ALBERT

Description: Statolith microstructure was studied in 162 specimens of Illex coindetii (mantle length ranging from 48 to 300 mm) captured on the shelves of Sierra Leone and Western Sahara between May and November 1987. Growth increments were revealed in all statoliths studied, but they were not well-resolved as in other congeneric species Illex illecebrosus and Illex argentinus . Age and growth rates were estimated assuming that growth increments within statoliths were produced. In both regions, differences in growth rates between sexes (females become larger than males) occurred first in length and then in weight. Geographical differences in sex-specific growth (Western Sahara squid grow faster and attain larger sizes than Sierra Leone squid) was apparent after 150 d in females and 120 d in males. Illex coindetii in Sierra Leone started maturing at smaller sizes but at approximately the same age as the Western Sahara squid. Small sized early maturing groups of I. coindetii with a life span of ~0.5 y occurred in both regions. However, large sized late maturing squid with a life span of ~1 y were encountered only in the Western Sahara. In waters of the west African shelf, I. coindetii spawn throughout the year.

Description: Stomach contents of Todarodes sagittatus caught by trawlers working from 100 to 800 m depth in the Balearic Sea (western Mediterranean) were studied. From the 348 stomachs examined (153 males and 195 females) 33.62% were empty (39.21% in males and 29.74% in females). The diet of the squid was composed of 58 different prey items belonging to four major groups: Osteichthya, Crustacea, Cephalopoda and Chondrichthya. Osteichthyes, crustaceans and cephalopods were the most common prey, with a frequency of occurrence value of 84.85, 48.92 and 29.87% respectively. A change in the diet as the squid grows was observed, since juveniles feed basically on fishes while adults prey more actively on crustaceans. Analysis of the diet by size-classes reflected an ontogenetic migration to deeper waters since, parallel to the increase of size, a raise in the percentage of prey species inhabiting deeper waters was detected. Cannibalism was quite frequent, since T. sagittatus was the second most common cephalopod prey. Females had higher fullness-weight index and lower emptiness index than males, which reflects their major energetic demand for egg production.

Description: The size of fish and squid prey of Loligo forbesi was investigated using otoliths, beaks and statoliths collected from stomach contents analysis of samples obtained from Scottish and Irish waters between 1990 and 1993. Loligo forbesi was found to consume a large range of prey sizes, but prey size was always less than the predator size. Season was shown to significantly influence the predator size-prey size relationship for sprat and sandeel prey, but this itself could be influenced by seasonal changes in the size of prey. Fish prey size increased with increased predator size up to a mantle length (ML) of 200 mm. Loligo forbesi of mantle length 〉200 mm consumed a range of prey sizes, with no clear increase in the size of prey. For most prey taxa the relationship between prey size and squid size was similar, the exceptions being dragonets and silvery pout. Cannibalism by L. forbesi was mostly limited to larger L. forbesi (〉150 mm ML) feeding on smaller (20–50 mm ML) conspecifics.

Description: The population structure of the European flying squid Todarodes sagittatus was studied using data of about 5000 squid caught in waters off the western Sahara between 1969 and 1997. The bulk of the population consisted of winter-spawned squid, which occurred as juveniles of 80–120 mm mantle length (ML) over the slope in spring, and as immature squid of 160–180 mm ML both on the shelf and slope in summer. In autumn, the squid attained 220–280 mm ML, matured, and shifted to the slope, where the spawning was supposed to occur in winter. Age and growth of T. sagittatus was studied using statolith ageing techniques. Assuming daily production of putative growth increments within statoliths, as well as sizes and proportions of immature and maturing females, the lifespan of the west Saharan populations of T. sagittatus lasts ∼1 y. Todarodes sagittatus is a fast growing squid at juvenile and immature ontogenetic phases. Early maturation (at ages 220–230 d in males and 250–260 d in females, respectively) and subsequent decrease of somatic growth rates caused rather small modal sizes of mature squid (250–300 mm ML) compared with those of their northern counterparts (350–420 mm ML). Both hatching dates and seasonality in occurrence of mature females shows that in waters off the western Sahara T. sagittatus spawns throughout the year with a well-pronounced winter peak.

Description: Size of ripe eggs and potential fecundity are described in the squid of the subfamilies Todaropsinae and Todarodinae (Oegopsida: Ommastrephidae)— Todaropsis eblanae from West Africa, Todarodes angolensis from Namibia, Todarodes sagittatus from north-west Africa and the Mediterranean Sea, Todarodes sp. from the south-east Pacific, Nototodarus hawaiiensis from the south-east Pacific and West Indian Ocean and Martialia hyadesi from the south-west Atlantic. Females of both subfamilies are characterized by a wide range of ripe egg size (0.7–2.4 mm) and low and medium values of potential fecundity (20,000–2,500,000).

Description: Records of unusual cephalopods, taken as by-catch in Irish and Scottish waters in the years 1985–1995, are presented. Of most interest are three specimens of giant squid (Architeuthis) that were caught in bottom trawls off the west of Ireland between April and June 1995, all were mature males of mantle length ∼1000 mm. Other records include a large mature female Histioteuthis bonnellii from the west of Ireland and three specimens of the gelatinous incirrate octopus, Haliphron atlanticus.

Description: Octopuses ( Eledone cirrhosa (Octopoda: Cephalopoda)) held in an aquarium were subjected to varying conditions of feeding and starvation to evaluate putative indices of feeding and growth. Specific growth rate (%d −1 ) was linearly related to feeding rate (% of the body mass d −1 ) in animals with a mean body mass of 250 g at 15°C. Maximum growth rates varied between 〉 2% d −1 (body weights 〈 300 g) to 〈 1% d −1 (body weights ≤ 900 g) but specific growth rates were not related to body weight. Growth rates became negative (weight loss) after one week without food. The digestive gland index (DGI) was significantly correlated with short and long-term feeding and specific growth rates, and with body weight. Muscle RNA concentration was linearly correlated with growth rate during the previous 1–3 weeks but not with feeding rate. RNA:protein ratios were not different between mid-arm and mantle sample sites but arm tip values were significantly higher. RNA:protein ratio was related to body weight only in feeding animals. It is concluded that DGI is an index of feeding rate and that RNA:protein ratio can be used as an index of recent (~ 4 weeks) growth rate.

Description: A specimen of Chaunoteuthis mollis (Cephalopoda: Onychoteuthidae), a mature mated female, from the Straits of Messina was examined. Measurements and indices were compared with those of Onychoteuthis banksii reported by previous authors. The differences found were attributed to the degeneration of the C. mollis specimen, due to sexual maturation. Two subocular (one under each eye) and two visceral photophores, similar to those of O. banksii , were present in the C. mollis specimen. The size of the posterior visceral photophore of C. mollis fit the regression line correlating posterior photophore diameter to mantle length in O. banksii. These results corroborate the hypothesis, put forward by other authors, that C. mollis represents the mature female of O. banksii.

Description: Three specimens of the giant squid Architeuthis were by-caught in demersal trawls to the west of Ireland, between April and June 1995. All three specimens were mature males, of mantle length 975–1084 mm. Although some intraspecific variation in fin, beak and gill were noticed, all three specimens were tentatively assigned to the species A. dux. The three specimens had food remains in their stomachs and food items identified included Micromesistius poutassou, Trachurus trachurus, Nephrops norvegicus and Eledone cirrhosa . Age estimates were made by counting putative daily growth increments in ground statoliths, and ranged from 294–122 d, giving percentage daily growth rates of 2·96–4·25% indicating a short life cycle and extremely rapid growth.

Description: Deep-water trawling for fish species on the shelf slope off Scotland results in a by-catch of rare cephalopod species. The cirrate octopus Opisthoteuthis grimaldii is normally present below 800 m; at least one other species of cirrate and five species of incirrate octopus are found; significant extensions of range and depth are recorded. An incidental catch of the ommastrephid squid, Todarodes sagittatus , included a fully mature specimen and is indicative of its probable breeding locations. The results highlight the many taxonomic and systematic uncertainties surrounding the rarer cephalopod forms, and emphasize the utility of the fishing by-catch.

Description: Two large (dorsal mantle length 42.5 and 47.5 cm), mated spent females of circum‐Antarctic bathypelagic cranchiid squid Galiteuthis glacialis were caught early in March 1992 at the surface of the ice hole in the western Weddell Sea over depths 1915‐1920 m by the team of the U.S.A.‐Russian Ice Station Weddell‐I. The structure of the reproductive system of adult females is described for the first time in detail. Both were gelatinous, devoid of tentacles, with empty or almost empty stomachs. The empty spermatangia (sperm reservoirs of spermatophores) 30‐35 mm in length were distributed in the mantle tissues parallel to the mantle surface and to each other in the dorso‐anterior part of the mantle: 13 in one female, parallel to the body axis, and 20 in the other, parallel (13) or perpendicular (7) to the body axis. In the latter case, they represented probably two mating events. The spermatangia lay nearer to the inner than the outer mantle side and opened by a round window on the inner side; the skin with chromatophores above them remained intact. The spermatozoa had one flagellum and rod‐like heads, length 5.0‐5.3 μm, width 1.2‐1.5 μm. The most characteristic features are: a very simple type of blood vessel branching making each micro‐gonad currant‐like, not grape‐like; a very compact disposition of oviducal, nidamental glands and gill, forming a united complex located on both sides of the mantle cavity; and an ovary connected by mesentery along all its length with the continuation of the stomach from the caecum to the end of the gastrogenital ligament. Only immature degenerating trophoplasmatic oocytes, length 0.9‐1.4, av. 1.0‐1.2 mm, were contained in ovaries; only one mature egg (length 3.3 mm, width 2.4‐2.5 mm) was found in each female. The absence of oocytes 〈0.9 mm and 1.5‐3.2 mm indicates that the maturation of oocytes proceeds rather synchronously, one large portion of eggs (some tens of thousands) matures in a short time while others degenerate. The residual fecundity is assessed to be approximately 20,000 eggs. It is hypothesized that mating occurs shortly before spawning and that mature males do not undergo gelatinous degeneration and do not lose tentacles. Spermatophores are placed on the inner side of the female's mantle with the aid of the male's tentacles and/or arms (less probably by the penis), but the exact mode of implantation is unclear. Spawning probably occurs at depths of adult habitat (approx. 500–2500m), may be multiportional but short; the exhausted female loses neutral buoyancy, rises to the surface and dies. Rising to the surface after spawning is a common feature of females of many meso‐ and bathypelagic squids undergoing gelatinous degeneration during maturation (Onychoteuthidae, Gonatidae, Histio‐teuthidae, Cranchiidae, etc.) and may explain the common occurrence of large deep‐water squids in the stomachs of seabirds, including those incapable of diving, and marine mammals.

Description: Growth in Loliolus noctiluca (Myopsida: Loliginidae) in Western Port, Victoria, Australia was studied from statolith growth increments. Tetracycline staining experiments verified previous work on tropical forms of this species that showed growth increments to be deposited daily. A logistic growth function described the relationship between length and increment number. There appear to be major differences in the form of growth, longevity and life history pattern between tropical and temperate forms of this species. These are probably attributable to differences in environmental conditions.

Description: Cephalopod prey of several cetacean species from Hawaiian waters were identified and quantified from the beaks in stomachs of stranded individuals. The different species of cetaceans all appear to target different species and sizes of cephalopods. Beaks from two sperm whales ( Physeter catodon ) included a total of 312 upper and 292 lower beaks (mandibles) of cephalopods. All of the cephalopods represented by lower beaks were oceanic squid belonging to 20 or more species in 14 families. The major constituents of the whale diet were Histioteuthis hoylei (45% by number and 10·9% by dry weight), Ommastrephes bartrami (7·6% by number, 30·6% by weight) and Architeuthis sp. (only 0·7% by number, but 26·5% by weight). In a Risso's dolphin ( Grampus griseus ) stomach, 1051 upper beaks and 1349 lower beaks were present. Eighteen or more genera of cephalopods in 15 families were present. The major constituents of the Grampus diet were Enoploteuthis spp. (36·1% by number and 30·1% by dry weight) and Abraliopsis spp. (23·6% by number, but only 4·9% by weight) and O. bartrami (1·3% and 32·1% respectively). The melon-headed whale ( Peponocephala electra ) eats fish as well as cephalopods which are represented by only six lower beaks belonging to five genera, Enoploteuthis, Teuthowenia, Abraliopsis, Abralia sp. and Bathyteuthis abyssicola . The beaked whale (unidentified) contained two lower beaks which were O. bartrami. Stenella attenuata also eats fish and the sample contained lower beaks, one of Enoploteuthis sp. and seven of an Abraliopsis sp.

Description: Observations on samples from Spanish trawlers between September and November of 1995 revealed the presence of mated females of Loligo gahi (Cephalopoda: Loliginidae) from 164–285 m depth, in the Western area of the Falkland Islands Conservation Zone. 93.8% of the mature females, and 31.0% of the maturing ones, were mated. Deposition of spermatophores always took place in the oral membrane between the connectives of arms IV. The relationships between sexual maturation and copulation have been analysed, and the hypothesis of mating acting as a ‘trigger’ of the final sexual maturation in Loligo gahi females is proposed and discussed.

Description: This paper gives morphometric variations and dorsal mantle length–total weight (DML-TW) relationships for Illex coindetii in the eastern Central Atlantic area. Positive allometry was observed in males and negative in females of the species. The most variable body measurements between males and females were width and perimeter of the head. In the study area, divergence of morphometric measurement starts at 95 mm. The point of divergence, however, varies with latitude; fluctuating from 104 mm in the north (Morocco and Sahara) to 76 mm in the central area (Mauritania and north of Senegal) and 73 mm in the south (Gulf of Guinea). The relationships between DML and TW showed that sexual dimorphism due to differential growth between males and females starts to occur at 56 mm ML. The starting point of sexual dimorphism (56 mm) varies according to the zone; the higher the latitude, the later it occurs. Thus, sexual dimorphism occurs at 49 mm in the south (Gulf of Guinea), at 54 mm in the central area (Mauritania and north of Senegal) and at 74 mm in the north (Morocco and Sahara). Females grow larger than males, but males were heavier at any given length. As latitude decreased, a slow down in the increase in weight-at-length was observed in both sexes.

Description: A total of 57 comparative hauls using a rectangular midwater trawl with a fishing mouth area of 50 m 2 (RMT 50) were carried out along the sides of an imaginary triangle south of Madeira in 1986. A total of 1258 cephalopods were caught, giving a mean of 22 per haul with a range from 0 to 67. The nets were used with a diver's light on the top bar which was either switched off or was operated with a 20, 70 or 150 W bulb, powered by a car battery. A significantly greater number of individuals per haul was caught with lights on than without lights, increasing from a mean of 13·5–25·1, a factor of 1·8. Similarly, the number of species caught was increased from a mean of 7 to 10·4, a factor of 1·5 and the volume of cephalopods was increased from a mean of 41·1–162·3ml, a factor of 3·9. Similar comparisons made for catches during day or night separately and on the three courses separately also showed marked increases with the lights. Samples show that increase in power of the lights increased the total number of cephalopod individuals caught. In the 12 species with more than ten individuals, in 33 of the 36 comparisons (of number of individuals, species and volumes) there is an increase with the light. The most influenced species was Taonius pavo which increased in numbers by a mean factor of 3·9 times with 20W, 4·0 times with 70W and 6·1 times with 150W when compared with the numbers caught with no light.

Description: Recent attention to members of the sepiolid squid genus Euprymna and symbiotic associations with luminescent bacteria ( Vibrio fischeri strains) has prompted a review of this poorly-resolved group of squids. Twelve nominal species have been placed in this genus of which the majority are ill-defined, known only from their original descriptions and separated on the basis of inadequate characters. As a first step in resolving this group, a temperate Australian species, the Southern dumpling squid, Euprymna tasmanica , is here redescribed in detail. As the genus Euprymna currently stands, most members are only distinguished on the number and position of enlarged suckers in mature males. No diagnostic characters are available to identify females. All nominal species placed in this genus are reviewed and a key to proposed valid species is presented. Six species are considered here to be valid: Euprymna berryi, E. hoylei, E. morsei, E. scolopes, E. tasmanica and an undescribed species treated here as Euprymna sp. 1. Euprymna similis is a synonym of E. morsei of Japan. Due to inadequate original descriptions, and lost or poor type material, two species are considered here to be nomen dubia ( E. schneehageni and E. pusilla ), while the taxonomic status of four additional species remain unresolved ( E. albatrossae, E. bursa, E. phenax and E. stenodactyla ).

Description: The statolith microstructure was studied in 142 females (mantle length, ML, ranging from 77–402 mm) and 119 males (72–328 mm ML) of Martialia hyadesi caught on the Patagonian and Falkland shelves and at the Antarctic Polar Frontal Zone between 1989–94. The statolith microstructure dark zone in this species, contains narrower and more numerous growth increments than the dark zones of other ommastrephid squids. Assuming daily production of putative growth increments within statoliths males live up to 12 months, and females live up to 13 months. It is likely that the life cycle lasts c. 1 yr, but immature squids with ages 〉330–340 d suggest that a part of M. hyadesi populations could have life span 〉1 yr. Growth in length was best described by the Gompertz function, whereas growth in weight was best described by the logistic function. M. hyadesi is characterized by slow juvenile growth (〈100 mm ML), fast growth of immature squids and a sharp decrease in growth rates during maturation. M. hyadesi mature later (at ages 〉270 d) than other temperate ommastrephids, but maturation is rather rapid (2–3 months). In the south-west Atlantic, M. hyadesi hatch throughout the year.

Description: The current status of Antarctic Odontocetes – sperm whales Physeter catodon , killer whales Orcinus orca , long-finned pilot whales Globicephala melaena , hourglass dolphins Lagenorhynchus cruciger and poorly known species of beaked whales (family Ziphiidae)–were studied in Anatarctic waters using data gathered in sighting surveys conducted from 1976/77 to 1987/88. Temporal variation in density demonstrated the different migration patterns by species, especially between sperm whale and killer whale. Spatial distributions during mid-summer demonstrated different peaks of occurrence for each species by latitude that suggest possible segregation between the species. Killer whales occur mainly in the very southernmost areas, sperm whales in the southern half of the study area, beaked whales (mostly southern bottlenose whales Hyperoodon planifrons ) ranged over a wide area, and long-finned pilot whales and hourglass dolphins were mainly in the northern regions of Antarctic waters. Several longitudinal peaks of occurrence and apparent distribution gaps were identified for sperm, beaked and killer whales. Abundance estimates for south of the Antarctic Convergence in January are based on line transect theory and were 28 100 animals (coefficient of variation CV 0.18) sperm whales, 599 300 (0.15) beaked whales (mostly southern bottlenose whales), 80 400 (0.15) killer whales, 200 000 (0.35) long-finned pilot whales, and 144 300 (0.17) hourglass dolphins. Based on this, biomass of these species were estimated as 0.77 (sperm whales), 2.70 (beaked whales), 0.32 (killer whales), 0.16 (long-finned pilot whales) and 0.01 (hourglass dolphins) million tonnes. Consumption of food (mostly squid) by the Odontocetes is estimated as 14.4 million tonnes with 67% of the total consumed by beaked whales. Indirect consumption of Antarctic krill through the predation of squid by beaked whales is estimated to be c . 24 million tonnes. This value is similar to the estimate of krill consumption by penguins in the Antarctic (33 million tonnes). Odontocetes, especially southern bottlenose whales, are suggested to have a much greater role in the Antarctic ecosystem than has previously been considered.

Description: Subantarctic Marion Island is one of the few localities where the congeneric albatrosses Phoebetria fusca and P. palpebrata breed sympatrically. Chicks of both species at Marion Island were induced to regurgitate their stomach contents after being fed. Liquid formed over half the diet by mass. Cephalopods occurred most frequently in both species' diets. In terms of mass, cephalopods formed the larger part of the diet of sooty albatrosses, whereas fish was more important to light-mantled sooty albatrosses. Crustaceans and birds were also recorded for both species. Squid of the families Onychoteuthidae, Histioteuthidae, Chiroteuthidae and Cranchiidae occurred most abundantly in both species. Most squid taken by both albatrosses were of species known to float after death, suggesting that scavenging plays an important role in the species' foraging behaviour. Light-mantled sooty albatrosses consumed more squid restricted to the south of the Antarctic Polar Front than did sooty albatrosses, supporting a trend to latitudinal segregation of the two species while foraging.

Description: The El Niño/Southern Oscillation (ENSO) phenomenon is the strongest natural interannual climate fluctuation1. ENSO originates in the tropical Pacific Ocean and has large effects on the ecology of the region, but it also influences the entire global climate system and affects the societies and economies of manycountries2. ENSO can be understood as an irregular low-frequency oscillation between a warm (El Niño) and a cold (La Niña) state. The strong El Niños of 1982/1983 and 1997/1998, along with the more frequent occurrences of El Niños during the past few decades, raise the question of whether human-induced 'greenhouse' warming affects, or will affect, ENSO3. Several global climate models have been applied to transient greenhouse-gas-induced warming simulations to address this question4, 6, but the results have been debated owing to the inability of the models to fully simulate ENSO (because of their coarse equatorial resolution)7. Here we present results from a global climate model with sufficient resolution in the tropics to adequately represent the narrow equatorial upwelling and low-frequency waves. When the model is forced by a realistic future scenario of increasing greenhouse-gas concentrations, more frequent El-Niño-like conditions and stronger cold events in the tropical Pacific Ocean result

Description: Recent captures of two female giant squid ( Architeuthis ) off southern Australia have provided the first record of a mated female specimen of these almost mythical deepsea creatures. We found sperm packages (spermatophores) embedded within the skin of both ventral arms of the larger of the two specimens. It seems that male giant squids may use their muscular elongate penis to ‘inject’ sperm packages under pressure directly into the arms of females.

Description: Changes in the molecular structure of a highly ordered kaolinite, intercalated with urea and potassium acetate, have been studied using Raman microscopy. A new Raman band, attributed to the inner surface hydroxyl groups strongly hydrogen bound to the acetate, is observed at 3605 cm (super -1) for the potassium acetate intercalate with the consequential loss of intensity in the bands at 3652, 3670, 3684 and 3693 cm (super -1) . Remarkable changes in intensity of the Raman spectral bands of the low-frequency region of the kaolinite occurred upon intercalation. In particular, the 144 and 935 cm (super -1) bands increased by an order of magnitude and were found to be polarized. These spectroscopic changes provide evidence for the inner surface hydroxyl group-acetate bond being at an angle approaching 90 degrees to the 001 face. Decreases in intensity of the bands at 243, 271 and 336 cm (super -1) were observed. The urea intercalate shows additional Raman bands at 3387, 3408 and 3500 cm-1 which are attributed to N-H vibrations after formation of the urea-kaolinite complex. Changes in the spectra of the inserting molecules were also observed.

Description: Squid (Lolliguncula brevis) were exercised in a tunnel respirometer during a stepwise increase in water velocity in order to evaluate the anaerobic treshold, i.e. the critical swimming speed above which anaerobic metabolism contributes to energy production. The average anaerobic treshold was found at speeds of 1.5-2 mantle lenghts s-1. Above this velocity, α-glycerophosphate, succinate and levels fell and phospho-L-arginine was progressively depleted, while the levels of glucose 6-phosphate and inorganic phosphate rose. The finding of a simultaneous onset of anaerobic metabolism in the cytosol and the mitochondria indicates that a limited oxygen supply to the mitochondria elicits anaerobic energy production. This finding is opposite to the situation found in many other vertebrate and invertebrate species, in which energy covered by anaerobic energy production. This finding is opposite to the situation found in many other vertebrate and invertebrate species, in which energy requirements in exvess of aerobic energy production are covered by anaerobic metabolism, with mitochondira remaining aerobic. In L. brevis, swimming at higher speeds is associated with a small factorial increase in metabolic rate based on a high resting rate of oxygen cnsumption. Pressure recordings in the mantle cavity support this finding, indicating a high basal level of spontaneous activity at rest and a small rise in mean pressure at higher swimming velocity. Bursts of higher pressures from the jet support elevated swiming speeds and may explain the early transition to anaerobic energy production which occurs when pressure rises above 0.22-0.25kPa. The finding f mitochondrial hypoxia at a low critical speed in these squid is interpreted to be related to their life in shallow coastal and bay waters, which limits the necessity to maintain high swimming velocities. At increased swimming velocities, the animals oscilliate between periods of high and lo muscular activity. This behaviour is interpreted to reduce transport cost and to permit a longer-term net use of anaerobic resources when speed exceeds the critical value or when the squid dive into toxic waters. The simultaneous onset of anaerobic metabolism in the cytosol and the mitochondria emphasizes that squid generally make maximal use of available requirements are the highest among marine invertebrates.

Description: A mineralogical and 4OAr/39Ar study of 13 amphibole samples in the Kamila Amphibolite Belt and Kamila Shear Zone in northern Pakistan has found a correlation between the degree of greenschist facies alteration and quantity of excess 40Ar. Additionally, there is a north–south divide with amphibole samples from the northern region showing larger degrees of gree schist facies alteration, brittle deformation, and excess 40Ar incorporation compared to the predominantly plastically deformed, less altered, amphibole samples from the Kamila Shear Zone in the south. Acid leaching of two amphiboles from the Kamila Amphibolite Belt indicates that a large proportion of the excess 40Ar is correlated with later greenschist facies alteration hases, and can be easily removed by acid etching, thus revealing acceptable regional 40Ar/39Ar plateau ages.

Description: The results of an examination of one quarry within the gneisses of the Bartica Assemblage are described and the validity of these results throughout the whole of the Bartica Assemblage is discussed. The foliation and banding of the gneisses at Kereti Quarry are mainly the results of tectonism; concomitant metamorphism within the Amphibolite Facies was succeeded by a two-phase metasomatism involving successive additions of Na and K. These results appear valid throughout the Bartica Assemblage and it is further suggested that the main rock type distribution in the Bartica Assemblage is also controlled by tectonism.

Description: Textural differences in the occurrence of microcline define augen gneisses, subhedral porphyroblastic gneisses and euhedral porphyroblastic gneisses within the Bartica Assemblage, British Guiana. The introduction of microcline is metasomatic and the nature of the porphyroblasts appears to be a reflection of tectonic control during their development.

Description: In this paper we examine the steady, two-dimensional convective motion which occurs in a horizontal circular cylinder whose wall is non-uniformly heated. One observes several qualitatively different physical phenomena depending on the wall temperature distribution and the value of the Rayleigh number. The low-Rayleigh-number behaviour for the single convective cell heated from below is related to the classical Rayleigh stability problem. The critical Rayleigh number for the single circular cell is approximately five times the value for Rayleigh's multi-celluar configuration. The flow which exhibits a nearly parabolic velocity profile near the critical Rayleigh number, gradually changes to a rigidly-rotating-core behaviour as the Rayleigh number increases. The speed of core rotation is a function of the Prandtl number, whereas the boundary-layer thickness is primarily a function of the Rayleigh number. When the heating is from side to side, the solution shows that as the Rayleigh number increases the core motion is progressively arrested leaving a narrow circulating band of fluid adjacent to the wall. An oblique heating produces a hybrid phenomenon, a low-Rayleigh-number behaviour which is characteristic of the sideways heating case and a high-Rayleigh-number interior motion characteristic of the bottom heating case. To determine the core motion in the high-Rayleigh-number limit, Batchelor's work concerning the uniqueness of incompressible, exactly steady, closed streamline flows with small viscosity is extended to include flows with small thermal conductivity.

Description: The energy flux in a finite-depth gravity-wave spectrum resulting from weak non-linear couplings between the spectral components is evaluated by means of a perturbation method. The fifth-order analysis yields a fourth-order effect comparable in magnitude to the generating and dissipating processes in wind-generated seas. The energy flux favours equidistribution of energy and vanishes in the limiting case of a white, isotropic spectrum. The influence on the equilibrium structure of fully developed wave spectra and on other phenomena in random seas is discussed briefly.

Description: The genus Illex is likely to constitute a large portion of the annual world ommastrephid squid catches (Roper et al, 1984), even though specific official statistics are difficult to obtain. The broad-tail short-fin squid Illex coindetii is a widespread species ranging from the western to the eastern Atlantic (Roper et al., 1984) and east through the whole Mediterranean Sea (Mangold & Boletzky, 1987). Usually a by-catch of important fisheries, it is caught mainly by trawlers. Although its economic value is lower than that of other squid species (i.e. Loligo spp.), in the Sicilian Channel Illex coindetii may represent a valuable resource due to its abundance. In Italian waters, the available statistics (Cingolani et al., 1986) report that 2680 tonnes of ommastrephid squid were landed in 1982 (0.5% of the total landed catch). The main component of these was landed in Sicily (2183 tonnes), a consistently large part of which was no doubt Illex coindetii (Ragonese & Jereb, 1992). The catches came mainly from southern Sicilian waters (Sicilian Channel) where one of the major Mediterranean landing places is in Mazara del Vallo. Large trawlers (up to 200 gross tonnage) usually carry out long fishing trips (15–20 d), and Illex coindetii is caught mainly by those targeting Parapenaeus longirostris and Merluccius merluccius (Jereb & Ragonese, 1991).

Description: The natural feeding of the two most abundant ommastrephid squid (Cephalopoda: Ommastrephidae) in Galician waters was studied and compared. A sample of 334 stomach contents of Todaropsis eblanae (34–222 mm ML) and 267 stomach contents of Illex coindetii (50–379 mm ML) caught by commercial trawlers was examined. A total of 21 (T. eblanae) and 23 (I. coindetii) different prey items, belonging to three zoological groups (Teleostei, Crustacea and Cephalopoda), were taken by these cephalopods. However, 43% of the T. eblanae diet comprised only one fish species, Micromesistius poutassou. The diet of these squid species was significantly influenced by the geographical area (both species), size (T. eblanae) and maturation (I. coindetii). Feeding rate of both species decreased with size, but the percentage of stomachs with food remains increased in maturing and mature females. Weight of prey captured was dependent on available prey sizes and, in small individuals, maximum prey weight was very close to the squid weight. Both squid species are mainly neritic nekto-benthic predators, but I. coindetii seems to have a broader and more pelagic diet.

Description: This communication presents the first records of mated female Loligo gahi in Falkland Island waters. In October 1993 fully mature mated female L. gahi were identified in samples taken from the commercial fishery in waters east of Lively Island, East Falkland, at depths of 145–174 m. Spermatophores were found in both the mantle cavity and buccal sites of deposition. These records, combined with past records of spent females, suggest spawning periods in late October/early November and April/May. These concur with two of the three periods of spawning suggested from previous studies of juvenile and adult L. gahi.

Description: Age, growth and maturation of Loligo vulgaris were studied by examination of growth increments within statoliths of 294 specimens (mantle length, ML, ranging from 31 to 498 mm) caught on the west Saharan shelf between 1985 and 1988. Maximum age was 335 d (290 mm ML) for females, and 396 d (498 mm ML) for males. Growth rates varied considerably among individuals with a greater range in males. Sexual dimorphism in length was apparent after about 210 d. Males and females diverged considerably in weight, with males reaching a greater weight after about 180 d. Growth in length between 124 and 396 d was best described by a power function, while growth in weight was best described by the Gompertz function. Males started maturing at 180–210 d and mature males ranged in age between 250 and 396 d; while females started maturing at 240–270 d and mature females ranged between 285 and 335 d. Loligo vulgaris hatched throughout the year with two distinct peaks; in winter (December - early March) and summer (June-July). The life cycle of L. vulgaris populations on the west Saharan shelf lasts ~1 y, with large males (〉450 mm ML) living slightly longer.

Description: The reproductive strategies of male and female Loligo forbesi Steenstrup, 1856 were investigated from samples obtained from commercial catches and research cruises in Irish waters. In females maturity increased with size, but in males two modes in the size at maturity were identified, with approximately 40% mature at small size (180–200 mm mantle length), and the remainder mature at 〉250 mm mantle length. The difference in estimated age of the two modes of mature males was small, so size differences were probably due to different growth rates. Growth and maturation proceeded together in both sexes over much of the life-cycle. The effect of maturation on relative growth of somatic tissues was examined using analysis of covariance and multivariate regressions. In males there was a significant decline in total mass, and in mass of mantle, head and viscera, relative to mantle length with maturation. In females total mass was not significantly affected by maturation, but relative masses of head, mantle and viscera declined with maturation, indicating that energy was diverted from somatic growth to gonad production. Potential fecundity estimates were obtained by counting eggs and ova in the oviduct and ovary of mature females and were in the range 2500 to 10,500 (mean 5800). Fecundity was positively related to mantle length. The ovaries of mature females contained a range of egg sizes and developmental stages, indicating that spawning probably occurs intermittently.

Description: A total of 57 comparative hauls using a rectangular midwater trawl with a fishing mouth area of 50 m2 (RMT 50) were carried out along the sides of an imaginary triangle south of Madeira in 1986. A total of 1258 cephalopods were caught, giving a mean of 22 per haul with a range from 0 to 67. The nets were used with a diver's light on the top bar which was either switched off or was operated with a 20, 70 or 150 W bulb, powered by a car battery. A significantly greater number of individuals per haul was caught with lights on than without lights, increasing from a mean of 13·5–25·1, a factor of 1·8. Similarly, the number of species caught was increased from a mean of 7 to 10·4, a factor of 1·5 and the volume of cephalopods was increased from a mean of 41·1–162·3ml, a factor of 3·9. Similar comparisons made for catches during day or night separately and on the three courses separately also showed marked increases with the lights. Samples show that increase in power of the lights increased the total number of cephalopod individuals caught. In the 12 species with more than ten individuals, in 33 of the 36 comparisons (of number of individuals, species and volumes) there is an increase with the light. The most influenced species was Taonius pavo which increased in numbers by a mean factor of 3·9 times with 20W, 4·0 times with 70W and 6·1 times with 150W when compared with the numbers caught with no light.

Description: PEEM allows ‘real-time’ observations to be made of solid-state transformations and other high-temperature processes taking place during vacuum-heating up to c. 2000°C The solid state transformations of an amphibole-rich hornblendite specimen have been observed in the temperature range of 750–1000°C (± 50°C Between c. 970–990°C a rapid change in orientation contrast was observed, indicating a structural rearrangement from an oxyhornblende crystal lattice to a clinopyroxene structure. This phase retains the original amphibole shape and texture (including two 120°C intersecting cleavage traces), but possesses a clinopyroxene crystal structure. At higher temperatures this phase is seen to decompose, forming iron oxides and other fine-grained products. PEEM has provided useful information on both the nature and rates of transformation of natural amphiboles which has proved invaluable in our understanding of the mineralogically-controlled mechanisms of argon release during 40Ar/39Ar dating of amphibole samples.

Description: Density-dependent effects of Anguillicola crassus larval infections in the copepod intermediate host were examined experimentally. Three species of copepods (Cyclops vicinus, C. viridis and C. fuscus) were subjected to a range of doses of larval A. crassus within infection arenas. Prevalence, intensity and parasite dispersion (variance: mean abundance) values increase and then approach an asymptote as infection dose increases. Infection parameters differ between species of copepod. Increasing temperature has a negative effect on the establishment of the parasite population within the intermediate host. Parasite-induced host mortality increases with dose. These mechanisms have the potential to regulate populations of A. crassus larvae within the copepod population and hence the whole suprapopulation.

Description: The size distribution of benthic nematodes was investigated along different gradients of food availability in various regions of the north-eastern Atlantic: I, across the continental margin and II, with increasing distance from the continental rise. An overall trend for miniaturization with increasing distance from the food source was found. Moreover, our results indicate that seasonally varying food supply or a periodically pulsed input of organic matter to the sea floor affects nematode size spectra. The hypothesis is proposed that the life cycle of deep-sea nematode species and hence the size structure of their populations are related to seasonal energy availability. This dependence might result in one year life spans of deep-sea nematodes and probably other meiofauna.

Description: Arabian Sea sediments record changes in the upwelling system off Arabia, which is driven by the monsoon circulation system over the NW Indian Ocean. In accordance with climate models, and differing from other large upwelling areas of the tropical ocean, a 500,000-yr record of productivity at ODP Site 723 shows consistently stronger upwelling during interglaciations than during glaciations. Sea-surface temperatures (SSTs) reconstructed from the alkenone unsaturation index (U K′ 37) are high (up to 27°C) during interglaciations and low (22-24°C) during glaciations, indicating a glacial-interglacial temperature change of 〉3°C in spite of the dampening effect of enhanced or weakened upwelling. The increased productivity is attributed to stronger monsoon winds during interglacial times relative to glacial times, whereas the difference in SSTs must be unrelated to upwelling and to the summer monsoon intensity. The winter (NE) monsoon was more effective in cooling the Arabian Sea during glaciations then it is now.

Description: In steady state, the export of photosynthetically fixed organic matter to the deep ocean has to be balanced by an upward flux of nutrients into the euphotic zone1. Indirect geochemical estimates2 of the nutrient supply to surface waters have been substantially higher than direct biological and physical measurements3, particularly in subtropical regions. A possible explanation for the apparent discrepancy is that the sampling strategy of the direct measurements has under-represented episodic nutrient injections forced by mesoscale eddy dynamics, whereas geochemical tracer budgets integrate fluxes over longer time and space scales. Here we investigate the eddy-induced nutrient supply by combining two methods potentially capable of delivering synoptic descriptions of the ocean's state on a basin scale. Remotely sensed sea-surface height data from the simultaneous TOPEX/Poseidon and ERS-1 satellite missions are assimilated into a numerical eddy-resolving coupled ecosystem–circulation model of the North Atlantic Ocean. Our results indicate that mesoscale eddy activity accounts for about one-third of the total flux of nitrate into the euphotic zone (taken to represent new production) in the subtropics and at mid-latitudes. This contribution is not sufficient to maintain the observed primary production in parts of the subtropical gyre, where alternative routes of nitrogen supply will have to be considered.

Description: Metazoan meiofauna were studied in replicated multiple-corer samples obtained at a bathyal site (1320–1360 m depth) in the Porcupine Seabight (51°36′N 13°00′W) before (April) and after (July) the delivery to the seafloor of a phytodetrital pulse originating from the 1982 spring bloom. In all samples the metazoan meiofauna was dominated by nematodes; harpacticoid copepods and their nauplii were the second most abundant taxon. Population densities and biomass were very similar in both sample sets, the only significant differences being in the numbers of ostracods (higher in April) and nauplii (higher in July). Furthermore, vertical distribution patterns in the top 5 cm of sediment indicate that the meiofauna did not migrate towards the sediment surface following the phytodetrital pulse. The lack of a metazoan meiofaunal response contrasts with published evidence, based on the same samples, for a substantial increase in the foraminiferal abundance following the sedimentation event. Thus our results suggest that metazoans (as a whole) fail to exploit and utilize phytodetritus as rapidly as foraminifera. This probably reflects the energetic expense of egg production coupled with frequently slower rates of somatic growth among metazoans. In addition, foraminifera may outcompete metazoans for detrital food because they possess extremely efficient food-gathering organelles (granuloreticulate pseudopodia) and are able to raise their levels of metabolic activity very rapidly. However, metazoan responses at the species level, or over longer time periods (〉3 months), would not have been detected and so remain a possibility.

Description: The nitrogen-isotope record preserved in Southern Ocean sediments, along with several geochemical tracers for the settling fluxes of biogenic matter, reveals patterns of past nutrient supply to phytoplankton and surface-water stratification in this oceanic region. Areal averaging of these spatial patterns indicates that reduction of the CO2 'leak' from ocean to atmosphere by increased surface-water stratification south of the Polar Front made a greater contribution to the lowering of atmospheric CO2 concentration during the Last Glacial Maximum than did the increased export of organic carbon from surface to deep waters occurring further north.

Description: The overflow and descent of cold dense water from the Denmark Strait sill-a submarine passage between Greenland and Iceland-is a principal means by which the deep ocean is ventilated, and is an important element in the global thermohaline circulation. Previous investigations of its variability-in particular, direct current measurements(1,2) in the overflow core since 1986-have shown surprisingly little evidence of long-term changes in now speed. Here we report significant changes in the overflow characteristics during the winter of 1996-97, measured using two current-meter moorings and an inverted echo sounder located at different depths in the fastest part of the now. The overflow warmed to the highest monthly value yet recorded (2.4 degrees C), and showed a pronounced slowing and thinning at its lower margin. We believe that the extreme warmth of the overflow caused it to run higher on the continental slope off east Greenland, so that the lower current meters and the echo sounder were temporarily outside and deeper than the fast-flowing core; model simulations appear to confirm this interpretation, We suggest that the extreme warmth of the overflow is a lagged response to a warming upstream in the Fram Strait three years earlier (caused by an exceptional amplification of the winter North Atlantic Oscillation). If this is so, over-now characteristics may be predictable.

Description: The ability to monitor the heat content of oceans over long distances is becoming increasingly important for understanding the role of oceans in climate change, for determining the variability of the state of the oceans, for operational ocean observing systems, and for studying large-scale ocean processes such as water-mass formation. Although the properties of the upper layers of the ocean can be routinely measured on large scales by satellite remote sensing (providing altimetric and infrared data) and with expendable probes dropped from commercial vessels, the deep interior of the ocean is more difficult to monitor. Ocean acoustic tomography1 is a promising technique for such applications, as it has the potential to provide systematic, instantaneous and repeated measurements of the ocean interior over large parts of an ocean basin. Here we demonstrate the capability of this technique for measuring the heat content across an entire (albeit small) ocean basin—the western Mediterranean Sea.

Description: The conventional model whereby plume volcanism forms linear age-progressive volcanic chains, with the youngest activity occurring nearest a spreading axis (at a 'hotspot'), has been challenged for the Easter seamount chain1–4. Whereas early work suggested the existence of a linear melting anomaly (a 'hotline')1,2, more recent studies3,4 have proposed a hotspot near Salas y Gomez island, connected with the Easter microplate spreading system by an ~800-km-long, volcanically active plume channel. Here we use geochemical, geological and geochronological data to argue that the hotspot lies close to Easter Island. Moreover, new isotopic data for lavas from the seamount chain provide evidence for bidirectional flow between the spreading axis and the plume, thus supporting geophysical and fluid-dynamical models of mantle flow in a plume/spreading axis system5–7. Material balance and flux considerations show the Easter plume to be weak and cool compared with those beneath larger features such as Iceland, Hawaii and the Galápagos islands.

Description: A knowledge of past changes in the biological productivity of the oceans is important for understanding the interactions between carbon cycling and climate. Phytoplankton productivity in today's oceans can be estimated from the concentrations of chlorophyll in sea water1, but chlorophyll is not preserved in the sediments. Existing proxies for past algal productivity do not represent total productivity; for example, biogenic opal2 reflects the contribution of only part of the phytoplankton community, and the organic carbon record can be subject to contamination from terrestrial inputs2,3. Although chlorins, the pigment-transformation products of chlorophyll, are widespread in Quaternary marine sediments, their potential as proxy measures of past variations in primary productivity has not been convincingly demonstrated. Here we report a high-resolution molecular stratigraphic record of chlorin concentrations over the past 350,000 years in a sediment core from the subtropical Atlantic continental margin. Maxima in the chlorin accumulation rate coincide with significant peaks in the accumulation rates of biogenic opal (at the end of glacial terminations) and organic carbon (between terminations). These results suggest that chlorins, unlike other proxies, can serve as a measure of total primary productivity variations.

Description: The ocean stores and transports vast quantities of heat, fresh water, carbon and other materials, and its circulation plays an important role in determining both the Earth's climate and fundamental processes in the biosphere. Understanding the development of climate and important biological cycles therefore requires detailed knowledge of ocean circulation and its transport properties. This cannot be achieved solely through modelling, but must involve accurate observations of the spatio-temporal evolution of the global oceanic flow field. Estimates of oceanic flow are currently made on the basis of space-borne measurements of the sea surface, and monitoring of the ocean interior. Satellite altimetry and acoustic tomography are complementary for this purpose1, as the former provides detailed horizontal coverage of the surface, and the latter the requisite vertical sampling of the interior. High-quality acoustic-tomographic2 and altimetric3 data are now available to test the combined power of these technologies for estimating oceanic flows. Here we demonstrate that, with the aid of state-of-the-art numerical models, it is possible to recover from these data a detailed spatio-temporal record of flow over basin-scale volumes of fluid. Our present results are restricted to the Mediterranean Sea, but the method described here provides a powerful tool for studying oceanic circulation worldwide.

Description: A fundamental issue in marine science is the identification of the factors controlling biological uptake of CO2, in high-nitrate, low-chlorophyll regions. A recent in situ iron fertilization experiment demonstrated that iron limitation is responsible for low phytoplankton stocks in the equatorial Pacific4. Here we show that flavodoxin, a biochemical marker of iron limitation, can be used to map the degree of iron stress in natural populations. Flavodoxin assays along a 900-km east-west transect in the northeastern subarctic Pacific revealed a pronounced increase in iron stress in the region west of the 135° W meridian. Addition of dissolved iron alleviated this stress. Immunostaining of single cells from the most western station showed that flavodoxin is present specifically within the chloroplasts of diatoms. Our approach provides a rapid means of defining the extent of iron stress in the ocean5 and supports the hypothesis that diatoms are iron stressed in the northeast Pacific.

Description: Seismic tomography and the isotope geochemistry of Cenozoic volcanic rocks suggest the existence of a large, sheet-like region of upwelling in the upper mantle which extends from the eastern Atlantic Ocean to central Europe and the western Mediterranean. A belt of extension and rifting in the latter two areas appears to lie above the intersection of the centre of the upwelling region with the base of the lithosphere. Lead, strontium and neodymium isotope data for all three regions converge on a restricted composition, inferred to be that of the upwelling mantle.

Description: THE hydrothermal circulation of sea water through permeable ocean crust results in rock–water interactions that lead to the formation of massive sulphide deposits. These are the modern analogues of many ancient ophiolite-hosted deposits1–4, such as those exposed in Cyprus. Here we report results obtained from drilling a series of holes into an actively forming sulphide deposit on the Mid-Atlantic Ridge. A complex assemblage of sulphide–anhydrite–silica breccias provides striking evidence that such hydrothermal mounds do not grow simply by the accumulation of sulphides on the sea floor. Indeed, the deposit grows largely as an in situ breccia pile, as successive episodes of hydrothermal activity each form new hydrothermal precipitates and cement earlier deposits. During inactive periods, the collapse of sulphide chimneys, dissolution of anhydrite, and disruption by faulting cause brecciation of the deposit. The abundance of anhydrite beneath the present region of focused hydrothermal venting reflects the high temperatures ( 〉 150 °C) currently maintained within the mound, and implies substantial entrainment of cold sea water into the interior of the deposit. These observations demonstrate the important role of anhydrite in the growth of massive sulphide deposits, despite its absence in those preserved on land.

Description: The fully developed flow in a vertical cavity or duct subject to horizontal heating is considered. Solutions of the Boussinesq equations are obtained for rectangular and elliptic sections, in terms of Fourier series and polynomials, respectively. Both generalize the familiar odd-symmetric cubic profile of the plane cavity. Uniqueness is demonstrated under the restriction that the flow is independent of height. For cavities with rectangular sections, it is predicted and verified that the flow in the plane of spanwise symmetry is practically independent of the span if this exceeds 1.7 times the breadth.

Description: An experimental investigation on the initial instability of nonlinear deep-water wave trains including wind effects is reported. The experiment was conducted at the Ocean Engineering Laboratory wind-wave facility (50m long, 4.2m wide, 2.1m deep), with a fully computer-controlled mechanical wave generator to explore the parameter space: steepness; sideband frequency; wind speed. The estimated growth rates of the Benjamin-Feir instability from seeded wind-free experiments agreed well with the theoretical prediction derived from Krasitskii's four-wave reduced equation as computed here. Wind was added to the same wave system ; the growth rates of the sidebands were reduced for weak, and enhanced for strong wind forcing. Experiments with naturally selected sidebands, i.e. unseeded, were conducted as well; measurements showed that wind did not inhibit the growth of sidebands in the case of either two-dimensional or three-dimensional instabilities. A comparison of the results with earlier work suggests that there are two independent effects of wind: first, the alteration of the inviscid growth for a given modulational frequency as shown by comparison with the seeded experiments without wind; second, a change in the natural modulational frequency appearing in the presence of wind which is a function of the wave age, as observed in unseeded experiments. Both effects combined will determine whether the modulational instability is enhanced or suppressed; comparison of experimental results with theoretical predictions suggests that the effect of wind on the natural selection of the modulational frequency is the dominant effect. It was shown that for moderate to old waves, the net effect of wind on the modulational instability is small. For all the experiments except a few unseeded cases with weak breakers, the modulation was small and no breaking was observed within the tank.

Description: A coherent-vortex analysis is made of a computational solution for the free decay of homogeneous, Charney-isotropic geostrophic turbulence at large Reynolds number. The method of analysis is a vortex detection and measurement algorithm that we call a vortex census. The census demonstrates how, through non-conservative interactions among closely approaching vortices, the vortex population evolves towards fewer, larger, sparser, and more weakly deformed vortices. After emergence from random initial conditions and a further period of population adjustment, there is a period of approximately self-similar temporal evolution in the vortex statistics. This behaviour is consistent with a mean-vortex scaling theory based on the conservation of energy, vortex extremum, and vortex aspect ratio. This period terminates as the population approaches a late-time non-turbulent end-state vortex configuration. The end state develops out of merger and alignment interactions among like-sign vortices, and even during the scaling regime, local clusters of nearly aligned vortices are common.

Description: When two drops of radius R touch, surface tension drives an initially singular motion which joins them into a bigger drop with smaller surface area. This motion is always viscously dominated at early times. We focus on the early-time behaviour of the radius rm of the small bridge between the two drops. The flow is driven by a highly curved meniscus of length 2πrm and width Δ ≪ rm around the bridge, from which we conclude that the leading-order problem is asymptotically equivalent to its two-dimensional counterpart. For the case of inviscid surroundings, an exact two-dimensional solution (Hopper 1990) shows that Δ ∝ rm3 and rm ∼ (tγ/πη) ln [tγ/(ηR)]; and thus the same is true in three dimensions. We also study the case of coalescence with an external viscous fluid analytically and, for the case of equal viscosities, in detail numerically. A significantly different structure is found in which the outer-fluid forms a toroidal bubble of radius Δ ∝ rm3/2 at the meniscus and rm ∼ (tγ/4πη) ln [tγ/(ηR)]. This basic difference is due to the presence of the outer-fluid viscosity, however small. With lengths scaled by R a full description of the asymptotic flow for rm(t) ≪ 1 involves matching of lengthscales of order rm2 rm3/2, rm, 1 and probably rm7/4.

Description: In this paper we investigate the linear stability of detonations in which the underlying steady one-dimensional solutions are of the pathological type. Such detonations travel at a minimum speed, which is greater than the Chapman-Jouguet (CJ) speed, have an internal frozen sonic point at which the thermicity vanishes, and the unsupported wave is supersonic (i.e. weak) after the sonic point. Pathological detonations are possible when there are endothermic or dissipative effects present in the system. We consider a system with two consecutive irreversible reactions A→B→C, with an Arrhenius form of the reaction rates and the second reaction endothermic. We determine analytical asymptotic solutions valid near the sonic pathological point for both the one-dimensional steady equations and the equations for linearized perturbations. These are used as initial conditions for integrating the equations. We show that, apart from the existence of stable modes, the linear stability of the pathological detonation is qualitatively the same as for CJ detonations for both one- and two-dimensional disturbances. We also consider the stability of overdriven detonations for the system. We show that the frequency of oscillation for one-dimensional disturbances, and the cell size based on the wavenumber with the highest group velocity for two-dimensional disturbances, are both very sensitive to the detonation speed for overdriven detonations near the pathological speed. This dependence on the degree of overdrive is quite different from that obtained when the unsupported detonation is of the CJ type.

Description: This paper considers the role of long finite-amplitude Rossby waves in determining the evolution of flow along a rapidly rotating channel with an uneven floor. The Rossby waves travel on a potential vorticity interface in a channel with a cross-channel step change in depth, where step position varies slowly along the channel. A nonlinear wave equation is derived describing the evolution of the potential vorticity interface. To leading order this is the hydraulic equation derived by Haynes, Johnson & Hurst (1993). Dispersion appears at the next order. Various solution regimes are identified. As well as slowly varying hydraulic solutions, two further types of steady solutions appear: approach-controlled flows and twin supercritical leaps. Both these solutions are characterized by leaps between supercritical branches of the hydraulic function. It is shown how the position and size of these 'supercritical leaps' can be determined within the context of hydraulic theory. To fully resolve the internal structure of these leaps dispersive effects must be included and leaps are shown to correspond to kink soliton solutions of the steady unforced problem. It is also shown that increasing dispersion (decreasing topographic length scale) causes the loss of the subcritical solution branch in some subcritical flows. The only candidate for a steady solution in these regimes is then an approach-controlled flow. Integrations of initial value problems show that in general flows evolve towards the dispersive form of the solution predicted by hydraulic theory, at least near the topographic perturbation. However, in those subcritical flows where sufficiently large dispersion causes the subcritical branch to disappear, unsteady integrations evolve to approach-controlled flows even when the dispersion is sufficiently small that the subcritical branch still exists.

Description: The transient evolution of the bubble-size probability density functions resulting from the breakup of an air bubble injected into a fully developed turbulent water flow has been measured experimentally using phase Doppler particle sizing (PDPA) and image processing techniques. These measurements were used to determine the breakup frequency of the bubbles as a function of their size and of the critical diameter Dc defined as Dc = 1.26 (σ/ρ)3/5∈-2/5, where ∈ is the rate of dissipation per unit mass and per unit time of the underlying turbulence. A phenomenological model is proposed showing the existence of two distinct bubble size regimes. For bubbles of sizes comparable to Dc, the breakup frequency is shown to increase as (σ/ρ)-2/5 ∈3/5 √D/Dc-1, while for large bubbles whose sizes are greater than 1.63Dc, it decreases with the bubble size as ∈1/3 D-2/3. The model is shown to be in good agreement with measurements performed over a wide range of bubble sizes and turbulence intensities.

Description: Direct numerical simulations of turbulence resulting from Kelvin-Helmholtz instability in stratified shear flow are used to examine the geometry of the dissipation range in a variety of flow regimes. As the buoyancy and shear Reynolds numbers that quantify the degree of isotropy in the dissipation range increase, alignment statistics evolve from those characteristic of parallel shear flow to those found previously in studies of stationary, isotropic, homogeneous turbulence (e.g. Ashurst et al. 1987; She et al. 1991; Tsinober et al. 1992). The analysis yields a limiting value for the mean compression rate of scalar gradients that is expected to be characteristic of all turbulent flows at sufficiently high Reynolds number. My main focus is the value of the constant q that appears in both the Batchelor (1959) and Kraichnan (1968) theoretical forms for the passive scalar spectrum. Taking account of the effects of time-dependent strain, I propose a revised estimate of q, denoted qe, which appears to agree with spectral shapes derived from simulations and observations better than do previous theoretical estimates. The revised estimate is qe = 7.3±0.4, and is expected to be valid whenever the buoyancy Reynolds number exceeds O(102). The Kraichnan (1968) spectral form, in which effects of intermittency are accounted for, provides a better fit to the DNS results than does the Batchelor (1959) form.

Description: A methodology termed the 'filtered mass density function' (FMDF) is developed and implemented for large-eddy simulation (LES) of variable-density chemically reacting turbulent flows at low Mach numbers. This methodology is based on the extension of the 'filtered density function' (FDF) scheme recently proposed by Colucci et al (1998) for LES of constant-density reacting flows. The FMDF represents the joint probability density function of the subgrid-scale (SGS) scalar quantities and is obtained by solution of its modelled transport equation. In this equation, the effect of chemical reactions appears in a closed form and the influences of SGS mixing and convection are modelled. The stochastic differential equations (SDEs) which yield statistically equivalent results to those of the FMDF transport equation are derived and are solved via a Lagrangian Monte Carlo scheme. The consistency, convergence, and accuracy of the FMDF and the Monte Carlo solution of its equivalent SDEs are assessed. In non-reacting flows, it is shown that the filtered results via the FMDF agree well with those obtained by the 'conventional' LES in which the finite difference solution of the transport equations of these filtered quantities is obtained. The advantage of the FMDF is demonstrated in LES of reacting shear flows with non-premixed reactants. The FMDF results are appraised by comparisons with data generated by direct numerical simulation (DNS) and with experimental measurements. In the absence of a closure for the SGS scalar correlations, the results based on the conventional LES are significantly different from those obtained by DNS. The FMDF results show a closer agreement with DNS. These results also agree favourably with laboratory data of exothermic reacting turbulent shear flows, and portray several of the features observed experimentally.

Description: The objective of this study is to develop a method of controlling vortex shedding behind a bluff body using control theory. A suboptimal feedback control procedure for local sensing and local actuation is developed and applied to the flow behind a circular cylinder. The location of sensors for feedback is limited to the cylinder surface and the control input from actuators is the blowing and suction on the cylinder surface. Three different cost functionals to be minimized (J1 and J2) or maximized (J3) are investigated: J1 is proportional to the pressure drag of the cylinder, J2 is the square of the difference between the target pressure (inviscid flow pressure) and real flow pressure on the cylinder surface, and J3 is the square of the pressure gradient on the cylinder surface, respectively. Given the cost functionals, the flow variable to be measured by the sensors and the control input from the actuators are determined from the suboptimal feedback control procedure. Several cases for each cost functional have been numerically simulated at Re = 100 and 160 to investigate the performance of the control algorithm. For all actuations, vortex shedding becomes weak or disappears, and the mean drag and drag/lift fluctuations significantly decrease. For a given magnitude of the blowing/suction, reducing J2 provides the largest drag reduction among the three cost functionals.

Description: Using a combination of bifurcation theory for two-dimensional dynamical systems and numerical simulations, we systematically determine the possible flow topologies of the steady vortex breakdown in axisymmetric flow in a cylindrical container with rotating end-covers. For fixed values of the ratio of the angular velocities of the covers in the range from -0.02 to 0.05, bifurcations of recirculating bubbles under variation of the aspect ratio of the cylinder and the Reynolds number are found. Bifurcation curves are determined by a simple fitting procedure of the data from the simulations. For the much studied case of zero rotation ratio (one fixed cover) a complete bifurcation diagram is constructed. Very good agreement with experimental results is obtained, and hitherto unresolved details are determined in the parameter region where up to three bubbles exist. For non-zero rotation ratios the bifurcation diagrams are found to change dramatically and give rise to other types of bifurcations.

Description: The high Reynolds number flow through a circular pipe divided along a diameter by a semi-infinite splitter plate is considered. Matched asymptotic expansions are used to analyse the developing flow, which is decomposed into four regions: a boundary layer of Blasius type growing along the plate, an inviscid core, a viscous layer close to the curved wall and a nonlinear corner region. The core solution is found numerically, initially in the long-distance down-pipe limit and thereafter the full problem is solved using down-pipe Fourier transforms. The accuracy in the corners of the semicircular cross-section is improved by subtracting out the singularity in the velocity perturbation. The linear viscous wall layer is solved analytically in terms of a displacement function determined from the core. A plausible structure for the corner region and equations governing the motion therein are presented although no solution is attempted. The presence of the plate has little effect ahead of the bifurcation, but wall shear on the curved wall is found to increase from its undisturbed value downstream.

Description: The behaviour of the long-time self-diffusion tensor in concentrated colloidal dispersions is studied using dynamic simulation. The simulations are of a suspension of monodisperse Brownian hard spheres in simple shear flow as a function of the Péclet number, Pe, which measures the relative importance of shear and Brownian forces, and the volume fraction, φ. Here, Pe = γ̇a2/D0, where γ̇ is the shear rate, a the particle size and D0 = kT/6πηa is the Stokes-Einstein diffusivity of an isolated particle of size a with thermal energy kT in a solvent of viscosity η. Two simulations algorithms are used: Stokesian Dynamics for inclusion of the many-body hydrodynamic interactions, and Brownian Dynamics for suspensions without hydrodynamic interactions. A new procedure for obtaining high-quality diffusion data based on averaging the results of many short simulations is presented and utilized. At low shear rates, low Pe, Brownian diffusion due to a random walk process dominates and the characteristic scale for diffusion is the Stokes-Einstein diffusivity, D0. At zero Pe the diffusivity is found to be a decreasing function of φ. As Pe is slowly increased, 0(Pe) and 0(Pe3/2) corrections to the diffusivity due to the flow are clearly seen in the Brownian Dynamics system in agreement with the theoretical results of Morris & Brady (1996). At large shear rates, large Pe, both systems exhibit diffusivities that grow linearly with the shear rate by the non-Brownian mechanism of shear-induced diffusion. In contrast to the behaviour at low Pe, this shear-induced diffusion mode is an increasing function of φ. Long-time rotational self-diffusivities are of interest in the Stokesian Dynamics system and show similar behaviour to their translational analogues. An off-diagonal long-time self-diffusivity, Dxy, is reported for both systems. Results for both the translational and rotational Dxy. show a sign change from low Pe to high Pe due to different mechanisms in the two regimes. A physical explanation for the off-diagonal diffusivities is proposed.

Description: Based on energy principles, we propose a statistical model to describe the bubble size probability density function of the daughter bubbles resulting from the shattering of a mother bubble of size D0 immersed in a fully developed turbulent water flow. The model shows that the bubble size p.d.f. depends not only on D0, but also on the value of the dissipation rate of turbulent kinetic energy of the underlying turbulence of the water, ∈. The phenomenological model is simple, yet it predicts detailed experimental measurements of the transient bubble size p.d.f.s performed over a range of bubble sizes and dissipation rates ∈ in a very consistent manner. The agreement between the model and the experiments is particularly good for low and moderate bubble turbulent Weber numbers, Wet = ρ Δu2(D0)D0/σ where the assumption of the binary breakup is shown to be consistent with the experimental observations. At larger values of Wet, it was found that the most probable number of daughter bubbles increases and the assumption of tertiary breakup is shown to lead to a better fit of the experimental measurements.

Description: Using the continuum model of Pedley, Hill & Kessler (1988) for bioconvection in a suspension of swimming, gyrotactic micro-organisms, we investigate the existence and stability of a two-dimensional plume in tall, narrow chambers with stress-free sidewalls. The system is governed by the Navier-Stokes equations for an incompressible fluid coupled with a micro-organism conservation equation. These equations are solved numerically using a conservative finite-difference scheme. In sufficiently deep chambers, the plume is always unstable to both varicose and meandering modes. A linear stability analysis for an infinitely long plume predicts the growth rates of these instabilities, explains the mechanisms, and is in good agreement with the numerical results.

Description: A three-dimensional mathematical model based on the Brinkman extended Darcy equation has been used to study double-diffusive natural convection in a fluidsaturated porous cubic enclosure subject to opposing and horizontal gradients of temperature and concentration. The flow is driven by conditions of constant temperature and concentration imposed along the two vertical sidewalls of the cubic enclosure, while the remaining walls are impermeable and adiabatic. The numerical simulations presented here span a wide range of porous thermal Rayleigh number, buoyancy ratio and Lewis number to identify the different steady-state flow patterns and bifurcations. The effect of the governing parameters on the domain of existence of the three-dimensional flow patterns is studied for opposing flows (N 〈 0). Comprehensive Nusselt and Sherwood number data are presented as functions of the governing parameters. The present results indicate that the double-diffusive flow in enclosures with opposing buoyancy forces is strictly three-dimensional for a certain range of parameters. At high Lewis numbers multiple dipole vortices form in the transverse planes near the horizontal top and bottom surfaces, which the two-dimensional models fail to detect. The dipolar vortex structures obtained are similar to those created in laboratory experiments by the injection of fluid into a stratified medium.

Description: The influence of inertia and elasticity on the onset and stability of Taylor-vortex flow (TVF) is examined for an Oldroyd-B fluid. The Galerkin projection method is used to obtain the departure from Couette flow (CF). Only axisymmetric flow is examined. The solution is capable of capturing the dynamical behaviour observed experimentally for viscoelastic fluids in the inertio-elastic and purely elastic ranges. For flow with dominant inertia, the bifurcation picture is similar to that for a Newtonian fluid. However, transition from CF to TVF is oscillatory because of fluid elasticity. Steady TVF sets in, via supercritical bifurcation, as Re reaches a critical value, Rec. The critical Reynolds number decreases with fluid elasticity, and is strongly influenced by fluid retardation. As elasticity exceeds a critical level, a subcritical bifurcation emerges at Rec, similar to that predicted by the Landau-Ginzburg equation. It is found that slip along the axial direction tends to be generally destabilizing. The coherence of the formulation is established under steady and transient conditions through comparison with exact linear stability analysis, experimental measurements, and flow visualization. Good agreement is obtained between theory and the measurements of Muller et al. (1993) in the limit of purely elastic overstable TVF.

Description: We present simulation results of vortex-induced vibrations of an infinitely long flexible cylinder at Reynolds number Re = 1000, corresponding to a 'young' turbulent wake (i.e. exhibiting a small inertial subrange). The simulations are based on a new class of spectral methods suitable for unstructured and hybrid grids. To obtain different responses of the coupled flow-structure system we vary the structure's bending stiffness to model the behaviour of a vibrating inflexible (rigid) cylinder, a cable, and a beam. We have found that unlike the laminar flow previously studied, the amplitude of the cross-flow oscillation is about one diameter for the cable and the beam, close to experimental measurements, but is lower for the rigid cylinder. We have also found that for the latter case the flow response corresponds to parallel shedding, but for the beam and cable with free endpoints a mixed response consisting of oblique and parallel shedding is obtained, caused by the modulated travelling wave motion of the structure. This mixed shedding pattern which alternates periodically along the span can be directly related to periodic spatial variation of the lift force. In the case of structures with pinned endpoints a standing wave response is obtained for the cylinder; lace-like flow structures are observed similar to the ones seen in the laminar regime. Examination of the frequency spectra in the near wake shows that at Re = 1000 all cases follow a -5/3 law in the inertial range, which extends about half a decade in wavenumber. However, these spectra are different in all three cases both in low and high frequencies, with the exception of the beam and cable, for which the high-frequency portion is identical despite the differences in the displacement time history and the large-scale features of the corresponding flow.

Description: The incompressible boundary layer in the corner formed by two intersecting, semi-infinite planes is investigated, when the free-stream flow, aligned with the corner, is taken to be of the form U∞F(x), x representing the non-dimensional streamwise distance from the leading edge. In Dhanak & Duck (1997) similarity solutions for F(x) = xn were considered, and it was found that solutions exist for only a range of values of n, whilst for ∞ 〉 n 〉 -0.018, approximately, two solutions exist. In this paper, we extend the work of Dhanak & Duck to the case of non-90° corner angles and allow for streamwise development of solutions. In addition, the effect of transpiration at the walls of the corner is investigated. The governing equations are of boundary-layer type and as such are parabolic in nature. Crucially, although the leading-order pressure term is known a priori, the third-order pressure term is not, but this is nonetheless present in the leading-order governing equations, together with the transverse and crossflow viscous terms. Particular attention is paid to flows which develop spatially from similarity solutions. It turns out that two scenarios are possible. In some cases the problem may be treated in the usual parabolic sense, with standard numerical marching procedures being entirely appropriate. In other cases standard marching procedures lead to numerically inconsistent solutions. The source of this difficulty is linked to the existence of eigensolutions emanating from the leading edge (which are not present in flows appropriate to the first scenario), analogous to those found in the computation of some two-dimensional hypersonic boundary layers (Neiland 1970; Mikhailov et al. 1971; Brown & Stewartson 1975). In order to circumvent this difficulty, a different numerical solution strategy is adopted, based on a global Newton iteration procedure. A number of numerical solutions for the entire corner flow region are presented.

Description: A generalized model of flow in meandering subaqueous and subaerial channels is developed. The conservation equations of mass and momentum are depth/layer integrated, normalized, and represented as deviations from a straight base state. This allows the determination of integrable forms which can be solved at both linear and nonlinear levels. The effects of various flow and geometric parameters on the flow dynamics are studied. Although the model is not limited to any specific planform, this study focuses on sine-generated curves. In analysing the flow patterns, the turbidity current of the subaqueous case is simplified to a conservative density flow with water entrainment from above neglected. The subaqueous model thus formally corresponds to a subcritical or only mildly supercritical mud-rich turbidity current. By extension, however the analysis can be applied to a depositional or erosional current carrying sand that is changing only slowly in the streamwise direction. By bringing the subaqueous and subaerial cases into a common form, flow behaviour in the two environments can be compared under similar geometric and boundary conditions. A major difference between the two cases is the degree of superelevation of channel flow around bends, which is modest in the subaerial case but substantial in the subaqueous case. Another difference concerns Coriolis effects: some of the largest subaqueous meandering systems are so large that Coriolis effects can become important. The model is applied to meander bends on the youngest channel in the mid-fan region of the Amazon Fan and a mildly sinuous bend of the North-West Atlantic Mid-Ocean Channel. In the absence of specific data on the turbid flows that created the channel, the model can be used to make inferences about the flow, and in particular the range of values of flow velocity and sediment concentration that would allow the growth and downfan migration of meander bends.

Description: The collisional flow of a slightly inelastic granular material down a rough inclined plane is usually described by kinetic theories. We present an experimental study aimed at analysing the assumptions and the quantitative predictions of such theories. A two-dimensional channel coupled to a model granular material and image analysis allow detailed and complete measurement of the kinematics and structure of the flows. We determine the range of inclination and particle flux for which the flow is stationary and uniform. The characteristic profiles of solid fraction, mean velocity and granular temperature are systematically measured. Both the true collisional and the dilute kinetic regimes are examined. We show that a quasi-hydrodynamic description of these regimes seems relevant, and that the pressure and the viscosity terms are in good qualitative agreement with the prediction of the kinetic theory. The profiles are well described by the kinetic theory near the top of the flow, at low solid fraction. Conversely there are large discrepancies near the rough plane, where the material is structured in layers.