ALBERT

Beschreibung: Changes in tree form and taper over time, as affected by changes in tree, stand, and site factors for interior lodgepole pine (Pinuscontorta var. latifolia Engelm.) were investigated using detailed stem analysis data from interior British Columbia. It was found that tree shape and taper change along the stem at one time and over time with changes in tree and stand factors, particularly the diameter at breast height to total tree height ratio, crown length, and crown ratio, and with predicted quadratic mean diameter at age 50 years, a stand density measure. At young ages, the trees were parabolic in shape from ground to top. However, as they increased in size over time, different portions of the stem took different shapes because of unequal growth in diameter along the stem. Changes in tree shape and taper over time were closely related to the crown size, which is related to stand density.

Beschreibung: Estimates of individual-tree narrow-sense heritability and additive genetic coefficient of variation of seven traits of forest trees were compiled from 67 published papers. Distributions of the values for each trait were characterized and compared by calculating medians and running Kolmogorov–Smirnov and Wilcoxon signed-rank tests. Generalizations are possible about at least some of the traits examined. Heritability of wood specific gravity was almost always above 0.3 (median 0.48). Heritabilities for other traits tended to be low: medians ranged from 0.185 to 0.26, and individual values generally ranged from 0.1 to 0.4. Evidence that heritabilities of form traits tend to be higher than those of growth traits was weak. The analysis of additive genetic coefficients of variation suggested that specific gravity tends to have lower values than other traits (median 5.1%), while height and diameter (medians 8.5 and 8.6%, respectively) had lower values than straightness (median 11.65%). Individual-tree volume showed the highest levels of additive genetic coefficient of variation (median 20.3%). The levels of additive genetic variation and heritabilities suggest that reasonable levels of genetic gain can be achieved by screening relatively low numbers of trees.

Beschreibung: By measuring incident precipitation, throughfall, and stemflow chemistry, the roles of coniferous- and deciduous-dominated forest canopies as a source of and sink for ions in precipitation were examined. A regression technique for distinguishing between external (dry deposition) and internal (canopy leaching) sources of ions in the throughfall flux was evaluated. The effect of seasonal changes in the forest canopy on throughfall and stemflow chemistry was also examined. Throughfall comprised 74 and 84%, respectively, of the hydrologic flux at the coniferous and deciduous sites. Sulphate fluxes were highest at the coniferous site during both growing and dormant seasons, suggesting either a higher scavenging efficiency of the needles for atmospheric SO42−, or higher SO42− leaching from the foliage. The deciduous site neutralized acidic inputs, as demonstrated by its net negative H+ flux year round. The buffering capacity of the coniferous forest was exceeded by the higher amount of acid interception by the canopy. Nitrate behaved conservatively and base ions were exported from the canopy. Stemflow contributions of ions, although low, were generally higher than the contribution of stemflow to the hydrologic flux (2–3%). Independent dry deposition measurements for the growing season, when compared with net SO42− flux, overestimated dry deposition collected by the deciduous canopy, but were comparable to the flux at the coniferous site. These data suggest that dry SO2−SO42− deposition may be responsible for all SO42− enrichment seen in throughfall at these sites. A regression technique for separating internal and external ion sources in throughfall yielded inconsistent results, and attributed virtually all ion enrichment to internal sources. Problems with false assumptions and spurious correlations are discussed. We conclude that this method is not satisfactory for separating ion sources. Seasonal patterns in throughfall chemistry are present. During the growing seasons bases exchange for H+ and are exported similarly with SO42−. Hydrogen retention mirrors SO42− export. Base cations (particularly K+) are leached from the canopy primarily during senescence, but from the stem of the tree primarily during the dormant period. This was most evident at the deciduous site. Chloride behaved in a similar manner, while NH4+ and H+ were retained during the senescent period.

Beschreibung: A simple system for the estimation of stem volume is presented based on the compatible stem profile and volume equations. This system can directly predict the stem volume above breast height from measurements of stem diameter at breast height and at an another point along the upper stem, and does not require any sample data for determining a parameter of volume equation. In comparison with the prediction accuracy of existing volume equations from the literature, using data from Cryptomeriajaponica D. Don, Chamaecyparisobtsusa Endl., and Pseudotsugamenziesii (Mirb.) Franco, this system has the advantage of reducing prediction error.

Beschreibung: Thirty years of cone production records for subalpine fir (Abieslasiocarpa (Hook.) Nutt.) and mountain hemlock (Tsugamertensiana (Bong.) Carr.) (two sites each) in the Cascade Mountains of Washington and Oregon were compared with basal area increment and weather records to determine relationships among weather, radial growth, and cone crop. Results show that the size of subalpine fir cone crops was negatively related to large crops and positively related to radial growth in the previous 2 years. Mountain hemlock cone crops were negatively related o a large cone crop and positively related to July or August temperature in the previous year. Radial growth in heavy cone years was inhibited more for subalpine fir than for mountain hemlock. Results are explained by differences in the location of cone production between species. It is concluded that global climate warming could result in fewer and more irregular cone crops for these species.

Beschreibung: We describe a visual survey technique for evaluating acorn production. In contrast with previously proposed methods, our technique yields ratio-level data on annual productivity that are analyzable with standard statistics and, by sampling the same trees each year, data on the reproductive patterns of individual trees. We compared this technique with two independent sets of acorn-trap data acquired on oaks of three species at Hastings Reservation in central coastal California. Correlations between acorns counted by the visual surveys and collected from acorn traps under the same trees were significant for all three species. Most scatter in the data appeared to be attributable to three causes: (1) sampling error, especially among trees with very small crops, (2) finite counting speed, leading to a lack of discrimination among trees with very large crops by the visual surveys, and (3) arboreal acorn removal by animals. This latter factor can be particularly large, rendering visual surveys more reliable than the use of traps. Furthermore, only the high efficiency of visual surveys allows for the practical assessment of samples large enough to accommodate high within-population variation and detect widespread geographic variation in acorn production. Visual surveys offer a method of assessing the fruit or cone crops of many hardwood and conifer species that is not only more efficient but also more accurate than the use of traps.

Beschreibung: Geostatistics provides tools to model, estimate, map, and eventually predict spatial patterns of tree size and growth. Variogram models and kriged maps were used to study spatial dependence of stem diameter (DBH), basal area (BA), and 10-year periodic basal area increment (BAI) in an old-growth forest stand. Temporal variation of spatial patterns was evaluated by fitting spatial stochastic models at 10-year intervals, from 1920 to 1990. The study area was a naturally seeded stand of southwestern ponderosa pine (Pinusponderosa Dougl. ex Laws. var. scopulorum) where total BA and tree density have steadily increased over the last decades. Our objective was to determine if increased stand density simply reduced individual growth rates or if it also altered spatial interactions among trees. Despite increased crowding, stem size maintained the same type of spatial dependence from 1920 to 1990. An isotropic Gaussian variogram was the model of choice to represent spatial dependence at all times. Stem size was spatially autocorrelated over distances no greater than 30 m, a measure of average patch diameter in this forest ecosystem. Because patch diameter remained constant through time, tree density increased by increasing the number of pine groups, not their horizontal dimension. Spatial dependence of stem size (DBH and BA) was always much greater and decreased less through time than that of stem increment (BAI). Spatial dependence of BAI was close to zero in the most recent decade, indicating that growth rates in 1980–1990 varied regardless of mutual tree position. Increased tree crowding corresponded not only to lower average and variance of individual growth rates, but also to reduced spatial dependence of BAI. Because growth variation was less affected by intertree distance with greater local crowding, prediction of individual growth rates benefits from information on horizontal stand structure only if tree density does not exceed threshold values. Simulation models and area estimates of tree performance in old-growth forests may be improved by including geostatistical components to summarize ecological spatial dependence.

Beschreibung: Application of base-cation fertilizers has been shown to increase tree growth and vigour in declining sugar maple (Acersaccharum Marsh.) stands in southern Quebec but little is known about the effects of such fertilizers on litter quality or decomposition. Sugar maple foliage litters from fertilized and unfertilized plots on a base-poor site and from a naturally base-rich site were incubated in litterbags of 1- and 3-mm mesh sizes on fertilized and unfertilized plots at the base-poor site. Mass loss of unfertilized litter was slower in fertilized than in unfertilized plots, suggesting a negative effect of fertilization on the decomposer community. Faster mass loss of fertilized than unfertilized litter incubated in the same plot indicated that changes in litter quality brought about by fertilization enhanced decomposition. Mass loss of fertilized litter on fertilized plots did not differ from that of unfertilized litter on control plots, indicating that although decomposition processes are affected by fertilization the overall effect on decomposition is negligible. Mass loss was significantly, but only slightly, higher in large mesh than in small mesh bags indicating that larger soil fauna play a limited role in litter decomposition in this forest.

Beschreibung: Radial growth patterns, canopy recruitment characteristics, and disturbance histories were examined in a shade-tolerant species, Nyssasylvatica Marsh., and a shade-intolerant species, Liriodendrontulipifera L., to determine the influence of canopy gaps in species with contrasting life histories. Tree cores of these co-occurring species were taken from three mixed-Quercus forests in northern Virginia. Most N. sylvatica individuals became established prior to 1850 and experienced multiple release and suppression periods coinciding with logging during the late 1800s and early 1900s. Many L. tulipifera became established during the early 1900s following logging, and only a few individuals experienced prolonged suppression periods. Regardless of site, L. tulipifera grew faster than N. sylvatica (average radial growth 〉1.70 mm/year for L. tulipifera vs. 2 mm (15–37 years for L. tulipifera vs.

Beschreibung: We test one of the fundamental assumptions of most dendroclimatological research, that the radial growth response of trees to climate does not vary with age once the biological growth trend has been removed. Piceaglauca (Moench) Voss from three sites in the western Northwest Territories, Canada, are disaggregated into age-classes, and their response to climate examined through response function and linear regression analyses. These data are then used in multiple regression analyses to estimate June-July temperatures at Norman Wells, N.W.T., from 1909 to 1989 using both age-dependent and standard (age-independent) models. The response function and regression analyses suggest that the response of Piceaglauca radial growth to climate differs between trees greater than 200 years old and less than 200 years old. These results suggest that the assumption of an age-invariate climate–growth function is therefore invalid at these sites. These apparent age-dependent responses are site specific and may reflect physiological changes related to aging. One possible causal factor of age dependence is that the trees are becoming increasingly moisture stressed with age owing to a reduction in the efficiency of water and nutrient translocation mechanisms. The estimation of June–July temperatures based on an age-dependent model produced improved calibration and verification statistics as compared with a reconstruction based on standard dendroclimatic modelling. If present, age-dependent climate–growth relationships may result in less accurate reconstructions of past climate, particularly during the early portions. However, age-dependent responses could also be used to increase the number of tree-ring based predictor variables for dendroclimatic reconstructions.

Beschreibung: The reliability of a leaf-disk assay to assess resistance of Populusdeltoides Bartr. to Melampsoramedusae Thuëm. f.sp. deltoidae, the causal agent of poplar leaf rust, was evaluated. Leaf disks of eight host genotypes were inoculated in a spore settling tower with 11 isolates of the pathogen in all possible combinations, and the latent period, infection probability, sporulation, and progeny/parent ratio were recorded. Correlations were established between these measures and measures derived from field epidemics, namely the relative area under the disease progress curve, the apparent rate of infection, the final disease severity, and the number of days before defoliation. Four genotypes were highly resistant to all 11 isolates tested with the leaf-disk assay and to the local inoculum in field tests. Three genotypes were highly susceptible in inoculation assays and were also susceptible in the field. One genotype was highly resistant in inoculation assays but had intermediate resistance in the field. Significant correlations (p 〈 0.05) were found between all leaf-disk and all field parameters.

Beschreibung: A combination of antiaggregation and aggregation pheromones was tested for protecting stands of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) at high risk for infestation by the Douglas-fir beetle (Dendroctonuspseudotsugae Hopkins). The antiaggregation pheromone, 3-methylcyclohex-2-en-1-one (MCH), was applied in a bubble capsule formulation to the perimeter of 1-ha circular plots at a rate of 60 g/plot. Treated plots also had three or four clusters of four Lindgren funnel traps baited with frontalin, seudenol, 1-methylcyclohex-2-en-1-ol, and ethanol located outside of the plot but within 160 m of the boundary. Mean (±SE) accumulated catches in all traps per plot were 73 658 ± 19 721 Douglas-fir beetles and 12 892 ± 2 513 Thanasimusundatulus (Say), a predator of the Douglas-fir beetle. The mean percentage of Douglas-fir trees ≥20 cm DBH that were mass attacked was reduced by 80% within the treated plots compared with the untreated plots. However, there was an eightfold increase in the percentage of trees mass attacked in the area outside the treated plots in the vicinity of the funnel traps. The net effect of the treatment was to concentrate mass-attacked trees within a limited area outside of the protected stand. Our results indicate that Douglas-fir beetle antiaggregation and aggregation pheromones can be used effectively to reduce the probability of infestation in small, high-value stands.

Beschreibung: Subalpine fir (Abieslasiocarpa (Hook.) Nutt.) regeneration following fire was studied at two locations that burned in 1902 northeast of Mount Rainier, Washington. Tree establishment dates were compared with local climatic records using multiple and logistic regression to identify potential relationships between seasonal climate and annual tree establishment. The influence of microsite features on forest regeneration was also explored. Little regeneration occurred in the first 30 years after the fires, and most trees established in the 1950s, 1977, 1983, and 1989. The dominance of trees

Beschreibung: This study presents an individual tree height prediction model for white spruce (Piceaglauca (Moench) Voss) and trembling aspen (Populustremuloides Michx.) grown in boreal mixed-species stands in Alberta. The model is based on a three-parameter Chapman–Richards function fitted to data from 164 permanent sample plots using the parameter prediction method. It is age independent and expresses tree height as a function of tree diameter, tree basal area, stand density, species composition, site productivity, and stand average diameter. This height-prediction model was fitted by weighted nonlinear regression for spruce and unweighted nonlinear regression for aspen. Almost all estimates of parameters were significant at α = 0.05 and model R2-values were high (0.9192 for white spruce and 0.9087 for aspen). No consistent underestimate or overestimate of tree heights was evident in plots of studentized residuals against predicted heights. The model was also tested on an independent data set representing the population on which the model was to be used. Results showed that the average prediction biases were not significant at α = 0.05 for either species, indicating that the model appropriately described the data and performed well when predictions were made.

Beschreibung: Beech bark disease occurs when either Nectriagalligena Bres., or Nectriacoccinea var. faginata Lohman, Watson, and Ayers kills bark that is or has been infested and altered by the beech scale, Cryptococcusfagisuga (Lind.). Introduced to Nova Scotia around 1890, this insect now occurs as far southwest as Ohio, West Virginia, Virginia, and North Carolina. I determined the relative occurrence of the two pathogens in forests affected for varying times and the temporal changes in their relative occurrence in recently affected stands of the Monongahela National Forest, West Virginia (MNF). Stands in the Canadian Maritimes, New England, New York, and Pennsylvania were sampled in 1985–1986; and in the Catskill Park, New York in 1988 and 1991. Stands on the MNF were sampled in 1982, 1983, 1984, 1986, 1988, and 1991. The following trends were indicated: N. galligena dominated recently affected stands in Pennsylvania and West Virginia, and N. coccinea var. faginata dominated long-affected stands in Canada, New England, and New York. On the MNF, only N. galligena was isolated in 1982; by 1988, N. coccinea var. faginata was isolated from 8 of 16 stands and from all 16 stands in 1991. Stands where N. galligena persisted had higher proportions of tree species highly susceptible to N. galligena than stands dominated quickly by N. coccinea var. faginata. Once present, however, N. coccinea var. faginata eventually will replace N. galligena as the dominant pathogen.

Beschreibung: The problem of fitting height–diameter curves for repeated measurements on growth plots is addressed. The context of the problem is fitting historical data with varying sampling protocols and varying measurement accuracy. A key consideration is obtaining good estimates of top height and top-height increment. A particular model and objective function for fitting are presented. The model has two parameters for each measurement and one common parameter; limited crossovers in the height–diameter curves for the various measurements are allowed. The objective function minimizes errors in predicted height and in predicted change in height. The programming is described, and the availability of code is announced. Examples show both the strengths and weaknesses of this approach.

Beschreibung: Reciprocal cross pollinations between western larch (Larixoccidentalis Nutt.) and alpine larch (L. lyallii Parl.) were done in Spring, 1991. The cross to alpine larch females was highly successful; 63% of the seeds developed mature embryos and 79% germinated. However, the cross to western larch females resulted in only 4% filled seed of which 68% germinated. Open-pollinated western larch and alpine larch averaged 26 and 32% filled seed with 48 and 44% germination, respectively. Less than 1% of the alpine larch hypocotyls were reddish in color; most of them were green. About 15% of the western larch and about 17% of the hybrids from alpine larch females had reddish hypocotyls, whereas hybrids from western larch females were intermediate between western larch and alpine larch. Hybrid seed began germinating before the parental types and initial height growth exceeded that of the parental types. However, 4 months following germination, western larch seedlings were ca. four times taller than the hybrids and 10 times taller than alpine larch. Stems of hybrids were significantly thicker than those of either western larch or alpine larch. Hybrid seedlings are robust, stocky, and may be useful in revegetating cold, moist sites between the elevational ranges of alpine and western larch.

Beschreibung: The effects of DiPel® 176, a commercially available Bacillusthuringiensis Berliner subsp. kurstaki (B.t.k.) formulation, on microbially mediated carbon and nitrogen mineralization processes, and the persistence of B.t.k. following application of DiPel® 176 to an acidic, coniferous forest soil were evaluated in the laboratory using simple microcosms. Litter (L) and fermentation–humus (FH) material were exposed to DiPel® at the recommended field application rate (FA), DiPel® at 1000× the field application rate (1000× FA), or left untreated. Respiration, substrate induced respiration (SIR), microbial biomass C, metabolic quotients (qCO2), NH4-N, NO3-N, cellulose decay, and B.t.k. viability were monitored regularly over 8 weeks. The FA treatment had no significant impact on soil processes in either the L or FH. The 1000× FA treatment increased SIR and biomass C and decreased qCO2 consistently in both the L and FH. No other effects of the 1000× FA treatment were evident in the L, while in the FH this treatment stimulated respiration initially, then reduced it below control levels; it enhanced cellulose decay; and it inhibited ammonification and nitrification after 8 weeks incubation. In both the L and FH there was no significant loss in viability of B.t.k. in either of the DiPel® treatments over 8 weeks. The microcosms used in this study were simple, inexpensive, and effective, with respiration, SIR, biomass C, and qCO2 being the least variable measurements and the most sensitive to perturbation. This approach is recommended for ecotoxicological and fate testing as outlined in the GuidelinesforRegistrationofNaturallyOccurringMicrobialPestControlAgents.

Beschreibung: Microbial isolates from conifer foliage and strawberry were evaluated for biocontrol of Botrytiscinerea Pers.:Fr. in container-grown seedlings of black spruce (Piceamariana (Mill.) B.S.P.). In growth room assays, seedlings were environmentally predisposed to the pathogen and coinoculated with the isolates and B. cinerea. Sporulation incidence of the pathogen was later estimated on needle segments that were cut from the seedlings and incubated on paraquat–chloramphenicol agar medium. The isolates suppressed sporulation by 0–100%. Those of Alternaria, Cladosporium, Epicoccum, yeasts, and bacteria were of low to moderate effectiveness, but those of Gliocladium, Myrothecium, Trichoderma, and Trichothecium were moderately to highly suppressive. In two greenhouse tests, spore suspensions of Gliocladium roseum Link:Bainier and of Myrotheciumverrucaria (Alb. & Schw.) Ditm. ex Stendel. (applied three times) at 2- to 3-week intervals suppressed incidence of B. cinerea by 50–69% and 42–60%, respectively, and were at least as effective as recommended fungicide treatments. Two or three additional applications of the antagonists did not further suppress the pathogen. Isolates of Fusarium sp. and Penicillium sp. were generally ineffective in the greenhouse tests. The four fungi ranked similarly in biocontrol effectiveness in the growth room and greenhouse. It is concluded that the growth room assay provided a strong indication of biocontrol performance in the greenhouse and that G. roseum and M. verrucaria have potential for managing B. cinerea in black spruce in greenhouses.

Beschreibung: Four species of boreal forest conifers share a similar southern limit of natural distribution in the three Prairie Provinces of western Canada. The southern boundaries of boreal forest and aspen parkland were compared with geographic patterns of several climate variables to provide a preliminary assessment of how global climate change could affect forest distribution in the future. Forest zonation corresponded most closely with climatic moisture regimes (annual precipitation minus potential evaporation). In contrast, thermal characteristics of climate (mean July and annual temperature, growing degree-days) showed an inconsistent relationship with forest zonation. It is postulated that moisture limitations prevent conifer regeneration south of the present limit of natural distribution. Alternatively, the more arid climates south of the boreal forest may have promoted higher fire frequencies historically, thus preventing conifers from achieving sufficient longevity to regenerate. The driest areas of boreal forest in the region occur at low elevations in west-central Manitoba, throughout Saskatchewan and Alberta, and the southwestern Mackenzie District, Northwest Territories; these areas may be most vulnerable to increased climatic dryness. Climatically induced losses of forest cover from these low-elevation areas could eventually lead to the fragmentation of the boreal forest in western Canada.

Beschreibung: Parent and preflight-adult Douglas-fir beetles (Dendroctonuspseudotsugae Hopk., Coleoptera: Scolytidae) were collected from felled Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) trees. Fungi isolated from the beetles included blue-stain fungi, such as Ophiostomapseudotsugae (Rumbold) von Arx, Leptographiumterebrantis Barras & Perry, and Leptographiumabietinum (Peck) Wingfield; other ascomycetes, such as Graphium spp. and Leptographium spp.; yeasts; and unidentified basidiomycetes. All fungal cultures derived from parent beetles, preflight adults, beetle eggs, larvae, frass, and bark from galleries had yeasts present. Ophiostomapseudotsugae was isolated from 67% of the parent females, 19% of the parent males, and 100% of the preflight adults of both sexes collected from their pupal cells. Rinsing the beetles with 70% ethanol prior to fungus isolations reduced the frequency of all fungi, except yeasts. Parent and preflight adult beetles (of both sexes) were examined with a scanning electron microscope, where spores of O. pseudotsugae were visible in shallow pits on the elytra of both male and female beetles. Spores of other fungi were observed in shallow pits on elytra and in much smaller but deeper pits on the scutellum of both sexes. The results indicate a close association between Douglas-fir beetles and fungi, accompanied by anatomical differentiation on the beetles that allows the dissemination of blue-stain fungi to new Douglas-fir hosts and substrates.

Beschreibung: The biochemical bases of water-stress tolerance in a pedigree consisting of black cottonwood (Populustrichocarpa Torr. & Gray female) and eastern cottonwood (Populusdeltoides Bartr. male) parental clones and four hybrid progeny were investigated. Trees were grown outdoors in pots; well-watered trees (soil water potential greater than −0.03 MPa) were kept moist in trays, and stressed trees (soil water potential less than −2.0 MPa) were subjected to repeated cyclical stress of 1 or 2 days duration over the 14-week study. Analysis of the major metabolites and ions in fully expanded leaves demonstrated that the greatest degree of osmotic adjustment was displayed by male hybrid 242, the P. deltoides male parent, and male hybrid 239 to a lesser extent. Osmotic adjustment in leaves of both hybrid 242 and the P. deltoides male parent was primarily constituted by malic acid, K, sucrose, and glucose, with the same metabolites also increasing in fine roots of hybrid 242, the only clone to display osmotic adjustment in roots. Female clone 240 and P. deltoides displayed organic solute-based adjustments to water stress that were offset by declines in inorganic ions, particularly Na and Mg. Given that the P. trichocarpa female parent did not display osmotic adjustment in either tissue, the hybrids' capacity for adjustment was likely conferred by the P. deltoides male parent.

Beschreibung: Some physical, thermal, and chemical properties of bark of 16 tree species native to the central hardwood region were measured to determine their potential to protect the vascular cambium from damage by fire. The relationship between DBH and bark thickness for each of 16 species was determined. For purposes of monitoring seasonal trends, two species (Quercusmacrocarpa Michx. and Acersaccharinum L.) were sampled periodically during one growing season. Temperature response to bark surface heating of 11 species was monitored at the cambial layer during simulated fires conducted in the field. Bark samples were analyzed for moisture content, specific gravity, dry weight, volatile matter content, and time until ignition. Overall, during simulated fires, temperature gradients were decreased and maximum cambial temperatures were reduced as bark thickness increased. Thick-barked species had lower maximum cambial temperatures, longer times to reach peak temperatures, slower rates of heat loss, and shorter time until surface ignition. Populusdeltoides Marsh, was the most heat resistant among species tested, while Acersaccharinum was the least. Higher specific gravities were associated with higher rates at which cambial temperatures rose as well as with increased time required for surface ignition.

Beschreibung: We tested the hypothesis that naturally occurring nitrogen (N) isotope ratios in foliage (from plants that do not symbiotically fix atmospheric N2) are an indicator of soil N dynamics in forests. Replicate plots were established at eight locations ranging in elevation from 615 to 1670 m in Great Smoky Mountains National Park in eastern Tennessee, U.S.A. The locations selected ranged from N-poor (low-elevation) to N-rich (high-elevation) forest stands. Soils were sampled in June 1992; plants, forest floors, and upper mineral soils were sampled in August 1992. Net N mineralization and net nitrification potentials for surface mineral soils and organic matter layers at each site were determined by aerobic laboratory incubations. Soils and organic layers from high-elevation sites had greater net N mineralization and nitrification potentials than soils from low-elevation sites. There were significant (P ≤ 0.05) differences between study sites in soil 15N abundance. Therefore, we examined correlations between measures of soil N availability and both mean foliar δ15N values and mean enrichment factors (εp−s = δ15Nleaf − δ15Nsoil). In evergreens, maples, and ferns, mean foliar δ15N values and mean enrichment factors were positively correlated with net N mineralization and net nitrification potentials in soil. The observed relationships between natural 15N abundance in plant leaves and soil N availability were explained by a simple model of soil N dynamics. The model predicts how the isotopic composition of plant N is affected by the following factors: (i) varying uptake of soil NH4-N and NO3-N, (ii) the isotopic composition of different soil N pools, and (iii) relative rates of soil N transformations.

Beschreibung: A 1992 study of serotiny in lodgepole pine (Pinuscontorta Dougl. ex Loud. var. latifolia Engelm.) in Yellowstone National Park asked four questions: (i) are there morphological characteristics that can be used to estimate pre-fire proportion of serotinous trees in forests that burned in 1988?; (ii) at what spatial scale does percent serotinous trees vary across the landscape?; (iii) which environmental factors are correlated with serotiny?; and (iv) what is the relationship between prefire serotiny and postfire lodgepole pine seedling density? We first sampled cone characteristics in serotinous and nonserotinous trees along four 2950-m transects in unburned forests, and examined burned trees nearby. Results indicated that asymmetrical cones and an acute angle of cone attachment to the branch were reliable indicators of serotiny even in burned trees. We then sampled nine patches of lodgepole pine forest that had burned in 1988, and varied in size from 1–3600 ha. We sampled serotiny at varying intervals along two perpendicular transects that crossed in the center of each patch. At each sample point, the 12 nearest canopy lodgepole pines were classified as serotinous or nonserotinous. We concluded that the percentage of serotinous trees is most variable at intermediate scales of 1–10 km, and is relatively homogeneous at both fine scales (

Beschreibung: Silvicultural practices such as thinning and fertilization can affect both canopy foliage quantity and distribution, altering stand growth. The objectives of this research were to quantify the effects of tree size and silvicultural treatment on the vertical distribution of foliage of individual trees of loblolly pine (Pinustaeda L.) and to estimate foliage quantity and distribution using easily measured tree data. In three stands sampled in North and South Carolina, fertilization and (or) thinning treatments had been applied 2 years prior to sampling. A fourth stand was untreated. Nonlinear and linear regression models were developed to test the effects of silvicultural treatment on individual branch foliage biomass and whole tree foliage biomass. Vertical distributions of foliage and branches were modelled using a Weibull probability density function. Analyses indicated that individual branch foliage biomass was positively related to branch size but negatively related to distance from the top of the tree. Fertilization with nitrogen and phosphorus, or thinning, increased the foliage biomass carried by a given sized branch. Silvicultural treatment effects on individual branches translated into whole-tree foliage biomass with thinning and fertilization increasing the crown size of individual trees. Though treatment affected crown size, the distribution of foliage (and branches) remained unaffected. Because silvicultural treatments change the size of crowns for trees of given dimensions, any estimation of loblolly pine crown biomass must be site and treatment specific.

Beschreibung: The level of genetic diversity of natural populations of eastern white pine (Pinusstrobus L.) from Quebec was estimated from allozyme variants of 18 loci coding 12 enzyme systems. On average, a white pine population was polymorphic at 50.6% of loci, had 1.96 alleles and 1.22 effective alleles per locus, and observed and expected heterozygosities of 0.176 and 0.180, respectively. The level of genetic diversity was lower in the populations of the St. Lawrence lowlands than in those of western Quebec. This observation will help in guiding the selection program of the eastern white pine improvement program under way in Quebec. Genetic differentiation among sampled populations was weak and accounted for only 2% of the total diversity. The estimate of gene flow was very high, resulting in low values for genetic distances among populations. Only one locus showed a heterogeneity of allelic frequencies among populations after the Bonferroni procedure was applied for simultaneous statistical tests. A cluster analysis based on genetic distances among populations revealed that the Anticosti and Abitibi populations, located at the limit of the natural range of white pine, were similar to populations from regions that were geographically the most distant.

Beschreibung: Crown class, site class, and breast-height age were incorporated into Kozak's variable-exponent taper equation (A. Kozak. 1988. Can. J. For. Res. 18: 1363–1368) for three species: Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco), western red cedar (Thujaplicata Donn), and aspen (Populustremuloides Michx.). For lodgepole pine (Pinuscontorta Dougl.), crown ratio, breast-height age, and quadratic mean diameter were incorporated into Kozak's taper equation. The effects of adding these variables to the exponent part of the taper equation on the prediction abilities of the taper model were assessed for prediction of diameter inside bark along the stem, total tree volume, and tree merchantable height. It was found that apart from the use of crown ratio for lodgepole pine, the additional variables resulted in only marginal improvements to the published version of Kozak's taper function. Therefore, the cost of measuring these additional variables is not justifiable.

Beschreibung: This study investigated the effect of different cold storage conditions on (i) root freezing tolerance of Norway spruce (Piceaabies (L.) Karst.) and Scots pine (Pinussylvestris L.) and (ii) the vitality of seedlings that suffered freezing injury to roots prior to storage. Container-grown seedlings, 1 year old, were stored from the end of October to April in three environments with different root temperatures: outdoor storage (−0.5 to 11.0 °C), cool storage (0.7 to 3.7 °C), or frozen storage (−5.0 to −3.8 °C). Root freezing tolerance was determined prior to storage in October and during storage in January and March. Maximum root freezing tolerance for both species occurred in January, when over 50% of spruce and pine seedlings survived 2 h exposure to −25 and −20 °C, respectively. At this time, roots of frozen-stored spruce were significantly more freezing tolerant than outdoor-stored seedlings, whereas storage environment had no significant effect on pine. Freezing tolerance in roots of both species decreased from January to March in all test environments but to a lesser extent at the subzero temperatures in the frozen storage. Root freezing to −10, −15, or −20 °C in late October before storage resulted in reduced poststorage survival of seedlings in April. Pine was more adversely affected (0–13% survival) than spruce (0–85% survival). Freezing of roots prior to storage caused the lowest survival with frozen storage.

Beschreibung: The crown development of red pine (Pinusresinosa Ait.) plantations originating from different initial spacings was studied between 13 and 33 years of age. First, the effect of spacing on models used to predict crown width and crown ratio from diameter at breast height (DBH) and height was examined. Models for trees of different ages that included all the spacings were found to predict crown growth measures as well as separate models derived for each spacing. Second, the following crown relative growth measures were studied: crown width/crown length (crown shape ratio), crown surface/crown volume, and foliage biomass/crown volume. The way such measures changed over time under different initial spacings was studied; these findings were compared with changes in relative growth rate (RGR), which can be used to evaluate the effect of competitive stress. Crown shape ratio decreased with an increase in DBH in the absence of severe competition, and increased with DBH under severe competitive stress. The other two crown relative growth measures were always negatively correlated with DBH; this shows that large trees use their aerial growing space less efficiently than small trees at all stages of stand development. Only crown shape ratio changed in the same way as RGR.

Beschreibung: Age structures and growth curves were used to determine the origin and to follow the development of 17 balsam fir (Abiesbalsamea (L.) Mill.) stands in the middle of the Boreal Zone in the north of Lake Saint-Jean, Quebec. Every growth curve from the studied sites presented a more or less long period of suppression in the early growth of the balsam fir, followed by a marked release synchronous between the trees of a given site. Generally, a balsam fir cohort established just before the growth release. Characteristic growth releases generally occurred after the formation of exceptionally narrow rings, which took place around 1952, 1914, and between 1860 and 1890 in the sampled trees. Unimodal even-aged stands regenerated from balsam fir advanced growth following spruce budworm (Choristoneurafumiferana (Clem.)) outbreaks that rapidly defoliated the trees and caused openings, sometimes associated with blowdown; these outbreaks occurred between 1944 and 1953 (one stand), between 1909 and 1923 (six stands), and possibly between ca. 1860 and 1900 (seven stands). Bimodal and uneven-aged stands were associated with two spruce budworm outbreaks. In one stand, some of the trees regenerated from the spruce budworm outbreak that occurred between 1909 and 1923, but the 1944–1953 outbreak was intense enough to cause important defoliation and mortality, which released a second cohort. Finally, two stands regenerated from the possible spruce budworm outbreak that occurred around 1860–1900 and from the 1909–1923 outbreak. The presence of a fir cohort and associated growth release between 1860 and 1890 strongly suggests that an outbreak occurred at that time in the studied region. Every sampled stand from the Boreal Zone north of Lake Saint-Jean regenerated from a balsam fir seedling bank following a spruce budworm outbreak that affected the mature balsam fir canopy. These results are in good agreement with the hypothesis that spruce budworm outbreaks and balsam fir forests form an interdependent self-regulating system. Our study shows the drastic effect of spruce budworm outbreaks in the boreal forest, where black spruce (Piceamariana (Mill.) B.S.P.) dominates.

Beschreibung: Water-stress tolerance of six clones in a pedigree consisting of black cottonwood (Populustrichocarpa Torr. & Gray, female) and eastern cottonwood (Populusdeltoides Bartr., male) parental clones and four hybrid progeny was investigated. Trees were grown outdoors in pots; well-watered trees were kept moist (soil water potential greater than −0.03 MPa), and stressed trees (soil water potential less than −2.0 MPa) were subjected to repeated cyclical stress of 1 or 2 days duration over the 14-week study. Male P. deltoides and the male clone 242 displayed the greatest degree of stress tolerance, as evidenced by greater osmotic adjustment at saturation (0.25 MPa) and maintenance of relative growth rate of the main stem under water stress at 100 and 69% of that of well-watered trees, respectively, compared with reductions to 50–58% for the other hybrid clones. However, differences in total plant dry weight under water stress were less obvious, with female clones allocating more carbon to branch production under well-watered conditions, which was further increased under water stress. Three of the four hybrids displayed some degree of osmotic adjustment at saturation after bud set, which was likely conferred by male P. deltoides. Screening clones of Populus for drought tolerance should take into account the segregating tendency of hybrids to allocate carbon to lateral meristems under stress.

Beschreibung: Woody vegetation was surveyed in 58 forest stands in northern Virginia to examine the effects of previous land-use history on past and present-day forest composition and dynamics. Stands were separated using detrended correspondence analysis (DCA) and overstory importance values into three forest groups: (i) white oak (Quercusalba L.)–tulip-poplar (Liriodendrontulipifera L.) (ii) white oak–scarlet oak (Quercuscoccinea Muenchh.) and (iii) Virginia pine (Pinusvirginiana Mill.) The first DCA axis represents a successional continuum from more recently disturbed areas containing young pine forests to less disturbed mature oak stands, and is negatively correlated with stand age and species diversity. White oak and red oak (Quercusrubra L.) dominated presettlement forests in the area. Following European settlement, forests experienced intense logging associated with the charcoal iron industry, large-scale clearing for agriculture, and subsequent land abandonment. By coupling radial growth analysis with age–diameter figures, we evaluated the responses of stands to disturbances associated with various land-use practices. This analysis indicated that many Virginia pine stands resulted from agricultural abandonment during the early 1900s, while a majority of oak stands experienced peak recruitment and radial growth following periodic logging disturbances in the 1800s. Canopy closure, forest protection, and reduced fire and logging disturbance this century led to increases in dogwood (Cornusflorida L.) and blackgum (Nyssasylvatica Marsh.) in area forests. The oldest stands exhibited a lack of tall oak regeneration; however, they also contained a scarcity of potential oak replacement species. Therefore, oak will seemingly share future dominance with several mixed-mesophytic species, although the exact successional status of these stands is unresolved.

Beschreibung: This paper reports the nuclear DNA content estimates obtained by flow cytometry for a group of twelve Eucalyptus species and five fast-growing hybrids that includes those most widely planted throughout the world. Estimates of nuclear (2C) DNA content for the species surveyed ranged from 0.77 pg/2C for Eucalyptuscitriodora Hook. (subgenus Corymbia) to 1.47 pg/2C for Eucalyptussaligna Smith (subgenus Symphyomyrtus). This range corresponds to a haploid genome size range of 370–700 megabase pairs. The average physical equivalent of a 1 cM distance could be as low as 200 kilobase pairs in Eucalyptus, an attractive feature for positional cloning efforts in woody plants. The closer the species were in phylogenetic relationship the more similar were their nuclear DNA content values. All the interspecific hybrids surveyed displayed a nuclear DNA content in the expected intermediate range between the respective parental species, with the exception of one originating from Rio Claro, Brazil, whose exact parentage is unknown. No evidence of polyploidy was observed in any of the hybrids. The flow cytometry procedure employed in this study is an efficient method for investigating ploidy levels of high yielding hybrids of Eucalyptus.

Beschreibung: The productivity of trees under short-rotation intensive culture is influenced by herbaceous competition and drainage conditions. During the first year of establishment of Salixdiscolor Mühl. and Salixviminalis L., four weed-suppression treatments were applied to two sites showing different drainage conditions, one well drained and the other is poorly drained. On the well-drained site, the productivity of the trees increased in all the plots under treatment, whereas on the poorly drained site, only the use of a plastic mulch increased the biomass production. The mean productivity on the well-drained site was always superior to the productivity measured on the poorly drained site except when the vegetation was controlled by the use of a plastic mulch. The analyses related to nutrition revealed that the trees on the well-drained site absorbed more nutrients than those on the poorly drained site. Salixdiscolor showed a higher uptake of potassium and S. viminalis, a higher concentration of calcium. Our results indicate that weed suppression is essential to the establishment of trees in short-rotation intensive culture and that the use of a plastic mulch proved to be particularly profitable on marginal sites.

Beschreibung: Canopy nutrition, leaf chlorophyll concentration, and leaf CO2 assimilation capacity (Amax) were examined in sugar maple (Acersaccharum Marsh.) trees exhibiting symptoms of crown dieback in four stands on acid soils (pH ≈ 4.0) in northern Vermont. Leaf CO2 assimilation capacity was measured on foliage from detached and rehydrated branches harvested from the upper portion of the canopy. Leaf calcium (Ca) and magnesium (Mg) concentrations were among the lowest reported for sugar maple in its natural range. Total leaf chlorophyll concentrations of canopy leaves were lowest on the sites exhibiting the lowest leaf nitrogen (N) and Ca, and CO2 assimilation capacity was correlated with chlorophyll concentration among canopy leaves from all sites. Strong linear relationships were observed between leaf CO2 assimilation capacity per unit leaf mass and leaf N (r2 = 0.60) as well as leaf Ca (r2 = 0.51) among the four sites. On the basis of the observed strong correlation between leaf Ca and leaf N (r2 = 0.64) and the lack of clear enhancement of leaf CO2 assimilation capacity in trees fertilized with base cations (K, Ca, and Mg), it appears that leaf CO2 assimilation capacity and leaf Ca may not necessarily be functionally related. However, since low leaf CO2 assimilation capacity and photosynthetic N-use efficiency were common in unfertilized trees with low Ca (Ca 〈 0.6%), CO2 assimilation processes in sugar maple on acid soils may be limited by N and Ca × Mg interactions. The strongly acidic nature of the soils in these stands and the magnitude of acidic deposition in the region may precondition sugar maple trees on some sites to levels of cation deficiency that may be associated with low CO2 assimilation in the forest canopy.

Beschreibung: One of the most common ways to account for investment risk is to add a risk premium to the risk-free discount rate when computing present values of expected revenues which are uncertain. Using certainty-equivalent analysis, we show that the correct risk premium for short-term investments can easily be in the commonly used 7-percentage-point range. But for such risk premiums to be appropriate for long-term forestry investments, the necessary certainty-equivalent conditions often seem to be unreasonably restrictive. Results suggest that the appropriate risk premium may decline with lengthening payoff period for many forest investments. Limited empirical data provide tentative support, but more research is needed to resolve the issue. We review policy implications and suggest areas for further research.

Beschreibung: Quantitative assessments of the incidence and severity of Mycosphaerella leaf disease were made on nine provenances (encompassing the four subspecies) of Eucalyptusglobulus Labill. over three seasons in 1990 in a trial in Victoria, Australia. Defoliation was also assessed and tree height and diameter measured. Mycosphaerella leaf disease increased rapidly from winter to spring with disease incidence reaching 100% in most provenances by summer. There were highly significant differences in disease severity among provenances, with provenances from E. globulus ssp. globulus Kirkpatrick and E. globulus ssp. bicostata (Maid, et al.) Kirkpatrick being the most severely affected, while provenances from E. globulus ssp. maidenii (F. Muell.) Kirkpatrick and E. globulus ssp. pseudoglobulus Kirkpatrick were only slightly affected. There were also highly significant differences among provenances within E. globulus ssp. globulus and E. globulus ssp. pseudoglobulus. There was a high correlation between disease severity in summer and defoliation, suggesting that Mycosphaerella leaf disease was the main causal agent in leaf fall. There was also a high negative correlation between disease severity and growth rate. This study shows the potential benefits of selecting resistant provenances of E. globulus to maximise production from plantations in areas where Mycosphaerella leaf disease is a problem.

Beschreibung: We used a new technique to examine the response of the trunk and structural roots of white spruce (Piceaglauca (Moench) Voss) to an increase in wind exposure. Ring widths were measured in the roots and trunks of trees located within a 120-year-old, boreal mixedwood stand (control) and at the edge of a road built through the stand 16 years before sampling (released). The observed ring widths were divided by widths predicted from regression to produce ring indices. Response indices were produced by subtracting the ring indices of control trees from those of released trees. Allocation indices were produced by subtracting the ring indices of trunks from those of roots. A final index, quantifying the change in allocation to the root and trunk after road clearing, was produced by subtracting allocation indices of control trees from those of released trees. Following the road clearing, the rate of trunk diameter growth remained unchanged for 3–9 years, while root diameter growth increased. These observations suggest that trunk growth may be suppressed for some years following road clearing as a result of increased root growth. The increase in root growth may help stabilize trees after exposure to increased wind stress by increasing the amount of root wood anchoring and supporting them.

Beschreibung: A common-garden study of Populustrichocarpa Torr. & Gray was initiated in 1985 when clonal material from 128 trees was collected from sites distributed along two mesic (Hoh and Nisqually) and two xeric (Dungeness and Yakima) river valleys. This material was grown for 1 year at Puyallup, Wash. In spring 1986, cuttings from this material were used to establish two replicate plantations, one at Puyallup and one at Wenatchee, Wash. Over the 2 year period, trees were assessed for survival, damage, and growth. Two-year survival was 86% at Puyallup and 59% at Wenatchee; survival was higher for mesic-origin trees at both sites. At Wenatchee, mortality was mainly due to a droughty soil and hot, dry climate in the first year, and damage was due to the tarnished plant bug (Lygus spp.), field voles (Microtus spp.), and grasshoppers (Family Acrididae). At both locations, Melampsoraoccidentalis Jacks. leaf rust was found mainly on trees originating from Yakima. Mean 2-year height and diameter at Puyallup were 457 cm and 40 mm, respectively; corresponding values at Wenatchee were 320 cm and 29 mm. At Puyallup, 2-year stem volume (diameter2 × height) decreased significantly in this order: Nisqually (13 500 cm3) = Hoh 〉 Dungeness 〉 Yakima (4700 cm3). Within the Nisqually trees, clones from lower, milder climate elevations grew more than those from the upper elevations. The reverse was true for the Yakima trees, presumably because the lower elevation trees are adapted to an arid climate and are very susceptible to Melampsora rust. At Wenatchee, high microsite heterogeneity masked much of the genetic variation. The genetic variance component (among rivers, sites, and clones) for stem volume at Puyallup increased from 27 to 51% over the second year. In analyses of each river valley, genetic components (elevational group and clone) were also high at 27–63%; however, for the Yakima trees, the elevational group variance (40%) was much larger than for the other valleys (0–7%) and suggests a steep selection gradient midway along the river transect.

Beschreibung: Field exclosure studies have shown that mammalian browsers such as white-tailed deer (Odocoileusvirginianus) can have pervasive effects on forest dynamics in eastern North America. Direct experimental tests of the effects of browsing on growth and survival of a wide range of tree species, however, have yielded conflicting results. This study was designed to assess the effects of variation in the frequency, seasonal timing, and intensity of browsing (simulated by mechanical clipping) on the growth and mortality of three of the major tree species of the Hudson Valley, New York. The clipping treatments were applied to seedlings grown under two different light regimes (full sun and 8% of full sun) to examine seedling responses under different levels of shade-induced carbon stress. Our results demonstrate that even 2 successive years of heavy winter clipping (75% of new shoot growth removed) has little immediate effect on growth or survival of any of the three species. It is possible that winter browsing only has significant negative effects when seedlings are browsed repeatedly over long periods of time. However, comparable levels of summer browsing for only 2 years significantly reduced both growth and survival of all three species. While most natural browsing occurs in the dormant season, our results suggest that it is the less frequent browsing during late spring and early summer that has the greatest immediate effect on tree seedlings. Shading reduced growth and increased mortality in all three species; however, there was only a limited interaction between light level and the simulated browsing treatments. The effects of browsing on survival were similar in all three species; however, the effects of browsing on cumulative height and annual growth varied enough among the species to suggest that browsing could cause significant variation among these species in their rate of invasion in old fields and rights of way, and their rate of regeneration following logging or disturbance of forests.

Beschreibung: Radial and height growth are characterized for saplings of 10 dominant tree species in a transition oak–northern hardwoods forest in southern New England. Growth of saplings in the field is regressed against measures of whole-season light availability, soil moisture, and sapling size. Statistical tests show strong effects of light availability on growth, but no significant effects of soil moisture. Comparison of the light-dependent growth functions for the 10 species revealed three apparent interspecific trade-offs. (i) Species growing quickly at high light tended to grow slowly at low light and vice versa. The order of species from fast growing at high light to fast growing at low light did not correspond to traditional classifications of shade tolerance, and variation along this axis was approximately continuous. (ii) There was substantial variation off the species continuum defined in i. At any point along the continuum from fast growth at high light to fast growth at low light, some species grew faster than others, and these faster growing species had lower survivorship during periods of suppression than the slower growing species. (iii) Height growth at high light was inversely related to survivorship when suppressed. This variation was again continuous (species did not cluster into discrete categories), but the order of the species did correspond closely to a traditional ordering of shade tolerance. There was little correspondence between our estimated growth functions and the growth functions assumed in the JABOWA–FORET class of forest simulation models. These results raise serious concerns about the current practice of assigning growth functions to species in simulation models using traditional classifications of shade tolerance.

Beschreibung: We have analyzed the light transmission characteristics of the nine deciduous and coniferous species that dominate the transition oak–northern hardwood forests of southern New England. Maximum likelihood techniques were used to estimate species-specific light extinction coefficients, using fish-eye photography combined with data on the locations and geometry of trees in the neighborhood around each photo point. Quantum sensors were also used to quantify interspecific variation in the importance of sunflecks and beam enrichment. Variation in light extinction was closely correlated with shade tolerance and successional status of the species. The most shade-tolerant species (Fagusgrandifolia Ehrh. and Tsugacanadensis (L.) Carr.) cast the deepest shade (5% full sun). These differences were more closely related to differences in crown depth than to differences in light extinction per unit depth of crown. Sunflecks contributed relatively little radiation beneath late successional species (

Beschreibung: We conducted soil moisture manipulation experiments in a red pine (Pinusresinosa Ait.) plantation at the Harvard Forest (Petersham, Mass.) in August 1992 and May 1993. To manipulate soil moisture, we added 10 cm of groundwater to 1-m2 plots and allowed the soils to dry down to their pretreatment moisture contents. We measured methane (CH4) flux, soil moisture, and temperature prior to and after the water addition. Soils in both the control and watered plots were usually sinks for atmospheric CH4. Average consumption rates by control soils ranged from 0.12 to 0.17 mg CH4-C•m−2•h−1. Methane consumption rates by watered soils ranged from 0 to 0.12 mg CH4-C•m−2•h−1 and were inversely related to the moisture content of the upper 10 cm of mineral soil. Linear regression between soil moisture and CH4 consumption explained 78% of the variability (CH4 consumption = 0.001 75 (percent water filled pore space)–0.1957). Using this empirical relationship, we predicted CH4 consumption by soils at three other locations in the Harvard Forest, which agreed closely (r2 = 0.7574) with rates measured in the spring, summer, and fall of 1988–1992. Results from our study suggest that soil moisture is a good predictor of methane uptake by these forest soils and may be used to predict how future changes in soil moisture resulting from alterations in regional precipitation patterns will affect the strength of this terrestrial CH4 sink.

Beschreibung: Residual stand damage was evaluated following partial cutting with a mechanical and a conventional harvesting system in a northern hardwood stand in Maine. Damage from harvesting using a swing to bunch feller–buncher and grapple skidder was compared with damage from a chainsaw and cable skidder operation. Harvesting resulted in actual residual basal areas of 10 and 16.5 m2/ha in a heavy and light conventional treatment, respectively, and in 9 and 6.9 m2/ha in two mechanized treatments. The proportion of stems wounded averaged 22–44% in the conventional, and 20–31% in the mechanized operation. Wounds were significantly (p 〈 0.05) longer in the heavy conventional [Formula: see text] treatment than in light conventional [Formula: see text] and mechanized ([Formula: see text] and 60.2 cm) treatments. Wounds were located higher on the stems in the 9 m/ha mechanized [Formula: see text] than in the 10 m2/ha conventional treatment [Formula: see text]. A separate survey of the mechanized harvest showed that season of harvest (winter vs. summer) significantly affected wound location and the wound width to tree circumference ratio. A model was developed that relates harvesting treatment (light and heavy conventional and two heavy mechanical), species, distance of a stem from the edge a skid trail or access corridor, and initial stand basal area to the probability of injuring an individual stem. Mechanized harvesting using equipment with long-reach swing to bunch felling capability appears to be better than some other mechanical systems, and no worse than conventional harvesting systems, but the need to further reduce damage levels is evident.

Beschreibung: The export of mass and nutrients associated with the formation of fungal sporocarps during the first 7 years of decomposition of logs of four conifer species (Abiesamabilis Dougl. ex Forbes, Pseudotsugamenziesii (Mirb.) Franco, Thujaplicata D. Don, and Tsugaheterophylla (Raf.) Sarg.) was investigated in western Oregon. Abundance of the most common fungal species, Naematolomacapnoides (Fr.:Fr.) P. Kumm, differed significantly with log species; the fungus was most abundant on Abies and least abundant on Thuja. Fungi increased concentrations of N, K, and P over those found in associated logs by as much as 38, 115, and 136 times, respectively. Thus, a fair proportion of the initial N (0.9–2.9%), K (1.8–4.5%), and P (1.9–6.6%) was transported out of logs via sporocarps at a time when immobilization would have been predicted from critical element ratios (e.g., C/N).

Beschreibung: Mineral soil and forest floor C and N contents were determined in 154 Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and western hemlock (Tsugaheterophylla (Raf.) Sarg.) stands in western Oregon and Washington ranging in age from 16 to 64 years. Relationships between site index and mineral soil and forest floor C, N, and C/N ratios were examined. Douglas-fir data were analyzed by geographic province (Puget Sound, Washington Cascades, Oregon Cascades, coastal Washington, coastal Oregon, and southwest Oregon). Average mineral soil C in Douglas-fir stands ranged from 102 Mg/ha to 177 Mg/ha in Puget Sound and Washington Cascades provinces, respectively. Soil N ranged from 3708 kg/ha in Puget Sound province to 9268 kg/ha in the Washington Cascade province. Western hemlock data were analyzed in three provinces (Washington Cascades, coastal Washington, and coastal Oregon). Average mineral soil C in western hemlock stands ranged from 241 Mg/ha in the Washington Cascades to 309 Mg/ha in coastal Washington and was higher than Douglas-fir mineral soil C. Western hemlock mineral soil N was also higher than Douglas-fir mineral soil N ranging from 10 495 kg/ha in the Washington Cascades to 15 216 kg/ha in coastal Oregon. Forest floor C and N contents were also higher in western hemlock than Douglas-fir stands. Nonlinear regression analysis revealed a weak positive relationship between site index and total mineral soil C in Douglas-fir (r2 = 0.19). A similar relationship was observed between Douglas-fir site index and total soil N (r2 = 0.19). Relationships were weak because of the large variability in mineral soil C and N within as well as across provinces. Maximum Douglas-fir site indexes occurred across a broad plateau of mineral soil and forest floor C/N ratios ranging from 15–25 and 35–45, respectively. Minimum site indexes also occurred in these C/N ranges. No increase in Douglas-fir productivity occurred above mineral soil C levels of 125 Mg/ha. There were no relationships between site index and mineral soil C and N or C/N ratios in western hemlock stands.

Beschreibung: Tabulated Student's t-values and climatic insensitivity among inner tree-ring widths can bias estimates of statistical significance for cross correlations relating "floating" and master tree-ring series. These biases can be removed by (i) directly computing significance levels for cross-correlation coefficients at dating positions and (ii) deleting insensitive inner rings from a dated floating sample before final correlation analysis. The number of early rings to delete can be determined from plots of cross-correlation coefficients linking a dated floating series of artificially decreasing length with a master series. These modifications improve the precision of Yamaguchi and Allen's approach (D.K. Yamaguchi and G.L. Allen. 1992. Can. J. For. Res. 22: 1215–1221) for estimating significance.

Beschreibung: The fertilizer efficacy of a variety of organic wastes was tested in a 9-year-old plantation of western red cedar (Thujaplicata Donn ex D. Don) growing on a cutover of cedar–hemlock (Tsugaheterophylla (Raf.) Sarg.) forest on northern Vancouver Island. Seven treatments were compared: municipal sewage sludge, sewage sludge plus pulp sludge, fish silage and wood ash, silage and ash plus pulp sludge, wood ash alone, ammonium nitrate with triple super phosphate, and control (untreated). Each treatment was replicated three times. Rates of application were 225 kg N•ha−1 in the inorganic fertilizer and about 500 kg N•ha−1 in the organic wastes (except wood ash). The height and diameter of the cedar trees 2 years after fertilization were greatest in the plots treated with inorganic fertilizer (average height was 274 cm, vs. 211 cm in control plots; average diameter at 30 cm was 49 mm, vs. 34 mm in control plots). Smaller but significant growth responses were achieved with sewage sludge and fish silage plus ash. Mixtures of sewage sludge or silage with pulp sludge produced smaller height growth responses, but did not affect diameter growth. Wood ash alone had no effect on tree growth. All treatments except wood ash increased the concentrations of macronutrients in foliage. Foliar N concentrations were greatest in trees treated with fish silage or inorganic fertilizer. Stagnated plantations of western red cedar appear to provide an opportunity for recycling these organic wastes.

Beschreibung: Height growth at 10 years from striking was assessed for clonally replicated full-sib black spruce (Piceamariana (Mill.) B.S.P) families tested at three locations in central Nova Scotia. Variance components were interpreted according to an additive–dominance–epistasis genetic model and used to derive comparative estimates of gain from various selection and deployment strategies. Field performance at 5 and 10 years was compared with that of the original ortets and families growing in a 25-week greenhouse study, by means of phenotypic and genetic correlation, and rank-change analyses. Between age 5 and 10, the additive portion of the total genetic variance for height decreased from 66 to 38%, while the dominance portion increased from less than 3 to 13%, and the epistatic portion from 31 to almost 49%. As a consequence, narrow-sense heritability estimates were lower at age 10 and gain estimates also decreased, particularly for those strategies that capture gain primarily from additive effects. Although correlations between field performance and early growth measurements were generally poor, the strongest were found at the half-sib level; full-sib correlations were somewhat weaker and those between clone means and early ortet performance were small and not statistically significant. The strongest age–age correlations were those that involved family mean seedling weight in the greenhouse. Family rankings based on early oven-dry biomass production also showed the most consistent agreement with ranking after 5 and 10 years of field testing, although the analyses suggest that effective early selection is probably limited to culling the worst 25% of the families based on biomass. Genotype–environment interactions were statistically significant, although these were limited to only 2% of the phenotypic variance in height growth; furthermore, strong genetic correlations between environments suggested that these interactions have little impact on selection efficiency.

Beschreibung: Sitka spruce, Piceasitchensis (Bong.) Carr., from 27 provenances and five sites in British Columbia, were examined for traits of leader morphology that might be involved in resistance to the white pine weevil, Pissodesstrobi (Peck). Trees from the Usk Ferry provenance at Head Bay and the Kitwanga provenance at Nass River both had low incidence of weevilling, and had needles pressed more closely against the stems by up to 20° than trees from other provenances. At Fair Harbour, trees from the two most resistant provenances, Haney and Cedarvale, had inner resin ducts that were approximately twice the diameter of those in trees from the most susceptible provenances. Resistant trees from these two provenances also had significantly more outer resin ducts than those from any other provenance planted at any of the five sites. Values ranged from 28 outer ducts per centimetre for Haney trees, to

Beschreibung: Age and size structures were determined in 40 stands of black spruce (Piceamariana (Mill.) B.S.P.) growing on peatlands and transitional peatlands in northeastern Ontario, Canada. Natural and second-growth stands representing several site types were examined. The age structure of natural black spruce stands varied from even aged in young stands (

Beschreibung: Owing to the significance of shade, wildlife habitat, soil stabilization, and water-filtering effects of streamside vegetation on the riparian zone, spatial restrictions are imposed on both public and private harvesting areas along the stream by state and federal agencies. Analysis of both short– and long–term impacts of the riparian zone spatial restrictions as well as spatial restrictions on harvest units is presented. The heuristic model called SSMART, for spatially constrained harvest scheduling problems, is used to solve the proposed problems. The Green River sub-basin on the Alsea Ranger District, Siuslaw National Forest, Oregon, is used for the analysis. Our computational experiments show that the marginal reduction of the total present net worth in dollars is about 6.2% per 100-foot (1 foot = 0.305 m) width of the riparian management area, and about 6.7% for the average annual harvest flow. It is also shown that increasing the exclusion period to two or three periods could cause major reduction (25–40%) in both the total present net worth and the average annual harvest flow. The linear programming relaxation is revealed to provide a good approximation for impacts of the size of riparian management areas, while it cannot be used for examining impacts of the exclusion period.

Beschreibung: We measured fluxes of N2O, CH4 and CO2 from control and urea-nitrogen fertilized soils of a mature slash pine (Pinuselliottii var. elliottii Englem.) plantation in Alachua County, Florida. The fertilization did not affect CO2 emissions, but significantly increased the emissions of N2O and lowered the uptake of atmospheric CH4. Daily average N2O emissions from the fertilized soils were 8–600 times higher (12–74 μg N2O-N•m−2•h−1) than daily average N2O emissions from control soils (0.02–4.0 μg N2O-N•m−2•h−1). Daily average CH4 uptake by the fertilized soils were 5–20 times lower (0.001–0.007 mg CH4-C•m−2•h−1) than daily average CH4 uptake by control soils (0.015–0.035 mg CH4-C•m−2•h−1). We also measured the relative activities of the bacteria populations that were responsible for CH4 oxidation in the control and fertilized soils. Results from these measurements suggest that fertilization shifted the relative activities of the CH4 oxidizing bacteria from those dominated by methanotrophs in the control soils to those dominated by nitrifying bacteria in the surface (0–2 cm) of the fertilized soils. The shift in relative activities of these bacteria may have been responsible for the lower CH4 uptake by the fertilized soils.

Beschreibung: The relationships of sapwood radial width and transverse area to tree diameter, age, and growth rate were investigated in Piceaabies (L.) Karst. A total of 125 trees growing with (suppressed trees) and without (dominant trees) competition for light were sampled. Both sapwood and heartwood amounts showed an increase with diameter at the stem base, with the heartwood portion increasing more rapidly. In young trees sapwood prevails both in terms of diameter and transverse area. After trees have reached a certain age, the width of the sapwood band remains more or less constant (on average 7.8 cm for dominant and 2.0 cm for suppressed trees), and the heartwood amount exceeds that of sapwood. The percentage of heartwood in suppressed trees is substantially greater than in dominant trees of the same age. Sapwood amount is closely correlated with the tree diameter, but not with age. Tree age controls the number of rings in sapwood, while the sapwood width depends on the tree's radial growth rate as well.

Beschreibung: Dendroecological techniques were used to study the influence of climate on the growth of subalpine fir (Abieslasiocarpa (Hook.) Nutt.), Engelmann spruce (Piceaengelmannii Parry), and subalpine larch (Larixlyallii Parl.) in the North Cascade Mountains of Washington state. Study sites were selected on different topographic features at three points along an elevation gradient (ridgetop, valley slope, and valley floor) to characterize site influences on growth response to climate. Mixed species stands were sampled to identify possible differences in species response to climate on a common site. Species differences account for most of the variability in radial growth patterns and response to climate. The greatest differences are between subalpine fir and subalpine larch, while the greatest similarities are between subalpine fir and Engelmann spruce. After species, aspect is the most significant factor affecting growth response to climate. Spring snowpack and summer temperature are the primary climatic factors influencing growth. Spring snowpack is negatively correlated with growth for all three species, but the relationship is strongest for subalpine fir and weakest for subalpine larch. Subalpine larch growth is positively correlated with June temperature. Subalpine fir growth is positively correlated with July–August temperature. Engelmann spruce growth is positively correlated with June–August temperature, but unusually warm July–August temperatures are associated with reduced growth the following year. The response of subalpine forests in the North Cascades to future climate change will depend on winter snowpack accumulations and spring snowmelt rates. Earlier meltouts and warmer summers would benefit growth on north aspect sites, but could also increase the frequency and severity of summer drought conditions on ridges and south-aspect sites, where summer soil moisture may already be limiting.

Beschreibung: Our previous work has demonstrated a relationship between polarity index and organic carbon referenced partition coefficients (Koc) of organic chemicals. Here we compare sorption of benzene, toluene, or o-xylene by protein with non-protein organic sorbents. Koc for benzene, toluene, or o-xylene sorption by collagen was much higher than by lignins, humic acid, chitin, or cellulose. Sorption of benzene, toluene, or o-xylene was not only influenced by organic sorbent polarity and functionality (e.g., aromaticity), but also by additional properties which we inferred related to physical conformation of the organic sorbents. In this respect, collagen exhibits unique sorptive behavior toward removal of nonpolar organic chemicals in the aqueous phase, which was not shared by humic acids of similar C, O or N content. Key words: Sorption, collagen, lignin, polarity, hydrophobic domains

Beschreibung: The 5-year height and diameter growth response of a frost-tolerant species, (lodgepole pine, Pinuscontorta var. latifolia Dougl. ex Loud.) and a frost-susceptible species (hybrid spruce, Piceaglauca (Moench) Voss × Piceasitchensis (Bong.) Carr.) to different vegetation-control site-preparation treatment and timing combinations was examined using a factorial experiment. The site chosen for the experiment is subject to frequent summer growing season frosts. The vegetation control treatments were: an untreated control, manual cutting, and glyphosate applied at the rates of 1.4 and 2.1 kg active ingredient (a.i.) per hectare. Each vegetation-control treatment was applied at four dates (timing) that covered the active growing season of the vegetation on the study area. Vegetation-control method had a major impact on the growth of the planted seedlings. The influence of timing of control treatments on growth was found to be minor. Fifth-year height and diameter in the manual cutting and control treatments were equally poor. Growth was significantly improved by both levels of chemical vegetation control and after 5 years, no differences were apparent between the two. Pine height and diameter and spruce diameter continually improved as the level of thimbleberry (Rubusparviflorus Nutt.) was reduced. Below 5% thimbleberry cover, this growth increased dramatically, suggesting a response threshold. As vegetation cover was lowered, however, spruce seedlings were damaged by frost, resulting in reduced total height. In this case, better height growth was associated with either very low or moderately high thimbleberry cover than with moderate to low levels.

Beschreibung: To assess the impact of atmospheric deposition on soil acidification and base cation supplies in montane spruce–fir forest soils at Whiteface Mountain, New York, base cation and proton fluxes were determined for organic and mineral horizons from measurements made at four stands (1020–1090 m above sea level) over a 4-year period. Our best estimates indicate an annual accumulation of H+ and a net loss of base cations from the forest floor of about 0.71 kmolc/ha, a 2.8% per year loss of the total forest floor base cation pool. This high rate of acidification is attributable to base cation leaching by sulfate and organic anions, and uptake by living biomass. From 1986 to 1990, the annual net loss rate of forest floor Ca, the most abundant base cation, was several times greater than historical loss rates as determined by 50-year comparisons of forest floor Ca in nearly identical forests of the Adirondack High Peaks region. Published data on long-term trends in Ca deposition in the U.S. Northeast suggest that the difference between historical and current net loss rates of forest floor Ca may be due to sharply reduced atmospheric inputs of Ca after about 1970, exacerbated by sulfate leaching. In mineral horizons where the total base cation pool (mostly mineral bound) is very large, the net losses of base cations were substantially lower and in the range where losses due to leaching and uptake can be countered by mineral weathering.

Beschreibung: Plant mixtures were established that differed in both proportion and density of loblolly pine (Pinustaeda L.), sweetgum (Liquidambarstyraciflua L.), and broomsedge (Andropogonvirginicus L.). Soil moisture availability to the pine seedlings was quantified every 2 weeks by measuring predawn xylem pressure potentials. Temporal variation in pine water potential was accounted for by a water stress integral approach. Cumulative water stress integral values were calculated over four overlapping periods, from May to June, May to July, May to August, and May to September and compared with the mean seedling stem volume index at each period to determine competitive responses at the whole plant scale. Diurnal measures of stomatal conductance were taken each month to compare competitive responses at the leaf scale. In addition, environmental and plant responses that may control stomatal behavior were quantified. The pine water stress integral was strongly influenced by competing vegetation after the onset of a period of drought in early summer. The correlation between the water stress integral and pine growth increased after a significant drying period, accounting for more than half of the variation in stem volume index at the end of the first growing season. Stomatal conductance was also influenced by competition, with competitive effects more evident during times of drought. Conductance was most often related to bulk leaf water potential, which in turn was related to competitive effects on soil moisture availability. Vapor pressure deficit also influenced stomatal conductance, but this was largely unrelated to competitive effects.

Beschreibung: Long-term (17–31 year) density estimates of jack pine budworm, Choristoneurapinuspinus Free. (Lepidoptera: Tortricidae), populations from 31 townships in northwestern Wisconsin were analyzed to investigate variation in population behaviour. Populations had varying combinations of cyclic components with periods of 5, 6, and 10 years. Populations that fluctuated with higher frequencies (shorter periods between outbreaks) tended to be found in locations where the habitat type was indicative of nutrient-poor and extremely dry soils. Many populations in these habitat types had high mean densities, and the coefficient of variation in density was smaller than that in other populations. Just under half (14) of the populations were regulated by statistically significant second-order density-dependent processes. Nevertheless, second-order processes were present to some extent in all populations examined. These results demonstrated that detection of density dependence and population regulation of jack pine budworm depends on the local site where studies are undertaken. Population fluctuations encountered in these populations are of the phase-forgetting quasi-cyclic kind. Studies of mechanisms that account for these cycles and cause populations to fluctuate at three different characteristic frequencies promise to be rewarding.

Beschreibung: Application of 224 kg N/ha to young, thinned stands of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) at 35 sites in western Oregon and Washington significantly increased basal area and volume increment over 8 years following treatment. However, response varied considerably between sites, and relative volume increment exceeded 10% at only 19 of the 35 sites. Response to applied N was evaluated in relation to forest floor and soil variables as well as to levels of N in foliage. Relative responses in basal area and volume were significantly correlated with total N concentration and the C/N ratio of the soil. However, these relationships explained only part (18–22%) of the observed variation in response. In contrast, relative response was strongly correlated with the level of N in the foliage of nonfertilized trees at 11 sites, accounting for 94% of the variation between sites. Use of foliar N data clearly has potential to predict growth responses to N fertilization of young thinned Douglas-fir stands, although further work is needed to test the relationship for a wider range of sites and stands.

Beschreibung: Soil water limits plant growth in the Canadian Prairie Provinces. Efficient use of soil water is, therefore, paramount in crop production. Two 2-yr field studies were conducted (i) to determine the effects of crop selection and cropping practice on the temporal and spatial distribution of soil water and (ii) to quantify water use and water-use efficiencies (WUE) of alternative crop/cropping systems for a subhumid region of Alberta. The first study was at the Ellerslie Research Station on an Orthic Black Chernozemic and the second at the University of Alberta Breton Plots on an Orthic Gray Luvisol. At each site three annual crops/cropping systems [barley (Hordeum vulgare L.), barley intercropped with field pea (Pisum sativum L.), and faba bean (Vicia faba L.)] and a perennial forage [creeping red fescue (Festuca rubra)] were grown in 1987 and in 1988. At Ellerslie, tillage methods to grow annual crops were conventional and zero tillage. At Breton, the two tillage methods used were: (i) conventional and (ii) deep tillage (to enrich the surface horizon with clay from the subsoil). The experimental design used was a split plot with four replications. Soil water was measured by neutron attenuation. Evapotranspiration was calculated as the change in soil water to 0.80-m depth plus precipitation. Soil water changes were more closely associated with the kind of crop grown than with the method of tillage used. The barley/field pea intercrop exhibited a pattern of water use similar to barley. Greater water-use efficiency measured in the intercrop system was attributed to differences in canopy structure and plant biomass production. Faba bean and red fescue had lower WUE than barley and the intercrop. The water requirement by faba bean was close to that of red fescue and related to leaf area development. Small tillage effects on soil water were observed during 1988. Soil water under zero tillage was greater than under conventional tillage. Faba bean and red fescue are likely to succeed more in agro-ecological regions such as Breton. Further studies are required to improve our understanding of the effects of these cropping systems on water resources when they are used in rotational-production systems. Key words: Barley, field pea, faba bean, creeping red fescue, conventiional tillage, zero tillage, deep tillage, water-use efficiency

Beschreibung: Freeze-thaw and flooding of usually well-drained soils occur in the spring in the prairie and boreal regions of Canada. We studied the impact of these conditions on nitrogen leaching in a Black Chernozemic soil (Udic Boroll). Soil samples, subjected to different treatments, were incubated for 12 wk in the laboratory and leached every 2 wk with 0.001 M CaCl2 solution. The cumulative leaching loss of total N (mg kg−1 soil) was reduced by freeze-thaw (76.0), flooding (41.4) and a superimposition of the two treatments (28.8) compared to the control (109). All treatments affected the distribution of the forms of N leached. The total loss of water soluble organic N (SON) and ammonium-N was in the order of flooded 〉 flooded-freeze-thaw 〉 freeze-thaw = control. In the leachates from the flooded treatments, SON accounted for 71.5–77.4% of the total N leached. Nitrate- and nitrite-N dominated the total leachable N in the unflooded treatments following an order of control 〉 freeze-thaw 〉 flooded = flooded-freeze-thaw. During the incubation, the Eh of the flooded soils decreased from 344 to −46 mV, compared to a variation in Eh from 355 to 301 mV for the unflooded soils. The maximum rate of leaching of organic nitrogen from the flooded treatment (0.53 mg N kg−1 d−1) coincided with a sharp decrease in Eh, from 131 to 42 mV. It is concluded that climatic events will have a significant impact on the dynamics of soil nitrogen. Flooding, in particular, may promote the loss of N in water soluble organic matter. Key words: Flooding, freeze-thaw, organic and inorganic nitrogen leaching, redox potential

Beschreibung: Soil solutions from four salt-affected soils in Alberta were extracted by immiscible displacement from field moist soils and saturated pastes after air-drying to examine the implications of using saturated paste extracts as a proxy of soil solutions. Solutions ranged in pH from 7.8 to 10.6 and in electrical conductivity (EC) from 6 to 45 dS m−1. Sodium dominated the cation suite, with Mg and K concentrations varying by several orders of magnitude between sites. Calcium was the major cation in lowest abundance at all sites. Sulfate was the predominant anion. Phosphorus concentrations ranged from 1 to 247 ppm. Considerable variation was seen between the properties of saturated paste extracts and immiscibly displaced solutions, with the saturated paste extracts having lower conductivities and Na, Mg, K and anion concentrations. The differences in composition between the two extraction methods resulted in differences in ion speciation, SAR and ESP values and mineral stabilities. Saturation indices for evaporite minerals were poor predictors of minerals formed in evaporated soil solutions. It was also found that even in chemically similar solutions, different minerals precipitated, and therefore the prediction of evaporite mineral formation from extracted soil solutions is not recommended. Key words: Salinity, saturated paste extracts, soil solutions, evaporites, mineral stability

Beschreibung: Concentrations of Ca, Mg, K, Na, Al, Fe, Mn, and Si extractable with 1 N ammonium chloride (NH4Cl, pH 4.5) and 1 N ammonium acetate (NH4OAc, pH 4.5) were determined for forest soil samples as follows: (1) before drying, and (2) at several time intervals after air-drying (1, 5, 11 and 14 wk). Values for CEC were obtained for the same samples by determining (1) the sum of cations (Al3+, Ca2+, Mg2+, K+, Na+, Fe3+, and Mn2+) in the extracts [denoted [Formula: see text] and [Formula: see text]], and (2) the amount of ammonium retained by the soil samples against water washing [denoted CEC(NH4OAc) and CEC (NH4Cl)]. The soils used in this investigation were taken from four New Brunswick upland forest sites (two sugar maple sites, one mixed wood site, and one spruce site). It was round that (1) extractable Mg, K, Na, and Mn levels were generally not affected by drying, storing, and type of extradant; (2) extractable Al and Fe levels increased immediately after drying; (3) NH4OAc-extracted Al, Fe, and Si exceeded NH4Cl-extracted Al, Fe, and Si; (4) extracted Al and Fe levels tended to drop after 11 wk of storage; (5) small drying effects were also noticed for NH4Cl-extracted Ca; (6) CEC(NH4OAc) and CEC(NH4Cl) values decreased with increasing time of storage; this effect was noticed most for soil samples with high levels of organic matter (Ah, Ahe, Bm, Bf, and Bfh), and was noticed least for sod samples taken from leached horizons (Ae) and subsoil horizons (BC and C); (7) in some cases, storage time increased CEC(NH4OAc) in subsoils; (8) values for [Formula: see text] remained fairly independent or increased slightly with storage time and were closely related with CEC(NH4Cl) values obtained with non-dried samples; (9) values for [Formula: see text] did not relate well with CEC(NH4OAc) and CEC(NH4Cl). Differences for extractable Al were likely due to Al complexation by acetate ions. Drying effects on extractable Al and Fe (and possibly Ca) were likely due to drying-induced fragmentation of soil organic matter. Drying and storage effects on CEC(NH4OAc) and CEC(NH4Cl) were likely due to (1) water-washing and related loss of organic matter, and (2) sensitivity of subsoil minerals to air exposure. Apparent drying and storage effects on CEC were most noted with [Formula: see text] and were least noted with [Formula: see text]. Key words: Cation exchange capacity, ion exchange, drying, storage, ammonium acetate, ammonium chloride extractions

Beschreibung: There exists a need (i) to test, whether equal or better cereal yields could be obtained using cropping systems which rely on renewable resources rather than on fertilizer nitrogen; and (ii) to discover the condition of the soil resource under these systems.The long-term cropping systems on a Gray Luvisol at Breton were studied. They included: (i) an agro-ecological 8-yr rotation (AER), established in 1981, which involved addition of both fababean green manure and manure from livestock fed with forages and fababeans grown in the rotation: (ii) a continuous grain (barley) system (CG), with fertilizer N at 90 kg ha−1 y−1, established in 1981; (iii) a classical Breton 5-yr rotation (CBR) involving forages and cereals, with no return of crop residues or manure, established in 1930. Mean barley yields were 16–19% higher in the AER (P ≤ 0.05) than in the CG system, and yield on either was about double that of the CBR. Within 9 yr, there was evidence of increased total C, N, and P; available N, P and K, CEC; microbial biomass, microbial respiration; and counts of bacteria, fungi, and mycorrhizae in the AER compared with the CG system.We conclude that biological fixation of N by legumes can be used as the sole source of N for barley production on Luvisolic soils of low fertility such as the Breton loam, without sacrificing yield or soil quality. Barley yields in the AER (38% of the rotation time) exceeded those of barley grown under continuous cereal cropping. The soil resource was maintained or improved during a 10-yr period under AER compared to the CG or CBR systems. Further research is needed to discover the mechanisms involved in regulating biological activity and availability of plant nutrients other than N in the AER system. Key words: Barley, Breton loam, cropping systems, Gray Luvisol, soil quality, fababeans

Beschreibung: Improving efficient use of applied nutrients in crop production is a desirable agronomic, economic, and environmental goal. Field experiments with corn (Zea mays L.) were carried out at two sites with contrasting soil textures in southwestern Quebec from 1988 to 1990. Objectives were to establish an optimum N fertilization rate for grain production, and to study fertilizer N use efficiency on a Chicot sandy clay loam (Grey Brown Luvisol) and a Ste. Rosalie clay (Humic Gleysol). Under favourable heat units and rainfall, yields of grain corn were quadratic functions of fertilizer N rates on both soils. Maximum grain yields of corn were calculated to require from 300 to 350 kg N ha−1 fertilizer application. However, large amounts of N fertilizer at both 285 kg ha−1 and 400 kg ha−1 were not recovered by the crop. Optimum economic grain yields at a price ratio of 10:1 N:grain were found at 273 kg N ha−1 on the Ste. Rosalie site in 1990, and 286 and 179 kg N ha−1 for 1989 and 1990, respectively, on the Chicot site. A normal fertilizer rate (170–44–141 kg N–P–K ha−1) and a high fertilizer rate (400–132–332 kg N–P–K ha−1) were used in 1989 and 1990 for 15N microplot experiments. Grain and stover fertilizer N recovery (FNR) varied with N rate and year from 9 to 58%. Favourable climatic conditions substantially increased crop yield, and thus crop FNR. Residual FNR the second year was less than 5% of the initial applied N. Key words:15N, grain yield, Zea mays L., nitrogen use efficiency, fertilization

Beschreibung: The production and utilization of root-derived C is fundamental to the functioning of ecosystems. The objectives of this experiment were to quantify the amount of root-released C produced by two barley (Hordeum vulgare L.) cultivars, to evaluate the direct and indirect effects of shoot C, root C and root length on the root-released C and to quantify the kinetics of the decomposition of root-released C in soil for two barley cultivars. Seedlings of two barley cultivars, Abee and Samson, were cultured in nutrient solution for 20 d and then pulse-labelled with 14C for 5 d. Samson released more C than Abee during the entire experimental period. Root length had the greatest direct effect on root-released C for the two barley cultivars. Kinetics analysis of the decomposition of root-released C added to soil showed that the labile components of the added material was 87.3% for Abee and 74.4% for Samson with half-lives of 4.3 d and 4.5 d, respectively. The half-lives of the resistant components of the root-released C added to soil and microbially-derived material were 37.7 d for Abee and 29.6 d for Samson, respectively. The amount of root-released C and its decomposition rate in soil varied with cultivars used. Key words: Root-released C, path analysis, direct effect, indirect effect, 14C, decomposition, Typic Cryoboroll

Beschreibung: A study was conducted on three Manitoba soils (Marquette heavy clay, Fortier silty clay loam, and Souris loamy sand) with the objectives of: (a) assessing the effects of contrasting tillage systems on various soil physical properties; and (b) relating root penetration of cereals to these properties. The zero tillage (ZT) and conventional tillage (CT) treatments had been established between 3 (Marquette and Souris) and 10 (Fortier) years prior to the initiation of the study. Penetrometer resistance (PR), bulk density (BD) and pore size distribution (PSD) were measured at various depths and times during two growing seasons. The effects of tillage systems on soil properties were generally small, particularly in the poorly structured Souris loamy sand. PR in the top 10 cm of soil tended to be higher under ZT than under CT. PR varied markedly with time and was closely related to changes in soil water content. The proportion of macropores (〉 100 μm in diameter) near the soil surface tended to be higher under ZT than under CT. There was no evidence of any detrimental effect of poor aeration on the final root penetration profiles of wheat. The proportion of roots penetrating the soil was inversely related to PR. Soil strength critical for root penetration was determined to be 2 MPa and was independent of soil type. In most situations, however, roots were able to grow into soil with mechanical impedance greater than 2 MPa, possibly by making use of biochannels and spatial and temporal heterogeneity in the soil structure. Key words: Wheat, barley, zero tillage, conventional tillage, root penetration, penetration resistance

Beschreibung: The thick folic (mainly upland forest) materials (〉 40 cm of accumulated organic material) that occur in the Coastal Western Hemlock Biogeoclimatic Zone in British Columbia have not been described with regard to the spatial interrelationships of the soil constituents in context with the chemical composition of the different horizons. Micromorphological assessment and solid-state 13C NMR were used to characterize the accumulated folic materials from a Lignic Folisol (northern Vancouver Island), Histic Folisol (Prince Rupert, BC) and a Humic Folisol (Queen Charlotte Islands, BC). Micromorphology provided information on the spatial relationships of the soil constituents and 13C NMR provided data on the chemical components of the folic materials. Soil faunal activity, primarily from mites, was the dominant soil-forming process observed in the organic horizons of the Folisols, being especially prominent in the Lignic Folisol with the breakdown of woody materials. Solid state 13C CPMAS NMR spectra facilitated distinguishing three main types of horizons: (1) Horizons derived from accumulated residues (L, Fr, and Hr) showing higher carbohydrate-like C and O-alkyl C values and lower total aromatics; (2) Horizons with advanced decomposition (Hr2, Oh1, and Hd) which were higher in alkyl C; and (3) Horizons derived from ligneous material (Fw and Hdw) where carbohydrate-like C was less than total aromatic C. Implications for adequate nutrient content and forest growth were inferred from the observed micromorphology and chemical composition of the folic materials. Key words: Folisol, forest soils, micromorphology, 13C NMR, faunal activity

Beschreibung: The amount of available nitrogen in fresh and composted manures is required to adjust inorganic fertilizer applications for crops when using manures and composts. Corn (Zea mays L.) plants were grown for 8 wk at temperatures of 17–27 °C and 12–18 °C in soil amended with two rates of fresh and composted solid beef cattle manure. Apparent N uptake of corn grown in soil amended with fresh solid beef cattle manure, composted beef cattle manure and (NH4)2SO4 at the higher temperature was 2, 8, and 36%, respectively, during this 8-wk period. Nitrogen uptake by corn in fresh solid beef cattle manure-amended soil at low temperature was affected by factors other than N availability. A 12-wk laboratory experiment was performed in which 15NH4+ was added to fresh and composted manure-amended soil to determine the fate of the inorganic N in the manure and to measure N mineralization rates. Net immobilization of inorganic N occurred with all manures; however, after 3 wk, net mineralization occurred with the solid and composted beef cattle manure. Immobilization of 15N continued with all manure amendments as a result of mineralization-immobilization turnover. The amount of unrecovered 15N after 12 wk was 10.5, 2 and 1% of the added 15N in soil amended with solid beef cattle manure, composted beef cattle manure and manure composted with bark, respectively. Key words: Composted manure, nitrogen availability, 15N, immobilization, remineralization, temperature

Beschreibung: Illuviated mineral horizons (Bf) from a Podzol on the Precambrian Shield, Ontario, Canada were collected, placed in columns and treated with distilled water, cation amended water, acidified water or cation and acid amended water over a 4-mo period. While the cation treatment had little effect relative to the control, the acid treatment removed large quantities of Al, Si and base cations. The source of the leached Al was primarily (~ 70%) dithionite–citrate–bicarbonate (DCB) extractable Al, apparently some form of amorphous gibbsite. Organic Al (pyrophosphate extractable) contributed only about 20%. However, even more pronounced was a major within-column migration of oxalate extractable Al–Si, perhaps allophane or imogolite, which was leached from the top of the acidified columns and precipitated below. The weathering of primary minerals and possibly interlayer Al in vermiculite contributed little to the leaching of Al from the acidified columns. Key words: Acid, weathering, Podzol, aluminum, silica

Beschreibung: With the reduction in S inputs through atmospheric deposition and high analyses fertilizers, soil organic S will become a more important source of S to crops. Sulfur mineralization in 20 meadow soils was investigated using a long-term aerobic incubation and a greenhouse bioassay with alfalfa (Medicago sativa L.). The cumulative S mineralization (Sm) was linear with time for 13 soils whereas 7 soils showed an exponential phase over the first 10 wk followed by a linear release phase. Podzolic soils dominated the latter group. The data were described by an incremental zero-order model where an initial flush was accounted for. The total amount of mineralizable S (Sm), the amount of S mineralized in the first 1.4 wk (Se), the S mineralization potential (So) and the rate constant (k) were all very closely correlated to the N mineralization potential and to the amount of N mineralized in the first 1.4 wk of incubation (r = 0.64–0.85; P 

Beschreibung: The practice of planting potatoes (Solanum tuberosum) in rows and hilling the plants at some stage of growth has been universally adopted. Using a rainfall simulator and runoff-erosion plots (0.9 m wide × 1 m long), the effects of hilling on runoff, infiltration, and soil loss were examined on an Orthic Dystric Brunisol. Tests of the rainfall simulator revealed that variations in operating pressure (36.5–62.1 kPa) and soil slope conditions (0–15%) did not significantly (P 

Beschreibung: This paper presents information on pedogenic processes, weathering status and geochemical evolution of ultramafic soils at three areas in southwestern British Columbia characterized by climatic conditions ranging from relatively wet–cool to relatively dry–cool. The soils of the Coquihalla serpentine belt have Podzolic profiles that resulted from intense weathering in a moist environment. The Tulameen area has a moderately dry climate, and Brunisolic soils have developed in serpentinized peridotite and dunite. Brunisolic soils with composite profiles of tephra overlying serpenite developed in a dry environment at the Shulaps area. We investigated the weathering status of five soils using a variety of chemical and mineralogical techniques. Chlorite and mica weathering was indicated by XRD analysis of clay fractions for the Podzolic soils at the Coquihalla area. Serpentine dissolution was also indicated by geochemical analysis of these upland soil profiles. Weathering and leaching depleted Mg from B horizons relative to parent materials. Chromium accumulated in B horizons of the soils, and was associated with high levels of Fed. The distribution of total and extractable Ni and Co indicated that these elements were transported to the C horizons in all but the driest environments. Redistribution of Mn also occurred, but to a lesser extent. The distribution patterns of Cr, Mn, Co, and Ni were useful indicators during the early stages of ultramafic pedogenesis. Key words: Serpentine, ultramafic rocks, mineralogy, soil processes, trace minerals

Beschreibung: Pore size, distribution and continuity are important characteristics for the exchange and storage of air and water in artificial mixes. Saturated hydraulic conductivity (Ks) measurements can be used to obtain such a characterization. However, two difficulties are encountered when using Ks in potting media. First, the validity of Ks may be limited because it may not apply in media composed of coarse material or peat. Second, the structure of peat substrates is very sensitive and in situ measurements of potted peat substrates (i.e. measurements made directly in the pots) should be carried out to avoid any disruptive effect due to handling. Such a measurement, when made in pots, may require the evaluation of the water flux reduction resulting from the container outflow configuration. The objectives of this study were therefore to check the validity of Darcy’s law for peat substrates and to propose an approach for estimating the saturated hydraulic conductivity from flow measurements made in nursery containers. For three different substrates, water flow in artificial mixes followed Darcy’s law for hydraulic gradients ranging from 1.1 to 1.6 cm cm−1. Experimental results showed that the measured fluxes in 5-L nursery container filled at five different substrate heights (9, 11.5, 14, 16.5 and 19 cm) with laterally located drainage holes were significantly different from those measured in pots with the bottom removed (therefore equivalent to measurement currently made in cylinders) at P = 0.0022. Fluxes in containers with bottoms removed were 7–31% higher than in intact pots. Water flux measurements may therefore need to be corrected for this flux reduction in order to accurately estimate hydraulic conductivity from flow experiments run in pots. A correction factor based on the results obtained from a finite difference model was derived and calibrated. Then, this correction factor was used to convert flux measurements made in pots with lateral holes into equivalent flux that would have been obtained had the pot had an open bottom. After correction, no significant flux reductions were found between pots with open bottoms and pots with lateral holes (P = 0.55). A correction factor estimated from Laplace’s equation, once calibrated, can therefore be applied to flux measurements obtained from pots to obtain estimates of Ks of undisturbed potted media. Key words: Hydraulic conductivity, peat substrates, container

Beschreibung: The spatial distribution of soil properties, erosion and crop yield was studied along a cultivated transect and an adjacent transect in virgin grassland. The mapping unit was in hummocky moraine in the Dark Brown soil zone of central Saskatchewan. Erosion was affected by an interaction between elevation and surface curvature which affected the spatial and statistical distribution of soil properties and yield in the landscape. For the cultivated transect, percent sand was higher on knolls than in depressions, furthermore several soil properties were nonlinearly related to elevation. Decreases in grain yield on convex or planar lower slopes were correlated with erosion and high carbonate contents. Carbonate concentrations were high in lower slope positions in the cultivated transect, particularly at a relative elevation of 1 m, and appeared to be an important soil property influencing yield. Future studies of soil properties and yield on hillslopes may require careful evaluation of both elevation and microtopography in order to elucidate the effects of erosion. Key words: Erosion, yield, spatial variability, topography

Beschreibung: The risk of surface-water contamination by herbicides is greatest following application to cropland when the active ingredients are at the maximum concentration and the soil is the most vulnerable to erosion following cultivation. This study determined the magnitude of surface runoff losses of herbicide and nutrients at, and subsequent to, application. The first of three weekly 10-min, 2.6-cm rainfalls were simulated on triplicated 1-m plots (a set) on which corn had been planted and the herbicide (metolachlor/atrazine, 1.5:1.0) and fertilizer (28% N at 123 kg ha−1) had just been applied. Identical simulations were applied to two other adjacent plot sets (protected from rainfall) 1 and 2 wk following herbicide application. Runoff (natural, simulated) was monitored for soil, nutrient and herbicide losses. Concentrations of total phosphorus in surface runoff water and nitrate N in field-filtered samples were not significantly influenced by the time of the rainfall simulation but exceeded provincial water-quality objectives. Atrazine and metolachlor runoff losses were greatest from simulated rainfall (about 5% loss) immediately following application. Subsequent simulated rainfall usually resulted in

Beschreibung: The influence of 40 years of red pine and Norway spruce growth on forest floor and soil nutrient status was examined in a well-replicated series of plantation blocks established on abandoned agricultural land. Concentrations of N, P, K and Ca, and mass of organic matter and all nutrients in the forest floor were higher under spruce than under pine. In the mineral soil, concentrations of exchangeable K and Ca were higher under spruce whereas Mg, extractable P and mineralizable N did not differ between the species. Forest floor pH was higher under spruce but mineral soil pH did not differ between the species. The soil characteristics reflected litter chemistry of the two species. Relative to pine, spruce foliage litter was consistently higher in nutrient concentration and had lower acidity and higher ash bases. The results are inconsistent with the reputation of Norway spruce as a species that strongly acidifies soils, an observation that may be the result of elevated levels of Ca and K in our soils relative to those in other studies where acidification has been observed. This research demonstrates that soil fertility can be altered significantly by tree species effects over the period of a single rotation. Key words: Forest soil fertility, plantation, acidity, forest floor, leaf litter

Beschreibung: Lupin (Lupinus albus L. ’Ultra’) was evaluated as a green manure crop the year preceding potato production on Prince Edward Island for 3 yr. One pass of tandem disc was used to incorporate lupin green manure on 1 September (early) or 1 October (late). Barley residue following combine harvest was also incorporated in the same way in early September. One treatment of lupin was left unharvested with no fall incorporation (none), and in another treatment of lupin the seed was fall harvested with no fall incorporation. All treatments were moldboard plowed the following spring. Potato (Solanum tuberosum L. ’Russet Burbank’) was planted with three rates of nitrogen (0, 50, and 200 kg ha−1) band applied. Tuber yield and tuber N content increased with increasing rates of N applications in all incorporation treatments. Following late incorporated lupin, potato had higher tuber yield at 0 N than following the early incorporated lupin, while both treatments were higher than barley. Potato petiole nitrate-N levels and the mean spring nitrate-N concentrations in the soil were lower following barley than following lupin, and generally lower following early compared to late incorporation of lupin green manure. These data indicate the time of fall incorporation of a legume can have a significant effect on the N response of a succeeding crop. Early fall incorporation can result in significant fall N mineralization and winter leaching. Research on N response after legumes in rotation should consider the impact of time and method of residue incorporation. Key words: Petiole nitrate-N, nitrate leaching, green manure

Beschreibung: The soil solution provides the majority of essential nutrients needed for plant growth. Obtaining this solution in situ presents logistic problems and therefore most results are obtained from air-dried soils, even though chemical properties of soils can be substantially modified by drying. Water-saturated extracts obtained from dried soils were tested for their efficiency to detect the effects of in situ fertilization with P-TSP, K-K2SO4 and Ca-Ca(OH)2 on the Ahe horizon of a maple stand. Results obtained with water-saturated extracts on dried soils were compared with those obtained from conventional (exchangeable) extraction on dried soils and to those obtained from extraction with a soil:water ratio of 1:5 on dried soils. A similar experiment comparing water-saturated extracts with the soil:water ratio of 1:5 was carried out using soils fertilized in vitro. The effects of fertilization were clearly evident with saturated extracts when these could be observed by conventional analysis. Furthermore, the saturated extract indicated potential toxicity of Al in relation to pH. For the same samples, the saturated extracts were more sensitive than the extracts with a soil:solution ratio of 1:5, especially with regard to element ratios not easily expressed by conventional analysis. The originality of this method rests on a combination of efficient centrifugation with a double bottom container, the convenience in the utilization of air-dried soils and the use of near water field capacity as a representative water content. Key words: Saturated extracts, fertilization, toxicity, maple stand

Beschreibung: The spatial distribution of cesium-137 was examined at two uncultivated sites in the Black soil zone of Saskatchewan. Although considerable variability occurred at both sites, the variability was not systematically related to landscape position. Between 9 and 16 samples need to be taken from uncultivated grassland sites to reliably estimate the mean 137Cs concentration at such sites. Key words: Cesium, soil redistribution, landscape

Beschreibung: The distribution and dynamics of N following green manuring of full bloom field pea (Pisum sativum ’Sirius’) were investigated in the soil mineral, microbial and non-microbial organic (NMO N) fractions at two contrasting field sites in central Alberta; one on a Chernozemic (Dark Brown) soil near Provost and the other on a Luvisolic (Gray Luvisol) soil near Rimbey. Soils were sampled four times during a 1-yr period. The accumulations of N in the whole soil and in the soil mineral and microbial fractions were similar between sites. Net mineralization rates under controlled environments were strongly influenced by pre-incubation soil conditions. The short-term dynamics of N were distinguished best between the two soils by mineralization rates normalized to selected soil fractions rather than on the basis of N accumulation in these fractions. Net N mineralization rates expressed on the basis of soil N, microbial N or NMO N were greater in the Luvisolic soil indicating a more rapid internal N cycling system and greater activity of microbial biomass. These observations were consistent with the hypothesis that higher rates are associated with soils of lower clay content. Key words: Soil N dynamics, N mineralization rate, legume green manuring, Chernozemic, Luvisolic

Beschreibung: Studies were conducted to determine whether KCl application alone or in combination with Mn fertilizer would increase the bioavailability of indigenous and applied Mn and to study the mechanisms by which KCl could enhance Mn uptake by crops. The effect of rates of application (0, 50, 100 and 300 μg K g−1) of two KCl carriers (fertilizer grade and reagent grade KCl), as well as the effects of various salts (KCl, MnSO4, K2SO4, K2CO3, KCl + MnSO4, and KCl + MnSO4 + KI) on Mn uptake by wheat were examined. The effect of KCl on the solubility of soil Mn and applied Mn and the effect of K2SO4 on solubility of soil Mn were examined in calcareous and noncalcareous soils.The two KCl carriers behaved similarly in affecting Mn uptake by wheat. On the calcareous soil only the highest rate of KCl significantly enhanced Mn uptake, whereas on the slightly acidic soil both 100 and 300 μg K g−1 treatments significantly increased Mn uptake. Different salts differed significantly in affecting Mn uptake by wheat, and their effect was observed in the order KCl 〉 K2SO4 〉 〉 K2CO3 in both soils, with Mn uptake with K2CO3 being less than in the control. Application of KCl alone, at the higher rates, was as effective as MnSO4 alone in increasing Mn uptake by wheat. Mn uptake by wheat with dual application of MnSO4 and KCl was greater than with separate applications of MnSO4 or KCl. Addition of KCl to MnSO4 fertilizer bands increased amounts of water-soluble Mn in the fertilizer bands. K2SO4 was almost as effective as KCl in increasing water-soluble Mn in MnSO4 fertilizer bands but only for short periods of time (1–3 d). In contrast, KCl increased Mn solubility in the MnSO4 bands for periods up to 56 d. Key words: KCl fertilizer, KCl-enhanced Mn uptake, Mn availability, Mn solubility

Beschreibung: Distribution of NO3, P, K, Cl, pH and conductance through the soil profile were measured on two soil types after 4 yr of crop production using zero tillage (ZT) or conventional tillage (CT), with or without addition of KCl. All plots received N and P fertilizer each year as banded applications. Surface concentrations of NO3-N were higher under ZT than CT, particularly on the fine sandy loam soil. Accumulation of NO3-N also occurred in the 60- to 120-cm zone, under both tillage systems in both soils. Carryover of NO3-N was substantially greater on the silty clay than the fine sandy loam soil. Phosphate accumulated at the depth of band application in both soils under both tillage systems. Potassium concentration was generally higher under ZT than CT in the surface 15 cm of both soils, presumably due to surface retention of K from fertilizer applications and crop residues. Chloride was higher under ZT than CT in the surface 5 cm of both soils, but was higher under CT than ZT in the 30- to 60-cm and 60- to 120-cm depths in the silty clay soil, if KCl had been applied. The pH on both soils under both tillage systems was reduced in the 10- to 12.5-cm soil depth, corresponding to the zone of fertilizer application. On the silty clay soil, pH was higher under ZT than CT in the 10- to 15-cm depth and tended to be higher under ZT than CT at all depths below 15 cm. Conductance was not influenced by tillage in either soil. Application of KCl increased K and Cl concentrations in the surface 15 cm on both soils. Concentration of Cl was increased to 120 cm in both soils, indicating the mobility and leaching potential of this anion. Conductance and pH were increased in the 2.5- to 5.0-cm and 10- to 12.5-cm depths by KCl application in the fine sandy loam soil, but on the silty clay soil, only conductance was increased. Key words: Zero tillage, nutrient stratification, pH stratification

Beschreibung: Winter wheat, Triticum aestivum, is a new crop in south coastal British Columbia. The purposes of this study were to characterize plant development, dry matter accumulation and N uptake under low input and intensively managed systems as well as to assess the capability of some of the region’s soils to supply N to the crop. Grain yields, crop development and dry matter and N accumulation were similar to those reported from southern England. High amounts of winter rainfall (November–April precipitation ranged from 523 to 1111 mm) leach virtually all residual NO3 from south coastal B.C. soils and, without N fertilization, result in uniformly N deficient winter wheat. The low input N regime, 75 kg N ha−1 at Zadoks growth stage 31, plus soil N mineralized subsequent to the winter leaching period were sufficient in this study to maximize grain and total aboveground crop dry matter yields, but not to achieve adequate grain protein contents. The soils in the study were capable of supplying N in amounts sufficient to support only 30–53% of the maximum N uptake between growth stages 31 and 78. Appropriate quantities and timing of N are critical to successful production of high-yielding, good-quality wheat in south coastal British Columbia. Nitrogen management is likely to be most efficient when guided by the stage of crop development and demand and not by spring soil sampling and mineral N analysis. Key words: Winter wheat, N demand, soil N supply, crop development, intensive crop management, low input

Beschreibung: We investigated the nature of the layer charge of the different expandable 2:1 phyllosilicates present in glacial till, marine sediment, beach sand, loess, weathered sandstone and acid shale parent materials of selected soils from the arctic, alpine and interior plateau regions of Canada. Detailed characterization of 2:1 clay minerals, using the layer charge, was done by intercalation of the clay fractions with different chains (nc = 6 to 18) of n-alkylammonium cations (AC).Results indicated that AC were able to detect a mixture of smectite and vermiculite that was not detected by previous studies, using the glycerol (GLY) and ethylene glycol (EG) solvation techniques. The mean unit cell layer charge [in mol(+)/Si,Al)4O10)] varies from 0.30 to 0.39 for the smectite and from 0.71 to 0.74 for vermiculite. Vermiculite from the beach sand is trioctahedral in which most of the charge is located in the tetrahedral sheet, while vermiculite in the weathered sandstone is dioctahedral in which part of the charge originates from the octahedral layers. The presence of smectite, associated with the weathering of chlorite and an expanding component associated with the degradation of biotite, was also observed. Key words:n-alkylammonium cations, mean layer charge, 2:1 phyllosilicates

Beschreibung: The field persistence of trifluralin from fall treatments of both emulsifiable concentrate (EC) and granular formulations was investigated at two sites in Saskatchewan over a 2-yr period. At both locations, there were no significant differences (P = 0.05) in trifluralin persistence from fall applications of either formulation. In addition, the carryover of trifluralin residues in farmers’ fields following fall and spring applications of EC and granular formulations was monitored at several Saskatchewan locations to determine their persistence under typical farming practices. Trifluralin residue data indicated that following fall applications, the percent carry-over to the first spring was 83 ± 21(15 fields sampled) and 37 ± 15 (28 fields sampled) to the second spring; and that following spring applications, the percent carry-over to the next spring was 33 ± 12 (14 fields sampled). Key words: Field studies, persistence, formulations, trifluralin, herbicide

Beschreibung: The Mn release by M KCl from two Oxisols from Indonesia and one Inceptisol from Oregon, USA was studied at different temperatures (25, 35 and 45 °C) to understand the kinetics and mechanism of Mn release from these soils, which were reported to have a Mn toxicity problem. The Mn release by M KCl continued to increase with time up to at least 772 h without reaching equilibrium. The kinetics of Mn release followed a parabolic diffusion model during the 0.25–772 h reaction period. The activation energy of Mn release, calculated from the overall diffusion coefficients, varied from 34 to 47 kJ mol−1. Even after removing the exchangeable Mn from the soils with M calcium nitrate, significant amounts of Mn were released by M KCl. Data presented indicate that ionic strength effect coupled with Cl complexation was the main mechanism governing the KCl-induced Mn release. The Mn release from the soils during the initial reaction period (0.25 h) was fast and accounted for 34–39% of the total amount of Mn released during the 772 h reaction period. The significantly high Mn release during the initial 0.25 h reaction time, the high overall diffusion coefficients of Mn release, and the high values of the pre-exponential factor (calculated from the Arrhenius equation) of Mn release from Mn-toxic soils apparently contributed to the KCl-induced Mn toxicity in these soils. Key words: Potassium chloride-induced Mn toxicity, pre-exponential factor, ionic strength, complexation

Beschreibung: Mineral composition of the parent material is a major factor controlling natural radioactivity of soils. The objective of this study was to investigate the relationship between the natural radioactivity of 18 till and six lacustrine parent materials and their properties measured during soil surveys. Gamma spectroscopy was used to estimate 40K and daughters of 238U and 232Th; neutron activation analysis was used to determine total U; and alpha counting was used to measure 222Rn release from the soil. The amounts of the various radioisotopes were positively correlated with each other and with clay content; the correlation with sand content was negative; and the correlations with silt and inorganic C content were generally not significant. Adding the fine/total clay ratio as a further independent variable considerably improved the correlation between 40K and clay content. The data suggested considerable disequilibrium in the 238U decay chain, which appeared to be in part related to the mode of deposition of the parent materials. Soil survey information — in particular, texture — thus appears a readily available tool for preliminary screening of terrestrial radiation in Saskatchewan. Key words: Radioactivity, 40K, uranium, thorium, radon, texture

Beschreibung: Barley receiving unlabelled nitrogen fertilizer was grown among treatments receiving Ca(15NO3)2 fertilizer (5 atom %) in a growth cabinet with high air exchange and in a greenhouse. In both environments, the unlabelled plants and soil were significantly enriched with 15N, apparently, in part, by foliar exchange of atmospheric ammonia. This exchange may warrant consideration in the selection of appropriate controls for 15N calculations. Key words:15N, ammonia, atmospheric exchange

Beschreibung: Palaeosols, formed prior to deposition of Mazama tephra (6600 yr B.P.), are widespread in Alberta. Palaeo-Ahb horizons are readily identifiable, with colors ranging from very dark gray to gray and brown. Establishment of fundamental and stable properties of the original organic matter would facilitate the classification of these buried soils. Use of a suite of eight monosaccharides divided the 15 palaeosols examined in this study into one group of three, possibly formed under Brown Chernozemic soil-forming conditions or drier, and the remainder, possibly formed under Black Chernozemic soil-forming conditions. The former had C/N ratios approaching those of microorganisms and low xylose to mannose ratios. It is concluded that, because of the long time-window of surface stability between deglaciation and the Mazama tephra fall, pre-Mazama palaeosols can only be studied on a site-specific basis. Grouping these palaeosols, based on their pre-Mazama tephra deposition only, is not realistic. Synthesis of a unified hypothesis of palaeoenvironmental conditions is, therefore, not possible using only their monosaccharide status. Key words: Holocene soils, palaeosols, buried soils, monosaccharides, landscape

Beschreibung: In order to assess to what extent soil nutrient properties support differentiation of field-identified soil nutrient regimes, composite samples from each forest floor and 0–30 cm of the mineral soil were collected from 116 forest stands in central British Columbia. The samples were analyzed for acidity, total carbon (tC), total nitrogen (tN), mineralizable-N (min-N), and extractable Ca, Mg, K, P, and SO4-S (eCa, eMg, eK, eP, eSO4); and the results were expressed as concentrations on a dry-mass basis. Mineralizable-N of the mineral soil showed (1) the largest amount of variation in the population of sampled soils, (2) significant differences (P 

Beschreibung: Kochia scoparia (L.) Schrad. naturally invades mechanically disturbed soils in the Brown and Dark Brown soil zones of the Canadian Prairies and offers potential as a forage crop in saline environments. A 3-yr study conducted on severely saline soil (~ 11 dS m−1) near Swift Current, Saskatchewan, evaluated the production of kochia forage and plant-N uptake following applications of 0, 56, 112 and 168 kg N ha−1 of ammonium nitrate. The yield response in aboveground, dried forage (Y) followed a curvilinear function based on the quantity of nitrogen applied (N): Y = 4740 + 38.5N − 0.121N2. The N-fertilizer requirement to produce 7500 kg ha−1 of dried forage (96% of the function maximum) equalled 110 kg N ha−1 and reflected average growing conditions at the study site during 1987–89. The kochia assimilated protein N (Kjeldahl) within its tissue in direct proportion to the fertilizer dosage applied, reaching theoretical maxima which varied yearly: 18 g kg−1 in 1987, 12 g kg−1 in 1988 and 22 g kg−1 in 1989. Although NO3-N concentrations increased with each addition of fertilizer N, the maximum accumulation of NO3 N (0.5 g kg−1) among all the tests and treatments was within the safe limit of 1.1 g kg−1 to avoid livestock poisoning. The efficiency with which the fertilizer N was assimilated by the kochia ranged between 44 and 69% over the test years and fertilizer treatments. Key words: Saline soil, N fertilization, kochia forage, soil fertility, salinity