ALBERT

Description: The sedimentation regime off Northwest Africa is shaped by: (1) structur~al factors. which result in generallv low relief on land. shelf widths between 40 and more than 120 km. and av-erage sfope inclinations between 10 30' and 30; (2) land climates. which contral the delivery of terrigenous particles to the margin: (3) water movements including boundary currents and upwelling; and (4) the post- Pleistocene sea level rise. This chapter combines published and new results arising from research into the sedimentation processes off Northwest Africa. and emphasizes particularly the activities of the Kiel marine geological group during the past few years. Reviews of cruise activities and results were given in Closs et al. (1969) (Meteor cruise 8. 1967. off Morocco) . Seibold (1972) (Meteor cmise 25 . 1971. off Sahara to Central Senegal). Seibold and Hinz (1976) (Meteor cmise 39,1975 . and Valdivia cruise 10. 1975, from Morocco to South Senegal), and Waiden et al. (1974) (Meteor cmise 30, 1973, off Sierra Leone). Some of these cmises were used for pre- or post-site surveys for the Deep-Sea Drilling Project, or to add undisturbed Quaternary cores to the Glomar Challenger cores (leg 41, ] 975; Lancelot, et al .• 1978); leg 47 A, Arthur er al .• 1979; Lutze et al., 1979). We have concentrated our geological investigations on a number of standard profiles from the shelf to the upper continental rise as given in Figure 1. The manuscript was finished May 1979.

Description: https://www.researchgate.net/publication/230891291_The_Orbital_Theory_of_Pleistocene_Climate_Support_frim_a_Revised_Chronology_of_the_Marine_d18O_Record

Description: Summary Holocene sediments of the North Lagoon, Bermuda, were studied with shallow seismic reflection profiles (200 km CSP-survey, UNIBOOM-system) and vibration coring (40 sediment cores, pneumatic vibration corer, Meischner et al., 1981). Seismic Stratigraphy Four seismic sequences are distinguishable by seismic stratigraphy. All seismic sequences correspond to depositional sequences built up during high sea levels in interglacial times. The seismic sequences are separated by unconformities which are often strongly reflective and correspond to emersion planes during glacial phases. The upper sequence (sequence 4) is related to Holocene sediments. The pre-Holocene bedrock is divided into three different seismic sequences (Kuhn et al., 1981): Sequence 1: oldest Pleistocene sequence (pre-Sangamon sea-level highstands), upper boundary with levelled relief (lower boundary not discernible), composed of strongly cemented carbonate sediments, forms the bedrock below Three Hill Shoals Sequence 2: Sangamon (125 ky sea-level highstand), distinct surface morphology, forms the bedrock of a large area below Holocene sediments, Holocene reefs grew up on elevations of the sequence 2 surface, the Holocene reef rim was developed on an elevated rim of sequence 2 Sequence 3: youngest Pleistocene sequence (Sangamon, 105 and 85 ky sealevel highstands lower than recent), deposited mainly in depressions of the bedrock deeper than -15 m below recent Mean Sea Level, levelling the older relief, peat sedimentation in places The distribution of recent reef areas and lagoonal basins is strongly controlled by pre-Holocene topography and geology of the bedrock. During the Holocene approx. 1050 x 106 m3 of carbonate sediments were deposited in the North Lagoon (290 km2) and approx. 1350 x 106 m3 in the reef rim area (170 km2). Sedimentology There are no larger oscillations of the Holocene sea level identifiable in the sedimentological record. The pre-Holocene topography was gradually drowned during the Holocene sea-level rise. At first, the depositional depressions were separated and landlocked. Fresh water peat marshes, fresh water ponds, marine ponds and bays were formed. With rising sea level, the land barriers were more and more eroded, drowned and lost their influence on the back-barrier sedimentation area. Autochthonous and allochthonous peat, lime gyttja and carbonate mud are a typical transgressive back-barrier sediment sequence. After destruction of the barrier, the depositional milieu changed from restricted marine to normal marine, open lagoonal. Sea-grass sediments and nearly mud-free carbonate sand were deposited in shallow water in an exposed environment. Hydrodynamic energy decreases with increasing water depth in the lagoonal basin. A more densely growing reef rim and intralagoonal reef growth added to the protection of the deeper lagoonal floors. Fine-grained sediments were deposited in this environment. They are distributed over a large area of the North Lagoon and form the top of the transgressive lagoonal sediment sequence. Holocene reefs mainly developed on rises of the pre-Holocene surface. In the early Holocene, solid reef build-ups were able to keep up with the rapid rise of sea level. Sand pockets in the reefs were left behind and filled up mainly in the later Holocene. The percentage of fine-grained sediments, produced and resuspended in the reef rim and deposited in the near lagoonal back-reef zone, increased during the Holocene. Two models of Holocene sedimentation in a depression and on an elevation of the pre-Holocene surface illustrate the dependence of vertical facies gradation on pre-Holocene topography. Trends of the mostly polymodal grain-size distributions of the Holocene sediments are a coarsening-upward in the back-barrier and a fining-upward in the lagoonal sediment sequences. Change in the composition of the molluscan fauna in the Holocene sediments (particle size 〉 2000 µm) is an Indication for fades changes. Gastropods are abundant in the basal backbarrier sediments. Bivalves are rare and their diversity 1s low. Sea-grass sediments contain Codakia orbicularis and Astraea phoebia shells. In the sheltered lagoonal environment shell fragments 〉 2000 µm become rare, common species are Gouldia cerina, Pitar fulminata and Finella sp. (approx. 1000 µm). Fine-grained reef-rim derived sediments differ from lagoonal sediments by a higher percentage of Homotrema rubrum fragments and Alcyonaria spicules.

Description: Twelve species of terricolous microlichens from the Angmagssalik District, Southeast Greenland, are reported: Caloplaca friesii, C. livida, Lecanora boligera, Lecidea oligotropha and Leciophysma arctophila, which are new to the lichen flora of Greenland, Rinodina conradi, which is new to the eastcoast, and Baeomyces roseus, B. rufus, Buellia geophila, B. punctata, Caloplaca tornoensis and Mycoblastus tornoensis, new to Southeast Greenland. In a discussion of the greenlandic distribution, unpublished records from the herbarium of Copenhagen (C) are incorporated. Notes on the habitats are given and the pertinent phytosociological units indicated. Some morphological and anatomical characters are commented upon briefly.

Description: The three species Galium silvaticum L., Galium aristatum L. and Galium schultesii Vest show differences in morphology, cytology and geographical distribution. These differences are described and discussed. Crossing experiments between the three species were without results. No hybrid could be obtained. Galium silvaticum, Galium aristatum and Galium schultesii must be considered as separate species.

Description: Although a large number of Tortricoid species and several genera from the Indo-Malayan region have been described by earlier authors (Walker, Snellen, Walsingham, Meyrick, and a few others), no survey of the present group has ever been made. Edward Meyrick, the author of most of the new names, has never attempted a synopsis of the Olethreutinae. He made surveys of the Australian and New Zealand Tortricoidea (1911), but the results are too superficial for our modern standards. During a long sojourn, working and collecting in Java, I became greatly fascinated by that fauna. Having completed a number of preliminary studies of the subfamily Tortricinae (1939 et seq.), I turned next to the South Asiatic, especially Javanese, Olethreutinae. After a long delay due to World War II, their revision has been initiated by the study of the two then least known and most confused genera, Bactra Stephens and Lobesia Guenée (Diakonoff, 1950 et seq.).

Description: The 16 recognized species of Sorocea are listed with their synonyms and distribution. Two new taxa are described: S. steinbachii C.C. Berg and S. hirtella Mildbread ssp. oligotricha Akkermans & Berg. Three new combinations are made: S. muriculata Miquel ssp. uaupensis (Baillon) C.C. Berg, S. trophoides W. Burger ssp. rhodorachis (Cuatrecasas) C.C. Berg, and S. sprucei (Baillon) Macbride ssp. saxicola (Hassler) C.C. Berg. A key to the species is presented.

Description: From a bryological point of view Greenland is not well known in detail. Unlike Siberia or Arctic America, it is not known through a few large collections, but through multitudes of mostly relatively small gatherings. In this arctic island that is almost 2000 miles long and extends from below 60° N.lat. to nearly 84° N.lat., travel and logistics are difficult. Consequently no over-all study of the whole island has been made, although many collections have been achieved through casual or intensive studies of small areas. Like the other contributions to the knowledge of the bryoflora of Greenland this paper will only deal with a rather limited area; the Angmagssalik district on the East coast of Greenland, ranging from 65° N.lat. to 67°20' N.lat. In 1887 Lange and Jensen published the first and until now the only comprehensive review of specimens and publications on the Musci of Greenland. In their paper the first moss collection from the Angmagssalik area was reported, made in 1884-1885 in the course of ethnographical studies by Gustav Holm (Sphagnum girgensohnii and Polytrichum juniperinum), the first European to visit this part of East Greenland.

Description: Some rain and savanna forests of western Suriname (Corantijn R., Winana Creek; Upper Marataka R.; Upper Nickerie R;) were studied and their composition was compared with that of forests of other parts of Suriname and Guyana. The savanna forests of western Suriname proved to be much related to Guyanan ( Walabaand Dakama-) savanna forests as described by Davis & Richards (1934) and Fanshawe (1952). On the other hand, there was less relationship as regards rain forests of western Suriname when compared with ones of Guyana and other parts of Suriname, except for the Demerara greenheart forest of the Upper Marataka R., which was closely related to the Demerara greenheart forests of Guyana as described by Davis & Richards (1934). In addition an upland rain forest was studied near Blanche Marie falls, Upper Nickerie R., which proved to be very much like that of the Stofbroekoe Mts., eastern Suriname, as described by Schils (1960). Species/area curves for some rain and savanna forests are given. The geographical distribution of some common western Surinam tree species was studied; of the seventeen species studied one was endemic for Suriname. An annotated list of all species of trees and palms occurring in the explored areas is provided.

Description: Contrary to Europe, with only one Caltha species, North America has at least three species of this genus. These are the polymorphic C. palustris L., also widely distributed in Europe, the floating aquatic C. natans Pall, and the polymorphic C. leptosepala-biflora group. Two previous papers (Smit 1967, 1968) dealt with taxonomic aspects of C. palustris, that in North America were not essentially different from those of European material.

Description: Protium Pullei Swart n.sp. Arbor circ. 12 m alta. Ramuli robusti 4 mm diam. teretes glabri fusci lenticellis oblongis ferrugineis muniti. Folia trifoliolata 17 (16—21) cm longa glabra, petiolis robustis semiteretibus 4.5 cm longis basi incrassatis demum transverse rimosis, petiolulis semiteretibus robustis utrinque subincrassatis 1 cm longis sed terminalibus 2.25 cm longis, foliolis oblongo-ellipticis II (7.5—13) cm longis 5 (3.75—5.5) cm latis, apice abruptius acuminatis, acumine sublineari 8 (5—10) mm longo 2.5 (2—3) mm lato, basi cuneata, margine integro, coriaceis utrinque nitidis laevibus supra glaucescentis infra viridis, nervis secundariis utrinque II, nervis prim. et sec. utrinque prominentibus. Inflorescendae axillares breves pauce ramosae pauciflorae circ. 1 cm longae. Ramuli teretes striati cum pedicellis teretibus flore aequilongis bracteis bracteolisque triangularibus obtusis densiuscule puberulis. Flores 5-meri glabri. Calyx cupuliformis lobis oblongo-triangularibus acutis tubo aequilongis. Petala valvata oblongo-triangularia acuto apiculo inflexo carnosa. Stamina 10. Discus 10-lobis glaber. Pistillum glabrum, ovario late ovoideo stigmate 5-lobo coronato. Type: Maguire 24784 in herb. NY, 17 Sept. 1944, Suriname, Tafelberg, mixed transition high-low bush, 5 km S.W. of Savanna I.

Description: The chromosome numbers of 16 species of Angiosperms, collected in Cameroun and the Ivory Coast, were determined. The numbers given for 14 species are new, in the remaining species the results of other authors could be confirmed.

Description: Three sections with a total number of four species of the genus Phyllanthus have been examined. The pollen grains show a strong resemblance to each other and also the taxonomic arguments to differentiate between the three sections proved to be rather weak. Because of both palynological and taxonomic reasons the sections Ceramanthus Baillon, Cluytiopsis Mueller Arg. and Anisolobium Mueller Arg. have been united into one section; viz. section Ceramanthus Baillon s.l.

Description: Neohattoria Kamim. is a monotypic genus of the Jubulaceae (= Frullaniaceae) with a single species, N. herzogii (Hatt.) Kamim., known from central to northern Japan and the southern part of the Kurile Islands. The present genus was segregated from Frullania by Kamimura (1961; sub. nom. Hattoria Kamim. nom. illeg., non Schust., 1961) on the basis of the branching type, the shape of the first leaf and underleaf on branch, the total lack of secondary pigmentation, the uniform cell structure of the stem in cross section, and the strongly toothed leaf lobes. The generic concept of Neohattoria was greatly expanded by Schuster (1970), who included eight species and classified them into two subgenera, subgen. Neohattoria (with a single species) and subgen. Microfrullania Schust. (with seven species); however, Hattori et al. (1972) transferred all species of subgen. Microfrullania to a newly segregated genus Schusterella Hatt. et al., thus retaining the monotypic status of Neohattoria. As already described and illustrated by Hattori (1955), Kamimura (1961), Mizutani (1961), Ladyzhenskaja (1963), Schuster (1970), and Hattori et al. (1972), Neohattoria herzogii is closely related to species of both Jubula and Frullania. Regarding the taxonomic desposition of Neohattoria, Mizutani (1961) and Mizutani & Hattori (1969) placed it with Jubula in a subfamily Jubuloideae of Lejeuneaceae and Hattori et al. placed it in Jubulaceae (s. lat.). But, Kamimura (1961), Schuster (1970, 1979), and Guercke (1978) placed it more close to Frullania, e.g. in a subfamily Frullanioideae of Jubulaceae (s. lat.); more recently, Asakawa et al. (1979b), admitting three distinct families, Jubulaceae, Frullaniaceae, and Lejeuneaceae, placed Neohattoria and Jubula in the Jubulaceae (s. str.) but Frullania and Schusterella in the Frullaniaceae.

Description: Zeer geachte Toehoorderessen en Toehoorders, Bij het beginnen van een wetenschappelijk onderzoek zal meestal degene die zich daaraan gezet heeft, allereerst het antwoord dienen te vinden op enkele fundamentele vragen. Zijn deze primaire vragen beantwoord, dan is de weg gebaand voor verder onderzoek en voor algemene theoretische beschouwingen. Deze fundamentele vragen zijn echter niet voor elke onderzoeker en ook niet voor elk onderzoek in een zelfde tak van wetenschap steeds gelijk. Dit hangt af van vele factoren, zoals: uiteindelijk doel van de studie, aard van het materiaal, geaardheid vooral van de onderzoeker, enz. Vandaag wil ik met U behandelen de hoofdvragen, die zich bij mij, voor de aan mij toevertrouwde onderdelen van de botanie steeds op de voorgrond plaatsen en de wijze waarop ik die beantwoorden pleeg te interpreteren voor het verdere onderzoek. Hierdoor zal ik tevens de gelegenheid hebben, om aan te stippen in welke richting wij op het gebied van de bijzondere plantkunde en de plantengeografie nog onderzoekingen kunnen verrichten, die ons inzicht in het geheel aanmerkelijk kunnen verruimen. Voor ik met mijn eigenlijke onderwerp begin, moet ik toch iets zeggen over wat „bijzondere plantkunde” is. Ik zal er niet te veel over uitweiden, daar, zoals Koningsberger en Reinders in het voorwoord van het eerste deel van het Leerboek der Algemeene Plantkunde terecht opmerken, de scheiding tussen „algemene” en „bijzondere” plantkunde uiteraard onscherp is. Volgens de letter van de gebruikelijke terminologie zou eigenlijk alles wat niet „algemeen” is thuis horen onder de bijzondere plantkunde. Zover wil ik niet gaan, want dat zou mijn taak wel heel omvangrijk maken en afgezien van het feit dat het buiten mijn kunnen zou komen te vallen, denk ik ook dat mijn collega voor de algemene plantkunde ernstige bezwaren zou maken indien ik datgene van de plantenphysiologie dat zeker niet algemeen is te noemen, voor mijzelf zou gaan opeisen. Ik wil daarom beginnen de physiologie, hoe bijzonder deze hier en daar ook moge zijn, maar onmiddellijk in zijn geheel bij de algemene plantkunde te plaatsen. Voor de rest zou ik mijn bovengenoemde definitie, dus „bijzonder” is alles wat niet „algemeen” is, in grote trekken willen volgen, met dien verstande dat ik mij natuurlijk wil houden aan de veelal gebruikelijke taakverdeling, zodat b.v. de „algemene” morphologie en anatomie van de Angiospermen, die in feite in het plantenrijk als geheel, „bijzonder” is, bij de „algemene plantkunde” wordt ondergebracht. De speciale en vergelijkende morphologie van deze groep reken ik echter zeer zeker tot de mij toegewezen tak van wetenschap. Ook de afgrenzing met de genetica is niet scherp. Indien men elk onderzoek waarbij niet uitsluitend op het phaenotype maar ook op het genotype gelet wordt, tot de genetica wil rekenen, dan zal de betrekkelijk jonge experimentele plantensystematiek of biosystematiek geen deel kunnen uitmaken van de bijzondere plantkunde. De genetici zullen het mij wel niet euvel duiden, dat ik ook deze tak van onderzoek laat staan bij de bijzondere plantkunde, waaruit zij is voortgekomen en waarvoor zij van zoveel betekenis is.

Description: Xylopia surinamensis R. E. Fr. n. sp. — Ramuli novelli dense aureo-ferrugineo-sericei, vetustiores plus minus glabrescentes et cortice densissime lenticellifero punctato vestiti; internodia 0,5—1 cm longa. Foliorum petiolus sericeo-tomentosus, 5—7 mm longus; lamina rigida, anguste lanceolata, basi rotundato-truncata, apicem versus sensim longeque attenuata, summo apice obtusa, supra ab initio glaberrima sed densissime verruculoso-punctata, subtus dense argenteosericea, 8—11 cm longa et 2—2,5 cm lata; costa supra valde impressa glaberrima, subtus prominens teres; nervi secundarii cum venulis vix conspicui. Flores in inflorescentiis densis nonnulli; bracteae numerosae, late ovatae, 1,5—2 mm longae. Sepala fere omnino coalita, calycem cupuliformem semiglobosum argenteo-sericeum 2—3 mm altum et 5—6 mm latum formantia. Petala exteriora linearia obtusa, extus argenteo-sericea, circ. 13 mm longa et 2 mm lata, prope basin subito dilatata; interiora linearia, acuta, quadrangulari-prismatica, utrinque cinereo-puberula, 12 mm longa et 1—1,2 mm crassa. Staminum filamenta 0,2 mm longa; antherae circ. 1 mm longae, locellatae, connectivi discus glaber; stamina exteriora plus minus sterilia. Pistilla numerosa (fere 30); ovaria dense sericea, circ. 1 mm longa, styli 1 mm et stigmata 1 mm longa. (Fructus ignotus). Suriname: Boschreserve, Sectio O (florifera Junio 1944. — Wood Herbarium Surinam No. 139). Typus speciei in Herb. Utrecht).

Description: As an introduction to a number of researches of his own the author wishes to give the following data: „Veen” has two meanings in Dutch: 1. in a petrographic sense (peat) Von Büllow’s definition was accepted: „Torf” ist zu deflnieren als ein meist dunkles, kohlenstoffreiches und ± saures Gemenge unvollständig spezifisch-zersetzter Pflanzenteile, das erdgeschichtlich jüngste Glied der Verwantschaftsreihe der Kohlen, dessen Bildung noch heute andauert.” 2. in a plant-sociological-geographic sense (bog) the following definition has been suggested: a bog is a plot, the surface of which consists of a layer of peat, either covered or not with vegetation, with which that layer is genetically connected. The classification of bogs according to their position with regard to the water-level of the surroundings (Staring) and that of the geological chart were rejected on account of their ambiguous character. The classification suggested by Van Baren according to the environment in which the bogs have been formed, was likewise thought insufficient. Preference was given to the classification according to the plants which gave rise to the peat (eutrophic, mesotrophic and oligotrophic bogs) and according to the origin of the water needed for peat formation (topogenous, ombrogenous and soligenous bogs). The conditions of peat-formation are of a botanical (presence of a vegetation and micro-organisms), climatologic (presence of a certain temperature and moisture) and geological nature (presence of a basin, valley or dead river-branch, certain level of ground water, a possible impervious layer). With reference to a number of authors (Picardt; Van Lier; Grisebach, Venema and Staring; Weber) the alteration in conception as to peatformation from the 17th via the 18th and 19th to the 20th century has been given. The word „Peel” cannot be derived from „palus”. Nothing is certain about its origin. It may mean the low land, bog or marsh. The bogs of the Peel lie on the Brabant-Limburg border-plateau (fig. 2). Lorié and Pannekoek van Rheden have shown that the peatformation of the Peel is likely to have occurred in channels, which have been formed by the Meuse, in co-operation with wind and rain (fig. 4). The bogs were therefore in the first instance topogenous formations, which afterwards developed into ombrogenous bogs. For his own research the author collected peat in three ways: 1. by cutting lumps of peat from open profiles; 2. by boring with a simple peat-bore (photograph 1); 3. by boring with the Utrecht peat-bore, an improvement on Dachnowski’s (fig. 5). To assist in the pollen-analytic examination the samples were treated according to Erdtman’s method. The latter has the following advantages compared with the usual treatment with a 10% KOH-solution: 1. the surface-structures of the pollen-grains are more distinct and as a result the grains themselves can be recognized better; 2. the pollen is more concentrated, so that in spite of the method taking up much time, a saving of time is possible. How the method is applied may be found in the chapter concerned (p. 38 and following). For the stratigraphic examination the samples were broken apart in a glass-bowl of water and viewed with a binocular microscope. Dry sandy samples were broken in water, when seeds and other vegetative parts came floating to the top; next they were put with a brush on thick blotting paper and studied through the binocular microscope. The designations for the sediments and species of peat have been derived from Fægri & Gams. For Scheuchzeria peat a new designation has been added. A plea was made for replacing the word pollen-analysis by „palynology”. A survey of the observations and examinations up to abt. 1935 closes the introduction (see the diagrams of Weber, Erdtman and Duyfjes in the figs. 6, 7, 8 and 9). The author’s own research refers to the Southern and Astense Peel, as in the remaining grounds of the geological chart indicated I 4v (= raised bog) no samples could be taken owing to the digging off having progressed too far. 10 profiles were examined. The situation of the bore-sites has been given in the geological chart of the grounds (fig. 3). The result of the examination (figs. 10—27) and the discussion on it may be summerized as follows: Zoning of pollen-diagrams The sub-zoning of the late- and post-glacial periods according to Blytt & Sernander has proved useful as a zoning of pollen-diagrams, provided atlantic and sub-boreal are joined. It is desirable to replace Blytt & Sernander’s terminology by a different one, because the authors gave a climatologic connotation to their names of periods. The limit between pleistocene and holocene was drawn between preboreal and boreal as Florschütz did. As phases of the holocene the following names were suggested: young post-glacial = sub-atlantic mid post-glacial = sub-boreal and atlantic old post-glacial = boreal. Neither in the Peel nor elsewhere in Holland have Allerød-deposits been found. They are not likely to be found either, as on account of the long distance from the land-ice-margin the flora will have been hardly or not at all influenced by the Allerød interstadial period. For Holland therefore the zoning of the late-glacial according to Firbas (1935) may be considered sufficient. The names of the periods do not bear a climatologic connotation as those of the post-glacial phases do. For the sake of a unity the following names have been suggested: young late-glacial = pre-boreal mid late-glacial = sub-arctic period old late-glacial = arctic period. Forest-history In a table (p. 98), in which likewise the Peel diagrams of Weber, Erdtman and Duyfjes have been inserted, the examined profiles have been arranged from North to South. From each profile it has been stated whether it originated in a certain period (+) or not (—). The sub-arctic phase was characterized by forests of Betula and Pinus and was followed by the pre-boreal phase, in which Corylus and Alnus occurred. Also from the other Dutch diagrams (see list on p. 99) it appeared that in the Netherlands the Alnus pollen occurs with an equal frequency before, during and after that of the Quercetum mixtum. The old post-glacial zone of the diagrams shows a peak in the Pinusline. In contrast with the from Mid-Europe there is not always a maximum in the Corylus-curve after the Pinus-peak. In other Dutch diagrams this phenomenon is likewise found. Only in 28% of all Dutch profiles with a boreal zone does a hazel-maximum succeed a Pinus one. They often co-incide (16%), while in the remaining cases no hazelpeak has been established. There is no fixed order of sequence in the occurrence of the components of the Quercetum mixtum, either in the Peel or elsewhere in Holland. The mid post-glacial is the phase of culmination of warmth-loving forest elements: Alnus pollen shows the highest percentage in this zone. Quercus pollen also occurs in great quantities, while Ulmus and Tilia take up an important place up to the „Grenzhorizont”. The absolute and empiric Fagus pollen limits are found at different heights in the mid post-glacial zone of the diagrams, the rational limit lies somewhere near the „Grenzhorizont”. In the young post-glacial phase the Fagus pollen attains fairly high percentages (up to 30%). The maxima in the East and South-east of the Netherlands are between 20% and 38%; they decrease towards the coast and increase towards the South-east (Hautes Fagnes, Belgium) and East (Germany). It seems incorrect to class the Netherlands almost entirely among the oak-alderterritory poor in beeches, as Firbas did. An attempt has been made to fit the Peel-diagrams into Overbeck & Schneider’s zonation system. For the territory for which it has been made there are already difficulties (p. 104), for use in the Peel and other Dutch diagrams there are even more objections (p. 68, 104). Godwin’s zonation system appeared to be a little less forced, but not quite useful on account of too many details. From his horizons that of Ulmus proved useless for the continent. Neither for the Peel nor for the Netherlands and its surrounding territory can a detailed zonation system be designed. It has proved difficult to proceed any farther than Rudolph’s „Grundsukzession”: birch, pine-hazel-mixed oak-forest-beech, in which the alder generally joins the mixed oak-forest and the hornbeam the beech. Before drawing far-reaching conclusions from the course of the curves (as has been done by some authors) more palynological researches are needed in accordance with the actuality principle, known from geology. Pollen-grains from warmth-loving trees in seemingly sub-arctic spectra In profile 4 (Deurnse Peel II) pollen-grains of Abies, Alnus, Picea, Tilia, Ulmus and Corylus were found in the „late-glacial” zone (figs. 14, 15). Investigations were made as to which of the following possibilities would be the cause of their appearance: 1. in taking and preparing the samples pollution occurred; 2. pollen-transport over long distances has taken place; 3. the pollen-grains found have got secondarily into the deposit; 4. warmth-loving trees have occurred in favourable circumstances in the late-glacial phase or 5. in an interstadial period or in an interglacial phase. The said pollen-grains probably hail from a Würm interstadial or interglacial phase. Interglacial peat On the site of the bore-point 7 it was possible to collect samples from the layers under the peat. The upper 40 cm of the diagram Griendtsveen IX (fig. 27) of this profile proved a repetition of the lower 40 cm of the Griendtsveen I profile (fig. 18). The diagram shows that pollen of Carpinus, Picea and Abies occurs showing the deposit to be of interglacial age. The pollen-curves, however, pass unnoticed from an interglacial into a post-glacial portion. The limit is likely to be found between the two, about 30 cm below the mowing field. There is therefore a great stratigraphic hiatus. Pollen-analytically it could not be decided from which interglacial period the profile hails; on account of its situation on the middle terrace, it was deemed likely that it was an Eem sea deposit. The examined profile probably corresponds to Jessen & Milthers’ zone g; showing it to have been formed at the end of the Eem sea period. The Meuse therefore cannot have flowed through this part of the Astense Peel after the mid Eemean phase. Stratigraphy This is difficult to summarize. Compare various profiles. Individual mention may be made here of: 1. peat on a podsol layer; this was found in two places (Deurnse Peel I Kraaienhut and Griendtsveen VIII). Peat-formation may be thought to have occurred in the following way: heather started growing on drift-sand giving rise to a podsol layer. As the latter is impervious the vegetation surface became marshy. The heath was replaced by a Caricetum from which peat arose. Gradually more Eriophorum occurred, from which almost pure vaginatum peat arose. The bog-surface grew moister and moister, Sphagnum cuspidatum and Scheuchzeria could grow on it and formed a „Vorlaufstorf”. Only then could non-extremehydrophile Sphagna join in peat-formation. 2. the occurrence of Scheuchzeria-peat after the „Grenzhorizont” period. This species of peat, which is often found at the basis of the old Sphagnum-peat as a mesotrophic transition vegetation, has for the Netherlands only been found in the young post-glacial phase in the Peel (Deurnse Peel I Kraaienhut, Griendtsveen V and VIII and Nederweerd). At present the plant is very rare. The severe decline of this plant was also observed elsewhere. Probably it is caused by the gradual drying up or reclaiming of the raised bogs. Of the present station of Scheuchzeria near Ommen a short description has been given (p. 59 and photographs 2, 3, 4). 3. the „Grenzhorizont”. Where the young Sphagnum-peat has not been dug for the preparation of moss-litter, the Peel bogs show a clear „Grenzhorizont” (photograph 8). The conceptions about its origin have been discussed. The distinct separation between the old and the young Sphagnum-peat was not considered sufficiently explained. Though on the whole the „Grenzhorizont” is synchronous in the North-west European profiles, the point of transition from old to young Sphagnumpeat was fairly unstable and easily changeable as to time. Generally the date of the „Grenzhorizont” is fixed at about 500 A.D., though there are differences in opinion. There is a lack of archeological correlation which renders a correct dating impossible. Interference of man in the Peel Three ways of interference were stated: 1. peat has been dug off for the greater part in the territory of the Peel: young Sphagnum-peat for the preparation of moss-litter, old Sphagnum-peat for fuel. The trees which appeared when the bog was dug up in the „Veenderij der Maatschappij Griendtsveen” are sometimes in so good a condition, that they are used for building sheds. The 1 st, 2nd and 4th beam in the foreground of the shed in photo 5 has been sawn from a 30 m long subfossil pine. 2. in a native peat-digging it was possible to collect recent young Sphagnum-peat. 40 to 50 years ago the peasants living there had dug peat in holes, which were afterwards left to themselves. Sphagnum started growing again and the holes were filled in again. The diagram (fig. Griendtsveen VII) represents the surrounding heath with scattered pines and birches, sown by the wind, and a pine-plantation close by. 3. in the profiles Nieuwe Peel, Griendtsveen VI and VII it has been fixed by the indications given by Firbas, that only in the surface layers of the bog has corn-pollen occurred. So in these parts cultivation of cereals will be of recent date. This also appeared from the history of the reclamation of the said territory.

Description: The chromosome number of 157 species of Angiospermae occurring in the Netherlands is dealt with.

Description: Dicranella riparia (H. Lindb.) Mårt. & Nyh. is reported for the first time from Greenland, where it was found on a fluvioglacial delta in the Angmagssalik District in plant communities belonging to the association Calamagrosto-Ditrichetum (all. Calamagrostion neglectae). This is the sixth locality known, and the first outside Fennoscandia.

Description: Name: Wahlenbergia marginata (Thunb.) DC. Monogr. Camp. (1830) 143. Family: Campanulaceae.

Description: Alangium LAMK.—M. M. J. van Balgooy, Pac. Plant Areas 2: map 72. Complete; Old World, also incl. Indo-Malesia, E. Australia, Pacific (Solomons, New Caledonia, New Hebrides, Fiji); delineated except in Africa and Madagascar, localities indicated only in the Pacific, species density; monograph.

Description: The Upper Hunter palynological assemblage described includes 54 species, assigned to 29 genera. Seven genera (Scabratisporites, Guttatisporites, Lapposisporites, Pseudogravisporites, Paralundbladispora, Taeniaepollenites and Tubantiapollenites) and 39 species are new; three generic descriptions are emended (Apiculatasporites, Colpectopollis and Angustisulcites) and three new combinations are proposed. Bisaccate pollen grains average 84 %; within this group no Upper Permian elements could be demonstrated. The assemblage is compared with other European Lower and Middle Triassic assemblages; there are no close similarities to assemblages outside Europe.

Description: El género Plagiochila (hepatica) esta representada en las Islas Galapagos por ocho (8) especies diferentes: P. bursata (Desv.) Lindenbg., P. galapagona Inoue, P gradsteinii Inoue, P. guilleminiana Mont., P. inouei Grolle, P. scabrifolia Inoue, P. spinifera Ångstr. y P. subplana Lindenbg. El endemismo en este género es más alto que en otros géneros de las hepaticas, con cinco (5) especies que comienzan a conocerse solamente de los Galapagos ( P. galapagona, gradsteinii, scabrifolia, inouei, y spinifera). Las otras tres (3) son comunes y ampliamente distribuidas a lo largo de la America tropical. La mayoría de las especies estan restringidas a las zonas altas-húmedas de vegetación de las Islas Galapagos (matorrales de Zanthoxylum, Miconia y pampa) excepto P. guilleminiana muy común, la cual puede presentarse en la zona seca de transición de bosque. La más amplia variación de Plagiochila ha sido vista en Isabela (Cerro Azul), San Cristobal y Santa Cruz.

Description: The species Polypodium banaense C.Chr. is transferred to Crypsinus. The recognition of a genus Phymatopteris Pic. Ser. (= Phymatopsis J.Sm.) separate from Crypsinus is discussed.

Description: The six species of Curtia, including a hitherto undescribed species published here, as well as the monotypic genus Hockinia can be distinguished from each other by the seed coat structure. The anticlinal walls and the cuticle provide the most useful information. Curtia tenuifolia appears to be a complex species, but subsp. tenella can be readily separated from this complex by the seed coat structure. Heterostyly has been found in C. tenuifolia subsp. tenuifolia, C. obtusifolia, and Hockinia montana, but differences in seed coat structure can not be correlated with long-, short-, and equal-styled flowers. The differences in seed coat structure, the length of the seeds, and the number of cells per seed plead for maintaining Hockinia (closely related to Curtia) as a distinct genus. One new species of Curtia is described and a new combination is made.

Description: SETTEN, A. K. van & KOEK-NOORMAN, J.: Studies in Annonaceae. VI. A leafanatomical survey of genera of Annonaceae in the Neotropics. — Bot. Jahrb. Syst. 108: 17—50. 1986. — ISSN 0006-8152. Within the scope of the multidisciplinary research project on systematics of Annonaceae, a survey of the leafanatomical features and their distribution in the neotropical Annonaceae is presented. The studied specimens form a rather homogeneous group, as may appear from the family description given here. A detailed study of the leafanatomical features reveals, that differences are mainly found in the indument, the position and contents of the idioblasts, the structure of the primary vein, the type of crystals in the epidermal cells, and the type of sclereids. Based on character states, phenetic similarities and differences are discussed and compared with the classifications of FRIES (1959) and WALKER (1971).

Description: One new species of Dorstenia from Brazil is described: D. carautae C.C. Berg, and four new combinations are made: D. cayapia Vellozo subsp. asaroides (Hooker) C.C. Berg, D. cayapia Vellozo subsp. paraguariensis (Hassler) C.C. Berg, D. cayapia Vellozo subsp. vitifolia (Gardner) C.C. Berg, and D. ramosa (Desvaux) Carauta, Valente & Sucre subsp. dolichocaula (Pilger) C.C. Berg. A list of and a key to the 22 Dorstenia species distinguished in south-eastern tropical America are presented, together with synonymy and distributional data.

Description: The samples of the genus Calypogeia in the dutch institutional herbaria and private collections, those of C. arguta excluded, have been re-identified, according to the revision of the Swiss Calypogeias by Bischler (1957); distribution maps are given for all the taxa. More exact circumscriptions are given of several differentiating characters which were already established by previous authors. In C. fissa and C. sphagnicola the areolation of the leaves appeared to be a new differentiating character: in C. fissa the cells in the middle of the leaf show a great variation in length, whereas in C. sphagnicola the cell size is uniform. These differences are shown in histograms. C. muelleriana appeared to be restricted to the diluvial parts of the country, whereas C. fissa is common on both alluvium and diluvium; c. neesiana, C. sphagnicola and C. trichomanis are very rare, so that no clear geographical distribution can be given.

Description: This study deals with the taxa of the section Rubus of the genus Rubus L., so far as they are found in the Guelders district within the flora of the Netherlands. It concerns fifty species and some subspecies and varieties, mainly of the subsections Fruticosi Wimm. et Grab. and Discolores P. J. Müller. The similarity with the bramble-flora of northern Germany is obvious. A number of species, that occur in the latter region are absent however. Species of Central-European hills and mountains are as good as limited to the southern border of the Veluwe, which is mostly considered to belong to the Subcentreuropean district. South-European, often calciphilous species are absent. The nomenclature in the genus Rubus is very confused. There is an abundance of homonyms and synonyms. The number of misidentifications is rather large, meaning that a great deal of the literature is unreliable. The descriptions with many authors are absolutely insufficient, and type-specimens are often with difficulty or not at all to be traced. The difficulties arise from the fact that many taxa are not clearly separated. Some of them are well distinguishable, others are related by transitions. From a geographical point of view there is much difference as well. Some species have as their area almost the whole of Europe, others are limited to a very restricted area. In addition there is a difference in chromosome numbers (from diploids (2n = 14) to hexaploids (2n = 42). Most taxa are tetraploid. The abundance of forms within the section Rubus arises from a partly apomictic, partly amphimictic propagation. To set up some order in all those differences, the author has made the following distinctions: morphologically there are the different ranks of species and infraspecific taxa. Geographically distinctions have been made by means of a code of the capitals A to D inclusive: A indicates the taxa with the largest area, D the local taxa. Cytologically a code of Roman numerals has been given: I for diploids, II for polyploids. Beside the introductory theoretical part a short description of all taxa of the section above the specific rank has been given. All species and infraspecific taxa of this section, that are found to occur in the Guelders district have been described in detail, with mention of the type-specimen. Pictures have been added of the newly described taxa, and of some others as well. Maps of the distribution in the Netherlands of all the taxa have been inserted.

Description: Dames en Heren, In een universitair blad kwam onlangs de mededeling voor, dat aan een hoogleraar, die zich in dezelfde moeilijkheid bevindt als ik, nl. dat hij in de loop van deze cursus 70 jaar is geworden, een afscheidscollege zou worden aangeboden. Ik vond dat een sympathiek plan. Als men met college geven, ondanks de daaraan verbonden bezwaren, de 70-jarige leeftijd heeft gehaald, is het werkelijk geen overbodige weelde dat een ander de taak voor deze laatste keer van hem overneemt.

Description: In a previous palynological study in Cocoid palms the present authors (Punt and Wessels Boer, 1966) were able to demonstrate a rather strong but not absolute correlation between the pollen types and the staminate flower types within the genus Attalea in the broad sense. The existence of partly apparently primitive, partly very advanced flower types within the otherwise close related group of Geonomoid palms made it worthwhile to investigate the same feature in this group. The Geonomoid palms are usually considered to comprise 7-9 genera of monoecious Arecoid palms which share a large number of characteristics (Burret, 1930; Moore, 1966). The group is very obviously a most natural one. The generic distinctions are mainly based on the flower morphology, notably on differences found in the androeceum and pistil.

Description: It is the fate of most historic personalities that in the course of time their work sinks almost completely into oblivion, and that the few lingering reminiscences of their achievements are transmitted to later generations in the form of second- or third-hand quotations, usually mixed with more or less anecdotic episodes from their life. It must be admitted that LINNÉ occupies in this respect a comparatively favourable position, for most educated people will remember that they heard in their school days of at least three things which are credited to him, in the first place that he produced a classification of the plant kingdom which is based on the number of stamens and carpels, the so-called sexual system, in the second place that he was the first who consistently applied the binomial nomenclature, i.e. the custom to designate an organism by a combination of two names, viz. a generic and a specific one, and thirdly that he was the originator of the pronouncement “Species to numeramus quot diversae formae in principio sunt creatae” (We count so many species as in the beginning different forms were created). Other achievements of LINNÉ may have been of greater importance, but it are these three things for which he is most generally remembered. The pronouncement quoted above, which means that the groups of individuals which form the species are descended from ancestors that owed their origin to an act of creation, derives its historic importance from the part it played in the debates on the theory of evolution. As it implies that the species are constant, it became the watchword of the antagonists. It is, however, rather strange that this pronouncement has so often been quoted, for it is found in LINNÉ’s earlier works only, and was in the later ones replaced by another statement that flatly denies the constancy of the species.

Description: This paper is an addendum to the author’s (1971) paper. At the time that the latter paper was finished, there were difficulties in taking photographs of the newly described male fructifications. Subsequently those difficulties have been solved, and the present paper contains the photographs of the male fructifications of the type specimens of Hastystrobus muirii v. Kon., Masculostrobus harrisii v. Kon., and Pityanthus scalbiensis v. Kon., and the photographs of the male fructifications, as described in the above-mentioned paper, of Ginkgo huttoni (Heer) Sternberg and Brachyphyllum crucis Kendall. All specimens are preserved in the Division of Palaeobotany and Palynology, Botanical Museum and Herbarium, State University, Utrecht, The Netherlands. Most of the photographs were taken with the specimens illuminated obliquely in air, but some were taken with the specimens flooded with oil. This procedure is generally applied when the specimen requires enhancement of contrast, so that details are more evident than if the specimen was photographed dry.

Description: Recently a multidisciplinary investigation program on the systematics of Annonaceae was started at Utrecht with special emphasis on the Neotropics. This project will be carried out largely within the framework of the UNESCO-project Flora Neotronica. The first goal is to provide a modern classification of the family as a whole, the second is the publication of a series of monographs for Flora Neotropica. The project has been planned and started in close consultation with leading botanists on the Neotropical flora. The Annonaceae are a family of pantropical distribution with between 2000 and 2500 species in ca. 130 genera as presently understood. In the Neotropics the family is represented by ca. 750 species and 35 genera. It is a family of trees, shrubs, and lianas. Its place is within the order of the Magnoliales and its supposedly closest relative is the family of the Myristicaceae. The Annonaceae, although generally considered primitive in many features, nevertheless offer a number of specialized features as well This makes it a promising object using various kinds of comparative morphological, karyological, and anatomical data. Besides, many species are of medicinal or commercial value, such as various species of Annona and Rollinia, the fruits of which are commonly eaten in most countries of Central America and South America; the Soursop (Annona muricata) is widely cultivated throughout the tropics.

Description: Since the completion of Radlkofer’s monumental work on the Sapindaceae in Engler’s series “Das Pflanzenreich” 50 years have now elapsed, almost 40 since its publication. It is still the basis of virtually all taxonomic studies in the family. Some of the gerontogean genera have since been the subject of revisional work (Leenhouts 1969, 1971), but for the neogean representatives there are only some regional treatments (e.g. Rambo 1952; Barkley 1957; Reitz 1962; Soukup 1969), apart from descriptions of new taxa scattered through the literature. When studying the taxa native to Suriname in connection with the preparation of a supplement to the family treatment published previously in the “Flora of Suriname” (Uittien 1937) it soon became apparent to me that the genus Talisia was particularly incompletely known when Uittien published his account of the family, actually not much more than an extract from Radlkofer’s work. The number of species known or to be expected from Suriname proved to have doubled; this is not due to inadequateness of Uittien’s work but to much more extensive collecting. Two of the species met with since could not be identified with any species dealt with by Radlkofer or described after his time: these are described as new below. In order to establish that they were truly undescribed the descriptions and, where possible, types and/or other authentic specimens of all species described after Radlkofer were checked. A list of these follows; it may serve as a kind of bibliographic supplement to Radlkofer’s monograph. The two species marked with an asterisk have been posthumously listed in the supplement to his work.

Description: The chromosome numbers of 31 species of Angiospermae collected in S. Brazil were determined. Of these species 5 were studied before, the other numbers are new, 11 are first counts for genera and one even for a family. Some notes on the cytology and morphology are added.

Description: Dicranella staphylina Whitehouse, a species recently described from Great Britain, is now recorded from Belgium, Denmark and The Netherlands. A new combination, Anisothecium staphylinum (Whitehouse) Sipman, Rubers & Riemann, is proposed. A study of the costal anatomy revealed that A. staphylinum in this respect most resembles A. rufescens.

Description: The author studied the morphology of Blackstonia perfoliata s.l. and compared its variability with that of the other representatives of the genus. She also carried out ecological studies of “Blackstonia perfoliata ssp. serotina” on the Dutch island Voorne and compared her results with those in the literature relating to B. perfoliata in some adjacent regions, notably the Upper Rhine area. On morphological and ecological grounds B. perfoliata ssp. perfoliata and ssp. serotina are to be regarded as two distinct species, B. perfoliata and B. acuminata.

Description: In a forthcoming publication (Kramer, in prep.) floristic and taxonomic data of the pteridophyte flora of Suriname will be assembled, with keys and notes on their local distribution and ecological preference. The present paper deals with the geographical distribution of Suriname pteridophytes beyond the boundaries of Suriname (Fig. 2), a subject that lies beyond the scope of a local fern Flora. In the past, some (but relatively not very many) authors of fern Floras included a paragraph on the distribution of the taxa (Posthumus, 1928; Christensen, 1932; Backer & Posthumus, 1939). In some other fern Floras some space is devoted to ecology, but very little to geography (Holttum, 1954). In still others, considerations of a general kind on ecology and geography are altogether lacking (Vareschi, 1969). Lyell (1870), in his rather little-known book on the distribution of ferns, tried to bring together all the data known at his time; his work is now, of course, almost exclusively of historical significance.

Description: Erismadelphus Mildbr. is remarkable because it is the only African genus of the Vochysiaceae, a family represented in tropical America by no less than 5 genera and 180 species. Erismadelphus was discovered in 1913 by Prof. J. MILDBREAD and has hitherto been represented by only one species: E. exsul Mildbr. Recent examination of the African collections has, however, revealed the existence of two other taxa. Unfortunately the type of E. exsul (Mildbraed s.n. from Elon, French Cameroons) was destroyed at Berlin during the 1939-45 war and no duplicates or cotypes are known to exist. In response to an enquiry Prof. MILDBREAD, to whom we are very grateful, informed one of us that in his view Corbisier 1362 from Eala in Belgian Congo was identical with his original type. Prof. MILDBRAED and KEAY have, in fact, together examined Corbisier 1362 in the Herbarium of the Jardin Botanique de l’Etat at Bruxelles through the courtesy of Prof. ROBYNS. Duplicates of Corbisier 1362 are at Kew and Paris, they agree in every respect with MILDBRAED’s original description and figure and we therefore propose that this specimen be adopted as the neotype (lectotype).

Description: A new species of Asterophorum, A. mennegae, is described from the Sipaliwini Savanne (Suriname). The position of Asterophorum within the family is briefly reviewed, and a key to the 2 species is given.

Description: The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.

Description: In this preliminary investigation attention was paid to pollen morphology of West-European species of the Rosaceae. Some new terms were used like fastigium, endocingulus etc. The terminology of Iversen and Troels-Smith has been followed in addition to improvements by Erdtman. A key is given to the types and subtypes for the use of pollen analytical investigators. Sanguisorba officinalis appeared to be always 3-colporate and not 6-colporate.

Description: Nanocyperion communities (s.l.) are considered here as “warp-and-woof” communities; the Nanocyperion components are described separately as synusiae. On the Netherlands Frisian Islands, four main synusiae have been recognized. Raunkiaer life form spectra show few differences between the communities. Life strategy spectra of the Nanocyperion synusiae, based on systems for phanerogams (modified after Bakker 1966) and bryophytes, yield the clearest patterns. A comparison of the ecology of the communities and an interpretation of the spectra in terms of avoidance of stress or competition suggest that inundations and standing crop of the communities are the main factors determining the distribution of the synusiae. Winter inundations overrule the influence of differences in productivity level, which becomes prominent in drier situations.

Description: A palynological assemblage obtained from marly limestones of the Dutch Lower Muschelkalk is discussed. A qualitative analysis has disclosed its great resemblance to Upper Bunter (Röt) assemblages ; however, differences in quantitative composition were demonstrated. Utilizing palynological data new information can be added to the knowledge of the European Middle Triassic flora.

Description: The water economy, the mineral content of the soil, and human influence are the principal ecological factors governing the variation of the heath vegetation of a limited region. Sloping of the surface is also an important factor. In hilly country it is of a twofold nature: on the one hand the difference between high and low altitudes, based on the water economy, on the other hand differences in (micro-) climate. If the hills are higher, this results in greater climatic differences. In extremely oceanic and in boreal regions a rise in altitude of 100 m is sufficient for creating a noticeable decrease in temperature and an increase in precipitation, aerial moisture, and wind force. This results in the occurrence on the hills of heath communities that have their main distribution more to the North. The same observation was made by Gimingham (1961). On Slieve League on the Donegal coast (Ireland) Salix herbacea and Lycopodium selago occur in the heath at an altitude of 600 m, near Tongue on the Scottish north coast Dryas octopetala, Saxifraga oppositifolia, Alchemilla alpina and Thalictrum alpinum at an altitude of 60 m. West of Apeldoorn in the Netherlands are found extensive stretches of heath with abundant Vaccinium myrtillus and V. vitis-idaea at an elevation of 60-80 m, even on south-facing slopes. This is an area with high precipitation due to ascending air west of the hill ridge of the Eastern-Veluwe. Here the Vacciniums, elsewhere requiring the protection of the forest, can tolerate the habitat of the open heath (Stoutjesdijk, 1959; De Smidt, 1966). Higher elevation combined with north-facing slopes creates extreme conditions e.g. on Roc Trévézel (300—360 m) in Brittany, with Vaccinium myrtillus, Melampyrum pratense, Hymenophyllum wilsonii and Rhytidiadelphus loreus. These species are virtually lacking in the surrounding plains where the heath consists of such South Atlantic species as Erica cinerea, E. ciliaris, Ulex gallii, Lobelia urens, Lithospermum prostratum and Symethis planifolia.

Description: Frutex epiphyticus, ramis quadrangularibus, angulis lenticellatis. Folia ramorum fertilium petiolis 2-5 mm longis, 1.5-2 mm latis; lamina chartacea vel subcoriacea, oblanceolato-oblonga vel interdum elliptico-oblonga, 9-14 cm longa, 3-4.5 cm lata, ápice acuminata, acumen 1-2 cm longum, basi attenuata, costa subtus prominente, nervis lateralibus supra et subtus prominentibus vel prominulis, glandulis hypophyllis aliquot patelliformibus vel foveolatis, parvis, 3-5 in folii parte inferiore oblique seriatis, aliis minutis, punctiformibus, nigricantibus, aequaliter dispersis. Flores in racemis umbelliformibus (20-) 30-45-floris; rhachis ad circa 1 cm longa; nectaria clavato-cucullata, stipitata, stipes 5-8 mm longus, cucullus 1-1.5 cm longus, circa 4-5 mm diametro, ore late-rotundata, margine plerumque recurvo, apiculata; pedicelli 5-7 cm longi, lenticellati; bracteolae sepaloideae, circa 1 mm longae, 2-3 mm latae; sepala suborbicularia vel reniformia, circa 2-3 mm longa, circa 4-5 mm lata, margine glandulosa; corolla oblongo-subconoidea, circa 1 cm longa, circa 4-5 mm diametro; stamina 18-33, filamentis applanatis, liberis, inaequalibus, in alabastro 4-6 mm longis, antheris linearibus, 3-5 mm longis, circa 1 mm latis, basi subsagittatis; ovarium circa 2-3 mm diametro, 6-11-loculare. Fructus globosus, circa 9 mm diametro, stylo persistenti ornatus. Typus : Costa Rica, vicinity of Vara Blanca, North slope of Central Cordillera, between Poás and Barba Volcanoes, alt. 1700 m, April 1938, Skutch 3762 (holotype US; isotypes GH, MO, NY, S). Paratypes; Costa Rica: Heredia: Cerro de las Caricias, North of San Isidro. Standley & Valerio 52202, 52248,52375 (US); Panama:Chiriquí: Boquetedistrict, Bajo Chorro, Davidson 398 (GH, US).

Description: Although algology deals with a large group of plants, widespread and of a great morphological diversity, the history of this branch of botany is fairly young. Linnaeus (1753) listed in his Species Plantarum under the heading “Cryptogamia—Algae” only five genera of plants which are still accepted as algae at the present time. Under the same heading he also described a number of liverworts, lichens and sponges and a few other things.

Description: The pollen analyse of a raised-bog on the High Vosges crest shows the vegetation regional development since 3200 years. A prehistoric civilization, the Gallo-roman period, the great migrations and the Carolingian period are reflected in the pollen diagram by N.A.P. minima and maxima. A discussion on curves fluctuations of the main A.P. follows.

Description: The subject of this study are soil samples taken in the “Makkumer Waard”, a wide expanse of low-lying land, which follows the Frisian coast. Stratigraphical and palynological investigations showed that in the beginning of the Atlanticum the area lay below the level of the sea, but that gradually the influence of the sea decreased and peat formation became possible. From the transition from marine deposits to Sphagnum peat (— 4,55 m to — 3,50 m) we must conclude that there has been a temporary standstill in the transgression, or even a regression, in the middle of the Atlanticum. Towards the end of the Atlantic period a sudden marine transgression followed, which deposited a layer of sand and clay on the Sphagnum peat (— 3,50 m to — 3,30 m). Shortly before the beginning of the Subboreal (which probably sets in at — 3,10 m) an important regression began and an Eriophorum peat was formed directly on the clay (—3,30 m to —3.00 m). It is probable that the peat formation went on in the Subatlanticum, but the younger Sphagnum peat is no longer present, for a third marine transgression, which lead to the formation of the “Zuiderzee”, washed away the peat and deposited the younger sea sand. The data obtained from the Makkum profile proved to agree very well with the results of other investigators who worked in the area round the North-sea.

Description: Joseph Gaertner (1732-1791) was the first to develop a carpological taxonomy in his book De fructibus et seminibus plantarum (1788-1791). The scope and background of this work are discussed; its history is sketched on the basis of the Banks correspondence at the British Museum; the main sources of material are listed. A brief outline of Gaertner’s life is given, also mainly based on letters from him and his contemporaries to Joseph Banks.

Description: Previously, the chromosome numbers of some species of Loganiaceae were dealt with (Gadella, 1961, 1962, 1963). The chromosome numbers of 7 species are reported in this paper, of which 4 species, all belonging to the genus Strychnos, had not been investigated cytologically before. The materials, kindly supplied to me by Dr. A. J. M. Leeuwenberg and by Ir. F. Breteler, were collected in the form of seed-samples in the Ivory Coast and in Cameroun. The plants of 2 species originate from botanical gardens. Living material of all species (except for Strychnos lernata Gilg. ex Lwb.) is grown in the botanical garden of Wageningen (WAG). The determination of the chromosome numbers was based on the study of roottipmitoses. Roottips of the plants were fixed in Karpechenko, embedded in paraffin, sectioned at 15 micron and stained according to Heidenhain’s haematoxylin method. The results may be summarized as follows: 1. Nuxia floribunda Benth.: 2n = 38 Origin of the material: S. Africa, obtained from the University of Stellenbosch. Seeds probably collected in the wild. Herbarium material of the mother-plant: J. J. Bos, no. 310 (WAG). Herbarium material of seedling: A. J. M. Leeuwenberg no. 3665 (WAG). References: Gadella (1963): 2n = 38. 2. Strychnos dinklagei Gilg.: 2n = 44 Origin of the material: Ivory Coast, Forêt d’Abouabou, between Abidjan and Grand Bassam. Herbarium material of the mother-plant: R. A. A. Oldeman no. 845 (WAG). Herbarium material of the seedling: A. J. M. Leeuwenberg no. 3561 (WAG). References: Gadella (1963); 2n = 44. 3. Strychnos innocua Del. subsp. innocua: 2n = 44 Origin of the material: Ivory Coast, 16 km S. of Ferkéssédougou. Herbarium material of the mother-plant: A. J. M. Leeuwenberg no. 4435 (ABI, WAG). 4. Strychnos millepunctata Leeuwenberg: 2n= 441) Origin of the material: Ivory Coast, Forêt d’Abouabou, between Abidjan and Grand Bassam. Herbarium material of the mother-plant: J. J. F. E. de Wilde & A. J. M. Leeuwenberg no. 3447 (ABI, WAG). 5. Strychnos samba Duvign.: 2n = 44 Origin of the material: Cameroun, 4 km S. of Nguélémendouka. Herbarium material of the mother-plant: F. J. Breteler no. 2051 (WAG). 6. Strychnos spinosa Lam.: 2n = 44 Origin of the material: obtained from the botanical garden of Groningen, the Netherlands, origin unknown. Herbarium material of cutting: A. J. M. Leeuwenberg no. 3564 (WAG). References: Mangenot and Mangenot (1958): 2n = 44. Miège (1960): 2n = 44. Gadella (1962): 2n = 44. 7. Strychnos ternata Gilg. ex Leeuwenberg: 2n = 44 1) Origin of the material: Cameroun, 27 km from Bertoua on road to Bétaré Oya. Herbarium material of the mother-plant: F. J. Breteler no. 2196 (WAG). Herbarium material of the seedling: F. J. Breteler no. 2994 (WAG). Twenty-one species of the genus Strychnos have been studied up to the present. The following chromosome numbers have been counted: 2n = 24 (3 species, counted by Mohrbutter, 1936); 2n = 44 (16 species); 2n = 88 (2 species). From these data the conclusion may be drawn that the more common basic number of the genus Strychnos is X = 11 or 1 = 22.

Description: A suitable subtitle for this paper would have been “The rise and fall of a family”. What is usually called the Cyphellaceae is an instructive example of a situation not uncommonly encountered in the current systematics of mycology: a family retained in a traditional sense by some mycologists and considered by them as good a family as any, while others are convinced that it is nothing but a handy bin from which part of the contents has already been taken out and disposed of by scattering it over various groups, but which is still needed for keeping what remains. We do not yet know what to do with this considerable remainder, mainly because the published accounts are inadequate and the species have not yet been scrutinized anew in the light of present-day taxonomic requirements. In order to understand the basic idea of the Cyphellaceae the type species may be briefly introduced. The fact that Cyphella digitalis was originally described as Peziza digitalis is telling, and one could not do better than characterize it as a ‘discomycete’ with basidia, viz. a cup-shaped fruit-body with the hymenium lining the smooth inside or ‘disk’. If one were pressed to form an opinion about its taxonomic position from a dried, not annotated collection and without the aid of the microscope, one would even now, very likely, dispose of it as a discomycete. However, there is little doubt that in nature the cup is directed downward at least when mature, in contradistinction to the average discomycete in which the hymenium containing the asci is directed upward. This difference is a reflection of the two modes of violent spore discharge inherent in the hymenomycetous basidium and ascus; it has been explained through Buller’s well-known researches. The cups in the various species are not always typically cup-shaped; in a number they are more or less tubular or else more flattened and even disk-like.

Description: Cytological investigations within Galium palustre L. showed the occurrence of three cytotypes, a diploid with 2n=24 chromosomes, a tetraploid with 2n=48 and an octoploid with 2n=96. Comparative morphological investigations, together with transplantation and crossing experiments confirmed the complexity of the species. The cytotypes are here considered to be subspecies of Galium palustre L.

Description: La systématique scientifique, qui classe tous les êtres vivants, actuels et fossiles, d’après leurs affinités naturelles ou présumées telles, doit pouvoir donner à chaque taxon un nom stable, permettant de le désigner avec son rang taxonomique et de l’identifier facilement. La nomenclature botanique n’échappe pas à cette exigence et depuis que C. Linné généralisa, en 1753, la nomenclature binominale, le besoin s’est fait sentir de compléter et de préciser, sur le plan international, les principes et les règles établis par l’illustre botaniste suédois.

Description: The wood descriptions of Juniperus communis L. ssp. communis are compared with those of earlier authors. The average and maximum tracheid lengths and the ray height distribution frequencies offer a means of separating the wood of the erect J. communis L. ssp. communis from that of the subspecies nana Syme with an entirely different habit.

Description: From the general discussion of the effect of mass selection on the genotypic array of the next generation we have seen that under certain conditions mass selection can lead to homozygosity, but does not do so necessarily. Outcrossing, mutations and disadvantages of certain genotypes may prevent reaching equilibrium condition with complete homozygosity even in the simplest case of one locus with two alleles. It depends on the magnitude of these factors and the degree of heterozygosity of the variety how close to genetic uniformity we ultimately can come. The advancing of one generation has only a small effect, which becomes less as the frequency of one of the genotypes becomes less. Mass selection can reduce segregation in a variety, but only in ideal situations and only in small steps. Obviously the most effective way to promote genetic uniformity is to begin with non-segregating material obtained through careful inbreeding and within-family selection. Then, if this is available, mass selection is hardly necessary and seed collection should be done so as to prevent a return to a heterozygous condition. When only segregating populations are available, some changes can be expected if the selection intensity is low, but they will not be great and may not be noticed until after several generations. Returning to the original question of the stability of the tobacco variety, the conclusion can be drawn that when large numbers of plants from a field are selected as seedplants the changes in the next generation as a whole will be small, regardless of the selection procedure used. When 10-30 % of all plants are allowed to produce seed we cannot expect important changes. Also, when the environmental variations are as great as in shade tobacco, the possibilities of selecting against certain genotypes for a number of characteristics simultaneously becomes virtually impossible. Concerning whether or not mass selection can lead to improvements in the variety, it was explained that the selection intensity and the heritability of a character determine the selection response. Perhaps this response can be predicted in the case of one character, but it becomes difficult, if not impossible, to forecast the results when many characters are involved at the same time. Reference was also made to a selection index which has been effective in livestock breeding. Even if an index could be determined for shade tobacco, its use is not necessarily effective, as Kempthorne (1957) pointed out. The genotype-environment interaction makes questionable the choice of certain fields over others. Without experiments, such questions cannot be answered. It is likely that mass selection with low selection pressure changes a variety very little. Again, in a field where up to 30 % of the plants are selected for seed, the pressure cannot be very great and we should not expect great changes. Only when a few plants are carefully selected for certain characters should progress become noticeable, as is the experience of single plant selection in plant breeding. We conclude that mass selection for seed and plant breeding to improve a variety should not be confused with each other. Each has its own aims and methods, which are not interchangeable. Where uncertainty exists about seed, progeny tests usually are made to compare a seed with the parent seed. Bolsunov (1959) has described a number of such procedures for tobacco, though the more elaborate of them appear impractical.

Description: What KIAERSK wrote in 1893 in the preface of his “Enumeratio Myrtacearum Brasiliensium” is still largely valid. It is often most difficult to define a species belonging to this family, not only because, in the absence of ripe seeds, the genus is not easily ascertainable, but also because of the strong variability shown by the vegetative characters. Thanks to the examination of the rich Guiana material preserved in the herbaria of Genève, Kew, Leiden, New York, Paris and Utrecht, I have usually been able to delimit the species in a satisfactory way; their allocation to a definite genus, however, is often a difficult problem. During the preliminary stage of this investigation, which was interrupted by the war, it was of great advantage to me that I could study the Guiana specimens of the Leiden herbarium. In order to avoid misinterpretations, I have tried to base my conclusions as far as possible on an examination of either the types themselves or of duplicates of the latter. Several of these types, especially those that form part of the earlier collections of Guiana plants, e.g. of the collection Aublet, and of the collections Desfontaines (herb. Florence) and De Candolle (Genève) had never before been reexamined, and BERG, the last monographer of the South American Myrtaceae (in Linnaea XXVII (1855—56), XXIX (1858) and XXX (1861) has either neglected these species or given an, often incorrect, interpretation based on the description alone. For this reason the second part of this paper will be devoted to a short survey of these earlier types. My best thanks are due to the directors of all herbaria mentioned. Moreover, I have to thank the “Van Eedenfonds”, whose financial aid enabled me to pay a visit to Kew and to the British Museum.

Description: Caudalejeunea grolleana Gradst. spec. nov. from Madagascar is referred to this genus with some doubt because of the absence of gemmiparous branches. Ptychocoleus cristilobus (Steph.) Steph. from S. E. Asia has gemmiparous branches and therefore is a true Caudalejeunea: C. cristiloba (Steph.) comb. nov. This species is remarkable by its complicated ciliate leaflobule and by its polystratose rhizoid-disc. Two subspecies are distinguished: ssp. cristiloba from Burma, Andaman Is., Thailand, and Singapore, and ssp. samoana (Steph.) comb. nov. (Caudalejeunea samoana Steph.) from Samoa. Both C. grolleana and C. cristiloba have a 4-5-carinate perianth, which shows that the trigonous perianth present in most species of Caudalejeunea is not a stable character of this genus.

Description: Eén van de grote giften van A. A. Pulle, Lanjouw’s voorganger als Hoogleraar in de Bijzondere Plantkunde en de Plantengeographie te Utrecht, was zijn goede keus van medewerkers. Hij kon die keus maken dank zij een andere gift, die van leermeester. Pulle’s stimulerende activiteit als hoogleraar luidde de wedergeboorte in van de Nederlandse plantensystematiek door een stroom van leerlingen waarvan er velen ook nu nog een belangrijke rol spelen in de botanische wereld. Vele van Pulle’s leerlingen uit de eerste tijd moesten Utrecht verlaten omdat er in de twintiger jaren nu eenmaal nog geen sprake was van een redelijk gesalariëerde wetenschappelijke staf. Eén leerling echter werd zo gefascineerd door de vele mogelijkheden die het Utrechtse instituut bood, dat hij bleef, niettegenstaande de soms zeer ongunstige economische en organisatorische omstandigheden. Deze leerling, Lanjouw, toonde reeds toen de taaie volhardendheid die zijn medewerkers en leerlingen in later jaren zo goed leerden kennen en waarderen. Reeds van de eerste jaren van zijn assistentschap aan stonden Lanjouw de idealen voor ogen die hij in de loop van zijn lange loopbaan van assistent tot directeur voor het grootste gedeelte zou weten te realiseren. Deze loopbaan begon toen hij op 1 januari 1926 benoemd werd tot assistent bij de Bijzondere Plantkunde. In het jaar waarin we het feit herdenken dat het veertig jaar geleden is dat Lanjouw verbonden werd aan het Botanisch Museum valt ook de honderdenvijftigste verjaardag van het herbarium. Plantensystematiek is te Utrecht in meerdere of mindere mate beoefend van de stichting van de Universiteit af. Op 30 oktober 1816 echter werd het eerste herbarium voor de Universiteit verworven. In 1966 herdenken we dus eigenlijk twee jubilea. De periode van werkelijk actief systematisch onderzoek is echter veel korter geweest: de korte jaren van Miquel’s werkzaamheid (1859-1871) en de periode die in 1906 begon met de aanstelling van Pulle tot Lector in de plantensystematiek. Pulle gaf, voor het eerst na Miquel, en mede dank zij de stimulerende invloed van Went, een nieuwe stoot aan het plantensystematisch onderzoek in Nederland en wel vooral aan het onderzoek van de tropische flora’s. Een reeks van leerlingen wijdde zich onder zijn leiding aan de studie van de flora’s van Suriname en Nederlands Indië. Het is voldoende hierbij de namen van van Sloten, Lam, van Steenis, en Uittien te noemen om deze ontwikkeling te illustreren. Lanjouw koos voor zijn proefschrift de Surinaamse Euphorbiaceae en het was uit dit werk dat de „Flora of Suriname” voortkwam. Zonder ook maar iets tekort te doen aan de rol van Pulle kan gezegd worden dat van het begin van zijn loopbaan af Lanjouw actief heeft medegewerkt aan de opbouw van het Botanisch Museum en dat hij bij de ontwikkeling van de zo belangrijke „Flora of Suriname” een doorslaggevende rol heeft gespeeld. Zijn dissertatie „The Euphorbiaceae of Suriname” vormde, ten dele, tegelijk de eerste aflevering van de Flora.