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  • 1
    Publication Date: 2010-07-01
    Description: A synthesis was carried out to examine Alaska’s boreal forest fire regime. During the 2000s, an average of 767 000 ha·year–1 burned, 50% higher than in any previous decade since the 1940s. Over the past 60 years, there was a decrease in the number of lightning-ignited fires, an increase in extreme lightning-ignited fire events, an increase in human-ignited fires, and a decrease in the number of extreme human-ignited fire events. The fraction of area burned from human-ignited fires fell from 26% for the 1950s and 1960s to 5% for the 1990s and 2000s, a result from the change in fire policy that gave the highest suppression priorities to fire events that occurred near human settlements. The amount of area burned during late-season fires increased over the past two decades. Deeper burning of surface organic layers in black spruce ( Picea mariana (Mill.) BSP) forests occurred during late-growing-season fires and on more well-drained sites. These trends all point to black spruce forests becoming increasingly vulnerable to the combined changes of key characteristics of Alaska’s fire regime, except on poorly drained sites, which are resistant to deep burning. The implications of these fire regime changes to the vulnerability and resilience of Alaska’s boreal forests and land and fire management are discussed.
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  • 2
    Publication Date: 1991-08-01
    Description: In an open-pollinated progeny trial of Pinusradiata D. Don, stem diameter assessments were cross-referenced for 410 families for ages 5, 10, and 17 years from planting. Also cross-referenced were Cyclaneusma needle cast (CYCLA) and wood density (PILO) measured by Pilodyn needle penetration. Estimated narrow-sense heritability for stem diameter declined mildly from 0.34 at age 5 to 0.25 at age 17. Estimated heritability of family means, however, only declined from 0.59 to 0.55. CYCLA and PILO gave, respectively, narrow-sense heritability estimates of 0.32 and 0.40, with repeatabilities of family means of 0.57 and 0.67. The genetic age-age correlations for stem diameter were all positive and somewhat higher than phenotypic (family-mean) age–age correlations. Such correlations indicated comparable or slightly slower rank changes among progeny families than had been reported previously for diameter, basal area, or stem volume in P. radiata and Pinustaeda L., but faster rank changes than the literature reports for tree height. A considerable contribution of CYCLA to rank changes in stem diameter was evident from path coefficients and partial correlations. PILO made no evident contribution to rank changes. Predicted gains for stem diameter at age 17 were almost maximal using year-10 data, while using CYCLA as an auxiliary selection criterion enhanced expected gain, particularly with selection at year 5. Predicted gains for stem diameter, with age–age correlations extrapolated according to the Lambeth relationship, indicated maximal gains per annum with selection at 7–8 years for rotations of 25–30 years.
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  • 3
    Publication Date: 2010-11-01
    Description: This study investigates geographic patterns of genetic variation in aspen (Populus tremuloides Michaux.) spring phenology with the aim of understanding adaptation of populations to climatic risk environments and the practical application of guiding seed transfer. We use a classical common garden experiment to reveal genetic differences among populations from western Canada and Minnesota, and we present a novel method to seamlessly map heat-sum requirements from remotely sensed green-up dates. Both approaches reveal similar geographic patterns: we find low heat-sum requirements in northern and high-elevation aspen populations, allowing them to take full advantage of a short growing season. High heat-sum requirements were found in populations from the central boreal plains of Saskatchewan and Alberta, and populations from Minnesota exhibit moderately low heat-sum requirements for budbreak. Analysis of corresponding climate normal data shows that late budbreak is strongly associated with the driest winter and spring environments, which suggests selection pressures for late budbreak due to both frost and drought risks in early spring. We therefore caution against long-distance seed transfer of Minnesota provenances to the boreal plains of Alberta and Saskatchewan. Although such transfers have been shown to increase tree growth in short-term field tests, this planting material may be susceptible to exceptional spring droughts.
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  • 4
    Publication Date: 2010-12-01
    Description: There is increasing recognition that forestry provides a low cost and robust means of climate change abatement through carbon sequestration and substitution. However, current understanding of forest ecosystem carbon exchange and forest–atmosphere interactions are often inadequately characterized by existing empirical growth models with resulting poor representation for regional extrapolations. In this paper, we describe the parameterisation and independent validation, against both eddy covariance and forest growth experimental data, of a process-oriented model 3PGN to provide assessments of carbon sequestration of Sitka spruce (Picea sitchensis (Bong.) Carrière) plantations across Scotland. In comparison with eddy covariance measurements, the model predicted all of the major annual carbon fluxes, i.e., gross primary production (PG), net ecosystem production (PE), and ecosystem respiration (RE), with biases lower than 10%. At a monthly time step, only PG and PE were accurately estimated, whereas RE was not. At longer time scales (i.e., several decades), the model reliably represented the major patterns of the carbon balance. Soil type was identified as the important factor influencing site productivity; fertilization practices did not alter long-term site nutritional status. The analyses also highlighted the potential impact of carbon loss from carbon-rich soils, which can result in differences between optimal rotation length for carbon sequestration and for timber production.
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  • 5
    Publication Date: 2010-10-01
    Description: Understanding of the natural factors that lead to complex changes in forest ecosystems is limited. Worldwide, there are only a few forests as pristine and isolated as the Sierra de La Laguna in the southernmost range of the arid Baja California, Mexico. Its outstanding trait as a model system is that anthropogenic stressors are notably absent, which facilitates the study of natural ecological processes of the forest because separating human-induced ecological changes from natural ones is not a simple matter. In this study, we sampled sites and defined vegetation units on the basis of dominance of the canopy by the main tree species. We identified three forest types: the pine and encino forests that occupy the higher areas and the roble forest at lower elevations. For each living tree in the sampling plots, we measured height, canopy coverage per tree, diameter at breast height, as well as the amount of deadwood, leaf litter, and abundance of young trees. A succesional competition occurs between Pinus and Quercus sensu lato; we conclude that the encino forest represents a climax condition, the pine type represents an early succesional stage, and the roble forest type is a simple climax community.
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  • 6
    Publication Date: 1991-04-01
    Description: The internal transport of carbon dioxide by water flow has until now been omitted when interpreting measurements of photosynthesis and respiration. Theoretical and empirical analyses of the behaviour of carbon dioxide within a tree show that the consideration of dissolved carbon transportation may be important when estimating the respiration rate, but not so important when considering photosynthetic production.
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  • 7
    Publication Date: 2010-04-01
    Description: Forest structure, as measured by the physical arrangement of trees and their crowns, is a fundamental attribute of forest ecosystems that changes as forests progress through suc;cessional stages. We examined whether LiDAR data could be used to directly assess the successional stage of forests by determining the degree to which the LiDAR data would show the same relative ranking of structural development among sites as would traditional field measurements. We sampled 94 primary and secondary sites (19–93, 223–350, and 600 years old) from three conifer forest zones in western Washington state, USA, in the field and with small-footprint, discrete return LiDAR. Seven sets of LiDAR metrics were tested to measure canopy structure. Ordinations using the of LiDAR 95th percentile height, rumple, and canopy density metrics had the strongest correlations with ordinations using two sets of field metrics (Procrustes R = 0.72 and 0.78) and a combined set of LiDAR and field metrics (Procrustes R = 0.95). These results suggest that LiDAR can accurately characterize forest successional stage where field measurements are not available. This has important implications for enabling basic and applied studies of forest structure at stand to landscape scales.
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  • 8
    Publication Date: 2010-02-01
    Description: Sustainable forest management requires timely, detailed forest inventory data across large areas, which is difficult to obtain via traditional forest inventory techniques. This study evaluated k-nearest neighbor imputation models incorporating LiDAR data to predict tree-level inventory data (individual tree height, diameter at breast height, and species) across a 12 100 ha study area in northeastern Oregon, USA. The primary objective was to provide spatially explicit data to parameterize the Forest Vegetation Simulator, a tree-level forest growth model. The final imputation model utilized LiDAR-derived height measurements and topographic variables to spatially predict tree-level forest inventory data. When compared with an independent data set, the accuracy of forest inventory metrics was high; the root mean square difference of imputed basal area and stem volume estimates were 5 m2·ha–1 and 16 m3·ha–1, respectively. However, the error of imputed forest inventory metrics incorporating small trees (e.g., quadratic mean diameter, tree density) was considerably higher. Forest Vegetation Simulator growth projections based upon imputed forest inventory data follow trends similar to growth projections based upon independent inventory data. This study represents a significant improvement in our capabilities to predict detailed, tree-level forest inventory data across large areas, which could ultimately lead to more informed forest management practices and policies.
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  • 9
    Publication Date: 2010-10-01
    Description: Over the last decades, continuous signs of sugar maple ( Acer saccharum Marsh.) dieback in stands of northeastern North America have promoted the experimentation of corrective measures to restore sugar maple vitality. To verify the hypothesis that K–Mg antagonism may have limited the full response of sugar maple to dolomitic lime application in a previous experiment (CaMg(CO3)2, 12% Mg), two Ca fertilizers (CaCO3 and CaSO4·2H2O), having negligible Mg content, were applied at rates of 1, 2, and 4 t Ca·ha–1 on sugar maple trees adjacent to the limed area. After 3 years, most of the foliar nutrient concentrations of treated trees were improved, particularly Ca, for both Ca fertilizers, in line with published ranges for healthy sugar maple trees, except for Mg. Moreover, no persistent nutrient antagonism was observed. The crown dieback rate of treated sugar maple was ≤5.8% after 3 years, while it reached 12% for the controls. Also, relative basal area growth showed that both Ca sources can improve growth rate. Growth response following Ca treatments was, however, lower than for the former lime experiment after the same period of time. In this context, our results suggest that Mg nutrition could be more important for sugar maple in this ecosystem than initially thought.
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  • 10
    Publication Date: 2010-09-01
    Description: Soil compaction often limits conifer regeneration on sites degraded by landings and roads, but inadequate understanding of the relationship between compaction and tree growth could lead to inappropriate soil conservation and rehabilitation efforts. We tested liquid and plastic limits, oxidizable organic matter, total carbon, particle size distribution, and iron and aluminum oxides on soil samples collected from five forest experiments in interior British Columbia. These data were used to estimate soil maximum bulk density (MBD) and relative bulk density (RBD); our objective was to relate RBD to tree growth. Height of interior Douglas-fir ( Pseudotsuga menziesii var. glauca (Bessin) Franco) was limited when RBD was 〉0.72. For lodgepole pine ( Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) and hybrid white spruce ( Picea glauca (Moench) Voss × Picea engelmannii Parry ex Engelm.), RBDs of 0.60–0.68 corresponded to maximum height, whereas RBDs of 0.78–0.87 appeared to limit height growth. The presence of surface organic material mitigated compaction and was often associated with lower RBD. Our results illustrate the usefulness of RBD to assess compaction and suggest that soil rehabilitation should be considered on disturbed sites where soil RBD is 〉0.80.
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  • 11
    Publication Date: 1991-03-01
    Description: Radial growth following a shelterwood seed cut in a 174-year-old stand of white spruce (Piceaglauca (Moench) Voss) in interior Alaska was compared with growth in an adjacent undisturbed stand of the same age. After a 2-year lag, radial growth of residual trees accelerated an average of 27% in 5 of the next 6 years. Net mean increase in growth after 8 years was 164%. Basal area growth of individual shelterwood trees increased 26.8% over the 14-year posttreatment period, while control trees increased 16.5%.
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  • 12
    Publication Date: 2010-12-01
    Description: A novel “varying-centroid” method is presented for predicting whole-tree, aboveground stem volume (i.e., bole plus branch volume) to bole volume ratios from changes in the centroid of tree bole volume associated with branching of the bole. The method was derived from a simple fractal-like tree model based on a conceptualization of tree branching architecture by Leonardo da Vinci. The method recognizes that the centroid of bole volume (the point at which one half of bole volume is above and one half is below) is always lower than the centroid of whole-tree volume and that shifts in the centroid of bole volume should be predictably related to the size of a tree’s crown. The method assumes that branch-displaced bole volume profiles can be compared with reference bole profiles that are not significantly influenced by branching, at the centroid of bole volume, and that the magnitude of bole centroid displacement predicts the branch volume necessary to cause it. When the method was applied to hardwood trees representing diverse species, sizes, and stand conditions across Michigan, the centroid of bole volume was found to vary predictably with measurable tree crown attributes and bole plus branch wood to bole wood volume ratios were generally predicted within 10% of the true value using the new method.
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  • 13
    Publication Date: 2010-11-01
    Description: Because of the importance of seed surface area, volume, and fill to hydraulic and thermal exchanges with the soil, mechanistic simulation of seed physiological processes associated with tree migration dynamics and the spread of invasive species require accurate equations to model seed shape. Seed dimensions have previously been described with measurements of the three principal axes, assuming an implied single ellipsoid. However, conifer seeds often exhibit anisotropy that results from bilaterally symmetric pairing on cone scales. We developed a method for measuring and modeling conifer seed shape as a sum of 2jpartial ellipsoids fused at their equators, where j = 0, …, 3. We demonstrate the use of the methods in the study of shape characteristics of ponderosa pine (Pinus ponderosa P.& C. Lawson) seeds from four families in Montana and among commercial lots of ponderosa pine, lodgepole pine (Pinus contorta Dougl. ex Loud.), and Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). The shapes of 92%, 73%, and 47% of seeds in commercial lots studied had eight unique ellipsoids when classified with 1%, 5%, and 10% difference classification rules, respectively. Ponderosa pine seeds with longer minor axes were less well filled with storage reserves. Three-dimensional surface areas of lodgepole and ponderosa pine were approximately 2 and 3.4 times larger, respectively, than previously reported one-sided surface areas.
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  • 14
    Publication Date: 2010-09-01
    Description: Small-scale disturbance is a significant process in all major forest biomes. Some silvicultural practices, particularly group selection harvesting, intend to emulate natural small-scale disturbance by harvesting small clearcuts in the continuous forest. We conducted a meta-analysis on the effects of small-scale harvesting on North American breeding forest birds. We extracted species richness and relative abundance of several functional bird groups and guilds from published studies and compared them between gap-dominated and unlogged forest as a function of forest type and the size and age of the gap. The abundance of many bird groups was higher in the gap-dominated than in the continuous forest. Species preferring interior parts of the forest had the most negative association with the presence of gaps but this relationship was not statistically significant. Abundances of many bird groups increased with increasing gap size, while its effect on abundance of some bird groups disappeared quickly. Our review suggests that silvicultural practices that bring about small gaps do not negatively affect the abundances of most forest birds and often even enhance it. However, more studies are needed to examine optimal size and abundance of gaps in a forest and whether emulated small-scale disturbance effectively mimics natural processes.
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  • 15
    Publication Date: 1991-12-01
    Description: Biomass allocation to roots was studied in holm oak (Quercusilex L.), a dominant evergreen tree in broad-leaved sclerophyllous Mediterranean forests. The root systems of 32 single-stemmed holm oaks growing in shallow soils on largely unfissured bedrock were excavated in a mesic site and a xeric site in the Montseny Mountains (northeast Spain). Individual root:shoot biomass ratios (roots with diameter
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  • 16
    Publication Date: 1991-10-01
    Description: The responses of CO2 assimilation rate (A), transpiration rate (E), and leaf conductance (g) to increasing leaf to air water vapor concentration difference (ΔW) were investigated (i) using excised shoots from mature trees of Abiesalba, Abiescephalonica, Abiesmarocana, and Abiesnordmanniana and (ii) in situ on a mature tree of Abiesbornmulleriana. Gas-exchange responses to increasing soil drought were also studied in plants of A. bornmulleriana, A. cephalonica, and Cedrusatlantica. Stable carbon isotope composition measurements were carried out on annual growth rings of A. bornmulleriana to estimate the time-integrated values of the ratio of intercellular leaf (Ci) to ambient (Ca) CO2 concentration. Increasing ΔW around the shoots reduced A and g in such a way that either Ci remained constant or its decrease was not pronounced enough for the changes in A to be accounted for by changes in g only. This suggests a direct effect of ΔW on photosynthesis. The different Abies species showed clear differences in water-use efficiency. Abiescephalonica and A. marocana had lower water costs of CO2 assimilation (E/A) than A. nordmanniana and A. alba. It has also been shown that A. cephalonica and A. marocana are characterized by an optimal stomatal control of leaf gas exchange. Stomata closed very rapidly in A. bornmulleriana in response to water supply being withheld, even prior to there being any important decrease in leaf predawn water potential. The stomatal response in C. atlantica was more gradual. In A. bornmulleriana, drought adaptation appears to be linked to the ability to avoid internal water stress, whereas drought adaptation in C. atlantica involves the ability to tolerate internal water stress. The high stomatal sensitivity mA. bornmulleriana is also supported by the isotopic carbon composition data, as shown by the substantial interannual variations in the estimates of Ci/Ca, ranging from 0.48 for the dryest years to 0.61 for the rainy years.
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  • 17
    Publication Date: 2010-04-01
    Description: A system of equations for compatible prediction of total and merchantable volumes that allows for different definitions of tree volume was developed in this study. The use of the developed system will allow the conversion and subsequent comparison of results from forest inventories using different definitions of tree volume (e.g., including or not the top material of the tree and (or) the stump, inside or outside bark). The compatibility between taper, total volume, and volume ratio equations is ensured by properly integrating the taper equation. The diameter under the bark at any height is modeled with the Demaerschalk taper equation, and the corresponding diameter over the bark is obtained by assuming that bark thickness is also modeled with Demaerschalk’s function. The set of equations that has contemporaneous cross-equation error correlation (known as nonlinear seemingly unrelated regression equations) was fit using nonlinear joint generalized least squares regression. The predictive ability was evaluated using an independent data set. The system is consistent and performs well when applied to maritime pine ( Pinus pinaster Ait.) trees in Portugal, showing better performance than do other total volume equations for maritime pine used in the latest Portuguese national forest inventories.
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  • 18
    Publication Date: 2010-12-01
    Description: Loblolly pine (Pinus taeda L.) is a major plantation species grown in the southern United States, producing wood having a multitude of uses including pulp and lumber production. Specific gravity (SG) is an important property used to measure the quality of wood produced, and it varies regionally and within the tree with height and radius. SG at different height levels was measured from 407 trees representing 135 plantations across the natural range of loblolly pine. A three-segment quadratic model and a semiparametric model were proposed to explain the vertical and regional variations in SG. Both models were in agreement that a stem can be divided into three segments based on the vertical variation in SG. Based on the fitted models, the mean trend in SG of trees from the southern Atlantic Coastal Plain and Gulf Coastal Plain was observed to be higher than in other physiographical regions (Upper Coastal Plain, Hilly Coastal Plain, northern Atlantic Coastal Plain, and Piedmont). Maps showing the regional variation in disk SG at a specified height were also developed. Maps indicated that the stands in the southern Atlantic Coastal Plain and Gulf Coastal Plain have the highest SG at a given height level.
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  • 19
    Publication Date: 2010-07-01
    Description: Subsistence harvesting and wild food production by Athabascan peoples is part of an integrated social–ecological system of interior Alaska. We describe effects of recent trends and future climate change projections on the boreal ecosystem of interior Alaska and relate changes in ecosystem services to Athabascan subsistence. We focus primarily on moose, a keystone terrestrial subsistence resource of villages in that region. Although recent climate change has affected the boreal forest, moose, and Athabascan moose harvesting, a high dependence by village households on moose persists. An historical account of 20th century socioeconomic changes demonstrates that the vulnerability of Athabascan subsistence systems to climatic change has in some respects increased while at the same time has improved aspects of village resilience. In the face of future climate and socioeconomic changes, communities have limited but potentially effective mitigation and adaptation opportunities. The extent to which residents can realize those opportunities depends on the responsiveness of formal and informal institutions to local needs. For example, increases in Alaska’s urban population coupled with climate-induced habitat shifts may increase hunting conflicts in low-moose years. This problem could be mitigated through adaptive co-management strategies that project future moose densities and redirect urban hunters to areas of lower conflict.
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  • 20
    Publication Date: 2010-12-01
    Description: Forest management planners usually treat potential fire loss estimates as exogenous parameters in their timber production planning processes. When they do so, they do not account for the fact that forest access road construction, timber harvesting, and silvicultural activities can alter a landscape’s vegetation or fuel composition, and they ignore the possibility that such activities may influence future fire losses. We develop an integrated fire and forest management planning methodology that accounts for and exploits such interactions. Our methodology is based on fire occurrence, suppression, and spread models, a fire protection value model that identifies crucial stands, the harvesting of which can have a significant influence on the spread of fires across the landscape, and a spatially explicit timber harvest scheduling model. We illustrate its use by applying it to a forest management unit in the boreal forest region of the province of Alberta in western Canada. We found that for our study area, integrated fire – forest management planning based on our methodology could result in an 8.1% increase in net present value when compared with traditional planning in which fire loss is treated as an exogenous factor.
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  • 21
    Publication Date: 1991-12-01
    Description: The renewal of boreal fir stands after harvesting is related to the abundance of fir advanced regeneration. The objective of this study was to compare the advanced regeneration in 45-years-old balsam fir stands of second growth to determine if ecological site conditions could explain some regeneration problems noted in the balsam fir – white birch ecoclimatic domain. Seven ecological phases were studied; these are among the most common in the Laurentians north of Québec. On the basis of fir seedling densities (2 years old and more), three groups could be distinguished using a cluster analysis method for grouping means. Dry balsam fir – herb-and-moss type on well drained tills formed a first group characterized with very high seedling densities (〉 60 000 seedlings/ha). A second group, characterized with high seedling densities (25 000 – 40 000 seedlings/ha), was constituted of three phases with an important moss cover:the mesic balsam fir – moss-and-herb type on moderately well drained tills, the humid balsam fir – moss-and-herb type on imperfectly drained tills with seepage, and the humid balsam fir – moss type on imperfectly drained tills. A third group, with low seedling densities (
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  • 22
    Publication Date: 2010-10-01
    Description: White poplar ( Populus alba L.) is a native species in Europe, but its growth potential is largely unknown. The general objectives of our study were to determine the impact of contrasted environments across Europe and the influence of parental characteristics on the growth potential of an intraspecific F1 white poplar family originating from a cross between parents native from the south and the north of Italy. The growth of the family was monitored at three sites located in the north of Italy, in central France, and in the southern United Kingdom. The family showed a highly superior productivity in Italy. A pronounced plasticity among sites was found for the male parent only. Indeed, for this parent, the highest growth was observed in northern Italy, its area of origin. A positive heterosis was observed mainly in France and in the United Kingdom. Broad-sense heritability values were moderate in most cases. However, the growth of the family was in some cases superior to the one of several other interspecific hybrid families growing under the same conditions, underlying the poorly known growth potential of such intraspecific hybrids for biomass production under European conditions.
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  • 23
    Publication Date: 2010-03-01
    Description: Declines of whitebark pine ( Pinus albicaulis Engelm.) have occurred across much of the species’ range over the last 40 years due to mountain pine beetle outbreaks and white pine blister rust infection. Management efforts to stem these declines are increasing, yet the long-term success of whitebark pine depends on the species itself adapting to the modern environment. Natural regeneration will be a critical part of this process. We examined patterns in natural whitebark pine regeneration as related to the biophysical environment in sixty 0.1 ha plots in Montana, Idaho, and Oregon. Whitebark pine regeneration was present in 97% of our plots and varied widely in density from 0 to 17 000 seedlings/ha and 0 to 2680 saplings/ha. Using nonparametric correlation analysis and ordination techniques, we found whitebark pine regeneration abundance was unrelated to stand age but significantly related to several biophysical site characteristics, including positive relationships with elevation and canopy tree mortality caused by mountain pine beetle and negative relationships with moisture availability, temperature, and subalpine fir importance. Our findings indicate that whitebark pine is regenerating in many areas and that the widespread mortality from recent mountain pine beetle outbreaks may provide suitable settings for whitebark pine regeneration given sufficient seed sources.
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  • 24
    Publication Date: 2010-01-01
    Description: We evaluated spruce budworm ( Choristoneura fumiferana (Clem.)) outbreak effects in nine study areas (60–86 ha each) located in the boreal forest of eastern Quebec (Canada). In each area, spruce budworm outbreak effects were measured from vegetation plots, dominant canopy and understory tree age structures, retrospective analysis of aerial photographs, defoliation records, and host tree growth reductions (dendrochronology). Large-scale synchronous outbreaks were detected across the region around the years 1880, 1915, 1950, and 1980. Overall, contrarily to what was expected for a region where host species (balsam fir ( Abies balsamea (L.) Mill.), Picea spp.) content is relatively high, these spruce budworm outbreaks seemed to have a relatively minor influence on stand dynamics, with the exception of the most recent outbreak (1980). This outbreak resulted in major stand mortality in the southern part of the region and favored the establishment of extensive tracts of young even-aged stands with few residual mature trees. This very abrupt increase in outbreak severity compared with earlier outbreaks, perhaps due to climatic or random factors, suggests that historical trends in successive outbreak severity should be extrapolated very cautiously and that the study of several outbreak cycles is needed to establish a range of natural variability that can be used to develop an ecosystem forest management strategy.
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  • 25
    Publication Date: 2010-03-01
    Description: Forest managers are seeking strategies to create stands that can adapt to new climatic conditions and simultaneously help mitigate increases in atmospheric CO2. Adaptation strategies often focus on enhancing resilience by maximizing forest complexity in terms of species composition and size structure, while mitigation involves sustaining carbon storage and sequestration. Altered stand age is a fundamental consequence of forest management, and stand age is a powerful predictor of ecosystem structure and function in even-aged stands. However, the relationship between stand age and either complexity or carbon storage and sequestration, especially trade-offs between the two, are not well characterized. We quantified these relationships in clearcut-origin, unmanaged pine and aspen chronosequences ranging from 130 years in northern Minnesota. Complexity generally increased with age, although compositional complexity changed more over time in aspen forests and structural complexity changed more over time in pine stands. Although individual carbon pools displayed various relationships with stand age, total carbon storage increased with age, whereas carbon sequestration, inferred from changes in storage, decreased sharply with age. These results illustrate the carbon and complexity consequences of varying forest harvest rotation length to favor younger or older forests and provide insight into trade-offs between these potentially conflicting management objectives.
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  • 26
    Publication Date: 1991-01-01
    Description: Stand tables from forest inventories representing more than 22 × 106 ha of forests in tropical Asia were used to estimate aboveground biomass (point and 99% confidence interval). The mean inventory-based biomass for moist forests (225 Mg/ha) was lower than that reported by direct measurements for mature forests in the same region (350 Mg/ha), whereas the mean inventory-based biomass for dry forests (82 Mg/ha) was higher than estimates based on direct measurements (55 Mg/ha). Our analyses demonstrated that human use of the forests in tropical Asia is intense, leading to degradation. Between two national forest inventories of Peninsular Malaysia in 1972 and 1981, the total area and biomass of forests declined by 18 and 28%, respectively. Modeling land-use changes and carbon dynamics of tropical Asian forests must take into consideration human impact on vegetation because such use of forests reduces their biomass and may stimulate forest growth and carbon uptake.
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  • 27
    Publication Date: 1991-08-01
    Description: Measuring the mass of snow on cut branch tips soon after snowfalls during two winters provided comparisons of catch by Engelmann spruce (Piceaengelmannii Parry), subalpine fir (Abieslasiocarpa (Hook.) Nutt.), and lodgepole pine (Pinuscontorta var. latifolia Engelm.). Analysis of these and other reported measurements confirmed (i) snow bridging by cohesion, (ii) bouncing of snow crystals by elastic rebound, and (iii) branch bending as mechanisms that determine the sigmoidal growth curves characterizing snow interception relative to snowfall. The fraction of snowfall intercepted by the branches was largest when storm accumulations were 3–4 mm water equivalent, with low specific gravity (0.04–0.07). Percent catch in snowfalls with 10 mm water and low specific gravity was near 50% for Engelmann spruce and about 45% for subalpine fir and lodgepole pine, but values decreased to near 30% in 20-mm storms. Catch was inversely proportional to the density of snow accumulations in the specific gravity range 0.04–0.13. Average branch catch was only about 10% of a storm with 10 mm water equivalent at 0.13 specific gravity. Meteorological conditions were more important than branch growth form in determining snow interception amounts on the conifers tested. The results suggest, as a hypothesis, a computational function for the fraction of snowfall caught on conifer crowns.
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  • 28
    Publication Date: 1991-02-01
    Description: Western hemlock (Tsugaheterophylla (Raf.) Sarg.) seedlings from four dormancy induction treatments (DIT) (i.e., long-day dry, long-day wet, short-day dry, short-day wet) were planted on a coastal reforestation site in British Columbia and monitored for physiological response and morphological development over the first growing season. Seedlings in all DIT showed a similar net photosynthetic, needle conductance, and shoot water potential response to seasonal low temperature, non droughty, and late summer drought conditions. Pressure–volume analysis at the beginning of the growing season (late February) showed short-day wet seedlings to have lower saturated and turgor loss point osmotic potentials than other DIT, while at the end of the growing season (October) there was no osmotic potential difference between DIT. At the end of the growing season, short-day wet seedlings had a much lower maximum modulus of elasticity than other DIT, indicating their shoots were still elongating. Morphological assessment 1 month after field planting showed short-day DIT seedlings had less needle damage and greater root development in response to continuous low temperature exposure. Four months after planting, long-day wet seedlings had the greatest shoot (i.e., height, root collar diameter) and root development in response to mild temperatures and high soil moisture conditions. Eight months after planting, survival was between 95 and 97% for all DIT. Long-day DIT seedlings had the greatest height and all DIT had similar root collar diameter and root development. Short-day DIT seedlings had the best shoot/soil roots ratio. Short-day wet seedlings had the least number of stem units per centimetre of new shoot development throughout the growing season. Results are discussed in reference to stock quality assessment presented in the first paper of this series.
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  • 29
    Publication Date: 2010-09-01
    Description: An integrated forest management optimization model was developed to calculate potential spruce budworm ( Choristoneura fumiferana Clemens) effects on forest and wood product carbon (C) from 2007 to 2057 and to evaluate potential C sequestration benefits of alternative management strategies (salvage, biological insecticide application). The model was tested using simulated spruce budworm outbreaks on a 210 000 ha intensively managed forest in northwestern New Brunswick, Canada. Under a severe spruce budworm outbreak scenario from 2007 to 2020, harvest volume and forest and wood product C storage in 2027 were projected to be reduced by 1.34 Mm3, 1.48 Mt, and 0.26 Mt, respectively, compared with the levels under no defoliation. Under the same severe outbreak scenario, implementation of salvage and harvest replanning plus a biological insecticide applied aerially to 40% of susceptible forest area, reduced harvest, forest C, and wood product C impacts by 73%, 41%, and 56%, respectively. Extrapolation of these results to all of New Brunswick suggests that a future severe spruce budworm outbreak could effectively increase total provincial annual C emissions (all sources) by up to 40%, on average, over the next 20 years. This modeling approach can be used to identify to what extent insecticide application, as a forest-C-offset project, could result in additional C storage than without forest and pest management.
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  • 30
    Publication Date: 1991-05-01
    Description: Seedlings of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco), lodgepole pine (Pinuscontorta Dougl.), and white spruce (Piceaglauca (Moench) Voss) were grown in a container nursery from February to July 1988 and then exposed to three temperatures and three levels of drought stress applied factorially during mid-July to October 1988. Seedlings were retained in a shelter house until January 1989, when they were cold-stored until early May. Measurements of stomatal conductance (gs), transpiration (E), and specific leaf area (SLA) were made at the end of the treatment period in September 1988 and again after growth the following year at the end of June. Root growth capacity (RGC) was tested in early May 1989. Results were considered in conjunction with performance of other samples of the same plants that had been planted in sand beds in April 1989, where irrigation was regulated to provide three levels of moisture stress. Low temperature (13 °C) generally reduced gs and E, which were adjusted for xylem pressure potential, and SLA in all species by the time nursery treatment was completed at the end of September. No effect of nursery temperature treatment on gs and E could be detected when new needles were measured in June and July (after 9 to 12 weeks of growth), but SLA of lodgepole pine increased with nursery temperature treatment, and SLA of white spruce decreased with nursery temperature treatment. RGC was higher for the 13 °C treatment than for the 16 and 20 °C treatments. Survival of outplanted seedlings was mainly inversely related to nursery temperature. Low nursery temperature reduced gs, E, and SLA and increased RGC. SLA of planted lodgepole pine increased with level of nursery drought treatment, and severe nursery drought increased gs under stress, when measured in June. No other effects of drought were detected, although drought treatment was effective in increasing survival of planted seedlings.
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  • 31
    Publication Date: 2010-05-01
    Description: Managed coniferous forest dominates much of the black-backed woodpecker’s ( Picoides arcticus Swainson) breeding range. Despite this, little is known about the fine-scale foraging behaviour of this focal species in unburned managed forest stands in the absence of insect outbreaks. To investigate the foraging substrates used in such a habitat, we employed radio-telemetry to track a total of 27 black-backed woodpeckers. During two successive summers (2005–2006), 279 foraging observations were recorded, most of which were on dying trees, snags, and downed woody debris. Individuals frequently foraged by excavation, suggesting that in the absence of insect outbreaks the black-backed woodpecker forages mainly by drilling. The majority of foraging events occurred on recently dead snags with a mean dbh (±SE) of 18.3 ± 0.4 cm. Our results suggest that in unburned boreal forest stands, substrate diameter and decay class are important predictors of suitable foraging substrates for black-backed woodpeckers. We suggest that conservation efforts aimed at maintaining this dead-wood dependent cavity nesting species within the landscape, should endeavour to maintain 100 ha patches of old-growth coniferous forest. This would ensure the continuous production of a sufficient quantity of recently dead or dying trees to meet the foraging needs of this species.
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  • 32
    Publication Date: 1991-01-01
    Description: The net annual flux of carbon from south and southeast Asia as a result of changes in the area of forests was calculated for the period 1850 to 1985. The total net flux ranged from 14.4 to 24.0 Pg of carbon, depending on the estimates of biomass used in the calculations. High estimates of biomass, based on direct measurement of a few stands, and low estimates of biomass, based on volumes of merchantable wood surveyed over large areas, differ by a factor of almost 2. These and previous estimates of the release of carbon from terrestrial ecosystems to the atmosphere have been based on changes in the area of forests, or rates of deforestation. Recent studies have shown, however, that the loss of carbon from forests in tropical Asia is greater than would be expected on the basis of deforestation alone. This loss of carbon from within forests (degradation) also releases carbon to the atmosphere when the products removed from the forest burn or decay. Thus, degradation should be included in analyses of the net flux of carbon from terrestrial ecosystems. Degradation may also explain some of the difference between estimates of tropical forest biomass if the higher estimates are based on undisturbed forests and the lower estimates are more representative of the region. The implication of degradation for estimates of the release of carbon from terrestrial ecosystems is explored. When degradation was included in the analyses, the net flux of carbon between 1850 and 1985 was 30.2 Pg of carbon, about 25% above that calculated on the basis of deforestation alone (with high estimates of biomass), and about 110% above that calculated with low estimates of biomass. Thus, lower estimates of biomass for contemporary tropical forests do not necessarily result in lower estimates of flux.
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  • 33
    Publication Date: 2010-11-01
    Description: New sensor-based approaches for assessing the quantity, quality, and value of timber are being developed with the goals of improving the accuracy and economics of forest measurements. One new approach is based on terrestrial laser scanning (TLS). Thirty-three plots in six radiata pine (Pinus radiata D. Don) stands were scanned using TLS. Tree locations were automatically detected. Stem profiles were measured using three methods: (i) TLS scans, (ii) Atlas Cruiser inventory procedures, and (iii) manual measurement after harvesting. Stems were optimally bucked based on log specifications and prices for Australian markets. Tree values and log product yields were estimated for the TLS data and compared with estimates based on Cruiser and actual manual measurements of stem profiles. TLS volume and value recovery were within 8% and 7%, respectively, of actual harvester recovery for five of the six stands in which it was used. Cruiser volume and value estimates were both within 4% of actual harvester recovery. Plot preparation procedures, tree characteristics, and taper equations used to model diameters on hidden stem sections affected the accuracy of automated stem detection and profile measurements for the TLS system. Improvements in data capture and analytical procedures should improve the accuracy of TLS-based volume and value estimates.
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  • 34
    Publication Date: 2010-06-01
    Description: Declining biodiversity is a critical component of global change owing to its influence on ecosystem functioning. Decomposition rate frequently increases with fungal species number, but the responses of extracellular enzymes to fungal species number have not been tested. To test the effect of biodiversity on decomposition and enzyme activities, quaking aspen ( Populus tremuloides Michx.) litter was inoculated with mixtures of one, two, four, or eight fungi from a pool of 16 fungi that had been isolated from a boreal forest in Alaska. Total CO2 release and the activities of β-glucosidase, which targets cellulose, and polyphenol oxidase, which targets lignin and other recalcitrant phenolic compounds, were observed across the range of species numbers in the mixtures. Total CO2 release and β-glucosidase activity increased with number of species but were only weakly correlated with each other; polyphenol oxidase activity had no correlation with number of species or CO2 release. The results indicate that, over 4 months, decomposition of labile carbon is positively correlated with number of species.
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  • 35
    Publication Date: 2010-07-01
    Description: In recent years, the annual imports of wooden bedroom furniture by the United States have been over five billion dollars, with more than two billion dollars of that coming from China. This trend led to an antidumping action against China in October 2003. Since January 2005, antidumping duties of 0.83% to 198.08% have been imposed on individual Chinese firms. To assess the impact of this antidumping action, intervention analysis was employed to examine the import values of four furniture commodities and the prices of two of them over 1997–2008. China and six other major competing countries were included in the analysis. With regard to import values from China, significant trade investigation effects were identified: the petition announcement generated a positive impact in March 2004; the preliminary less-than-fair-value (LTFV) determination had a negative impact from July to December 2004. However, the final implementation did not show any expected trade duty effect. The aggregate impact of the antidumping action on import values from China over 2003–2008 was approximately equivalent to a 1-month import reduction. The impact on the unit prices for China was insignificant. For the six competing countries, intervention analyses revealed that the antidumping action generated a positive trade diversion effect, with the magnitude smaller than the trade depression effect on China.
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  • 36
    Publication Date: 1991-07-01
    Description: Measurements of the spatial mean values of global irradiance, photosynthetic photon flux density, and the downward longwave irradiance under a 26-year-old, second-growth Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) forest canopy on a 26° south-facing slope were made with a tramway system, which scanned a 20-m transect in 12 min. The diffuse solar irradiance under the canopy was measured with a stationary pyranometer equipped with a shadow band. The extinction coefficients for the direct, global, and photon components were derived as functions of the solar incident angle over the range of 15° to 85°. The extinction coefficient for the diffuse radiation was found to correlate well with the ratio of the direct to diffuse irradiance above the canopy. Complete diurnal cycles of the downward longwave irradiance were simulated with a simple model based on the air temperature inside the stand. Analyses of the measurements of all the shortwave and longwave components were made using an effective leaf area index, which was derived from the measurements of the direct irradiance above and below the stand. It was found that the distribution of the leaf inclination angle of a Douglas-fir canopy has strong planophile characteristics, and that in the case of a forest stand on a slope, it is critical to obtain the characteristics of the light transmission through the canopy over the entire incident angle range before effective leaf area index is calculated. Warren Wilson's 57.5° approximation did not hold for the Douglas-fir canopy, which had distinct foliage clumping features.
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  • 37
    Publication Date: 2010-07-01
    Description: The comparative analysis of a large set of long-term fertilization and thinning studies in the major forest types of interior Alaska is summarized. Results indicate that nutrient limitations may only occur during the early spring growth period, after which moisture availability is the primary control of tree growth on warm sites. The temperature dynamics of both air and soil set seasonal bounds on the nutrient and moisture dynamics for all forest types. Air and soil temperature limitations are the primary control of intraseasonal growth in the colder topographic locations in interior Alaska. These locations are usually dominated by black spruce (Picea mariana (Mill.) Britton, Sterns, Poggenb.) vegetation types. The seasonal progression of factors controlling growth is strongly tied to the state factor structure of the landscape.
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  • 38
    Publication Date: 2010-03-01
    Description: Restoration by imitating natural disturbances is widely practised in boreal forests to increase the availability of habitats for specialized species. We studied the abundance and species richness of saproxylic beetles on different types of created dead wood during 2 years after restoration. The study was conducted on areas of a large-scale experiment in which Norway spruce ( Picea abies (L.) Karst.) forests were restored by controlled burning and partial harvesting with down wood retention in southern Finland. More beetle species were attracted to spruces than to birches and more species were attracted to burnt trees than to unburnt trees killed by girdling. Birch-living species consistently benefited from fire, but on spruce, the abundance of cambium consumers and their associates was negatively affected by fire. Trees at harvested sites attracted more beetles in the first year, but the volume of down wood retention had only minor effects. Beetle assemblages were strongly altered by burning and harvesting. We conclude that burning and harvesting are efficient tools to promote species richness within a short time period, but there is a risk that the dead wood resource may be rapidly exhausted. Moreover, many saproxylic species of spruce forests may not be adapted to open habitats formed by stand-replacing disturbances.
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  • 39
    Publication Date: 2010-04-01
    Description: The current mountain pine beetle infestation in British Columbia’s lodgepole pine forests has raised concerns about potential impacts on water resources. Changes in forest structure resulting from defoliation, windthrow, and salvage harvesting may increase snow accumulation and ablation (i.e., spring runoff and flooding risk) below the forest canopy because of reduced snow interception and higher levels of radiation reaching the surface. Quantifying these effects requires a better understanding of the link between forest structure and snow processes. Light detection and ranging (lidar) is an innovative technology capable of estimating forest structure metrics in a detailed, three-dimensional approach not easily obtained from manual measurements. While a number of previous studies have shown that increased snow accumulation and ablation occur as forest cover decreases, the potential improvement of these relationships based on lidar metrics has not been quantified. We investigated the correlation between lidar-derived and ground-based traditional canopy metrics with snow accumulation and ablation indicators, demonstrating that a lidar-derived forest cover parameter was the strongest predictor of peak snow accumulation (r2 = 0.70, p 〈 0.001) and maximum snow ablation rate (r2 = 0.59, p 〈 0.01). Improving our ability to quantify changes in forest structure in extensive areas will assist in developing more robust models of watershed processes.
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  • 40
    Publication Date: 1991-09-01
    Description: The temporarily increased fire hazard that is believed to result from the process of thinning is included in a single-stand model for assessing the economic benefits of juvenile spacing. Formulas for the expected net present value and the land expectation value are given along with methods for determining the age of financial maturity and the optimal rotation age. A numerical example is given to illustrate the degree of loss due to the increased fire risk. The problem of commercial thinning when the risk of fire is present is addressed using continuous-time models. It is shown how, when the fire hazard is exogenous to the thinning activity, the problem reduces to one of deterministic optimal control with the discount rate adjusted upward by an amount equal to the fire hazard rate. In the case when the fire hazard increases whenever thinning is taking place, it is shown that in general the optimal thinning policy is qualitatively different from that which is optimal in the no-risk case and involves periods of thinning at the maximum rate interspersed with periods of no thinning activity.
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  • 41
    Publication Date: 2010-06-01
    Description: Automated individual tree detection and delineation from high spatial resolution imagery provides good opportunities for forest inventory at a large scale. However, the accuracy of delineated crown size compared with ground measurements may not be sufficient. Thus, ordinary least squares (OLS) regression is no longer an appropriate approach to estimating and predicting variables from the delineated tree crown because both response variable and regressor are subject to measurement errors. In this study, we describe the functional and structural relationships between field-measured tree variables (i.e., tree diameter and crown width) and delineated tree crown width from remotely sensed imagery. We investigated the performance of OLS and three error-in-variable regression techniques including maximum likelihood estimator (MLE), major axis (MA) regression, and reduced major axis (RMA) regression using field-measured data and simulated data under different conditions. Our results indicated that MLE was desirable for estimating unbiased model coefficients. However, the adjustment assumption of the MLE model should be checked for predicting tree variables from remotely sensed imagery. When the assumption holds, the MLE model performed better for predicting the response variables than did the OLS model. Otherwise, the MLE model produced biased predictions for the response variables.
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  • 42
    Publication Date: 1991-07-01
    Description: An aseptic technique was used to collect 227 sap samples from 84 tapholes, including several initial and successive collections, of 27 sugar maples (Acersaccharum Marsh.) over six sap flow seasons (mid-February to late April). In this technique, all material used is sterilized, and the bark as well is sterilized with alcohol. Of the 227 sap samples, 62.5% were sterile or essentially sterile (0–10 colony forming units of microorganisms per millilitre), and of the 74 sap samples from initial collections, 83.8% were sterile or essentially sterile, in contrast with successive collections from the same taphole (P = 0.001). The 35 composite sap samples collected from 1800 nonaseptically tapped tapholes on 900 trees, which were our control, were all contaminated with greater than 800 colony forming units of microorganisms per millilitre. Sap obtained with the aseptic tapping technique produced 92.8% light amber grade syrup and 7.2% medium amber grade syrup. Sap obtained with the traditional tapping technique produced 61.8% light amber syrup, 11.8% medium amber syrup, 14.7% dark amber syrup, and 11.8% syrup that was darker than the dark amber grade. The grade was determined both visually and by a spectrophotometric method. We have substantiated that maple sap is sterile in maple xylem. If sap microbial contamination is minimized and cold sap storage is used, lighter color grade syrup than what is achieved with traditional practices (P = 0.001) can be produced to the end of the season. To avoid the usage of a chemical taphole sanitizer, we recommend cleaner ways for tapping maple trees.
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  • 43
    Publication Date: 2010-09-01
    Description: It is widely accepted that N limits primary production in temperate forests, although colimitation by N and P has also been suggested, and on some soils, Ca and base cations are in short supply. I conducted a meta-analysis to assess the strength of existing experimental evidence for limitation of primary production by N, P, and Ca in hardwood forests of the northeastern United States and southeastern Canada using data from 35 fertilization experiments in deciduous forests on glaciated soils across the region. There is strong evidence for N limitation (formal meta-analysis weighted mean response ratio = 1.51, p 〈 0.01; simple mean = 1.42, p 〈 0.001). Forest productivity also tended to increase with additions of P (simple mean = 1.15, p = 0.05) and Ca (simple mean = 1.36, p 〈 0.001). Across all treatments, 85% of response ratios were positive. Multiple-element additions had larger effects than single elements, but factorial experiments showed little evidence of synergistic effects between nutrient additions. Production responses correlated positively with the rate of N fertilization, but this effect was reduced at high rates of ambient N deposition.
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  • 44
    Publication Date: 2010-07-01
    Description: The resilience and vulnerability of permafrost to climate change depends on complex interactions among topography, water, soil, vegetation, and snow, which allow permafrost to persist at mean annual air temperatures (MAATs) as high as +2 °C and degrade at MAATs as low as –20 °C. To assess these interactions, we compiled existing data and tested effects of varying conditions on mean annual surface temperatures (MASTs) and 2 m deep temperatures (MADTs) through modeling. Surface water had the largest effect, with water sediment temperatures being ~10 °C above MAAT. A 50% reduction in snow depth reduces MADT by 2 °C. Elevation changes between 200 and 800 m increases MAAT by up to 2.3 °C and snow depths by ~40%. Aspect caused only a ~1 °C difference in MAST. Covarying vegetation structure, organic matter thickness, soil moisture, and snow depth of terrestrial ecosystems, ranging from barren silt to white spruce ( Picea glauca (Moench) Voss) forest to tussock shrub, affect MASTs by ~6 °C and MADTs by ~7 °C. Groundwater at 2–7 °C greatly affects lateral and internal permafrost thawing. Analyses show that vegetation succession provides strong negative feedbacks that make permafrost resilient to even large increases in air temperatures. Surface water, which is affected by topography and ground ice, provides even stronger negative feedbacks that make permafrost vulnerable to thawing even under cold temperatures.
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  • 45
    Publication Date: 2010-07-01
    Description: This paper assesses the resilience of Alaska’s boreal forest system to rapid climatic change. Recent warming is associated with reduced growth of dominant tree species, plant disease and insect outbreaks, warming and thawing of permafrost, drying of lakes, increased wildfire extent, increased postfire recruitment of deciduous trees, and reduced safety of hunters traveling on river ice. These changes have modified key structural features, feedbacks, and interactions in the boreal forest, including reduced effects of upland permafrost on regional hydrology, expansion of boreal forest into tundra, and amplification of climate warming because of reduced albedo (shorter winter season) and carbon release from wildfires. Other temperature-sensitive processes for which no trends have been detected include composition of plant and microbial communities, long-term landscape-scale change in carbon stocks, stream discharge, mammalian population dynamics, and river access and subsistence opportunities for rural indigenous communities. Projections of continued warming suggest that Alaska’s boreal forest will undergo significant functional and structural changes within the next few decades that are unprecedented in the last 6000 years. The impact of these social–ecological changes will depend in part on the extent of landscape reorganization between uplands and lowlands and on policies regulating subsistence opportunities for rural communities.
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  • 46
    Publication Date: 2010-02-01
    Description: Multiple-use forestry requires comprehensive planning to maximize the utilization and sustainability of many forest resources whose growth and productivity are interconnected. Forest fungi represent an economically important nonwood forest resource that provides food, medicine, and recreation worldwide. A vast majority of edible and marketed forest mushrooms belong to fungi that grow symbiotically with forest trees. To respond to the need for planning tools for multiple-use forestry, we developed empirical models for predicting the production of wild mushrooms in pine forests in the South-Central Pyrenees using forest stand and site characteristics as predictors. Mushroom production and species richness data from 45 plots were used. A mixed modelling technique was used to account for between-plot and between-year variation in the mushroom production data. The most significant stand structure variable for predicting mushroom yield was stand basal area. The stand basal area associated with maximum mushroom productivity (15–20 m2·ha–1) coincides with the peak of annual basal area increment in these pine forests. Other important predictors were slope, elevation, aspect, and autumn rainfall. The models are aimed at supporting forest management decisions and forecasting mushroom yields in forest planning.
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  • 47
    Publication Date: 2010-02-01
    Description: Exotic Pinus taeda L. plantations may be more productive than native ones. Several hypotheses may explain this difference; however, process models with a light-interception-driving variable cannot test these hypotheses without foliage display first being quantified in native and exotic trees. We quantified leaf area duration in North Carolina, USA (natural), and Gobernador Virasoro, Argentina (exotic), with no additional nutrients and optimum fertilizer treatments. More (60%–100%) foliage was displayed but for a shorter (∼86 fewer days) time per fascicle in the exotics than in the naturals. Study inference was limited, with only one native and one exotic site. However, while the sites were markedly different in soils, climate, resource availability, and genetics, and we observed significant differences in fascicle display and longevity, most fascicles at both sites survived two growing seasons: the one in which they were produced and the subsequent one. This robust finding indicates it would be reasonable to use two growing seasons for fascicle longevity in process modeling to test hypotheses explaining growth differences in native and exotic loblolly. Fertilization had no effect on any exotic tree parameter, but it increased natural tree fascicle number (24%) and length (30%).
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  • 48
    Publication Date: 2010-12-01
    Description: We studied the effect of water availability on basal area growth and wood properties of 11-year-old loblolly pine ( Pinus taeda L.) trees from contrasting Florida (FL) (a mix of half-sib families) and South Carolina coastal plain (SC) (a single, half-sib family) genetic material. Increasing soil water availability via irrigation increased average whole-core specific gravity (SG) and latewood percentage (LW%) by 0.036 and 6.93%, respectively. Irrigation did not affect latewood SG or wood stiffness, but irrigated FL and SC trees had more latewood due to a 29 day longer growing season. Irrigation did not affect the length of corewood production, but irrigated trees had earlier transition ages, producing outerwood ~3 years before rainfed trees. The increase in whole-core SG and LW% was moderate because irrigation promoted earlywood growth in corewood formed before canopy closure, but after year 7, rain-fed and irrigated trees had similar earlywood growth but irrigated trees had more latewood growth, increasing ring SG and LW%. The SC half-sib family had higher SG and greater LW% than trees from FL independent of irrigation due to greater yearly latewood growth. Thus, absence of soil water stress extended seasonal diameter cessation date but did not change latewood SG or wood stiffness.
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  • 49
    Publication Date: 2010-11-01
    Description: Wildfire risk assessment research has made considerable progress towards estimating the probability of wildfires but comparatively little progress towards estimating the expected consequences of potential fires. One challenge with estimating wildfire consequences has been to identify a common metric that can be applied to consequences measured in different units. In this paper, we use the preferences of representatives of local fire management agencies as the common consequences metric and apply it to a case study in the southern Gulf Islands, British Columbia, Canada. The method uses an expert survey and a maximum-difference conjoint analysis to establish the relative importance of specific fire consequences. A fire with a major potential for loss of life was considered to be about three times worse than major damage to houses and 4.5 times worse than loss of a rare species. Risk ratings were very sensitive to changes in fire consequences ratings. As the complexity of values at risk and number of stakeholders increase, the most efficient allocation of wildfire prevention, protection, and suppression resources becomes increasingly challenging to determine. Thus, as the complexity of stakeholder representation and values at risk increases, we need to pay increasing attention to quantitative methods for measuring wildfire consequences.
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  • 50
    Publication Date: 2010-06-01
    Description: In their recent review of arrested succession, Royo and Carson ( A.A. Royo and W.P. Carson. 2006. Can. J. For. Res. 36: 1345–1362 ) demonstrate that “recalcitrant understory layers” are widespread and pervasive modifiers of ecosystems and disruptors of forest regeneration. They rightly point out that many plant species associated with arrested succession are characterized by rapid vegetative spread. Extending their review, we point out that most of such species are clonal or thicket-forming and suggest that an additional reason why these plants so effectively suppress succession for extended periods is their long life-spans.
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  • 51
    Publication Date: 2010-03-01
    Description: Although considerable research has focused on the influences of logging debris treatments on soil and forest regeneration responses, few studies have identified whether debris effects are mediated by associated changes in competing vegetation abundance. At sites near Matlock, Washington, and Molalla, Oregon, studies were initiated after timber harvest to quantify the effects of three logging debris treatments (dispersed, piled, or removed) on the development of competing vegetation and planted Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco var. menziesii ). Each debris treatment was replicated with initial and annual vegetation control treatments, resulting in high and low vegetation abundances, respectively. This experimental design enabled debris effects on regeneration to be separated into effects mediated by vegetation abundance and those independent of vegetation abundance. Two to three years after treatment, covers of Scotch broom ( Cytisus scoparius (L.) Link) at Matlock and trailing blackberry ( Rubus ursinus Cham. & Schltdl.) at Molalla were over 20% greater where debris was piled than where it was dispersed. Debris effects on vegetation abundance were associated with 30% reductions in the survival of Douglas-fir at Matlock (r2 = 0.62) and the stem diameter at Molalla (r2 = 0.39). Douglas-fir survival and growth did not differ among debris treatments when effects were evaluated independent of vegetation abundance (i.e., with annual vegetation control), suggesting negligible short-term effects of debris manipulation on soil productivity.
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  • 52
    Publication Date: 2010-07-01
    Description: Disturbance–succession models describe the relationship between the disturbance regime and the dominant tree species of a forest type. Such models are useful tools in ecosystem management and restoration, provided they are accurate. We tested a disturbance–succession model for the oak–pine ( Quercus spp. – Pinus spp.) forests of the Appalachian Mountains region using dendrochronological techniques. In this model, fire promotes pines, while fire suppression, bark beetle outbreaks, and ice storms encourage oaks. We analyzed nine Appalachian oak–pine stands for species establishment dates and the occurrence of fires and canopy disturbances. We found no evidence that fire preferentially promoted the establishment of pine more than oak, nor did we find any evidence that canopy disturbances or periods of no disturbance facilitated the establishment of oak more than pine. Rather, we found that both species groups originated primarily after combined canopy and fire disturbances, and reduction of fire frequency and scope coincided with the cessation of successful oak and pine regeneration. Currently, heath shrubs are slowly dominating these stands, so we present a revised disturbance–succession model for land managers struggling to manage or restore oak–pine forests containing a dense ericaceous understory.
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  • 53
    Publication Date: 2010-08-01
    Description: Based on individual tree damage data dating back to the gale “Lothar” (winter 1999) in Baden-Württemberg, Germany, a statistical model was developed to estimate the risk of storm damage for individual trees. The data were compiled from the National German Forest Inventory. The model attempts to separate the effects of tree-specific variables, topography, site conditions and flow field related effects on damage probability. The crucial problem of missing information on the actual flow field parameters was solved by applying a generalized additive model that enables the simultaneous fit of a spatial trend function. The geographical location of risk hotspots as predicted by the model correspond well to the actual distribution pattern of storm damage as assessed by the forest service. Tree height proved to be one of the most important factors affecting the level of damage, while height to diameter at breast height ratio influences damage probability to a much lesser extent. The Norway spruce ( Picea abies (L.) Karst.) group has the highest potential to be damaged followed by the silver fir ( Abies alba Miller) – Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco) group and the Scots pine ( Pinus sylvestris L.) – larches ( Larix spp.) group. Predicted probabilities for deciduous trees are generally lower than those of conifers. West- to south-exposed locations bear a considerably higher damage risk and waterlogged soils show an increased predicted probability compared with slightly or not waterlogged soils.
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  • 54
    Publication Date: 2010-04-01
    Description: Fire suppression has facilitated the spread of red maple ( Acer rubrum L.), a fire-sensitive, yet highly adaptable species, in historically oak-dominated forests of the eastern United States. Here, we address whether a shift from upland oaks to red maple could influence forest hydrology and nutrient availability because of species-specific effects on precipitation distribution and inorganic nitrogen (N) cycling. In eastern Kentucky, we measured seasonal variations in red maple, chestnut oak ( Quercus montana Willd.), and scarlet oak ( Quercus coccinea Münchh.) throughfall and stemflow quantity and quality following discrete precipitation events, and we assessed net N mineralization rates in underlying soils over a 2-year period (2006–2008). Throughfall was 3%–9% lower underneath red maple than both oaks, but red maple generated 2–3× more stemflow. Consequently, NH4+ throughfall deposition was less under red maple than chestnut oak, whereas stemflow-derived nutrient inputs were substantially larger for red maple than both oaks. Soils underlying red maple had 5–13× greater winter net nitrification rates than soils under both oaks and 20%–30% greater rates of seasonal net ammonification than soils under chestnut oak. These findings suggest a spatial redistribution of water and nutrients via precipitation as red maple dominance increases and point to stemflow as an important mechanism that may foster red maple competitive success, further bolstering the mesophication process in the United States.
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  • 55
    Publication Date: 2010-04-01
    Description: The effects of precommercial thinning on the understory vegetative cover of 16- to 18-year-old spruce–hemlock ( Picea sitchensis (Bong.) Carrière – Tsuga heterophylla (Raf.) Sarg.) stands were studied in seven replicate areas over seven growing seasons postthinning. Vegetative cover was analyzed at the class level, but species-specific effects were examined in relation to their value as food for Sitka black-tailed deer ( Odocoileus hemionus sitkensis Cowan). When compared with unthinned controls, thinned stands (3.6–6.3 m spacing) had significantly greater understory cover. However, all thinned spacings led to similar understory cover. Conifer cover recovered to about two-thirds of its prethinning level within seven growing seasons posttreatment. Understory nonconiferous cover increased during the first 2–4 years postthinning but began to decline with increasing conifer cover during the next 3 years, nearly reaching pretreatment levels by year 7. In unthinned understories, vegetative cover had declined and was significantly lower than that beneath thinned stands. Summer food resource values for deer were increased by thinning. Winter food resource values were increased by thinning for snow-free conditions but were unaffected for conditions when herb-layer forbs were buried by snow.
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  • 56
    Publication Date: 1991-03-01
    Description: EM (expectation–maximization) algorithm procedures were used to estimate mating-system parameters in four natural populations of Thujaorientalis L. from China using seven allozyme marker loci (Fest1, Idh1, Idh2, Mr, Pgi2, Skdh1, and 6Pg2). The mean single-locus outcrossing rate was 0.70 for the species, but estimates showed that there was significant heterogeneity among loci within populations. Multilocus estimates of the outcrossing rate showed that there was significant heterogeneity among populations and among trees within populations. The mean multilocus outcrossing rate (0.75) was higher than the outcrossing rate (0.63) observed in a closely related conifer, Thujaoccidentalis L., but lower than estimates that have been reported for most other conifers. Significant heterozygote deficiencies, relative to Hardy–Weinberg and mating-system equilibria, were observed in all filial populations. In contrast, the maternal populations conformed to Hardy–Weinberg and mating-system equilibria at most loci. Self-fertilization and other forms of inbreeding (e.g., sibling mating) within the sampled populations are important contributors to the low outcrossing estimates in this conifer.
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  • 57
    Publication Date: 2010-02-01
    Description: Ecological land classification (ELC) is central to forestry and environmental management. Few methods exist for the statistical confirmation of the distinctness and continued integrity of the ecological character of ELC regions. Consequently, forest managers lack the tools to measure the impact of ecosystem stressors such as harvest practices and climate change. We develop a framework for tracking the distinctness and modification of vegetative communities of ELC natural regions. We base the framework on principles of numerical taxonomy using the Kolmogorov–Smirnov measure of distributional difference. We demonstrate the utility of the framework using data from a 1986 Forest Development Survey of tree species abundances on 13 508 sample plots from natural regions of the New Brunswick, Canada, ELC. Using the framework, we found vegetative communities of the ELC statistically distinct in the 1986 sampling. We also investigated the impacts of plantations on forest composition using vegetation profiles from a 1999 Forest Development Survey of planted stands. Simulated planting to 20% levels suggests that past and projected planting practices will modify the vegetative character of several natural regions on a scale comparable with interregion variation. The results demonstrate the potential of the framework to track changes to a variety of biotic communities impacted by natural and anthropogenic disturbance.
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  • 58
    Publication Date: 2010-07-01
    Description: La coupe avec protection des petites tiges marchandes est un type de coupe partielle qui consiste généralement à récolter toutes les tiges d’un diamètre à hauteur de poitrine (dhp) supérieur à 15,0 cm, tout en conservant les tiges de plus petites dimensions. Le succès du traitement, appliqué à des forêts résineuses mûres dominées par le sapin baumier ( Abies balsamea (L.) Mill.) ou l’épinette noire ( Picea mariana (Mill.) Britton, Sterns & Poggenb.), repose en partie sur la capacité des tiges protégées à survivre. Un modèle logistique mixte a été calibré à partir de 27 blocs expérimentaux établis au Québec. Ce modèle identifie les variables qui conditionnent les probabilités de pertes des tiges individuelles protégées de 5,1 cm et plus de dhp, par mortalité sur pied ou par chablis, 5 ans après des coupes avec protection des petites tiges marchandes. Les résultats indiquent que les probabilités de pertes après traitement sont largement tributaires des caractéristiques du peuplement avant coupe (surface terrière marchande, densité de gaules, proportion de pin gris ( Pinus banksiana Lamb.)), de la qualité des opérations (procédé de récolte, taux de protection des petites tiges marchandes) et des caractéristiques des tiges protégées au moment de la coupe (inclinaison, dhp, essence). Les variables associées à l’exposition aux vents et à l’écologie des stations n’ont pas permis d’améliorer le modèle. Afin d’éviter des pertes trop élevées, il importe de bien cibler les peuplements à traiter et de réaliser un suivi rigoureux des opérations de récolte.
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  • 59
    Publication Date: 2010-02-01
    Description: A beech bark disease infested American beech tree ( Fagus grandifolia Ehrh.) and two uninfested trees were selected in a mature natural stand in Michigan, USA, and mated to form two full-sib families for evaluating the inheritance of resistance to beech scale ( Cryptococcus fagisuga Lind.), the insect element of beech bark disease. Four half-sib families from both infested and uninfested trees were also evaluated for resistance. Using an artificial infestation technique, adult and egg count data were collected over 2 years and analyzed with generalized linear mixed methods to account for nonnormal distributions of the response variables. A significant effect for family was found for each variable. Family least squares means were computed as a measure of resistance and repeatabilities were calculated to provide an upper limit estimate of broad-sense heritability. The two families that ranked highest for resistance were the full-sib family from two uninfested parents and the half-sib family from a stand where all diseased trees had been removed. Together, the results suggest that selection and breeding may be an effective means to improve populations for artificial regeneration, and silvicultural treatments may provide an effective management option for mitigating beech bark disease through managing the genetic composition of natural regeneration.
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  • 60
    Publication Date: 1991-06-01
    Description: An experiment to examine the use of sewage sludge as a fertiliser for pole-stage Corsican pine (Pinusnigra var. maritima (Ait.) Melville) is described. Dominant and codominant trees responded markedly to additions of sludge at rates of 200 and 400 m3•ha−1, increasing sawlog volume by over 50% in treated plots. Foliar analyses suggest that the trees have responded to phosphorus in the sludge. The greatest impact of sludge on the soil occurred in the litter layer; concentrations of nitrogen, phosphorus, calcium, zinc, copper, cadmium, and lead were all greater compared with concentrations of untreated controls. Nevertheless, at rates of sludge application appropriate to fertilising (200 m3•ha−1), additions of heavy metals are considered small and are unlikely to cause adverse environmental effects.
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  • 61
    Publication Date: 2010-10-01
    Description: Shrubland biomass is important for fire management programmes and for carbon estimates. Aboveground biomass and the combustible portion of biomass, the fuel load, in the past have been measured using destructive techniques. These techniques are detailed, highly labour intensive, and costly; hence, an alternative approach was sought. The new approach used linear mixed-effects models to estimate biomass and fuel loads from easily measured field variables: shrub overstorey height and cover, and understorey height and cover. Site was regarded as a random effect. Sampling sites were located throughout New Zealand and included a range of shrubland vegetation types: manuka ( Leptospermum scoparium J.R. Forst. et G. Forst.) and kanuka ( Kunzea ericoides (A. Rich.) J. Thomps.) scrub and heath, pakihi (mixed low heath, fern, and rushes), and gorse ( Ulex europaeus L.). The approach was extended and confidence intervals were constructed for the regression models. Statistical analysis showed that understorey height and overstorey cover were significant (at the 5% level) in some cases. Overstorey height was highly significant in all cases (p 〈 0.0001), allowing development of models useful to the operational user. The models allow rapid estimation of average fuel loads or biomass on new sites, and double sampling theory can be applied to calculate the error in the resultant biomass estimate.
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  • 62
    Publication Date: 1991-01-01
    Description: Roots of 5-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco var. menziesii) on three Oregon sites were excavated with explosives and analyzed for the effects of competition on root biomass and for planting-induced root deformities. The plantations were in Nelder designs with graduated spacing from 300 to 15 250 cm2 per tree. Competition treatments consisted of weed-free intraspecific competition, grass cover seeded after 1 year of seedling growth, and red alder (Alnusrubra Bong.) interplanted 1:1 among the Douglas-fir. All plantations were kept at low water stress in year 1. The ratio of standing aboveground to belowground biomass was the same for each competitor type. Shoot:root ratios averaged about 4:1, except in severely suppressed trees, where ratios dropped toward 1:1 in those near death. Neither shoot:root ratio nor tree size was affected by planting-induced root deformities such as J- or L-rooting. It appears that after 5 years all root systems have fully compensated for planting deformities and future growth and stability have not been jeopardized.
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  • 63
    Publication Date: 2010-10-01
    Description: Knowledge of past forest fire regimes is important for developing management plans for conservation areas and for predicting the probable effects of forest management and climate change on the structure and dynamics of forests. In this study, fire scars on living and dead trees were systematically sampled on 256 study plots in three landscapes in northeastern Finland dominated by Scots pine ( Pinus sylvestris L.). A total of 1030 disks or partial cross sections from different trees, including scars from 98 distinct forest fires, were dendrochronologically dated with an accuracy of 1 year or better. The extraordinarily well-preserved old Pinus snags and stumps allowed us to reconstruct annual tree-ring and fire chronologies beginning from the year AD 653. The fire cycles in the studied landscapes were exceptionally long for a boreal region dominated by Pinus, on average 350 years during the last millennium. This demonstrates that the fire regimes of poorly studied remote regions cannot be extrapolated from fire regimes of sites examined in more detail. Based on statistics on lightning-ignition densities, we suggest that most of the fires detected in this study were ignited by humans. The reconstructed past fire cycles were probably shortened by human influence.
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  • 64
    Publication Date: 1991-02-01
    Description: Western hemlock (Tsugaheterophylla (Raf.) Sarg.) seedlings were grown in a greenhouse and subjected to four dormancy induction treatments (DIT) (i.e., long-day dry, long-day wet, short-day dry, and short-day wet) during midsummer. In the late summer and fall, seedling development was monitored and it was found that (1) short-day DIT caused a rapid cessation of shoot growth; (2) short-day DIT caused rapid fall development of needle primordia, while long-day DIT took until midwinter to produce the same number of needle primordia; and (3) short-day DIT seedlings had lower saturated and turgor loss point osmotic potentials and greater maximum modulus of elasticity in October than long-day DIT. Seedlings were tested with a comprehensive stock quality assessment procedure just before late winter field planting. These tests showed the following: (1) morphological parameters: short-day DIT reduced shoot to root ratios; (2) pressure–volume analysis: short-day wet seedlings had the lowest osmotic potentials at saturation and turgor loss point; (3) soluble sugar analysis: greater levels of total soluble sugars were found in non water stressed DIT compared with water stressed DIT seedlings; (4) seedling water movement: short-day DIT seedlings had the lowest resistance to water movement at low root temperature (5 °C); (5) low root temperature response: short-day compared with long-day DIT seedlings had greater photosynthesis and stomatal conductance at low root temperatures; (6) root growth capacity: seedlings from all DIT were capable of growing roots at optimum root temperature (22 °C), but short-day compared with long-day DIT had greater root growth at low root temperatures; (7) drought stress response: short-day wet seedlings had the highest photosynthesis and stomatal conductance levels as predawn shoot water potentials decreased; and (8) frost hardiness: short-day wet seedlings had the least needle damage when tested at temperatures of −15 and −18 °C.
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  • 65
    Publication Date: 1991-05-01
    Description: Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco), lodgepole pine (Pinuscontorta Dougl.), and white spruce (Piceaglauca (Moench) Voss) seedlings were grown in Styroblock containers in a container nursery from February to July 1988 and then exposed to three temperatures and three levels of drought stress applied factorially during 18 July to 29 September 1988. Mean temperatures of 13, 16, and 20 °C were imposed in growth chambers, in a cooled plastic house, and in an ambient plastic house, respectively. Control, medium, and severe levels of drought stress were imposed in a series of eight cycles, resulting in mean xylem pressure potentials of −0.32, −0.50, and −0.99 MPa, respectively. Seedlings were kept in the ambient plastic house until January, when they were lifted and cold-stored until planting. Between 11 and 18 April 1989, seedlings were planted in 0.5 m deep sand beds, which provided hygric, mesic, and xeric conditions for testing all species and treatments. At the end of nursery growth, increase in nursery temperature increased height and height:diameter ratio in all species and shoot:root dry weight ratio in Douglas-fir and lodgepole pine. Increase in temperature also increased the number of seedlings with large well-formed buds in white spruce, but reduced the number in Douglas-fir. Drought stress reduced height and dry weight in all species and bud length in lodgepole pine. After 9 weeks in sand beds, low nursery temperature increased survival (19% for lodgepole pine and white spruce grown in the xeric bed), except for Douglas-fir grown in the xeric bed. Nursery drought stress also increased survival (16% for Douglas-fir and lodgepole pine in the xeric bed), but had little effect on white spruce. Low temperature and drought stress treatments that increased survival also reduced height and dry weight of lodgepole pine and white spruce after one growing season in sand beds. Survival showed significant negative correlations with height, dry weight, and height:diameter and shoot:root weight ratios. Low nursery temperature continued to affect growth after planting, increasing relative growth rate and allometric ratio (K) of Douglas-fir and decreasing K of white spruce.
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  • 66
    Publication Date: 2010-02-01
    Description: Anthropogenic influences have altered most fire regimes. Fire management programs often try to mimic natural fire regimes to maintain fuels and sustain native fire-dependent species. Lightning is the natural ignition source in Florida, substantiating the need for understanding lightning fire incidence. Sixteen years of lightning data (1986–2003, excluding 1987 and 2002 due to missing data) from the NASA Cloud to Ground Lightning Surveillance System and fire ignition records were used to quantify the relationship between lightning incidence and ignitions on Kennedy Space Center, Merritt Island National Wildlife Refuge, and Cape Canaveral Air Force Station. There were 230 lightning fires with an average of 14 ignitions per year, primarily in July, and only one winter ignition. Precipitation influenced the efficiency of lightning ignitions, particularly July precipitation. We found that negative polarity strikes caused the majority of ignitions. Pine flatwoods was ignited more frequently than expected given equal chance of ignition among landcover types. About half (51%) of detected fires were instantaneous ignitions and the other 49% were delayed an average of 2 days. This information is useful for paramaterizing fire regime models and for mimicking the natural fire regime through fire prescriptions on these properties and throughout the southeastern United States. These methods may be useful in fire-maintained systems globally.
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  • 67
    Publication Date: 2010-12-01
    Description: Inventories of the necromass of coarse woody debris typically involve measurements of density (e.g., kilograms per cubic metre) on a sample of logs, with densities of other logs estimated based on assignment to decay classes. Here, we compare two new devices for assessing density of woody debris, a spring penetrometer and a dynamic penetrometer, with the traditional decay classification and knife test in terms of the strength of the relationship with measured density and the consistency in measurements by four different people. Our evaluation was conducted in a diverse tropical forest and involved only a brief training period in each method. Classifications or scores from all four methods were only weakly correlated with measured density, and consistency among technicians in the measurement–density relationship was highest for the dynamic penetrometer. Therefore, we conclude that when training time is limited and the sampled logs can reasonably be assumed to be representative of all of the logs (e.g., an inventory of one site at one time), it is best to simply assume that the average density of the sampled logs is representative of nonsampled logs. For inventories involving multiple people, limited training, and cases where the sample average is likely to be unrepresentative, we recommend the dynamic penetrometer.
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  • 68
    Publication Date: 2010-07-01
    Description: In the boreal forests of Alaska, recent changes in climate have influenced the exchange of trace gases, water, and energy between these forests and the atmosphere. These changes in the structure and function of boreal forests can then feed back to impact regional and global climates. In this manuscript, we examine the type and magnitude of the climate feedbacks from boreal forests in Alaska. Research generally suggests that the net effect of a warming climate is a positive regional feedback to warming. Currently, the primary positive climate feedbacks are likely related to decreases in surface albedo due to decreases in snow cover. Fewer negative feedbacks have been identified, and they may not be large enough to counterbalance the large positive feedbacks. These positive feedbacks are most pronounced at the regional scale and reduce the resilience of the boreal vegetation – climate system by amplifying the rate of regional warming. Given the recent warming in this region, the large variety of associated mechanisms that can alter terrestrial ecosystems and influence the climate system, and a reduction in the boreal forest resilience, there is a strong need to continue to quantify and evaluate the feedback pathways.
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  • 69
    Publication Date: 2010-02-01
    Description: In forest ecosystems, litterfall that collects in trapping devices, to quantify organic matter and nutrient inputs, is exposed to periods of wetting, drying, freezing, and thawing. These fluctuating environmental conditions may influence the microbial community structure inhabiting the leaves and may result in the loss of mobile nutrients, leading to an underestimation of actual organic matter and nutrient inputs. The objectives of this study were to evaluate the influence of (i) different quantities of moisture (LOW = 30 mm, MED = 60 mm, HI = 100 mm) and (ii) freeze–thaw (FT) on leaf (sugar maple ( Acer saccharum Marsh.), American basswood ( Tilia americana L.), and American beech ( Fagus grandifolia Ehrh.)) microbial activity and community structure. There was a significantly greater (p 〈 0.05) CO2 production rate in LOW and FT treatments for sugar maple and beech, and in HI and FT treatments for basswood. A similar trend occurred for leaf nitrogen concentration but not for carbon (C). Utilization of C substrates was up to 10% greater in the FT treatments. Principal components analysis on the activity of C source utilization showed a distinct clustering between leaf species and between treatments following a pattern similar to that of microbial respiration. Results from this study suggested that the collection of litter should take place more frequently during seasons when frost is imminent.
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  • 70
    Publication Date: 2010-07-01
    Description: Warming in the boreal forest of interior Alaska will have fundamental impacts on stream ecosystems through changes in stream hydrology resulting from upslope loss of permafrost, alteration of availability of soil moisture, and the distribution of vegetation. We examined stream flow in three headwater streams of the Caribou–Poker Creeks Research Watershed (CPCRW) in interior Alaska over a 30-year period to determine (i) how stream flow varied among streams draining watersheds with varying extents of permafrost and (ii) evaluate if stream hydrology is changing with loss of permafrost. The three streams drained subcatchments with permafrost extents ranging from 4% to 53%. For each stream, runoff data were analyzed by separating base and storm flow contributions using a local-minimum method and with analysis of flood recession curves. Mean daily runoff during the ice-free season did not significantly vary among streams (mean = 0.57 mm·d–1), although the watersheds with lower permafrost had a greater contribution of base flow. Across years, flow was variable and was related with summer temperature in the watershed with low permafrost and with precipitation in the watershed with high permafrost. With climate warming and loss of permafrost, stream flows will become less responsive to precipitation and headwater streams may become ephemeral.
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  • 71
    Publication Date: 1991-10-01
    Description: Three experiments were conducted to determine effects of N and P fertilizers on growth and levels of plant-tissue nutrients of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and western hemlock (Tsugaheterophylla (Raf.) Sarg.). Both pole-size trees in closed-canopy stands and potted seedlings were used. Soil series were Bunker for Douglas-fir and Klone for western hemlock in experiments 1 and 3, and Vesta in experiment 2. For each species in experiments 1 and 2, P and N fertilizers were tested in six or eight treatments using factorial design. In experiment 3, N and P fertilizers were individually tested on seedlings, at one rate of application each. Nitrogen fertilizers used were urea in experiment 1 and ammonium nitrate in the other two experiments; P was applied as triple superphosphate in all three experiments. In general, fertilization changed levels of some plant-tissue nutrients of the pole-size trees and potted seedlings. Neither height nor basal-area growth of the trees was significantly affected by any of the fertilization treatments in the first two experiments. Seedling growth of both Douglas-fir and western hemlock was dramatically improved by the P fertilizer, but was negatively affected by the N fertilizer. Results clearly show differences between pole-size trees and seedlings in response to N and P fertilizers. They also suggest that N not be applied where soils are high in N and low in P and that P applications be confined to sites with low-P soils, when trees are young, before canopy closure.
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  • 72
    Publication Date: 1991-12-01
    Description: Starch is the main carbohydrate storage form in conifers and is derived from the translocation of photosynthate (soluble sugars) in the phloem. We examined seasonal patterns in concentrations of both carbohydrate forms in the needles, twigs and branches, stems, and coarse and fine roots of mature Pinuselliottii Engelm. var. elliottii trees in a north Florida plantation. Starch showed marked and similar seasonality in all the tissues, with maxima in the late winter near the time of the inception of new growth in the spring. Sugars showed little seasonality, except in the fine roots. Repeated fertilization had little effect on either sugar or starch concentrations in all tissues. Using published biomass data from these same stands, we estimated that coarse roots store more than half of the starch in these stands over the year, with foliage and fine roots storing less than 5%.
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  • 73
    Publication Date: 2010-01-01
    Description: Large trees with hollows are an important component of stand structural complexity worldwide. Understanding their population dynamics is needed to manage cavity-dependent biota. We quantified long-term rates of collapse of 302 measured trees with hollows in 1939-aged regrowth mountain ash ( Eucalyptus regnans F. Muell.) forest in southeastern Australia. We identified time-dependent dynamics in which the collapse rates of trees slowed from ∼4% annually between 1983 and 1993 to ∼2.2% between 1993 and 2007. Transitions of trees between different decay states (forms) also slowed over time. Nevertheless, during the 24-year period of our study, over half of our marked and measured trees had fallen, but there was no recruitment of new trees with hollows. Under current projections, few trees with hollows will occur on our field sites by ∼2050, although more had been forecast in earlier investigations. Such a paucity of trees with hollows in extensive areas of regrowth mountain ash forests will lead to a shortage of nesting and sheltering sites for cavity-dependent biota. We suggest a short–medium (10- to 100-year) focus on the conservation of old growth and multi-aged stands will be needed to maintain populations of those species strongly associated with trees with hollows in mountain ash forests.
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  • 74
    Publication Date: 1991-03-01
    Description: Inherently rapid growth in black spruce (Piceamariana (Mill.) B.S.P.) and radiata pine (Pinusradiata D. Don) could be characterised at a relatively early age (e.g., 3–6 months from germination) by growing progeny of family crosses in near-optimal phytotron or glasshouse environmental conditions. This observation was retrospective in nature, in that the studies of very early performance utilized stored seed from families that had been already tested to age 10+ years in field progeny trials. The very early traits that show the most significant family correlation with the field performance (height or stem volume) are total height, height growth, stem volume, or stem dry weight. The possibility that inherently rapid growth in trees may be causally related to concentration of endogenous plant hormones of the gibberellin class is discussed in relation to the recent finding (S.B. Rood, R.I. Buzzell, L.N. Mander, D. Pearce, and R.P. Pharis. 1988. Science (Washington, D.C.), 241: 1216–1218.) that heterotic growth (hybrid vigour) in maize was strongly related to the concentration of gibberellin A1 (a shoot growth effector in maize) and gibberellin A19 (a precursor to gibberellin A1). Preliminary evidence, based on analysis of hybrid and parental poplar tissue, and on the positive and significant growth response of black spruce slow-growing families (but not fast-growing families) to applied gibberellin (A4/7 mixture), supports the possibility that rapid-growing conifer families may have near-optimal (high) concentrations of endogenous gibberellins, and conversely, that some slow-growing families, at least, may have lower endogenous gibberellin concentrations.
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  • 75
    Publication Date: 2010-06-01
    Description: The arboreal forage lichen Bryoria fremontii (Tuck.) Brodo & D.Hawksw. appears sensitive to conditions of prolonged hydration in wet forests of British Columbia. I estimated the abundance of this lichen in mixed-conifer forest canopy in the contrasting Mediterranean climate of the southern Sierra Nevada in relationship to the vertical gradient of vapor pressure deficit. Abundance was estimated by biomass in 5 m strata from the ground to the tops of 50 trees. Transplants of Bryoria thalli were installed in 18 fir trees for 1 year to assess their growth relative to distance from perennial streams. VPD generally increased with height, being significantly greater at 45 m than 5 m. Bryoria biomass averaged across tree heights was estimated as 15.9, 0.60, 0.15, 0.25, and 0.19 g·m–1 in red fir (Abies magnifica A.Murray), white fir (Abies concolor (Gordon & Glend.) Hildebr. var. lowiana (Gordon) Lemmon), incense cedar (Calocedrus decurrens (Torr.) Florin), Jeffrey pine (Pinus jeffreyi Balf.), and sugar pine (Pinus lambertiana Douglas), respectively. Transplant growth was significantly greater in trees 25 m from water (9.7%). Bryoria had strong positive associations with red fir, proximity to streams, and decreasing VPD. The sensitivity of Bryoria to prolonged hydration does not appear to be important in this climate characterized by extended summer dryness.
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  • 76
    Publication Date: 1991-12-01
    Description: Effects of K and P deficiency on polyamines (putrescine, spermine, spermidine, and agmatine) in leaves and roots were investigated in poplar (Populusmaximowiczii Henry) cuttings growing in half-strength Hoagland solution with varying levels of K (0, 0.3, and 3 mM) and P (0, 0.05, and 0.5 mM). Putrescine contents increased in the leaves of plants grown without K and in roots of plants grown in 0.3 mM K and without K. In the absence of K, the putrescine levels in the roots and leaves were 80 and 25 times, respectively, those of the control (3 mM K). The zero-P treatment increased the putrescine content in the leaves and decreased the putrescine content in the roots. The accumulation of putrescine did not appear to be regulated solely by the K status of the plants. The spermine content in the leaves of 0.3 mM K and zero-P treatments decreased compared with the control, as did spermine in the leaves of the zero-P treatment. The spermine content in the leaves of 0.05 mM P treatment was similar to that of the control (0.5 mM P). This work shows that in poplar polyamine levels in leaves and roots are strongly influenced by the nutrient status of the plants.
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  • 77
    Publication Date: 1991-08-01
    Description: The Diagnosis and Recommendation Integrated System (DRIS) was used to describe the variation of 12 elements in woody tree tissue of balsam fir (Abiesbalsamea (L.) Mill.), sugar maple (Acersaccharum Marsh.), jack pine (Pinusbanksiana Lamb.), red pine (Pinusresinosa Ait.), and aspen (Populustremuloides Michx.) across Minnesota, Wisconsin, and Michigan, United States. DRIS indices of elemental balance for the growth decades 1956–1965 and 1966–1975 were compared with standards developed from the growth decade 1976–1985. The DRIS analysis indicated that older wood of most species was relatively depleted of N, P, K, S, Fe, Cu, and Al. In at least one of the five species, however, K, S, Cu, or Al was relatively more abundant in older than in younger wood. The older wood of all species was relatively enriched in Ca, Mg, Mn, B, and Zn. Sulfur in older wood became relatively more enriched from west to east across a gradient of wet sulfate deposition; the trend was strongest for hardwood species. These results support the potential use of DRIS for monitoring stoichiometry of tissue from woody increment cores as an indicator of environmental stresses such as air pollution.
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  • 78
    Publication Date: 2010-06-01
    Description: Airborne Heterobasidion annosum (Fr.) Bref. sensu lato infections can be controlled by winter thinning or by mechanically spreading urea or Phlebiopsis gigantea (Fr.) Jülich spores on stump surfaces during summer thinning operations. The long-term outcomes of these control methods when applied as part of the conventional forest operations are unclear. We studied the rot incidence and population structure of H. annosum in plots of Picea abies (L.) Karst. thinned in winter or thinned in summer with and without treatment of the stumps. Plots were distributed among 11 stands in Sweden representing two different land use histories: forest and agricultural. After 13 years, the effect of stump treatment on rot incidence was only evident in stands on former agricultural land. In stands planted on former forest land with higher levels of preexisting rot than on former agricultural land, the expansion of preexisting genets of H. annosum might have masked the effects of stump protection. In former forest land, unprotected summer plots showed a greater diversity of H. annosum genotypes and a smaller number of trees infected by each genet than in protected plots, suggesting that protection treatments prevented the establishment of new genets, which may result in a reduced rot incidence in the future.
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  • 79
    Publication Date: 1991-09-01
    Description: In the present study we describe biomass, productivity, and nutrient cycling in an 8-year-old Eucalyptustereticornis Sm. (Eucalyptus hybrid) plantation and compare them with those of a Populusdeltoides Bartr. plantation of the same age and area, a natural sal (Shorearobusta Gaertn. F.) forest, and other natural forests of the central Himalaya. The total vegetation biomass of the Eucalyptus plantation (126.7 t•ha−1) was lower than that of the P. deltoides plantation (176 t•ha−1) and natural forests (163.4–786.7 t•ha−1). The net primary productivity of the Eucalyptus plantation (23.4 t•ha−1•year−1) was similar to that of the P. deltoides plantation (25 t•ha−1•year−1) and the natural sal forest (22 t•ha−1•year−1). The net nutrient uptake of Eucalyptus was lower than that of Populus plantation and natural forests.
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  • 80
    Publication Date: 1991-03-01
    Description: For many types of forest studies, it is essential to identify the exact years of formation of annual rings in increment cores taken from living trees. To accomplish this, dendrochronologists employ cross dating, which involves both ring counting and ring-width pattern matching, to ensure against counting error, or errors, caused by missing or false rings. To date, published accounts of the cross-dating process generally describe a graphical method for achieving cross dating, known as skeleton plotting. However, when working with cores from living trees, skeleton plotting is seldom necessary. Such cores can commonly be cross-dated more quickly and easily by listing the narrow rings that are present in each core in a laboratory notebook and then comparing core notes for shared narrow rings. The latter approach permits faster recognition of ring-width patterns because calendar-year, rather than relative-year, dates are assigned to rings in cores. It also allows cross-dating records to be stored in a more concise manner.
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  • 81
    Publication Date: 2010-05-01
    Description: We investigated the frequency of root grafting in naturally and artificially regenerated stands of jack pine ( Pinus banksiana Lamb.) in the western boreal forest of Quebec, Canada. Twelve 30–60 m2 plots were hydraulically excavated to determine effects of site characteristics on frequency and timing of root grafting. Naturally regenerated stands had grafted tree percentages similar to artificially regenerated stands (21%–71% across plots) but greater numbers of root grafts per tree (naturally regenerated, 0.73 graft·tree–1; artificially regenerated, 0.52 graft·tree–1). Mean percentages of grafted trees, number of grafts per tree, and the speed of graft formation were greater in sandy soils (61%, 0.71 graft·tree–1 and 2.43 years, respectively) compared with clay soils (44%, 0.54 graft·tree–1 and 2.97 years, respectively). Proximity of trees was a better predictor of root grafting than stand density, despite many root grafts being found with distant trees (〉2 m) in artificially regenerated stands. Our results suggested that root grafts form early in stand development. Even if trees are initially separate entities, this relatively high level of root grafting produces stands where trees are extensively interconnected.
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  • 82
    Publication Date: 1991-01-01
    Description: This study investigates the impact of postulated greenhouse warming on the severity of the forest fire season in Canada. Using CO2 levels that are double those of the present (2 × CO2), simulation results from three general circulation models (Geophysical Fluid Dynamics Laboratory, Goddard Institute for Space Studies, and Oregon State University) were used to calculate the seasonal severity ratings for six stations across Canada. Monthly anomalies from the 2 × CO2 simulation results were superimposed over historical sequences of daily weather. Then, seasonal severity ratings of the present were compared with those for 2 × CO2 using five variations involving temperature, precipitation, and relative humidity. The relationship between seasonal severity rating and annual provincial area burned by wildfire was explored. The results suggest a 46% increase in seasonal severity rating, with a possible similar increase in area burned, in a 2 × CO2 climate.
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  • 83
    Publication Date: 2010-08-01
    Description: Ditch networks in drained peatland forests are maintained regularly to prevent water table rise and subsequent decrease in tree growth. The growing tree stand itself affects the level of water table through evapotranspiration, the magnitude of which is closely related to the living stand volume. In this study, regression analysis was applied to quantify the relationship between the late summer water table depth (DWT) and tree stand volume, mean monthly summertime precipitation (Ps), drainage network condition, and latitude. The analysis was based on several large data sets from southern to northern Finland, including concurrent measurements of stand volume and summer water table depth. The identified model demonstrated a nonlinear effect of stand volume on DWT, a linear effect of Ps on DWT, and an interactive effect of both stand volume and Ps. Latitude and ditch depth showed only marginal influence on DWT. A separate analysis indicated that an increase of 10 m3·ha–1 in stand volume corresponded with a drop of 1 cm in water table level during the growing season. In a subsample of the data, high bulk density peat showed deeper DWT than peat with low bulk density at the same stand volume.
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  • 84
    Publication Date: 2010-06-01
    Description: We embedded a linear programming timber harvest scheduling model into an aspatial stochastic simulation model of a flammable forest to evaluate two fire risk mitigation strategies. The harvest scheduling model is solved repeatedly to produce harvest schedules within a rolling planning horizon framework. The risk mitigation strategies we examined were (1) whether or not to account for fire in the planning model and (2) replanning interval. We evaluated those strategies under four representative fire regimes. We found that accounting for fire in the planning model reduced the harvest volume variability as fire activity increased (i.e., for average annual burn fractions ≥0.45%), but replanning intervals over a range of 1 to 10 years had little impact on harvest volume variability. We also developed a risk analysis decision-making aid that forest managers can use to help deal with fire-related uncertainty. Our results suggest that risk-averse forest managers should account for fire while planning, especially when burn fractions exceed 0.45%.
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  • 85
    Publication Date: 2010-08-01
    Description: Pine needle blight, caused by Dothistroma septosporum (Dorog.) M. Morelet, is one of the most serious foliar diseases of Pinus spp. in Australia and New Zealand. In 16 Pinus radiata (D.Don.) progeny trials in northeastern Victoria, Australia, Dothistroma-caused defoliation varied widely among trials and assessment years, ranging from 5% to 65%. The estimated narrow sense heritability ranged from nonsignificant to as high as 0.69 with a median of 0.36. Spatial autocorrelation of residuals accounted for a significant proportion of residual variance, and that increased heritability estimates. Genetic correlation between defoliation scores at an early age and growth at a later age was negative with a median value of –0.39. Phenotypic correlation between defoliation and survival was low and negative with a median value of –0.11. Economic analyses indicated that at sites with a high risk of infection, the effect of reducing defoliation on profitability was comparable with that of increasing growth at sites free from infection. The genetic parameters and economic impacts of Dothistroma were used to derive selection indices and include resistance to defoliation into the current breeding objective for radiata pine.
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  • 86
    Publication Date: 1991-08-01
    Description: Pollen contamination causes major losses to genetic improvement from selection and breeding of "plus" trees in conifer seed orchards. Genetic losses arise by the influx of "wild" conspecific pollen into seed orchards and its deleterious fertilization of superior genetic lines. This review firstly addresses the basis of the problem: pollen, conifer reproduction, and the concept of seed orchard management, especially in regard to reduction of contamination. Secondly, the physical processes of pollen liberation, dispersal, and deposition are described, and examples of previous studies illuminating these phenomena given. Thirdly, past research on measuring pollen dispersal in natural stands and seed orchards in discussed in the light of modelling techniques used to predict these types of dispersal pattern. Work on the other facets of contamination measurement, gene-flow studies, are listed. It is concluded that a detailed study that combines both the physical and gene-flow aspects of pollen dispersal should be initiated to compare and contrast the two methods, and that attempts to model pollen contamination should be sought.
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  • 87
    Publication Date: 2010-08-01
    Description: Optimal breeding zones were developed for white spruce ( Picea glauca (Moench) Voss) in Ontario under present and future climate conditions to examine potential shifts due to climate change. These zones were developed by (i) determining a set of candidate breeding zones based on the relationship between measured performance variables and climate and (ii) employing a decision support model to select subsets of breeding zones that maximize geographic coverage subject to a constraint on the maximum number of zones. Current optimal breeding zones were based on 1961–1990 climate normals, and future breeding zones were based on three general circulation model (CGCM2, HADCM3, and CSIRO) predictions of 2041–2070 climate. Based on a maximum adaptive distance of 2.0 least significant difference values between sites within zones, 14 zones were required to cover the Ontario range of white spruce for the 1961–1990 data. Compared with breeding zones of other boreal conifers, current optimal breeding zones for white spruce were quite large, spanning up to 3° latitude and 10°–12° longitude and indicating large distances of effective seed transfer. Of the three general circulation models used to simulate future climate, HADCM3 B2 and CGCM2 B2 predicted 2041–2070 breeding zones that largely coincide with 1961–1990 zones. In contrast, CSIRO B2 indicated much narrower 2041–2070 breeding zones.
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  • 88
    Publication Date: 2010-02-01
    Description: Acacia auriculiformis A. Cunn. ex Benth. is an important planting tree species, but little attention has been paid to its wood properties, such as shrinkage, stiffness, strength, and basic density, which are important for use in structural and appearance-grade timber applications. Here we report the genetic variation in static bending stiffness and strength of wood in a 5½-year-old clone trial in southern Vietnam and the genotypic correlations among these traits and tree diameter, wood shrinkage, and basic density. There was significant variation in stiffness and strength among 40 randomly selected clones. Clonal repeatability (H2) was high for stiffness and moderate for strength. There was no consistent pattern of difference between heartwood and sapwood for the estimates of H2 for stiffness and strength, whereas the estimates of H2 were lower for heartwood density than for sapwood density. Diameter showed a significant negative genotypic correlation with stiffness but a nonsignificant correlation with strength. Genotypic correlations between density and stiffness, and density and strength, were positive, but only the latter was significant. Stiffness and strength were not significantly correlated with wood shrinkage. There is potential to simultaneously improve tree growth, wood basic density, and the mechanical properties of juvenile wood of this species.
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  • 89
    Publication Date: 1991-06-01
    Description: A comparison of four types of litter traps was conducted during three growing seasons in an old-growth Acersaccharum Marsh, dominated forest on rugged topography in northern Ontario, Canada. The comparison involved traps of two rim heights (0.25 and 1.0 m) and two surface areas (0.25 and 1.0 m2). A supplementary study, also described, compared the two smaller sizes of traps over the same time period. Analysis of variance showed significant effects of both height and surface area on the mass of litter trapped. In a four-way comparison, however, only small traps positioned close to the ground produced estimates that differed significantly from the others. The higher catches associated with these traps are probably the result of a more favorable microsite for entrapment and accumulation of windblown leaves. The number of traps of each type required to constitute a representative sample ranged from ca. 6 to 14 when calculated over three growing seasons. Over the course of a single growing season, the number of traps required increased substantially. No trap type exhibited a distinct advantage in this regard. Generally, however, traps that present a favorable microsite for accumulation of windblown leaves should be avoided in A. saccharum forest.
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  • 90
    Publication Date: 2010-07-01
    Description: Along the Tanana River floodplain, several turning points have been suggested to characterize the changes in ecosystem structure and function that accompany plant community changes through primary succession. In the past, much of this research focused on a presumed chronosequence that uses space for time substitutions. Within this chronosequence, permanent vegetation plots repeatedly measured over time provide an excellent test of the turning points model. We analyzed both canopy and understory vegetation data collected since 1987 in the Bonanza Creek Experimental Forest at the Bonanza Creek Long Term Ecological Research site to address the following questions: (i) Do long-term changes in the densities of seedling, sapling, and mature trees and shrubs of the dominant woody taxa at each successional stage support the turning points model? (ii) How does the entire plant community change with time at each hypothesized turning point? (iii) Do we see evidence of directional and synchronous shifts in species composition across successional stages? We conclude that some aspects of vegetation change during the last 25 years were consistent with the turning points model; however, many changes were not consistent, indicating the potential roles of biological, environmental, landscape, and climate controls in vegetation patterns.
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  • 91
    Publication Date: 2010-03-01
    Description: Litter quality is often considered the main driver of decomposition rate. The objective of this study was to investigate the relative contribution of two other tree-driven mechanisms, litter mixing and forest floor conditions, to foliar litter decomposition and nutrient dynamics for trembling aspen ( Populus tremuloides Michx.) and black spruce ( Picea mariana (Mill.) BSP) using a microcosm approach. Results based on mixed linear models show that the greater influence over these processes was obtained through litter quality followed by forest floor conditions and litter mixing. Specifically, the results indicate that significantly more C and nutrients were mineralized (i) from aspen than from spruce litter, (ii) from spruce litter in mixture with aspen litter than from spruce litter applied singly, and (iii) from litter incubated on forest floor from the aspen stand rather than from the spruce stand, except for nutrients in the spruce litter. Collectively, our results show that the litter and forest floor material from aspen both favour decomposition and nutrient mineralization processes. Hence, we provide evidence that the effect of tree species on litter decomposition may not only be caused by the properties of its litter but also, indirectly, by the specific conditions and the decomposer community that tree species develop in their forest floor.
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  • 92
    Publication Date: 2010-02-01
    Description: Using in situ light curves and understory seasonal light measurements the importance of canopy avoidance was evaluated for a population of sugar maple ( Acer saccharum Marsh.) seedlings living at the boreal ecotone in Lake Superior Provincial Park, Ontario, Canada. At higher latitudes, the time period associated with canopy avoidance is shorter and occurs at cooler temperatures, increasing the risk of frost damage. In 2008, leaf-out began 5 days prior to the last frost of the season. By modeling the potential carbon gain of seedlings, it is clear that the short time period prior to canopy leaf-out still contributes a disproportionate amount of carbon to the overall budget of seedlings. Of the total seasonal carbon gain, 80.6% was assimilated in the initial 15 days following sugar maple seedling leaf-out. Based on our model, by leafing out only 6 days earlier than the average seedling leaf-out date, ∼200% more carbon could be assimilated during the course of the growing season. Vegetation phenology is cued by climatic triggers — as climate changes so too will phenological responses. These mechanisms add to the overall carbon budget of seedlings; recognizing these mechanisms is essential to thoroughly understand the natural history of this species.
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  • 93
    Publication Date: 1991-11-01
    Description: We report the results of a study of the near infrared reflectance spectra of decaying forest foliage. During the decay process, a broad absorbance feature develops in the 1100–2000 nm region of the near infrared spectrum. The magnitude of this feature is directly related to the age of the material (or to degree of decomposition) and may be useful in determining degree of decay in field samples. More specifically, multiple linear regression equations derived from second-derivative near infrared reflectance spectra are presented that predict the concentrations of nitrogen, lignin, and cellulose in decaying foliage. We conclude that near infrared reflectance spectroscopy is a very viable and attractive method for the simultaneous determination of these components in decaying foliage.
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  • 94
    Publication Date: 2010-12-01
    Description: The extent and severity of overstory lodgepole pine ( Pinus contorta var. latifolia Engelm. ex Wats.) mortality from mountain pine beetle ( Dendroctonus ponderosae Hopkins) has created management concerns associated with forest regeneration, wildfire risk, human safety, and scenic, wildlife, and watershed resources in western North America. Owing to the unprecedented nature of the outbreak and associated management in the southern Rocky Mountains, it is unknown if the forests that regenerate after this current period of extensive change will differ from those that regenerated in the past. Here, we compare the density and species composition of post-harvest seedling recruits in pre-outbreak (1980–1996) and outbreak stands (2002–2007). Lodgepole pine accounted for more than 95% of post-harvest seedling recruitment and the density of seedlings colonizing clearcuts was equal during both the pre-outbreak and outbreak periods. Compared with harvested areas, the density of tree regeneration was 75% lower in uncut forests and was more evenly distributed among subalpine fir ( Abies lasiocarpa (Hook.) Nutt.) and lodgepole pine. This comparison provides evidence that the density of seedling recruitment will be at least as high after extensive pine beetle caused mortality as under healthy, pre-outbreak conditions and that the species composition of stands regenerating after this outbreak will differ between treated and untreated areas.
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  • 95
    Publication Date: 2010-04-01
    Description: CO2 efflux from forest soils is an important process in the global carbon cycle; however, effects of stand age and successional status remain uncertain. We compared soil respiration and its relationship to soil carbon content, forest floor mass, root biomass, soil temperature, and soil moisture content among three temperate forest ecosystems in Changbai Mountains, northeastern China, from 2003 to 2005. Forest types included an old-growth, mixed coniferous and broad-leaved primary forest (MN), a middle-aged, broad-leaved secondary forest (BL), and a young coniferous plantation forest (CP). Average annual soil CO2 efflux at BL (1477.9 ± 61.8 g C·m–2·year–1) was significantly higher than at CP (830.7 ± 48.7 g C·m–2·year–1) and MN (935.4 ± 53.3 g C·m–2·year–1). Differences in soil temperature among those sites were not statistically significant but contributed to the differences in annual CO2 efflux. In addition, the temperature response of soil CO2 efflux was higher at MN (Q10 = 2.78) than that at BL (Q10 = 2.17) and CP (Q10 = 2.02). Our results suggest that successional stage affects soil respiration by the differences in substrate quantity and quality, environmental conditions, and root respiration.
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  • 96
    Publication Date: 1991-07-01
    Description: Treatment effects over time are frequently investigated using repeated measures designs, but analyses of these experiments frequently fail to address a primary objective of collecting data over time, namely description of the response curve. The analysis advocated in this paper utilizes the intrinsic continuity of the repeated measures factor by focusing on response curves. Treatments are compared by analyzing estimated coefficients of response curves proposed by the investigator. This approach provides more information on treatment effects than analyses that compare treatments separately at each time period. Analysis of estimated coefficients is easier to interpret than multivariate analyses of variance and does not require often biologically implausible assumptions of split-plot analyses currently in vogue. An example describing effects of aluminum on sugar maple (Acersaccharum Marsh.) seedling growth illustrates the method.
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  • 97
    Publication Date: 2010-04-01
    Description: As a hand-held, portable tool capable of assessing wood density of standing trees quickly and inexpensively, the Resistograph holds considerable potential application for evaluating forest genetics field tests. However, phenotypic correlations between Resistograph density index values and values from conventional wood density assessment techniques are weak. In an effort to investigate the extent to which environmental, operator, or instrument factors may affect density index values, we evaluated the sensitivity of Resistograph measurements to seven experimental factors. Drill bit flexion (a measure of operator steadiness), moisture content of wood, and air temperature significantly affected Resistograph density index values, while the influence of knots is minimized at a vertical distance of 3 cm. Battery type, sharpness of the drill bit (at least up to 350 uses), and battery charge (at least up to 310 uses with a 12 V motorcycle battery) had no significant effect on density index. By ensuring that the operator remains steady while drilling, sampling only live trees, only when air temperatures are above freezing, and by drilling at least 3 cm vertical distance from knots, measurement error should be minimized. Three measurements per tree are required to estimate density index to within 3 units 19 times out of 20.
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  • 98
    Publication Date: 2010-03-01
    Description: Predicted climate change is expected to significantly affect tree growth in many areas. We used a process-based model (Physiological Principles for Predicting Growth, 3-PG) to evaluate how climatic variation might alter growth of Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco var. glauca (Beissn.) Franco and Pseudotsuga menziesii (Mirb.) Franco var. menziesii ) across biogeoclimatic zones in British Columbia. The results indicate that there will be significant changes in site index (defined as the height (in metres) of dominant trees at 50 years) over this century. In the interior, a reduction in site index is likely, particularly in stands with mid-range values of site index (25–30 m), with many of the interior bioecoclimatic zones predicted to experience a gradual mean decrease in site index by up to 10%. Individual sites may decrease by as much as 40% from current values. In contrast, site index along the coast overall is predicted to increase to a maximum of 43 m by 2080. In the Coastal Western Hemlock zone, however, mean site index is likely to increase from 26 m to only 34 m. We believe that combining process-based models with fine-spatial resolution climate forecasts offers a viable approach to assess future changes in forest productivity.
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  • 99
    Publication Date: 2010-09-01
    Description: This study examines the potential impacts of changes in federal timber harvest, acting through regional log markets, on the sequestration of carbon in forests and forest products in western Oregon. We construct a dynamic model of the region’s log markets in which market prices, log consumption at mills, and timber harvests and timber inventories on private, federal, and state forests are endogenous. Absent any policies regulating forest carbon sequestration, simulations show that regional carbon flux in forests and forest products would gradually decline as federal harvest rises from recent historical levels. If regional forest carbon flux were constrained to meet some minimum target, however, projections indicate that there would be opportunities for substituting carbon sequestration between federal and nonfederal lands through coordination of harvests across ownerships. We find that relatively small reductions in average private harvest could offset substantial losses of carbon flux on federal timberlands caused by increased federal harvest. One mechanism for achieving the changes needed in private harvest to meet a regional carbon flux target would be a carbon tax/subsidy program or a carbon offset market. For example, if federal owners offered timber for sale equal to the maximum sustainable level under the Northwest Forest Plan, our analysis indicates that a carbon price of roughly $US 19 per tonne of carbon would be sufficient to induce private owners to undertake the harvest and management modifications necessary to maintain regional forest carbon flux at its level in the early 2000s.
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  • 100
    Publication Date: 2010-05-01
    Description: Living trees in declining Austrocedrus chilensis (D.Don) Pic. Sern. et Bizarri forests exhibit cambial mortality or death of part of the cambium causing incomplete formation of rings around the circumference, which represents a potential source of error when determining tree ages and years of death. We sampled 12 stands to quantify the incidence of cambial mortality and tested whether its occurrence is independent of the health condition and canopy position of living trees. Trees were cored and statistically cross-dated. Cambial mortality was identified when the outer-ring date differed from the year of sampling. Of the 811 trees sampled, 307 exhibited cambial mortality with duration ranging from 1 to 39 years. Cambial mortality was most common in subcanopy trees independent of their health condition. Symptomatic canopy trees also exhibited a greater than expected incidence of cambial mortality. The presence of cambial mortality represented a source of error when ages were estimated from ring counts, reiterating the importance of cross-dating for determining accurate tree ages. Cambial mortality also poses complex challenges for estimating the years of tree death. Using the cumulative frequency of cambial mortality duration, we present a novel method for estimating error in tree ages from ring counts and selecting class widths to more accurately depict age structures and mortality rates.
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