that anthers are not poricidal. Both species ofKielmeyera possess strong xenogamous breeding systems. The presence of staminate flowers and andromonoecy inK. coriacea, as well as the longevity ofK. speciosa flowers are discussed as alternative strategies to improve pollination success and reproductive efficacy.
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    Chemotaxonomic study of the genusTabernaemontana (Apocynaceae) based on their indole alkaloid content (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 13-34 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Apocynaceae ; Tabernaemontana ; Indole alkaloids ; chemotaxonomy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract According to their alkaloidal products species of the “new” genusTabernaemontana can be partly differentiated. This differentiation is in agreement with the “old” genera classification. From the chemotaxonomic point of view a subdivision of subfam.Plumerioideae of theApocynaceae is proposed.
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    A phenetic comparison of someFumaria spp. (Fumariaceae) (1990)
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    Plant systematics and evolution 172 (1990), S. 51-63 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Fumariaceae ; Papaveraceae ; Fumaria ; Phenetics ; numerical taxonomy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The taxonomy of the genusFumaria has not been considered in detail sincePugsley's work in 1919 ff., and few modern methods have been applied to it. In a phenetic study, 33 populations of 11Fumaria spp. were grown in uniform conditions, and seven morphological characters measured. After re-expression and transformation the data were analysed by cluster analysis and principal components analysis. Alternative analyses did not indicate contradictory taxonomic conclusions. Artificial crosses gave some evidence on interfertility, and suggestedF. occidentalis to be an allopolyploid ofF. bastardii ×F. capreolata. Pugsley's subsectional classification is supported within sect.Parviflora, but not within sect.Grandiflora. His two sections are seen to be meaningful, but not discrete.
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    A phylogenetic analysis of chloroplast DNA restriction site variation inPoaceae subfam.Pooideae (1990)
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    Plant systematics and evolution 172 (1990), S. 83-97 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Pooideae ; Phylogenetics ; evolution ; chloroplast DNA ; restriction site variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A phylogenetic analysis was conducted on chloroplast DNA restriction site variation in 34 genera of grasses (familyPoaceae), including 28 genera from subfam.Pooideae (representing tribesAveneae, Brachypodieae, Bromeae, Meliceae, Poeae, Stipeae, andTriticeae) and representatives of three other subfamilies,Arundinoideae, Oryzoideae, andPanicoideae. Analyses of all 34 genera always distinguishedPooideae as monophyletic, regardless of which nonpooid genus functioned as outgroup; six separate analyses of all 28 pooid genera, each including one of the six nonpooid genera as outgroup, resolved five identically-constituted clades withinPooideae (in four cases), or (in the other two cases) yielded results that were less well resolved, but not in conflict with those of the other four analyses. The four best-resolved analyses distinguishedMeliceae as the earliest diverging lineage withinPooideae, andStipeae as the next. Above the point of divergence ofStipeae is a dichotomy between supertribeTriticodae (including tribesBrachypodieae, Bromeae, andTriticeae), and a clade comprisingPoeae andAveneae. The analysis supports some tribal realignments, specifically the assignment ofBriza, Chascolytrum, Microbriza, andTorreyochloa toAveneae, andArctagrostis, Catabrosa, andSesleria toPoeae. The analysis also suggests that the pooid spikelet (i.e., glumes shorter than lemmas and florets two or more) is plesiomorphic inPooideae, and that spikelets with one floret, and those with glumes longer than the first lemma, each have evolved more than once withinPooideae. Results also indicate that small chromosomes and chromosome numbers based on x=c. 10−12 are plesiomorphic withinPooideae. Alternative states of these characters (chromosomes large, chromosome numbers based on x=7) are interpreted as synapomorphies or parallelisms of clades that includeTriticodae, Aveneae, andPoeae. Lanceolate lodicule shape may be a synapomorphy of the clade that includesStipeae, Triticodae, Aveneae, andPoeae, and loss of lodicule vascularization a synapomorphy of the entirePooideae.
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    A phylogenetic and allozyme approach to understanding rarity in three “green ash” eucalypts (Myrtaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 99-118 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Myrtaceae ; Eucalyptus ; Rarity ; allozymes ; phylogeny
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Phylogenetic relationships among the 12 species of the “green ash” group of eucalypts were examined using allozyme data, to investigate the causes of rarity in three localized endemics of the group. The relationships suggested by the allozyme data showed both similarities to and differences from those estimated from morphological data byLadiges and coworkers. The phylogenetic relationships suggest that rarity inEucalyptus burgessiana may be related to recent divergence, whileE. paliformis andE. rupicola are relatively old species, more likely to be relicts, and/or restricted to a rare habitat.
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    Self-incompatibility inCrocus vernus subsp.vernus (Iridaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 77-82 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Iridaceae ; Crocus vernus subsp.vernus ; Embryology ; mentor effect ; pollen tube ; self-incompatibility
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Outcross, self- and mixed pollinations were performed inCrocus vernus subsp.vernus, a species with bicellular pollen, dry stigmas and hollow style. No differences were noted among the above pollinations concerning the germination of pollen and the growth of pollen tubes until the top of ovary. Within 45 min after pollinations 62% of pollen grains germinated. Pollen tubes penetrated the papilla cuticle extending along the papilla wall; on entry into stigmatic lobes they continued growth in the stylar secretion to ovarian locules. Here, however, self-pollen tubes failed to reach or to enter the ovule micropyle; while pollen tubes from either outcross- or mixed pollinations grew until fertilizing ovules. These observations gave evidence of a self-incompatibility system inCrocus, which appeared to be neutralized by mentor effect. The ovary as site of incompatibility response is discussed.
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    Giemsa C-banded karyotypes of diploid and tetraploidHordeum bulbosum (Poaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 141-150 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Hordeum bulbosum ; Cytotypes ; chromosome analysis ; Giemsa C-banding patterns ; autopolyploidy ; relationships
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The similar-looking basic genomes ofHordeum bulbosum (2x and 4x) have five rather similar metacentric, one submetacentric, and one satellited choromosome. C-banding patterns are characterized by one or two centromeric, or juxtacentromeric, small to larger bands in most chromosomes, by bands at the nucleolar organizers, by small or very small telomeric bands, and by the nearly complete lack of intercalary bands. Banding pattern polymorphism is widespread. Banding patterns supported by chromosome morphology enable identification of homologues, and discrimination between non-homologues inH. bulbosum (2x). The C-banded karyotype ofH. bulbosum (4x) supports an autopolyploid origin, but it was possible to identify only homologues of submetacentrics and SAT-chromosomes.
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    Restriction fragment length polymorphisms in the genusActinidia (Actinidiaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 193-203 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Actinidiaceae ; Actinidia ; Chloroplast genome ; kiwifruit ; molecular evolution ; phylogenetic trees ; restriction fragment length polymorphism
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A series of chloroplast and nuclear probes were used to examine restriction fragment length polymorphisms (RFLPs) in kiwifruit (Actinidia deliciosa) and three of its closest relatives. The four species fell into two pairs, withA. chinensis andA. deliciosa closely related but some distance away from the other two species,A. latifolia andA. eriantha. The results are consistent with the hypothesis that the diploid species,A. chinensis, is a precursor ofA. deliciosa, which is hexaploid.
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    Ribosomal RNA genes inQuercus spp. (Fagaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 127-139 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Fagaceae ; Quercus ; Gene mapping ; ribosomal RNA genes ; rRNA/DNA hybridization ; taxonomy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The taxonomy of the genusQuercus is still unclear. In order to elucidate the taxonomy of Mediterranean oaks we have analyzed ribosomal RNA genes ofQuercus cerris, Q. coccifera, Q. trojana, Q. ilex, Q. suber, andQ. macrolepis by means of Southern blot hybridization. Oak nuclear DNA was extracted from root tips of 300 acorns and from catkins of single plants. EcoRI and BamHI restriction endonucleases were used. DNA electrophoresis and rRNA/DNA hybridization were performed usingVicia faba rRNA 18 S and 25 S as probes. The rRNA genes of all the species studied have an identical restriction mapping in the 18 S and 25 S regions, while differences in length are present in the intergenic regions.Q. cerris possesses at least four types of genes of 12.1, 11.5, 8.5, and 8.3 kb;Q. coccifera at least three types of 12.4, 10.4, and 10.1 kb;Q. trojana possesses the same rRNA genes asQ. cerris plus another gene type 12.0 kb long, with EcoRI and BamHI restriction sites in the intergenic spacer;Q. ilex at least three types of 12.4, 10.85, and 9.5 kb;Q. suber at least five types of 11.5, 11.0, 8.6, 8.5, and 8.3 kb;Q. macrolepis, finally, at least seven types of 11.5, 11.0, 10.2, 8.6, 8.5, 8.3, and 8.15 kb.Q. coccifera andQ. ilex rDNA appears quite different respect to other species examined, while high similarity seems to exist betweenQ. cerris, Q. trojana, Q. suber, andQ. macrolepis. These results are in agreement with the taxonomic model proposed bySchwarz for the genusQuercus.
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    Evolution of the tetraploidCapsella bursa-pastoris (Brassicaceae): Isoelectric focusing analysis of Rubisco (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 205-213 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Brassicaceae ; Capsella ; Ribulose-1,5-bisphosphate carboxylase/oxygenase ; isoelectric focusing ; evolution ; polyploidization
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Ribulose-1,5-bisphosphate carboxylase/oxygenase (Rubisco, EC 4.1.1.39) and its subunits (large subunits = LSU, small subunits = SSU) were isolated from threeCapsella spp. by gel electrophoresis and polypeptide composition was analyzed by isoelectric focusing (IEF) in the presence of 8M urea. The described techniques are recommended for large scale systematic studies. Multiple IEF banding patterns of the SSU are probably the outcome of a heterogenous multigene family. The two diploid speciesC. rubella andC. grandiflora show an identical IEF pattern and could be differentiated from the putative allotetraploidC. bursa-pastoris only by the SSU banding pattern. Uniqueness of some SSU bands in the tetraploid and in the two diploid species, respectively, may indicate an ancient alloploid origin of tetraploidC. bursa-pastoris followed by events leading to divergences in the genomes of the allotetraploid and its presumed diploid progenitors after the hybridization event (SSU gene elimination, acquisition of new SSU genes).
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    Heterostyly in species ofNarcissus (Amaryllidaceae) andHugonia (Linaceae) and other disputed cases (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 215-227 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Amaryllidaceae ; Narcissus ; Linaceae ; Hugonia ; Bauhinia ; Cleome ; Aneilema ; Agelaea ; Heterostyly ; tristyly ; distyly
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The hypothesis ofHenriques andFernandes that several Iberian species ofNarcissus (Amaryllidaceae) are tristylous is reconsidered. Contrary to the opinion ofBateman and most subsequent authors, we believe that the available evidence indicates that some populations ofN. triandrus andN. fernandesii, at least, are tristylous; other populations ofN. triandrus are distylous.Hugonia cf.penicillanthemum (Linaceae) from new Caledonia is distylous, but it remains possible that other species ofHugonia are tristylous. The disputed occurrence of heterostyly in S. African species ofBauhinia (Leguminosae),Cleome (Capparaceae) andAneilema (Commelinaceae), and inAgelaea (Connaraceae) is discussed.
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    Rumex subgen.Rumex sect.Axillares (Polygonaceae) in South America (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 151-192 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Polygonaceae ; Rumex subg.Rumex sect.Axillares ; Taxonomy ; systematics ; hybridization ; Flora of S. America
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resumen Rumex secciónAxillares fue descrito por primera vez por el autor en 1937. Se conoce en América del Norte con 20 especies endémicas. De América del Sur se conocían sólo 5 especies, todas ellas endemicas; este número asciende a 15 en la presente revision. Fuera de las Américas se encuentran pequeños grupos de especies endémicas en Sudáfrica, en las Islas de Hawaii y en Australia; especies aisladas se encuentran en Africa Central, en las lejanas Islas Atlánticas de Tristan de Cunha y Gough Island y en Asia Oriental. La sección no tiene representantes en Europa, con excepción de tres especies americanas introducidas, poco frecuentes. La secciónAxillares se caracteriza bien, tanto morfológicamente por su forma de creciemiento—sin rosetas foliares, período floral prácticamente ilimitado por la formación de inflorescencias axilares parciales, flores a menudo monoicas—como por su estructura genética que se manifiesta en un comportamiento diferente de hibridación. Híbridos son frecuentes en las especies de la secciónRumex (syn.Simplices) y reconocibles por su esterilidad que se expresa en cierta particularidad del habito. Al contrario, no hay seguridad de híbridos espontáneos en la secciónAxillares. La mayor parte de los híbridos artificiales son totalmente fértiles. Hasta ahora se conocían solo tres híbridos entre las especies de ambas secciones. Otros tres han sido encontrados por el autor en Chile. Todos completamente estériles y con las características de hábito típicas resultantes la esterilidad, como ocurre en los híbridos de la secciónRumex.
    Notes: Abstract Rumex sect.Axillares first described by the author in 1937 is known to be represented in N. America by 20 endemic species. From S. America only five species, all endemic, were previously known; this number is raised in the present revision to 15. Outside America there exist small groups of endemic species in S. Africa and in the islands of Hawaii as well as in Australia and scattered single species in C. Africa, on the remote Atlantic islands of Tristan de Cunha and Gough Island, and in eastern Asia.—In Europe the section is completely absent, except for three American species which occur as rare aliens. TheAxillares are not only morphologically well-characterized by their growth—no leaf rosettes, flowering period practically unlimited by formation of later flowering axillary shoots, flowers often monoecious—but also by their genetic structure expressed by a different behaviour in hybridization. Hybrids are frequent between species of sect.Rumex (syn.Simplices) and are easy to recognize by their sterility which results in a certain peculiarity in habit. In contrast, no spontaneous hybrids are known with certainty among species of sect.Axillares. Most artificial hybrids are fairly fertile. So far only three hybrids between species of the two sections have been known. Three more have been found by the author in Chile, all being highly sterile and showing the growth characteristics resulting from sterility as occur in hybrids between species of sect.Rumex.
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    Robertsonian change in allies ofZebrina (Commelinaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 263-271 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Commelinaceae ; Tradescantia soconuscana ; Zebrina ; Chromosome evolution ; Robertsonian change
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Meiotic pairing inTradescantia soconuscana 2n=26 (6M+16A+4T) suggests that it has a tetraploid constitution which is not apparent in its chromosome number. Its “nombre fondamental” of 32 indicates that it could have evolved from an ancestor with x=8 by a combination of Robertsonian fusion, hybridization and polyploidy. The 2n=16 (8A+8T) karyotype of a closely related diploid supports this. The allied genusZebrina may have followed a similar method of chromosome evolution.
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    Aspects of floral structure and phenology in the genusConophytum (Mesembryanthemaceae) (1990)
    Springer
    Plant systematics and evolution 172 (1990), S. 229-240 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Mesembryanthemaceae ; Conophytum ; Floral morphology ; floral phenology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The complex floral structure in the southern African genusConophytum (Mesembryanthemaceae; 77 spp.) is described in detail and assigned to three basic floral types, two of which can be divided into two subtypes. Correlations between structural features and phenological patterns are demonstrated and discussed in the family context as well as in relation to the systematic subdivision of the genus.
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    The chemotaxonomy ofGeranium (Geraniaceae) (1990)
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    Plant systematics and evolution 173 (1990), S. 1-15 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Geraniaceae ; Geranium ; Chemotaxonomy ; seed proteins ; essential oils ; nectar amino acids ; additivity in hybrids and allopolyploids ; chemical divergence and mutation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Relationships amongGeranium species constituting sectt.Anemonifolia, Lucida (monotypic) andRuberta together with representatives of sect.Unguiculata were investigated by gas-chromatographic study of essential oils, electrophoretic comparison of seed proteins and chromatographic separation of nectar amino acids. — Essential oil study gave little information.G. macrorrhizum (sect.Unguiculata) had far greater quantities of essential oils in its foliage than other species and differed from them qualitatively. — Species of sectt.Anemonifolia andRuberta, together withG. cataractarum (sect.Unguiculata), between them yielded 19 seed protein bands; the distribution of these indicated close relationship among the species and was consistent with hypotheses for the origin of certain species by alloand autopolyploidy partly within the group. Involvement of an unknown species in the origin of two allopolyploids was implied. Separate origins for the two octoploid species in this set are also inferred. In two instances there was evidence for the transformation of one band into another subsequent to the separation of related species. The inference of allopolyploidy was supported by the occurrence of additivity of parental bands shown by an artificial hybrid between two of the species. A model for the evolutionary divergence of the seed protein patterns is presented. Two species outside the above set appeared less closely related; they wereG. lucidum andG. macrorrhizum (sect.Unguiculata) and between them they showed 6 additional bands, four of which were shared. — Of 18 nectar amino acids found, 4 to 15 occurred in any one species, with low numbers (4 and 8) occurring in the most extreme inbreeders. The spectra of nectar amino acids of two hybrids showed additivity of those of the respective parents. The results echo rather closely those provided by seed proteins, but in the absence of data from outside the group their taxonomic significance is uncertain. However, the divergence between the two octoploids was again evident.
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    DNA “fingerprints” detect genetic variation inAcer negundo (Aceraceae) (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 49-56 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Aceraceae ; Acer negundo ; box elder ; RFLPs ; minisatellite DNA ; M13 probe ; DNA analysis ; genetic variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Genomic DNA samples from 21 box elder plants collected in Missouri (U.S.A.) were digested with restriction enzyme and southern blot hybridized with the M13 minisatellite probe. Each plant was found to have a unique DNA fragment pattern. Moreover, levels of genetic variation estimated from a similarity index appear to be related to sampling distances. However, size of the fragments utilized in the analysis affects the estimates of genetic variation to a considerable degree.
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    Prolamins in systematics ofPoaceae subfam.Arundinoideae (1990)
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    Plant systematics and evolution 173 (1990), S. 57-70 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Arundinoideae ; Protein ; prolamin ; immunology ; immunoblotting ; electrophoresis ; numerical systematics ; phylogeny
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Circumscription of theArundinoideae and the taxonomic treatments of its tribes have long been disputed. In the present study, 21 species representing the tribesArundineae, Danthonieae, Cortaderieae, andAristideae were examined for prolamin size heterogeneity and immunological cross-reactivities. The immunmological data were analyzed phenetically and cladistically. Prolamins of species examined were similar in size and size diversity except forPhragmites. Structural similarities as measured by ELISA and immunoblotting were very high among all genera exceptAristida andPhragmites. Based on prolamin structure,Aristida is not similar to the core genera of theArundinoideae, the remaining genera could not be distinguished as distinct tribes, and theArundinoideae as represented by the taxa tested appeared monophyletic.
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    A numerical taxonomic study of thePolygonum aviculare complex (Polygonaceae) in Belgium (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 71-89 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Polygonaceae ; Polygonum aviculare ; Numerical taxonomy ; weeds ; ruderal strategists ; population variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract 16 morphological characters were assessed in 300 plants sampled from 27 populations ofPolygonum aviculare sensu lato representative of the whole ecological range of the species in Belgium. The three multivariate treatments performed, namely principal component analysis, cluster analysis and discriminant analysis provide evidence thatP. aviculare can be divided in 4 units, roughly corresponding to the four taxa recognized byChrtek. However, the results point out that subsp.monspeliense and subsp.calcatum should be included at the varietal rank within respectively subsp.aviculare (=P. heterophyllum) and subsp.aequale (=P. arenastrum) as defined byLindman. Fruit dimensions, fruit shape and tepal length are the most discriminant characters for separating the four taxa, while several other characters are discriminant at the population level only. The evolutionary significance of the variation pattern of the whole complex is discussed in terms of life history differentiation and ecological specialization.
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    Flavonoid aglycones from the leaf surfaces of someLabiatae species (1990)
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    Plant systematics and evolution 173 (1990), S. 109-118 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Labiatae ; Exudate flavonoids ; ecological biochemistry ; biochemical systematics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The distribution of excreted flavonoid aglycones within the familyLabiatae was studied and differences were found, especially in the A-ring substitution patterns. Thus, 5,7-dihydroxy-6-methoxyflavones with substituted B-rings are characteristic of species ofSalvia (sect.Salvia),Rosmarinus andOcimum; 5,7-dihydroxy-6,8-dimethoxyflavones occur only inOcimum and 5,6-dihydroxy-7,8-dimethoxyflavones inThymus and related species. Members of the two subfamiliesLamioideae andNepetoideae produce exudate flavonoids, but some genera are devoid of these compounds. There is a correlation between the habitat where the plant grows and production of these compounds, the species from (semi-)arid habitats being those which generally accumulate external flavonoids.
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    Studies on theStellaria longipes complex (Caryophyllaceae): Isozyme variability and the relationship betweenStellaria longipes andS. longifolia (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 129-141 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Caryophyllaceae ; Stellaria longipes ; S. longifolia ; Isozyme variability ; allozyme analysis ; allopolyploidy ; population differentiation ; genetic variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Genetic variation within and the relationship betweenStellaria longipes Goldie andS. longifolia Muhl. were studied. Ten enzyme systems were assessed in eight natural populations ofS. longipes (25 loci) and three ofS. longifolia (20 loci) using starch and polyacrylamide gel electrophoresis. Patterns of population differentiation corresponded to geographic distance. There was no evidence that polyploidS. longipes had greater electrophoretic variability than diploidS. longipes. The isozyme data confirmed extensive population differentiation in these species and, within that context, a relatively close relationship betweenS. longipes andS. longifolia. It was postulated that diploids of these two species might be the progenitors of tetraploidS. longipes.
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    Pollination of New CaledonianWinteraceae: Opportunistic shifts or parallel radiation with their pollinators? (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 143-157 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Winteraceae ; Zygogynum ; Lepidoptera ; Micropterigidae ; Sabatinca ; Coleoptera ; Pollination biology ; coevolution ; floral volatiles ; parallel radiation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Flower visitors on 12 species of New CaledonianWinteraceae were studied. The visitors were two species of ancestral moths (Sabatinca; Micropterigidae), three species of weevils (Palontus; Curculionidae), and a species of thrips. Behavior observations and pollen records suggest that the beetles and occasionally the moths serve as pollinators ofZygogynum and Exospermum, andBelliolum is pollinated primarily by thrips. The floral volatiles are simple in composition, usually dominated by short esters. Preliminary experiments showed that ethyl acetate elicited alighting, and a distinctive huddling behavior was elicited by artificially mixed fragrance. The host associations of otherSabatinca andPalontus spp. do not support the hypothesis that theWinteraceae have radiated in association with their pollinators. The available evidence supports the notion of opportunistic isolated host colonizations at some point in the radiation of the pollinator groups.
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    Nuclear DNA variability withinPisum sativum (Leguminosae): Cytophotometric analyses (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 179-185 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Leguminosae ; Pisum sativum ; Cytophotometry ; nuclear DNA content ; DNA variability
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The cultivars or experimental lines ofPisum sativum were analyzed cytophotometrically for nuclear DNA content of early prophases after ethanol-acetic acid fixation. Wide variability was found (from 3.93 to 5.07pg per haploid nucleus). This result was confirmed by the cytophotometric analysis of interphase nuclei isolated from leaf tissues fixed in formalin. Analysis of interphase nuclei at different thresholds of optical density showed that certain chromatin fractions are involved in the variations.
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    The pollen morphology ofOdontites (Scrophulariaceae) and its taxonomic significance (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 159-178 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Scrophulariaceae ; Odontites ; Odontitella ; Macrosyringion ; Bornmuellerantha ; Omphalothrix ; Euphrasia ; Bartsia ; Parentucellia ; Pollen morphology ; exine sculpturing
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The pollen morphology of 29 species ofOdontites and related genera was examined by light and scanning electron microscopy. Three major pollen types differing fundamentally in exine sculpturing were found. In the parasitic tribePedicularieae retipilate sculpturing is the most widespread type representing a plesiomorphic character state from which the two other types are derived. Of these, reticulate sculpturing is confined toOdontites, whereas a complex retirugulate pattern was found only in the monotypic Near East genusBornmuellerantha. Within the retipilate and the reticulate major types eight minor types were distinguished based on the differential correlation of exine surface morphology, size, shape and amb form. The pollen data are generally well correlated with macromorphological features and furnish important taxonomic characters at the genus—as well as at the specieslevel.
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    Ribosomal RNA genes structure in somePopulus spp. (Salicaceae) and their hybrids (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 187-196 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Salicaceae ; Populus ; Ribosomal RNA genes ; restriction enzyme maps ; rDNA
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The tandemly repeated multigene families encoding 18S and 25S rRNAs were studied at the restriction enzyme level inPopulus alba L.,Populus deltoides Bartr. exMarsh.,Populus trichocarpa Torr. & Gray and in the hybrids between the last two mentioned species. The analysis of single and double digestion with EcoRI, BamHI, XbaI, and SstI endonucleases showed the presence of single repetitive unit types of 12.25 and 11.75kb inP. alba andP. trichocarpa, respectively.P. deltoides showed two rDNA gene types having the same length (12.25Kb) but different nucleotide sequence in the IGS. The rDNAs genes ofP. deltoides andP. triochocarpa are inherited codominantly in their hybrids.
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    Morphological and anatomical differentiation ofVellozia hirsuta populations (Velloziaceae) (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 197-208 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Velloziaceae ; Vellozia hirsuta ; Morphological and anatomical variation ; geographical differentiation ; taxonomy ; Flora of the campos rupestres ; Brazil
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The morphological and leaf anatomical differentiation ofVellozia hirsuta is analysed and classified into several types (A1, A2, A4, B3, B5, C3). The species has a relatively wide distribution in the campos rupestres of Minas Gerais in Brazil. The variation of the isolated populations on different mountain ranges is complex, does not follow a clear geographical pattern, and defies taxonomic classification.
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    The production of floral oils byMonttea (Scrophulariaceae) and the function of tarsal pads inCentris bees (1990)
    Springer
    Plant systematics and evolution 173 (1990), S. 209-222 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Scrophulariaceae ; Monttea ; Hymenoptera ; Anthophoridae ; Centris ; Oil-flowers ; oil-bees ; Flora of Argentina
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Plant species that secrete oil as their primary floral reward are rare and sporadically found in the angiosperms. We report here thatMonttea, a genus previously unsuspected of being an oil-plant, produces lipids from trichome elaiophores on the inside of the lower (anterior) lip. The discovery of the production of oils by species of this S. American genus explains the occurrence of unusual dual-function collecting structures in ArgentineCentris (Hymenoptera: Anthophoridae) and explains the presence of oil-collecting bees in regions where oil-secreting flowers were previously thought to be absent. The behavior of these centridine pollinators onMonttea flowers parallels that of oil-collecting bees onDiascia (Scrophulariaceae) in S. Africa.
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    Globba unifolia andG. corneri stat. et nom. nov. (Zingiberaceae; Peninsular Malaysia) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 1-4 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Zingiberaceae ; Globba ; Taxonomy ; Flora of Malesia and Malaysia
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Based on personal collections ofGlobba unifolia Ridl. andG. unifolia var.sessiliflora Holtt., the latter is recognized as a distinct species. As the nameG. sessiliflora is already occupied, the nomen novumG. corneri is proposed in honour of Prof. DrE. J. H. Corner, who was the first collector of the plant.
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    The effect of crossing distances and population subdivision on floral morphology inScleranthus annuus (Caryophyllaceae), a selfing annual (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 5-16 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Caryophyllaceae ; Scleranthus ; Floral morphology ; within-population crosses
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract I used a discontinuous population ofScleranthus annuus (Caryophyllaceae) to study the effect of crossing distances on flower morphology of the progeny. Four types of progeny were produced by artificial selfing, crossing with pollen-donors from the same patch in the population, crossing with pollen from donors from other patches in the population and inter-population crosses. The size of gynoecium parts and 12 sepal characters in this petal-lacking species were significantly influenced by the type of cross and the patch in the population from where the seed-parents originated. All comparisons of progeny types except selfed vs progeny produced by within-patch crosses were significantly separated from each other, while all four seed-parent patches used were significantly separated in a multidimensional space.
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    Pollination biology ofNierembergia (Solanaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 17-35 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Solanaceae ; Nierembergia ; Tapinotaspis ; Centris ; Lanthanomelissa ; Paratetrapedia ; Pollination biology ; oil-flowers
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A morphological and anatomical description of the floral syndrome ofNierembergia is given, including a survey of the presence of elaiophores in most of its species, chromatography of the floral oils and observations on the pollinator behaviour. The principal pollinators areTapinotaspis spp. includingT. chalybaea which is endowed with oil collectors on the middle legs. Other exomalopsines (Lanthanomelissa andParatetrapedia) and centridines (Centris) also visitNierembergia for pollen and oil. Based on these observations and on a morphological study of the fertile parts, different pollination modes are recognized and the biogeography and evolution of the genus is discussed.
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    Medicago truncatula f.laxicycla (Leguminosae), a new taxon with loose fruit coiling promoting rapid germination of the fruit-retained seeds (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 37-46 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Leguminosae ; Medicago truncatula f.laxicycla ; M. laxispira ; Evolution ; adaptation ; seed germination ; chromosome number ; Flora of Libya
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Medicago truncatula is a member ofMedicago sect.Spirocarpos, in which all 34 species have coiled fruit, usually tightly coiled, that permanently retain the seeds. A variant ofM. truncatula from Libya is unique for the species in possessing very loosely coiled fruits, and is here recognized as a new forma,M. truncatula f.laxicycla E. Small. In germination tests of a number of species ofMedicago, the unusual Libyan variant proved to be the only one in which rapid and simultaneous germination of most of the seeds occurred while still within a fruit. The loose coiling of the fruit of the Libyan variety appears to facilitate water entry and seedling escape. Its rapid germination might adapt it to respond quickly to intermittently available moisture, but at the cost of abandoning the strategy of intermittent germination adopted by other populations ofMedicago sect.Spirocarpos.
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    Subgroups in theHordeum patagonicum complex (Poaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 47-66 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Hordeum patagonicum complex ; barley ; Classification ; numerical methods ; evolution
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Seeds of theHordeum patagonicum complex were collected from the field and grown in the greenhouse. The aim was to take a sample of members of the complex, and on the basis of the phenotypic similarities in some morphological and physiological characters, determine whether distinct groups exist. When cluster analyses, to generate hypotheses, and orthodox statistical procedures, for hypotheses obtained a priori, were applied to the reproductive morphology, germination and flowering patterns, onlyH. patagonicum subsp.magellanicum, out of the five recognized taxa, could be distinguished consistently. The remaining four taxa, which overlapped considerably, could be re-formed into three groups whose centroids were different but whose ranges of variation were not distinct from each other. We conclude that the highly cross-compatible members of theH. patagonicum complex, first defined as species and later redefined as subspecies are probably no more than biotypes.
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    Genome analysis ofElymus angulatus andE. patagonicus (Poaceae) and their hybrids with N. and S. AmericanHordeum spp. (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 75-82 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Triticeae ; Elymus ; Hordeum ; Intergeneric hybrids ; genome analysis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The results of genome analysis of five hybrids, viz.Elymus patagonicus ×Hordeum procerum, E. patagonicus ×H. tetraploidum, E. angulatus ×H. jubatum, E. angulatus ×H. lechleri, andE. angulatus ×H. parodii, are reported. The genomic constitution ofHordeum tetraploidum andH. jubatum is best given as H1H1H2H2, ofH. lechleri andH. parodii as H1H1H2H2H4H4, ofH. procerum as H1H1H2H2H3H3, and ofElymus patagonicus andE. angulatus as SSH1H1H2H2.
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    Electrophoretic study and genetic affinity in theCampanula elatines andC. fragilis (Campanulaceae) rock-plants group from Italy and W. Jugoslavia (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 67-73 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Campanulaceae ; Campanula isophylla ; C. elatines ; C. elatinoides ; C. fragilis subsp.fragilis and subsp.cavolinii ; C. garganica ; C. fenestrellata subsp.istriaca ; C. portenschlagiana ; Allozyme variation ; heterozygosity ; Flora of Italy and W. Jugoslavia
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Variability and genetic divergence of 11 field populations of seven species of theCampanula elatines andC. fragilis rock-plants group from the Mediterranean and pre-Alpine areas have been studied by starch-gel electrophoretic techniques.Campanula isophylla, C. elatines, C. elatinoides, C. fragilis subsp.fragilis and subs.cavolinii, C. garganica were collected in Italy, whilstC. fenestrellata subsp.istriaca andC. portenschlagiana came from W. Jugoslavia. Twelve enzymatic loci for each population were genetically analysed: PGI-1 and 2, PGM-2, IDH-1 and -2, SKDH, ME, ADH, GOT-2, MPI-1 and -2, SOD-1. The genetic distances among the above mentioned entities have been calculated by Nei's index and depicted in a dendrogram.
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    Morphology and distribution of trichomes in Italian species ofTeucrium sect.Chamaedrys (Labiatae) — a taxonomical evaluation (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 83-91 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Labiatae ; Teucrium sect.Chamaedrys ; Taxonomy ; trichomes ; Flora of Italy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Micromorphology and distribution of trichomes (both secretory and not) in different parts of plants of the five Italian species belonging toTeucrium sect.Chamaedrys have been studied. These characters neatly evidence the differences between the taxa considered; of particular interest are the observations concerning the dense coverage of peltate hairs on the abaxial face of the leaves ofT. flavum subsp.glaucum and the characters contributing to a better identification ofT. marum andT. subspinosum. The importance of the calyx indumentum as distinctive taxonomical character of the genusTeucrium is confirmed, stressing at the same time the necessity of completing the observations at micromorphological level.
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    Pollinator limitation and position dependent fruit set in the high Andean orchidMyrosmodes cochleare (Orchidaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 93-101 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Orchidaceae ; Myrosmodes cochleare ; Pollination ; breeding system ; Andes of Venezuela
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Myrosmodes cochleare is a terrestrial orchid restricted to high elevations of the northern Andes. It is self-compatible but not autogamous. Flowering begins at the apex of the inflorescence, but flower size increases steadily towards the base. Fruit set of selfed flowers was found to increase significantly from the apex to the base of the inflorescence. Open-pollinated fruit set was significantly lower than hand-pollinated fruit set overall, but the highest levels occurred in the middle of the inflorescence. Eumenid wasps and a calliphorid fly were observed to pollinate the flowers, but visits were rare and occurred only during infrequent warm periods.
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    Kengyilia habahenensis (Poaceae: Triticeae) — a new species from the Altai mountains, China (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 103-108 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Poaceae ; Triticeae ; Kengyilia habahenensis ; spec. nova ; Taxonomy ; genome analysis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Kengyilia habahenensis, spec. nova, from the Altai mountains, China, is described morphologically and cytologically. It has 2n = 42 chromosomes, and the genome formula PYS.
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    Phytochemical variation inCondalia microphylla (Rhamnaceae) (1991)
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    Plant systematics and evolution 174 (1991), S. 109-114 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Rhamnaceae ; Condalia ; Lipid analysis ; seed protein electrophoresis ; chemical variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Seed oil composition (sterol esters and sterols) and seed protein profiles ofCondalia microphylla were investigated. The chemical differences observed may be attributed to genotypic changes and could support the existence of infraspecific taxa.
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    Karyological differentiation and speciation in C. MediterraneanAnacamptis (Orchidaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 115-123 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Orchidaceae ; Anacamptis pyramidalis ; A. urvilleana ; Polyploidy ; isolation mechanisms ; speciation ; Flora of the Mediterranean ; Malta
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Anacamptis pyramidalis is a variable and wide-spread European-Mediterranean taxon. Beside a dominant cytotype with 2n = 36 it includes cytotypes with 2n = 54 and 63 in northern Tuscany (and the Eastern Pyrenees) and one with 2n = 72 on Malta. In contrast,A. urvilleana, formerly often misidentified and included inA. pyramidalis, is a monomorphic and distinct species, endemic to the Maltese Islands. It has 2n = 36, can be clearly separated by morphological and anatomical features and is isolated from partly sympatric populations ofA. pyramidalis with 2n = 72 by differences in chromosome number, flowering time and habitat preference.
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    Chromosomal C-band variation inAllium schœnoprasum (Liliaceae) in eastern North America (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 125-137 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Liliaceae ; Allium schœnoprasum ; C-banding ; karyotype polymorphism ; geographical variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Variation in C-banding was studied in seven populations ofAllium schœnoprasum from eastern N. America, including populations referable to var.sibiricum, var.laurentianum, and ± intermediate. 23 bands were recognized on five pairs of chromosomes, and were treated as 23 loci. No banding site was monomorphic throughout the plants studied. The level of polymorphism per population was 〉60%, and the average heterozygosity values varied from 0.21 to 0.27. The various “banding patterns” of chromosomes were shown to depend on the random combination of individual bands. Nei's genetic distances between populations varied from 0 to 0.070 (mean: 0.033). The matrix of genetic distances was analysed by non-metric multidimensional scaling, and the results showed a significant relationship between longitude and population scores on the ordination. The chromosomal data did not clearly discriminate between the two native varieties ofA. schœnoprasum, but were interpreted as a longitudinal cline. It is suggested that studies of C-banding variation in vascular plants should focus on individual banding sites, rather than on whole chromosome banding patterns.
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    Aestival and autumnal vicariads ofGentianella (Gentianaceae): A myth? (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 139-158 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Gentianaceae ; Gentianella germanica ; Gentianella campestris ; Seasonal dimorphism ; seasonal differentiation ; seasonal ecotypes ; flowering time ; aestival and autumnal vicariads ; grazing
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Plant samples were collected from 13 populations of the two speciesGentianella germanica andG. campestris. In addition, two samples from populations of grazed plants lacking their main stems were collected. A number of relevant morphological characters were measured, the flowering time, and the habitats of each population were studied. Some ecotypes ofG. germanica with clear differences in flowering time were recognized. Differences in flowering time and morphology between populations ofG. campestris seem to be clinal. The generally accepted theory of seasonal dimorphism with the assumption of separate aestival and autumnal vicariads proves to be inadequate. A hypothesis based upon the effect of human influences on the origin of this ecotypes is suggested.
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    An analysis of the B genome speciesArachis batizocoi (Fabaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 159-169 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Fabaceae ; Arachis batizocoi ; A. hypogaea ; Genome analysis ; phylogenetic relationships
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Arachis batizocoi Krap. & Greg. is a suggested B genome donor to the cultivated peanut,A. hypogaea L. Until recently, only one accession of this species was available in U.S.A. germplasm collections for analyses and species variability had not been documented. The objective of this study was to determine the intraspecific variability ofA. batizocoi to better understand phylogenetic relationships in sect.Arachis. Five accessions of the species were used for morphological and cytological studies and then F1 intraspecific hybrids analyzed. Some variation was observed among accessions—for example, differences in seed size, plant height and branch length. The somatic chromosomes of accessions 9484, 30079, and 30082 were nearly identical, whereas, the karyotypes of accessions 30081 and 30097 have several distinct differences. For example, 30081 had significantly more asymmetrical chromosomes 2 and 6 and more median chromosomes 7 and 10, and 30097 had significantly more asymmetrical chromosomes 3 and 10 and more median chromosomes 1 and 5 than accessions 9484, 30079, and 30082. All F1 hybrids among accessions were highly fertile. Meiotic observations indicated that hybrids among accessions 9484, 30079, or 30082 had mostly bivalents. However, quadrivalents were observed when either 30081 or 30097 was crossed with the above three accessions and 30081 × 30097 had quadrivalents, hexavalents and octavalents. The presence of translocations is the most likely cause of multivalent formation inA. batizocoi hybrids. Cytological evolution via translocations has apparently been an important mechanism for differentiation in the species.
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    Sex ratio variation and spatial segregation of the sexes in populations ofRumex acetosa andR. acetosella (Polygonaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 183-195 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Polygonaceae ; Rumex acetosa ; R. acetosella ; Sex ratios ; spatial segregation of the sexes ; environmental correlations ; density effects
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Populations ofRumex acetosa andR. acetosella were studied during two growth seasons. The ramet sex ratios ofR. acetosa were always female-biased. InR. acetosella the sex ratios expressed more variation but were mostly female-biased. In both species the sex ratios commonly varied between subpopulations reflecting a partial spatial segregation of the sexes. No marked differences between sexes in vegetative vigour were detected in either species. Interactions between sex ratios, various soil characteristics and population densities were determined. Possible mechanisms for causing biased sex ratios and partial spatial segregation of the sexes are discussed.
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    New taxa ofRubiaceae-Rubieae from Iran (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 197-211 
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    ISSN: 1615-6110
    Keywords: Angiosperms ; Rubiaceae ; Rubieae ; Asperula sect.Oppositifoliae ; sect. nova ; spec. novae:A. rezaiyensis ; A. rechingeri ; Galium spec. novae:G. anguineum ; G. azerbayjanicum ; G. plurifolium ; G. pseudomegalanthum ; G. kandavanense ; G. wendelboi ; G. sojakii ; G. subvelutinum var.oligotrichum ; var. nova ; G. decumbens ; G. problematicum ; stat. nov ; Flora of Iran
    Source: Springer Online Journal Archives 1860-2000
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    Notes: Abstract Asperula sect.Oppositifoliae is validated. 2 new species ofAsperula and 7 new species and 1 variety ofGalium from Iran are described, discussed in respect to their affinities and illustrated. 2 subspecies are raised to species rank.
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  • 97
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    Leaf anatomy inTigridieae (Iridaceae) (1991)
    Springer
    Plant systematics and evolution 175 (1991), S. 1-10 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Iridaceae ; Tigridieae ; Leaf anatomy ; systematics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Leaf anatomy is described in 32 species of 16 genera of the New World tribeTigridieae (Iridaceae, Iridoideae), including examples from both subtribesCipurinae andTigridiinae. Both plicate (pleated) and foliated (with flanges, or keel extensions) leaf types occur in both subtribes. However, amongTigridiinae, foliated leaves are restricted toTigridia and differ from the foliated leaves ofCipurinae in that they lack subepidermal marginal sclerenchyma, which is present in all the foliated species ofCipurinae. This indicates that the foliated leaf type has evolved at least twice in the tribe, since the plicate leaf, which most closely resembles the equitant leaf typical ofIridaceae, is probably plesiomorphic.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00942141
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  • 98
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    An embryological study ofPlatanthera bifolia (Orchidaceae) (1991)
    Springer
    Plant systematics and evolution 174 (1991), S. 213-220 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Orchidaceae ; Platanthera bifolia ; Embryology ; embryo sac ; megasporogenesis ; megagametogenesis ; confocal scanning laser microscopy (CSLM)
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Flowers ofPlatanthera bifolia were hand-pollinated and fixed in FPA50 after 2, 5, 7, 14, and 21 days. Ovules, made transparent in Herr's clearing fluid, were investigated using confocal scanning laser microscopy. Pollination initiates the megasporogenesis. Two days after pollination dyads are frequent. Three days later most embryo sacs contain two nuclei. Seven days after pollination the embryo sacs are 4–8-nucleate and some are organized, and a week later all embryo sacs are organized and fertilization takes place. The embryo sac development follows thePolygonum type. Twenty-one days after pollination the egg nuclei have been fertilized and the embryo sacs contain 2- to many-celled embryos. A suspensor is formed during early stages of embryo development but degenerates later. Fertilization of the central nucleus does not lead to endosperm development.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00940341
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  • 99
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    Allozyme variation and phylogeny in annual species ofCicer (Leguminosae) (1991)
    Springer
    Plant systematics and evolution 175 (1991), S. 11-21 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Leguminosae ; Cicer ; Allozymes ; isozyme gene duplication ; phylogenetic relations ; polymorphism
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Allozymic variation at 30 isozyme loci was examined electrophoretically in nine annual and one perennial species ofCicer. While most of the accessions examined were monomorphic, species can be differentiated on the basis of their enzyme phenotypes. Several groups of species were identified based upon genetic distance values. For example,C. arietinum, C. reticulatum, andC. echinospermum shared the same alleles for most of the loci exmained. PerennialC. anatolicum is also closely related to this group. Similarly,C. judaicum, C. bijugum, andC. pinnatifidum formed another group. Two annual species,C. chorassanicum andC. yamashitae clustered together, whereasC. cuneatum was the most distantly related species. Correlations were found between genetic distances and geographic distribution. Results from enzyme electrophoresis tend to support the previously reported taxonomic treatments based upon crossability and morphological similarity. However,C. yamashitae, which has been classified in the second crossability group, is quite distinct genetically and morphologically from the remaining species of the group. An isozyme gene duplication observed in the genus suggested the monophyletic origin of the species examined in the present study.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00942142
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  • 100
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    Restriction data from chloroplast DNA for phylogenetic reconstruction: Is there only one accurate way of scoring? (1991)
    Springer
    Plant systematics and evolution 175 (1991), S. 39-54 
    Details
    ISSN: 1615-6110
    Keywords: Angiosperms ; Rubiaceae ; Chloroplast DNA ; fragment analysis ; restriction analysis ; site analysis ; Wagner parsimony
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Information from the same restriction analysis of chloroplast DNA of 33 taxa ofRubiaceae was scored in four different ways, two of which were based on fragments, and two on restriction sites, and they were subsequently analysed with Wagner parsimony. The methods resulted in different phylogenetic trees. The inherent differences between the methods relate to the amount of non-homologous characters and dependent characters, but none of the methods will systematically bias the resulting cladograms. The fragment analyses are much less time-consuming, but probably less accurate, than the site analyses. The choice of method is dependent on a trade-off between accuracy and resources (time). One important recommendation is made: all phylogenetic analyses of chloroplast DNA data should be accompanied by a data matrix and contain information on how the matrix was compiled.
    Type of Medium: Electronic Resource
    URL: http://dx.doi.org/10.1007/BF00942144
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