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  • growth  (63)
  • Eddies
  • Turbulence
  • Springer  (73)
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  • 1
    Publication Date: 2022-10-12
    Description: Author Posting. © American Meteorological Society, 2022. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 52(10), (2022): 2325–2341, https://doi.org/10.1175/jpo-d-21-0015.1.
    Description: The ocean surface boundary layer is a gateway of energy transfer into the ocean. Wind-driven shear and meteorologically forced convection inject turbulent kinetic energy into the surface boundary layer, mixing the upper ocean and transforming its density structure. In the absence of direct observations or the capability to resolve subgrid-scale 3D turbulence in operational ocean models, the oceanography community relies on surface boundary layer similarity scalings (BLS) of shear and convective turbulence to represent this mixing. Despite their importance, near-surface mixing processes (and ubiquitous BLS representations of these processes) have been undersampled in high-energy forcing regimes such as the Southern Ocean. With the maturing of autonomous sampling platforms, there is now an opportunity to collect high-resolution spatial and temporal measurements in the full range of forcing conditions. Here, we characterize near-surface turbulence under strong wind forcing using the first long-duration glider microstructure survey of the Southern Ocean. We leverage these data to show that the measured turbulence is significantly higher than standard shear-convective BLS in the shallower parts of the surface boundary layer and lower than standard shear-convective BLS in the deeper parts of the surface boundary layer; the latter of which is not easily explained by present wave-effect literature. Consistent with the CBLAST (Coupled Boundary Layers and Air Sea Transfer) low winds experiment, this bias has the largest magnitude and spread in the shallowest 10% of the actively mixing layer under low-wind and breaking wave conditions, when relatively low levels of turbulent kinetic energy (TKE) in surface regime are easily biased by wave events.
    Description: This paper is VIMS Contribution 4103. Computational resources were provided by the VIMS Ocean-Atmosphere and Climate Change Research Fund. AUSSOM was supported by the OCE Division of the National Science Foundation (1558639).
    Keywords: Turbulence ; Wind shear ; Boundary layer ; Parameterization
    Repository Name: Woods Hole Open Access Server
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  • 2
    Publication Date: 2022-08-29
    Description: Author Posting. © American Meteorological Society, 2022. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 52(8), (2022): 1593-1611, https://doi.org/10.1175/jpo-d-21-0180.1.
    Description: This study presents novel observational estimates of turbulent dissipation and mixing in a standing meander between the Southeast Indian Ridge and the Macquarie Ridge in the Southern Ocean. By applying a finescale parameterization on the temperature, salinity, and velocity profiles collected from Electromagnetic Autonomous Profiling Explorer (EM-APEX) floats in the upper 1600 m, we estimated the intensity and spatial distribution of dissipation rate and diapycnal mixing along the float tracks and investigated the sources. The indirect estimates indicate strong spatial and temporal variability of turbulent mixing varying from O(10−6) to O(10−3) m2 s−1 in the upper 1600 m. Elevated turbulent mixing is mostly associated with the Subantarctic Front (SAF) and mesoscale eddies. In the upper 500 m, enhanced mixing is associated with downward-propagating wind-generated near-inertial waves as well as the interaction between cyclonic eddies and upward-propagating internal waves. In the study region, the local topography does not play a role in turbulent mixing in the upper part of the water column, which has similar values in profiles over rough and smooth topography. However, both remotely generated internal tides and lee waves could contribute to the upward-propagating energy. Our results point strongly to the generation of turbulent mixing through the interaction of internal waves and the intense mesoscale eddy field.
    Description: The observations were funded through grants from the Australian Research Council Discovery Project (DP170102162) and Australia’s Marine National Facility. Surface drifters were provided by Dr. Shaun Dolk of the Global Drifter Program. AC was supported by an Australian Research Council Postdoctoral Fellowship. AC, HEP, and NLB acknowledge support from the Australian Government Department of the Environment and Energy National Environmental Science Program and the ARC Centre of Excellence in Climate Extremes. KP acknowledges the support from the National Science Foundation.
    Keywords: Diapycnal mixing ; Eddies ; Fronts ; Inertia-gravity waves ; Ocean dynamics
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  • 3
    Publication Date: 2022-09-15
    Description: Author Posting. © American Meteorological Society, 2022. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 52(1),(2022): 75–97, https://doi.org/10.1175/JPO-D-21-0099.1.
    Description: Mesoscale eddies contain the bulk of the ocean’s kinetic energy (KE), but fundamental questions remain on the cross-scale KE transfers linking eddy generation and dissipation. The role of submesoscale flows represents the key point of discussion, with contrasting views of submesoscales as either a source or a sink of mesoscale KE. Here, the first observational assessment of the annual cycle of the KE transfer between mesoscale and submesoscale motions is performed in the upper layers of a typical open-ocean region. Although these diagnostics have marginal statistical significance and should be regarded cautiously, they are physically plausible and can provide a valuable benchmark for model evaluation. The cross-scale KE transfer exhibits two distinct stages, whereby submesoscales energize mesoscales in winter and drain mesoscales in spring. Despite this seasonal reversal, an inverse KE cascade operates throughout the year across much of the mesoscale range. Our results are not incompatible with recent modeling investigations that place the headwaters of the inverse KE cascade at the submesoscale, and that rationalize the seasonality of mesoscale KE as an inverse cascade-mediated response to the generation of submesoscales in winter. However, our findings may challenge those investigations by suggesting that, in spring, a downscale KE transfer could dampen the inverse KE cascade. An exploratory appraisal of the dynamics governing mesoscale–submesoscale KE exchanges suggests that the upscale KE transfer in winter is underpinned by mixed layer baroclinic instabilities, and that the downscale KE transfer in spring is associated with frontogenesis. Current submesoscale-permitting ocean models may substantially understate this downscale KE transfer, due to the models’ muted representation of frontogenesis.
    Description: The OSMOSIS experiment was funded by the U.K. Natural Environment Research Council (NERC) through Grants NE/1019999/1 and NE/101993X/1. ACNG acknowledges the support of the Royal Society and the Wolfson Foundation, and XY that of a China Scholarship Council PhD studentship.
    Keywords: Ageostrophic circulations ; Dynamics ; Eddies ; Energy transport ; Frontogenesis/frontolysis ; Instability ; Mesoscale processes ; Nonlinear dynamics ; Ocean circulation ; Ocean dynamics ; Small scale processes ; Turbulence
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  • 4
    Publication Date: 2022-09-01
    Description: Author Posting. © American Meteorological Society, 2022. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 52(8), (2022): 1677-1691, https://doi.org/10.1175/jpo-d-21-0269.1.
    Description: Oceanic mesoscale motions including eddies, meanders, fronts, and filaments comprise a dominant fraction of oceanic kinetic energy and contribute to the redistribution of tracers in the ocean such as heat, salt, and nutrients. This reservoir of mesoscale energy is regulated by the conversion of potential energy and transfers of kinetic energy across spatial scales. Whether and under what circumstances mesoscale turbulence precipitates forward or inverse cascades, and the rates of these cascades, remain difficult to directly observe and quantify despite their impacts on physical and biological processes. Here we use global observations to investigate the seasonality of surface kinetic energy and upper-ocean potential energy. We apply spatial filters to along-track satellite measurements of sea surface height to diagnose surface eddy kinetic energy across 60–300-km scales. A geographic and scale-dependent seasonal cycle appears throughout much of the midlatitudes, with eddy kinetic energy at scales less than 60 km peaking 1–4 months before that at 60–300-km scales. Spatial patterns in this lag align with geographic regions where an Argo-derived estimate of the conversion of potential to kinetic energy is seasonally varying. In midlatitudes, the conversion rate peaks 0–2 months prior to kinetic energy at scales less than 60 km. The consistent geographic patterns between the seasonality of potential energy conversion and kinetic energy across spatial scale provide observational evidence for the inverse cascade and demonstrate that some component of it is seasonally modulated. Implications for mesoscale parameterizations and numerical modeling are discussed.
    Description: This work was generously funded by NSF Grants OCE-1912302, OCE-1912125 (Drushka), and OCE-1912325 (Abernathey) as part of the Ocean Energy and Eddy Transport Climate Process Team.
    Keywords: Eddies ; Energy transport ; Mesoscale processes ; Turbulence ; Oceanic mixed layer ; Altimetry ; Seasonal cycle
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  • 5
    Publication Date: 2022-06-10
    Description: Author Posting. © American Meteorological Society , 2021. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Zippel, S. F., Farrar, J. T., Zappa, C. J., Miller, U., St Laurent, L., Ijichi, T., Weller, R. A., McRaven, L., Nylund, S., & Le Bel, D. Moored turbulence measurements using pulse-coherent doppler sonar. Journal of Atmospheric and Oceanic Technology, 38(9), (2021): 1621–1639, https://doi.org/10.1175/JTECH-D-21-0005.1.
    Description: Upper-ocean turbulence is central to the exchanges of heat, momentum, and gases across the air–sea interface and therefore plays a large role in weather and climate. Current understanding of upper-ocean mixing is lacking, often leading models to misrepresent mixed layer depths and sea surface temperature. In part, progress has been limited by the difficulty of measuring turbulence from fixed moorings that can simultaneously measure surface fluxes and upper-ocean stratification over long time periods. Here we introduce a direct wavenumber method for measuring turbulent kinetic energy (TKE) dissipation rates ϵ from long-enduring moorings using pulse-coherent ADCPs. We discuss optimal programming of the ADCPs, a robust mechanical design for use on a mooring to maximize data return, and data processing techniques including phase-ambiguity unwrapping, spectral analysis, and a correction for instrument response. The method was used in the Salinity Processes Upper-Ocean Regional Study (SPURS) to collect two year-long datasets. We find that the mooring-derived TKE dissipation rates compare favorably to estimates made nearby from a microstructure shear probe mounted to a glider during its two separate 2-week missions for O(10−8) ≤ ϵ ≤ O(10−5) m2 s−3. Periods of disagreement between turbulence estimates from the two platforms coincide with differences in vertical temperature profiles, which may indicate that barrier layers can substantially modulate upper-ocean turbulence over horizontal scales of 1–10 km. We also find that dissipation estimates from two different moorings at 12.5 and at 7 m are in agreement with the surface buoyancy flux during periods of strong nighttime convection, consistent with classic boundary layer theory.
    Description: This work was funded by NASA as part of the Salinity Processes in the Upper Ocean Regional Study (SPURS), supporting field work for SPURS-1 (NASA Grant NNX11AE84G), for SPURS-2 (NASA Grant NNX15AG20G), and for analysis (NASA Grant 80NSSC18K1494). Funding for early iterations of this project associated with the VOCALS project and Stratus 9 mooring was provided by NSF (Awards 0745508 and 0745442). Additional funding was provided by ONR Grant N000141812431 and NSF Award 1756839. The Stratus Ocean Reference Station is funded by the Global Ocean Monitoring and Observing Program of the National Oceanic and Atmospheric Administration (CPO FundRef Number 100007298), through the Cooperative Institute for the North Atlantic Region (CINAR) under Cooperative Agreement NA14OAR4320158. Microstructure measurements made from the glider were supported by NSF (Award 1129646).
    Keywords: Ocean ; Turbulence ; Atmosphere-ocean interaction ; Boundary layer ; Oceanic mixed layer ; In situ oceanic observations
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  • 6
    Publication Date: 2022-06-13
    Description: © The Author(s), 2022. This article is distributed under the terms of the Creative Commons Attribution License. The definitive version was published in Fine, E., MacKinnon, J., Alford, M., Middleton, L., Taylor, J., Mickett, J., Cole, S., Couto, N., Boyer, A., & Peacock, T. Double diffusion, shear instabilities, and heat impacts of a pacific summer water intrusion in the Beaufort Sea. Journal of Physical Oceanography, 52(2), (2022): 189–203, https://doi.org/10.1175/jpo-d-21-0074.1.
    Description: Pacific Summer Water eddies and intrusions transport heat and salt from boundary regions into the western Arctic basin. Here we examine concurrent effects of lateral stirring and vertical mixing using microstructure data collected within a Pacific Summer Water intrusion with a length scale of ∼20 km. This intrusion was characterized by complex thermohaline structure in which warm Pacific Summer Water interleaved in alternating layers of O(1) m thickness with cooler water, due to lateral stirring and intrusive processes. Along interfaces between warm/salty and cold/freshwater masses, the density ratio was favorable to double-diffusive processes. The rate of dissipation of turbulent kinetic energy (ε) was elevated along the interleaving surfaces, with values up to 3 × 10−8 W kg−1 compared to background ε of less than 10−9 W kg−1. Based on the distribution of ε as a function of density ratio Rρ, we conclude that double-diffusive convection is largely responsible for the elevated ε observed over the survey. The lateral processes that created the layered thermohaline structure resulted in vertical thermohaline gradients susceptible to double-diffusive convection, resulting in upward vertical heat fluxes. Bulk vertical heat fluxes above the intrusion are estimated in the range of 0.2–1 W m−2, with the localized flux above the uppermost warm layer elevated to 2–10 W m−2. Lateral fluxes are much larger, estimated between 1000 and 5000 W m−2, and set an overall decay rate for the intrusion of 1–5 years.
    Description: This work was supported by ONR Grant N00014-16-1-2378 and NSF Grants PLR 14-56705 and PLR-1303791, NSF Graduate Research Fellowship Grant DGE-1650112, as well as by the Postdoctoral Scholar Program at Woods Hole Oceanographic Institution, with funding provided by the Weston Howland Jr. Postdoctoral Scholarship.
    Keywords: Arctic ; Diapycnal mixing ; Diffusion ; Fluxes ; Instability ; Mixing ; Turbulence
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  • 7
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    American Meteorological Society
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(9), (2020): 2491-2506, doi:10.1175/JPO-D-20-0056.1.
    Description: An idealized two-layer shallow water model is applied to the study of the dynamics of the Arctic Ocean halocline. The model is forced by a surface stress distribution reflective of the observed wind stress pattern and ice motion and by an inflow representing the flow of Pacific Water through Bering Strait. The model reproduces the main elements of the halocline circulation: an anticyclonic Beaufort Gyre in the western basin (representing the Canada Basin), a cyclonic circulation in the eastern basin (representing the Eurasian Basin), and a Transpolar Drift between the two gyres directed from the upwind side of the basin to the downwind side of the basin. Analysis of the potential vorticity budget shows a basin-averaged balance primarily between potential vorticity input at the surface and dissipation at the lateral boundaries. However, advection is a leading-order term not only within the anticyclonic and cyclonic gyres but also between the gyres. This means that the eastern and western basins are dynamically connected through the advection of potential vorticity. Both eddy and mean fluxes play a role in connecting the regions of potential vorticity input at the surface with the opposite gyre and with the viscous boundary layers. These conclusions are based on a series of model runs in which forcing, topography, straits, and the Coriolis parameter were varied.
    Description: This study was supported by National Science Foundation Grant OPP-1822334. Comments and suggestions from two anonymous referees greatly helped to improve the paper.
    Description: 2021-02-17
    Keywords: Eddies ; Ekman pumping/transport ; Ocean circulation ; Ocean dynamics ; Potential vorticity ; Shallow-water equations
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  • 8
    Publication Date: 2022-05-27
    Description: Author Posting. © American Meteorological Society, 2021. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 51(1), (2021): 19-35, https://doi.org/10.1175/JPO-D-19-0233.1.
    Description: In the Beaufort Sea in September of 2015, concurrent mooring and microstructure observations were used to assess dissipation rates in the vicinity of 72°35′N, 145°1′W. Microstructure measurements from a free-falling profiler survey showed very low [O(10−10) W kg−1] turbulent kinetic energy dissipation rates ε. A finescale parameterization based on both shear and strain measurements was applied to estimate the ratio of shear to strain Rω and ε at the mooring location, and a strain-based parameterization was applied to the microstructure survey (which occurred approximately 100 km away from the mooring site) for direct comparison with microstructure results. The finescale parameterization worked well, with discrepancies ranging from a factor of 1–2.5 depending on depth. The largest discrepancies occurred at depths with high shear. Mean Rω was 17, and Rω showed high variability with values ranging from 3 to 50 over 8 days. Observed ε was slightly elevated (factor of 2–3 compared with a later survey of 11 profiles taken over 3 h) from 25 to 125 m following a wind event which occurred at the beginning of the mooring deployment, reaching a maximum of ε= 6 × 10−10 W kg−1 at 30-m depth. Velocity signals associated with near-inertial waves (NIWs) were observed at depths greater than 200 m, where the Atlantic Water mass represents a reservoir of oceanic heat. However, no evidence of elevated ε or heat fluxes was observed in association with NIWs at these depths in either the microstructure survey or the finescale parameterization estimates.
    Description: This work was supported by NSF Grants PLR 14-56705 and PLR-1303791 and by NSF Graduate Research Fellowship Grant DGE-1650112.
    Keywords: Ocean ; Arctic ; Internal waves ; Turbulence ; Diapycnal mixing
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  • 9
    Publication Date: 2022-05-27
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(11), (2020): 3267–3294, https://doi.org/10.1175/JPO-D-19-0310.1.
    Description: As part of the Flow Encountering Abrupt Topography (FLEAT) program, an array of pressure-sensor equipped inverted echo sounders (PIESs) was deployed north of Palau where the westward-flowing North Equatorial Current encounters the southern end of the Kyushu–Palau Ridge in the tropical North Pacific. Capitalizing on concurrent observations from satellite altimetry, FLEAT Spray gliders, and shipboard hydrography, the PIESs’ 10-month duration hourly bottom pressure p and round-trip acoustic travel time τ records are used to examine the magnitude and predictability of sea level and pycnocline depth changes and to track signal propagations through the array. Sea level and pycnocline depth are found to vary in response to a range of ocean processes, with their magnitude and predictability strongly process dependent. Signals characterized here comprise the barotropic tides, semidiurnal and diurnal internal tides, southeastward-propagating superinertial waves, westward-propagating mesoscale eddies, and a strong signature of sea level increase and pycnocline deepening associated with the region’s relaxation from El Niño to La Niña conditions. The presence of a broad band of superinertial waves just above the inertial frequency was unexpected and the FLEAT observations and output from a numerical model suggest that these waves detected near Palau are forced by remote winds east of the Philippines. The PIES-based estimates of pycnocline displacement are found to have large uncertainties relative to overall variability in pycnocline depth, as localized deep current variations arising from interactions of the large-scale currents with the abrupt topography around Palau have significant travel time variability.
    Description: Support for this research was provided by Office of Naval Research Grants N00014-16-1-2668, N00014-18-1-2406, N00014-15-1-2488, and N00014-15-1-2622. R.C.M. was additionally supported by the Postdoctoral Scholar Program at the Woods Hole Oceanographic Institution, with funding provided by the Weston Howland Jr. Postdoctoral Scholarship.
    Keywords: Tropics ; Currents ; Eddies ; ENSO ; Internal waves ; Mesoscale processes
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  • 10
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    American Meteorological Society
    Publication Date: 2022-05-27
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(11),(2020): 3331–3351, https://doi.org/10.1175/JPO-D-20-0035.1.
    Description: This study examines the generation of warm spiral structures (referred to as spiral streamers here) over Gulf Stream warm-core rings. Satellite sea surface temperature imagery shows spiral streamers forming after warmer water from the Gulf Stream or newly formed warm-core rings impinges onto old warm-core rings and then intrudes into the old rings. Field measurements in April 2018 capture the vertical structure of a warm spiral streamer as a shallow lens of low-density water winding over an old ring. Observations also show subduction on both sides of the spiral streamer, which carries surface waters downward. Idealized numerical model simulations initialized with observed water-mass densities reproduce spiral streamers over warm-core rings and reveal that their formation is a nonlinear submesoscale process forced by mesoscale dynamics. The negative density anomaly of the intruding water causes a density front at the interface between the intruding water and surface ring water, which, through thermal wind balance, drives a local anticyclonic flow. The pressure gradient and momentum advection of the local interfacial flow push the intruding water toward the ring center. The large-scale anticyclonic flow of the ring and the radial motion of the intruding water together form the spiral streamer. The observed subduction on both sides of the spiral streamer is part of the secondary cross-streamer circulation resulting from frontogenesis on the stretching streamer edges. The surface divergence of the secondary circulation pushes the side edges of the streamer away from each other, widens the warm spiral on the surface, and thus enhances its surface signal.
    Description: Authors W. G. Zhang and D. J. McGillicuddy are both supported by the National Science Foundation through Grant OCE 1657803.
    Keywords: Buoyancy ; Eddies ; Frontogenesis/frontolysis ; Mesoscale processes ; Transport ; Vertical motion
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  • 11
    Publication Date: 2022-05-27
    Description: © The Author(s), 2021. This article is distributed under the terms of the Creative Commons Attribution License. The definitive version was published in Spingys, C. P., Garabato, A. C. N., Legg, S., Polzin, K. L., Abrahamsen, E. P., Buckingham, C. E., Forryan, A., & Frajka-Williams, E. E. Mixing and transformation in a deep western boundary current: a case study. Journal of Physical Oceanography, 51(4), (2021): 1205-1222, https://doi.org/10.1175/JPO-D-20-0132.1
    Description: Water-mass transformation by turbulent mixing is a key part of the deep-ocean overturning, as it drives the upwelling of dense waters formed at high latitudes. Here, we quantify this transformation and its underpinning processes in a small Southern Ocean basin: the Orkney Deep. Observations reveal a focusing of the transport in density space as a deep western boundary current (DWBC) flows through the region, associated with lightening and densification of the current’s denser and lighter layers, respectively. These transformations are driven by vigorous turbulent mixing. Comparing this transformation with measurements of the rate of turbulent kinetic energy dissipation indicates that, within the DWBC, turbulence operates with a high mixing efficiency, characterized by a dissipation ratio of 0.6 to 1 that exceeds the common value of 0.2. This result is corroborated by estimates of the dissipation ratio from microstructure observations. The causes of the transformation are unraveled through a decomposition into contributions dependent on the gradients in density space of the: dianeutral mixing rate, isoneutral area, and stratification. The transformation is found to be primarily driven by strong turbulence acting on an abrupt transition from the weakly stratified bottom boundary layer to well-stratified off-boundary waters. The reduced boundary layer stratification is generated by a downslope Ekman flow associated with the DWBC’s flow along sloping topography, and is further regulated by submesoscale instabilities acting to restratify near-boundary waters. Our results provide observational evidence endorsing the importance of near-boundary mixing processes to deep-ocean overturning, and highlight the role of DWBCs as hot spots of dianeutral upwelling.
    Description: CS, ACNG, AF, and EFW were supported by the U.K. Natural Environment Research Council (NERC) Grant NE/K013181/1. ACNG was supported by the Royal Society and Wolfson Foundation. EPA and CEB were supported by NERC Grant NE/K012843/1. CEB was funded by an MSCA grant (No. 798319) from the European Union’s Horizon 2020 program. EPA was supported by NERC Grant NE/N018095/1. SL and KP were supported by U.S. National Science Foundation Grants OCE-1536453 and OCE-1536779. SL acknowledges support of Award NA18OAR4320123 from the National Oceanic and Atmospheric Administration, U.S. Department of Commerce. The statements, findings, conclusions, and recommendations are those of the authors, and do not necessarily reflect the views of the National Oceanic and Atmospheric Administration, or the U.S. Department of Commerce.
    Keywords: Bottom currents ; Diapycnal mixing ; Turbulence ; Southern Ocean
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  • 12
    Publication Date: 2022-05-27
    Description: Author Posting. © American Meteorological Society, 2021. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 51(7), (2021): 2087–2102, https://doi.org/10.1175/JPO-D-20-0255.1.
    Description: The boundary current system in the Labrador Sea plays an integral role in modulating convection in the interior basin. Four years of mooring data from the eastern Labrador Sea reveal persistent mesoscale variability in the West Greenland boundary current. Between 2014 and 2018, 197 middepth intensified cyclones were identified that passed the array near the 2000-m isobath. In this study, we quantify these features and show that they are the downstream manifestation of Denmark Strait Overflow Water (DSOW) cyclones. A composite cyclone is constructed revealing an average radius of 9 km, maximum azimuthal speed of 24 cm s−1, and a core propagation velocity of 27 cm s−1. The core propagation velocity is significantly smaller than upstream near Denmark Strait, allowing them to trap more water. The cyclones transport a 200-m-thick lens of dense water at the bottom of the water column and increase the transport of DSOW in the West Greenland boundary current by 17% relative to the background flow. Only a portion of the features generated at Denmark Strait make it to the Labrador Sea, implying that the remainder are shed into the interior Irminger Sea, are retroflected at Cape Farewell, or dissipate. A synoptic shipboard survey east of Cape Farewell, conducted in summer 2020, captured two of these features that shed further light on their structure and timing. This is the first time DSOW cyclones have been observed in the Labrador Sea—a discovery that could have important implications for interior stratification.
    Description: A. P. and R. S. P. were funded by National Science Foundation Grants OCE-1259618 and OCE-1756361. I. L. B. and F. S. were funded by National Science Foundation Grants OCE-1258823 and OCE-1756272. N. P. H. was supported by the Natural Environment Research Council U.K. OSNAP program (NE/K010875/1 and NE/K010700/1). M. A. S. was supported by NSF Grants OCE-1558742 and OPP-1822334.
    Description: 2021-12-08
    Keywords: Boundary currents ; Eddies ; Transport
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  • 13
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    American Meteorological Society
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(8),(2020): 2315-2321, doi:10.1175/JPO-D-19-0327.1.
    Description: Low-frequency currents and eddies transport sediment, pathogens, larvae, and heat along the coast and between the shoreline and deeper water. Here, low-frequency currents (between 0.1 and 4.0 mHz) observed in shallow surfzone waters for 120 days during a wide range of wave conditions are compared with theories for generation by instabilities of alongshore currents, by ocean-wave-induced sea surface modulations, and by a nonlinear transfer of energy from breaking waves to low-frequency motions via a two-dimensional inverse energy cascade. For these data, the low-frequency currents are not strongly correlated with shear of the alongshore current, with the strength of the alongshore current, or with wave-group statistics. In contrast, on many occasions, the low-frequency currents are consistent with an inverse energy cascade from breaking waves. The energy of the low-frequency surfzone currents increases with the directional spread of the wave field, consistent with vorticity injection by short-crested breaking waves, and structure functions increase with spatial lags, consistent with a cascade of energy from few-meter-scale vortices to larger-scale motions. These results include the first field evidence for the inverse energy cascade in the surfzone and suggest that breaking waves and nonlinear energy transfers should be considered when estimating nearshore transport processes across and along the coast.
    Description: Funding was provided by a Vannevar Bush Faculty Fellowship [from OUSD(R&E)] and NSF.
    Keywords: Ocean ; Coastlines ; Eddies ; Wave breaking
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  • 14
    Publication Date: 2022-05-26
    Description: © The Author(s), 2020. This article is distributed under the terms of the Creative Commons Attribution License. The definitive version was published in Cusack, J. M., Brearley, J. A., Garabato, A. C. N., Smeed, D. A., Polzin, K. L., Velzeboer, N., & Shakespeare, C. J. Observed eddy-internal wave interactions in the Southern Ocean. Journal of Physical Oceanography, 50(10), (2020): 3042-3062, doi:10.1175/JPO-D-20-0001.1.
    Description: The physical mechanisms that remove energy from the Southern Ocean’s vigorous mesoscale eddy field are not well understood. One proposed mechanism is direct energy transfer to the internal wave field in the ocean interior, via eddy-induced straining and shearing of preexisting internal waves. The magnitude, vertical structure, and temporal variability of the rate of energy transfer between eddies and internal waves is quantified from a 14-month deployment of a mooring cluster in the Scotia Sea. Velocity and buoyancy observations are decomposed into wave and eddy components, and the energy transfer is estimated using the Reynolds-averaged energy equation. We find that eddies gain energy from the internal wave field at a rate of −2.2 ± 0.6 mW m−2, integrated from the bottom to 566 m below the surface. This result can be decomposed into a positive (eddy to wave) component, equal to 0.2 ± 0.1 mW m−2, driven by horizontal straining of internal waves, and a negative (wave to eddy) component, equal to −2.5 ± 0.6 mW m−2, driven by vertical shearing of the wave spectrum. Temporal variability of the transfer rate is much greater than the mean value. Close to topography, large energy transfers are associated with low-frequency buoyancy fluxes, the underpinning physics of which do not conform to linear wave dynamics and are thereby in need of further research. Our work suggests that eddy–internal wave interactions may play a significant role in the energy balance of the Southern Ocean mesoscale eddy and internal wave fields.
    Description: Funding for DIMES was provided by U.K. Natural Environment Research Council (NERC) Grants NE/E007058/1 and NE/E005667/1. JMC acknowledges the support of a NERC PhD studentship, and ACNG that of the Royal Society and the Wolfson Foundation. NV acknowledges support from the ARC Centre of Excellence for Climate Extremes (CLEX) Honours Scholarship and the ANU PBSA Partnership - Spotless Scholarship. CJS acknowledges support from an ARC Discovery Early Career Researcher Award DE180100087 and an Australian National University Futures Scheme award. Numerical simulations were conducted on the National Computational Infrastructure (NCI) facility, Canberra, Australia. This study has been conducted using E.U. Copernicus Marine Service Information. We thank two anonymous reviewers for their comments which helped to improve the manuscript significantly. Codes and output files are available online at the project repository (https://github.com/jessecusack/DIMES_eddy_wave_interactions).
    Keywords: Southern Ocean ; Eddies ; Internal waves ; Turbulence
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  • 15
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(7), (2020): 1839-1852, https://doi.org/10.1175/JPO-D-19-0209.1.
    Description: The Lagrangian characteristics of the surface flow field arising when an idealized, anticyclonic, mesoscale, isolated deep-ocean eddy collides with continental slope and shelf topography are explored. In addition to fluid parcel trajectories, we consider the trajectories of biological organisms that are able to navigate and swim, and for which shallow water is a destination. Of particular interest is the movement of organisms initially located in the offshore eddy, the manner in which the eddy influences the ability of the organisms to reach the shelf break, and the spatial and temporal distributions of organisms that do so. For nonswimmers or very slow swimmers, the organisms arrive at the shelf break in distinct pulses, with different pulses occurring at different locations along the shelf break. This phenomenon is closely related to the episodic formation of trailing vortices that are formed after the eddy collides with the continental slope, turns, and travels parallel to the coast. Analysis based on finite-time Lyapunov exponents reveals initial locations of all successful trajectories reaching the shoreline, and provides maps of the transport pathways showing that much of the cross-shelf-break transport occurs in the lee of the eddy as it moves parallel to the shore. The same analysis shows that the onshore transport is interrupted after a trailing vortex detaches. As the swimming speeds are increased, the organisms are influenced less by the eddy and tend to show up en mass and in a single pulse.
    Description: IR and LP were supported by National Science Foundation (NSF) Grant OCE-1558806. DC was supported by NSF U.S. National Science Foundation’s Physical Oceanography program through Grants OCE-1059632 and OCE-1433953 as well as the Academic Programs Office, Woods Hole Oceanographic Institution. We acknowledge high-performance computing support from Yellowstone (http://n2t.net/ark:/85065/d7wd3xhc) provided by NCAR’s Computational and Information Systems Laboratory, sponsored by the National Science Foundation.
    Keywords: Ocean ; Eddies ; Nonlinear dynamics ; Transport
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  • 16
    Publication Date: 2022-05-27
    Description: Author Posting. © American Meteorological Society, 2021. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 51(8),(2021): 2425–2441, https://doi.org/10.1175/JPO-D-20-0317.1.
    Description: The frequency and latitudinal dependence of the midlatitude wind-driven meridional overturning circulation (MOC) is studied using theory and linear and nonlinear applications of a quasigeostrophic numerical model. Wind forcing is varied either by changing the strength of the wind or by shifting the meridional location of the wind stress curl pattern. At forcing periods of less than the first-mode baroclinic Rossby wave basin crossing time scale, the linear response in the middepth and deep ocean is in phase and opposite to the Ekman transport. For forcing periods that are close to the Rossby wave basin crossing time scale, the upper and deep MOC are enhanced, and the middepth MOC becomes phase shifted, relative to the Ekman transport. At longer forcing periods the deep MOC weakens and the middepth MOC increases, but eventually for long enough forcing periods (decadal) the entire wind-driven MOC spins down. Nonlinearities and mesoscale eddies are found to be important in two ways. First, baroclinic instability causes the middepth MOC to weaken, lose correlation with the Ekman transport, and lose correlation with the MOC in the opposite gyre. Second, eddy thickness fluxes extend the MOC beyond the latitudes of direct wind forcing. These results are consistent with several recent studies describing the four-dimensional structure of the MOC in the North Atlantic Ocean.
    Description: This study was supported by National Science Foundation Grant OCE-1947290.
    Description: 2022-01-13
    Keywords: Eddies ; Large-scale motions ; Meridional overturning circulation ; Ocean dynamics ; Planetary waves
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  • 17
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(6), (2020): 1557-1582, doi:10.1175/JPO-D-19-0239.1.
    Description: We examine various contributions to the vertical velocity field within large mesoscale eddies by analyzing multiple solutions to an idealized numerical model of a representative anticyclonic warm core Gulf Stream ring. Initial conditions are constructed to reproduce the observed density and nutrient profiles collected during the Warm Core Rings Program. The contributions to vertical fluxes diagnosed from the numerical simulations are compared against a divergence-based, semidiagnostic equation and a generalized omega equation to better understand the dynamics of the vertical velocity field. Frictional decay alone is found to be ineffective in raising isopycnals and transporting nutrients to the upper ocean. With representative wind forcing, the magnitude of vorticity gradient–induced Ekman pumping is not necessarily larger than the current-induced counterpart on a time scale relevant to ecosystem response. Under realistic forcing conditions, strain deformation can perturb the ring to be noncircular and induce vertical velocities much larger than the Ekman vertical velocities. Nutrient budget diagnosis, together with analysis of the relative magnitudes of the various types of vertical fluxes, allows us to describe the time-scale dependence of nutrient delivery. At time scales that are relevant to individual phytoplankton (from hours to days), the magnitudes of nutrient flux by Ekman velocities and deformation-induced velocities are comparable. Over the life span of a typical warm core ring, which can span multiple seasons, surface current–induced Ekman pumping is the most effective mechanism in upper-ocean nutrient enrichment because of its persistence in the center of anticyclones regardless of the direction of the wind forcing.
    Description: This work was supported by the National Science Foundation Ocean Science Division under Grant OCE-1558960. PG also acknowledges support of the NASA Physical Oceanography Program Grant NNX16H59G. KC would like to thank D. McGillicuddy Jr. for inspiring discussions and suggestions during the course of this study. Constructive comments from two anonymous reviewers are appreciated.
    Keywords: Ageostrophic circulations ; Eddies ; Ekman pumping/transport ; Mesoscale processes ; Upwelling/downwelling ; Vertical motion
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  • 18
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(4), (2020): 1045-1064, doi:10.1175/JPO-D-19-0137.1.
    Description: Three simulations of the circulation in the Gulf of Mexico (the “Gulf”) using different numerical general circulation models are compared with results of recent large-scale observational campaigns conducted throughout the deep (〉1500 m) Gulf. Analyses of these observations have provided new understanding of large-scale mean circulation features and variability throughout the deep Gulf. Important features include cyclonic flow along the continental slope, deep cyclonic circulation in the western Gulf, a counterrotating pair of cells under the Loop Current region, and a cyclonic cell to the south of this pair. These dominant circulation features are represented in each of the ocean model simulations, although with some obvious differences. A striking difference between all the models and the observations is that the simulated deep eddy kinetic energy under the Loop Current region is generally less than one-half of that computed from observations. A multidecadal integration of one of these numerical simulations is used to evaluate the uncertainty of estimates of velocity statistics in the deep Gulf computed from limited-length (4 years) observational or model records. This analysis shows that the main deep circulation features identified from the observational studies appear to be robust and are not substantially impacted by variability on time scales longer than the observational records. Differences in strengths and structures of the circulation features are identified, however, and quantified through standard error analysis of the statistical estimates using the model solutions.
    Description: This work was supported by the Gulf Research Program of the National Academy of Sciences under Awards 2000006422 and 2000009966. The content is solely the responsibility of the authors and does not necessarily represent the official views of the Gulf Research Program or the National Academy of Sciences. The authors acknowledge the GLORYS project for providing the ocean reanalysis data used in the ROMS simulation. GLORYS is jointly conducted by MERCATOR OCEAN, CORIOLIS, and CNRS/INSU. Installation, recovery, data acquisition, and processing of the CANEK group current-meter moorings were possible because of CICESE-PetróleosMexicanos Grant PEP-CICESE 428229851 and the dedicated work of the crew of the B/O Justo Sierra and scientists of the CANEK group. The authors thank Dr. Aljaz Maslo, CICESE, for assistance with analysis of model data. The Bureau of Ocean Energy Management (BOEM), U.S. Dept. of the Interior, provided funding for the Lagrangian Study of the Deep Circulation in the Gulf of Mexico and the Observations and Dynamics of the Loop Current study. HYCOM simulation data are available from the HYCOM data server (https://www.hycom.org/data/goml0pt04/expt-02pt2), MITgcm data are available from the ECCO data server (http://ecco.ucsd.edu/gom_results2.html), and the ROMS simulation data are available from GRIIDC (NA.x837.000:0001).
    Keywords: Ocean circulation ; Abyssal circulation ; Bottom currents/bottom water ; Eddies ; Ocean models
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  • 19
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(5),(2020): 1227-1244, doi:10.1175/JPO-D-19-0280.1.
    Description: The Nordic seas are commonly described as a single basin to investigate their dynamics and sensitivity to environmental changes when using a theoretical framework. Here, we introduce a conceptual model for a two-basin marginal sea that better represents the Nordic seas geometry. In our conceptual model, the marginal sea is characterized by both a cyclonic boundary current and a front current as a result of different hydrographic properties east and west of the midocean ridge. The theory is compared to idealized model simulations and shows good agreement over a wide range of parameter settings, indicating that the physics in the two-basin marginal sea is well captured by the conceptual model. The balances between the atmospheric buoyancy forcing and the lateral eddy heat fluxes from the boundary current and the front current differ between the Lofoten and the Greenland Basins, since the Lofoten Basin is more strongly eddy dominated. Results show that this asymmetric sensitivity leads to opposing responses depending on the strength of the atmospheric buoyancy forcing. Additionally, the front current plays an essential role for the heat and volume budget of the two basins, by providing an additional pathway for heat toward the interior of both basins via lateral eddy heat fluxes. The variability of the temperature difference between east and west influences the strength of the different flow branches through the marginal sea and provides a dynamical explanation for the observed correlation between the front current and the slope current of the Norwegian Atlantic Current in the Nordic seas.
    Description: We thank Ilker Fer and two anonymous reviewers whose comments improved this paper. S. L. Ypma and S. Georgiou were supported by NWO (Netherlands Organisation for Scientific Research) VIDI Grant 864.13.011 awarded to C. A. Katsman. M. A. Spall was supported by National Science Foundation Grants OCE-1558742 and OPP-1822334. E. Lambert is funded by the ERA4CS project INSeaPTION. The model data analyzed in this study are available on request from the corresponding author. This study has been conducted using E.U. Copernicus Marine Service Information. The altimeter products were produced by Ssalto/Duacs and distributed by Aviso+, with support from CNES (https://www.aviso.altimetry.fr).
    Description: 2020-10-27
    Keywords: Boundary currents ; Deep convection ; Eddies ; Fronts ; Instability ; Meridional overturning circulation
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  • 20
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2019. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 49(11), (2019): 2867-2881, doi: 10.1175/JPO-D-19-0072.1.
    Description: The Antarctic Circumpolar Current plays a central role in the ventilation of heat and carbon in the global ocean. In particular, the isopycnal slopes determine where each water mass outcrops and thus how the ocean interacts with the atmosphere. The region-integrated isopycnal slopes have been suggested to be eddy saturated, that is, stay relatively constant as the wind forcing changes, but whether or not the flow is saturated in realistic present day and future parameter regimes is unknown. This study analyzes an idealized two-layer quasigeostrophic channel model forced by a wind stress and a residual overturning generated by a mass flux across the interface between the two layers, with and without a blocking ridge. The sign and strength of the residual overturning set which way the isopycnal slopes change with the wind forcing, leading to an increase in slope with an increase in wind forcing for a positive overturning and a decrease in slope for a negative overturning, following the usual conventions; this behavior is caused by the dominant standing meander weakening as the wind stress weakens causing the isopycnal slopes to become more sensitive to changes in the wind stress and converge with the slopes of a flat-bottomed simulation. Eddy saturation only appears once the wind forcing passes a critical level. These results show that theories for saturation must have both topography and residual overturning in order to be complete and provide a framework for understanding how the isopycnal slopes in the Southern Ocean may change in response to future changes in wind forcing.
    Description: MKY and RF acknowledge support through NSF Awards OCE-1536515 and AGS-1835576. MKY acknowledges funding from NDSEG. GRF was supported by NSF OCE-1459702. We are very grateful for conversations with David Marshall, Andrew Stewart, and two anonymous reviewers that greatly improved the manuscript. The code for running the model is found at https://github.com/mkyoungs/JPO-QG-Channel.
    Description: 2020-04-30
    Keywords: Southern Ocean ; Eddies ; Storm tracks ; Quasigeostrophic models
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  • 21
    Publication Date: 2022-05-26
    Description: © The Author(s), 2019. This article is distributed under the terms of the Creative Commons Attribution License. The definitive version was published in Cusack, J. M., Voet, G., Alford, M. H., Girton, J. B., Carter, G. S., Pratt, L. J., Pearson-Potts, K. A., & Tan, S. Persistent turbulence in the Samoan Passage. Journal of Physical Oceanography, 49(12), (2019): 3179-3197, doi: 10.1175/JPO-D-19-0116.1.
    Description: Abyssal waters forming the lower limb of the global overturning circulation flow through the Samoan Passage and are modified by intense mixing. Thorpe-scale-based estimates of dissipation from moored profilers deployed on top of two sills for 17 months reveal that turbulence is continuously generated in the passage. Overturns were observed in a density band in which the Richardson number was often smaller than ¼, consistent with shear instability occurring at the upper interface of the fast-flowing bottom water layer. The magnitude of dissipation was found to be stable on long time scales from weeks to months. A second array of 12 moored profilers deployed for a shorter duration but profiling at higher frequency was able to resolve variability in dissipation on time scales from days to hours. At some mooring locations, near-inertial and tidal modulation of the dissipation rate was observed. However, the modulation was not spatially coherent across the passage. The magnitude and vertical structure of dissipation from observations at one of the major sills is compared with an idealized 2D numerical simulation that includes a barotropic tidal forcing. Depth-integrated dissipation rates agree between model and observations to within a factor of 3. The tide has a negligible effect on the mean dissipation. These observations reinforce the notion that the Samoan Passage is an important mixing hot spot in the global ocean where waters are being transformed continuously.
    Description: The authors thank Zhongxiang Xao and Jody Klymak, who provided earlier setups of the numerical model, and also Arjun Jagannathan for insightful discussions on the subject of flow over topography. We also thank John Mickett and Eric Boget for their assistance in designing, deploying, and recovering the moorings. In addition, we also thank the crew and scientists aboard the R/V Revelle and R/V Thompson, without whom the data presented in this paper could not have been gathered. Ilker Fer and two anonymous reviewers provided thoughtful feedback that improved the paper. This work was supported by the National Science Foundation under Grants OCE-1029268, OCE-1029483, OCE-1657264, OCE-1657795, OCE-1657870, and OCE-1658027.
    Keywords: Gravity waves ; Turbulence ; Abyssal circulation ; Mixing ; Tides
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  • 22
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(3), (2020): 715-726, doi:10.1175/JPO-D-19-0021.1.
    Description: Closing the overturning circulation of bottom water requires abyssal transformation to lighter densities and upwelling. Where and how buoyancy is gained and water is transported upward remain topics of debate, not least because the available observations generally show downward-increasing turbulence levels in the abyss, apparently implying mean vertical turbulent buoyancy-flux divergence (densification). Here, we synthesize available observations indicating that bottom water is made less dense and upwelled in fracture zone valleys on the flanks of slow-spreading midocean ridges, which cover more than one-half of the seafloor area in some regions. The fracture zones are filled almost completely with water flowing up-valley and gaining buoyancy. Locally, valley water is transformed to lighter densities both in thin boundary layers that are in contact with the seafloor, where the buoyancy flux must vanish to match the no-flux boundary condition, and in thicker layers associated with downward-decreasing turbulence levels below interior maxima associated with hydraulic overflows and critical-layer interactions. Integrated across the valley, the turbulent buoyancy fluxes show maxima near the sidewall crests, consistent with net convergence below, with little sensitivity of this pattern to the vertical structure of the turbulence profiles, which implies that buoyancy flux convergence in the layers with downward-decreasing turbulence levels dominates over the divergence elsewhere, accounting for the net transformation to lighter densities in fracture zone valleys. We conclude that fracture zone topography likely exerts a controlling influence on the transformation and upwelling of bottom water in many areas of the global ocean.
    Description: The data used in this study were collected in the context of several projects funded by the U.S. National Science Foundation (NSF), in particular BBTRE (OCE-9415589 and OCE-9415598) and DoMORE (OCE-1235094). Funding for the analysis was provided as part of the NSF DoMORE and DECIMAL (OCE-1735618) projects. Author Ijichi is a Japan Society for the Promotion of Science (JSPS) Overseas Research Fellow. Comments on an early draft of this paper by Jim Ledwell and Bryan Kaiser, as well as topical discussions with Jörn Callies and Trevor McDougall, are gratefully acknowledged. The paper was greatly improved during the review process, in particular because of the critical comments from one of the two anonymous reviewers.
    Keywords: Diapycnal mixing ; Topographic effects ; Turbulence ; Upwelling/downwelling ; Bottom currents/bottom water
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  • 23
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    American Meteorological Society
    Publication Date: 2022-05-26
    Description: Author Posting. © American Meteorological Society, 2020. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Physical Oceanography 50(1), (2020): 255-268, doi:10.1175/JPO-D-19-0166.1.
    Description: Regional connectivity is important to the global climate salinity response, particularly because salinity anomalies do not have a damping feedback with atmospheric freshwater fluxes and may therefore be advected over long distances by ocean circulation, resulting in nonlocal influences. Climate model intercomparison experiments such as CMIP5 exhibit large uncertainty in some aspects of the salinity response, hypothesized here to be a result of ocean dynamics. We use two types of Lagrangian particle tracking experiments to investigate pathways of exchange for salinity anomalies. The first uses forward trajectories to estimate average transport time scales between water cycle regimes. The second uses reverse trajectories and a freshwater accumulation method to quantitatively identify remote influences in the salinity response. Additionally, we compare velocity fields with both resolved and parameterized eddies to understand the impact of eddy stirring on intergyre exchange. These experiments show that surface anomalies are readily exchanged within the ocean gyres by the mean circulation, but intergyre exchange is slower and largely eddy driven. These dynamics are used to analyze the North Atlantic salinity response to climate warming and water cycle intensification, where the system is broadly forced with fresh surface anomalies in the subpolar gyre and salty surface anomalies in the subtropical gyres. Under these competing forcings, strong intergyre eddy fluxes carry anomalously salty subtropical water into the subpolar gyre which balances out much of the local freshwater input.
    Description: We acknowledge the World Climate Research Programme’s Working Group on Coupled Modelling, which is responsible for CMIP, and we thank the climate modeling groups (listed in Table 1 of this paper) for producing and making available their model output. We also thank the creators of the SODA and ECCO reanalysis products. This work was supported by NASA Headquarters under the NASA Earth and Space Science Fellowship Program Award 80NSSC17K0372, and by National Science Foundation Award OCE-1433132. The SODA outputs used here can be accessed at http://www.atmos.umd.edu/~ocean/, and the ECCO outputs at https://ecco.jpl.nasa.gov/. Data from the CMIP5 ensemble is available at https://esgf-node.llnl.gov/projects/esgf-llnl/. The particle tracking code used for these experiments can be found at https://github.com/slevang/particle-tracking.
    Description: 2020-07-20
    Keywords: North Atlantic Ocean ; Eddies ; Hydrologic cycle ; Lagrangian circulation/transport ; Transport ; Climate change
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  • 24
    ISSN: 1570-7458
    Keywords: butterfly ; Formicidae ; food ; growth ; mutualism ; myrmecophily ; parasitism
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Caterpillars of the lycaenid butterfly Maculinea rebeli Hirschke (Lepidoptera: Lycaenidae) live for 11–23 months as social parasites in Myrmica (Hymenoptera: Formicidae) red ant nests, a trait that is believed to have evolved from mutualistic myrmecophilous ancestry. Although Maculinea rebeli caterpillars harm Myrmica larvae, they simultaneously produce copious secretions which the adult worker ants imbibe, perhaps representing a vestige of the ancestral mutualism. We report the results of laboratory experiments designed to test alternative hypotheses: (i) Maculinea rebeli caterpillars provide a beneficial source of sugar in return for being tended by Myrmicaworkers; (ii) Maculinea rebeli harms its host by stressing the workers by competing for available sugar. Comparisons were made of Myrmica worker fitness after 90–450 days under all possible combinations of three experimental treatments: ± M. rebeli caterpillars, ± sucrose and ± ant brood. Caterpillars always reduced the survival of both ant workers and their larvae, even when sugar was not provided, suggesting that M. rebeli is wholly parasitic on all stages in its host colony. The results also confirmed the importance of sucrose in the diet of Myrmica, and showed that M. rebeli caterpillars which eat ant brood to supplement their normal trophallactic feeding by workers develop more quickly - but have the same survival and pupal weights – as caterpillars that are fed solely by worker ants.
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  • 25
    ISSN: 1572-879X
    Keywords: aluminum oxide ; hydroxyl groups ; rhodium ; growth ; scanning tunneling microscopy ; infrared spectroscopy
    Source: Springer Online Journal Archives 1860-2000
    Topics: Chemistry and Pharmacology
    Notes: Abstract In order to investigate how the presence of surface hydroxyl groups on oxide surfaces affects the interaction with the supported metal, we have modified a well-ordered alumina film on NiAl(110) by Al deposition and subsequent exposure to water. This procedure yields a hydroxylated alumina surface as revealed by infrared and high-resolution electron energy loss spectroscopy. By means of scanning tunneling microscopy, we have studied the growth of rhodium on the modified film at 300 K. Clear differences in the particle distribution and density are observed in comparison to the clean substrate. While, in the latter case, decoration of domain boundaries as typical defects of the oxide film governs the growth mode, a more isotropic island distribution and a drastically increased particle density is found on the hydroxylated surface. From infrared data, it can be deduced that the growth is connected with the consumption of the hydroxyl groups due to the interaction between the metal deposit and the hydroxylated areas. This finding is in line with photoemission results published earlier.
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  • 26
    Electronic Resource
    Electronic Resource
    Springer
    Small business economics 14 (2000), S. 195-210 
    ISSN: 1573-0913
    Keywords: growth ; manufacturing ; performance ; product innovation ; small firms
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract The paper considers the relative performance [along a number of parameters] of a sample of 228 small manufacturing firms categorised by level of innovation. Whilst innovators appear no more likely to have experienced some form of sales or employment growth, they are significantly more likely to have grown more. In other words, the innovators' growth rate distributions are highly negatively skewed. With regards to export intensities, profitability and productivity levels, the findings are less clear. On the whole, the results reported here are similar to those of other small firm studies, yet vary markedly from large firm equivalents; suggesting that the nature of the returns to innovation may be contingent, at least in part, upon firm size. Moreover, the high levels of variation in firm performance should caution us against proffering innovative imperatives. If we are to counsel firms to "innovate at all costs", we must be clear about, and clearly demonstrate, the nature of the returns they may reasonably expect and the processes through which these may be optimised.
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  • 27
    Electronic Resource
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    Journal of population economics 13 (2000), S. 403-424 
    ISSN: 1432-1475
    Keywords: JEL classification: O41 ; F22 ; Key words: Altruism ; education ; growth ; convergence ; capital mobility
    Source: Springer Online Journal Archives 1860-2000
    Topics: Sociology , Economics
    Notes: Abstract. The aim of this paper is to discuss the process of regional convergence within the framework of an overlapping generations model in which the engine of growth is the accumulation of human capital. In particular, we consider different education funding systems and compare their performance in terms of growth rates and pace of convergence between two heterogeneous regions. The analysis suggests that the choice of a particular education system incorporates a possible trade-off between long run growth rate and short run convergence. In such choice, the initial capital stock and the extent of regional human capital discrepancy appear as central variables.
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  • 28
    Electronic Resource
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    Ecological research 15 (2000), S. 101-106 
    ISSN: 1440-1703
    Keywords: comparative ecology ; growth ; marine fish ; patterns ; reproduction
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: A number of strong regularities characterize certain very basic biological parameters in marine fishes. For example, the ovulated eggs of fish usually measure approximately 1 mm in diameter. The small, relatively uniform size of the eggs means that almost all fish larvae experience environmental variability at very similar scales, which itself establishes strong constraints for, and links between reproduction and recruitment. Additional constraints emerge from seawater being a poor medium for respiration, which establishes further linkages between growth and mortality. These constraints have produced strongly convergent features, and thence the patterns in reproduction and growth of marine fishes that are presented.
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  • 29
    ISSN: 1573-143X
    Keywords: body composition ; catfish (Mystus nemurus) ; feeding rate ; growth
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract A 10 week experiment was conducted to determine theeffects of feeding rate on growth, feed utilizationand body composition of the tropical bagrid catfish,Mystus nemurus. Catfish fingerlings with anaverage initial body weight of 12 g were fed apractical diet (36.2% protein, 16.5 kJ/g diet) atrates of 1, 2, 2.5, 3, 3.5, 4 or 5% of their bodyweight (BW) per day in two equal meals. Watertemperature was approximately 29 °C throughoutthe experiment. Percent weight gain increased almostlinearly with increasing feeding rates up to 2.5%BW/day beyond which no significant (P 〉 0.05)improvement in weight gain was observed. The specificgrowth rate of catfish fed rations of 1% BW/day was0.72%/day and this increased significantly to anaverage of 1.39%/day for catfish fed 2.5% BW/day andbeyond. Feed utilization did not differ significantly(P 〉 0.05) between fish fed 1.0 to 2.5%BW/day but decreased when rations were increased to3.0% BW/day and above. Feed efficiency ratio was0.79 for catfish fed 1.0% BW/day compared to a ratioof 0.27 for fish fed at 5% BW/day. Catfish fed 1.0%BW/day had the lowest condition factor, hepato- andviscerosomatic indices, but the highest carcass tobody weight ratio. These fish also had lowerproportions of whole body dry matter, lipid andprotein, carcass dry matter and lipid, and visceraldry matter and lipid than fish in other groups. Therewere no significant differences in either conditionindices or relative body composition of fish fedrations of 2.0 to 5.0% BW/day. Based on the growth,feed efficiency and body composition data obtained, afeeding rate of 2.5% BW per day is recommended forM. nemurus fingerlings raised at 29 °C.
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  • 30
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    Aquaculture international 8 (2000), S. 531-542 
    ISSN: 1573-143X
    Keywords: feed intake ; growth ; protein utilisation ; proximate ; composition ; rainbow trout (Oncorhynchus mykiss)
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Rainbow trout Oncorhynchus mykiss grew from 44 to 326 g in 96days when held at ∼12 °C. Fish were fed to satiation twice dailywith either high (L1: 30.8%, L2:31.4%) or lower-lipid feeds (C1: 18.8%,C2: 21.8%). Four feeding treatments were studied.Group C1C2 received feed C1 for 43 days(days 0–43) and C2 thereafter (days 44–96).Groups L1L2, L1C2 andC1L2 were subjected to dietary changes asindicated by the feed designations. After a short period of feedadaptation, fish ingested similar amounts of feed energy i.e., they ateless by weight of the lipid-rich (L) feeds. Feed lipid content did notaffect growth but fish fed L-feed had reduced feed conversion ratio(FCR) compared to fish fed C-feed (0.731 vs. 0.773) during days0–43 (P 〈 0.01). After 96 days,L1L2-fish were lower in body protein(15.8%) than the C1C2-fish (16.8%)(P 〈 0.01). L-feeds also tended to increase percentage lipidand reduce percentage whole body moisture and ash. A higher net proteinutilisation (NPU) was recorded in fish fed L-feeds (43.6%)compared to fish fed C-feeds (38.8%) in days 0–43(P 〈 0.05). This seemed to be the result of a lower proteinintake rather than a protein-sparing effect of feed lipid. Above athreshold value of approximately 6.5 mg protein eaten·g bodywtminus 1·day−1, NPU decreased.
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  • 31
    ISSN: 1573-143X
    Keywords: aggression ; Atlantic salmon (Salmo salar) ; dominance ; growth
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Aggression in groups of 0+ Atlantic salmon (Salmo salar) was monitoredat weekly intervals in two tanks containing 100 fish each. Three 1 + salmonparr were added to one of these. After 5 weeks, fish weights were measuredin both tanks and the conditions reversed. At ten weeks, weights of fish inboth tanks were measured again. In both populations, levels of aggressionamong the smaller fish were significantly lower and growth ratessignificantly higher when the large fish were present. Although the largefish attacked the small ones, the rate at which they did so was an order ofmagnitude lower that the rate at which small fish attacked each other in theabsence of larger conspecifics. This raises the possibility that levels ofaggression among farmed salmon might be reduced by the addition of a fewlarge conspecifics.
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  • 32
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    Boundary layer meteorology 94 (2000), S. 461-493 
    ISSN: 1573-1472
    Keywords: Radiation fog ; Large-eddy simulation ; Turbulence ; Kelvin–Helmholtz instability ; Convection ; Mixed layer
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In order to study the three-dimensional structure of radiation fogand to obtain a basic understanding of its generation mechanism,a numerical experiment is performed with a large-eddysimulation model and compared with the observation at Cabauw in the Netherlands. After confirming that the results are insatisfactory agreement with the observations, the structure of thefog and its generation mechanism are examined in more detail. Before the fog forms, the atmosphere is stable and an inversionlayer exists almost adjacent to the ground surface. As the fog grows, however, the stratification is destabilized and a mixed layerdevelops gradually. The longwave radiative cooling near thefog top contributes to the destabilization more than thecondensational heating does. The evolution of the fog can be classified into three stagesaccording to the behaviour of turbulent kinetic energy (TKE):formation, development, and dissipation stages.The fog layer has different flow structures at each stage.During the formation stage, longitudinal rolls similar tostreaks in channel flows appear near the ground surface.The development stage is characterized by an initiation oftransverse bands due to Kelvin–Helmholtz instability anda sudden increase of TKE. During the dissipation stage, longitudinalrolls and polygonal cells due to convective instability are organized.
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  • 33
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    Boundary layer meteorology 97 (2000), S. 219-249 
    ISSN: 1573-1472
    Keywords: Conditional concentration statistics ; Mixing ; Scalar dissipation ; Turbulence
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We analyse cross-wind-integrated statistics of theconcentration field of a conserved scalar for pointand line sources in grid turbulence. In particular,using wind-tunnel measurements we calculate thecross-wind integrated probability density function(pdf) for the scalar concentration. We then use thatquantity in the exact evolution equation for the pdfto calculate the cross-wind integrated mean of therate of dissipation of scalar variance, conditional onthe scalar concentration. Much of the variation ofthese statistics with distance downstream is accountedfor by scaling with concentration, length and timescales based on the development of the mean plume.This scaling thus suggests some simple practicalparameterisations of these statistics in terms ofmean-field quantities. One of the motivations for thiswork is to find a simple parameterisation for thescalar dissipation that can be used for modellingchemical reactions in plumes. We also consider the cross-wind integral of the firstfew moments of the concentration field and show thatthe integration greatly simplifies the budgets forthese moments. Thus the first moment is just thedownstream flux of the scalar, which is constant. Thesecond moment budget provides a check on the meandissipation estimated directly from the pdf evolutionequation.
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  • 34
    ISSN: 1573-1472
    Keywords: Katabatic flow ; Spectra ; Stable boundary layer ; Turbulence ; Waves
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Observations obtained over a glacier surface in a predominantlykatabatic flow and with a distinctwind maximum below 13-m height are presented. The data werecollected using a 13-m high profilemast and two sonic anemometers (at about 2.5-m and 10-m heights).The spectra at frequencies belowthat of the turbulence range appear to deviate considerably fromthe curves obtained by Kaimal andco-workers during the 1968 Kansas experiment. The characteristicsof these deviations are compared tothe observations of others in surface-layers disturbed by anykind of large-scale outer-layer (orinactive) turbulence. In our case the disturbances arelikely to be induced by the highmountain ridges that surround the glacier. Moreover, the deviationsobserved in the cospectra seemto result from an, as yet, unspecified interaction between theinactive outer-layer turbulenceand the local surface-layer turbulence. Near the distinctwind maximum turbulence production ceasedwhile turbulence itself did not, probably the result ofturbulence transport from other levels. Consequently, we studied thelocal similarity relations using σw instead of u* as an alternative velocity scale. Wellbelow the wind maximum, and for relatively low stability(0〈 Rig 〈0.2), the flow behaves accordingto well established local-scaling similarity relationshipsin the stable boundary layer. For higherstability (Rig 〉 0.2), and near or above the wind maximum, the boundary-layer structure conforms tothat of z-less stratification suggesting that the eddy sizeis restricted by the local stability ofthe flow. In line with this we observed that the sensibleheat fluxes relate remarkably well to thelocal flow parameters.
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  • 35
    ISSN: 1573-1472
    Keywords: Coherent structures ; Numerical modelling ; Turbulence
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract In two preceding papers, coherent structures of theatmospheric boundary layer (ABL), such as rollvortices or cells, were investigated through radar andaircraft observations collected during the TRAC-93(Turbulence Radar Aircraft Cells) experiment held inFrance in June 1993. The analysis of this experimentaldata set provided information on the spatialcharacteristics of these organisations (length scale,orientation, type ... ), their temporal and verticalevolution, and their relation with the dynamic andthermodynamic conditions of the ABL. For the thirdpaper in this series, a large eddy simulation model is used to examine the impact of thecoherent structures on the ABL vertical fluxes. Theanalysis of the simulated horizontal fields is madewith two-dimensional auto and cross-correlationsapplied on different pertinent ABL variables. Theresults emphasise a directional anisotropy of theseorganised fields throughout the ABL, much morepronounced in the heat flux fields, not only at thelength scale of organisations but also at theturbulence scales. This finding has an importantconsequence for traditional ABL flux measurementsbased on the hypothesis of isotropic and homogeneousturbulence. It can explain part of the underestimationof the surface fluxes often mentioned in theliterature. This approach makes it possible tomodify the concept of diffusion time (in chemicalmodelling) and could also lead to revised ABLparameterisations in Range Scale models.
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  • 36
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    Journal of chemical ecology 26 (2000), S. 2015-2023 
    ISSN: 1573-1561
    Keywords: Allelopathy ; growth ; phenolics ; Hebeloma ; Hymenoscyphus ; Picea abies ; Vaccinium myrtillus
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Chemistry and Pharmacology
    Notes: Abstract Allelopathy due to humus phenolics is a cause of natural regeneration deficiency in subalpine Norway spruce (Picea abies) forests. If inhibition of spruce germination and seedling growth due to allelochemicals is generally accepted, in contrast there is a lack of knowledge about phenolic effects on mycorrhizal fungi. Thus, this work tested effects of a humic solution and its naturally occurring phenolics on the growth and respiration of two mycorrhizal fungi: Hymenoscyphus ericae (symbiont of Vaccinium myrtillus, the main allelochemical-producing plant) and Hebeloma crustuliniforme (symbiont of P. abies, the target plant). Growth and respiration of H. crustuliniforme were inhibited by growth medium with the original humic solution (−6% and −30%), respectively, whereas the same humic solution did not affect growth but decreased respiration of H. ericae (−55%). When naturally occurring phenolics (same chemicals and concentrations in the original humic solution) were added to the growth medium, growth of H. crustuliniforme was not affected, whereas that of H. ericae significantly increased (+10%). We conclude that H. ericae is better adapted to the allelopathic constraints of this forest soil than H. crustuliniforme and that the dominance of V. myrtillus among understory species could be explained in this way.
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  • 37
    ISSN: 1608-3237
    Keywords: poly-β-hydroxybutyric acid ; Yersinia pseudotuberculosis ; Listeria monocytogenes ; temperature ; growth
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A comparative investigation of the intracellular content of poly-β-hydroxybutyric acid showed that Yersinia pseudotuberculosis strains accumulated, on the average, lower amounts of this reserve substance than Listeria monocytogenes strains. The intracellular pool of poly-β-hydroxybutyric acid was responsible for the growth of the bacteria at low temperatures (4–6°C) in the absence of any exogenous carbon and energy source.
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  • 38
    ISSN: 1608-3407
    Keywords: Scenedesmus quadricauda ; growth ; cell size ; photosynthetic activity ; imazalil sulfate ; three-phase dose response
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Three-phase dose responses of biological systems of different levels of organization are often called “paradoxical” because the biological effects are clearly manifested under low- and high-intensity treatments, but are absent during moderate-strength treatments. In this work, we found anomalous changes in the cell number of a green alga Scenedesmus quadricauda (Turp.) Breb. grown in the presence of the fungicide imazalil sulfate. At low imazalil concentrations (2.5 × 10–9–2.5 × 10–6 M), the slow increase in the cell number as compared to an untreated culture was not related to cell death. As seen by the dynamics of the population structure and cell functional characteristics (photosynthesis, thermal stability of photosynthetic membranes, etc.), the decrease in the growth rate at low concentrations of imazalil (2–10 × 10–9 M) was due to a long-term arrest of cell division in a fraction of the cell population rather than to a decrease in the rate of division. The absence of a toxic effect or even a slight stimulation of culture growth at moderate concentrations (0.05–1.25 × 10–6 M) was due to the resumption of cell division after a temporal cessation. At these concentrations, imazalil induced cell stress and adaptive elevation of cell tolerance to the fungicide (acclimation). Cell death was observed only at a high fungicide content in the medium (6.25 × 10–6 and higher). Thus, the three-phase (bimodal) dose response corresponds to two regimes (steady-states) of cell functioning which differ in cell sensitivity to external stimuli. The low-sensitivity state, which is characteristic of cells that have experienced stress, is likely to be the state known as “hormesis.”
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  • 39
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    Journal of applied phycology 12 (2000), S. 185-189 
    ISSN: 1573-5176
    Keywords: blue-green alga ; cyanobacterium ; carbon dioxide ; culture ; growth ; Nostocflagelliforme ; rehydration, watering
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The terrestrial blue-green alga (cyanobacterium), Nostoc flagelliforme, was cultured in air at variouslevels of CO2, light and watering to see theireffects on its growth. The alga showed the highestrelative growth rate at the conditions of highCO2 (1500 ppm), high light regime (219–414μmol m-2s-1) and twice daily watering,but the lowest rate at the conditions of low light(58–114 μmol m-2s-1) and daily twicewatering. Increased watering had little effect ongrowth rate at 350 ppm CO2, but increased byabout 70% at 1500ppm CO2 under high lightconditions. It was concluded that enriched CO2could enhance the growth of N. flagelliformewhen sufficient light and water was supplied.
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  • 40
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    Aquatic ecology 34 (2000), S. 19-27 
    ISSN: 1573-5125
    Keywords: adaptation ; fitness ; food quality ; growth ; reproduction ; temporary ponds
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Temporary pond Daphnia flourish on relatively poor-quality food, suggesting adaptation to stringent temporary pond conditions. We conducted laboratory life history experiments on populations of Daphnia obtusa from a shaded woodland temporary pond (short hydroperiod, dystrophic) and an open farmland temporary pond (long hydroperiod, eutrophic), and compared a suite of physical/chemical conditions in the ponds to evaluate habitat conditions. We hypothesized that the shaded woodland pond population would be more fit in terms of life history variables for individuals (age and size at first reproduction, mean brood size, mean number of neonates) and populations (generation time, net reproductive rate R 0, and intrinsic reproductive rate r) given a standard, low-quality food (trout chow and yeast). Life history traits of woodland pond animals were mixed, relative to farm pond animals, and consistent with bet-hedging for an unpredictable habitat. Values of life history traits rivaled or exceeded those of other studies using phytoplankton as food, and were influenced by the pond water used for our study. Life histories clearly differ among local and regional temporary pond Daphnia obtusa populations, and should be valuable for examining the relative influences of local selection and metapopulation dynamics on population structure.
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  • 41
    ISSN: 1573-5176
    Keywords: desiccation ; growth ; growth model ; inorganic carbon ; nutrients ; photoperiod ; photosynthesis ; pigments ; Porphyra linearis ; PPF ; respiration ; temperature ; water velocity
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The effect of environmental parameters on the growthof Porphyra linearis gametophytes was examinedunder controlled conditions, and related to themultilinear regression growth model recently developedfor this seaweed under coastal conditions in theeastern Mediterranean. Growth chambers, a gradienttable, special culture devices and analytical methodswere combined for this culture study.The major factors significantly controlling thegrowth rate of the P. linearis gametophytein glass dishes were: photoperiod, temperature, agein culture, photosynthetic photon flux (PPF), salinityand water dynamics. Maximal growth occurred underdaylength of 12 h, medium temperature (15–20 °C), low PPF (70–140 μmol photon m-2s-1), ambient salinity (30–40 ppt), 1–3 h ofdaily air exposure, and water velocity of 4 cm s-1.Photosynthesis and respiration rates weredominantly affected by daylength and temperature,while the concentration of pigments was dominantlyaffected by PPF and temperature.These conditions correspond well to the optimalnatural growth environment of this local species andare in agreement with the optimum estimated throughthe recently developed outdoor mathematical growthmodel.
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  • 42
    ISSN: 1573-5125
    Keywords: crayfish ; Procambarus alleni ; growth ; Florida Everglades ; hydroperiod
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Environmental conditions influence crustacean growth by affecting molt intervals and incremental increases in length and weight. In the seasonally-flooded marl prairie wetlands of eastern Everglades National Park, U.S.A., hydropattern exerts considerable influence on aquatic primary productivity, and so may influence the availability of food resources for higher trophic levels. The seasonal hydroperiod has been drastically altered by anthropogenic factors, but the impacts on the aquatic community are not well known. We studied whether differences in growth of crayfish Procambarus alleni could be detected in habitats with different hydroperiods. We first described growth patterns based on incremental increases in length and weight of crayfish on a high protein diet in the laboratory. Regression analyses indicated that growth patterns in males and females were similar. Although the intermolt period increased with age, the proportional increases in length and weight were similar through successive molts. The relationship between length and weight of crayfish was best described by a power equation for allometric growth. We then compared growth curves for crayfish subpopulations from different areas of the marl prairie. In habitats with the longest hydroperiods, crayfish weight-at-size was not significantly different from that in laboratory crayfish on the high protein diet. However, weight gain per unit increase in length in short hydroperiod sites was significantly less than in long hydroperiod sites or in the laboratory. These results indicate that crayfish productivity may be associated with hydroperiod in these stressed wetlands, and this may contribute to observed source-sink population regulation.
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  • 43
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    Aquaculture international 8 (2000), S. 159-168 
    ISSN: 1573-143X
    Keywords: emersion ; growth ; Pecten maximus ; scallop spat ; survival ; transport
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Scallop spat production normally requires transfer between growthsystems. Simulated transport experiments were carried out in April, June,December and February to evaluate effects of transport time on greatscallop (Pecten maximus) spat growth and survival. The spat (1.7–1.8 mm in shell-height and 21–25 µg ash free dry weight [AFDW]) wereheld in moist coffee filters at a temperature of 10 °C for up to 24 h,before being replaced into sieves in rearing tanks at 15 °C. The studyshowed that by increasing air emersion time, survival and growthdecreased. No significant difference in the results between 0 and 4 h of airemersion was found, while the effects after 12 and 24 h differed betweenspat groups. Survival and growth rates showed seasonal differences. Meansurvival was 35–71% in April and 77–99% from June to February. In Junemean growth rates attained were 115–128 µm shell-height and 15–18µg AFDW per day compared with 49–69 µm and 3.8–7.0 µgper day for the other spat groups. Great scallop spat may survive atransfer time of 24 h, but transportation for longer than 12 h is notrecommended if subsequent high survival and growth rates are to beensured.
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  • 44
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    Aquaculture international 8 (2000), S. 207-225 
    ISSN: 1573-143X
    Keywords: growth ; mortality ; Pecten maximus ; suspended culture
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Growth and mortality of the king scallop, Pecten maximus, werecompared when grown in cages and by ear hanging in suspended culturein Fuengirola, Malaga, in southern Spain. Seed (juveniles) used in theexperiment was collected in September 1997 that had settled on collectorsin April-June, of that year. Culture in suspended cages began in January1998 when the seed measured 42.7 (3.3) mm shell height and ended inFebruary 1999. Significantly faster growth was found at a minimum culturedensity (16 scallops/cage) than at two other densities (24 and 36scallops/cage). Depth (1, 5 and 10 m from the bottom) influenced growth,poorest growth occurred closest to the bottom. Under optimum growingconditions, 16 scallops/cage suspended 10 m from the bottom, scallops grewto 10 cm shell length (legal size) by February 1999.In ear hanging culture, ropes were moored in April (51.3 (4.5) mm),June (58.2 (4.5) mm) and November 1998 (64.3 (4.9) mm).Initially, rapid shell growth was observed in all three cultures.Subsequently, the shells became covered with barnacles, Balanus sp.,that possibly caused total mortality of the April culture and led to highmortalities in the two other cultures.
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  • 45
    ISSN: 1573-143X
    Keywords: growth ; silver perch (Bidyanus bidyanus) ; size grading ; size variation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract This study presents two experiments addressing growth and size variation in fingerling silver perch (Bidyanus bidyanus. In the first experiment, fish close to mean population size were raised either in the presence or absence of five larger fish for 60 days. Mean specific growth rate (SGR) and increases in the coefficient of variation and skewness were lower in the presence of larger fish, indicating a negative effect of large fish on the growth of smaller ones. In the second experiment, fingerlings were graded into groups smaller and larger than the median size of the population and raised in size-sorted groups of 60 large or small fish and mixed groups of 30 fish of each size category, for 60 days. There was no difference in mean SGR among groups, nor between the mixed group ad the weighted mean of the small and large groups. Biomass gain was higher in the mixed groups than in the weighted small-and-large groups, probably due to a slightly lower survival in the groups comprised of large fish. The fact that the effect of large fingerlings on the growth of smaller ones was evident in the first, but not the second, experiment may be attributed to higher size disparity between large and small fingerlings in the first experiment.
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  • 46
    ISSN: 1573-143X
    Keywords: 17 β-estradiol ; fish meal ; gonadal development ; growth ; Nile tilapia (Oreochromis niloticus) ; plant proteins
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract This study was conducted to evaluate theeffect of plant protein‐based diets on gonadaldevelopment and plasma 17 β-estradiol (E2) levelin female Nile tilapia Oreochromis niloticus.Fish with a mean body weight of 6.7 (0.1) g were fedfour different diets with the same digestible protein(DP) and digestible energy (DE) containing gradedlevels of a mixture of plant ingredients as partial ortotal replacement of fish meal protein for 20 weeks.The control diet (D0) was based on fish meal, twodiets containing 33% (D33) and 66% (D66) of plantprotein, and one diet containing only plant protein(D100). Fish were sampled at 12 and 20 weeks. Nosignificant differences were found in different stagesof oocyte development and plasma E2 levels betweentilapia fed diets D0 and D100 at 12 weeks. Eight weekslater tilapia fed diet D0 showed a higher (P 〈 0.05)level of E2 than the D100 group. This difference andthe reduced proportion of vitellogenic and matureoocytes demonstrated that diets containing only plantprotein are less efficient in terms of tilapia growthand consequently ovarian development.
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  • 47
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    Aquaculture international 8 (2000), S. 455-461 
    ISSN: 1573-143X
    Keywords: floating cages ; growth ; pink dentex (Dentex gibbosus) ; sparids (Dentex dentex, Sparus aurata, Spondyliosoma cantharus, Diplodus puntazzo)
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
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  • 48
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    Aquaculture international 7 (2000), S. 369-382 
    ISSN: 1573-143X
    Keywords: growth ; haddock (Melanogrammus aeglefinus) ; larvae ; light intensity ; photperiod ; tank colour
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract In two separate experiments, haddock (Melanogrammus aeglefinus) larvae were raised under different photoperiods (24L : 0D or 15L : 9D), or different combinations of tank colour (black or white) and light intensity (1.1 μmol s−1 m−2 or 18 μmol s−1 m−2). Growth (0.8% day−1 in standard length; 2.9% day−1 in body area) and survival (2%) were not significantly different between photoperiod treatments after 35 days. Larval survival was greater in white versus black tanks after 41 days (2% versus l%, respectively). Growth of larvae was impaired in black tanks at low (1.1 μmol s−1 m−2) light intensity (0.8% day−1 in standard length and 2.2% day−1 in body area versus 1.1% day 21 in standard length and 3.1% day−1 in body area, for all other treatments). Transmission and reflection of light was low in black tanks at low incident light, and there was very little upwelling light. The resultant poor prey to background contrast probably resulted in larvae being unable to consume sufficient food to sustain a level of growth comparable to that in other treatments.
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  • 49
    ISSN: 1573-143X
    Keywords: bivalve ; fouling ; growth ; Newfoundland ; nursery ; sea scallop (Placopecten magellanicus) ; spat
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Hatchery-reared sea scallop (Placopecten magellanicus) spat weremonitored for growth and recovery in three experiments to determine themost suitable system for nursery culture. In Experiment I, four size classesof nursery-sized spat held at two depths from October to July exhibiteddeclining growth rates over the winter period and increased growth ratesin the spring. Overall, season, depth and initial size had a significantinfluence on the absolute and specific growth rates of scallops. Recovery,defined as number of scallops remaining after mortality and loss of spatthrough gear mesh, was influenced by season and initial size, but notdepth. Scallops in the 3.0 mm+ size class had higher growth rates andrecovery than those in the 1.4–1.6 mm, 1.7–1.9 mm and 2.0–2.9 mm sizeclasses. In Experiment II, two gear types containing similar size spat werecompared. Growth rates were significantly higher in 3.0 mm pearl nets thanin 3.0 mm collector bags, although recovery was similar between the twoequipment types. Experiment III, two stocking densities of nursery-sizedspat were compared in collector bags. Neither growth rate nor recoverywere significantly different for the two densities (2600 and 5200spat/collector bag) tested. Overall, these studies indicated that importantparameters for optimizing the growth and recovery of scallops in a farm-based nursery system include season, initial spat size, deployment depthand gear type.
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  • 50
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    Aquaculture international 8 (2000), S. 381-389 
    ISSN: 1573-143X
    Keywords: Arctic charr (Salvelinus alpinus) ; growth ; low temperature ; stocking density ; time restricted feeding,/kwd〉
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The effects of time restricted feeding, possibility of bottomfeeding and stocking density on the growth of Arctic charr(Salvelinus alpinus L.) were examined in fish held at lowtemperature (〈2 °C). Fish fed for a restricted time (1 h) hadsignificantly (p 〈 0.05) lower specific growth rate (0.15 vs0.32% per day) than those fed the same ration over an extendedtime period (12 h). Increasing stocking densities had a positive andsignificant effect (p 〈 0.05) on growth with SGR increasing from 0.27to 0.52% per day at 2–30 kg m-3. Fish withaccess to feed on the tank floor had a significantly higher (p 〈0.05) growth rate (0.3 vs 0.13% per day) than those without thepossibility to feed from the bottom. When fish were held underconditions without access to the bottom a doubling of the feed rationdid not result in a significant (p 〉 0.05) increase in growth rate(0.13 vs 0.12%percnt; per day).
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  • 51
    ISSN: 1573-5036
    Keywords: growth ; inflow ; magnesium ; potassium ; (Prunus)
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The effects (and interaction) of two solution concentrations of Mg (50, 500, μM) and two of K (250, 4250 μM) on the growth of micropropagated plants of “F. 12/1” and “Colt” were investigated using a flowing solution culture system. Magnesium inflow and growth of “Colt” and “F. 12/1” were inhibited to a similar extent by an increased concentration of K in the nutrient solution. However, the consequences of this inhibition were different. Reduced inflow of Mg in “F. 12/1” caused Mg deficiency symptoms at high and low concentrations of K, whereas this only occurred with a combination of high K concentration and low Mg concentration in “Colt”. The distribution of dry matter within the plant was significant in determining susceptibility to Mg deficiency. Since “F. 12/1” has a smaller root:shoot ratio than Colt it is unable to sustain the same concentration of Mg in leaves as “Colt” irrespective of external K concentration. The molar ratio of K:Mg in soil solutions should remain 〈8.5:1 in order to ensure maximum growth of “F. 12/1” and “Colt”.
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  • 52
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    Plant and soil 219 (2000), S. 177-185 
    ISSN: 1573-5036
    Keywords: alfalfa ; growth ; Medicago sativa L. ; nitrogen
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The dynamics of biomass and N accumulation following defoliation of alfalfa and the application of N fertilization has rarely been studied under field conditions, particularly in the seeding year. Our objectives were to determine the effect of N fertilization on the dynamics of biomass and N accumulation during the first regrowth of alfalfa in the seeding year, and to determine if a model describing critical N concentration developed for established stands could be used in the seeding year. In two separate experiments conducted in 1992 and 1993, the biomass and N accumulation of alfalfa grown with three N rates (0, 40 and 80 kg N ha-1) were determined weekly. Maximum shoot growth was reached with 40 kg N ha-1 in 1992, and maximum shoot growth was not reached with the highest N fertilization rate in 1993. Nitrogen fixation, root N reserves and soil inorganic N uptake when no N was applied were, therefore, not sufficient to ensure non-limiting N conditions, particularly when growth rates were the highest between 14 to 21 d after defoliation. Nitrogen fertilization increased shoot biomass accumulation in the first 21 d of regrowth, biomass partitioning to the shoots and shoot and taproot N concentrations. The model parameters of critical N concentration developed by Lemaire et al. (1985) for established stands of alfalfa were not adequate in the seeding year. The N requirements per unit of shoot biomass produced are greater in the seeding year than on established stands, and this was attributed to a greater proportion of leaves in the seeding year.
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  • 53
    ISSN: 1573-5036
    Keywords: growth ; light intensity ; Phaseolus vulgaris L. ; phosphorus–zinc interaction ; photosynthesis ; yield ; zinc deficiency
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Two experiments were conducted in a factorial combination of three Zn levels (0, 10 and 40 mg Zn kg-1 soil) and two P levels (0 and 200 mg P kg-1 soil). Experiment 1 was carried out during winter in a heated glasshouse, and experiment 2 during summer under a rain shelter. Plants of dwarf bean (Phaseolus vulgaris L., cv. Borlotto nano) were grown in pots filled with sandy soil. In both experiments, leaf Zn concentration was reduced by the addition of P to plants grown at low Zn supply. However, leaf Zn concentration lower than the critical level was observed only during experiment 2, and the main effects of low Zn were reductions of internode length, light use efficiency and maximum photosynthetic rate. In plants with leaf Zn concentration lower than the critical level, saturating irradiance levels fell from ∼1000 μmol m-2 s-1 PPFD to ∼300–400 μmol m-2 s-1 PPFD. Reduction of net photosynthesis was observed from the beginning of flowering and led to decreased seed production.
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  • 54
    ISSN: 1573-5036
    Keywords: AM fungi ; colonisation ; growth ; mineral nutrition ; red raspberry
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Growth, development and nutrient status of micropropagated Rubus idaeus cv. Glen Prosen in response to inoculation with nine species of arbuscular mycorrhizal (AM) fungi from three different genera was investigated. The nine species of AM fungi included, Glomus clarum, G. etunicatum, G. intraradices, Gigaspora rosea, Gi. gigantea, Gi. margarita, Scutellospora calospora, S. heterogama and S. persica. Plant responses to AM fungi varied from growth enhancement to growth depression. Depressive growth effects were specific to Gigaspora species. Furthermore, particular species of AM fungi had unique effects on the mineral status of the raspberry plants. Importance of isolate selection for inoculation of micropropagated raspberry plants is discussed.
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  • 55
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    Plant growth regulation 32 (2000), S. 77-81 
    ISSN: 1573-5087
    Keywords: bioenergetics ; growth ; nutrition ; nutrient deficiency ; plant stress ; respiration ; root ; root:shoot ratio ; root respiration ; source-sink relationship ; stress ; stress physiology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract Root respiration of kohlrabi (Brassica oleraceavar. gongylodes) was measured non-destructivelyin vivo by infrared gas analysis of completeroot systems, using potted plants in sand culture andnutrient solutions, for six weeks under (a) nutrientsufficiency, (b) deficiency of all mineral nutrients,(c) potassium deficiency or (d) phosphorus deficiency.This was to study the adaptation to nutrient stress interms of changes in root growth, root respiration,assimilate allocation and energy requirement fornutrient uptake. Both deficiencies of phosphorus andpotassium increased the root:shoot-ratio. This wasattributed to the plants transferring a largerrelative proportion of assimilates to the roots thanto the shoots relative to nutrient-sufficient plants.Roots of nutrient-sufficient kohlrabi respired 1.7 or7.7 mg CO2 h−1 per g fresh or dry matter, respectively. However, potassiumdeficiency enhanced root respiration to 2.4 mgCO2 h−1 or 12.2 mg CO2 h−1 on a per g fresh or dry weight basis respectively. This originated from an additional2.6 mg glucose g−1 dry matter h−1 allocated to the roots and provided 50 Joule additional energy(150 versus 100 Joule g−1 dry matter h−1)which may become available for the proposedK+:H+ symporter for potassium uptake.
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  • 56
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    Plant growth regulation 30 (2000), S. 163-170 
    ISSN: 1573-5087
    Keywords: ATPase ; bioenergetics ; Fragaria ananassa Duch ; growth ; ion transport ; photosynthesis ; proton transport ; respiration ; source-sink ; strawberry
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The biochemical, physiological and anatomical properties of strawberry (Fragaria ananassa Duch.) cv. 'Cambridge Favourite' stolons were studied during growth. ATPase activity was measured, in microsomal and plasma membrane fractions, along with chlorophyll determination, in-situ photosynthesis measurements and scanning electron microscopy (SEM) and X-ray microanalysis of stolon cross-sections. Potassium-stimulated ATPase activity and proton-pumping, both together indicating the presence of plasma membrane ATPase, was greatest in the stolon tip, the tissue with the fastest growth and respiratory activity. The enzyme activity and respiration gradient from the tip of the stolon to the base was concomitant with xylem development which was more differentiated in the base than in the tip. These cross-sections also showed 30% greater amounts of calcium and potassium of the cryo-preserved basal part relative to the stolon tip. This gradient existed independent of the presence of daughter plants. A hypothesis is presented which suggests that for the long-distance longitudinal transport of nutrients this gradient between stolon tip and base is likely to be involved in stolon growth.
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  • 57
    ISSN: 1573-5117
    Keywords: growth ; secondary production ; Cerastoderma edule ; stable isotope ratios ; microphytobenthos ; intertidal ; Marennes-Oléron Bay
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The contribution of natural food sources to the growth and secondary production of the suspension feeding bivalve Cerastoderma edule (L.) was estimated under field conditions in the Marennes-Oléron Bay (Atlantic coast, France). Monthly estimates of abundance, biomass and cockle growth were combined with seasonal analyses of δ13C and δ15N ratios of juvenile and adult cockles, together with their potential food sources [i.e. suspended particulate organic matter (POM), microphytobenthos, macroalgae and seagrass] sampled at mid-tide level in a muddy sandflat. Adult cockles grew mainly in spring, whereas juveniles grew in summer and autumn, following spat recruitment in early summer. Total annual production and elimination of cockles were estimated to be 32.5 and 34.7 g AFDW m−2 yr−1. Relative contributions of each year class to production were ca 40, 41, 11 and 6% for 0-group, 1-, 3- and 4-yr-old cockles in 1995, respectively. Quantitative assessment of proportions of food sources to the annual secondary production of cockles was obtained by using a simple carbon isotope-mixing model with microphytobenthos (δ13C = −16.0±0.6‰) and POM (δ13C = −22.2±1.1‰) as end-members. On average, more than 70% of the total annual cockle production originated from microphytobenthos, with a much higher contribution for the 0-group (88%) than for adult cockles (60%). The between-age difference was induced mainly by changes in the availability of food resources (benthic versus planktonic) during the non-synchronous growing seasons of juvenile and adult cockles.
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    Environmental biology of fishes 58 (2000), S. 439-446 
    ISSN: 1573-5133
    Keywords: competition ; substrate ; habitat preferences ; growth ; foraging efficiency
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The substrate preferences, growth rates and foraging efficiency of two small benthic fish species, juvenile burbot, Lota lota, and stone loach, Barbatula barbatula, were compared in combined outdoor mesocosm and indoor laboratory experiments. Both species preferred the same stony substrate when alone, but significant differences in habitat selection were found between the two species under food deprivation and competition conditions. In burbot, preference for the stony habitat was reinforced under food-deprivation conditions and became even stronger when a potential competitor, the stone loach, was present. In contrast, stone loach switched to the gravel substrate when either starving or in the presence of a heterospecific competitor. Growth rates and foraging efficiency of burbot were significantly highest in the stony substrate and decreased with finer substrates. In stone loach, neither growth rates nor foraging efficiency were significantly different among the different substrates. The results provide an example of habitat partitioning by means of different competition styles, with a stenoecious, dominant style of the burbot and an euryoecious, evasive style of stone loach allowing coexistence of two sympatric fish species by graded interactions at an individual level in the littoral zone of a large lake.
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  • 59
    ISSN: 1573-5117
    Keywords: activity patterns ; foraging behaviour ; growth ; oxygen content ; predation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Field and laboratory experiments were set up to obtain data on the reasons for different habitat selection of Enallagma cyathigerum and Platycnemis pennipes. (1) Rearing of larvae in two different ponds showed that while P. pennipes was not able to survive conditions of low oxygen content, 50% of the E. cyathigerum larvae survived. (2) In field predation experiments with sticklebacks and dragonflies as predators, we found that E. cyathigerum suffered highest predation by the fish. In P. pennipes, mortality was highest with Anax imperator. (3) Experiments regarding larval behaviour showed that E. cyathigerum was generally more active and had higher foraging success than P. pennipes. Both species reduced activity in the presence of fish, but E. cyathigerum did so to a minor extent. In contrast to P. pennipes, E. cyathigerum showed escaping behaviour. (4) In the laboratory, the growth of E. cyathigerum was faster than that of P. pennipes.
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  • 60
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    Hydrobiologia 438 (2000), S. 237-244 
    ISSN: 1573-5117
    Keywords: Sargochromis codringtonii ; growth ; reproduction ; Lake Kariba
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Age determination of Sargochromis codringtonii was undertaken using scalimetry. Marginal scale analysis, used for age validation, indicated that annulus formation occurred in October and November. The possible reasons for this are discussed. S. codringtonii is relatively long-lived, surviving up to 8 years. It is sexually dimorphic with males growing at a faster rate and attaining a larger size than females. It is suggested that the mouth brooding habits of the female fish is one possible reason for sexual dimorphism. A higher total mortality rate (Z) was recorded for females as compared to males. S. codringtonii reproduces throughout the year, but with a peak during the hot wet months. The right ovary is atrophied and the possible significance of this is discussed.
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  • 61
    ISSN: 1573-5117
    Keywords: Chironomus tentans ; culture ; toxicity testing ; growth ; emergence ; density
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The midge Chironomus tentans Fabricius is a commonly used freshwater invertebrate in sediment toxicity tests. Rigorous laboratory culturing techniques are needed to provide organisms of uniform quality and known age for use in testing and for the continuation of the culture itself. This study was conducted to determine the effect of initial culture stocking density on: (1) post-hatch (larval) dry weight, body length and head-capsule width at 10 and 20 days; (2) time to emergence; (3) number and sex of emergent adults; (4) number of larvae and pupae at test termination (day 42 post hatch); and (5) adult dry weight. Three egg stocking densities were used 690 (1.1 eggs cm−2), 1043 (1.7 eggs cm−2) and 1463 (2.4 eggs cm−2). Mean weight of larvae at 10 days in high density tanks (0.13 mg/organism) was significantly higher (P=0.003) than both the medium and low density tanks (0.10 and 0.09 mg/organism, respectively). No significant differences between the three stocking densities were observed for the body length or head-capsule width at either 10 or 20 days post-hatch. Although not statistically significant, larval dry weight decreased with increased stocking density at day 20. A significantly (P=0.02) greater number of females (173±28) emerged from the low stocking density compared to both the medium and high stocking densities (123±45 and 118±54, respectively). Peak adult emergence for the low and medium stocking densities occurred between days 22 and 25 post-hatch, whereas peak adult emergence occurred between days 30 and 33 for the high stocking density. Survival relative to the initial number of eggs stocked was significantly greater (P=0.007) in the low density treatment compared to that in either the medium or the high density treatments. Mean adult weight exhibited an inverse relationship with initial stocking densities. At test end, there was not a significant difference in the mean number of organisms surviving and emerging in the three density levels. The central tendency for number of organisms surviving for all three treatments was 504 organisms per tank (0.82 organisms cm−2). The results of this experiment suggest that an optimal egg stocking density of 1.0 egg cm−2 (∼600 eggs/tank) be used with the feeding rate identified. This would ensure uniform larvae at the appropriate developmental stage (2nd–3rd instar) needed for toxicological research/testing (e.g. 10 days post-hatch), as well as producing sufficient emergence of males and females for future culture establishment.
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  • 62
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    Environmental biology of fishes 59 (2000), S. 91-97 
    ISSN: 1573-5133
    Keywords: age ; growth ; reproduction ; mortality ; fish
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Atherina presbyter is a common fish off the Canary Islands. Age, growth, reproduction, and mortality of the species are studied based on sampling carried out from July 1995 to June 1996. The parameters of the total length–total weight relationship are: a=0.004521, and b=3.0771. Otoliths age readings indicate that the sampled population consists of four age groups (0–III years). The von Bertalanffy growth parameters for all individuals are: L∞=122 mm total length, k=0.79 year−1, and t0=−0.21 years. Individuals grow quickly in their immature first year, attaining approximately 60% of their maximum length. After the first year, the annual growth rate drops rapidly, because the energy is probably diverted to reproduction. It is a gonochoristic species with no evidence of sexual dimorphism. The gonad is present as a single diffuse testis in males and as a single discrete ovary in females. The overall ratio of males to females is not significantly different from 1:1. The reproductive period of the species is protacted (February to June). The peak of the reproductive effort occurs in April–May. The size at first maturity is 68 mm. The population is being heavily exploited.
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  • 63
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    Environmental biology of fishes 59 (2000), S. 199-209 
    ISSN: 1573-5133
    Keywords: fish behavior ; schooling ; size disparity ; density ; growth ; food availability
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Group membership can confer both advantages and disadvantages to growth in juvenile fishes. The balance between costs and benefits of social interactions can shift depending on such factors as the composition of the group (density and size disparity) and the availability of food. We examined the effect of these factors on absolute growth and growth depensation in juvenile sablefish, Anoplopoma fimbria. Increasing density and increasing size disparity had little influence on absolute growth rates of juvenile sablefish and the effects of these social factors were not modified by ration level. In experiments testing density effects, absolute growth did not differ among groups of 1, 3, or 10 fish held at high rations, but at low rations single fish exhibited a different pattern of size-dependent growth compared to fish in groups. In experiments testing disparity effects, absolute growth did not differ between groups with an even size distribution and groups with a mixed size distribution. The relative size of an individual within a group, i.e., small, medium, or large, also did not modify growth, despite evidence of higher chasing behavior in mixed size distributions. Although the growth of small fish was not diminished in the presence of large fish, negative impacts of size disparity were expressed in high levels of cannibalism, which occurred in 42% of groups with a mixed size distribution. Significant growth depensation over time occurred in the density experiment, but not in the size disparity experiment, possibly due to the shorter duration of the latter experiment. We suggest that growth depensation was generated by individual variability in growth capacity rather than social effects on growth rates. Schooling behavior, measured by group cohesion indices, increased with fish size and was higher in groups with an even vs. a mixed size distribution. These results for sablefish are consistent with other schooling species in which growth variability is determined by exploitative competition and/or genetic variability in growth capacity rather than interference competition.
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  • 64
    ISSN: 1573-5141
    Keywords: behavior ; grass shrimp ; growth ; longevity ; mummichog ; neurotransmitter ; predator ; prey ; serotonin ; top-down
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Behavior can demonstrate linkages ofcontaminant effects at different levels oforganization from the biochemical/cellular to theorganism, population, and community levels.Mummichogs, Fundulus heteroclitus, from acontaminated area were previously found to havereduced condition, growth, and longevity, comparedwith conspecifics from clean sites. Thispopulation-level observation may be due to theirimpaired predator/prey behavior, which is associatedwith altered levels of serotonin in their brains. Theyare slow, less able to capture live food, and eat muchnon-nutritious detritus, which may contribute to theirdecreased growth and condition. They are also lessable to avoid being captured by blue crabs, which canalso contribute to their reduced life span andsize-structure. Grass shrimp, Palaemonetespugio, a major prey species, are not behaviorallyimpaired at this polluted site, but are more numerousand larger in size than at the reference site,probably due to reduced mummichog predation.
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  • 65
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    Fish physiology and biochemistry 22 (2000), S. 297-302 
    ISSN: 1573-5168
    Keywords: Carnitine ; diet ; growth ; hybrid striped bass ; lipid ; Morone ; nutrition
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A study was undertaken to investigate the effects of graded dietary levels and different types of carnitine on hybrid striped bass (Morone chrysops ♀ × M. saxatilis ♂ %) fed different levels of lipid. An incomplete factorial design was utilized in which diets containing lipid at either 5 or 10% were supplemented with l-carnitine at 0, 500, or 1000 mg kg−1 diet, dl-carnitine at 1000 mg kg−1 diet, or carnitine chloride to provide 1000 mg carnitine kg−1 diet. Juvenile hybrid striped bass (3.3 g fish−1) were stocked into individual 38-l aquaria connected as a brackish water (6‰), recirculating system and fed each diet in triplicate for 9 weeks. Supplementation of the diet with 1000 mg carnitine kg−1 increased muscle carnitine from 35.5 to 47.7 μg g−1 tissue. Carnitine supplementation did not result in increased weight gain regardless of carnitine level or type; however, weight gain showed a significant (p〈0.05) response to dietary lipid with fish fed diets containing 10% lipid growing 34% more than fish fed diets with 5% lipid. The hepatosomatic index also was unaffected by diet, but the intraperitoneal fat (IPF) ratio was significantly elevated (5.1 vs 3.2%) in fish fed diets with 10% lipid compared to those fed diets with 5% lipid. Fish fed diets containing 1000 mg carnitine kg−1 had increased IPF ratio values at 4.7% compared to 3.9% for fish fed the basal diet. Liver lipid also was responsive to dietary treatment, increasing from 6.7 to 8.8% of wet weight as dietary lipid increased from 5 to 10%. The relative quantities of triglycerides, free fatty acids and phospholipids in muscle and liver were not influenced by carnitine level, carnitine type or dietary lipid level. Supplementation of carnitine does not appear to be beneficial to hybrid striped bass based on either growth performance or body composition.
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    Plant molecular biology 43 (2000), S. 569-581 
    ISSN: 1573-5028
    Keywords: cell cycle cell differentiation ; cell proliferation ; growth ; shoot apical meristem
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The shoot apical meristem is a group of rapidly dividing cells that generate all aerial parts of the plant. It is a highly organised structure, which can be divided into functionally distinct domains, characterised by specific proliferation rates of the individual cells. Genetic studies have enabled the identification of regulators of meristem function. These factors are involved in the formation and maintenance of the meristem, as well as in the formation of the primordia. Somehow, they must also govern cell proliferation rates within the shoot apex. Possible links between meristem regulators and the cell cycle machinery will be discussed. In order to analyse the role of cell proliferation in development, cell cycle gene expression has been perturbed using transgenic approaches and mutation. The effect of these alterations on growth and development at the shoot apex will be presented. Together, these studies give a first insight into the regulatory networks controlling the cell cycle and into the significance of cell proliferation in plant development.
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    Fish physiology and biochemistry 22 (2000), S. 119-123 
    ISSN: 1573-5168
    Keywords: food competition ; growth ; size matching ; Sparus aurata
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The effects of food competition and relative size of group members on growth of sea bream, Sparus aurata were tested in groups composed of four fish. Each group consisted of a medium sized test fish (approximately 250 mg) and three fish larger or smaller than the test fish. In the first experiment food competition was addressed directly by contrasting its presence with its absence. Neither food competition nor relative size of school members had a significant effect on growth. However, there was a highly significant interaction between these two main effects. Under conditions of food competition the medium sized test fish had a three times higher specific growth rate when surrounded by small vs large fish. Under conditions of no food competition when the test fish were separated during feeding a similar but reverse pattern occurred due to differences in food conversion efficiency. In the second experiment, the effects of food quantity and relative size of group members were tested. Both main effects and their interaction were highly significant. Under conditions of low food competition (i.e., surplus food supply) fish grew again better when surrounded by larger fish as compared to their growth rate when surrounded by smaller ones. Size matching via a differential growth rate in a schooling species such as sea bream was suggested to increase the confusion effect by reducing the ability of a predator to focus on an odd sized fish. The implications of direct competition for food for the efficient culture of this species are discussed.
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    Fish physiology and biochemistry 22 (2000), S. 255-262 
    ISSN: 1573-5168
    Keywords: CHD3 ; cell culture ; complement ; differential mRNA display polymerase chain reaction ; growth ; nucleolin ; protein kinase ; rainbow trout ; RTG-2 cell ; salinity
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The direct effect of osmolality on growth and mRNA population were investigated in the rainbow trout cell line (RTG-2). These cells can grow in the media of osmolalities ranging from 200 to 600 mosmol kg-1. With two-dimensional electrophoresis, the in vitro translation of poly(A+) RNA isolated from these cells showed osmoresponsive changes in the population of translatable mRNAs. Using differential mRNA display polymerase chain reaction, however, we identified inducible cDNA products in hyper-osmotic and hypo-osmotic media as third component of complement, and as homologues of known genes: an atypical protein kinase regulated by the thyrotropin-dependent mitogenic pathway, nucleolin and CHD3. The remaining cDNAs have no significant homology in GenBank. Northern blots demonstrate that their mRNA levels were induced in hyper-osmotic and hypo-osmotic media, but not by other stresses. The expressed proteins of these mRNAs may be involved directly or indirectly in the adaptation of RTG-2 cells to different osmolalities probably through the osmotic signal transduction and adjustment in cellular metabolism to osmotic stress.
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    Fish physiology and biochemistry 22 (2000), S. 245-254 
    ISSN: 1573-5168
    Keywords: swimming velocity ; thermal tolerance ; Oncorhynchus mykiss ; metabolism ; food consumption ; growth
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Although thermal influences on the physiology of rainbow trout (Oncorhynchus mykiss) have been widely studied, there is little information about the responses of different genetic strains to temperatures. The effects of water temperature (10, 14, 19, 22, and 25 °C) on food consumption rate, growth rate, gross conversion efficiency, resting routine oxygen consumption rate, upper critical thermal tolerance and critical swimming velocity were investigated in juvenile rainbow trout of the Eagle Lake (O. m. aquilarum) subspecies and the Mt. Shasta strain. No strain-related differences in conversion efficiency, oxygen consumption rates, thermal tolerance or swimming performance were observed in 1995 (19, 22, and 25 °C) or 1996 (10, 14, and 19 °C), but Mt. Shasta strain trout grew faster at highest temperatures than did Eagle Lake trout. Food consumption rates, growth rates, conversion efficiency, and oxygen consumption rates declined at the extreme temperatures (10 and 25 °C) in both Eagle Lake and Mt. Shasta trout. Swimming performance was temperature-independent between 10 and 19 °C (overall mean: 5.43 body lengths s−1).
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    Journal of economic growth 5 (2000), S. 5-32 
    ISSN: 1573-7020
    Keywords: inequality ; growth ; Kuznets curve ; Gini coefficient
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract Evidence from a broad panel of countries shows little overall relation between income inequality and rates of growth and investment. For growth, higher inequality tends to retard growth in poor countries and encourage growth in richer places. The Kuznets curve—whereby inequality first increases and later decreases during the process of economic development—emerges as a clear empirical regularity. However, this relation does not explain the bulk of variations in inequality across countries or over time.
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  • 71
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    Journal of economic growth 5 (2000), S. 185-206 
    ISSN: 1573-7020
    Keywords: innovation ; growth ; inequality ; hierarchic demand ; multiple equilibria
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract This article analyzes the impactof inequality on growth when consumers have hierarchic preferencesand technical progress is driven by innovations. With hierarchicpreferences, the poor consume predominantly basic goods, whereasthe rich consume also luxury goods. Inequality has an impacton growth because it affects the level and the dynamics of aninnovator's demand. It is shown that redistribution from veryrich to very poor consumers can be beneficial for growth. Ingeneral, the growth effect depends on the nature of redistribution.Due to a demand externality from R&D activities, multipleequilibria are possible.
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  • 72
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    Journal of economic growth 5 (2000), S. 341-360 
    ISSN: 1573-7020
    Keywords: growth ; investment ; human capital ; financial development
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    Topics: Economics
    Notes: Abstract Thisarticle decomposes the well-documented relationship between financialdevelopment and growth. We examine whether financial developmentaffects growth solely through its contribution to growth in ``primitives'' or factor accumulation rates or whether it alsohas a positive impact on total factor productivity growth. Ourresults suggest that indicators of financial development arecorrelated with both total factor productivity growth and investment.However, the indicators that are correlated with total factorproductivity growth differ from those that encourage investment.In addition, many of the results are sensitive to the inclusionof country fixed effects, which may indicate that the financialdevelopment indicators are proxying for broader country characteristics.
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  • 73
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    Journal of economic growth 5 (2000), S. 361-384 
    ISSN: 1573-7020
    Keywords: capital ; growth ; productivity ; public sector
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract The cost of public investment is not the increment to the value ofpublic capital. Unlike with private investors, there is no plausiblebehavioral model in which every dollar that the public sectorspends as ``investment'' creates economically valuable ``capital.''While this simple analytic point is obvious, it has so far beenuniformly ignored in the empirical literature on economic growth,which uses—at best—cumulated, depreciated, investmenteffort (CUDIE) as a proxy for capital stocks. However, particularlyfor developing countries the difference between investment costand capital value is of first-order empirical importance: governmentinvestment is half of more of total investment, and calculationspresented here suggest that in many countries government investmentspending has created little useful capital. This has implicationsin three broad areas. First, none of the existing empirical estimatesof the impact of public spending has identified the productivityof public capital. Even where public capital has a potentiallylarge contribution to production, public-investment spendingmay have had a low impact. Second, it implies that all estimatesof total factor productivity in developing countries are deeplysuspect as there is no way to empirically distinguish betweenlow growth because of investments that create no factors andlow growth due to slow productivity growth. Third, multivariateregressions to date have not adequately controlled for capitalstock growth, which leads to erroneous interpretations of regressioncoefficients.
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  • 74
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    Journal of economic growth 5 (2000), S. 253-275 
    ISSN: 1573-7020
    Keywords: knowledge ; ideas ; growth ; knowledge set ; paradigm ; combination of ideas
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract This articlepresents a model of knowledge, seen as a set of ideas definedin a multidimensional idea space. Knowledge is created throughconvex combinations of older ideas and through paradigm shifts.When normal science has made the knowledge set convex, scientificopportunity is exhausted. Individual countries are endowed withdifferent knowledge sets, which gives rise to idea gaps. Thegrowth of a country's knowledge depends on diffusion from othercountries, on own production, and on the state of its human capitaland institutions. In the long run, economic growth will dependon knowledge growth, but only paradigm shifts can save R&Dfrom diminishing returns
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  • 75
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    Review of industrial organization 17 (2000), S. 229-248 
    ISSN: 1573-7160
    Keywords: Age ; firms' ; growth ; jobs ; size
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract This paper relates recent empiricalresearch on the growth of U.K. companies to the maineconomic theories of firms' growth and to empiricalresults for the U.S.A. Smaller and younger firms havebeen growing more quickly than larger and older firms, thus generating proportionately more new jobs. Theseresults do not support the various theories of staticand dynamic economies of scale. Serial correlation ofgrowth is very low, so success does not persist. Thesystematic tendency for small and younger firms togrow more quickly is the main reason why firm growthis not entirely stochastic.
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  • 76
    ISSN: 1440-1703
    Keywords: growth ; interspecific interactions ; patch structure ; seagrass ; Thailand
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Seagrass beds in South-east Asia sometimes consist of a mosaic of different species in monospecific patches. We examined whether the magnitude of within-patch variation in the seagrass Halophila ovalis is affected by the presence or absence of surrounding vegetation consisting of another seagrass species Thalassia hemprichii in an intertidal flat in Thailand waters. We measured biomass and growth rates of H. ovalis at the edges and centers of two different types of patches: (i) H. ovalis patches adjoining T. hemprichii vegetation (HT patches), and (ii) H. ovalis patches adjoining unvegetated sand flats (HS patches). Furthermore, we examined the possible effects of interspecific interactions on the growth of H. ovalis by experimentally removing adjoining T. hemprichii at the edges of HT patches. The biomass of H. ovalis was greater at the patch centre than the patch edge in both types of patches. For the growth rate of H. ovalis, significant interactions were detected between patch types and positions in patches. The difference in growth was significant and more than 4-fold between edges and centers of the HS patches, whereas the growth was not significantly different between edges and centers of the HT patches. The removal of T. hemprichii did not significantly affect the growth rate of H. ovalis at the edge of the HT patches. These findings demonstrate that the magnitude of within-patch variation in H. ovalis growth is affected by the conditions of adjoining habitats. However, any effects of local competition with T. hemprichii on H. ovalis growth were not evident in this short-term manipulative experiment.
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  • 77
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    Boundary layer meteorology 94 (2000), S. 139-163 
    ISSN: 1573-1472
    Keywords: Turbulence ; Coherent eddies ; Plant canopy ; Plane mixing layer ; Canopy shear scale ; Atmospheric stability
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract An analogy has been established between a plane mixing layer and the atmospheric flow near the top of a vegetation canopy. It is based on a common feature, a strong inflection in the mean velocity profile, responsible for hydrodynamical instabilities that set the pattern for the coherent eddies and determine the turbulence length scales. In an earlier study, this analogy was tested using a small data set from thirteen experiments, all in near-neutral conditions. It provided a good prediction of the streamwise spacing Λw of the dominant canopy eddies (evaluated from time series of vertical velocity) that appears to depend on a shear length scale Ls = U(h)/U'(h), where h is canopy height, U is mean velocity and U' the vertical gradient dU/dz. The present analysis utilizes an extensive data set of approximately 700 thirty-minute runs, from six experiments on two forest sites and a maize crop, with a large range of stability conditions. Λw was estimated for each run using the wavelet transform as an objective, automated detection method. First, the variations of Λw and Ls with atmospheric stability are discussed. Neutral and unstable values exhibit a large scatter whereas in stable conditions both variables decrease with increasing stability. It is subsequently found that Λw is directly related to Ls, in a way close to the neutral prediction Λw /h = 8.1Ls/h.The Strouhal number Str = Ls /Λw is then shown to vary with atmospheric stability, weakly in unstable conditions, more significantly in stable conditions. Altogether these results suggest that, to some extent, the plane mixing-layer analogy can be extended to non-neutral conditions. It is argued that the primary effect of atmospheric stability, at least in stable conditions, is to modify the shear length scale Ls through changes in U(h) and U'(h), which in turn determines the streamwise spacing of the active, coherent motions.
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  • 78
    ISSN: 1573-1472
    Keywords: Field measurements ; Thinned forests ; Turbulence ; Wind tunnel
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract Many forest management methods alterstand density uniformly. The effectsof such a change on the wind andturbulence regimes in the forest arecritical to a number of processes governingthe stability of the stand and itsmicroclimate. We measured wind speed andturbulence statistics with a Dantec tri-axialhot-film probe in model forests of variousdensities (31–333 trees m-2), created byremoving whole trees in a regular pattern in awind tunnel, and compared them with similarmeasurements made with propeller anemometers insimilarly thinned plots (156–625 trees ha-1)within a Sitka spruce stand in Scotland. The results agree well, in general, with measurements made inother such studies with diverse canopy types.The systematic variations with density and verticalleaf-area distribution (which differed betweenwind-tunnel and field trees) in our work can explainmuch of the variability shown in scaled profiles ofbasic turbulence statistics reported in theliterature. The wind tunnel and field results are shown to be in good agreement overalldespite the difference in vertical leaf-areadistribution. Within-canopy and isolated-treedrag coefficients in the wind tunnel showthat tree-scale shelter effects increase astree density increases. The measurements indicatethat turbulence in the canopy is dominated bylarge-scale structures with dimensions of the sameorder as the height of the canopy as found inother studies but suggest that inter-tree spacing also modulates the size of these structures. These structures are associated with the sweeps that dominatemomentum exchange in the canopy and it is thisfact that allows the tri-axial probe to operate sowell despite the relatively narrow range of anglesin which the wind vector is correctly measured. Theratio of streamwise periodicity of these structuresto vorticity thickness varies systematically withtree density in the range 2.7–5.1, which spans theexpected range of 3.5–5 found in a laboratorymixing-layer, suggesting that tree spacing imposes another relevant length scale. This test andothers show that the results are in agreement withthe idea that canopy turbulence resembles that of a mixing layer even though they disagree with, and challenge the linear relationship between, streamwise periodicity andshear length scale presented recently in theliterature. The measurements are also in goodoverall agreement with simple drag models presented recently by other researchers.
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    Boundary layer meteorology 96 (2000), S. 211-255 
    ISSN: 1573-1472
    Keywords: Canopy ; Characteristic eddy ; Empirical-orthogonal-function ; Shear flow ; Turbulence ; Wind tunnel
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract We have conducted an Empirical Orthogonal Functionanalysis (EOF) of a three dimensional, 2-pointvelocity covariance field, measured in a wind tunnel. The rate of convergence of the EOF sequence was usedas an objective test for the presence of distinctlarge turbulent structures. We found that in theroughness sublayer (2h 〉 z 〉 0) the sequence convergedmuch more rapidly than in the lower surface layer(6h 〉 z 〉 0), 75% of the total velocity variancebeing captured by the first three of 42 eigenmodes;h is the canopy height. The analysis was extended to three dimensions, whereover 50% of the variance and most of the spatialstructure of the covariance fields were captured by aneven smaller fraction of the total number ofeigenmodes. With some relatively weak additionalassumptions we were able to construct the velocityfield of a ‘characteristic eddy’ or large coherentstructure. This consisted of a pair ofcounter-rotating streamwise vortices centred above thecanopy. The sense of rotation of the vortex pair wasopposite to that found in the wall region of boundarylayers but matched that found in plane mixing layers. A strong gust or sweep motion generated between thevortices was responsible for most of the shear stresscarried by the large eddies. The region of significanttransport of streamwise momentum by the characteristiceddy is much smaller than the region of coherence of theeddy's velocity field.
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  • 80
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    Boundary layer meteorology 97 (2000), S. 331-357 
    ISSN: 1573-1472
    Keywords: Aircraft measurements ; Entrainment ; Stratocumulus ; Structure functions ; Turbulence
    Source: Springer Online Journal Archives 1860-2000
    Topics: Geosciences , Physics
    Notes: Abstract During the Dynamics and Chemistry of the MarineStratocumulus (DYCOMS) experiment in July–August 1985, the NCAR Electra aircraft flew a series of flight legs just at the top of the marinestratocumulus cloud decks that cap the mixed layer off the coast of southernCalifornia. Because of the corrugated structure of the cloud-top, the aircraft, which was flown at a nearly constant level and adjusted only to maintain its altitude at the average cloud-top height, was alternately within and above the clouds – roughly half the time in each domain. These legs were used to examine the structure of the cloud-top by compositing the segments on either side of the cloud/clear-air interface, which was identified by the transitions of liquid water measured by the Forward Scattering Spectrometer Probe (either increasing or decreasing) through a threshold of 0.04 × 10-3 kg m-3.An equivalent vertical distance (EVD) from the cloud-top was obtained from the horizontal flight legs by estimating the average slope of the cloud-top from the cloud-top radiation temperature. The results show that a near discontinuity occurs in variables across cloud top over an EVD of 0.3 m, but that above this, the air has already been modified by boundary-layer air. Thus, cloud-top is not the limit of mixing of boundary-layer air. This mixing may extend to tens of metres or more. The bulk Richardson number in the vicinity of cloud-top increases from near zero within the cloud to about 1.2 at an EVD of 3–6 m above cloud. Fluctuations of the three velocity components within cloud are nearly equal; above cloud the vertical component structure function is about half the horizontal components. The scalar structure functions are about an order of magnitude higher above cloud than in cloud. The structure parameters of temperature and humidity measured just below cloud-top agree reasonably well with predicted values based on a previously-developed model for the clear convective boundary layer. Above cloud, the scalar structure parameters are much larger, but their interpretation is questionable, since this region does notcontain isotropic turbulence.
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    World journal of microbiology and biotechnology 16 (2000), S. 297-301 
    ISSN: 1573-0972
    Keywords: Anaerobic bacteria ; growth ; protease ; psychrotrophs ; temperature ; volatile fatty acids
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Process Engineering, Biotechnology, Nutrition Technology
    Notes: Abstract Five anaerobic proteolytic bacteria were isolated from water bodies of Leh, India, where the ambient temperature varies from −25 to 25 °C. Isolates showed growth at all temperatures ranging from 5 to 37 °C except SPL-4 and SPL-5 which showed no growth at 5 °C. The cultures could grow and produce proteases on various protein substrates and the yield varied with the substrates. Two of the cultures showed the presence of spores. Acetate was the dominant VFA during hydrolysis of protein substrates.
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    Environmental and resource economics 17 (2000), S. 353-373 
    ISSN: 1573-1502
    Keywords: environmental transition ; growth ; Kuznets ; pollution ; transboundary externalities
    Source: Springer Online Journal Archives 1860-2000
    Topics: Energy, Environment Protection, Nuclear Power Engineering , Economics
    Notes: Abstract The Environmental Kuznets Curve (EKC) is a hypothesis which implies that it is possible to “grow out of environmental degradation”. Most theoretical models of the EKC relation have not accounted for transboundary and intergenerational externalities nor have empirical studies provided evidence that validates an inverted U shaped relation between environmental degradation and economic growth for pollution problems where the effects are far-displaced or are long-delayed. This paper integrates the theory of transboundary externalities into the most common theoretical framework applied to the EKC hypothesis. It shows that where a significant proportion of the environmental impacts of economic activity occurs outside the territories in which those activities take place, the de-linking of growth and environmental degradation is less likely to happen. This proposition is demonstrated by assuming that decisionmakers have a Nash-type non cooperative strategic behavior.
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  • 83
    ISSN: 1573-1561
    Keywords: Nicotiana sylvestris ; nitrogen ; nicotine ; allocation ; growth ; reproduction ; induced defenses ; costs of defense
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Chemistry and Pharmacology
    Notes: Abstract We present the first evidence for a fitness cost of an inducible response that is detectable in a nitrogen (N) currency. Nicotine is an induced defense in Nicotiana sylvestris that can utilize 5–8% of the plant's total N, an investment that apparently cannot be recouped by metabolism. Induced nicotine production is endogenously regulated by jasmonic acid (JA), and we treated leaves with the methyl ester of this wound hormone (MeJA) in quantities (0, 25, 250 μg) known to elicit changes in endogenous JA and subsequent nicotine responses comparable to those elicited by mechanical wounding and herbivory in this species. We grew plants in competition chambers (CCs) in which three same-sized plants could compete for a communal but fixed pool of 15NO3 to quantify the outcome of competition for this fitness-limiting resource that is used both in defense and seed production. Competition profoundly increased all measures of growth and reproductive performance measured per milligram of N acquired. While plants acquired all the N supplied to them in the hydroponic solution, plants grown in CCs (as compared those grown in individual chambers—ICs) retained more of this N and produced more biomass, had larger nicotine contents, allocated less of their N to nicotine, produced larger floral stalks with more flowers, aborted fewer flowers, matured more capsules, and produced a greater mass of seed. Plants grown in ICs produced heavier seed, but this difference did not translate into a difference in seed viability. MeJA treatment increased nicotine concentrations in proportion to the amount applied and significantly reduced growth (13–23%) and reproductive (31–44%) performance for plants grown with uninduced competitors, reflecting a large opportunity cost of induction. The effects of MeJA treatment on growth and reproduction were significantly less pronounced for plants grown in ICs. MeJA treatment significantly reduced the ability of plants to compete for [15N]KNO3 (reducing uptake by 9.5% and 23.7% for 25- and 250-μg MeJA-treated plants, respectively); no reductions in N acquisition were found in IC grown plants treated with MeJA. This impairment of competitive ability could account for 41–47% of the jasmonate-induced reductions in biomass by the day 15 harvest and 12–20% of the reductions in seed set and, in addition, created by "opportunity benefit" for neighboring uninduced plants, which grew larger, aborted fewer flowers, and matured more seed (a 28% increase) than did uninduced plants competing with similarly uninduced plants. Competition dramatically increased plant growth and reproductive performance, and MeJA treatment of these high-performing plants significantly reduced their competitive ability, which translated into opportunity costs for induced plants and opportunity benefits for neighboring uninduced plants. Induced plants minimized these fitness costs by reducing their use of recently acquired N for nicotine biosynthesis when growing with competitors. MeJA treatments also altered stalk length, flower production, flower abortion, and allocation to seed mass. In spite of all this plasticity, induced responses incur large fitness costs, costs that could be in part attributed to reductions in competitive ability for N. We conclude that inducibility functions to minimize these costs.
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  • 84
    ISSN: 1435-1536
    Keywords: Key words Drag reduction ; Poly(ethylene oxide) ; Hydrodynamic volume ; Ocean thermal energy conversion ; Rotating disk flow ; Turbulence
    Source: Springer Online Journal Archives 1860-2000
    Topics: Chemistry and Pharmacology , Mechanical Engineering, Materials Science, Production Engineering, Mining and Metallurgy, Traffic Engineering, Precision Mechanics
    Notes: Abstract  Polymer-induced turbulent drag reduction in a rotating disk apparatus was investigated using nonionic poly(ethylene oxide) (PEO) in a synthetic saline solution with novel application to ocean thermal energy conversion technology. A maximum total (skin friction plus form) drag reduction of 30% was obtained with 50 wppm of PEO with molecular weight 5.0 × 106. The concentration dependence of the percentage drag reduction for the PEO/saline solution system is found to fit Virk's empirical correlation, and a universal correlation for various molecular weights and Reynolds numbers is also presented. Furthermore, hydrodynamic volume fraction was introduced to correlate drag reduction efficiency with molecular parameters in this PEO/saline solution system.
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    Meccanica 35 (2000), S. 1-15 
    ISSN: 1572-9648
    Keywords: Wind ; Aeroelasticity ; Bridge ; Turbulence ; Mechanics of structures
    Source: Springer Online Journal Archives 1860-2000
    Topics: Mechanical Engineering, Materials Science, Production Engineering, Mining and Metallurgy, Traffic Engineering, Precision Mechanics , Physics
    Notes: Abstract Non-stationary aeroelastic wind force processes on a streamlined bridge deck are modelled for use within computations in the range of statistically stationary girder oscillation and for the mathematical investigation of the limit of aeroelastic stability. Time Domain (TD) methods, as the direct integration on a finite-element (FE) model, are applied in structural analyses for taking into consideration both geometric and physical nonlinearities. The FE method requires a continuous form of the force model, which permits an arbitrary discretization in time. The present model is based on continuous step response functions of Wagner-type, taking advantage of modern advances of that theory. Special emphasis is put on the continuity and the existence of equivalent representations in time and frequency domains. A realistic wind load model must include measured force functions and effects from the wind turbulence. For that reason, the force functions are calibrated in wind tunnel tests on section models in grid turbulence and boundary layer flow. The aeroelastic in-phase and out-of-phase forces are identified from highly accurate calibrations experiments on section models up to high frequencies which permits to decrease the time step increment. Turbulence effects on the aeroelastic coefficients are included in mean terms.
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  • 86
    ISSN: 1572-9702
    Keywords: Gallotia ; Lacertidae ; Ophionyssus ; parasitic mites ; ontogeny ; growth ; Canary Islands
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Wild-caught specimens of the lacertid lizard Gallotia galloti eisentrauti from the Canary Island of Tenerife were checked for ectoparasites. The parasitic gamasid mite Ophionyssus galloticolus Fain and Bannert (2000) was very abundant on these lizards. Additionally, parasitism by larvae of two species of Trombiculidae (Prostigmata: Parasitengona) was observed. O. galloticolus was reared in the laboratory on its natural host in order to investigate its life cycle, reproductive biology, and development. The life history of O. galloticolus is documented in detail and compared to literature data of other Ophionyssus species. O. galloticolus was found to be similar to other species of the same genus with respect to the duration of development, the precopulatory association of protonymphs, and the arrhenotokous development of eggs. However, it seems to be more tolerant towards low relative humidity and longer starvation periods than other Ophionyssus species. Evolutionary transformations of the life-history pattern of this genus and other parasitic mites in comparison to its predatory precursors involve a reduction or partial suppression of ontogenetic instars in order to decrease mortality during host-seeking phases, and a compensating increase in growth capacity of the remaining feeding instars facilitated by replacement of sclerites through elastic cuticle or by growth of new cuticle unrelated to a moult (neosomy).
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  • 87
    ISSN: 1573-5087
    Keywords: betaine ; ethanolamine ; growth ; Helianthus annuus L. ; salt stress
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The accumulation of compatible solutes is one of the strategies that plants have developed to tolerate salt stress. Proline and betaine are the main metabolites that accumulate in various species of higher plants in response to salt stress. In Helianthus annuus L., pre-treatment of seeds with ethanolamine led to enhanced seedling tolerance to conditions of saline stress during germination, as evidenced by the greater growth of pretreated seedlings (EAS group) versus untreated seedlings (S group), evaluated through such parameters as length, water and chlorophyll content. During the germination period, a considerable increase was observed in proline levels (up to 300%) in seedlings subjected to saline stress, whereas in the EAS group, the proline increment was much smaller (20%). Starting from the fourth day of germination, betaine levels in seedlings pretreated with ethanolamine and then with water (EAW group) and in EAS showed a significant increase versus C (control) and S seedlings, possibly because such a precursor promotes betaine biosynthesis. This could be responsible for the enhanced growth observed in EAS versus S seedlings, as well as for preventing the decrease in chlorophyll content in the EAS group. The accumulation of betaine seems to correlate with the greater tolerance of these seedlings against stress induced by sodium chloride.
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  • 88
    ISSN: 1573-5036
    Keywords: growth ; ion-uptake ; salinity ; salt tolerance ; sugarcane ; transpiration
    Source: Springer Online Journal Archives 1860-2000
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
    Notes: Abstract The effects of salinity on leaf growth, initiation and senescence, on transpiration rates, on leaf water potential and on uptake and distribution of several ions were studied in two sugarcane cultivars differing in salinity sensitivity. Plants, growing in a growing mixture in pots, were exposed to salinized irrigation water for 68 days, starting 60 days after planting. EC values of the irrigation water were 1.0, 2.0, 4.0, 8.0 and 12 dS/m, obtained by using a mixture of NaCl and CaCl2. Plants were also grown in nutrient solution and were at a similar age when exposed to a salinity level of 3 dS/m for 30 days followed by 6.0 dS/m for an additional 30 days. Two Na:Ca ratios of 18:1 and 1:2 were used for salinization of the nutrient solution. Both leaf dry weight and area decreased with increasing salinity, but in the more salinity tolerant cultivar H69-8235, the decrease was moderate. Salinity hardly reduced average area per leaf in H69-8235, while the number of leaves declined sharply. This decline was caused by enhanced senescence of mature leaves and not by a decreased rate of leaf initiation. In the more sensitive cultivar, H65-7052, leaf area and initiation of new leaves were sharply reduced by salinity while leaf senescence was less affected. Leaf water potential decreased during the early stages of salinity exposure, and the reduction in water potential was larger in H69-8235. Salinity also decreased the rate of transpiration rate but to a lesser extent than leaf development and growth. The accumulation of Cl and Na in the TVD (top visible dewlap) leaf of the tolerant cultivar H69-8235 was greater than in the sensitive cultivar H65-7052. The concentration of Cl in the TVD leaf was more than 10 times that of Na in both cultivars. The concentration of both ions, but not of K, increased during the early stages of salinity exposure and then remained constant. A gradient in concentration of Cl and Na over the plant was found in both cultivars at all salinity levels, and was steepest between the TVD and younger leaves. No specific Na effect on leaf growth or transpiration could be detected. The accumulation of Cl and Na but not of K occurred primarily in the roots rather than in the leaves and stalks.
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  • 89
    ISSN: 1573-5044
    Keywords: carbon source ; grape berries ; in vitro culture ; growth
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Using in vitro culture of isolated small berries of Vitis vinifera L. cv. Sultana, it was possible to study the effect of different carbon sources and sucrose concentration on fruit growth, hexose accumulation and soluble invertase activity during the first stage of berry development by eliminating the source tissue. Berries cultured in vitro lack stage III of berry development which is characterised by massive accumulation of water and sugars, and thereby berries reached only 30% of the weight of those grown in the plant. Sucrose and glucose were both good carbon sources for berry growth, while fructose was not as good. Berry growth, hexose accumulation and invertase activity increased as sucrose concentration increased up to 15% in the medium. Furthermore, the onset of hexose accumulation in cultured berries depended on the concentration of sucrose in the medium, starting earlier at higher concentrations.
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    Hydrobiologia 441 (2000), S. 149-153 
    ISSN: 1573-5117
    Keywords: aquatic hyphomycetes ; temperature and light effect ; growth ; sporulation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Nine isolates of the aquatic hyphomycetes Dactylella aquatica, Flagellospora penicillioides, Flagellospora saccata, Helicomyces sp., Lunulospora curvula, Phalangispora constricta, Tetracladium setigerum, Vermispora cauveriana and Wiesneriomyces laurinus were incubated at different temperatures (5–35 °C) to study their growth on MEA medium. Maximum growth was observed between 20 and 30 °C. Growth rate was highest in Vermispora cauveriana and lowest in Tetracladium setigerum. Colonies on submerged leaves showed maximum spore production at 25 °C. Light was confirmed as a stimulus to sporulation.
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    Hydrobiologia 440 (2000), S. 331-338 
    ISSN: 1573-5117
    Keywords: Bivalvia ; growth ; Arctic ; Clinocardium ciliatum ; Svalbard
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The Svalbard Islands are influenced by warm Atlantic water in the south and west, and cold Arctic water in the east. Ice cover, and hence the location of the highly productive marginal ice zone, varies both intra and interannually. Part of the primary production accumulates on the bottom and is utilized by the benthos. In this study, the annual growth of the cockle Clinocardium ciliatum (Fabricius, 1780) from three sites in Svalbard waters is reported. Moffen, the site in the north (80° 01′ N, 13° 48′ E) is located in the northernmost areas influenced by Atlantic water. The Storfjorden site (77° 10′ N, 20° 09′ E) is situated in cold Arctic water masses, and the Bear Island site (74° 50′ N, 18° 54′ E) is in the Polar front area where Atlantic and Arctic water masses meet. Annual growth of cockles was analysed retrospectively by measuring external growth increments, which gave annual growth records from the 1970s to 1996. Shell height for age for different year classes was highest at the Storfjorden site, and lowest at Bear Island. Periods of high growth occurred at Storfjorden and Bear Island during the 1980s while the beginning of 1990s was characterized by low growth. At Moffen, growth was more variable between single years. Several factors are influencing the growth of C. ciliatum in the Svalbard area and growth cannot be coupled to only one environmental factor like ice cover.
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  • 92
    ISSN: 1573-0778
    Keywords: cDNA ; differentiation ; gene expression ; growth ; humancells ; human tissues ; mitochondrial DNA ; mitochondrial RNA ; polyadenylated RNA
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology , Medicine , Process Engineering, Biotechnology, Nutrition Technology
    Notes: Abstract In the mitochondrion, essential genetic elements for replication and transcription are mostly housed within a shortsegment of its DNA located between tRNAPhe and tRNAPro genes, which is called mitochondrial regulatoryregion (mrr). RNAs are known to be transcribed from mrr, thestructures and the functions of which are yet to be fullycharacterized.We detected ca. 1.3 kb H-strand transcripts of mrr (mrrH-RNAs),and 0.2 kb L-strand transcripts of mrr (mrrL-RNAs) in varioushuman cultured cells and tissues using double stranded mrrDNAprobes. The steady state levels of mrrL-RNAs were generally highin cultured cells, while they varied among tissues. On the otherhand, the levels of mrrH-RNAs varied among tissues and amongcultured cells. A tendency was observed in these cells andtissues that a high level of mrrL-RNA is associated with cellproliferation, and a high level of mrrH-RNA withdifferentiation. Several cDNA clones to 1.3 kb mrrH-RNA were obtained from humanskeletal muscle polyadenylated RNAs. The 5′ terminus of the 1.3 kb RNA was determined to be at nucleotide position 15953 whichis immediately downstream of tRNAThr sequence.Polyadenylation site for most of the clones was demonstrated tobe at nucleotide position 576 which is immediately upstream oftRNAPhe sequence. The longest cDNA insert obtained was 1177 bps long spanning from nucleotide positions 15969 to 576 which could code for a peptide of 76 amino acids. The cDNAs isolatedhere are the first cDNA clones reported to human mrrH-RNAs.These results, together with previous results, furthersubstantiate that polyadenylated mrrH- and mrrL-RNAs are commonly present at varying levels among human tissues andcells. The 3′ end sequences of the cloned mrrH-cDNA provideswith insights into the mechanisms of transcription termination.The cDNA clones will provide tools to further the study of thefunction of mrr RNAs.
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  • 93
    ISSN: 1573-2932
    Keywords: 14C uptake ; ATP ; cadmium ; growth ; microalga ; toxicity ; ultrastructure
    Source: Springer Online Journal Archives 1860-2000
    Topics: Energy, Environment Protection, Nuclear Power Engineering
    Notes: Abstract Cadmium toxicity to the diatom Phaeodactylum tricornutum Bohlin has been studied on the basis of the effect of this metal on growth, ATP content, 14C uptake and cellular ultrastructure. The microalga was exposed to 1, 5, 10, 25, 50, 75 and 100 mg l-1 of cadmium and showed a concentration-dependent inhibition in those physiological parameters. At cadmium concentrations of 5 mg l-1 or higher, a significant effect on growth of P. tricornutum was observed. After 8 hours of exposure to the different cadmium concentrations, the decrease in the ATP content was significant only at cadmium concentrations of 25 mg l-1 or higher. 14C uptake was more susceptible to cadmium than the decrease in the ATP content, as 5 mg l-1 of cadmium caused a severe decrease in this parameter. Cadmium also caused ultrastructural changes in P. tricornutum cells: deposition of cadmium on the surface of cell, increase in the chloroplast size, appearance of electrodense granulations, and reduction in lipid inclusions.
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  • 94
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    Journal of elasticity 59 (2000), S. 175-193 
    ISSN: 1573-2681
    Keywords: biomechanics ; growth ; residual stresses
    Source: Springer Online Journal Archives 1860-2000
    Topics: Mechanical Engineering, Materials Science, Production Engineering, Mining and Metallurgy, Traffic Engineering, Precision Mechanics , Physics
    Notes: Abstract The constitutive functions of soft biological tissues during growth are studied. A growth, treated as addition (often non-uniform) of material points, results in deformation, residual stresses, and evolution of the constitutive functions. A theory based on the concept of equivalent material points is developed with the current configuration taken as the reference. The residual stresses developed in a spherical shell undergoing spherical growths are studied.
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  • 95
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    Biologia plantarum 43 (2000), S. 603-606 
    ISSN: 1573-8264
    Keywords: growth ; Indian mustard ; lead uptake
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The effects of different concentrations of lead nitrate (10−5 to 10−3 M) on root, hypocotyl, and shoot growth of Indian mustard (Brassica juncea L. var. megarrhiza), and the uptake and accumulation of Pb2+ by its roots, hypocotyls, and shoots were investigated. Lead had no significant inhibitory effect on the root growth at concentrations of 10−5 to 10−4 M during the entire treatment, while at 10−3 M, Pb slightly inhibited the root and shoot growth. B. juncea has ability to take up Pb from solutions and accumulate it in its roots, and transport and concentrate it. The Pb contents in the parts of plants treated with 10−3 M Pb were greater than those of untreated plants, by factors of 230 in the roots, 170 in the hypocotyls, and 3 in the shoots.
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  • 96
    Electronic Resource
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    Springer
    Biologia plantarum 43 (2000), S. 441-444 
    ISSN: 1573-8264
    Keywords: abscisic acid ; Cucumis sativus ; growth ; proline ; soluble proteins
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The effect of lead and cadmium on biomass accumulation of cucumber seedlings (Cucumis sativus L.) as well as the contents of abscisic acid (ABA), free proline and soluble proteins in leaves were studied. Seedlings were subjected to lead nitrate or cadmium bromide in low concentrations (1 – 5 µM) for 1, 4 or 7 d, and then to the action of the same substances in high concentrations (500 – 1000 µM). The pretreatments of the seedlings with heavy metals in low concentrations enabled them to tolerate the subsequent high concentrations of cadmium and lead without injury. The plant responses to heavy metal treatment were accompanied by the accumulation of ABA, free proline and soluble proteins in leaf tissues.
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