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  • 1
    Electronic Resource
    Electronic Resource
    Springer
    Pure and applied geophysics 140 (1993), S. 1-14 
    ISSN: 1420-9136
    Keywords: Crustal phases ; earthquake locations ; re-identification ; first arrival ; upper mantle ; granitic layer
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract For crustal earthquakes of moderate or large size, the first reported phases at distant stations are usually the first predicted by the crustal model used. For smaller events, however, or for stations at larger distances, the first phase detected is often a later crustal phase of larger amplitude. This may be eitherPg which travels entirely in the upper crustal layer with a velocity of about 5.6 km/s, orP * with a path mainly in the lower crustal layer at a velocity of about 6.7 km/s. Many earthquake location programs do not take account of these phases, and treat their arrivals as if they were the earlier phasePn. At the International Seismological Centre we re-identify up to 200 crustal phases each month. This often results in significant improvement in position, the direct determination of depth or even the obtaining of a formal solution where none was possible before. We find that these re-identifications are needed in most continental areas of the world, and that the original crustal model of Jeffreys and Bullen still remains a useful standard.
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  • 2
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    Pure and applied geophysics 140 (1993), S. 63-78 
    ISSN: 1420-9136
    Keywords: Attenuation ; coda waves ; lapse-time dependence ; frequency dependence ; north Greece
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The single scattering model has been applied for the estimation of codaQ values for local earthquakes that occurred in northern Greece during the period 1983–1989 and recorded by the telemetered network of the Geophysical Laboratory of the University of Thessaloniki. CodaQ estimations were made for four frequency bands centered at 1.5 Hz, 3.0 Hz, 6.0 Hz and 12.0 Hz and for the lapse time windows 10–20 sec, 15–30 sec, 20–45 sec, 30–60 sec and 50–100 sec. The codaQ values obtained show a clear frequency dependence of the formQ c =Q 0 f n , whileQ 0 andn depend on the lapse time window.Q 0 was found equal to 33 andn equal to 1.01 for the time window of 10 to 20 sec, while for the other windowsQ 0 increased from 60 to 129, withn being stable, close to 0.75. This lapse time dependence is interpreted as due to a depth dependent attenuation. The high attenuation and the strong frequency dependence found are characteristic of an area with high seismicity, in agreement with studies in other seismic regions.
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  • 3
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    Pure and applied geophysics 140 (1993), S. 301-316 
    ISSN: 1420-9136
    Keywords: Time-predictable model ; probabilities ; South and Central America region
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The repeat times,T, of strong shallow mainshocks in fourteen seismogenic sources along the western coast of South and Central America have been determined and used in an attempt at long-term forecasting. The following relation was determined: $$\log T = 0.22M_{\min } + 0.21M_p + a$$ between the repeat time,T, and the magnitudes,M min, of the minimum mainshock considered andM p , of the preceding mainshock. No dependence of the magnitude,M f , of the following mainshock on the preceding intervent time,T, was found. These results support the idea that the time-predictable model is valid for this region. This is an interesting property for earthquake prediction since it provides the ability to predict the time of occurrence of the next strong earthquake. A strong negative dependence ofM f onM p was found, indicating that a large mainshock is followed by a smaller magnitude one, andvice versa. The probability for the occurrence of the expected strong mainshocks (M s ≥7.5) in each of the fourteen seismogenic sources during the next 10 years (1992–2002) is estimated, adopting a lognormal distribution for earthquake interevent times. High probabilities (P 10〉0.80) have been calculated for the seismogenic sources of Oaxaca, Chiapas and Southern Peru.
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  • 4
    ISSN: 1420-9136
    Keywords: Fayalite ; elastic moduli ; olivine ; resonance ; pulse-superposition ; Brillouin scattering
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We present new elasticity measurements on single-crystal fayalite and combine our results with other data from resonance, pulse superposition interferometry, and Brillouin scattering to provide a set of recommended values for the adiabatic elastic moduliC ij and their temperature variations. We use a resonance method (RPR) with specimens that were previously investigated by pulse superposition experiments. The nineC ij of fayalite are determined from three new sets of measurements. One set of our newC ij data is over the range 300–500 K. We believe that the relatively large discrep ncies found in someC ij are due in large part to specimen inhomogeneities (chemical and microstructural) coupled with differences in the way various techniques sample, rather than only systematic errors associated with experimental procedures or in the preparations of the specimens. Our recommendeaC ij's (GPa) and (∂C ij/∂T) p (GPa/K) are: The resulting values for the isotropic bulk and shear moduli,K s and μ, and their temperature derivatives are:K s=134(4) GPa; μ=50.7(0.3) GPa; (∂K s/∂T) p =−0.024(0.005) GPa/K; and (∂μ/∂T) p =−0.013(0.001) GPa/K. An important conclusion is thatK s increases as the Fe/(Fe+Mg) ratio in olivine is increased.
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  • 5
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    Pure and applied geophysics 141 (1993), S. 379-392 
    ISSN: 1420-9136
    Keywords: Elastic constant ; thermodynamics ; equation of state ; acoustic velocity ; thermal expansivity ; heat capacity ; Grüneisen constant ; high pressure ; high temperature
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The complete travel-time equation of state (CT-EOS) is presented by utilizing thermodynamics relations, such as; $$K_T = K_S (1 + \alpha \gamma T)^{ - 1} , \gamma = \frac{{\alpha K_S }}{{\rho C_P }}, \left. {\frac{{\partial C_P }}{{\partial P}}} \right)_T = - \frac{T}{\rho }\left[ {\alpha ^2 + \left. {\frac{{\partial \alpha }}{{\partial T}}} \right)_P } \right], etc.$$ The CT-EOS enables us to analyze ultrasonic experimental data under simultaneous high pressure and high temperature without introducing any assumption, as long as the density, or thermal expansivity, and heat capacity are also available as functions of temperature at zero pressure. The performance of the CT-EOS was examined by using synthesized travel-time data with random noise of 10−5 and 10−4 amplitude up to 4 GPa and 1500 K. Those test conditions are to be met with the newly developed GHz interferometry in a gas medium piston cylinder apparatus. The results suggest that the combination of the CT-EOS and accurate experimental data (10−4 in travel time) can determine thermodynamic and elastic parameters, as well as their derivatives with unprecedented accuracy, yielding second-order pressure derivatives (∂2 M/∂P 2) of the elastic moduli as well as the temperature derivatives of their first-order pressure derivatives ∂2 M/∂P∂T). The completeness of the CT-EOS provides an unambiguous criterion to evaluate the compatibility of empirical EOS with experimental data. Furthermore because of this completeness, it offers the possibility of a new and absolute pressure calibration when X-ray (i.e., volume) measurements are made simultaneously with the travel-time measurements.
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  • 6
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    Pure and applied geophysics 141 (1993), S. 467-484 
    ISSN: 1420-9136
    Keywords: Polycrystals ; hot-pressing ; multi-anvil appratus ; acoustic velocities
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In the 1960s, E. Schreiber and his colleagues pioneered the use of hot-pressed polycrystalline aggregates for studies of the pressure and temperature dependence of the elastic wave velocities in minerals. We have extended this work to the high-pressure polymorphs of mantle minerals by developing techniques to fabricate large polycrystalline specimens in a 2000-ton uniaxial split-sphere apparatus. A new cell assembly has been developed to extend this capability to pressures of 20 GPa and temperatures of 1700°C. Key elements in the new experimental design include: a telescopic LaCrO3 forT〉1200°C; Toshiba Tungaloy grade F tungsten carbide anvils; and the use of homogeneous glasses or seeded powder mixtures as starting material to enhance reactivity and maximize densities. Cell temperatures are linearly related to electrical power to 1700°C and uniform throughout the 3 mm specimens. Pressure calibrations at 25°C and 1700°C are identical to 15 GPa. Cylindrical specimens of the beta and spinel phases of Mg2SiO4, stishovite (SiO2-rutile), and majorite-pyrope garnets have been synthesized within their stability fields in runs of 1–4 hr duration and recovered at ambient conditions by simultaneously decompressing and cooling along a computer-controlledP-T path designed to preserve the high-pressure phase and to relax intergranualar stress in the polycrystalline aggregate. These specimens are single-phased, fine-grained (〈5 micron), free of microcracks and preferred orientation, and have bulk densities greater than 99% of X-ray density. The successful fabrication of these high-quality polycrystalline specimens has made possible experiments to determine the pressure dependence of acoustic velocities in the ultrasonics laboratory of S. M. Rigden and I. Jackson at the Australian National University.
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  • 7
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    Pure and applied geophysics 141 (1993), S. 497-507 
    ISSN: 1420-9136
    Keywords: High pressure ; high temperature ; thermal analysis ; diamond anvil cell ; laser heating radiometry
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A new technique actively controls thermal radiation and monitors sample properties during laser-heating in a diamond anvil cell. The technique can be described as a qualitative application of thermal analysis. Discontinuities in temperature, laser power, visible thermal radiation, or in their derivatives as functions of time can be associated with the enthalpy of phase transitions (such as melting) or with changes in maternal properties (such as emissivity). The technique is illustrated with melting experiments on iron-magnesium-silicate perovskite. Temperature corrections associated with these experiments are discussed and the results are briefly reviewed.
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  • 8
    ISSN: 1420-9136
    Keywords: Pressure ; temperature ; stress ; measurement ; acoustic emissions ; deformation ; techniques
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract New and improved techniques and apparatus for testing the mechanical properties of materials at high presures and temperatures are described. These include an improved Griggs-type deformation apparatus designed to operate to 5 GPa and associated servo-controlled hydraulic drive and electronics, the design of hydrostatic (molten alkali halide mixtures) pressure assemblies to measure flow stresses as low as a few MPa, the characterization of temperature gradients and friction in such assemblies, measurement of the melting curve of an alkali halide mixture used as a confining pressure medium, and the measurement of acoustic emissions.
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  • 9
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    Pure and applied geophysics 141 (1993), S. 545-577 
    ISSN: 1420-9136
    Keywords: Tungsten carbide ; strength ; rheology ; high pressure design
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Uniaxial compressive stress-strain curves have been measured on a suite of 26 commercial grades of tungsten carbide cermets and three maraging steels of interest for use in high-pressure apparatus. Tests were conducted on cylindrical specimens with a length to diameter ratio of two. Load was applied to the specimens by tungsten carbide anvils padded by extrudable lead disks. Interference fit binding rings of maraging steel were pressed on to the ends of the specimens to inhibit premature corner fractures. Bonded resistance strain gages were used to measure both axial and tangential strains. Deformation was exremely uniform in the central, gauged portion of the specimens. Tests were conducted at a constant engineering strain rate of 1×10−5s−1. The composition of the specimens was principally WC/Co with minor amounts of other carbides in some cases. The Co weight fraction ranged from 2 to 15%. Observed compressive strengths ranged from about 4 to just above 8 GPa. Axial strain amplitude at failure varied from ∼1.5% to ∼9%. Representative stress-strain curves and a ranking of the grades in terms of yield strength and strain at failure are presented. A power law strain hardening relation and the Ramberg-Osgood stress-strain equation were fit to the data. Fits were very good for both functions to axial strain amplitudes of about 2%. The failure of these established functions is accompanied by an abrupt change in the trend of volumetric strain consistent with the onset of substantial microcrack volume.
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  • 10
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    Pure and applied geophysics 141 (1993), S. 615-629 
    ISSN: 1420-9136
    Keywords: Experimental techniques ; calorimetry ; high-pressure ; mineral physics ; phase transitions
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We have designed and calibrated a piston-cylinder cell assembly suitable for conductingin situ measurements of enthalpies of phase transitions at elevated pressures by heat-flux differential scanning calorimetry (DSC). The high-pressure DSC detector consists of a Pt-Pt13%Rh thermopile wrapped around a frame of fired pyrophyllite. Four thermocouple junctions, arranged radially around the sample capsule, are connected in series, with four reference thermocouple junctions located 3–4 mm above the sample and embedded in thermally inert ceramic. A W-W25%Re control thermocouple is situated directly above the top of the sample; the whole detector assembly is enclosed in a 1.5 mm thick cylindrical ceramic sleeve located at the center of a 8–10 mm long “hot-zone” in the tapered graphite furnace. Using this detector design and cell assembly, we have observed the thermal signal associated with the fusion of Au at 0.5 and 1.2 GPa, and have calculated a calibration factor (K) for this detector based on the gold melting curve ofMirwald andKennedy (1979). Detector sensitivity decreases by a factor of four over this pressure-temperature interval. The reproducibility of the enthalpy of fusion of gold at 0.5 GPa suggests that detector geometry is reproducible from one experiment to the next, and thus confirms the viability of this particular detector design for quantitative DSC measurements. Subsequent experiments will assess the dependence of (K) on temperature and pressure by measuring the enthalpies of fusion of additional metals (e.g., Ag, Cu, Al, Ge) and salts (e.g., NaCl, CsCl).
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  • 11
    ISSN: 1420-9136
    Source: Springer Online Journal Archives 1860-2000
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  • 12
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    Pure and applied geophysics 140 (1993), S. 331-364 
    ISSN: 1420-9136
    Keywords: Tectonics ; subduction ; plate segmentation ; Kermadec ; earthquake ; rupture process
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract To evaluate the tectonic significance of the October 20, 1986 Kermadec earthquake (M w =7.7), we performed a comprehensive analysis of source parameters using surface waves, body waves, and relocated aftershocks. Amplitude and phase spectra from up to 93 Rayleigh waves were inverted for centroid time, depth, and moment tensor in a two-step algorithm. In some of the inversions, the time function was parameterized to include information from the body-wave time function. The resulting source parameters were stable with respect to variations in the velocity and attenuation models assumed, the parameterization of the time function, and the set of Rayleigh waves included. The surface wave focal mechanism derived (ϕ=275°, δ=61°, λ=156°) is an oblique-compressional mechanism that is not easy to interpret in terms of subduction tectonics. A seismic moment of 4.5×1020 N-m, a centroid depth of 45±5 km, and a centroid time of 13±3 s were obtained. Directivity was not resolvable from the surface waves. The short source duration is in significant contrast to many large earthquakes. We performed a simultaneous inversion ofP andSH body waves for focal mechanism and time function. The focal mechanism agreed roughly with the surface wave mechanism. Multiple focal mechanisms remain a possibility, but could not be resolved. The body waves indicate a short duration of slip (15 to 20 s), with secondary moment release 60s later. Seismically radiated energy was computed from the body-wave source spectrum. The stress drop computed from the seismic energy is about 30 bars. Sixty aftershocks that occurred within three months of the mainshock were relocated using the method of Joint Hypocentral Determination (JHD). Most of the aftershocks have underthrusting focal mechanisms and appear to represent triggered slip on the main thrust interface. The depth, relatively high stress drop, short duration of slip, and paucity of true aftershocks are consistent with intraplate faulting within the downgoing plate. Although it is not clear on which nodal plane slip occurred, several factors favor the roughly E-W trending plane. The event occurred near a major segmentation in the downgoing plate at depth, near a bend in the trench, and near a right-lateral offset of the volcanic are by 80 km along an E-W direction. Also, all events in the region from 1977 to 1991 with CMT focal mechanisms similar to that of the Mainshock occurred near the mainshock epicenter, rather than forming an elongate zone parallel to the trench as did the aftershock activity. We interpret this event as part of the process of segmentation or tearing of the subducting slab. This segmentation appears to be related to the subduction of the Louisville Ridge, which may act as an obstacle to subduction through its buoyancy.
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  • 13
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    Pure and applied geophysics 140 (1993), S. 641-653 
    ISSN: 1420-9136
    Keywords: Seismic waves ; travel time ; hypocentral surface ; southern Tyrrhenian ; Japanese islands
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A method for the calculation of seismic waves velocities at focal depth is here proposed. A stratified earth model with spherical symmetry and the analytical relationship between the epicentral distance and the travel times of seismic waves are used. This method, applied to the southern Tyrrhenian region and to the Japanese islands, allows to reduce the observed travel times to the focal depth independently of a particular velocity model.
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  • 14
    ISSN: 1420-9136
    Keywords: Complex terrain ; sea breeze evolution ; sea breeze propagation ; sea breeze rotation ; synoptic effects ; numerical approach
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Field measurements of wind, air temperature and humidity were taken at the eastern part of the Attika district in June 1991, to examine the topographic influences exerted on the local sea breezes. These influences are due either to the elongated Evia island, faced by the northern half of Attica coastline some tens of kilometers offshore, or to the coast-parallel range of Hymettos mountain, rising steeply 12 km onshore. The instrumentation consisted mainly of three tethered meteorological balloons released at characteristic sires (i.e., the coast, a location between shoreline and mountain foot and the mountain top) and three autographic ground-based anemometers operating at selected locations. Data from the ground-based and upper air stations of the Hellenic National Meteorological Service, as well as the diurnal weather maps were also obtained and analyzed. Observations were made under different synoptic wind and the latter was found to determine remarkably the significance of the topographic effects. A preliminary two-dimensional numerical approach was also made concerning the sea breeze capability to reach the Hymettos mountain top in the case of a weak opposing geostrophic flow.
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  • 15
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    Pure and applied geophysics 141 (1993), S. 1-24 
    ISSN: 1420-9136
    Keywords: Slant-stack velocity analysis ; upper mantle structure
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In this paper, regionalP-wave upper mantle structure is investigated using slant-stack velocity analysis of short-period earthquake data recorded at station MAJO (Matsushiro, Japan). Shallow earthquakes from 1980–1986 within 35° of MAJO are used to construct a common receiver gather. Processing of the wavefield data includes focal depth and static time corrections, as well as deterministic deconvolution, in order to equalize pulse shapes and align wavelets on the first arrivals. The processed wavefield data are slant stacked and interatively downward continued to obtain a regional upper mantle velocity model. The model includes a low velocity zone between 107 and 220 km. Beneath the LVZ, the velocity increases smoothly down to the discontinuity at 401 km. In the transition zone, the velocity model again increases linearly, although there is some suggestion of further complexity in the downward continued wavefield data. At the base of the transition zone, a second velocity discontinuity occurs at 660 km, with a linear velocity gradient below. In addition to slant-stack analysis, travel times and synthetic seismograms are computed and compared with the processed and unprocessed wavefield data.
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  • 16
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    Pure and applied geophysics 141 (1993), S. 249-268 
    ISSN: 1420-9136
    Keywords: Exploration tool ; physical properties ; core characterization
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We will describe a new laboratory system which was designed to be highly automated and portable while maintaining quality. Driving this design was the recognition of the temporal dependence of physical properties. It becomes apparent that some sedimentary rocks, particularly shales, degrade and disaggregare so completely that mechanical or elastic properties cannot be measured. This temporal dependence displays a time scale much shorter than normal weathering but greater than the time for stress relief. A system was designed to permit field characterization of freshly recovered core material. A benefit of automation and portability is a marked increase in measurement efficiency. The attributes of this system permit rapid characterization of a large number of fresh cores in remote, frontier exploration areas. This feature can significantly reduce prospect evaluation time. Statistically significant rock property databases can be created in a short period of time.
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  • 17
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    Pure and applied geophysics 141 (1993), S. 287-323 
    ISSN: 1420-9136
    Keywords: Sonic velocity ; carbonates ; physical properties ; porosity ; diagenesis ; compaction
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Compressional and shear-wave velocities (V p andV s) of 210 minicores of carbonates from different areas and ages were measured under variable confining and pore-fluid pressures. The lithologies of the samples range from unconsolidated carbonate mud to completely lithified limestones. The velocity measurements enable us to relate velocity variations in carbonates to factors such as mineralogy, porosity, pore types and density and to quantify the velocity effects of compaction and other diagenetic alterations. Pure carbonate rocks show, unlike siliciclastic or shaly sediments, little direct correlation between acoustic properties (V p andV s) with age or burial depth of the sediments so that velocity inversions with increasing depth are common. Rather, sonic velocity in carbonates is controlled by the combined effect of depositional lithology and several post-depositional processes, such as cementation or dissolution, which results in fabrics specific to carbonates. These diagenetic fabrics can be directly correlated to the sonic velocity of the rocks. At 8 MPa effective pressureV p ranges from 1700 to 6500 m/s, andV s ranges from 800 to 3400 m/s. This range is mainly caused by variations in the amount and type of porosity and not by variations in mineralogy. In general, the measured velocities show a positive correlation with density and an inverse correlation with porosity, but departures from the general trends of correlation can be as high as 2500 m/s. These deviations can be explained by the occurrence of different pore types that form during specific diagenetic phases. Our data set further suggests that commonly used correlations like “Gardner's Law” (V p-density) or the “time-average-equation” (V p-porosity) should be significantly modified towards higher velocities before being applied to carbonates. The velocity measurements of unconsolidated carbonate mud at different stages of experimental compaction show that the velocity increase due to compaction is lower than the observed velocity increase at decreasing porosities in natural rocks. This discrepancy shows that diagenetic changes that accompany compaction influence velocity more than solely compaction at increasing overburden pressure. The susceptibility of carbonates to diagenetic changes, that occur far more quickly than compaction, causes a special velocity distribution in carbonates and complicates velocity estimations. By assigning characteristic velocity patterns to the observed diagenetic processes, we are able to link sonic velocity to the diagenetic stage of the rock.
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  • 18
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    Pure and applied geophysics 140 (1993), S. 179-181 
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  • 19
    ISSN: 1420-9136
    Keywords: Aftershocks ; Guerrero ; seismic gap ; Mexico ; subduction
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Aftershock activity following the April 25, 1989 (M S =6.9) earthquake near San Marcos, Guerrero, Mexico, was monitored by a temporary network installed twelve hours after the mainshock and remaining in operation for one week. Of the 350 events recorded by this temporary array, 103 were selected for further analysis in order to determine spatial characteristics of the aftershock activity. An aftershock area of approximately 780 km2 is delimited by the best quality locations. The area of highest aftershock density lies inside an area delimited by the aftershocks of the latest large event in the region in 1957 (M S =7.5) and it partially overlaps the zone of maximum intensity of the earlier 1907 (M S =7.7) shock. Aftershocks also appear to cluster close to the mainshock hypocenter. This clustering agrees with the zone of maximum slip during the mainshock, as previously determined from strong motion records. A low angle Benioff zone is defined by the aftershock hypocenters with a slight tendency for the slab to follow a subhorizontal trajectory after a 110 km distance from the trench axis, a feature which has been observed in the neighboring Guerrero Gap. A composite focal mechanism for events close to the mainshock which also coincides with the zone of largest aftershock density, indicates a thrust fault similar to the mainshock fault plane solution. The San Marcos event took place in an area which could be considered as a mature seismic gap. Due to the manner in which strain release has been observed to previously occur, the occurrence of a major event, overlapping both the neighboring Guerrero Gap and the San Marcos Gap segments of the Mexican thrust, cannot be overlooked.
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  • 20
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    Pure and applied geophysics 140 (1993), S. 415-426 
    ISSN: 1420-9136
    Keywords: Site ; frequency ; ratio
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The spectral ratio method is used here for evaluating the frequency dependence of a site amplification factor in the Messina Strait area. All stations exhibit the same features of spectral ratios evidencing two peaks at about 10 Hz and 16 Hz. We relate this observation to the same geological structures. In fact, all the stations (except the referenced one) are situated over pleistocenic sediments with a similar grain size. This causes the disappearing of any site effect when the average spectrum is used as a reference. MES station, situated in the city of Messina, presents a clear site effect at a different frequency (6 Hz) which cannot be related to any geological structure because no substantial difference is observed between the sediments on which this station is situated and the sediments on which the other stations are situated. MT1 station exhibits a spectral peak at about 2 Hz which can be explained with oscillations of a little sedimentary basin.
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  • 21
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    Pure and applied geophysics 140 (1993), S. 493-501 
    ISSN: 1420-9136
    Keywords: Gravity anomalies ; sedimentary basin ; modelling ; parabolic density function
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract For modelling sedimentary basins of large thickness from their gravity anomalies, the concept of parabolic density function which explains the variation of true density contrast of the sediments with depth in such basins is introduced inBott's (1960) procedure. The analytical expression the gravity anomaly of a two-dimensional vertical prism with parabolic density contrast needed to estimate the gravity effect of the basin in modelling procedure is derived in a closed form. Two profiles of gravity anomalies, one across San Jacinto Graben, California and the other across Tucson basin, Arizona where the density of sediments is found to vary with depth are interpreted.
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  • 22
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    Pure and applied geophysics 140 (1993), S. 317-330 
    ISSN: 1420-9136
    Keywords: Seismicity ; spatio-temporal variations ; seismic gaps ; southern Peru ; northern Chile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The spatio-temporal variation of seismicity in the southern Peru and northern Chile seismic gaps is analyzed with teleseismic data (m b ≥5.5) between 1965 and 1991, to investigate whether these gaps present the precursory combination of compressional outer-rise and tensional downdip events observed in other subduction zones. In the outer-rise and the inner-trench (0 to 100 km distance from the trench) region, lower magnitude (5.0≤m b 〈5.5) events were also studied. The results obtained show that the gaps in southern Peru and northern Chile do not present compressional outer-rise events. However, both gaps show a continuous, tensional downdip seismicity. For both regions, the change from compressional to tensional regime along the slab occurs at a distance of about 160 km from the trench, apparently associated with the coupled-uncoupled transition of the interplate contact zone. In southern Peru, an increase of compressional seismicity near the interplate zone and of tensional events (5.0≤m b ≤6.3) in the outer-rise and inner-trench regions is observed between 1987 and 1991. A similar distribution of seismicity in the outer-rise and inner-trench regions is observed with earthquakes (m b 〈5.5). In northern Chile there is a relative absence of compressional activity (m b ≥5.5) near the interplate contact since the sequence of December 21, 1967. After that, only a cluster of low-magnitude compressional events has been located in the area 50 to 100 km from the trench. The compressional activity occurring near the interplate zone in both seismic gaps represents that a seismic preslip is occurring in and near the plate contact. Therefore, if this seismic preslip is associated with the maturity of the gap, the fact that it is larger in southern Peru than in northern Chile may reflect that the former gap is more mature than the latter. However, the more intense downdip tensional activity and the absence of compressional seismicity near the contact zone observed in northern Chile, may also be interpreted as evidence that northern Chile is seismically more mature than southern Peru. Therefore, the observed differences in the distribution of stresses and seismicity analyzed under simple models of stress accumulation and transfer in coupled subduction zones are not sufficient to assess the degree of maturity of a seismic gap.
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  • 23
    ISSN: 1432-041X
    Keywords: Trichostatin A ; n-Butyric acid ; Histone acetylation ; Embryos ; Echinodermata
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary External application of 10 rig/ml (R)-trichostatin A (TSA), a potent and specific inhibitor of mammalian histone deacetylase, to the embryo of the starfish Asterina pectinifera inhibited development during the early gastrula stage before formation of mesenchyme cells. The TSA-sensitive period was limited to the mid-blastula stage before hatching. The pulse-chase experiment clearly demonstrated that TSA induced an accumulation of acetylated histone species in blastulae through inhibition of historic deacetylation. Similar blockage of development at the early gastrula stage was observed with n-butyrate, which has been known as a weak inhibitor of historic deacetylase. These results suggest an intimate role for historic acetylation-deacetylation equilibria in starfish development.
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  • 24
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    Development genes and evolution 202 (1993), S. 159-169 
    ISSN: 1432-041X
    Keywords: Drosophila ; Choline acetyltransferase ; cis-Regulatory element ; lacZ reporter gene ; Colinergic neuron
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Choline acetyltransferase (ChAT, EC 2.3.1.6) catalyzes the production of the neurotransmitter acetylcholine, and is an essential factor for neurons to be cholinergic. We have analyzed regulation of the Drosophila ChAT gene during development by examining the β-galactosidase expression pattern in transformed lines carrying different lengths of 5′ flanking DNA fused to a lacZ reporter gene. The largest fragment tested, 7.4 kb, resulted in the most extensive expression pattern in embryonic and larval nervous system and likely reflects all the cis-regulatory elements necessary for ChAT expression. We also found that 5′ flanking DNA located between 3.3 kb and 1.2 kb is essential for the reporter gene expression in most of the segmentally arranged embryonic sensory neurons as well as other distinct cells in the CNS. The existence of negative regulatory elements was suggested by the observation that differentiating photoreceptor cells in eye imaginal discs showed the reporter gene expression in several 1.2 kb and 3.3 kb transformants but not in 7.4 kb transformants. Furthermore, we have fused the 5′ flanking DNA fragments to a wild type ChAT cDNA and used these constructs to transform Drosophila with a Cha mutant background. Surprisingly, even though different amounts of 5′ flanking DNA resulted in different spatial expression patterns, all of the positively expressing cDNA transformed lines were rescued from lethality. Our results suggest that developmental expression of the ChAT gene is regulated both positively and negatively by the combined action of several elements located in the 7.4 kb upstream region, and that the more distal 5′ flanking DNA is not necessary for embryonic survival and development to adult flies.
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  • 25
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    Development genes and evolution 202 (1993), S. 170-175 
    ISSN: 1432-041X
    Keywords: Axolotl ; Cell culture ; Regeneration ; Positional information
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    Topics: Biology
    Notes: Summary Previous grafting experiments have demonstrated that cells from non-contiguous positions within developing and regenerating limbs differ in a property referred to as positional identity. The goal of this study was to determine how long the positional identity of axolotl limb blastema cells is stable during culture in vitro. We have developed an assay for posterior positional properties such that blastema cells can be cultured and then grafted into anterior positions in host blastemas, to determine if they can stimulate supernumerary digit formation. We report that posterior blastema cells are able to maintain their positional identities for at least a week in culture. In addition, we observed that blastema cells are able to rapidly degrade collagenous substrates in vitro, a property that apparently distinguishes them from limb cells of other vertebrates. These results provide information regarding the time boundaries within which the positional properties of blastema cells can be studied and manipulated in vitro.
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  • 26
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    Development genes and evolution 202 (1993), S. 176-180 
    ISSN: 1432-041X
    Keywords: Juvenile hormone ; Corpora allata activity ; Sexual dimorphism ; Metamorphosis ; Apis mellifera
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Juvenile hormone synthesis in drone larvae of the honey bee was measured by an in vitro radiochemical assay. The developmental profile of corpora allata activity in male larvae showed considerable differences from queen larvae, the presumptive reproductive females, and was comparable to workers, the sterile female morph. Drone and worker larvae, however, differed drastically in the regulation of juvenile hormone biosynthesis, as revealed by the addition of farnesoic acid to the culture medium. This precursor stimulated juvenile hormone synthesis of drone glands nearly eightfold, whereas in worker larvae it is known to lead to an accumulation of methyl farnesoate. The sex-specific differences in endocrine activity indicate a role for juvenile hormone in the expression of genetically determined sexually dimorphic characters during metamorphosis, a role not currently accounted for in models describing endocrine regulation of insect development.
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  • 27
    ISSN: 1432-041X
    Keywords: Monoclonal antibodies ; Acetylcholinesterase ; Cell line ; Differentiation ; Spiralia
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    Topics: Biology
    Notes: Abstract In this study we describe the site and moment of histospecific differentiation in developmental stages of the annelid Platynereis dumerilii by use of biochemical markers. The monoclonal antibody (mab) OI7 and uncloned hybridoma supernatants (pAb's) OI8, OI10, OI46 and OI69 recognize neural antigens that appear asynchronously during development. By an enzymatic test, acetylcholinesterase (AChE) was found specific for nervous tissue as well. The patterns of neural structures labelled by antibodies differ, however, from those revealed by AChE staining. Experimental inhibition of transcription (with actinomycin D) and of translation (using puromycin) demonstrate that the expression of histospecific neural markers depends on both zygotic transcription and subsequent translation. The mAb OI64 labels epidermal (and neural) gland cells. The antibody 4D9, raised against the engrailed protein of Drosophila, labels single rows of ciliated cells at the posterior border of segments.
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  • 28
    ISSN: 1432-041X
    Keywords: Cleavage arrest ; Competence ; Neurogenesis ; Larval gland cell ; Spiralian embryogenesis
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    Topics: Biology
    Notes: Abstract This paper is devoted to the role of cell divisions for the establishment of histospecificity in the embryo of the spiralian, Platynereis dumerilii (Annelida). We have incubated successive cleavage stages in cytochalasin B (CCB) and observed whether the cells thereafter were able to acquire the competence for expressing histospecific antigens of larval gland cells (labelled by the monoclonal antibody OI64) and of neural components of the ventral nerve cord (labelled by mAb OI7 or by testing acety1cholinesterase activity), respectively. Incubation in CCB results in permanent cleavage arrest, but does not necessarily interfere with biochemical differentiation of such markers. Synthesis of the differentiation marker specific for larval gland cells does not require any cleavages but this capacity becomes restricted to the 1a and 1b cell lines if cleavages are allowed to occur. In contrast, the progenitors of neural cells need at least 6.5 h of normal development before they acquire the competence to synthesize two neural differentiation markers, which can be demonstrated after at least two more days of development. Thus, prespecifying and diversifying cleavages are a prerequisite for neurogenesis and production of the investigated neural markers. Competence for the expression of histospecific antigens may also depend on cell-cell interactions. If 20–24 h old embryos are treated with puromycin, pioneering fibres fail to grow out from a pair of posterior nerve cell progenitors as they would have done normally 24–48 h after fertilization. Concomitantly, a number of potential nerve cells which now do not come into contact with pioneering fibres do not express the neural antigen. This suggests that a local inductive stimulus from the pioneering fibres normally imprints cell fate onto ventral plate cells and turns them into neuroblasts.
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  • 29
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    Development genes and evolution 202 (1993), S. 276-295 
    ISSN: 1432-041X
    Keywords: Embryogenesis ; Muscle development ; Drosophila melanogaster ; Genetic analysis
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    Topics: Biology
    Notes: Abstract We have begun a genetic analysis to dissect the process of myogenesis by surveying the X chromosome of Drosophila melanogaster for mutations that affect embryonic muscle development. Using polarised light microscopy and antibody staining techniques we analysed embryos hemizygous for a series of 67 deletion mutations that together cover an estimated 85% of the X chromosome, or 16.5% of the genome. Whereas the mature wild type embryo has a regular array of contractile muscles that insert into the epidermis, 31 of the deletion mutants have defects in muscle pattern, contractility or both, that cannot be attributed simply to epidermal defects and identify functions required for wild type muscle development. We have defined mutant pattern phenotypes that can be described in terms of muscle absences, incomplete myoblast fusion, failure of attachment of the muscle to the epidermis or mispositioning of attachment sites. Thus muscle development can be mutationally disrupted in characteristic and interpretable ways. The areas of overlap of the 31 deletions define 19 regions of the X chromosome that include genes whose products are essential for various aspects of myogenesis. We conclude that our screen can usefully identify loci coding for gene products essential in muscle development.
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  • 30
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    Development genes and evolution 202 (1993), S. 313-315 
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  • 31
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    Development genes and evolution 202 (1993), S. 371-381 
    ISSN: 1432-041X
    Keywords: Neurogenesis ; Drosophila ; Neurogenic genes ; PNS ; Lineage
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    Topics: Biology
    Notes: Summary In Drosophila, mutations in a class of genes, the neurogenic genes, produce an excess of neurons. This neural hyperplasia has been attributed to the formation of more than the normal number of neuronal precursor cells at the expense of epidermal cells. In order to find out whether the neurogenic genes only act at this intial step of neurogenesis, we studied the replication pattern of the sensory organ precursor cells by monitoring BrdU incorporation in embryos mutant for Notch (N), Delta (Dl), mastermind (mam), almondex (amx), neuralized (neu), big brain (bib) and the Enhancer of split-Complex (E(spl)-C). Using temperature sensitive alleles of two of the neurogenic genes, DI and N, we also induced an acute increase of replicating sensory precursors by shifting briefly to the restricted temperature. We have found that the loss of function of all the seven neurogenic loci that were tested causes an increase in replicating sensory precursor cells, consistent with the model that these neurogenic genes normally participate in the process of restricting the number of neuronal precursors. Whereas the temporal pattern of replication appeared normal in mutants of five of the seven neurogenic loci, in N and mam embryos replicating PNS cells are present beyond the time when they normally undergo replication. Experiments with colchicine suggest that many of these late replicating cells may be newly emerging precursors and probably not additional cell divisions of already recruited precursors. Thus, different neurogenic genes may be required over different periods of time for the specification of sensory precursor cells.
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  • 32
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    Development genes and evolution 203 (1993), S. 3-9 
    ISSN: 1432-041X
    Keywords: Ectoderm ; Neural tube ; Chordamesoderm ; Archenteron ; Anamnia
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    Topics: Biology
    Notes: Abstract The paper considers the appearance of differences in the structure of “primary ectoderm”, i.e. of the presumptive ectoderm and chordamesoderm material in different chordates, and offers evidence that these differences foreshadow the structural peculiarities of early embryogenesis in different groups of Anamnia. These differences are: presence or absence of the archenteron, presence or absence of the hypochordal plate, formation of a neural tube with a cavity generated by epithelial folding or by splitting a compact rudiment.
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  • 33
    ISSN: 1432-041X
    Keywords: Drosophila neurogenesis ; Helix-loop-helix protein ; Enhancer of split
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    Topics: Biology
    Notes: Abstract To assess the functional domains of the proteins encoded by E(spl) and HLH-m5, two genes of the Enhancer of split complex [E(SPL)-C] of Drosophila melanogaster, a number of variants have been made by in vitro mutagenesis, transformed into the germ line of the wild-type, and genetically combined with a chromosomal deletion lacking four of the genes of the E(SPL)-C. All constructs used attenuated the neurogenic phenotype associated with this deletion. However, constructs encoding proteins with truncated carboxy-termini exibited in all cases a higher activity than constructs encoding the full length version of the protein. Neutralization of the basic domain severely reduced, but did not completely abolish the rescuing activity of E(spl), while proteins in which a proline residue within the basic domain had been changed to either threonine or asparagine were slightly less efficient in their rescuing activity than the corresponding wild-type versions. We discuss the possible significance of these results for the function of the protein domains.
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  • 34
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    Development genes and evolution 203 (1993), S. 28-33 
    ISSN: 1432-041X
    Keywords: Mesoderm differentiation ; Marginal zone ; Animal region ; Amphibia
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    Topics: Biology
    Notes: Abstract Embryos of Ambystoma mexicanum from the late morula to the late blastula stage were dissected and cultivated in varying combinations. The marginal zone (presumptive mesoderm) when isolated together with the vegetal region differentiated to notochord after dissection from early blastulae, but did not differentiate to other tissues. When isolated from middle to late blastulae, in addition myoblasts and mesenchyme were formed. The marginal zone isolated together with the animal region (presumptive ectoderm) differentiated to notochord, muscle, mesenchyme, renal tubules and mesothelium irrespective of the stage of dissection. Combination of isolated animal and vegetal regions did lead to the induction of mesodermal organs. The experiments suggest that further steps in the differentiation of mesodermal organs after the induction of mesoderm by the vegetalizing factor depend on factors from the animal region, which are involved in pattern formation.
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  • 35
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    Development genes and evolution 203 (1993), S. 18-27 
    ISSN: 1432-041X
    Keywords: Oogenesis ; Germ line cell cluster ; Oocyte determination ; Ultrastructure ; Mayflies
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    Topics: Biology
    Notes: Abstract Germ line cell cluster formation in ovarioles of three different stages, each from a different mayfly species, was studied using ultra-thin serial sectioning. In the analysed ovariole of Cloeön sp., only one linear, zigzag germ line cell cluster was found, consisting of sibling cells connected by intercellular bridges which represent remnants of preceding synchronized mitotic cycles followed by incomplete cytokinesis. A polyfusome stretched through all sibling cells. At the tip of the ovariole, cytokinesis occurred without preceding division of nuclei; thus, intercellular bridges were lined up but the remaining cytoplasm between the bridges had no nuclei. The analysed Siphlonurus armatus vitellarium contained five oocytes at different stages of development. Each oocyte in the vitellarium was connected via a nutritive cord to the linear cluster of its sibling cells in the terminal trophic chamber. Each cluster had the same architecture as was found in Cloëon. The 3-dimensional arrangement and distribution of closed intercellular bridges strongly suggest that all five clusters are derived from a single primary clone. The position of oocytes within each cluster is random. However, each oocyte is embraced by follicular or prefollicular cells whilst all other sibling cells are enclosed by somatic inner sheath cells, clearly distinguishable from prefollicular cells. In the analysed ovariole of Ephemerella ignita, two small linear clusters were found in the tropharium beside two single cells, two isolated cytoplasmic bags with intercellular bridges but no nuclei, and some degenerating aggregates. One cluster was still connected to a growing oocyte via a nutritive cord. In all species the nurse cells remained small and no indications of polyploidization were found. We suggest that this ancient and previously unknown telotrophic meroistic ovary has evolved directly from panoistic ancestors.
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  • 36
    ISSN: 1432-041X
    Keywords: Bicoid activity ; Heterospecific transplantation ; bicoid orthologous homeobox fragments ; Calliphoridae
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    Topics: Biology
    Notes: Abstract In order to test for bicoid-like activity in insects other than Drosophila melanogaster, anterior egg cytoplasm from the following species was injected into cleavage stage embryos from mutant D. melanogaster lacking a functional bicoid (bcd) product: six other Drosophila species, the housefly, three blowfly species, the primitive cyclorrhaphic dipteran Megaselia, and the honeybee Apis mellifera; preliminary tests were made with four lower dipterans (Nematocera). Rescue effects were only observed with the drosophilids, housefly, and two of the three blowfly species. Rescue was stronger with the drosophilids than with the other flies as donors. Where checked (D. pseudoobscura), a positive correlation was found between the amount of cytoplasm injected and the number of pattern elements formed, suggesting threshold effects upon target genes as with the endogenous bcd product. By polymerase chain reaction, fragments from a bcd-orthologous homeobox were cloned from the three blowfly species. The derived sequence of 43 amino acids was identical in all blowflies and the housefly but differed at 4 positions from the orthologous D. melanogaster sequence. Localization of the mRNA recognized by the respective fragments in the blowflies Lucilia and Phormia resembled that known from D. melanogaster, while Calliphora — the blowfly species lacking rescue activity —showed remarkable differences of localization in both ovarian follicles and the deposited egg cell. This surprising divergence within a morphologically rather uniform family of cyclorrhaphic dipterans should be of interest from both functional and evolutionary points of view.
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  • 37
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    Development genes and evolution 203 (1993), S. 44-50 
    ISSN: 1432-041X
    Keywords: Insect egg ; Cell fusion ; Nuclear injection ; Cell cycle ; Pattern formation
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    Topics: Biology
    Notes: Abstract Explanted oocytes and eggs of different developmental stages from the Hymenopteron Pimpla were fused in pairs as “parabiotic tandems”. Interactions within the tandem were analysed by time lapse films. Except for the exchange of nuclei, no joint development was observed, and each partner followed its own time pattern. The rapid cell cycles of the cleavage energids switched over to longer cycles according to the local developmental stage of the different egg regions, although all of the nuclei were still contained in a single plasmodium. In a second experimental series, nuclei in newly deposited eggs were X-rayed and replaced by cleavage energids from later stages injected into the wrong (=posterior) egg pole. Even injected blastoderm nuclei immediately took up mitotic activities and underwent rapid cell cycles characteristic of early cleavage. Normal embryos could be formed, although the nuclei had populated the egg in a reverse direction.
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  • 38
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    Development genes and evolution 203 (1993), S. 51-59 
    ISSN: 1432-041X
    Keywords: Insect egg ; Nuclear injection ; Activation ; Cell cylce length ; Pattern formation
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    Notes: Abstract Oocytes explanted from adult ovaries of the arrhenotokous Hymenopteron Pimpla turionellae remain in an inactive state, because development has not been initiated by mechanical deformation during natural oviposition. However, they could be induced to enter development by injecting cleavage energids into the posterior pole. After lag phases of up to 32 h, the implanted nuclei initiated a normal cleavage process, except that the polarity of its progress was reversed. In other oocytes, the injected energids congregated in a ring-shaped region at the egg surface to form a superficial “nuclear front”, which slowly advanced towards the anterior egg pole, thereby successively stimulating portions of the quiescent ooplasm to take part in development. Up to 41 rapid cell cycles started from that front, each of them with an anaphase wave running backwards into the region already peripherally occupied by nuclei. Thus, the blastoderm was formed extremely metachronously and by rapid obviously biphasic cell cycles, which never occur at the egg surface during normal cleavage. A germ band, however, was only formed under the following conditions: (1) that cleavage did not follow the nuclear front mode, and (2) that ooplasm from the donor's posterior pole was co-injected with the graft nuclei. We conclude that embryonic differentiation requires some of the events which had been omitted in eggs where development failed, especially the exponential increase of the cell cycle length, and the activity of some posterior factor(s) during egg activation.
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  • 39
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    Development genes and evolution 203 (1993), S. 60-73 
    ISSN: 1432-041X
    Keywords: Head development ; Eye-antenna disc ; Drosophila
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    Notes: Abstract The embryonic development of the primordia of the Drosophila head was studied by using an enhancer trap line expressed in these structures from embryonic stage 13 onward. Particular attention was given to the question of how the adult head primordia relate to the larval head segments. The clypeo-labral bud to the stage 13 embryo is located at a lateral position in the labrum adjacent to the labral sensory complex (“epiphysis”). Both clypeo-labral bud and sensory complex are located anterior to the engrailed-expression domain of the labrum. Throughout late embryogenesis and the larval period, the clypeo-labral bud forms integral part of the epithelium lining the roof of the atrium. The labial disc originates from the lateral labial segment adjacent to the labial sensory complex (“hypophysis”). It partially overlaps with the labial en-domain. After head involution, the labial disc forms a small pocket in the ventro-lateral wall of the atrium. The eye-antenna disc develops from a relatively large territory occupying the dorso-posterior part of the procephalic lobe, as well as parts of the dorsal gnathal segments. Cells in this territory are greatly reduced in number by cell death during stages 12–14. After head involution, the presumptive eye-antenna disc occupies a position in the lateral-posterior part of the dorsal pouch. Evagination of this tissue occurs during the first hours after hatching. In the embryo, no en-expression is present in the presumptive eye-antenna disc. en-expression starts in three separate regions in the third instar larva.
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  • 40
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    Development genes and evolution 203 (1993), S. 74-82 
    ISSN: 1432-041X
    Keywords: Artemia ; Pattern formation ; Purine nutrition ; Guanylate ; Mycophenolic acid
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    Topics: Biology
    Notes: Abstract Mycophenolic acid (MA) reduces growth and survival rates of Artemia larvae fed hypoxanthine. The inhibitory effect of MA is suppressed by dietary guanosine as expected for inosinate dehydrogenase inhibition. Pulse MA treatments lasting 24 h imposed on larvae from 24 to 96 h posthatch result in the production of abnormal adults with thoracic, genital or abdominal defects. The MA-induced anomalies are interpreted as a disruption of pattern formation. Since dietary guanosine restores normal pattern in adults developing from larvae, MA-treated during early stages, it is concluded that guanylate production is required at critical developmental times to establish normal developmental fates.
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  • 41
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    Development genes and evolution 203 (1993), S. 83-91 
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    Keywords: Drosophila ; Monensin ; Extracellular matrix ; Membrane proteins ; Morphogenesis
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    Notes: Abstract Extracellular matrix and membrane proteins and their correct secretion probably are key elements in morphogenesis and differentiation in Drosophila. In this study, we have analysed the effects of monensin, a Na+-H+-ionophore which blocks normal secretion, applied during cellular blastoderm formation on further development. Normal cell morphology and intercellular contacts are lost and the extracellular matrix becomes disorganized. Gastrulation is blocked and abnormal foldings can be observed. Cuticle phenotypes showed different degrees of ventral, dorsal, head and posterior defects. The results are discussed in the context of what is known about membrane and extracellular matrix proteins in Drosophila.
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  • 42
    ISSN: 1432-041X
    Keywords: aFGF ; bFGF ; Palate development ; Extracellular matrix ; Growth factors
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    Notes: Summary The distribution of acidic and basic fibroblast growth factors (aFGF, bFGF) was mapped during mouse embryonic palate development. Generally, they localised most intensely in the basement membrane and epithelia rather than the mesenchyme. Localisation was predominantly restricted to the palatal nasal, and medial edge epithelia. Staining was particularly intense in the medial edge epithelia at the time of mid-line epithelial seam formation. Intense staining persisted in the epithelia of the degenerating seam and later in the oral and nasal epithelial triangles. Mouse embryonic palate mesenchyme (MEPM) cells cultured in vitro on a variety of substrata (on plastic, on the surface of a collagen gel and within a collagen gel) responded to treatment with aFGF or bFGF. These responses were modulated by the culture substratum. The FGFs stimulated MEPM cell proliferation on plastic and on collagen, but inhibited cell growth in collagen. The FGFs had little effect on protein production when cells were cultured on plastic, but caused a large reduction in on-collagen and incollagen cultures. This reduction was greater in collagenous than non-collagenous proteins. Generally, treatment with FGFs stimulated the production of glycosaminoglycans (GAGs), particularly hyaluronan (HA) and dermatan sulphate (DS). In addition, the size class of HA was shifted to a higher molecular weight form. These data indicate that aFGF and bFGF may play a role in modulating mesenchymal cell matrix biosynthesis, so facilitating palatal epithelial seam degeneration.
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  • 43
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    Keywords: Gastrulation ; Fibronectin ; Fish development
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    Notes: Summary The present report firstly describes a pilot study in which, during early development of embryos of the common carp, Cyprinus carpio, the cellular adhesion to fibronectin (FN) was blocked by administration of GRGDS peptide (which binds to the FN-receptor). As this treatment resulted in developmental aberrations, suggesting a functional role for FN, the major part of the work was focussed on the distribution of reactivity of anti-FN antibodies during epiboly and gastrulation. GRGDS treatment had a concentration dependent effect on development. Incubation of embryos in 1.5 mg/ml from the 32-cell stage onwards caused a retardation of epiboly, which did not proceed beyond 60%. The embryos did not show involution, as was confirmed by histological study. These preliminary results suggest that FN is involved in both epiboly and gastrulation of carp embryos. During cleavage, no specific extracellular binding of anti-FN antiserum could be observed. However, binding to a number of cell membranes took place from early epiboly onwards. After the onset of gastrulation, we observed a gradually increasing number of the deepest epiblast cells, showing immunostaining on part of their surface, facing the yolk syncytial layer (YSL) or the involuted cells. During early epiboly, anti-FN binding was restricted to areas in front of the migratory hypoblast cells. Later on, binding was found at the border of hypoblast and epiblast cells. At 100% epiboly, some contact areas of epiblast and hypoblast showed a discontinuous lining of reactivity, whilst other areas appeared devoid of anti-FN binding sites. The results indicate that FN is involved in the migration and guidance of hypoblast cells during gastrulation in carp.
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  • 44
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    Development genes and evolution 202 (1993), S. 224-232 
    ISSN: 1432-041X
    Keywords: Frog embryos ; Microtubules ; Dorsoventral polarity ; Sperm aster
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    Topics: Biology
    Notes: Abstract Two microtubule-containing structures are implicated in dorsoventral polarization of the frog egg, and we examined the relationship between them. The sperm aster provides a directional cue for a cortical rotation specifying polarity, and a vegetal cortical array of parallel microtubules is likely part of the rotational machinery. The growing aster has an accumulation of microtubules marking the path of the sperm pronucleus, and its microtubules extend into the egg cortex as well as the cytoplasm. To test whether the vegetal parallel array was an extension of astral cortical growth, fertilized or activated eggs were bisected into animal and vegetal fragments. The vegetal fragments formed parallel arrays, even when isolated within a few minutes of egg activation. Neither the sperm centrosome nor another microtubule organizing center in the animal half of the egg is required for formation of the parallel array, but some animal half activity is involved in its disappearance.
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  • 45
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    Development genes and evolution 202 (1993), S. 214-223 
    ISSN: 1432-041X
    Keywords: Drosophila embryogenesis ; Pharyngeal primordia ; cnc gene ; Cephalic patterning ; Segmentation gene
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    Notes: Abstract Genetic controls regulating the establishment of the pharyngeal primordia in the anterior region of the Drosophila embryo were investigated through the analysis of the expression of thecnc gene, which is continuously expressed specificially in three pharyngeal segments. The spatial regulation ofcnc gene transcription was analyzed by in situ hybridization of CNC transcript-specific probes to embryos mutant for other cephalic patterning genes. The anterior domain of CNC expression (corresponding to the labral segment primordium) was found to be activated bybicoid andtorso maternal pathways, independently of known zygotic gap genes, and sequentially constricted to its final size by repression from neighboring region-specific genes. Control of the posterior domain (corresponding to the intercalary and mandibular segment primordia) involved combinatorial regulation by zygotic gap genes: activation by thebtd gap gene and repression from theotd gap gene anteriorly and thesna gene ventrally. Surprisingly, the posterior domain was shifted relative to the segmentation plan in mutants of theems gap gene. These regulatory controls establishing the limits of CNC expression in the pharyngeal primordia suggest that one mechanism for patterning within the anterior terminal region may involve direct activation of region-specific gene(s) by maternal factors over a relatively broad domain followed by constriction of that domain by repression from adjacently activated zygotic genes.
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  • 46
    ISSN: 1432-041X
    Keywords: Determination/intermediate ; filament/gene ; expression/in situ ; hybridization/chick ; embryo/germ layers
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    Notes: Abstract Using in situ hybridization we show that a chick type II cytokeratin (CKsel) is differentially expressed within the developing ectoderm, mesoderm, and endoderm as early as the time of germ layer formation. CKsel expression demarcates specific regions that in many instances can be correlated with distinct presumptive developmental fates or potencies. In the epiblast of definitive streak stage embryos, CKsel hybridization is detected only in regions destined to form extraembryonic ectoderm. After this stage expression occurs within embryonic ectoderm but is largely restricted to regions that will form epidermis. It is not detected in presumptive neural ectoderm or in most regions of head ectoderm. The area of non-expression of CKsel in head ectoderm delineates ectoderm which has a strong bias for lens formation, and gives rise not only to lenses but to other placodally derived structures such as the nasal epithelium and the otic vesicles. In the developing mesoderm, CKsel transcripts become restricted to regions of lateral plate and extraembryonic mesodermal tissues at later stages. CKsel is expressed throughout the developing endoderm except in the region of the presumptive foregut roof. There is a unifying pattern to the expression of CKsel transcripts in all three germ layers: it is expressed in more lateral areas which generally have more ventral fates, beginning at the time of germ layer formation and indicating the existence of medial/lateral boundaries early in development.
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  • 47
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    Development genes and evolution 202 (1993), S. 364-370 
    ISSN: 1432-041X
    Keywords: Thrombomodulin expression ; Murine development ; Angiogenesis ; Monoclonal Antibodies
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    Topics: Biology
    Notes: Abstract Thrombomodulin is a transmembrane glycoprotein involved in the regulation of clot formation. Previous work has shown that in the adult mouse, thrombomodulin expression is primarily located on the luminal surface of endothelial cells; whereas during development, thrombomodulin has been localized to the placental parietal endoderm, the neuroepithelium, head mesenchyme, lung bud, and atrium of Day 10.5 post coitum embryos. We examined the expression of thrombomodulin in the adult mouse and throughout murine development from Day 7.5 to Day. 18.5 post coitum. The high expression of thrombomodulin in the lung bud, developing gonads, leptomeninges, and the parietal endoderm in comparison to the expression levels in the adult organs suggest that thrombomodulin may play an important role in development. The submitted manuscript has been authored by a contractor of the U.S. Government under contract No. DE-AC05-840R21400. Accordingly, the U.S. Government retains nonexclusive, royalty-free license to publish or reproduce the published form of this contribution, or allow others to do so, for US Government purposes.
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    Development genes and evolution 202 (1993), S. 383-383 
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  • 49
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    Development genes and evolution 203 (1993), S. 1-1 
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  • 50
    ISSN: 1432-041X
    Keywords: Porcine ; Embryo ; Inner cell mass ; Trophectoderm ; Segregation
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    Notes: Abstract The mammalian blastocyst consists of an inner cell mass (ICM) enclosed by the trophectoderm. The origin of these two cell populations lies in the segregation of inner and outer cells in the early morula. In the present study, the segregation of inner and outer cells has been studied in porcine embryos and is compared with segregation in mouse embryos. For this, nuclei of inner and outer cells were differentially labelled with two fluorochromes after partial complement-mediated lysis of the outer cells. In porcine and mouse embryos compaction and the first appearance of inner cells occur at different stages of development. In porcine embryos compaction was observed as early as the 4-cell stage, while in mouse embryos compaction occurred in the 8-cell stage. The first inner cells segregated in porcine embryos which were in the transition from four to eight cells and inner cells were added during two subsequent cell cycles. In mouse embryos inner cells segregated predominantly during the fourth cleavage division. From the results obtained we conclude that the segregation of inner and outer cells follows a different pattern in mouse and in porcine embryos.
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  • 51
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    Keywords: Ascidians ; Muscle actin gene ; Fusion gene construct ; Specific expression ; Muscle lineage cells
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    Notes: Abstract pHrMA4a-Z is a recombinant plasmid in which about 1.4 kb of the 5′ flanking region of a gene for muscle actin HrMA4a from the ascidian Halocynthia roretzi is fused with the coding sequence of a bacterial gene for β-galactosidase (lac-Z). In this study, we examined the expression of the fusion gene construct when it was introduced into eggs of another ascidian, namely Ciona savignyi. When a moderate amount of linearized pHrMA4a-Z was introduced into fertilized Ciona eggs, the expression of the reporter gene was evident in muscle cells of the larvae, suggesting that both species share a common machinery for the expression of muscle actin genes. The 5′ upstream region of HrMA4a contains several consensus sequences, including a TATA box at -30, a CArG box at -116 and four E-boxes within a region of 200 bp. A deletion construct, in which only the 216-bp 5′ flanking region of HrMA4a was fused with lac-Z, was expressed primarily in larval muscle cells. However, another deletion construct consisting of only the 61-bp upstream region of HrMA4a fused with lac-Z was not expressed at all. When pHrMA4a-Z or ΔpHrMA4a-Z (−216) was injected into each of the muscle-precursor blastomeres of the 8-cell embryo, expression of the reporter gene was observed in larval muscle cells in a lineage-specific fashion. However, expression of the reporter gene was not observed when the plasmid was injected into non-muscle lineage. Therefore, the expression of the reporter gene may depend on some difference in cytoplasmic constituents between blastomeres of muscle and non-muscle lineage in the 8-cell embyo.
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  • 52
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    Development genes and evolution 203 (1993), S. 121-123 
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  • 53
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    Development genes and evolution 203 (1993), S. 117-120 
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    Keywords: Endogenous lectins ; Morphogenesis ; Xenopus laevis
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    Notes: Abstract Galactoside-binding lectin has been isolated from whole Xenopus laevis embryos and tadpoles at four development stages: st. 24–26, 32, 41 and 47. The main lectin activity at st. 24–26 is β-galactoside specific, producing a 34/35.5K doublet on SDS-PAGE. Later in development, lectin activities specific for a wide range of other sugars appear concommitant with the detection of a number of new protein bands on SDS-PAGE gels. The greatest variety of new lectin activities exists at st. 32 when lectins specific for all of the main sugar families found in nature are detected. After this stage and up to st. 47 (the beginning of metamorphosis), fewer different lectin activities are again detected. The results suggest that a complex, developmentally regulated battery of different lectins are present during early Xenopus development, perhaps with stage-specific roles to play in the control of tissue morphogenesis.
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  • 54
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    Keywords: Wing imaginal discs ; Growth ; Metamorphosis ; Protein database ; two-dimensional gel electrophoresis
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    Notes: Abstract High-resolution two dimensional gel electrophoresis has been used to study the patterns of protein synthesis in imaginal discs of Drosophila melanogaster. In this paper we first compare the patterns of protein synthesis in wing, haltere, leg 1, leg 2, leg 3 and eye antenna imaginal discs of late third instar larvae. We have detected only quantitative changes: differences in 17 proteins among the different imaginal discs. In addition, we have analysed the variations in pattern of proteins in the wing disc of the last larval stage and early pupae as well as in wing discs cultured in vivo for 6 days. Variations in these patterns affect more than 20% of the proteins and involve both qualitative and quantitative changes. Some of the changes may correspond to protein phosphorylation. Correlations of these changes between discs and through development are also discussed.
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    Development genes and evolution 203 (1993), S. 124-130 
    ISSN: 1432-041X
    Keywords: Teleost embryogenesis ; Epiboly ; Enveloping layer ; Epidermis ; Mouth-breeding cichlids
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    Notes: Abstract The eggs of African mouth-brooders are of unusual size and shape. Studying their development may help to more clearly understand epiboly, gastrulation, and the relation between enveloping layer (periderm) and epidermis. When epiboly has progressed over just one fifth of the yolk mass, the germ ring and embryonic shield are already well established. Behind the germ ring very few deep cells are present at this early stage of epiboly, except in the embryonic shield. When the blastodisc covers the animal half of the yolk mass, the future body is already well established with notochord, somites and developing neural keel. Apart from these structures, no deep cells can be detected between enveloping layer and yolk surface; not even a germ ring remains behind the advancing edge of the enveloping layer. Epiboly over the greater part of the yolk is achieved only by the enveloping layer and the yolk syncytial layer. As the margin of the enveloping layer begins to reduce its circumference when closing around the vegetal pole, groups of cells in the advancing edge become spindle-shaped, with a single cell in between of each of these groups broadening along the edge. The enveloping layer (called periderm after epiboly) remains intact until after hatching, when, together with the underlying ectoderm, it forms the double-layered skin of the larval fish. Thereafter, cells deriving from the subperipheral ectoderm gradually replace the decaying periderm cells to form the final epidermis. Thus, in the cichlids studied, the enveloping layer alone forms the yolk sac to begin with, and it covers the larval body until some days after hatching.
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  • 56
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    Development genes and evolution 203 (1993), S. 151-158 
    ISSN: 1432-041X
    Keywords: Drosophila melanogaster ; Pattern formation ; Neurogenesis ; Peripheral nervous system ; Microchaetes
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    Notes: Abstract The small bristles (microchaetes) on the thorax of adult Drosophila are evenly spaced. We have analysed the development of this pattern using the enhancer trap line A101 where bacterial lacZ is expressed in the microchaete sensory mother cells (SMCs) and their progeny. We observed that the precursor cells appear in a stereotyped pattern of rows. Within each row, however, SMCs appear neither at a time nor in a restricted sequence: new SMCs are continuously intercalated between pre-existing SMCs until the distance between consecutive SMCs does not exceed a few cell diameters. In large individuals, additional SMCs may occasionally appear after the completion of the rows, in the largest empty spaces between the preexisting SMCs.
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  • 57
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    Keywords: Palate development ; Epidermal growth factor ; Transforming growth factor alpha ; Extracellular matrix
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    Notes: Abstract Mouse embryonic palatal mesenchymal (MEPM) cells were cultured either on plastic tissue culture dishes or on the surface of three-dimensional collagen gels or within collagen gel matrices in DMEM/F12 medium containing 2.5% donor calf serum. MEPM cells proliferated exponentially when cultured on collagen or on plastic. Cells cultured within collagen gels did not proliferate but remained viable. Addition of 10 ng/ml epidermal growth factor (EGF) or transforming growth factor alpha (TGFα) stimulated the proliferation of those cells cultured on plastic or on collagen but not those cultured within collagen gels. Immunocytochemical analysis revealed that MEPM cells synthesise collagen types I, III, IV, V, VI and IX; fibronectin, heparan sulphate proteoglycan, laminin and tenascin in vitro. These molecules are all present in the developing palate in vivo. EGF and TGFα produced a generalised stimulation of extracellular matrix (ECM) synthesis by MEPM cells in vitro. Biochemical analysis indicated that cells cultured within collagen gels had the highest intrinsic rate of protein synthesis. On all substrata neither EGF nor TGFα markedly altered the types of ECM molecules synthesised but rather caused a general increase in the total amount produced. This stimulation was most marked where the cells were cultured within collagen gels. The lack of stimulation of proliferation of MEPM cells cultured within collagen gels (i.e. in a physiologically-relevant environment) by EGF or TGFα together with the marked stimulation of ECM synthesis suggests that these factors may act as differentiation signals via their effects on ECM production.
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    Development genes and evolution 203 (1993), S. 159-163 
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    Keywords: Chick development ; Mesoderm induction ; Activin ; Vg1 ; Transforming growth factor-β
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    Notes: Abstract Transforming growth factor-β (TGF-β)-related genes have been postulated as possible mesoderm inducers in lower vertebrates. Activin, a member of the TGF-β family has been recently suggested to be directly involved in the induction of axial mesoderm in frogs and birds. Nodal, a novel TGF-β-like gene, seems to be essential for mesoderm formation in mice. In this work we have tried to characterize which growth factors of the TGF-β family are involved in the initial stages of differentiation of the chick embryo. Using degenerate primers to specific regions of the TGF-β super family in polymerase chain reactions (PCR), we have identified both TGF-β2 and TGF-β3 transcripts in early stages of chick development. In addition, we have identified a new gene which is transcribed during early chick development. We have called this gene Vgc1 since it is probably the chick homologue of the mouse Vg-related gene Vgrl. RNA coding for the Vgcl gene was detected by RNAse protection in early stage X blastoderms, gastrula and 3-day-old embryos. In situ hybridization experiments further indicate that Vgcl RNA is distributed uniformly throughout the various regions of the chick gastrula. The role of Vgcl in early chick development remains to be established.
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  • 59
    ISSN: 1432-041X
    Keywords: Retinoic acid ; Fish embryos ; Early development
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    Notes: Abstract The present study reports the early effect of exogenously applied retinoic acid (RA) on embryos of the Japanese medaka, Oryzias latipes. Furthermore, as not much is known about early medaka development, a morphological description of control embryos is given and compared with well studied early cyprinid development. In medaka embryos epiboly and convergence roughly start at the same stage. Gastrulation, resulting in an outer epiblast and an inner hypoblast layer, is observed from about 25% epiboly onwards and is restricted to the dorsal side, since at that stage converging cells are already located relatively dorsally. RA treatment leads, in a stage- and dose-dependent way, to a shortening of the embryonic axis. This may be caused by a complete or partial failure of extension movements which may occur during convergence. Convergence and gastrulation appear to be, at least partly, unaffected by RA since the embryo proper is located dorsally and consists of two separate germ layers. Observation of medaka embryos until 4 days post fertilization (dpf) revealed that RA treatment caused poorly developed anterior structures (eyes, brain, heart, blood vessels). However, pectoral fins and melanin granules were always present.
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    Development genes and evolution 203 (1993), S. 164-168 
    ISSN: 1432-041X
    Keywords: Rhodopsin ; Retina ; Development ; Amphibia ; Photoreceptors
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    Topics: Biology
    Notes: Abstract Notophthalmus (Triturus) viridescens, a urodele amphibian (newt) common to the Eastern United States, is a promising subject for developmental and regeneration studies. We have available a monoclonal antibody shown to be specific in many vertebrates for rod opsin, the membrane apoprotein of the visual pigment rhodopsin. This antibody to an N-terminal epitope, by rigorous biochemical and immunological criteria, recognizes only rod photoreceptor cells of the retina in light-and electron-microscopic immunocytochemistry. To determine the ontogeny and localization of rhodopsin in developing rods as an indicator of function in the embryonic urodele retina, we have utilized this antibody in the immunofluorescence technique on sections of developing N. viridescens. It was applied to serial sections of the eye region of Harrison stage 28 (optic vesicle) through stage 43 (most adult retina histology complete) embryos, and subsequently visualized with biotinylated species antibody followed by extravidin fluorescein isothiocyanate. The first positive reaction to rhodopsin could be detected in two to four cells (total) of the stage 37 embryonic eye, in the region of the central retinal primordium where the photoreceptors will be found. Some indications of retinal outer nuclear and inner plexiform layers could be seen at this time. Later embryonic stages demonstrated increasing numbers of positive cells in the future photoreceptor outer nuclear layer and outer and inner segments, spreading even to the peripheral retina. Nevertheless, by stale 43, no positive cells could be found at the dorsal or ventral retinal margins. Thus, biochemical differentiation of a photoreceptor population in the urodele retina occurs at a stage before retinal histogenesis is complete. The total maturation of retinal rods occurs topographically over a long period until the adult distribution is achieved.
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    Development genes and evolution 202 (1993), S. 67-69 
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  • 62
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    Development genes and evolution 202 (1993), S. 70-76 
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    Keywords: Arachidonic acid ; HETE ; Protein kinase C ; Lipoxygenase ; Hydra ; Coelenterates
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    Notes: Summary Repeated stimulation ofHydra magnipapillata with the diacylglycerol (DG) 1,2-sn-dioctanoylglycerol (diC8) induces an increase in positional value and eventually the development of ectopic heads. Upon stimulation, the polyps release [14C]-arachidonic acid from previously labelled endogenous sources. Arachidonic acid (AA) is not released into the external medium but remains within the animal, AA, linoleic acid and their lipoxygenase products were identified by gas chromatography-mass spectrometry. Several metabolites were found, most abundantly 12-HETE (hydroxy-eicosa-tetraenoic acid), 8-HETE, 9-HODE (hydroxy-octadecadienoic acid), and 13-HODE; this is the first evidence of their presence in coelenterates. Externally applied AA causes ectopic head formation, though less effectively than diC8. When administered simultaneously, (diC8) and AA, which both are known to activate protein kinase C (PKC), act synergistically in inducing ectopic head formation. Since released endogenous AA can spread in tissues, it may mediate a temporal and spatial extension of PKC activation and, hence, broaden the range in which positional value increases. However, in addition to the activation of PKC, the generation of AA metabolites appears to be essential for the induction of ectopic head formation, since not only a selective inhibitor of PKC, chelerythrine, but also an inhibitor of lipoxygenases, NDGA (nordihydroguaiaretic acid), significantly reduces the effectiveness of both AA and DG.
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    Development genes and evolution 202 (1993), S. 77-84 
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    Keywords: Actin ; Patella vulgata ; Development ; Differential expression ; F-actin
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    Notes: Summary The actin gene family of the marine molluscPatella vulgata was chosen as a model system to study the regulation of genes expressed during early development in molluscs. Using a hamster actin cDNA clone as a probe, we isolated nine actin cDNA clones from trochophore larvae. The total nucleic acid sequence of three of these clones has been determined. Each clone contains the whole protein encoding region. The deduced amino acid sequences resemble actin proteins from other species to a high extent. The nucleotide sequence from the 3′UTR (UnTranslated Region) and 5′UTR from all nine clones has been resolved. In this way we could identify four different subtypes. Southern blots with genomic DNA were probed with different 3′UTR's corresponding to each subtype to determine the genomic organization. One 3′UTR detected one band probably corresponding with one gene. Another 3′UTR detected one or two genes and the third 3′UTR between two and four genes. Northern blots were used to detect the presence of actin mRNA during different stages of development. In the mature oocyte, actin mRNA is present in low amounts. The level of actin mRNA starts to rise steadily from 8 h after fertilization (88-cell stage) onwards. The level of the different subtype mRNAs, as specified by their 3′UTR rises at different developmental stages and to various extents. This indicates that the expression of each type is regulated independently and in relation to the developmental stage of the embryo.
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    Development genes and evolution 202 (1993), S. 95-102 
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    Keywords: Developmental focus ; Genetic mosaics ; Myogenesis ; Pupa
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    Topics: Biology
    Notes: Summary InDrosophila, the temperature-sensitiveshibire (shi) mutation causes blockage of endocytosis. Disruption ofshi function by heat pulse at a sensitive pupal stage produces adults with altered flight muscles and motoneurons (Hummon and Costello 1988b). Thus, endocytosis is evidently required for normal development of flight muscle, but the site(s) with primary sensitivity to disruption ofshi function has not been shown. In Ring-X generated mosaic animals (wild type/shi), the inducedshi flight muscle phenotype maps to the area of the blastoderm fate map containing the presumptive thoracic mesoderm, the developmental foci for specific muscles (Hummon and Costello 1992). We use these wild type/shi mosaics to look for independence versus correlation between the pattern of muscle phenotypes and the genotype of other tissues that might control muscle phenotype, e.g. the motoneuron. For each of these tissues, we define the expected pattern of muscle phenotypes, and compare expected and observed patterns in each of a series of wild type/shi animals. We find that muscle phenotype is independent of the genotype of the motoneurons or muscle attachment sites. Control of the inducedshi muscle phenotype evidently lies within the muscle itself. The normalshi function, endocytosis, is therefore essential in this muscle tissue during a sensitive stage of myogenesis in early pupae (20 h).
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    Development genes and evolution 202 (1993), S. 85-94 
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    Keywords: Apterous mutation ; Corpus allatum ; Juvenile hormone ; Ring gland
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The ultrastructure of the corpus allatum of theapterous mutantsap 4 andap 56f ofDrosophila melanogaster during larval-pupal-adult metamorphosis and adult life was correlated with the gland's ability to synthesize juvenile hormone in vitro. During the early wandering period of the third instar of both mutants, a high concentration of smooth endoplasmic reticulum, mitochondria and mitochondrion-scalariform junction complexes are typical features of an active corpus allatum cell. Juvenile hormone biosynthesis by the glands is high at that time and, in fact, only slightly lower than that of wild type glands. In contrast to the wild type gland, the cells of the pupal and pharate adult corpus allatum of both mutants contains highly electron dense mitochondria with tubular cristae but no whorls of smooth endoplasmic reticulum nor glycogen clusters. The frequency and size of the lipid droplets, putatives depots of the juvenile hormone precursors, in cells of theap 56f gland is a function of the insect's age, but both are lower than in wild type gland cells. Juvenile hormone biosynthesis by both mutant glands remains at the basal level when compared to increased synthesis by the wild type gland. The frequency and density of lipid droplets in cells of theap 4 corpus allatum are much lower than in theap 56f glands. During adult life, the ultrastructural profile of theap 56f corpus allatum is similar to that of the wild type gland although the in vitro production of juvenile hormone by the former is much lower than that of the wild type gland. The ultrastructural features of the adult corpus allatum ofap 4 homozygotes reveal precocious degeneration and support the view that this non-vitellogenic mutant is a juvenile hormone deficient mutation.
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    Development genes and evolution 202 (1993), S. 129-131 
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    Development genes and evolution 202 (1993), S. 112-122 
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    Keywords: Axon guidance ; Drosophila ; Enhancer trap ; Kinesin-lacZ ; Neural development
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary We have analyzed the development of neuronal projections inDrosophila by fusing the gene encodingDrosophila kinesin, a microtubule-associated motor protein, toEscherichia coli lacZ, and employing the resulting chimeric protein as a reporter molecule for labelling cells by the “enhancer-trap” method. Expression of kinesin-β-galactosidase in neurons has afforded a detailed view of the morphologies and projections of neurons. The images of cells provided by this method will facilitate anatomical and genetic investigations of theDrosophila nervous system as well as other cell types.
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    Development genes and evolution 202 (1993), S. 103-111 
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    Keywords: Cell interactions ; Muscle cell specification ; Pigment cell specification ; Developmental autonomy ; Conditional development ; Restriction of developmental potential
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Muscle and brain pigment cell specification was studied by disrupting cell adhesion, cell dissociation, and reaggregation in embryos of the ascidianStyela clava. Treatment of embryos with Ca2+-free sea water between the 2-cell and gastrula stages disrupted blastomere adhesion but did not prevent acetylcholinesterase or muscle actin expression in presumptive muscle cells. Similar treatments initiated between the 2- and 32-cell stages caused more ectoderm cells to express tyrosinase and develop pigment granules than expected from the cell lineage. Whereas 2 pigment cells become the otolith and ocellus sensory organs in normal embryos, up to 33 pigment cells could differentiate in embryos after disruption of cell adhesion. Replacement of Ca2+-free sea water with normal sea water restored cell adhesion and usually resulted in development of embryos containing the conventional number of pigment cells. Dissociation of embryos into single cells between the 2- and 64-cell stages and culture of these cells beyond the fate restricted stage had no effect on the accumulation of muscle actin mRNA and muscle actin synthesis, but blocked pigment cell differentiation. Reaggregation of the dissociated cells did not enhance the number of cells that developed muscle features, but rescued pigment cell development. The results indicate that ascidian muscle cell specification occurs by an autonomous mechanism, whereas pigment cell specification occurs by a conditional mechanism involving cell interactions. In addition, the results suggest that negative cell interactions may restrict the potential for pigment cell development in the ectoderm of cleaving ascidian embryos.
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  • 69
    ISSN: 1432-041X
    Keywords: Transient expression ; Embryo ; Vitellophages ; Cytoplasmic actin gene ; Bombyx
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    Topics: Biology
    Notes: Summary Freshly deposited eggs ofBombyx mori were microinjected with supercoiled plasmid DNA which carried the β-galactosidase coding sequence ofEscherichia coli inserted in place of the coding sequence of theB. mori cytoplasmic actin A3 gene. Transient expression of this fusion gene in the embryo was determined by in situ histochemical detection of enzyme activity. After injection of the plasmid at different stages of embryonic development, β-galactosidase activity depending on the injected DNA was only detected in the vitellophages. This indicates the presence of active transactivators of the actin A3 gene promoter in this cell type. Tissue specificity of the fusion gene expression could be related to the early polyploidization of vitellophages, a process which would favour the stability of the nuclear pool of injected plasmids. The activity of the transgene in vitellophages was detectable at 24–33 h of egg development, the stage presumed for the onset of zygotic gene expression, up to the end of embryogenesis. This gene transfer system is thus promising to analyse thecis regulatory sequences of the actin A3 gene and could be utilized for other ubiquitous genes.
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  • 70
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    Development genes and evolution 202 (1993), S. 233-239 
    ISSN: 1432-041X
    Keywords: Mesoderm ; Growth factors ; Endoderm ; Xenopus laevis
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    Topics: Biology
    Notes: Abstract We have raised a monoclonal antibody, 4G6, against gut manually isolated from stage 42Xenopus laevis embryos. It is specific for endoderm and recognises an epitope that is first expressed at stage 19 and which persists throughout subsequent development. The antibody maintains gut specificity through metamorphosis and into adulthood. The epitope is conserved in the mouse, where it is also found in the gut. Isolated vegetal poles fromXenopus blastula stage embryos express the epitope autonomously after culturing to the appropriate stage. This shows that certain aspects of endoderm differentiation do not require germ layer interactions. Animal cap cells from stage 9 blastulae cultured in the presence of the mesodermal growth factors FGF, XTC-MIF and PIF form both endodermal and mesodermal tissues, assessed by the binding of tissue-specific monoclonal antibodies. Endoderm is typically found in those caps which form intermediate and ventral forms of mesoderm, that is muscle and lateral plate.
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  • 71
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    Development genes and evolution 202 (1993), S. 247-247 
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  • 72
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    Development genes and evolution 202 (1993), S. 240-245 
    ISSN: 1432-041X
    Keywords: Maturation ; Rearrangement ; Xenopus ; Maternal cytoplasm ; Cytoskeleton
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Using the monoclonal antibody (MoAb) Xa5B6 as probe, the authors examined the mechanisms of cytoplasmic rearrangement occurring during maturation of theXenopus oocyte. The antigen molecules recognized by the MoAb are arranged in radial striations of the oocyte cytoplasm. The radial striations were disorganized in vitro by progesterone treatment, and the antigen molecules were uniformly distributed, predominantly in the animal hemisphere. Even when the germinal vesicle was mechanically removed or when germinal vesicle breakdown was suppressed in a K+-free medium, progesterone induced a disorganization of the radial striations. This progesterone-induced disorganization was inhibited by the protein synthesis inhibitor cycloheximide. When full-sized oocytes were treated with cytochalasin B, the radial striations were also disorganized, but the antigen molecules did not disperse into the large mass. Colchicine treatment had little effect. Antigen molecules were no longer arranged in radial striations and were completely dispersed when the oocyte was simultaneously treated with both drugs. These results indicate that the two compartments in the oocyte cytoplasm, the yolk-free cytoplasm and yolk column, are organized by different types of cytoskeletal system. It is also suggested that the maturation-promoting factor (MPF) activated during progesterone-induced maturation disrupts these cytoskeletal systems and disorganizes the radial striations.
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  • 73
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    Development genes and evolution 202 (1993), S. 250-259 
    ISSN: 1432-041X
    Keywords: Zebrafish ; Neurulation ; Dye applications ; Neural anlage
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    Topics: Biology
    Notes: Abstract We have studied the process of neurulation within the anterior trunk region of the zebrafish by means of serial sectioning of staged embryos and labelling cells by applications of the dye Dil and intracellular injections of fluoresceine dextran amine. The first morphological manifestation of the prospective neural plate is a dorsomedial ectodermal thickening which becomes visible immediately after gastrulation. Within 1–2 h, by the time somatogenesis begins, two bilaterally symmetrical thickenings have appeared more laterally, which eventually fuse with the medial thickening to form the neural keel. The central canal forms next by separation of the cells on either side of the midline of the neural keel, beginning ventrally at the 17-somite stage and progressing towards dorsal levels. By means of fluorescent dye labelling in the late gastrula, we found that both the medial and lateral thickenings contribute to the nerve cord. The medial thickening was found to contain, exclusively, neural progenitor cells from the 90–100% epiboly stage on, whereas the adjacent regions contained a mixture of neural and epidermal progenitor cells, as well as prospective neural crest cells. Between the 90–100% epiboly and 2-somite stages, this heterogeneity of developmental capabilities is resolved into territories, with epidermogenic and neurogenic cells clearly separated from each other. To achieve this segregation into neural and epidermal anlagen, cells from the lateral thickenings have to move over a distance of roughly 400 μm within 1–2 h. Epidermal overgrowth of the nerve cord occurs during the morphogenetic movements that accompany nerve cord formation.
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  • 74
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    Development genes and evolution 202 (1993), S. 306-311 
    ISSN: 1432-041X
    Keywords: Gametogenesis ; Sterility ; Ovarian tumour ; Somatic ; Behaviour
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    Topics: Biology
    Notes: Abstract The 17-ethyl-methyl-sulphonate (EMS) induced female sterile alleles of the ovarian tumour (otu) locus show a wide spectrum of phenotypes and affect various processes of Drosophila oogenesis. These phenotypes have been previously studied in detail, but the exact molecular function of the otu locus in the different processes of oogenesis is only poorly known. To date, no effect of otu mutations have been reported in the males. However, separate species of otu mRNAs are expressed in the testes and the thorax of the adult male, but their role is not known. In this study we analysed the effects of EMS-induced otu mutations on male fertility. We observed that the proportion of totally sterile males is significantly higher in most of the tested otu strains as compared to the wild type. There was a strong correlation between male sterility and severity of impairment in the female phenotype. Spermatogenesis of these semi-sterile strains was analysed by phase contrast microscopy, Hoechst 33258 and Feulgen stain, and by in situ hybridisation with testis-specific probes. No changes which could account for the induction of sterility were recorded and normal amounts of motile sperm were observed in all strains. Sterility turned out to be a consequence of a failure in mating behaviour. The wild type females refused to react to the courtship attempts of the mutant males. We propose two alternative explanations for this. Either the otu locus may play some important role in male somatic tissue, or some germ line function is necessary for correct mating behaviour.
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  • 75
    ISSN: 1432-041X
    Keywords: Chimaera ; Mosaic ; Mouse ; Fetus ; Placenta
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    Topics: Biology
    Notes: Abstract Mouse chimaeras were produced by aggregating eight-cell embryos from two different F2 matings, abbreviated to AF2 and BF2 respectively: (C57BL/ OIa.AKR-Gpi-1s a, c/Ws female × BALB/c male)F2 and (C57BL/Ws female × CBA/Ca male)F2. Quantitative electrophoresis of glucose phosphate isomerase (GPI-1) was used to estimate the proportions of the two cell populations in different tissues of the 12 $${\raise0.7ex\hbox{$1$} \!\mathord{\left/ {\vphantom {1 2}}\right.\kern-\nulldelimiterspace}\!\lower0.7ex\hbox{$2$}}$$ day chimàeric conceptuses, with the % GPI-1A indicating the percentage of cells derived from the AF2 embryos. The % GPI-1A was found to be highly positively correlated within the primitive ectoderm lineage (between the fetus, amnion and yolk sac mesoderm) and within the primitive endoderm lineage (between the yolk sac endoderm and the parietal endoderm) but no correlation (either positive or negative) was seen between the two lineages. This confirms the results of a previous,study of chimaeras made between partially congenic strains and suggests the original conclusions have general validity. The % GPI-1A in the placenta was corrected for the expected contribution of maternal GPI-1, based on control experiments involving transfer of homozygous Gpi-1s b /Gpi-1s b embryos to the uteri of Gpi-1s a /Gpi-1s a pseudopregnant females. The corrected % GPI- lA in the placenta was positively correlated with that in each of the three primitive ectoderm derivatives. This suggests either (1) exchange of cells between the polar trophectoderm and the underlying part of the inner cell mass that forms the primitive ectoderm or (2) cells are incompletely mixed in the chimaeric blastocyst and patches of AF2 and BF2 cells straddle the boundary between the polar trophectoderm and the underlying primitive ectoderm. The second explanation does not imply the existence of shared developmental lineages between trophectoderm and primitive ectoderm in non-chimaeric embryos. Unlike that of any other tissue, the distribution of placental GPI-1A was U-shaped; in 17/28 placenta samples the proportion of the minor component was 10% or less. This suggests that the placental trophoblast is derived from a small number of coherenct clones of polar trophectoderm cells (either a small number of polar trophectoderm cells or a larger number if the two cell populations are not finely intermingled). Thus, although as a population the placentas of chimaeric conceptuses are balanced with respect to the % GPI-1A (mean close to 50%), individually most placentas are extremely unbalanced in their chimaeric composition (〈 10% or 〉 90% GPI-IA). This non-random composition of the chimaeric placentas is in contrast to the widely held assumption that the distribution of cells in chimaeric conceptuses is normally random.
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  • 76
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    Development genes and evolution 202 (1993), S. 316-320 
    ISSN: 1432-041X
    Keywords: Neuralizing inducing factor ; Vegetalizing inducing factor ; Activin A ; Proteoglycans ; Complex formation
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    Topics: Biology
    Notes: Abstract Proteoglycans from chicken embryos bind neuralizing and vegetalizing inducing factors. The proteoglycan-factor complexes have no inducing activity. Enzymatic cleavage of the core proteins of the proteoglycans abolishes inhibition of the inducing activity by proteoglycans. The possible significance of the formation of complexes of inducing factors with proteoglycans is discussed.
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  • 77
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    Development genes and evolution 202 (1993), S. 321-328 
    ISSN: 1432-041X
    Keywords: Acrosomal vesicles ; Fertilization ; Phallusia nigra ; Ascidiacea ; Tunicata
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    Topics: Biology
    Notes: Abstract The spermatozoa of Phallusia (Ascidia) nigra have an elongated head (approximately 5 μm in length) in which a nucleus and a single mitochondrion are located side by side. There is no midpiece. The apex of the head is wedge-shaped. Acrosomal vesicles (approximately 55–65 nm in diameter) and moderately electron-dense material (MEDM) are present between the plasmalemma and the nuclear membranes in the anterior tip of the head. The MEDM occupies a central position and three or four acrosomal vesicles are seen in a line alongside it. The acrosomal vesicles disappear as the sperm makes contact with the surface of the chorion. Gamete fusion most likely occurs between a small process extending from the peripheral margin of the sperm apex and the egg surface, resulting in incorporation of the sperm into the egg from the anterior region of its head.
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  • 78
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    Development genes and evolution 202 (1993), S. 329-340 
    ISSN: 1432-041X
    Keywords: organogenesis ; excretory organ ; nephridial development ; epithelial innervation ; immunocytochemistry ; transporting epithelium ; Hirudo medicinalis
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    Topics: Biology
    Notes: Abstract The formation of the definitive excretory system (nephridium and bladder complex) in Hirudo medicinalis during the last two thirds of embryonic development was observed with light- and electron microscopy, immunocytochemistry, and nuclear labeling. In jawed leeches, two excretory systems develop and function successively. The nephridia of the cryptolarva are associated with the larval sac and persist until the definitive nephridia are sufficiently developed to be functional. Development of the definitive excretory system begins with the differentiation of the (ectodermal) bladder and urethra. The cells from which they arise incorporate bacteria and are thereby recognizable at day 8. The (mesodermal) urine-forming tissues of the nephridium (canalicular cells and central canal cells) appear a day later. By day 17, the nephridia are in contact with the outlet region and structurally able to function. Each nephridium is individually innervated by a peripheral neuron, the nephridial nerve cell, which expresses FMR Famide-like immunoreactivity and begins growing into the nephridium on day 11. Organogenesis of the leech nephridium is compared with the formation of excretory organs in other species. The temporal correlation of innervation and the development of the transporting cells is discussed.
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  • 79
    ISSN: 1420-9098
    Keywords: Lasioglossum malachurum ; sweat bee ; Dufour's gland volatiles ; ontogenetic patterns of odorous compounds ; pheromones ; mating
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    Notes: Summary Ontogenetic patterns of volatile compounds identified in Dufour's gland extracts from queens and workers of the primitively eusocial sweat beeLasioglossum malachurum (K.) were compared. Only young unmated queens showed high proportions of isopentenyl esters, while macrocyclic lactones were dominant in old breeding queens, spring queens, and workers. In young queens the relative and absolute amounts of volatiles changed one day after mating. A discriminant analysis revealed significant differences in odor patterns of unmated and mated young queens. The fat body was the largest in young females, while eggs could be recorded only in breeding queens. Possible functions of different odor components in the investigated female groups are discussed.
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  • 80
    ISSN: 1420-9098
    Keywords: Ropalidia rufoplagiata ; Vespidae ; polygyny ; permanent reproductive division of labour ; eusociality
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    Notes: Summary Ropalidia rufoplagiata Cameron (Hymenoptera: Vespidae), a polistine species from penisular India, appears to be unique among all known primitively eusocial wasps. A total of 33 out of 46 identified females from an observed colony were found to oviposit on 1–17 occasions. No single predominant egg-layer could be identified during the 45-day period. Of the 17 dissected egg-layers, 12 were mated. All egg-layers showed several oviposition-related behavioural patterns including systematic, but indiscriminate, cannibalism of eggs and larvae, cleaning of empty cells, and guarding of freshly-laid eggs. There was no correlation between the egg-laying activity of the females (whether mated or not), oophagy, and their position in the dominance hierarchy. All nest-maintenance activities were performed exclusively by the egg-layers, while the non-egg-layers were mainly involved in the extranidal task of foraging. No significant morphometric differences between egg-layers and foragers could be discerned. Almost all the older individuals in the colony were egg-layers, while foragers were mainly younger animals. Such a temporal differentiation in reproductive labour suggests the absence of a permanent reproductive caste in this species.
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    Insectes sociaux 40 (1993), S. 87-94 
    ISSN: 1420-9098
    Keywords: Mating ; dealation ; attraction/aggregation pheromone ; queen aging ; Iridomyrmex humilis
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    Notes: Summary Queen power inIridomyrmex humilis during six arbitrarily chosen physiological stages of queens (virgin winged queens at the time of emergence, the same 4–5 days old, mated winged queens 5–6 days old, mated queens at the time of dealation, young egg-laying queens, and old egg-laying queens) was tested with regard to workers. Tested workers and queens were nestmates. The results were as follows. 1) Power of the old queens remained rather constant throughout the reproductive season. 2) Young queens were always less attractive than older ones. No changes were observed from the time of emergence to dealation. 3) At the time of egg-laying, these young queens became markedly attractive for workers but never as much as old egg-laying queens. Therefore, insemination and dealation do not trigger the increase in queen power. In contrast, oogenesis ending by egg-laying produces a significant increase in queen power.
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    Insectes sociaux 40 (1993), S. 107-109 
    ISSN: 1420-9098
    Keywords: Microstigmus ; trophallaxis ; primitively social wasp ; Sphecidae
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    Notes: Summary In a total of six hours of observation of severalMicrostigmus nigrophthalmus nests with two or more adult wasps, nine buccal contacts, six between females and three between females and males were observed. Except for two cases, these occurred soon after one of the females involved (the apparent donor) had returned to the nest and involved transfer from older to younger individuals. Experimental feeding of colored solution confirmed the occurrence of trophallaxis. InM. nigrophthalmus, trophallaxis seems to have arisen as a prolongation of maternal care since food is transferred from an older to a younger individual.
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  • 83
    ISSN: 1420-9098
    Keywords: Tetraponera ; bamboo ; ants ; nest flooding ; water removal
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    Topics: Biology
    Notes: Summary The Southeast Asian antTetraponera sp. nearattenuata F. Smith inhabits internodes of large bamboo species that it shares with symbiotic pseudococcids. It finds access to this nesting habitat via small holes made by wood-boring insects. During heavy rain, runoff water collects in these punctured internodes. TheTetraponera workers remove the water by ingesting it, walking to the entrance hole, and regurgitating it to the outside. In this way, they 1) reduce the danger of colony members drowing, 2) enable their symbionts to feed also on the internode floor, and 3) prevent excessive growth of microbes in the nest.
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  • 84
    ISSN: 1420-9098
    Keywords: Ants ; division of labor ; foraging ; behavioral flexibility ; site constancy
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    Topics: Biology
    Notes: Summary Social organization allows a division of labor between reproduction and foraging, as well as a task allocation among foraging individuals. Therefore, a colony simultaneously exploiting various resources can use different ways of getting them. This work analyzes wood ant foraging tactics both at the collective and the individual level. The main issues are: (1) How does a wood ant colony respond to stable vs temporally and spatially variable resources in terms of worker force allocation? (2) How is retrieval of ephemeral but rich resources effected at the individual level? The results show a correspondence between resource stability and behavior. Ants visited stable resources continually and recruited to ephemeral ones. They also visited empty food sites at a low frequency suggesting territoriality and constant scanning. The results suggest that recruitment may involve defence of the resource, since only a small fraction of all the workers available carry the food home. Additional ants arriving at the resource are recruited from inside the nest, not by reallocating outdoor workers. The foragers also form two separate groups, one site-persistent and another site-flexible. These two forager groups may reflect specialization on two different kinds of diet: honeydew or insect prey (spatially and temporally stable vs unstable resources, respectively).Formica truncorum is thus an example of how sociality allows the use of two different foraging strategies simultaneously: one that is based on recruitment to ephemeral resources and another that is based on search persistance and memory.
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    Insectes sociaux 40 (1993), S. 169-180 
    ISSN: 1420-9098
    Keywords: Halictidae ; sociality ; demography ; photoperiod ; temperature
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Several years' observations of population of a primitively social halictine bee,Halictus rubicundus, revealed the following: (1) there is a correlation between ambient temperatures during the spring provisioning phase and the sex ratio of the resulting brood, such that warmer temperatures are associated with an increase in male bias; (2) over the course of the season, the degree of male bias of eggs laid appears to correspond with photoperiod; and (3) increasing male bias in the first brood is associated with decreasing proportions of social colonies formed in the population, and this effect may be accentuated by small population size. These phenomena suggest that abiotic environmental conditions at the time of brood production may profoundly influence the demography of this population, and that the demography in turn determines the degree to which sociality is expressed. These findings are related to hypotheses dealing with caste determination in temperate halictine bees, viewed in the context of the evolution and maintenance of sociality, and it is proposed that these phenomena reveal a mechanism by which social behavior (the occurrence of a “worker” caste, in particular) is facultatively “fine-tuned” to suit the characteristics of the environment.
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    Insectes sociaux 40 (1993), S. 191-205 
    ISSN: 1420-9098
    Keywords: Termite ; Incisitermes ; genetics ; turnover ; caste
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Analysis of allozyme markers was carried out on 16 colonies of the termiteIncisitermes schwarzi in which one or both primary (founding) reproductives had been replaced by secondary reproductives. About one-quarter of field-collected colonies have replacement reproductives, and the genetic data suggest that in most of these a single replacement event had occurred. Genetic evidence for a second replacement event was found in one colony. Genetic analysis of the offspring allowed the following conclusions: (1) Soldiers are a relatively long-lived caste, so that even in colonies in which all other members have been replaced by offspring of the secondary reproductives, the soldiers remain the offspring of the primary reproductives; and (2) when primary reproductives are replaced by secondary reproductives, the nosoldier offspring of the former are completely replaced by those of the latter over a period of about two or three years, probably as a result of the normal maturation of colony members (workers, nymphs, and alates). The results provide no evidence for the existence of a true sterile worker caste in this species.
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  • 87
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    Insectes sociaux 40 (1993), S. 231-232 
    ISSN: 1420-9098
    Keywords: Marking ; immobilization ; ants
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    Topics: Biology
    Notes: Summary A method to immobilize ants or other small insects for marking using a vacuum pump is proposed.
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  • 88
    ISSN: 1420-9098
    Keywords: Orientation ; individual memory ; chemical communication ; Formicidae
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The relative contribution of visual and chemical components in the orientation ofLasius niger andIridomyrmex humilis (Argentine ant) workers during mass recruitment to newly discovered food sources is analyzed over short time intervals. While both species orient in response to the trail pheromone, a large number ofL. niger foragers rapidly switch to a more individual orientation, based on their memory of environmental cues.I. humilis workers, on the other hand, predominantly use collective chemical cues. The effect of the number of reinforcements on visual learning and its interference with chemical communication show that olfactory cues always prevail in the Argentine ant. InL. niger, the proportion of ants orienting to visual cues is independent of the trail concentration. Detailed observations of the trail-laying behavior of individually marked foragers show that nearly all theI. humilis workers initially lay a trail, whereas only half theL. niger foragers do so. This proportion decreases considerably with the number of trips performed byL. niger workers, while remaining constant for the Argentine ants. These results are interpreted with respect to the species' behavioral ecology.
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    Insectes sociaux 40 (1993), S. 391-402 
    ISSN: 1420-9098
    Keywords: Aphid ; defenders ; gall ; sociality ; soldiers
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    Topics: Biology
    Notes: Summary Gall-inhabiting individuals of the aphidPemphigus obesinymphae act as defenders, protecting other colony members against attack by dipteran and neuropteran larvae that are the primary predators of this species. As first instar nymphs, the progeny of the fundatrix patrol surfaces of galls and adjoining leaves. These first instar nymphs attack potential predators by mounting and grasping them and inserting their stylets. This defensive behavior, which is not exhibited by nymphs in later instars, appears to be effective in reducing predation. The fundatrix typically produces defenders throughout the extended gall-inhabiting phase, and her progeny delay development beyond the defensive first instar stage. By August, galls contain an average of 101 defenders. Early death of the fundatrix reduces the number of defenders in the gall and advances maturation of defenders into winged migrants, which otherwise mature in September and October. InPemphigus, defensive behavior by first instar nymphs appears to have evolved in the context of several types of derived life cycle, each involving an extended gall-inhabiting phase.
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  • 90
    ISSN: 1420-9098
    Keywords: Ant-repellent ; paper wasps ; latitudinal gradient ; rubbing behavior ; single foundress
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    Topics: Biology
    Notes: Summary Observations on pre-emergence, single-foundress colonies of Japanese paper wasps (Polistes) revealed that there was a latitudinal gradient in intensity of application of an ant-repellent substance (secreted by the metasomal sternum VI glands) to the nest petiole. Thus the lower the latitude, the more frequently a foundress rubbed ant repellent onto the nest petiole. Estimation of potential ant predation on wasp brood using bait traps showed that there was a positive correlation between the frequency of rubbing and potential predation pressure from ants which are guided predominantly by substrate cues for foraging. There was also a latitudinal gradient in the degree of temporal association of rubbing behavior with foraging: the lower the latitude, the more closely foundress departure from the nest was associated with rubbing. Enlargement of the nest petiole by applying oral secretion potentially obliterated previous coats of ant repellent; however, this behavior was not always followed by rubbing behavior. The ant-repellent chemical barrier around the nest petiole may have evolved in tropical regions of the world as a defense against ant predation on wasp brood. I argue that as ant predation pressure diminishes towards the cooler regions, so does selection maintaining the behavioral sequence where foundress departure from the nest is preceded by rubbing behavior.
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    Insectes sociaux 40 (1993), S. 457-457 
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  • 92
    ISSN: 1420-9098
    Keywords: Attraction ; sexual pheromone ; Formica lugubris ; ant ; mating communication
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Sexuals ofFormica lugubris fly to mating places, where females attract males by using a sex pheromone. Females collected on the nest surface before departing on a mating flight are much less attractive than those collected on the mating place after the mating flight, suggesting that the mating flight triggers the release of the sex pheromone. Olfactory cues are essential for males to locate females while they patrol. Males probably use visual cues to locate females once they have alighted nearby them. Males are also attracted by aggregations of other males on the ground, probably because one or several females are likely to be close to male aggregations.
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  • 93
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    Pure and applied geophysics 140 (1993), S. 137-137 
    ISSN: 1420-9136
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    Topics: Geosciences , Physics
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    Pure and applied geophysics 140 (1993), S. 79-94 
    ISSN: 1420-9136
    Keywords: Stick-slip ; rock friction ; instability ; precursor ; earthquake
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Stick-slips have been studied in the laboratory on granite, labrodorite and sandstone samples of two different sizes. Different roughness was achieved on the sawcut surfaces by finishing them with different grinding compounds ranging from grit 40 to grit 1000. Stick-slips occurred as a result of 1) slowly increasing the shear and normal stresses, 2) superimposing a sinusoidal stress modulation (0.1 and 10 Hz) on the slowly increasing stresses, 3) triggering by a stress impulse when the shear stress was well below the levels where stick-slips occurred without the impulse, and 4) foreshocks. Stick-slips triggered by impulses or foreshocks occurred “long” after the beginning of the triggering events, i.e., long in comparison with elastic wave travel times through the sample. All triggered events were very rich in high frequencies (corner frequency of ∼100 kHz). The untriggered stick-slips did not contain much energy at the high frequencies (corner frequency of ∼10 kHz). The dynamic friction coefficients for the triggered stick-slips were smaller than for the untriggered events. The “long” delay between the onset of the trigger and the stick-slip, and the high frequencies may be a consequence of corrosion of asperities. The ultimate triggering and the rate of corrosion are likely related to the interplay of the normal and shear stresses as they load and unload the fault surface. The consistent shape of the high frequency spectra is probably due to sample resonances which are excited rather than being characteristic of the details of the stick-slips. If these laboratory observations are directly applicable to earthquake seismology, the spectra of earthquakes which were triggered by other earthquakes should be anomalously rich in high frequencies.
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  • 95
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    Pure and applied geophysics 141 (1993), S. 445-466 
    ISSN: 1420-9136
    Keywords: Viscoeasticity ; anelasticity ; dispersion ; attenuation ; shear modulus ; creep ; forced oscillation methods
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract An apparatus is described which provides for the investigation of viscoelasticity/anelasticity in geologic and related materials under conditions of high pressure and temperature. Cylindrical specimens are tested in torsion—a geometry particularly well suited to shear mode observations at the low strain amplitudes of the linear regine. Forced oscillation experiments allow the measurement of disperision and attenuation at the low frequencies of teleseismic wave propagation. The conduct of complementary forced oscillation and creep tests allows recoverble anelastic strains to be distinguished from those of permanent viscous deformation. It has been demonstrated that robust measurements can be made at strain amplitudes below 10−5 and frequencies of 1 mHz–1 Hz, underP-T conditions to 300 MPa and 1200°C. The prospects for further development of this facility are outlined.
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  • 96
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    Pure and applied geophysics 141 (1993), S. 509-519 
    ISSN: 1420-9136
    Keywords: Stress ; pressure ; elasticity ; X-ray diffraction ; diamond anvil cell ; synchrotron radiation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Deviatoric stress in a diamond anvil cell with gold as a pressure and stress indicator is measured by two complementary techiques using synchrotron radiation. The first method employs a white X-ray beam using energy disperisive X-ray diffraction. The incident X-ray beam is parallel to the load axis and the diffraction pattern is recorded at a low two-theta angle. Using powder diffraction patterns of polycrystallin gold, we measured the elastic strain of two crystal planes oriented normal to the diffraction vector. Stresses nearly parallel and perpendicular to the load axis can be calculated by stress-strain tensor relatonship. The other method uses a monochromatic wiggler X-ray beam. In this case, the diamond cell is oriented so that the incident beam is perpendicular to the load axis. The diffraction pattern is recorded on an image plate area detector. Elastic strains responding to stresses perpendicular and parallel to the load axis can be measured and stresses of the same orientations can be calculated from the strain data. These measurements provide a lower bound of the actual differential stress in a diamond cell. With these techniques, we can measure stress distribution in a less deviatoric gasketted sample and determine yield strength of mantle materials at high pressures and temperatures.
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  • 97
    ISSN: 1420-9136
    Keywords: 6–8 double state ; synchrotron ; lower mantle condition
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A new multi-anvil type high-presure apparatus has been developed using sintered diamond anvils to generate pressures over 30 GPa and temperatures up to about 2000°C. A maximum sample volume of about 1 mm3 is available in this system. The pressure was confirmed by dissociation of forsterite into Mg-perovskite and periclase. The basic techniques and problems in utilizing sintered diamond in the MA8 type high-pressure apparatus are discussed with an emphasis on the future prospect of incorporating simultancous X-ray diffraction observation.
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  • 98
    ISSN: 1420-9136
    Keywords: High pressure ; pressure calibration ; ZnTe ; synchrotron X-ray source ; DIA type cubic anvil apparatus
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Pressure behavior of ZnTe at room temperature was studied using an X-ray energy dispersive method on a DIA type cubic anvil apparatus (SAM-85) at NSLS-X17B1. By using powdered polyethylene, the sample and NaCl for a pressure scale were held under quasihydrostatic conditions, which were confirmed by X-ray diffraction method. Two high-pressure phase transitions were confirmed using X-ray powder diffraction simultaneously with electrical resistance measurements. The phase transition pressures under quasihydrostatic conditions were determined to be 9.6 GPa, at which the resistance increased, and 12.0 GPa, which was the midpoint of a large resistance decrease. Errors in the pressure determinations were estimated to be less than 0.2 GPa. These pressure values may depend on grain size and anisotropic stress effects on the calibrant. From X-ray observation of ZnTe, the bulk modulus of the zinc blende structure was calculated to beK 0=51(3) GPa andK 0 ′ =3.6(0.8), and the first transition at 9.6 GPa was found to have about 9% volume change. It was consistent with an anomaly in the pressure generating curves.
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  • 99
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    Pure and applied geophysics 140 (1993), S. 183-210 
    ISSN: 1420-9136
    Keywords: Slip partitioning ; strain partitioning ; plate tectonics ; convergent plate boundaries
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Along plate boundaries characterized by oblique convergence, earthquake slip vectors are commonly rotated toward the normal of the trench with respect to predicted plate motion vectors. Consequently, relative plate motion along such convergent margins must be partitioned between displacements along the thrust plate interface and deformation within the forearc and back-arc regions. The deformation behind the trench may take the form of strike-slip motion, back-arc extension, or some combination of both. We observe from our analysis of the Harvard Moment Tensor Catalog that convergent arcs characterized by back-arc spreading, specifically the Marianas and New Hebrides, are characterized by a large degree of slip partitioning. However, the observed rates, directions, and location of back-arc spreading are not sufficient to account for degree of partitioning observed along the respective arcs, implying that the oblique component of subduction is also accommodated in part by shearing of the overriding plate. In the case of the Sumatran arc, where partitioning is accommodated by strike-slip faulting in the overriding plate, the degree of partitioning is similar to that observed along the Marianas, but the result is viewed with caution because it is based on a predicted plate motion vector that is based on locally derived earthquake slip vectors. In the case of the Alaskan-Aleutian arc, where back-arc spreading is also absent, the degree of partitioning is less and rotation of slip vectors toward the trench normal appears to increase linearly as a function of the obliquity of convergence. If partitioning in the Alaskan-Aleutian arc is accommodated by strike-slip faulting within the upper plate, the positive relationship between obliquity of convergence and the rotation of earthquake slip vectors to the trench normal may reflect that either (1) the ratio of the depth extent of strike-slip faults behind the trenchZ s to the subduction thrustZ t increases westward along the arc, (2) the dip of the subduction thrust increases westward along the arc, or (3) the strength of the subduction thrust decreases westward along the arc.
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  • 100
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    Pure and applied geophysics 140 (1993), S. 471-491 
    ISSN: 1420-9136
    Keywords: Sheart zone ; compressional and extensional boundary zone ; brittle-ductile transition ; viscoclastic model
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A model is proposed describing the mechanical evolution of a shear zone along compressional and extensional plate boundaries, subject to constant strain rate. The shear zones are assumed as viscoelastic with Maxwell rheology and with elastic and rheological parameters depending on temperature and petrology. Stress and strain are computed as functions of time and depth. For both kinds of boundaries the model reproduces the existence of a shallow seismogenic zone, characterized by a stress concentration. The thickness of the seismogenic layer is evaluated considering the variations of shear stress and frictional strength on faults embedded in the shear zone. Assuming that a fault dislocation takes place, the brittle-ductile transition is assumed to occur at the depth at which the time derivative of total shear stress changes from positive to negative values. The effects of different strain rates and geothermal gradients on the depth of the brittle-ductile transition are studied. The model predictions are consistent with values inferred from seismicity data of different boundary zones.
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