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  • 1990  (59,894)
  • 1
    Electronic Resource
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    Springer
    Pure and applied geophysics 132 (1990), S. 21-47 
    ISSN: 1420-9136
    Keywords: Wave scattering ; free surface ; heterogeneity ; body ; Rayleigh ; and Love waves
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The seismic coda is usually thought to be composed of waves scattered from the heterogeneities in the earth. Three classes of scattering mechanisms have been recognized: repeated specular reflection of primary waves in a uniformly layered structure, volume scattering of primary waves from localized volume heterogeneities, and scattering of primary waves from irregularities in an otherwise generally layered structure. The presence of the earth's surface complicates the description of all these scattering phenomena whenever the scattering obstacles are near or at the surface. In this paper I review work which demonstrates the effects of scattering near the earth's surface, emphasizing three general areas of investigation: scattering of body waves from an irregular free surface, scattering of body waves in irregular layers, and propagation of surface waves across irregular topography or in irregular wave guides. Most of the effects of importance have been recognized in model studies of idealized geometries. Observational evidence in support of the model studies exists, but is often inferred. Few controlled experiments to measure scattering have been performed. Topography can focus or defocus incident body waves and can convert body waves to surface waves and vice versa. Irregular surface layers can amplify incident body waves, couple body and surface waves, and produce resonances in spatially limited low velocity valley structures and highly irregular layers. Love wave propagation is highly sensitive to irregularities in a wave guide. Love wave dispersion measured over irregularly layered media can be quite different from the dispersion of the mean plane layered structure beneath the receiver array. Rayleigh wave dispersion is far less sensitive to smooth irregularities in a wave guide and is usually representative of the mean structure beneath the receiving array.
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  • 2
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    Pure and applied geophysics 132 (1990), S. 245-267 
    ISSN: 1420-9136
    Keywords: P wave amplitudes ; focussing ; three-dimensional wave propagation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Short-period teleseismicP waves from the Nevada Test Site (NTS) show systematic variations in amplitudes and travel times, with low amplitudes corresponding to fast travel times, suggesting elastic focussing-defocussing effects. Also, the azimuthal amplitude and travel time patterns for events at the Pahute Mesa subsite are systematically different from those at the Yucca Flat subsite, indicating the presence of a near-source component in both the amplitude and travel-time variations. This component is isolated by removing the mean station pattern for all of NTS from the observations. A very-near-source component in the Pahute Mesa observations is also isolated by removing subsite station means from the measurements, whereas the Yucca Flat observations exhibited no coherent very-near-source component. These anomalies are back-projected through laterally homogeneous structure to form thin lens models at various depths. Travel-time delays are predicted from the amplitude variations using the equation for wavefront curvature. The long-wavelength components of the predicted and observed time delays correlate well, at depths of 25 km for the very-near-source component under Pahute Mesa and 160 km for the regional component under NTS. The time delay surfaces predicted by the amplitudes at these depths are mapped into warped velocity discontinuities suitable for the calculation of synthetic seismograms using the Kirchhoff-Helmholtz integral formulation. Both the intersite (near-source) and intrasite (very-near-source) differences in amplitudes are qualitatively predicted very well, although the range of variation is somewhat underpredicted. This deficiency is likely due to the destructive interference of anomalies inherent in back-projection to a single layer.
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  • 3
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    Pure and applied geophysics 132 (1990), S. 363-400 
    ISSN: 1420-9136
    Keywords: Crustal scattering ; apparent attenuation ; anisotropy ; physical models
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We study wave propagation through isotropic and anisotropic scatterer distributions in order to observe azimuthal variations in velocity and apparent attenuation. Using thin aluminum plates as physical models, we obtained seismograms for compressional and shear wave propagation through heterogeneous media. Three random distributions of scatterers are studied: circular scatterers in isotropic distributions (modeling circular scatterers), elongated scatterers in isotropic distributions (modeling randomly oriented elliptical scatterers), and elongated scatterers in anisotropic distributions (modeling aligned elliptical scatterers). All scatterers had approximately the same cross-sectional area and were filled with epoxy in order to reduce the impedance contrast. In addition to seismograms recorded for no scatterers, seismograms were recorded for several scatterer volume fractions. Azimuths were measured relative to the long axis of the aligned elongated scatterers. Velocities were calculated using travel times and phase shifts at low frequencies. The velocities measured from the data were compared to simple low-frequency average-velocity theories based on thin lamellae or on distributions of penny-shaped cracks. The apparent attenuation for different scatterer distributions was computed using spectral ratios. Comparisons of the results for circular and randomly oriented elongated scatterers were made to determine the effects of scatterer shape. As expected, the circular and randomly oriented elongated scatterers showed no systematic azimuthal variation in velocity. The velocity anomalies were systematically larger for the randomly oriented elongated scatterers than for the circular scatterers. Both methods of theoretical estimation for the isotropic velocities produced velocities significantly larger than those measured. The spectral ratios showed more apparent attenuation for the randomly oriented elongated scatterers than for the circular scatterers. Comparisons of the results for the randomly oriented and aligned elongated scatterers were made to determine the effects of anisotropy in the scatterer distribution. Compressional waves for the aligned elongated scatterers with wave propagation parallel to the scatterers had larger velocities than for the aligned elongated scatterers with wave propagation perpendicular to the scatterers for all velocity calculations. Shear wave velocities were complicated by an anomalous phase change in the shear wave seismograms for azimuths less than 40° and were not as conclusive. The general trend of the theoretical velocities is similar to the velocities calculated from the data. There are, however, what appear to be significant differences. The spectral ratios showed more apparent attenuation for the randomly oriented elongated scatterers than for the aligned elongated scatterers with wave propagation parallel to the scatterers, and less attenuation than for the aligned elongated scatterers with wave propagation perpendicular to the scatterers.
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  • 4
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    Pure and applied geophysics 132 (1990), S. 457-479 
    ISSN: 1420-9136
    Keywords: Turbulence ; mesures aéroportées ; fluctuation de pression ; presso-corrélations
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Résumé Nous avons décrit dansBardeau et al. (1987), la réalisation, le fonctionnement et les tests d'un microdébitmètre basé sur l'anémométrie à fil chaud. Il a été particulièrement montré que cet appareil était bien adapté aux mesures de pression et pression différentielle avec une grande sensibilité et une très faible constante de temps. Nous présentons ici les applications qui ont été faites de ce capteur aux mesures à haute résolution à bord d'avions instrumentés pour la recherche atmosphérique. L'avion utilisé était spécialement équipé pour les mesures de turbulence et permettait donc l'acquisition d'un grand nombre de paramètres indispensables à l'analyse des données de ce capteur. Dans les applications aéronautiques qui ont été faites, le capteur a surtout été utilisé en tant que variomètre. Les comparaisons avec les mesures standards d'altitude par pression et couplages pression-données inertielles montrent que cet instrument pourrait apporter une contribution intéressante dans le domaine de la mesure à haute fréquence des fluctuations de pression.
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  • 5
    ISSN: 1420-9136
    Keywords: magnetic interpretation ; horizontal cylinders ; correlation factors ; least-squares residuals
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Procedures are formulated using the correlation factors between successive least-squares residual magnetic anomaly profiles due to long horizontal cylinders for interpreting the three principal anomalies (vertical, horizontal, and total). It is demonstrated that correlation values can be used to determine the depth to the center of the buried structure and the index parameter. Procedures are also formulated to estimate the amplitude coefficient. Two worked examples using theoretical data show the effectiveness of the present method.
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  • 6
    ISSN: 1420-9136
    Keywords: Hilbert transform ; S.P. anomalies ; convolution ; abscissae ; amplitude ; error analysis ; Cuddapah basin
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Self-potential anomalies due to two-dimensional inclined sheet of finite depth extent are analysed from their horizontal and vertical derivatives via the Hilbert transform. The depths to the top and bottom of the sheet and the inclination are obtained by means of simple mathematical expressions. The method is applied to a theoretical example and to a field S.P. anomaly over a sulphide deposit in the Kalava fault zone of Cuddapah basin, India. The presence of random noise is analysed for various levels and its effect is discussed. This interpretation process can be automated for all practical purposes by simple programming.
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  • 7
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    Pure and applied geophysics 132 (1990), S. 151-173 
    ISSN: 1420-9136
    Keywords: Scattering ; wave propagation ; seismic waves ; core-mantle topography ; inhomogeneities
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Statistical properties of small-scale inhomogeneities (wavelengths between 20 and 70 km) near the core-mantle boundary are inferred from scattered core waves. Observations of scattered core waves at large seismic arrays and worldwide networks indicate that the inhomogeneities have a global nature with similar characteristics. However, there may exist a few regions having markedly stronger or weaker strengths. Scattering by volumetric inhomogeneities of about 1% inP-wave velocity in the lower mantle or by about 300 m of topographic relief of the core-mantle boundary can explain the observations. At present it is not possible to rule out either of these two alternatives, or a combination of both.
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  • 8
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    Pure and applied geophysics 132 (1990), S. 175-196 
    ISSN: 1420-9136
    Keywords: Wave propagation ; seismic waves ; heterogeneities ; lithosphere
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Adopting the spectral approach, we derive the formulation of angular coherence and transverse coherence of transmission fluctuations. Our derivation and results provide new insight on transmission fluctuation analysis. A review of research work on fluctuation analysis using observations at large seismic arrays such as LASA and NORSAR-follows. We point out that the model of a single-layer Gaussian medium cannot explain the angular coherence of NORSAR data and a more general model of a non-Gaussian, multi-scale, vertically inhomogeneous random media is needed. The model of a two-layer power-law medium proposed by Flatté and Wu is among the simplest of such models.
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  • 9
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    Pure and applied geophysics 132 (1990), S. 495-504 
    ISSN: 1420-9136
    Keywords: Radon ; S.S.N.T.D. ; geochemical tracer ; fault ; atmospheric parameters ; enhancement technique
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The existence of a suspected geological fault has been confirmed using Solid State Nuclear Track Detectors (SSNTDs) by measuring radon concentration variations in the upper soil above its inferred position. The results obtained prompted us to increase the natural radon signal in the soil, using an additional radon source; this “enhancement technique,” has been experimentally checked with SSNTD detectors.
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  • 10
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    Pure and applied geophysics 132 (1990), S. 583-597 
    ISSN: 1420-9136
    Keywords: Reflection ; seismology ; conservation ; inversion
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We show that the time-dependent wave equation in both one and two spatial dimensions possesses quantities which are globally conserved. We show how these conserved quantities can be used to determine the characteristic impedance, the rock density and the elastic constant of the rock. We also demonstrate that the conserved quantities possess the capability of determining and/or bracketing the unknown component of the direct pressure response, which is required to begin downward continuation algorithms. Further, we demonstrate that the conserved quantities are always available irrespective of the source structure in time. Numerical instability, arising if the “filtering” due to the source structure is too harsh, can then be controlled to a degree by demanding that the conserved quantities be indeed conserved.
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  • 11
    ISSN: 1420-9136
    Source: Springer Online Journal Archives 1860-2000
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  • 12
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    Pure and applied geophysics 132 (1990), S. 533-543 
    ISSN: 1420-9136
    Keywords: Computer ; earthquake ; Hungary ; intensity ; macroseismic ; MSK scale ; questionnaire
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In this paper a computerized method for the intensity estimation from macroseismic observations is presented. The successive steps of the MSK intensity scale (2° to 10°) receives weights according to the observations marked on the earthquake questionnaires. These weights are summed for the individual scale steps. The distribution of these summed weights and their relation to an experimentally chosen function of the criterion make it possible to estimate the appropriate intensity value and its reliability. By computer evaluation of earthquake reports an objective comparison of estimated intensities given by different seismologists can be made, provided that uniform earthquake questionnaires are used. The application of this method will result in more homogeneous intensity data.
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  • 13
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    Pure and applied geophysics 132 (1990), S. 545-568 
    ISSN: 1420-9136
    Keywords: Cracks ; scattering ; attenuation ; phase velocity ; SH waves
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The effect of randomly distributed cracks on the attenuation and dispersion ofSH waves is theoretically studied. If earthquake ruptures are caused by sudden coalescence of preexisting cracks, it will be crucial for earthquake prediction to monitor the temporal variation of the crack distribution. Our aim is to investigate how the property of crack distribution is reflected in the attenuation and dispersion of elastic waves. We introduce the stochastic property, in the mathematical analysis, for the distributions of crack location, crack size and crack orientation. The crack size distribution is assumed to be described by a power law probability density (p(a) ∞ a −γ fora min≤a≤a max according to recent seismological and experimental knowledge, wherea is a half crack length and the range 1≤γ≤3 is assumed. The distribution of crack location is assumed to be homogeneous for the sake of mathematical simplicity, and a low crack density is assumed. The stochastic property of each crack is assumed to be independent of that of the other cracks. We assume two models, that is, the aligned crack model and the randomly oriented crack model, for the distribution of crack orientation. All cracks are assumed to be aligned in the former model. The orientation of each crack is assumed to be random in the latter model, and the homogeneous distribution is assumed for the crack orientation. The idea of the mean wave formalism is employed in the analysis, and Foldy's approximation is assumed. We observe the following features common to both the aligned crack model and the randomly oriented crack model. The attenuation coefficientQ −1 decays in proportion tok −1 in the high frequency range and its growth is proportional tok 2 in the low frequency range, wherek is the intrinsic wave number. This asymptotic behavior is parameter-independent, too. The attenuation coefficientQ −1 has a broader peak as γ increases and/ora min/a max decreases. The nondimensional peak wave numberk p a max at whichQ −1 takes the peak value is almost independent ofa min/a max for γ=1 and 2 while it considerably depends ona min/a max for γ=3. The phase velocity is almost independent ofk in the rangeka max〈1 and increases monotonically ask increases in the rangeka max〉1. While the magnitude ofQ −1 and the phase velocity considerably depend on the orientation of the crack in the aligned crack model, the above feature does not depend on the crack orientation. The accumulation of seismological measurements suggests thatQ −1 ofS waves has a peak at around 0.5 Hz. If this observation is combined with our theoretical results onk p a max, the probable range ofa max of the crack distribution in the earth can be estimated for γ=1 or 2. If we assume 4 km/sec as theS wave velocity of the matrix medium,a max is estimated to range from 2 to 5 km. We cannot estimatea max in a narrow range for γ=3.
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  • 14
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    Pure and applied geophysics 132 (1990), S. 719-731 
    ISSN: 1420-9136
    Keywords: Magnetic interpretation ; pair of sheets ; equivalence to dykes
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Magnetic anomaly profiles over two thin sheets separated by a small distance resemble those of dykes andvice versa. Interpretation of anomalies over a pair of sheets based on the magnetic properties of dykes predicts a dyke whose centre lies midway between the positions of the sheets. The dyke, on the other hand, is magnetically equivalent to a pair of sheets, both lying at the same depth and having the same magnetization. The magnetic anomalies due to a pair of sheets can be interpreted by framing linear equations between the anomalies and their distances measured from an arbitrary reference. Application of this method to anomalies of dipping sheets with a finite depth extent is indicated.
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  • 15
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    Pure and applied geophysics 132 (1990), S. 811-824 
    ISSN: 1420-9136
    Source: Springer Online Journal Archives 1860-2000
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  • 16
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    Pure and applied geophysics 133 (1990), S. 23-52 
    ISSN: 1420-9136
    Keywords: CodaQ −1 ; scattering ; mainshock area
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A temporal and spatial change of codaQ −1 associated with the occurrence of the North Palm Springs earthquake of July 8, 1986 was studied by using 242 small local earthquakes in the vicinity of the mainshock. We found that the codaQ −1 of earthquakes which occurred before the mainshock was significantly higher than that of the aftershocks in the mainshock area while the codaQ −1 for the surrounding area remained almost constant throughout 1986. CodaQ −1 was determined separately for the lapse time windows of 10 to 20 sec. and 15 to 40 sec. for the period from 1981 to 1987. The result for the time window 10 to 20 sec. showed a peak in codaQ −1 before the time of mainshock at all frequencies. The peak appeared earlier at lower frequencies. There was no significant change in codaQ −1 for the time window 15 to 40 sec., probably because the change was restricted to a small area.
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  • 17
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    Pure and applied geophysics 133 (1990), S. 103-115 
    ISSN: 1420-9136
    Keywords: Reflection travel time ; interval velocities ; layer thickness
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In reflection surveys and velocity analysis, calculations of interval velocities and layer-thicknesses of a multilayered horizontal structure are often based on Dix's equation which requires the travel times at zero offsets and a prior estimate of the root mean squared velocities. In this paper a method is presented which requires only the reflection travel-time data. A set of equations are derived which relate the interval velocity and thickness of a layer to the reflection travel time from the top and the bottom of that layer, the offset distances and the ray parameter. It is shown that the difference of the offset distances and the difference of the picked travel times of any reflected rays with the same value of ray parameter from the top and the bottom of a horizontal layer can be used to calculate the interval velocity and thickness of that layer.
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  • 18
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    Pure and applied geophysics 133 (1990), S. 213-227 
    ISSN: 1420-9136
    Keywords: Indoor radon ; soil radon ; aeroradioactivity ; uranium
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Attempts to predict which geographic areas should be associated with a high percentage of homes with unusually high indoor radon levels in Virginia and Maryland have been based on estimates of soil radon and soil permeability for geological units. This method is found to be less successful and probably less cost-effective than the use of total-gamma aeroradioactivity maps.
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  • 19
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    Pure and applied geophysics 133 (1990), S. 429-446 
    ISSN: 1420-9136
    Keywords: Attenuation ; maximum entropy ; surface waves
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract A new technique for calculating interstation Green's functions and attenuation coefficients for seismic surface waves is presented. The interstation Green's function is evaluated from the autocorrelation functions of the seismograms, which are obtained from a maximum entropy process. Since a data-invariant time window is not used, the evaluated Green's functions gives reliable information on both the amplitude and the phase spectra of the system. This new technique is compared with other methods by applying them to both synthetic and real data from a path in the Canadian shield.
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  • 20
    ISSN: 1420-9136
    Keywords: Surface waves ; transversely isotropic ; liquid saturated porous solid ; dispersion ; Rayleigh type waves ; nondissipative porous media ; frequency equation ; phase velocity
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Dispersion of Rayleigh-type surface wave is studied in a homogeneous transversely isotropic elastic layer overlying a nondissipative liquid-saturated porous solid half-space and lying under a uniform layer of homogeneous liquid. The frequency equation in the form of ninth-order determinant is obtained. Special cases have been deduced by reducing the depth of the layers to zero and by changing the transverse isotropic layer to an isotropic layer. Dispersion curves for the phase velocity have been plotted for a particular model.
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  • 21
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    Pure and applied geophysics 133 (1990), S. 691-712 
    ISSN: 1420-9136
    Keywords: Rossby waves ; ray theory ; oceanic mixed layer
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Rossby wave propagation in the presence of a nonseparable Brunt-Väisälä frequency,N(y,z), and the associated geostrophic zonal flow,U(y,z), is examined in this paper. The usual quasi-geostrophic potential vorticity equation only includes vertical variations in Brunt-Väisälä frequency (i.e.N(z)). We derive a linearised quasi-geostrophic potential vorticity equation which explicitly includesN(y, z), where variations inN may occur on the internal Rossby radius length scale. A mixed layer distribution that monotonically deepens in the poleward direction leads to a nonseparableN(y,z). The resulting meridional pressure gradient is balanced by an eastward zonal geostrophic flow. By assuming mixed layer depth changes occur slowly, relative to a typical horizontal wavelength of a Rossby wave, a local analysis is presented. The Rossby wave is found to have a strongly modulated meridional wavenumber,l, with amplitude proportional to |l|−1/2. To elucidate whether the modulations of the Rossby wave are caused by the horizontal variations inN orU we also consider the cases where eitherN orU vary horizontally. Mixed layer depth changes lead to largestl where the mixed layer is deepest, whereasl is reduced in magnitude whereU is nonzero. When bothU(y,z) andN(y,z) are present, the two effects compete with one another, the outcome determined by the size of |c|/U max, wherec is the Rossby wave phase speed. Finally, the slowly varying assumption required for the analytical approach is removed by employing a numerical model. The numerical model is suitable for studying Rossby wave propagation in a rectangular zonal channel with generalN(y, z) andU(y, z).
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  • 22
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    Pure and applied geophysics 133 (1990), S. 733-748 
    ISSN: 1420-9136
    Keywords: Nonlinear coherent structures ; modons ; IG eddies ; local multiple equilibria ; long Rossby waves ; potential vorticity
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract It is demonstrated that nonlinear Rossby modes, such as modons and IG eddies, can be excited in planetary fluids by a sufficiently strong forcing of potential vorticity. When a weak forcing is balanced with a weak dissipation, two (linear and nonlinear) equilibrium states can be produced, depending on the initial condition. When the fluid is inviscid, a sufficiently strong steady forcing may generate a sequence of propagating nonlinear eddies. A weak forcing, by contrast, only generates linear Rossby waves. The criterion which divides the high amplitude nonlinear state and the low amplitude linear state may be interpreted in terms of a ratio of a time necessary to force the eddy to a time for a fluid particle to circulate about the nonlinear eddy once.
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  • 23
    ISSN: 1420-9136
    Keywords: Rock failure ; fractal ; percolation ; rock modulus ; damage parameter ; rock mechanics
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The elastic properties of a physical model representing a damaged rock matrix were studied using a square lattice deformed under tensile stress. The elastic modulusM of such a system varies in agreement with percolation theory as|x−x c | f , wherex is the damage parameter andx c the threshold value of the damage parameter,f≃3.6. Atx≃x c the scale dependence ofM can be expressed asM∼L −f/v , whereL is the size of the sample andv the correlation exponent in percolation theory. The experimental results are of interest in assessing elastic properties in earthquake focal zones and fault zones in general.
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  • 24
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    Pure and applied geophysics 134 (1990), S. 117-133 
    ISSN: 1420-9136
    Keywords: Seismic refraction ; travel times and waveforms joint inversion ; annealing algorithms
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract An algorithm of annealing is applied to a joint inversion of travel times and waveforms belonging to a synthetically generated seismic refraction experiment. The medium (crust and upper mantle) is modelled by a set of plane stratified layers and a halfspace. The obtained structure (elastic parameters and depth of layers) shows that, in spite of contaminating seismic noise and poor knowledge of the seismic source, annealing methods are a good tool in these kinds of inversion problems. We think that many characteristics of the annealing process described here could be used with real data and more sophisticated media for the crust and upper mantle of the earth than the present example.
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  • 25
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    Pure and applied geophysics 134 (1990), S. 195-228 
    ISSN: 1420-9136
    Keywords: Earthquakes ; coda ; source attenuation ; temporal changes ; prediction site effects
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We analyse spectral ratio of the coda of doublets of microearthquakes. Our purpose is to find evidence for temporal changes of the attenuation in the crust before a large magnitude earthquake. A Moving Window Cross Spectral analysis of the coda of doublets gives a plot of the spectral ratio as a function of lapse time along the seismogram, for several frequency bands (SR(T, f) plot). From a certain pattern in theSR(T, f) plot, we should infer a temporal change in coda attenuation. Several doublets recorded in Central California by the USGS network are analysed. Using events very close in time from one another, we show that the radiation pattern can be different enough to induce important variations in the spectral ratio of the first arrivals and of the coda. Another doublet exhibits a strong variation of the low frequencies for stations in the region of Hollister (California), wherePhillips andAki (1986) have noted a strong amplification of low frequencies, that they attribute to site effects on unconsolidated sediments in the fault zone. These variations could be related to slight changes in local conditions (creep, or water table). On the other hand, some doublets, in the vicinity and close in time to the August 1979 Coyote Lake Earthquake (M=5.9), show no variation inSR(T, f) related to this earthquake: this proves that there was no major change in attenuation in the crust preceding this large shock. If a change occurred, it should have been confined to a very limited region, which was not sampled by the many paths we studied. The employed method probably provides today the most accurate estimation of spectral ratio in the coda of microearthquakes. It shows that there are numerous kinds of variations and that it is not straightforward to relate them to coda attenuation changes only. Also, we must be circumspect when dealing with the coda of microearthquakes in relation to forecasting earthquakes.
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  • 26
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    Pure and applied geophysics 134 (1990), S. 303-311 
    ISSN: 1420-9136
    Keywords: Maximum entropy ; autoregressive ; spectral analysis ; harmonic function
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract The maximum entropy (ME) spectrum, or its equivalent form of the autoregressive (AR) spectrum, has been used as a tool for harmonic analysis of time series in geophysics. This paper critically examines its usage in estimating the amplitude and the exponential decay rate of a harmonic function. The argument is based upon Prony's relation, which relates a complex-conjugate pair of poles for the AR model of the time series on one hand, to the complex frequency of one harmonic component in the time series on the other. It is found that: (i) the ME spectrum can be used as an estimator for the decay rate in a way similar to the Fourier spectral analysis; (ii) the ME spectrum contains no information whatsoever about the amplitude, contrary to what has been claimed and practiced in geophysical applications.
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  • 27
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    Pure and applied geophysics 132 (1990), S. 439-455 
    ISSN: 1420-9136
    Keywords: Temperature inversion ; katabatic wind ; Antarctica ; climate
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    Topics: Geosciences , Physics
    Notes: Abstract Intensive meteorological observations were carried out at three sites along the slope of Adélie Land, Antarctica, with the goal of documenting summertime meteorological features of the katabatic zone. Three observational sites were placed at 5 km (D10), 110 km (D47), and 210 km (D57) from the coast, and frequency distributions of meteorological parameters were obtained at each site. Some meteorological features at D47 and D57 (interior sites) were different than at D10 (the coastal site); namely, mean air temperatures averaged for the whole observational period showed more pronounced diurnal variations at the interior sites than at the coastal site. These variations could be represented by sinusoidal curves. Wind speeds also showed diurnal variations, whose extremes appeared several hours later than the temperature extremes. The prevailing wind direction for all sites was southerly, about 30° east of the fall line, with small variations in wind direction. The strength and the height of temperature inversions were examined in two different layers (0 to 50 m, 50 to 300 m) at D47. A strong, shallow inversion layer formed at night, however, completely vanished during the day. On the other hand, an upper level inversion whose strength was weak persisted. The katabatic characteristics of the surface wind found at D47 in summer could be due to this upper level temperature inversion.
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  • 28
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    Pure and applied geophysics 132 (1990), S. 401-415 
    ISSN: 1420-9136
    Keywords: Complex rays ; viscoelastic waves ; anelasticity ; attenuation ; Fermat's principle ; synthetic seismograms ; ray tracing
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In order to trace a ray between known source and receiver locations in a perfectly elastic medium, the take-off angle must be determined, or equialently, the ray parameter. In a viscoelastic medium, the initial value of a second angle, the attenuation angle (the angle between the normal to the plane wavefront and the direction of maximum attenuation), must also be determined. There seems to be no agreement in the literature as to how this should be done. In computing anelastic synthetic seismograms, some authors have simply chosen arbitrary numerical values for the initial attenuation angle, resulting in different raypaths for different choices. There exists, however, a procedure in which the arbitrariness is not present, i.e., in which the raypath is uniquely determined. It consists of computing the value of the anelastic ray parameter for which the phase function is stationary (Fermat's principle). This unique value of the ray parameter gives unique values for the take-off and attenuation angles. The coordinates of points on these stationary raypaths are complex numbers. Such rays are known as complex rays. They have been used to study electromagnetic wave propagation in lossy media. However, ray-synthetic seismograms can be computed by this procedure without concern for the details of complex raypath coordinates. To clarify the nature of complex rays, we study two examples involving a ray passing through a vertically inhomogeneous medium. In the first example, the medium consists of a sequence of discrete homogeneous layers. We find that the coordinates of points on the ray are generally complex (other than the source and receiver points which are usually assumed to lie in real space), except for a ray which is symmetric about an axis down its center, in which case the center point of the ray lies in real space. In the second example, the velocity varies continuously and linearly with depth. We show that, in geneneral, the turning point of the ray lies in complex space (unlike the symmetric ray in the discrete layer case), except if the ratio of the velocity gradient to the complex frequency-dependent velocity at the surface is a real number. We also present a numerical example which demonstrates that the differences between parameters, such as arrival time and raypath angles, for the stationary ray and for rays computed by the above-mentioned arbitrary approaches can be substantial.
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  • 29
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    Pure and applied geophysics 132 (1990), S. 505-520 
    ISSN: 1420-9136
    Keywords: Earthquake ; radon-222 ; soil measurements ; SSNTD technique ; forerunner signal
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    Topics: Geosciences , Physics
    Notes: Abstract A radon concentration measurement survey was carried out in Ecuador using the SSNTD technique. The data analysis shows a definite radon anomaly prior to March 6, 1987;M=6.9 earthquake. Anomalies found before October 18 and October 28th mild earthquakes in 1986 could be correlated with these events. Other anomalies that we cannot correlate with any geophysical event were also found.
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  • 30
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    Pure and applied geophysics 134 (1990), S. 93-108 
    ISSN: 1420-9136
    Keywords: Microearthquake ; Wadati plot ; station-by-station method ; velocity-ratio ; upper crust ; lower crust ; precursor
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    Topics: Geosciences , Physics
    Notes: Abstract The upper crustal (≤20 km)P-wave velocity beneath the Shillong Plateau and Nowgong area has been studied by the time-distance plot method. TheP-arrival data of the shallow (≤20 km) microearthquakes from three temporary networks are used, and the average velocity is found to be 5.55 km/s. The velocity ratio (V p /V s ) for the upper crust (0–20 km) as well as for the lower crust (21–40 km) are determined by the Wadati-plot method and station-by-station method. The average value obtained by the two methods is compatible; theV p /V s ranges between 1.74 to 1.76. A generalized seismic velocity model of the area is suggested by this study, which has been very useful for microearthquake location.
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  • 31
    ISSN: 1420-9136
    Keywords: Earthquake precursors ; clustering ; quiescence ; earthquake statistics ; seismicity
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    Topics: Geosciences , Physics
    Notes: Abstract This paper describes a new method, single-link cluster analysis (SLC), to evaluate percursory quiescence for shallow earthquakes in sixteen subduction zones, using data from the ISC catalog. To define quiescent regions, we divided the catalog into time intervals with a durationT, overlapping byT/2. We considered all earthquakes having magnitudes larger than some magnitudeM min, lying within a specified distance of a great circle which is approximately coincident with the trench near a subduction zone. Within each time interval we connected or ‘linked’ all earthquakes lying within some cutoff distanced of one another. We then projected all these links onto the great circle, and defined a region to be quiescent if it was not covered by the projection of any links. For this study,T was two years,M min wasm b =4.9, and we variedd from 100 to 400 km. We defined an earthquake as ‘following quiescence’ if it occurred within two years following, and within 75 km of a quiescent zone as defined above. The primary conclusion of this study was that earthquakes with surface wave magnitudes 7.2 and greater were about 5–15% more likely to follow quiescence than were the smaller background earthquakes withm b 〉-4.9. A chi-squared analysis shows that this result is significant at the 99% level. In contrast, earthquakes with surface wave magnitude of 6.7 to 7.1 were no more likely to follow quiescence than were background earthquakes. Of sixteen individual regions, Central America, Japan, and Peru-Chile were the only regions where large earthquakes were more likely to occur following quiescence than were background earthquakes. For a cutoff link length of 300 km, only in Central America was the difference between large earthquakes and background earthquakes significant at the 95% level of significance. For a cutoff link length of 250 km, the significance level exceeded 95% only in Japan. The SLC method is an objective, quantitative method for evaluating large data catalogs, or for monitoring quiescence in regions where quiescence is conjectured to precede large earthquakes.
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  • 32
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    Pure and applied geophysics 134 (1990), S. 109-115 
    ISSN: 1420-9136
    Keywords: Wide-angle reflections ; planar shift
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    Topics: Geosciences , Physics
    Notes: Abstract At angles other than normal incidence a pencil of plane acoustic waves, incident on a planar interface separating a low velocity medium from a high velocity medium, gives rise to a reflected pencil of waves which is laterally displaced from the illumination point of the incident pencil. The importance of this planar shift for understanding and interpreting wide-angle acoustic reflections is discussed, as is the variation of the equivalent depth below the interface at which a “point-like” reflection would be considered as an equivalent reflecting horizon.
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  • 33
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    Pure and applied geophysics 134 (1990), S. 163-174 
    ISSN: 1420-9136
    Keywords: Tsunami warnings ; automation ; tsunamis ; earthquake processing ; natural hazard ; ATWC
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    Topics: Geosciences , Physics
    Notes: Abstract The Alaska Tsunami Warning Center has the responsibility of providing timely tsunami warning services for Alaska and the west coasts of Canada and the United States. Recently, the ATWC implemented a new microcomputer system which is used for both automatic and interactive earthquake processing, and for disseminating critical information to the Tsunami Warning System recipients. Real-time seismic wave form data from 23 short-period and 9 long-period sites in Alaska, the lower 48 States, and Hawaii, are continually computer-monitored for the occurrence of an earthquake. Once detected from the short-period wave form data, pre- and post-earthquake data are displayed on a graphics terminal along with an indicator to identify the time of the onset of theP waves (P-picks). TheP-picks can easily be changed during or after data collection via a mouse. Magnitudes (M b ,M l ,M B ,M S ) are automatically computed from appropriate short- and long-period wave form data concurrently with the above processing. A second graphics terminal displays cycle-by-cycle long-period wave form data that was used to compute an earthquake'sM B andM S magnitudes. An earthquake's parametric data and other information are available and printed within tens of seconds after theP wave arrivals are recorded at the first 5 sites, then 7 sites, 9 sites, and a final parametric computation using all collected data. Three video display monitors are used for displaying the parameters, procedural aids, and a map showing the epicenter. Additionally, selected event parameters are immediately transmitted by VHF radio to alphanumeric beepers which are carried by standby duty personnel during those times that the Center is not manned. Using a dedicated video display terminal and printer, the interactive system can use data and parameters resulting from the automatic processes for concurrent parameter recomputations; perform additional computations; disseminate critical information; and generate procedural aids for duty geophysicists to facilitate an earthquake/tsunami investigation.
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  • 34
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    Pure and applied geophysics 134 (1990), S. 261-282 
    ISSN: 1420-9136
    Keywords: Greece ; earthquake ; intermediate prediction algorithm M8 ; TIP
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    Notes: Abstract The 3 strongest earthquakes,M≥7.0, which have occurred since 1973 in the area of Greece were preceded by a specific increase of the earthquake activity in the lower magnitude range. This activation is depicted by algorithm M8. This algorithm of intermediate term earthquake prediction was originally designed for diagnosis by Times of Increased Probability (TIPs) of the strongest earthquake,M≥8.0 worldwide (Keilis-Borok andKossobokov, 1984). At present the algorithm is retrospectively tested for smaller magnitudes in different seismic regions (Keilis-Borok andKossobokov, 1986, 1988). A TIP refers to a time period of 5 years and an area whose linear size is proportional and several times larger than that of the incipient earthquake source. Altogether the TIPs diagnosed by the algorithm M8 in the area of Greece occupy less than 20% and the Times of Expectation (TEs) about 10% of the total space-time domain considered. Also there is a current TIP for the southeastern Aegean sea and 1988–1992. It may specify the long-term prediction given inWyss andBaer (1981a,b). The results of this study are further evidence favoring applicability of algorithm M8 in diverse seismotectonic environment and magnitude ranges and support indirectly the hypothesis of self-similarity of the earthquake activity. It also implies the possibility of intermediate term prediction of the strongest earthquakes in the area of Greece.
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  • 35
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    Pure and applied geophysics 134 (1990), S. 333-354 
    ISSN: 1420-9136
    Keywords: Mantle magnitude ; Rayleigh waves ; deep sources
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    Topics: Geosciences , Physics
    Notes: Abstract We extend to the case of intermediate and deep earthquakes the mantle magnitude developed for shallow shocks byokal andTalandier (1989). Specifically, from the measurement of the spectral amplitude of Rayleigh waves at a single station, we obtain a mantle magnitude,M m, theoretically related to the seismic moment of the event through $$M_m = \log _{10} M_0 - 20.$$ The computation ofM minvolves two corrections. The distance correction is the same as for shallow shocks. For the purpose of computing the frequency-dependent source correction, we define three depth windows: Intermediate (A) (75 to 200 km); Intermediate (B) (200–400 km) and Deep (over 400 km). In each window, the source correctionC S is modeled by a cubic spline of log10 T. Analysis of a dataset of 200 measurements (mostly from GEOSCOPE stations) shows that the seismic moment of the earthquakes is recovered with a standard deviation of 0.23 units of magnitude, and a mean bias of only 0.14 unit. These figures are basically similar to those for shallow events. Our method successfully recognizes truly large deep events, such as the 1970 Colombia shock, and errors due to the potential misclassification of events into the wrong depth window are minimal.
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  • 36
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    Pure and applied geophysics 133 (1990), S. 349-365 
    ISSN: 1420-9136
    Keywords: Cascadia ; subduction ; receiver functions ; P-waves ; converted phases ; seismic structure
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    Topics: Geosciences , Physics
    Notes: Abstract The local subduction geometry at a site south of Puget Sound in western Washington is investigated using teleseismicP-waveforms recorded on a three-component event triggered seismograph. The data are processed using source equalization deconvolution in order to isolate locally convertedP-to-S arrivals and stacked to improve the signal-to-noise ratio. Stable arrivals in the radial component indicate an oceanic Moho within the subducted slab at a depth of about 53 km beneath the station. Observed amplitude variations with azimuth in the radial data, as well as qualitative aspects of the tangential data, are used to establish a slab dip of 16° to the southeast. Our results are compatible with previous results from a site 60 km to the west, and further confirm a substantial warp in the regional geometry of the subducted Juan de Fuca plate.
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  • 37
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    Pure and applied geophysics 133 (1990), S. 475-487 
    ISSN: 1420-9136
    Keywords: Earthquake warning ; earthquake alarm ; expert system ; San Andreas
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    Topics: Geosciences , Physics
    Notes: Abstract A prototype expert system has been developed to provide rapid warning of earthquakes while they are occurring. Warning times of up to 100 seconds will be possible. In the complete system, several accelerometers are distributed at intervals within a few kilometers of a known fault; data are telemetered to a central computer which implements the expert system. The expert system incorporates specific information about the type of fault to be monitored, and includes simple rules for estimating the fault slip, rupture length, and seismic moment, all in real time. If the seismic moment exceeds a preset value, an alarm may be issued. The prototype is designed for deployment on near-surface strike-slip faults such as the San Andreas and has been successfully tested with data from the 1979 Imperial Valley and 1984 Morgan Hill earthquakes. Crucial concepts have also been tested using synthetic data calculated for a model of the 1857 Fort Tejon earthquake. Parkfield, California, could be used as a test site.
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  • 38
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    Pure and applied geophysics 133 (1990), S. 547-551 
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  • 39
    ISSN: 1432-041X
    Keywords: Inner cell mass ; Blastocyst ; Morula ; Implantation ; Rat
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    Topics: Biology
    Notes: Summary The number of trophectoderm (TE) and inner cell mass (ICM) cells was determined by complementmediated lysis and differential staining in rat embryos collected at different times during in vivo preimplantation development. At 90 h after fertilization, two groups of morulae were discriminated according to the presence or absence of detectable ICM cells, and the analysis of their total cell number indicated that acquisition of a permeability seal between TE cells begins at the 14-cell stage. On the other hand, our data confirmed that blastocoele formation occurs after the fourth cleavage division in the rat. The total cell number increased exponentially with time in blastocysts recovered between 90 h and 127 h but the cell kinetics of TE and ICM cells were different. The proportion of ICM cells consequently varied throughout blastocyst development, with a peak value for expanded blastocysts at 103 h. Finally, a linear-quadratic relationship was found between the numbers of TE and ICM cells when all the embryos with a detectable ICM were analysed together.
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  • 40
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    Development genes and evolution 198 (1990), S. 295-302 
    ISSN: 1432-041X
    Keywords: Drosophila ; Oogenesis ; Embryogenesis ; Ecdysteroids ; Localized factors
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    Topics: Biology
    Notes: Summary We have produced monoclonal and polyclonal antibodies against an antigen that is asymmetrically distributed in mature oocytes of Drosophila melanogaster. During late oogenesis and early embryogenesis the antigen undergoes dramatic changes in its cellular localization: until about 2.5 h before completion of oogenesis it is homogeneously distributed in the cytoplasm, then it becomes localized in granules that are more numerous in posterior than in anterior peripheral positions of the ooplasm. The germ plasm is void of the antigen. Shortly after egg deposition the antigen is released from the granules and forms a shallow temporary gradient in the egg. Later during embryogenesis the antigen is associated with the yolk-containing cytoplasm. At the syncytial blastoderm stage it is also detected in the peripheral nuclei. Preliminary evidence suggests that the antigen is an ecdysteroid-related molecule. Five different anti-ecdysone antisera were found to bind to the same antigen or to an antigen with the same localization as our monoclonal antibody. In pattern mutants affecting anteroposterior polarity, the described asymmetrical distribution of the antigen is abnormal. In the mutant BicD, for example, which leads to the formation of two abdomina of opposite polarity, the antigen-containing granules are distributed homogeneously in mature oocytes.
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  • 41
    ISSN: 1432-041X
    Keywords: Amphibia ; Cleavage cycle ; Epiboly ; Time-lapse photography
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    Topics: Biology
    Notes: Summary Measurements were made of the external surface areas occupied by animal and vegetal blastomeres and their daughter cells at successive cleavage cycles in 15 embyros of Xenopus laevis. On the animal side, after each cleavage a general area increase (epiboly) occurs from cycle 4 (16-cell stage, stage 5) to cycle 10 (stage 8 1/2), while on the vegetal side there is a slight general area decrease after each cleavage from cycle 6 to cycle 10. The comparison between the external surface areas of individual animal blastomeres and those of their daughter cells, visible at the next cycle, shows a significantly larger increase after radial than after tangential cleavage, a difference that may be connected with the insertion of new membrane into the external surface at radial cleavage.
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  • 42
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    Development genes and evolution 198 (1990), S. 309-321 
    ISSN: 1432-041X
    Keywords: Development ; Integument ; Ecdysteroids ; Ixodid tick ; Amblyomma variegatum
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    Topics: Biology
    Notes: Summary The structure of the extensible (alloscutum) and inextensible (scutum) integument of the nymph, Amblyomma variegatum was examined during the whole bloodmeal and the nymphal-adult moulting cycle. Integumental events were tentatively correlated with the ecdysteroid levels measured by radioimmunoassay. We observed that all the integumental events were realised along an anteroposterior gradient. During the 5 days corresponding to the bloodmeal, although the hormone concentration was low, a new endocuticle was deposited on both the alloscutum and scutum. Furthermore, mitoses were initiated in the capitulum. On days 1–2 after the meal, ecdysteroid titres began to increase and reached a first peak corresponding to 4.1 ng 20-hydroxyecdysone equivalents/tick on the 4th day after the ticks dropped off their host. At this time the epidermis of the capitulum was detached and the outline of the adult capitulum was already visible. Mitotic activity in the alloscutum was initiated. On day 6 post-drop, the frontal apolysis was achieved and the ecdysteroid titres declined to basal values. A second peak much higher than the first one (maximum value of 33.7 ng/tick) and identified principally as 20-hydroxyecdysone by HPLC/RIA was noted on the 13th day post-drop. During the period of increase in the ecdysteroid levels (days 9–10 post-drop), the mitotic phase ended in the alloscutum and the apolysis began. Epicuticle was deposited after day 12 postdrop. Then, while the titre fell to low values (about 1.6 ng/tick, days 16–20 post-drop), the exocuticle was deposited and the nymphal cuticle was digested. All adult structures were functional 3 days before ecdysis. In young male as in female adults the mean value of the ecdysteroid levels corresponded to about 2.5 ng/tick. Finally, hydrolysis of tick whole extracts with esterase demonstrated a low increase of RIA-positive material, demonstrating the probable presence of natural ecdysteroid fatty-acid conjugates in this species.
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  • 43
    ISSN: 1432-041X
    Keywords: Mesoderm induction ; Activin A ; Erythroid differentiation factor ; Xenopus laevis
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    Topics: Biology
    Notes: Summary Recently the mesoderm-inducing effects of the transforming growth factor β (TGF-β) family of proteins have been widely examined. In an attemt to elucidate the functions of these proteins, porcine inhibin A and activin A (erythroid differentiation factor; EDF) were examined. Treatment of explants with activin A led to differentiation of mesodermal derivatives such as mesenchyme, notochord, blood cells and muscle, but inhibin A had a much lesser effect. The mesodermal differentiation induced by activin A was also comfirmed by analyses using a polyclonal antibody against muscle myosin. By indirect immunofluorescence analysis, the differentiation of muscle blocks was observed in the activin-A-treated explants, whereas no differentiation was observed in inhibin-A-treated and control explants. These findings confirm that this protein of the TGF-β family has mesoderm-inducing ability.
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  • 44
    ISSN: 1432-041X
    Keywords: Xenopus embryos ; Midblastula transition ; DNA injection ; CAT enzyme expression ; Actin-CAT fusion gene
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    Topics: Biology
    Notes: Summary Previous papers have reported that DNAs exogenously injected into Xenopus laevis fertilized eggs are expressed only at and after the midblastula transition (MBT). We have injected fertilized eggs of Xenopus laevis with circular plasmids that contained bacterial chloramphenicol acetyltransferase (CAT) genes connected to the promoter of viral genes (pSV2CAT and pAd12.E1aCAT) or the Xenopus cardiac actin gene (actin-CAT fusion gene), and examined whether these DNAs are expressed during the stage before the MBT. We found that expression of CAT enzyme can be detected before the MBT when CAT genes connected to viral promoters were injected. The activity was low during the cleavage stage on a per-embryo basis; however, the time course of the accumulation of the CAT enzyme activity roughly paralleled the increase in cell number. Therefore, CAT enzyme activity per cell was constant during cleavage and did not change dramatically before and after the MBT stage. CAT mRNA level, detected by CAT antisense RNA, was roughly proportional to the levels of CAT enzyme. Therefore, we assume that the observed enzyme activity reflects the transcriptional activity of injected CAT genes before and after the MBT. When the actin-CAT fusion gene was injected, however, no enzyme activity was detected until embryos had reached the neurula stage, a stage when endogenous actin genes are first activated. On the basis of these results, we conclude that the concept of an initial transcriptional activation of exogenous genes at amphibian MBT has to be changed. We propose that the expression of polymerase-II-transcribed genes is regulated by the nature of the promoters connected to the genes rather than by changes associated with MBT.
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  • 45
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    Development genes and evolution 198 (1990), S. 355-362 
    ISSN: 1432-041X
    Keywords: Wound healing ; Currents ; Electric fields ; Newt ; Epithelization
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    Topics: Biology
    Notes: Summary Wounded amphibian skin heals initially by a migration of epithelial cells from the cut edge towards the center of the wound. The density of currents leaving wounds made in Notophthalmus viridescens skin was manipulated in order to determine whether electrical fields associated with these currents might have a significant role in promoting this cell migration during wound healing. Wounds were made with either a needle (200 μm) or a biopsy punch (500 μm). Currents leaving the wounds were measured with a vibrating probe, and the wounds fixed at various times after wounding. When the Na+-dependent currents were reduced by blocking Na+ channels with benzamil, wound healing, as revealed by scanning electron microscopy and by paraffin histology, was impaired. These results are consistent with the hypothesis that there is an electrical component to wound healing.
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  • 46
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    Development genes and evolution 198 (1990), S. 402-410 
    ISSN: 1432-041X
    Keywords: Anteroposterior polarity ; Cytoplasmic factors ; Drosophila ; Cytoplasm transplant ; Maternal-effect mutants
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    Topics: Biology
    Notes: Summary Cytoplasm removal/transplant techniques applied to Drosophila cleavage-stage embryos induced changes in anteroposterior polarity. Removal of anterior cytoplasm or anterior transplantation of posterior cytoplasm caused the anterior formation of posterior (telson) structures, and the replacement of anterior cytoplasm with posterior cytoplasm induced double-abdomen embryos, as reported by Frohnhöfer et al. [J Embryol Exp Morphol 97 (suppl):169–179 (1986)]. Changing the conditions of anterior cytoplasm removal we showed that greater volumes, earlier stages, and removal from the periphery were efficient. In addition we found that double-cephalon embryos are induced by replacing posterior cytoplasm with anterior cytoplasm, while removal of posterior cytoplasm or the posterior transplantation of anterior cytoplasm was without effect. However, introduction of anterior cytoplasm into the posterior of nanos embryos, which are mutants not developing abdominal segments, caused the formation of double-cephalon embryos. Similarly, double-abdomen embryos are produced by introducing posterior cytoplasm into the anterior of bicoid embryos, which are mutants not forming cephalic and thoracic structures. These results are compatible with the initial involvement of separate anterior, posterior and terminal cytoplasmic factors deduced from mutant analysis (Nüsslein-Volhard and Roth 1989).
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  • 47
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    Development genes and evolution 198 (1990), S. 411-419 
    ISSN: 1432-041X
    Keywords: Drosophila ; Embryonic cells ; Ca2+-dependent cell aggregation ; Inhibiting antibodies ; Aggregation proteins
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    Topics: Biology
    Notes: Summary By using an in vitro functional assay, we have shown that Drosophila embryonic cells possess Ca2+-dependent adhesive sites, which resemble in many respects those described for vertebrate cells and tissues. The cells, obtained by mechanical disruption of gastrulastage embryos, form aggregates within 30 min when maintained under constant rolling. The aggregation is completely dependent on the presence of Ca2+ in the medium. In its absence, the cells remain dispersed but the process is reversible by readdition of Ca2+. In addition the aggregation is temperature-dependent. No aggregation occurs at 4° C but it can be restored by raising the temperature to 25° C. These properties are characteristic of these cells: established cell lines do not aggregate under the same conditions and mixing of cell lines and embryonic cells does not result in chimeric aggregates, thus pointing towards cell-type selectivity with respect to aggregability. Observations in electron microscopy have shown that the embryonic cells in the aggregates tightly adhere to one another and form, as early as after 30 min, maculae adherens junctions. Drosophila embryonic cells have adhesion sites that are protected from trypsin proteolysis in the presence of Ca2+ and sensitive in its absence. The cells' aggregation can be inhibited by a mouse antiserum directed against cell-surface components and a good correlation exists between neutralization of the inhibitory activity of the antiserum and the presence of trypsin-sensitive sites on the cells. These data are in favour of cell-cell adhesion mediated by specific adhesion proteins.
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    Development genes and evolution 198 (1990), S. 430-432 
    ISSN: 1432-041X
    Keywords: Ascidia ; Metamorphosis ; Ammonium ions ; Diacylglycerol
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    Topics: Biology
    Notes: Summary Larvae of the tunicate Ciona start metamorphosis between some hours and a few days after hatching. Several substances were found to reduce this time span, such as vital dyes [Cloney (1961) Am Zool 1:67–87; Hirani (1961) Bull Mar Biol Stn Asamushi 11:121–125], heavy-metal ions including copper [see review by Lynch (1961) Am Zool 1:59–66] and the hormone thyroxine [Patricolo et al. (1981) Cell Tissue Res 214:289–301]. This study shows that low concentrations of ammonium ions as well as the second messenger dicapryloylglycerol induce metamorphosis immediately after hatching. On the other hand, when the follicle cells are removed newly hatched Ciona remain larvae for days. Follicle cells are possibly degraded by bacteria, which thereby produce ammonia.
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  • 49
    ISSN: 1432-041X
    Keywords: Ambystoma mexicanum ; Melanin ; Germ cell marker ; Germ cells ; Transplantation
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    Topics: Biology
    Notes: Summary In urodele amphibians, the lack of a reliable germ cell marker restricts the experimental study of the germ lineage. In the present work, we conducted genetic and histological analyses in order to demonstrate that melanin from oocytes constitutes a germ cell marker available for intraspecific experiments in Ambystoma mexicanum. Then, using this marker, we implanted germ cells from undifferentiated gonads (stage 48) into the blastocoel of host embryos and investigated their fate and determined state. Our results show that, from this stage on, the donor cells do not differentiate into other cell types; therefore, they are restricted in developmental capacity and irreversibly determined as germ cells. On the other hand, exogenous germ cells were found in an isotopic position until the young tail-bud stage, and then were found in an ectopic position; these results suggest that, from the middle tail-bud stage on, an active process contributes to migration of primordial germ cells to the gonadal territory.
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  • 50
    ISSN: 1432-041X
    Keywords: Mesenchymal-epithelial interaction ; Stomach ; Heterotypic induction ; Pepsinogen ; Morphogenesis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Urogenital sinus endoderm of 16.5-day rat foetuses was combined with stomach mesenchyme and the recombinants were either treated with testosterone and grown in vitro or cultured beneath the kidney capsule of adult male rats of the same strain. It was found that testosterone stimulated mitosis in the urogenital endoderm. In recombinants grown under the kidney capsule a stratified squamous epithelium and stomach-like glands were induced under the influence of the forestomach and glandular stomach mesenchymes. However, the induced glands expressed neither rat pepsinogen nor rat ventral prostatic antigen. They did not produce mRNA for the prostatic steroid-binding protein C1. Thus, stomach mesenchyme of rat foetuses induces organ-specific morphogenesis but not functional differentiation in the heterologous endoderm, indicating that cytodifferentiation does not always accompany morphogenesis.
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  • 51
    ISSN: 1432-041X
    Keywords: Amphibian ; Gastrulation ; Dorsalization ; Mesodermal cell migration ; Lithium
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    Topics: Biology
    Notes: Summary The effect of lithium (Li+) on gastrulation movements was investigated during the development of the urodele amphibianPleurodeles waltl. Attention was focused on mesodermal cell migration. Under conditions of Li+ treatment providing a maximal enhancement of dorsoanterior structures, it was found that the dorsoventral polarity of gastrulation was abolished. In particular, vital staining and scanning electron microscopy observations on embryo fractures showed that mesodermal cells migrated radially after Li+ treatment, which led to the formation of rounded embryos. Epiboly movements thus were accelerated. Nevertheless, contrasting with the precocious disappearance of the early-formed yolk plug, archenteron invagination was constantly retarded and commenced with a delay of several hours as compared to control gastrulae. Cell-lineage analysis of the progenies from ventral or dorsal equatorial blastomeres of 32-cell-stage embryos provided evidence that both dorsal and ventral mesoderm contributed to notochordal tissue after Li+ treatment. Dorsalization of the entire marginal zone was confirmed by the ability of the entire mesoderm rudiment to behave as a dorsal organiser after Li+ treatment. Comparison of the migratory behaviour of isolated animal hemispheres from Li+-treated or control embryos cultured on fibronectin-coated substrate indicated that all marginal cells acquired the autonomous capacity for migration of dorsal marginal cells under the action of lithium.
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  • 52
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    Development genes and evolution 199 (1990), S. 14-30 
    ISSN: 1432-041X
    Keywords: Polychaete ; Development ; Dorsoventral polarity ; Segregation ; Morphometry
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    Topics: Biology
    Notes: Summary As in many spiralian embryos with unequal cleavage, cleavage inPlatynereis follows an invariant pattern. Preceding each cleavage the cytoplasm is reorganized, allowing the spiral cleavage mode to produce cells with different cytoplasmic composition. The fertilized egg undergoes a dramatic ooplasmic segregation after the completion of the cortical reaction. As a consequence, a plug of clear cytoplasm becomes located at the animal pole. Once the four quadrants of the embryo have been established, the cleavage sequence of the D quadrant differs clearly from that of the other three quadrants. The results presented here suggest that differential distribution of the clear cytoplasm governs this sequence. The first quartet of micromeres, which will form the ectoderm and the cerebral ganglia of the head, is clearly bilaterally symmetrical from the onset of the third cleavage. Dorsoventral polarity and bilateral symmetry in the ectoderm of the trunk is expressed most markedly by the dorsal location of the large 2d cell, whose rapid proliferation is bilaterally symmetrical with respect to the median plane. As a result of this proliferation it comes to fill most of the posttrochal region (ectoderm, three pairs of anlagen for the setal sacs, and the ventral plate which forms the nerve cord). The other micromeres contribute only a minor portion of the ventral ectoderm and are involved in the formation of the stomodaeum. The mesentoblast, 4d, i.e. the stem cell of the primary mesoderm, forms at the sixth cleavage, also in a position on the dorsal mid-line. The daughter cells, which arise from 4d by strictly bilaterally symmetrical cleavage, form the mesodermal germ bands, which lie beneath the ectoderm. The trochoblasts are formed by asynchronously cleaving founder cells, but further cleavages in these cells are synchronous. This suggests that cell-cell interaction is involved in the development of this alleged mosaic embryo.
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  • 53
    ISSN: 1432-041X
    Keywords: Glucose-6-phosphatase ; Endoplasmic reticulum (ER) ; Microsome ; Endoderm, sea urchin
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    Topics: Biology
    Notes: Summary Glucose-6-phosphatase (G-6-Pase) activity was analyzed during early embryogenesis of the sea urchinS. purpuratus. This activity is detected in very low levels in unfertilized eggs and early embryos but is present at high levels in preparations of endoplasmic reticulum (microsomes) from gastrula stage embryos. The approximately eight-fold increase in the relative activity of G-6-Pase associated with the ER occurs abruptly during a 12 h interval at gastrulation, and thereafter remains at a level comparable to that found in mammalian liver microsomes. The enzyme activity associated with the ER of gastrula stage embryos was completely eliminated from the microsomal pellet when cell lysates were first treated with non-ionic detergent. Analysis of germlayer tissues from late stage pluteus embryos revealed that G-6-Pase activity was more highly enriched in microsomes of endo/mesoderm tissues as compared to microsomes from ectoderm. The increase in ER associated G-6-Pase activity during embryonic development and its enriched activity in the ER of endo/mesoderm, as well as the observation that the signal recognition particle becomes associated with the ER at gastrulation (Le Blanc and Infante 1989), opens the question that this cellular organelle may be differentiating during embryogenesis in sea urchins.
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  • 54
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    Development genes and evolution 199 (1990), S. 107-113 
    ISSN: 1432-041X
    Keywords: Metamorphosis ; Phosphatidylinositol signalling ; K+ ionophores ; Protein kinase inhibitors ; Hydractinia echinata (Hydrozoa)
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    Notes: Summary Metamorphosis of planula larvae involves an activation of morphogenetically quiescent cells. The present work extends a previous study [Leitz T and Müller WA (1987) Dev Biol 121:82–89] on the participation of the phosphatidylinositol/diacylglycerol/protein kinase C system. Metamorphosis is stereospecifically induced by diacylglycerols, 1,2,-sn-dioctanoylglycerol (diC8) being by far the most effective substance. K-252a and sphingosine, inhibitors of mammalian protein kinases C, profoundly inhibited metamorphosis. Phorbolester-binding studies and the corresponding Scatchard plots revealed a specific and saturable binding of [3H]phorbol 12,13-dibutyrate to a single site of particulate fractions ofHydractinia with a specific binding affinityK d = 50 nM. K+ ionophores stimulated Cs+ — but inhibited diC8-induced metamorphoses, K+-channel blockers enhanced the inducing action of Cs+ or diC8. On the basis of these data and observations of others we propose that the activation ofHydractinia larvae takes place in some cells at the anterior end as a result of activation of a kinase-C-like enzyme, which directly or indirectly leads to the closure of K+ channels. Closure of these channels then causes depolarisation and, thus, release of an internal signal. This hypothesis unifies notions about the role of K+ channels and of the phosphatidylinositol system in initiation of metamorphosis inHydractinia.
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  • 55
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    Development genes and evolution 199 (1990), S. 114-121 
    ISSN: 1432-041X
    Keywords: Vitellogenesis ; Hemolymph proteins ; Fat body ; Oocyte ; Mosquito
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    Topics: Biology
    Notes: Summary The fat body of vitellogenic mosquitoes was found to synthesize and secrete another protein in addition to vitellogenin, that accumulated in developing oocytes. In the tissues, this protein has Mr = 53000 on SDS-PAGE under reducing or non-reducing conditions. This protein is glycosylated as shown by [3H]mannose incorporation and experiments with tunicamycin. Polyclonal antibodies were produced using the ovarian 53-kDa peptide. Immunoblot analysis demonstrated the immunological identity of 53 kDa peptides from the fat body and the ovary. Furthermore, the 53-kDa protein (53KP) is synthesized and secreted exclusively by the vitellogenic fat body. Radioimmunoassay showed that 53KP is produced by the female fat body as early as 4 h and reaches its peak near 24 h after the initiation of vitellogenesis. Synthesis then drops to low levels by 36 h and declines to background levels by 48 h. In vitro experiments conducted on fat bodies of previtellogenic females demonstrated that the synthesis and secretion of 53KP can be stimulated by a physiological dose of 20-hydroxyecdysone (10−6 M). Immunocytochemical studies of the ovary demonstrate that 53KP is present in channels between follicle cells, in the perioocytic space and in yolk granules of the developing oocytes. This suggests that 53KP is accumulated in the oocytes by a pathway similar to that of vitellogenin.
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  • 56
    ISSN: 1432-041X
    Keywords: Ammonia ; Methylamines ; Glutamine ; Nematocytes ; Nerve cells ; Coelenterates ; Hydra ; Hydractinia
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    Topics: Biology
    Notes: Summary Hydrozoa replace used-up nematocytes (cnidocytes) by proliferation and differentiation from interstitial stem cells (i cells). Repeated pulsed exposure ofHydra to elevated levels of unprotonated ammonia leads to successive loss of the various types of nematocytes: first of the stenoteles, then of the isorhizas and finally of the desmonemes. The loss is due to deficits in supply; the number of nematoblasts and differentiating intermediates is reduced. In the hydroidHydractinia the main process leading to numerical reduction was observed in vivo: mature nematocytes as well as precursors emigrate from their place of origin into the gastrovascular channels where they are removed by phagocytosis. This is a regular means by which these animals down-regulate an induced surplus of nematocytes. With lower effectiveness, pulses of methylamine, trimethylamine and glutamine also induce elimination of the nematocyte lineages. In the long term the population of nerve cells, which are permanently but slowly renewed from interstitial neuroblasts, decreases, too. After 2 months of daily repeated treatment the density of the Arg-Phe-amide-positive nerve cells was reduced to 50% of its normal level. Thus, ammonia induces down-regulation of all interstitial cell lineages. The temporal sequence of the ammonia-induced loss reflects the diverse rates with which the various i cell descendants normally are renewed.
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  • 57
    ISSN: 1432-041X
    Keywords: Micropylar apparatus ; Bradysia tritici ; Oogenesis ; Follicle cells ; Egg shell
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    Notes: Summary The ultrastructure and morphogenesis of the micropylar apparatus (MPA) have been studied in follicles of the fungus gnatBradysia tritici. The MPA is formed by a group of follicle cells located at the anterior pole of the single large nurse cell. In principle, the MPA consists of two thickened plates made of vitelline membrane material, the lower (LMP) and upper micropylar plate (UMP). The former is synthesized by 3 follicle cells, the latter by 4 different follicle cells. The micropylar channel system consists of a central channel with a single outer orifice and three branches which reach the plasma membrane of the oocyte. The branches are moulded by cellular extensions of the LMP-forming cells which are sandwiched between the two growing micropylar plates. Microtubuli and microfilaments were identified parallel to the long axis of the cellular extensions. At the time of MPA synthesis the nurse cell is still large and hence the MPA-forming cells have no contact to the oocyte. At the end of oogenesis when the regression of the nurse cell is completed, the MPA becomes connected to the other parts of the egg shell. At this time an ultrastructurally homogeneous region forms in the adjacent ooplasm (“cytoplasmic cone”). The possible relevance of these cytological observations for the control of development is discussed.
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  • 58
    ISSN: 1432-041X
    Keywords: Ca2+/Mg2+ ATPase ; Molluscan eggs ; Ionic currents ; Polarity ; Calcium
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    Topics: Biology
    Notes: Summary During extrusion of the first polar body in eggs ofLymnaea stagnalis andBithynia tentaculata a localized Ca2+ /Mg2+ ATPase activity was detected, using Ando's enzyme-cytochemical method for electron microscopy [Ando et al. (1981) Acta Histochem Cytochem 14:705–726]. The enzyme activity was distributed in a polar fashion, along the cytoplasmic face of the plasma membrane. In the eggs ofLymnaea it was found only in the vegetal hemisphere, whereas inBithynia eggs it was localized both in the vegetal hemisphere and at the animal pole. This pattern of enzyme activity corresponds to the polar pattern of transcellular ionic currents measured with the vibrating probe, which we showed to be partially carried or regulated by calcium [Zivkovic and Dohmen (1989) Biol Bull (Woods Hole) 176 (Suppl):103–109]. The characteristics of the ATPase were studied using a variety of approaches such as ion and substrate depletions and substitutions, addition of specific inhibitors of ATPase activity, treatment with EDTA/EGTA and electron energy-loss spectrometry. The results indicate that, inLymnaea, there are at least two enzymatic entities. The first one is a Ca2+ /Mg2+ ATPase localized along the membrane and in the cortex of the vegetal hemisphere. The second one is a Ca2+-stimulated ATPase (calcium pump of the plasma membrane) localized in a small region of the membrane at the vegetal pole. We speculate that in the eggs ofLymnaea andBithynia a functional relationship exists between the plasma-membrane-associated ATPase activity and the transcellular ionic currents measured in the same region.
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  • 59
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    Development genes and evolution 199 (1990), S. 246-250 
    ISSN: 1432-041X
    Keywords: F-actin ; Cell-to-cell interactions ; Molluscan embryos ; Induction ; TRITC-phalloidin
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The dorsal-ventral axis inPatella vulgata embryos is established at the 32-cell stage by an inductive interaction between the animal micromeres and one vegetal macromere. This vegetal macromere, once induced, is called the 3D macromere, and marks the future dorsal side of the embryo. We examined the pattern of filamentous (F) actin in such embryos using fluorescent phalloidin and found that this dorsal 3D macromere contains more F-actin than the remainder of the cells. In addition, only one of its two daughter cells, i.e. the 4D macromere, retains this higher density. In embryos in which the establishment of the dorsal-ventral axis has been experimentally inhibited via treatment with monensin, such differences in F-actin were not found. These results suggest that the appearance of an increased density of F-actin in the dorsal 3D and 4D macromeres of normal embryos requires the inductive interactions that establish the dorsal-ventral axis. We therefore conclude that F-actin is an early marker for dorsal induction in thePatella embryo.
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  • 60
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    Development genes and evolution 199 (1990), S. 237-245 
    ISSN: 1432-041X
    Keywords: Cellular interactions ; Cell migration ; Xenopus laevis ; Explants ; Cell division
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The consistency of the frog blastula's fate map is produced, in part, because the progeny of blastomeres located in dfferent regions do not intermix with one another. We examined the cause for this restriction of intermixing in two types of cultures. In one type of culture, two groups of cells were excised from blastulae and stuck together; the movement of cells between the groups was monitored. Cells migrated more extensively between groups derived from the same region than between groups derived from different regions. In the other type of culture, a single cell was implanted into a group of cells that was excised from the blastula. The rate of division and the extent of migration of the implanted cell's clone were monitored. The implanted cell divided more rapidly among cells from its own region than among cells from a different region. Both experiments show that the restriction of intermixing that occurs between regions of the intact embryo also occurs in vitro. These results indicate that the restriction does not result secondarily from normal morphogenetic movements, which are absent from the explants, but probably from cellular interactions that limit the extent of cell migration. This limitation is correlated with a reduction in the rate of cell division.
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  • 61
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    Development genes and evolution 198 (1990), S. 363-369 
    ISSN: 1432-041X
    Keywords: Type II collagen ; Subepithelial extracellular matrices ; Embryonic chick development
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    Topics: Biology
    Notes: Summary Type II collagen is a major component of hyaline cartilage but recent studies have demonstrated the presence of this protein in a variety of interfaces that separate epithelia from mesenchyme, particularly in early stages of embryonic chick development. In the present study an immunohistochemical analysis of the distribution of type II collagen was performed on closely staged wing buds of early chick embryo. This report describes how using two different monoclonal antibodies against type II collagen and the peroxidase or fluorescence staining technique reveals that deposition of type II collagen at the ectoderm-mesenchyme interface occurs in the proximal part of the limb coincidentally with the appearance of this protein in the proximal core region, where chondrogenesis begins (stage 25). Then the staining in the subepithelial region spreads distallly with time, following the progression of the formation of cartilage rudiments. At about 7 days of development type II collagen is present under the apical ectoderm ridge and surrounds completely the wing bud underneath the epithelium. At the same time, another antibody directed against the cartilage-specific proteoglycan core protein only stains the chondrogenic central core of the limb and not the subepithelium. Although type II collagen and cartilage-specific proteoglycan are closely associated in the cartilage, the observations presented here suggest that the deposition of these proteins can be regulated independently during limb formation. The role of type II collagen at the epithelium-mesenchyme interface during limb formation is still to be determined.
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  • 62
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    Development genes and evolution 198 (1990), S. 371-381 
    ISSN: 1432-041X
    Keywords: Chick embryo ; Extremities (limbs) ; Morphogenesis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The purpose of this study was to determine whether the organizer regions of early avian and amphibian embryos could induce supernumerary (SN) wing structures to develop when they were grafted to a slit in the anterior side of stage 19–23 chick wing buds. Supernumerary digits developed in 43% of the wings that received anterior grafts of Hensen's node from stage 4–6 quail or chick embryos; in addition, 16% of the wings had rods of SN cartilage, but not recognizable SN digits. The grafted quail tissue did not contribute to the SN structures. When tissue anterior or lateral to Hensen's node or lateral pieces of the area pellucida caudal to Hensen's node were grafted to anterior slits, the wings usually developed normally. No SN structures developed when Hensen's nodes were grafted to posterior slits in chick wing buds. Wings developed normally when pieces of the dorsal lip of the blastopore from stage 10–11.5 frog (Xenopus laevis and Rana pipiens) embryos were grafted to anterior slits. No SN digits developed when other tissues that have limb-inducing activity in adult urodele amphibians [chick otic vesicle, frog (Rana pipiens) lung and kidney] or that can act as heteroinductors in neural induction (rat kidney, lung, submaxillary gland and urinary bladder; mouse liver and submaxillary gland) were grafted to anterior slits in chick wing buds. SN digits also failed to develop following preaxial grafts of chick optic vesicles. These results suggest that although the anteroposterior polarity of the chick wing bud can be influenced by factors other than the ZPA (e.g., Hensen's node, retinoids), the wing is not so labile that it can respond to a wide variety of inductively-active tissues.
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  • 63
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    Development genes and evolution 198 (1990), S. 382-388 
    ISSN: 1432-041X
    Keywords: Hydra vulgaris ; Pattern control ; Lithium ; Foot formation
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    Topics: Biology
    Notes: Summary LiCl in concentrations exceeding 0.5 mM affects morphogenesis in Hydra vulgaris (formerly named H. attenuata) by interfering with the foot-forming system(s). Pulse treatment of Hydra bearing small buds or of animals that develop a bud within 14 h after the end of treatment prevented foot formation at the bud's base in a concentration-dependent manner. With increasing concentrations of Li+ or length of treatment in increasing percentage of the buds remained permanently connected to the parent by a bridge of tissue thus forming a stable secondary axis. Instead of the normal ring-shaped foot a patch of basal disc tissue developed or the bud failed to differentiate foot tissue at all. Long-term culture of animals in 1 mM LiCl inhibited budding from the second day of treatment onwards and detachment of existing buds was delayed. After 4 days of treatment 15%–30% of budless or bud-bearing animals developed up to three patch-like basal discs at various positions along the body axis; these usually grew out one above the other on the same side of the animal but never at the same transverse level. Besides these patch feet broad belts of foot tissue were observed in the lower gastric region. After 1 week of treatment half of the animals developed a constriction located usually in the lower two-thirds of the body axis. The tissue adjacent to this constriction and particularly above it differentiated into mucus-secreting foot tissue. Subsequent separation into two morphologically intact polyps occurred occasionally. When treatment was stopped, budding restarted within the next 3 days at several positions along the body axis whether or not secondary feet or a constriction existed. Buds grew out in different budding zones, which persisted for several days. This burst of budding led to up to 7 buds per animal within 3 days. After about 1 week the animals regulated to normality or became epithelial, i.e. they lost their stem cells during and after treatment.
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  • 64
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    Keywords: Cell surface ; Xenopus ; Glycoprotein ; Lectin ; Biotinylation
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    Notes: Summary The populations of cell surface proteins and total glycoproteins were investigated in early Xenopus embryos through lectin staining, affinity binding of glycoproteins to lectins, and use of a succinimide ester to biotinylate cell surface molecules. Lectin staining shows that the egg is endowed with a thick layer of surface glycoprotein, and that glycoprotein is immediately detected on the newly formed membranes of nascent blastomeres. The amount of glycoprotein found in eggs and early embryos remains constant, and electrophoretic analysis reveals no changes in abundant lectin-binding glycoproteins through the neurula stage. In contrast, the amount of cell surface protein increases dramatically from the 2-cell to the gastrula stages. Despite this quantiative increase, only a small number of differences in cell surface proteins were detected during this period. A series of bands was detected which appears to be specific to the outer surface of the embryo. Because the populations of surface proteins and of total glycoproteins overlap to a great extent, the increase in cell surface protein, in the absence of a change in total glycoprotein, indicates the presence of a maternal glycoprotein pool in the Xenopus egg, from which the cell surface proteins of embryonic blastomeres are recruited.
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  • 65
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    Development genes and evolution 198 (1990), S. 443-447 
    ISSN: 1432-041X
    Keywords: Limb regeneration ; Proliferation ; Blastema ; Epidermal cap ; Axolotl ; Growth factors
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The presence of a mitogenic activity in limb blastemas of axolotls was detected in crude extracts of blastemas at the mid-bud stage. The mitogenicity of the extracts was estimated from the mitotic index of blastema cells grown for 6 days in the presence of limb blastema extracts, with colchicine present for the last 2 days. All the extracts tested (whole blastema, blastemal mesenchyme, epidermal cap) significantly enhanced proliferation of blastema cells. The highest stimulation factors we observed were 7 × with 7 μg protein/ml whole blastema extracts, 5.2 × with 14 μg/ml blastemal mesenchyme extracts, and 11 x with 3.5 μg/ml epidermal cap extracts. Hence the epidermal cap extracts appeared to be the most mitogenic. Extracts from the blastemal mesenchyme, although less mitogenic than the epidermal cap extracts, were more potent than nerve extracts [Albert P, Boilly B (1986) Biol Cell 58:251–262]. These results are discussed with regard to the production of growth factors during limb regeneration.
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  • 66
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    Development genes and evolution 198 (1990), S. 448-459 
    ISSN: 1432-041X
    Keywords: Positional information ; Wing morphogenesis ; Lyra ; E1C antigen
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary A number of parameters characteristic of the wing margin precursor in imaginal discs of Drosophila are known: the zone of non-proliferating cells or ZNC (O'Brochta and Bryant 1985), aldehyde oxidase (AO) and other enzyme staining patterns (Sprey et al. 1982), E1C antigen localization in a narrow band along the margin (Piovant and Lena 1988). To test our hypothesis that such parameters, and others, act in concert to determine margin identity and the positional information that specifies the bristles and hairs appropriate to the anterior, posterior and distal margins, we have examined these parameters in the dominant mutant Lyra, in which much of the anterior and posterior margins is missing. After establishing that Lyra phenotype is already evident in the early pupal wing, we tested the known imaginal disc parameters and found that only Mab E1C (Piovant and Lena 1988) distribution differs from wild type, suggesting that E1C antigen may be a component of positional information. Sibatani's (1983) model for specification of positional information (PI) applied to wing discs predicts the Lyra adult wing shape as well as the reduced distribution of E1C antigen in Lyra wing discs. The model is based on the assumption that specification of positional information depends on interactions of multiple, independent factors. Clonal analysis with shaggy (Simpson et al. 1988 and Ripoll et al. 1988) indicates that factors in addition to E1C antigen contribute to margin PI in Lyra wings and should allow us to test the multi-component hypothesis further.
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  • 67
    ISSN: 1432-041X
    Keywords: Epithelial-mesenchymal interaction ; Pepsinogen expression ; Gland formation ; Stomach
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary To clarify the precise conditions under which chick embryonic proventricular mesenchyme can induce proventricular epithelial differentiation, transfilter experiments were carried out. Six-day proventricular epithelium formed glands and expressed pepsinogen when a Nucleopore filter with a pore size of more than 0.6 μm, but not 0.2 μm, was inserted between the epithelium and the proventricular mesenchyme. The larger the pore size of the filter, the more elongated the glands and the more pepsinogen was induced in the explants. The quail nuclear marker and scanning electron microscopy were used to examine penetration of mesenchymal cells through the Nuclepore filter. The filter of more than 0.2 μm pore size allowed cell processes of mesenchymal cells to pass through. However, only the filter with a pore size of more than 0.6 μm allowed actual migration of mesenchymal cells through the filter, and the larger the pore size of the filter, the more mesenchymal cells passed through. Under the same conditions 6-day and 4.5-day gizzard epithelium formed glands and expressed pepsinogen. These results indicate that a flow of diffusible substances through a Nuclepore filter and even direct contact of a few short cell processes of mesenchymal cells with epithelial cells are not sufficient for induction, and that direct contact of mesenchymal cell processes and/or mesenchymal cells with epithelial cells over a considerably wide area may be prerequisite for the induction.
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    Development genes and evolution 198 (1990), S. 474-478 
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    Keywords: Arginine kinase ; Imaginal discs ; 20-hydroxyecdysone ; Drosophila
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    Topics: Biology
    Notes: Summary Arginine kinase (AK) is present throughout the life cycle of Drosophila melanogaster, but there is a sharp, transient peak of AK activity during the prepupal period and a second period of elevated activity at the time of eclosion of the adult. Imaginal discs show the greatest increase in AK activity at the prepupal stage of those tissues assayed. The prepupal peak is not seen when the temperature-sensitive ecdysoneless mutant ecd-1 is shifted to 29° C at mid-third instar larval stage. The peak in activity reappears when ecd-1 is either shifted back to 20° C after 60 h at 29° C or is fed 20-hydroxyecdysone. At the restrictive temperature, imaginal discs from ecd-1 larvae progressively lose AK activity, whereas discs from 20-hydroxyecdysone-fed larvae have a marked increase in AK activity at stage P3 of the prepupal period. These data suggest that the prepupal peak is regulated by the hormone 20-hydroxyecdysone.
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    Development genes and evolution 199 (1990), S. 207-211 
    ISSN: 1432-041X
    Keywords: 83-kDa nuclear antigen ; Germinal vesicles ; Ascidian embryogenesis ; Monoclonal antibody
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Monoclonal antibodies were raised against germinal vesicles which were isolated from fully grown oocytes of the ascidianHalocynthia roretzi. Immunoblot analyses revealed that one of the antibodies, designated Hgv-2, recognized a single band with a molecular weight of about 83 kDa. The antibody, visualized by indirect immunohistochemistry, reacted only with the germinal vesicles of oocytes and did not react with test cells, follicle cells, and other somatic cells of the gonad. During embryogenesis the antigenicity was found in interphase nuclei of all embryonic cells. The antibody did not react with chromosomes or the mitotic apparatus. The antigenicity was retained by interphase nuclei of larval cells, but it disappeared from nuclei of juveniles about 7 days after metamorphosis.
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    Development genes and evolution 199 (1990), S. 219-227 
    ISSN: 1432-041X
    Keywords: Puparium ; Metamorphosis ; Irradiation ; Larva ; Developmental timer
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary In the insectDrosophila, formation of the puparium marks the onset of metamorphosis and serves as a useful marker for developmental progress. The cells of the adult remain diploid and divide during the larval stage while the larval cells become polytene and do not divide. We use a high dose of gamma-irradiation (10 krad) to selectively delete the imaginal lineage from the developing larvae ofDrosophila melanogaster. We find that animals depleted of imaginal cells including those of the imaginal brain pupariate only if the larval cells are allowed to mature, demonstrating that the larval cells harbor the primary developmental timer for this process. However, proliferating imaginal cells can exert a negative influence on the timing of pupariation.
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    Development genes and evolution 199 (1990), S. 212-218 
    ISSN: 1432-041X
    Keywords: Sea urchin ; Hatching ; A23187 ; Tetracaine ; Verapamil
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Pulse treatment of sea urchin embryos with 3 µM A23187 for 2 h at 20° C, starting from 3 to 6 h of development, prevented the embryos from hatching. Many embryos thus treated with A23187 produced mesenchyme cells and underwent gastrulation while still enclosed within the fertilization membrane. The pulse treatment in this pre-hatching period exerts markedly stronger inhibitory effects on hatching than on other events in early development. Treatment beginning at times earlier than 2 h and later than 8 h of development caused only a slight delay of hatching. The activity of hatching enzyme, known to increase between 6 and 8 h after fertilization, was quite low, if present at all, in embryos in which hatching was blocked by A23187. Hatching enzyme synthesis is probably blocked by the preceding pulse treatment. However, overall protein synthesis, estimated with methionine S 35 incorporation, was somewhat augmented in embryos by the pulse treatment. The blockage of hatching and the augmentation of overall protein synthesis by A23187 were appreciably reversed by procaine, tetracaine, ruthenium red or verapamil. Probably, an artificial Ca2+ signal induced by A23187 activates protein synthesis but blocks the induction of hatching enzyme synthesis.
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    Development genes and evolution 199 (1990), S. 251-280 
    ISSN: 1432-041X
    Keywords: Spermatogenesis ; Drosophilia ; Y chromosomal lampbrush loops ; Male sterile mutations ; Cellular differentiation
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    Topics: Biology
    Notes: Summary We constructed balancer-chromosomes for the large autosomes ofDrosophila hydei and screened more than 16000 chromosomes for male sterile mutations in order to dissect spermatogenesis genetically. 365 mutants on the X chromosome and the autosomes 2, 3, and 4 were recovered and analysed cytologically in squash preparations under phase-contrast optics. The majority of the mutations allows a rather advanced differentiation of the spermatozoa. At the light-microscopical level, it is possible to classify these mutations with respect to individualization, coiling or motility of the mutant spermatozoa. In contrast, a small number of mutants exhibits conspicuous, pleiotropic phenotypes. Gonial divisions, the shaping of the spermatocyte nucleus and male meiotic divisions are controlled by X chromosomal or autosomal genes which can mutate to male sterile alleles. A number of nonallelic 3rd chromosome male sterile mutations interfere with the unfolding of the Y chromosomal lampbrush loops. Other autosomal male sterile mutations modify the morphology of these lampbrush loops. Another group of mutations inhibits the formation of the nebenkern while the development of the spermatid nucleus and the flagellum can proceed. Such male sterile mutations can decouple the development of nucleus, protein body, nebenkern, and flagellum of the spermatid. Thus, we can describe spermatogenesis inDrosophila as the coordinate execution of the individual developmental programs of the different components of the spermatozoon.
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    Development genes and evolution 199 (1990), S. 281-288 
    ISSN: 1432-041X
    Keywords: Ceratitis capitata ; Integument ; Haemocyte proteins ; Cuticle sclerotization ; Tyrosine carrier polypeptides
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    Topics: Biology
    Notes: Summary Studies of the synthesis of integumental proteins during the feeding and non-feeding stages ofCeratitis capitata demonstrated stage specificity. The synthetic profile changed dramatically, showing a maximum of protein synthesis just before the larval wandering stage, followed by an abrupt decline. The comparison between synthetic and accumulation profiles indicated that some polypeptides must be internalized into the integument from the haemolymph. The major haemolymph proteins or arylphorins have already been documented to be incorporated into the integument. In the present work, we demonstrated the interalization of some haemocyte proteins into the integument. For that purpose, polyclonal antibodies were raised against total haemocyte proteins. Immunoblot analysis of haemocyte salt extractable proteins revealed that the protein bands at 36, 54, 58, 84, 110 and 130 kDa were immunoreactive with the total haemocyte antibodies. Cell-free protein synthesis, organ culture experiments and immunoblot analysis indicated that the 36-, 54- and 58-kDa polypeptides were synthesized only in the haemocytes and were probably internalized into the integument from the serum. The 36-kDa polypeptide was also demonstrated to be internalized into the fat body of white puparia. The immunofluorescence experiments suggested that the internalization of haemocyte proteins first occurs into the epidermal cells and then into the cuticle. The presence of haemocyte proteins in the integument was also demonstrated by immunofluorescence experiments in twoC. capitata mutants. These mutations affect the darkening and stiffening of the cuticle. The demonstration of 36-, 54- and 58-kDa haemocyte polypeptides in the integument reveals a hitherto unknown function of this cell type. Moreover, the demonstration of tyrosine binding to the 54- and 58-kDa polypeptides points to their potential involvement in the sclerotization process in the cuticle.
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    Development genes and evolution 198 (1990), S. 264-274 
    ISSN: 1432-041X
    Keywords: Development ; Visual system ; Optic ganglia
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    Notes: Summary The larval and early pupal development of the optic lobes in Drosophila is described qualitatively and quantitatively using [3H]thymidine autoradiography on 2-μm plastic sections. The optic lobes develop from 30–40 precursor cells present in each hemisphere of the freshly hatched larva. During the first and second larval instars, these cells develop to neuroblasts arranged in two epithelial optic anlagen. In the third larval instar and in the early pupa these neuroblasts generate the cells of the imaginal optic lobes at discrete proliferation zones, which can be correlated with individual visual neuropils. The different neuropils as well as the repetitive elements of each neuropil are generated in a defined temporal sequence. Cells of the medulla are the first to become postmitotic with the onset of the third larval instar, followed by cells of the lobula complex and finally of the lamina at about the middle of the third instar. The elements of each neuropil connected to the most posterior part of the retina are generated first, elements corresponding to the most anterior retina are generated last. The proliferation pattern of neuroblasts into ganglion mother cells and ganglion cells is likely to include equal as well as unequal divisions of neuroblasts, followed by one or two generations of ganglion mother cells. For the lamina the proliferation pattern and its temporal coordination with the differentiation of the retina are shown.
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    Development genes and evolution 198 (1990), S. 275-285 
    ISSN: 1432-041X
    Keywords: Neurogenesis ; Notch ; split ; daughterless ; Genetic interactions ; Second-site modifiers ; Drosophila
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary We have searched for dominant modifiers, i.e., enhancers and suppressors, of the compound eye phenotype of split, a recessive viable allele of Notch. Among the spl modifiers found, we have detected mutations in loci whose functions were previously known to cooperate with Notch in embryonic neurogenesis, such as daughterless, master mind, Delta and Hairless. In addition, other spl modifier mutations have been found in loci that were not previously known to interact with Notch, such as scabrous, glass, roughened eye, and several other genes that have not yet been assigned to known loci. The phenotypes associated with mutations in some of these latter loci suggest the participation of the corresponding genes in embryonic neurogenesis. We show that in some cases the observed interactions are due to genetic haplo-insufficent expression of the genes, whereas allele-specific interactions with spl are observed in master mind and Delta alleles. From this observation, we propose a direct functional association between the proteins encoded by Notch, Delta and master mind.
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  • 76
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    Keywords: Compound eye morphogenesis ; Enhancer of split ; Drosophila
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    Notes: Summary The spl mutation of the N gene causes, among other phenotypic traits, the lack of a few ommatidia, roughness and a general reduction in the size of the compound eye; these defects are drastically enhanced by the dominant mutation E(spl) D. We have studied cellular and developmental aspects of the phenotypic interaction between spl and E(spl) D. We found that the initial clustering of photoreceptor cells is affected in eye imaginal discs of spl larvae causing the defects visible in the adult eye. The degree of disorganization of the spl/Y; E(spl) D/ + eye disc is much higher, only a few photoreceptor cells are able to group with representatives of the other cell types and differentiate normally. BrdU incorporation shows that the proliferation pattern of the spl/Y; E(spl) D/ + disc cells during the third instar is normal. Abundant cell death occurs posteriorly in the mutant discs, which accounts for their small size. Finally, we found that in the eye imaginal disc the transcription of m8, the E(spl) gene, responsible for the enhancement of the spl phenotype caused by the E(spl) D mutation, is restricted to the morphogenetic furrow, where the ommatidial cells start grouping with each other to take on their future developmental fates; the m8 transcription rate is highly increased in E(spl) D eye discs. All these observations indicate that the assembly of the ommatidial cells is affected in the spl/Y; E(spl) D/ + disc and that the other abnormalities are morphogenetic consequences of the defective cell grouping.
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    Development genes and evolution 198 (1990), S. 336-354 
    ISSN: 1432-041X
    Keywords: Pattern formation ; Morphogenetic operation ; Genetic interaction ; Wing vein
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    Topics: Biology
    Notes: Summary Of the many mutations known to affect the wing vein pattern we have selected the most extreme in 29 genes for study. Their phenotype can be classified in two major classes: lack-of-veins and excess-of-veins, and in several internally coherent groups. The study of multiple mutant combinations, within groups and between groups, reveals several genetic operations at work in the generation of the vein pattern. The finding that some of these mutations also affect cell proliferation in characteristic ways has prompted a generative model of wing morphogenetic and pattern formation based on cell behaviour properties defined by the corresponding wild-type genes.
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  • 78
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    Keywords: Athalia rosae ; Hymenoptera ; Thelytoky ; Automixis ; Gynandromorph
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    Topics: Biology
    Notes: Summary Mature eggs dissected from ovaries of unmated females of Athalia rosae (Hymenoptera: Tenthredinidae), if placed on a filter-paper soaked with distilled water, are activated and develop to haploid males. Occasionally, however, diploid females develop from these artificially activated eggs. Treatment of mature unfertilized eggs dissected from diploid females with ice-cold temperatures immediately before activation and with a high temperature (36° C) upon and immediately after activation resulted in the production of diploid males, diploid females, triploid females and gynandromorphs at high frequency. The same treatment of mature unfertilized eggs dissected from triploid females resulted in the production of only triploid survivors. These results, together with the results on the segregation of a marker mutation, yellow fatbody (yfb), appear to indicate that meiotic divisions were complete in the treated eggs, and that all four nuclei became potentially capable of participating in development with or without automictic fusion.
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    Development genes and evolution 198 (1990), S. 395-401 
    ISSN: 1432-041X
    Keywords: Morphogenesis ; Epithelial morphogenesis ; Monoclonal antibody ; Molecular marker ; Cockroach
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary A molecular marker has been identified in embryos of the cockroach, Periplaneta americana, that is localized among epithelial cells to those directly involved in morphogenesis. A monoclonal antibody has been developed that selectively binds to epithelial cells undergoing any of three very different morphogenetic movements-invagination, evagination or epiboly. Neighboring cells not involved in these developmental processes are not labeled by the antibody. The antigen is transiently present on the cells for a period just prior to and during the morphogenetic activity. It is localized on the apical surface of the cells. The spatial, temporal and subcellular distributions of antibody binding during development indicate a role for the antigen in epithelial morphogenesis different from that of any previously described molecule.
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    Development genes and evolution 199 (1990), S. 31-47 
    ISSN: 1432-041X
    Keywords: Bristle ; Pattern formation ; Drosophila ; Gamma rays ; Mitomycin C ; Heat shock
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The development of a leg segment of the fruitflyDrosophila melanogaster was analyzed in order to determine whether the orderliness of the segment's bristle pattern originates via waves of cellular interactions, such as those that organize the retina. Fly development was perturbed at specific times by either teratogenic agents (gamma rays, heat shock, or the drug mitomycin C) or temperature-sensitive mutations (l(1)63, l(1) Notchts1, orl(1) shibire ts1 ), and the resulting abnormalities (e.g., missing or extra structures) were mapped within the pattern area. If bristles develop in a linear sequence across the pattern, then they should show sensitivity to perturbations in the same order, and wavefronts of cuticular defects should result. Contrary to this prediction, the maps reveal no evidence for any directional waves of sensitivity. Nevertheless, other clues were uncovered as to the nature and timing of patterning events. Chemosensory bristles show earlier sensitivities than mechanosensory bristles, and longer bristles precede shorter ones. The types and sequence of cuticular abnormalities imply the following stages of bristle pattern development: (1) scattered inception of bristle mother cells, each surrounded by an inhibitory field, (2) alignment of the mother cells into rows, (3) differential mitoses, (4) assignment of cuticular fates to the mitotic progeny, (5) polytenization of the bristle cells, (6) fine-tuning adjustments in bristle spacing, and (7) signalling from bristle cells to adjacent epidermal cells, inducing them to form “bracts”.
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    Development genes and evolution 199 (1990), S. E1 
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    Development genes and evolution 199 (1990), S. 48-62 
    ISSN: 1432-041X
    Keywords: Bristle ; Sensilla ; Pattern formation ; Drosophila ; achaete-scute complex
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The arrangement of bristles on a leg segment of the fruitflyDrosophila melanogaster was studied in various mutants that have abnormal numbers of bristles on this segment. Eighteen mutations at six different genetic loci were analyzed, plus five double or triple mutant combinations. Recessive mutations at theachaete-scute locus were found to affect distinct groups of bristles:achaete mutations remove mechanosensory bristles, whereasscute mutations remove mainly chemosensory bristles. Mechanosensory bristles remain uniformly spaced along the longitudinal axis unless their number decreases below a certain threshold, suggesting that spacing is controlled by cell interactions that cannot function when bristle cells are too far apart. Above a certain threshold, bristle spacing and alignment both become irregular, perhaps due to excessive force from these same interactions. Chemosensory bristles occupy definite positions that are virtually unaffected by removal of individual bristles from the array. Extra chemosensory bristles develop only near the six normal sites. At two of the six sites the multiple bristles tend to exhibit uniform longitudinal spacing — a property confined to mechanosensory bristles in wild-type flies. To explain the various mutant phenotypes the following scheme is proposed, with different mutations directly or indirectly affecting each step: (1) spots and stripes are demarcated within the pattern area, (2) one bristle cell normally arises within each spot, multiple bristle cells within each stripe, (3) incipient bristle cells inhibit neighboring cells from becoming bristle cells, and (4) the bristle cells within each stripe become aligned to form rows and then repel one another to generate uniform spacing.
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    Development genes and evolution 199 (1990), S. 89-96 
    ISSN: 1432-041X
    Keywords: Amphibian embryo ; Transcriptional regulation ; Serum response element ; Microinjection ; In vitro mutagenesis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary The Serum Response Element (SRE) is a sequence motif which activates transcription of certain genes in response to factors that stimulate cell proliferation. This motif is found in the promoter of aXenopus laevis cytoskeletal actin gene, which is transcriptionally activated very early in embryonic development. We tested whether the SRE plays a role in the developmentally-timed transcriptional activation of this gene by constructing an SRE replacement mutant and studying its transcription after microinjection intoXenopus embryos. Normal amounts of actin mRNA are transcribed at the normal time in development from this mutant, suggesting that the SRE is not the sole determinant of temporal specificity of actin gene transcription in the embryo.
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    Development genes and evolution 199 (1990), S. 80-88 
    ISSN: 1432-041X
    Keywords: Morphogenesis ; Chick embryo ; Monoclonal antibodies ; A7H2
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    Notes: Summary In order to identify transitory molecules involved in specific events during development, monoclonal antibodies were raised against various structures of the chick embryo at stages exhibiting important tissue reorganization. A7H2 is a monoclonal antibody reacting with an epitope suddenly expressed in select embryonic tissues during precise morphogenetic processes. Splanchnomesoderm and most mesothelia exhibited a strong immunoreactivity during organogenesis whereas non-mesodermal A7H2-activities were usually confined to invaginating or folding epithelia. Most embryonic reactivities were transient but the amnion demonstrated a strong and persistent reactivity and therefore was selected as a source of material for A7H2-antigen characterization. The antigenic determinant was shown to be on a huge molecular complex having a pI of 5. Immunoblots obtained after SDS-PAGE demonstrated a highly disperse pattern in the high MW region of the gel that could be altered or abolished by proteolytic treatments. Fingerprint analysis of the immunoreactive peptide bands demonstrated a clear relationship between them. Using different fixation and permeabilization procedures on amnion cultured cells, we have observed that A7H2-antigen was associated with the cytoplasmic face of the plasma membrane and was resistant to treatments removing most cellular components and producing substratum-attached ventral membranes. We suggest that this macromolecular complex plays a role in specific subsets of cells involved in precise morphogenetic processes, possibly as a link with the external milieu or as a membrane associated structure.
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    Development genes and evolution 199 (1990), S. 63-79 
    ISSN: 1432-041X
    Keywords: Hydrozoa ; Planula metamorphosis ; Voltage dependent calcium channels ; Calcium transients ; Epithelial conduction ; Protein kinase-C activation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Both the natural metamorphic stimulus (an unidentified bacterial product) and an artificial trigger of metamorphosis (Cs+) cause large calcium transients in planula cells of the hydrozoanMitrocomella polydiademata. When these transients are inhibited with calcium channel blockers, metamorphosis is also inhibited. All cells of theMitrocomella planula contain a calcium-specific photoprotein. The cells where the calcium transients occur during natural- and Cs+-induced metamorphosis have been visualized in normal and entoderm free planulae that lack ganglion cells, using a compound microscope coupled to an image intensifier and video camera. During bacteria- and Cs+-induced metamorphosis, groups of contiguous cells, occupying from about 10% to the entire visible surface of the planula, simultaneously exhibit calcium transients. When the cells that initiate a transient comprise only part of the planula surface, the calcium transient frequently propagates and can eventually involve every cell on the visible planula surface. There is no special site on the planula surface where calcium transients are more apt to be initiated. There is no indication that propagation of a flash in one direction is more likely than in another. The velocity of propagation is virtually the same in all directions. The only feature of the spatial distribution of bacteria- and Cs+-induced calcium transients that appears to be necessary for the induction of metamorphosis is that at least one transient must involve all of the surface cells of the planula. The spatial behavior of calcium transients is the same in entoderm free planulae (lacking ganglion cells) as in normal planulae. The propagation of these calcium transients most probably occurs via epithelial conduction. This metamorphic step involving calcium transients is probably the intercellular communication system that informs the cells of the planula that metamorphosis will commence. Metamorphosis inMitrocomella planulae can also be induced with phorbol esters. Calcium transients do not occur during phorbol ester-induced metamorphosis, indicating that they act at a different point in the metamorphic pathway. Calcium channel blockers do not inhibit phorbol ester-induced metamorphosis. Inhibitors of protein kinase-C, inhibit both phorbol ester-induced metamorphosis and Cs+- and bacteria-induced metamorphosis, but have no effect on the calcium transients induced by Cs+. This indicates that the calcium transient mediated step in the metamorphic pathway occurs prior to protein kinase-C activation. Calcium transients probably play a major role in activating protein kinase-C.
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    Development genes and evolution 199 (1990), S. 97-101 
    ISSN: 1432-041X
    Keywords: Neurogenesis ; cGMP ; Xenopus ; Signal transduction
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Recent evidence indicates an important role for cell-surface mediated signal transduction in embryonic induction. We, therefore, started a systematic search to identify signal transduction pathways which are activated during embryonic induction and specifically during neural induction. We showed previously that the protein kinase C and cAMP pathways mediate neural induction inXenopus laevis. Here, we investigated whether cGMP is also involved in the early development of the nervous system. We measured the cGMP content of whole embryos at embryonic stages which mark important events in the early development of the nervous system, as well as in the developing neural tissue itself, after this was induced from ectoderm by dorsal mesoderm. No changes in cGMP content were found, either in whole embryos at different developmental stages, or in developing neural tissue from these stages. We also found no evidence for the presence of nitroprusside stimulatable guanylate cyclase in these developmental stages. A cGMP analogue, 8-Br-cGMP, was not able to induce neural tissue, either alone or in combination with known neural inducers, the phorbol ester TPA and 8-Br-cAMP. 8-Br-cGMP also had no negative influence on the neural inducing ability of dorsal mesoderm or TPA, alone or in combination with 8-Br-cAMP. We conclude that cGMP has no role in the early development of the central nervous system inXenopus laevis. This conclusion underlines the specificity of the signal transduction pathways (PKC and cAMP pathways) that do mediate neural induction.
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  • 87
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    Keywords: Lectin ; Apolipoprotein ; Embryo ; Blastoderm
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary Affinity purified preparations of the galactose-binding lectin from gastrulating chick blastoderms consist of three main polypeptides. Two of these have been identified as the 14 kD and 16 kD galactose-binding lectins. A third one migrates in SDS-PAGE gels with a relative molecular weight of 6,500±500 and has been identified as an apolipoprotein (Apo) of plasma very low density lipoproteins, Apo-VLDL-II. We have studied the localization of these polypeptides using immunofluorescence and ultrastructural immunocytochemistry with peroxidase and protein-A gold. The 14 kD lectin occurs in the intracellular yolk where it is mainly present within the electron lucent component. The 16 kD is also present in the intracellular yolk platelets, but tends to predominate in the electron-dense component. In addition, the 16 kD lectin is also present in pleiomorphic yolk-associated organelles and in the extracellular matrix. Apo-VLDL-II is also localized in the electron-lucent component of the yolk platelet and in the extracellular matrix. Our results suggest that the lectin(s) are associated with Apo-VLDL-II in the yolk platelet, and may subsequently become externalized.
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  • 88
    ISSN: 1432-041X
    Keywords: Specific gene expression ; Myosin heavy chain ; Muscle lineage cells ; Ascidian embryos
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Summary In ascidians the lineage of embryonic precursor cells is well documented. In this study, we investigated temporal and spatial expression of myosin heavy chain (MHC) gene during embryogenesis of the ascidianHalocynthia roretzi. When the occurrence of MHC transcripts was examined by Northern blot hybridization and in situ hybridization with specific cDNA and antisense RNA probes, transcripts were undetectable in fertilized eggs and cleavage-stage embryos. This suggests that the maternal message for MHC is not directly associated with the synthesis of MHC protein in ascidian embryos. MHC transcripts were initially observed at the gastrula stage. In situ hybridization of whole-mount preparations demonstrated that the transcripts first appeared in nuclei of primary-lineage muscle cells of the early-to-middle gastrula and accumulated rapidly as development progressed. The occurrence of MHC transcripts was restricted to differentiating muscle cells. No hybridization signal was detected in mesenchyme cells which are thought to form adult body-wall muscle. After metamorphosis the amount of MHC transcripts decreased; they became undetectable in juveniles about 10 days after metamorphosis, suggesting an intermission of MHC gene activity prior to the formation of adult bodywall muscle. Thus the expression of MHC gene in ascidian embryos was strictly regulated in both spatial and temporal orders and occurred only in muscle lineage cells.
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    Insectes sociaux 37 (1990), S. 58-72 
    ISSN: 1420-9098
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    Topics: Biology
    Description / Table of Contents: Resume L'exécution de deux tâches, butinage et production de cire, a été comparée pour des colonies d'abeilles mellifères issues d'essaims principaux et d'essaims secondaires. Dans les colonies d'essaims secondaires, les ouvrières ont tendance à commencer le butinage à des âges plus avancés. Ce delai est généralement accompagné d'une plus longue production de cire. Les différences dans l'exécution des tâches des deux sortes d'essaims démontrent l'incorporation d'un système flexible de castes temporelles dans la vie d'une colonie. Aucune relation n'a pu être observée entre le nombre d'abeilles dans les essaims et les âges auxquels les tâches furent exécutées. Il est donc suggeré que les interruptions d'élevage du couvain et des changements dans la distribution des âges vers des ouvrières plus âgées furent responsables des délais de l'ontogenèse des castes temporelles de colonies originaires d'essaims secondaires.
    Notes: Summary The performance of two tasks, wax production and foraging, was compared between colonies founded by prime swarms and afterswarms of honey bees. Workers began foraging at older ages in afterswarms and this delay was generally coupled with a longer duration of wax production. Differences in task performance between swarm types demonstrate the integration of a flexible behavioral caste system into life history. No relationships were found between swarm population size and ages of task performance, suggesting that breaks in brood rearing and upward shifts in worker age distribution were responsible for delays in the ontogeny of temporal castes in afterswarm-founded colonies.
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    Insectes sociaux 37 (1990), S. 73-89 
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Zusammenfassung Die Biologie zweierPolyrhachis-Arten, die dieArachne-Artengruppe bilden, wird beschrieben.Polyrhachis arachne Emery 1896 undPolyrhachis hodgsoni Forel 1902 sind auf das Leben auf breitblättrigen Bambusarten spezialisiert, ohne jedoch mit diesen in mutualistischer Symbiose zu leben. Sie tragen verschiedene Homoptera in Blattpavillons und halten sie dort. Die evolutiven Vorteile dieser Einnischung werden diskutiert.
    Notes: Summary The biology of two species ofPolyrhachis which together form thearachne species-group is described. BothPolyrhachis arachne Emery 1896 andPolyrhachis hodgsoni Forel 1902 are confined to broad-leaved species of bamboo, but there is no ant-plant mutualism. They collect Homoptera on bamboo plants and cultivate them in leaf pavilions. The evolutionary advantages of this type of ecological specialization are discussed.
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  • 91
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    Insectes sociaux 37 (1990), S. 101-115 
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resume Les modalités d'orientation utilisées dans le retour au nid sont étudiées au moyen de labyrinthes en Y dans 3 sous-familles de fourmis:Atta laevigata, Acromyrmex octospinosus, Acromyrmex rugosus, Trachymyrmex urichi, Ectatomma ruidum etPseudomyrmex termitarius. Les résultats montrent que les insectes utilisent plusieurs stratégies pour s'orienter dans le labyrinthe. Pour chaque espèce, les différentes modalités sont classées par ordre d'importance dans leur utilisation lors du retour au nid. Outre les repères visuels et chimiques, apparaît l'existence d'une orientation topochimique.P. termitarius utilise dans son orientation un sens kinesthétique. L'apprentissage des repères par la fourmi au laboratoire n'est pas en relation directe avec la hiérarchie des modalités d'orientation utilisées par chaque espèce dans le labyrinthe et apparaît en corrélation inverse avec la complexité sociale des espèces; le nombre de repères utilisés augmente par contre avec le degré évolutif de l'insecte. Une évolution divergente de ces stratégies d'orientation, même dans des espèces proches phylogénétiquement, est proposée.
    Notes: Summary The cues used for orientation during homing were examined in ants from 3 subfamilies, i.e.Atta laevigata, Acromyrmex octospinosus, Acromyrmex rugosus, Trachymyrmex urichi; Ectatomma ruidum andPseudomyrmex termitarius, by means of a Y-maze experiment. The results showed that ants use various cues for orientation through a maze. For each ant species, the cues may be classified in a kind of species-specific hierarchy according to their importance during homing. In addition to visual and chemical orientation, empirical evidence supporting the existence of a topochemical is presented.P. termitarius uses a kinesthetic sense for orientation. Learning of the cues in the laboratory is not directly related to the respective hierarchy used for orientation through a maze by each species, but seems to be inversely correlated with the social complexity of the species. The number of cues used for orientation correlates directly with the social complexity of the species. A divergent evolutionary development of orientation strategies, even among closely related species, is proposed.
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  • 92
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    Insectes sociaux 37 (1990), S. 131-145 
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resume L'activité alimentaire des fourmis champignonnistesAtta sexdens etA. cephalotes a été étudiée par l'observation de deux colonies de chaque espèce dans une forêt dense humide primaire d'Amazonie Centrale. Ces deux espèces utilisent des pistes marquées visuellement et chimiquement. Il a été démontré que l'activité alimentaire est orientée sur ces pistes: le nombre de nouveaux sites d'alimentation est inversement lié à la distance à une piste déjà existante. En conséquence, 15 à 23 % seulement du rayon du territoire d'alimentation des colonies est effectivement utilisé. Les deux espèces attaquent de manière préférentielle de grands arbres (diamètre du tronc à hauteur d'homme 〉20 cm). Cette préférence semble liée à la qualité du matériel végétal récolté et à la stratégie de recherche sur une aire restreinte employée par ces fourmis. Bien que ces fourmis aient exploité un grand nombre d'espèces végétales, l'activité alimentaire est concentrée sur un petit nombre de celles-ci. Les jeunes feuilles sont le principal élément exploité parA. cephalotes cependant queA. sexdens ne montre aucune préférence par rapport au type de matériel végétal récolté. Cette dernière espèce présente également une plus grande variation temporelle des taux d'alimentation qui est liée, en partie, au type de matériel récolté. Il est suggéré que le comportement non spécialisé présenté parA. sexdens est un des facteurs qui contribuent au fait qu'elle occupe un plus grand nombre d'habitats queA. cephalotes.
    Notes: Summary The foraging activity of the leaf-cutting antsAtta sexdens andA. cephalotes was studied observing two colonies of each species in a primary forest of the Central Amazon. These two ant species utilize chemically and visually marked trunk trails, and it was show that foraging is centered on these trails. The number of new foraging sites was inversely reated to the distance from existing trails. As a consequence, the area effectively exploited by the studied colonies was only 15 to 23% of their respective foraging ranges. Both species attacked large trees (stem diameter 〉20 cm) in a proportion higher to the available one and this preference seems to be related with quality of the harvested material and with the “area restricted search” strategy employed by these ants. Although these ants were able to exploit a wide range of plant species, harvest activity was concentrated on only a few. New leaves was the main item foraged byA. cephalotes whereasA. sexdens did not show a pattern of plant part preference.A. sexdens also presented a higher temporal variation in foraging rates, and this variation was partiallly related to the type of plant part harvested. It is suggested that the more generalistic behaviour presented byA. sexdens is one of the factors that contributes to its diverse habitat distribution as compared toA. cephalotes.
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  • 93
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    Insectes sociaux 37 (1990), S. 158-168 
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resume Des nids adultes deMacrotermes subhyalinus contenant 6,6 kg de poids sec d'adultes stériles, ont produit 2,35 kg de poids sec d'ailés chaque année. Les petits nids availent une plus grande proportion de larves, et une distribution de stades larvaires compatible avec une croissance rapide de la population du nid. Ils avaient aussi une plus grande proportion de soldats dans la population stérile adulte que les nids adultes. Les relations entre les dimensions extérieures du nid et les paramètres de la colonie sont approximativement linéaires, positives et significatives.
    Notes: Summary Mature nests ofMacrotermes subhyalinus contained a mean standing crop of 6.6. kg dry weight of sterile adults, and produced 2.35 kg dry weight of alates each year. Small nests contained a much larger proportion of larvae, and somewhat different proportions of the different larval instars, both features being consistant with a rapid increase in population. They also contained a larger proportion of soldiers in the adult population than in mature nests. Mound dimensions and nest parameters were all linearly related, and positively and significantly correlated.
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  • 94
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Summary In incipient laboratory colonies ofKalotermes flavicollis, the young female lays only after male stimulation. The laying goes on only if male and female hold together or if the male is replaced by larvae after the stimulation time. When the young mated female is isolated, it lays only its first eggs. A virgin female does not lay when kept only with additional larvae. The castrated male stimulates the female. The ablation of the male genital tract, after the first mating, has no consequence on the laying in the first months. When a young female has been stimulated by a castrated male, a few parthenogenetic larvae are born: they are capable of living.
    Notes: Resume Dans les jeunes colonies débutantes deKalotermes flavicollis, les femelles ne pondent que si elles ont été stimulées par un mâle. La ponte ne se poursuit que si le couple se maintient, ou si le mâle est remplacé par des larves après la période initiale, de stimulation. Une jeune femelle fécondée isolée, ne pond que ses premiers oeufs. Une femelle vierge accompagnée de larves ne pond pas. Un mâle castré stimule la femelle. L'ablation de l'appareil génital mâle, apès la première copulation, n'entraîne aucune conséquence sur la ponte dans les premiers mois. Les oeufs pondus par une femelle vierge stimulée par un mâle castré, peuvent donner des larves parthénogénétiques viables.
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  • 95
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resume Des expériences sur une piste circulaire artificielle montrent que le coléoptèreEdaphopaussus favieri suit la piste de sa fourmi hôte,Pheidole pallidula. Si le coléoptère a été préalablement maintenu isolé, la mise en présence d'ouvrières améliore son suivi de la piste. E. favieri réagit de façon optimale à une concentration de 0.03 glandes à poison d'ouvrière minor par cm. Les ouvrières minor et major sont respectivement 10 et 4 fois plus sensibles que le coléoptère à la phéromone de piste. Contrairement aux ouvrières, le coléoptère est repoussé par des concentrations égales ou supérieures à 0.06 glandes/cm. Cet effet répulsif s'observe aussi pour des glandes à poison et des glandes mandibulaires. entières, dePh. pallidula, ainsi quue pour les produits volatils de la glande de Dufour et certaines concentrations d'extraits de glandes à poison présentés ponctuellement en face du coléoptère.E. favieri suit uniquement la piste de son hôte. La valeur adaptative des différentes caractéristiques du suivi de la piste parE. favieri est discutée.
    Notes: Summary Laboratory experiments on circular artificial trails show that the beetleEdaphopaussus favieri follows the trail of its host ant,Pheidole pallidula. After having been isolated for a certain time, the beetle greatly improves its trail-following ability once it is again in contact with workers. The optimal response ofE. favieri occurs for a concentration of 0.03 minor worker's poison glands per cm. Minor and major workers are respectively 10 and 4 times more sensitive. High trail pheromone concentrations, from 0.06 glands/cm onwards, repel the beetle, not the workers. This avoidance effect is also found with wholePh. pallidula's poison and mandibular glands, with volatiles from the Dufour's gland and with poison gland extracts presented in the form of a dot, in front of the beetle. The trail following ofE. favieri is host-specific. The adaptive value of the trail-following characterisitcs ofE. favieri is discussed.
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  • 96
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Zusammenfassung C. borneensis (Myrmicinae) lebt in enger Gemeinschaft mit myrmekophytischen Arten der südostasiatischen PionierbaumgattungMacaranga (Euphorbiaceae). Die Ameise ist so eng an die Pflanze adaptiert, daß sie getrennt von ihr nicht lebensfähig ist. Die Nahrung beziehtC. borneensis in Form von Nährkörperchen ausschließlich von der Pflanze. Im Sproßachseninnern gehaltene spezifische Schildläuse bieten eine weitere Nahrungsquelle. Die Adaptationen erstrecken sich bis auf die Anatomie des Verdauungstraktes: Der Kropf ist sehr klein und kann nur geringe Nahrungsmengen speichern. C. borneensis besiedelt spezifisch nurMacaranga-Pflanzen. Sowohl Königinnen als auch Arbeiterinnen sind in der Lage, die Wirtspflanze zu erkennen, wobei offenbar chemische Reize eine Rolle spielen. Die koloniegründenden Königinnen schwärmen das gesamte Jahr über, das Schwärmen erfolgt überwiegend während der Dunkelheit. Um die besiedlungsfähigenMacaranga-Pflanzen herrscht ein starker Konkurrenzdruck. Die beteiligten Schildlausarten sind spezifisch für die Assoziation. Sie werden nicht von der Königin beim Hochzeitsflug mitgenommen. Die Arbeiterinnen sind tag- und nachtaktiv. Die meisten Tiere halten sich im jüngsten Drittel der Pflanze auf, wo sie patrouillieren und Nährkörperchen sammeln. Mittels eines spezifischen Säuberungsverhaltens entfernen die Arbeiterinnen alle Fremdobjekte von der Pflanze. Trotz ihrer geringen Größe attackieren die Ameisen eine Vielzahl phytophager Insekten und sind dabei besonders effektiv in der Abwehr kleiner, wenig sklerotisierter Tiere wie z.B. Raupen. Sie verfügen zwar nicht über einen funktionsfähigen Stachel, setzen aber wehrsekrete ein und beißen sich hartnäckig fest. Mit Hilfe eines effektiven Alarmierungssystems, das einen Massenangriff ermöglicht, gewährleisten sie eine Verteidigung ihrer Kolonien und damit gleichzeitig ihrer Wirtspflanze. Eine Vergleich mit einer anderenCrematogaster-Art demonstriert die besonderen Adaptationen vonC. borneensis an ihre Wirtspflanze.
    Notes: Summary C. borneensis (Myrmicinae) lives in close association with several myrmecophytic species of the South East Asian pioneer tree genusMacaranga (Euphorbiaceae). The ants are adapted to the plants so closely that they do not survive away from it. The only food they utilize is provided as food bodies by the plant and honeydew from specific scale insects kept inside the hollow internodes. The anatomy of the digestive tract is also adapted to life on the host plant: the crop is very small and can store only minute food quantities. C. borneensis exclusively colonizes certainMacaranga species. Queens as well as workers are able to recognize their host plant species, probably by chemical cues. Colony founding queens swarm throughout the year, mostly during darkness. There is strong competition among queens for host plants. Queens do not carry scale insects on their nuptial flight. Worker ants are active day and night. Most of them patrol and collect food bodies on the younger parts of the host plant. An important characteristic is their cleaning behaviour, which results in removal of all foreign objects. Even though they are rather small, workers respond very aggressively to certain kinds of disturbance of the host plant. The ants attack most phytophagous insects and are especially effective in killing and removing small, softbodied herbivores (e.g. caterpillars). They do not possess a functional sting, but apply defensive secretion and—once biting an intruder—will not let go. Their effective alarm system results in a mass attack, which provides adequate defence for the colony and the host plant. A comparison with anotherCrematogaster species further illustrated the special adaptations ofC. borneensis to its host plant.
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  • 97
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Summary The development of colonies in the paper wasp,Polistes dominulus, was studied in two experimental situations: in the first, the temperature and the daytime were progressively increased (situation A: 5 colonies); and in the second, these two factors were progressively decreased (situation B: 6 colonies). These two experimental situations give rise to different types of reproductive patterns. The duration of the annual cycle was found to be shorter with colonies raised under condition B (4–5 months) than with those raised under condition A (5–9 months). In the colonies with a shortened cycle (situation B), males and long lived females (250–500 days) were found to emerge earlier than in those with a longer cycle. In experimental situation B, the males emerged almost exclusively from eggs laid before the foundresses disappeared; whereas in the situation A, the male production occured later and the males were produced partly by the foundresses, and partly by the workers after the foundresses died.
    Notes: Resume L'étude du développement des colonies dePolistes dominulus dans deux situations environnementales distinctes, caractérisées l'une par un accroissement progressif de la température et de la durée du jour (situation A: 5 colonies), l'autre par une diminution progressive de ces deux facteurs (situation B: 6 colonies), a mis en évidence au niveau des guêpiers deux configurations reproductrices. La durée du cycle des colonies a été plus courte en situation B (4 à 5 mois) qu'en situation A (5 à 9 mois). Parallèlement au raccourcissement du cycle reproducteur, on a observé, dans la situation B, une apparition plus précoce des femelles à vie longue (250–500 jours) et des mâles. Concernant la production mâle, elle a résulté, dans la situation B, exclusivement d'œufs pondus avant la disparition des fondatrices, alors que, dans la situation A, la production mâle, plus tardive, a résulté en partie de la ponte des fondatrices, en partic de la ponte des ouvrières, après la disparition des reines.
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  • 98
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    Insectes sociaux 37 (1990), S. 268-269 
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    Topics: Biology
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  • 99
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    Insectes sociaux 37 (1990), S. 258-267 
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Resume Nous présentons une série d'expëriences qui montrent comment la fourmiLasius niger peut utiliser son système des recruitement par piste afin de sélectionner une dfes deux sources de nourriture. Sil on leur offire simultanément deux solution 1M de saccharose, la sojciét'e concentre son activité sur l'une des deux. Si l'on offre deux solutions, une de 1M et l'autre de 0.1M, elle sélectionne la plus riche, à mons que la piste qui mène à la source 0.1M soit déjà bien développée au moment où l'on introduit la source 1M. Face à la même situation, la fourmiTetramorium caespitum, qui recrute par groupe/masse, utilise son mode de transmission d'information plus, individuel pour changes son exploitation vers la source 1M. Un modèle mathématique décrit ces processus, dont la dynamique correspond bien aux observations expérimentales.
    Notes: Summary A series of experiments shows how the andLasius niger uses its trail recruitment system to select between two food sources. Simultaneously presented with to 1M sucrose solution it concentrates on one of them. When offered a 1M solution together with a 0.1M solution it selects the richer source, unless the trait to the 0.1M source had become well-developed before the 1M source was introduced. In the same situation, however, the group/mass recruiting antTetramorium caespitum uses its more individual transmission of information to switch to the 1M source. A mathematical model describes these processes and its dynamics reflect the experimental results.
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  • 100
    ISSN: 1420-9098
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Description / Table of Contents: Summary The respiration of both castes ofFormica polyctena (Foerster) during the larval development was investigated manometrically. The distribution of mitochondria and their structure in the fat body cells were measured simultaneously by electron-microscopy to ascertain if caste-specific differences of the intensity of respiration were recognizable by the content of mitochondria. The following results were obtained: At 20°C larvae of sexuals weighing 1 to 5 mg per individual consumed 860–910 μl O2/g/h, which was double the oxygen amount used by larvae of workers weighing up to 1.5 mg per individual. At the end of the larval development this difference disappeared, with values stabilizing at 410–420 μl O2/g/h. The volume density of mitochondria in 8 mg-sexual larvae was 12.2%, and in 60 mg-larvae or 1.9 mg — and 12.5 mg — worker larvae only 8.2–8.6% of the cytoplasm (without nucleus). The numeric density of the mitochondria was calculated as 840 per mm3 in larvae of 8 mg-sexuals, and in those of 60 mg or in 1.9 mg — and 12,5 mg — worker larvae only as 570–625 per mm3. No caste-specific and stage-dependent differences could be found in the size and microstructure of the mitochondria. An increased respiration was associated with a higher numeric and volume density of the mitochondria.
    Notes: Zusammenfassung Die Atmung wurde während der Larvalentwicklung beider Kasten vonFormica polyctena (Foerster) manometrisch untersucht. Parallel dazu wurde nach elektronenoptischer Präparation die Mitochondrienverteilung und Mitochondrienstruktur in Fettkörperzellen ermittelt, um festzustellen, ob kastenspezifische Unterschiede in der Atmungsintensität auch im Mitochondriengehalt zu erkennen sind. Folgende Resultate wurden erhalten: Geschlechtstierlarven von 1–5 mg/Tier verbrauchten mit 860–910 μl O2/g/h bei 24°C mehr als doppelt so viel Sauerstoff wie Arbeiterinlarven bis 1,5 mg/Tier. Am Ende der Larvalentwicklung war dieser Unterschied bei 410–420 μl O2/g/h ausgeglichen. Die CO2-Abgabe verhielt sich ähnlich. Die Volumendichte der Mitochondrien betrug bei 8 mg-Geschlechtstierlarven 12,2%, bei solchen von 60 mg sowie Arbeiterinlarven von 1,9 mg und 12,5 mg nur 8-2-8,6% des Cytoplasmas (ohne Kern). Die numerische Dichte der Mitochondrien betrug bei 8 mg-Geschlechtstierlarven 840/mm3, bei solchen von 60 mg sowie Arbeiterinlarven von 1,9 mg und 12,5 mg nur 570–625/mm3. In der Größe und Feinstruktur der Mitochondrien konnten keine kastenspezifischen und stadiumabhängigen Unterschiede festgestellt werden. Eine erhöhte Atmungstätigkeit war verbunden mit höherer numerischer und Volumendichte der Mitochondrien.
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