ALBERT

Description: Trees and their environment were studied in floodplain forests in the glaciated region of northern Missouri. Ordination of tree vegetation samples by detrended correspondence analysis indicated a primary vegetation continuum of decreasing Acersaccharinum L. and increasing Caryalaciniosa (Michx. f.) Loud with several other species associated with secondary vegetation gradients, TWINSPAN classification of tree vegetation identified three groups of plots that were dominated by A. saccharinum in varying degrees of association with other species, most importantly Populusdeltoides Bartr. and Ulmusamericana L.; two groups dominated by species of Carya and Ulmus; and two groups where dominance was more broadly distributed among lowland Quercus spp., U. americana, Aesculusglabra Willd., Fraxinuspennsylvanica Marsh., Platanusoccidentalis L., and Betulanigra L. Importance of A. saccharinum was greatest in plots where the leading dominants were young, while Quercus and Carya spp. were more common in plots with older leading dominants, suggesting that the predominant environmental influence on vegetation composition was frequency and severity of disturbance associated with flooding. Plots with younger dominant trees had lower species richness and diversity than plots with older dominants. Higher soil pH and slough location were also positively correlated with A. saccharinum importance, and Acernegundo L. was more frequent in sloughs. Analysis of overstory and understory relationships indicated that A. saccharinum is likely to remain important in the immediate future in many forests currently dominated by this species because of its abundance in subcanopy positions. Potential canopy trees of F. pennsylvanica, C. laciniosa, Caryacordiformis (Wangenh.) K. Koch, and Celtisoccidentalis L. may, in the absence of major flooding disturbance, cause long-term shifts in composition in some of these forests. While Ulmus spp. are abundant in the understory, they are unlikely to become important canopy species because of disease.

Description: The effects of climate on the growth of sugar maple (Acersaccharum Marsh.) were studied at five sites along an 800-km acidic deposition gradient from Cook County, Minnesota, to Oceana County, Michigan. Fifty increment cores were taken from 25 dominant and codominant individuals at each site in the spring of 1990. Annual ring widths (1940–1989) were measured to 0.01 mm, standardized by taking the first logarithmic differences, and averaged into chronologies using the biweight mean. The five resulting chronologies were then related to climate using least squares regression techniques. The analyses indicate that temperature is associated with sugar maple growth to a greater degree than precipitation, though there were differences in the relationships among the five study sites. Growth was also found to be significantly affected by prior growing season conditions. No evidence of an overall decline or increase in sugar maple growth rates was observed over the 50-year study period.

Description: Calamagrostiscanadensis (Michx.) Beauv. is a widely distributed rhizomatous grass that can seriously inhibit growth of white spruce (Piceaglauca (Moench) Voss) seedlings in the boreal forests of North America. We review the dynamics of this grass during four successional stages: the colonization of disturbed sites; dominance of the site by the grass a few years after disturbance; gradual loss of dominance with overstory development; and maintenance of the grass at low levels in the understory of the mature forest. We also describe C. canadensis in relation to recruitment from clonal growth and seed, environmental conditions for growth, the effects of grass litter buildup on conifer seedling microclimate, and overall competitive abilities. Control strategies for C. canadensis are as follows. If the grass is found in nearly every square metre in the understory prior to logging, there will be rapid spread when the stand is clear-cut unless clones are killed using herbicides or a deep burn. Large spruce seedlings, planted on large soil scalps or mounds, coupled with release by way of herbicides or sheep grazing, may be necessary for plantation establishment under conditions of encroachment by C. canadensis. Alternatively, the shade provided by a partial canopy may inhibit the grass sufficiently to allow spruce seedlings to establish. If grass is not abundant in the understory, we recommend (i) minimizing forest floor disturbance to reduce sites for grass seedling colonization or (ii) a slash burn with the hope of encouraging colonization by herbaceous species that have less impact on conifer seedlings.

Description: Bulk density of forest soils from nine locations in New England was closely and inversely related to the organic fraction of the soil. Measured data over the whole range of organic fractions followed the theoretical relationship Db = DbmDbo/[FoDbm + (1 − Fo)Dbo] where Db is the bulk density (Mg/m3), Fo is the organic fraction (kgo/kg), Dbo is the bulk density when Fo = 1, and Dbm is the bulk density when Fo = 0. The relation arises from assuming that (i) Dbo, the bulk density of "pure" organic matter, and Dbm, the bulk density of "pure" mineral matter, are constant and (ii) in a mixture, the volumes occupied by the organic mass and the mineral mass are additive. For forest soils on coarse-textured till in New England, Dbo = 0.11 Mg/m3 and Dbm varied from 1.45 Mg/m3 for sandy loams to 2.19 Mg/m3 for silt loams. When these parameters are known, Db can be estimated from Fo, which is more easily measured. When Fo is greater than 0.1 kgo/kg, the organic mass per unit soil volume (FoDb), or organic density, is approximately constant at 0.1 Mgo/m3. For many nutrients, separate evaluation of the organic density and the amount of nutrient per unit organic mass may facilitate intersite comparisons for studies of nutrient availability and leaching.

Description: Boles of quaking aspen (Populustremuloides Michx.), white spruce (Piceaglauca (Moench) Voss), red pine (Pinusresinosa Ait.), and jack pine (Pinusbanksiana Lamb.) were sampled after decomposing for 11–17 years. Mass loss and changes in chemical composition were determined. Density decreased by 40–73%, and the decomposition constant (k) decreased in the order aspen 〉 spruce 〉 red pine 〉 jack pine. Although the decomposition rate was strongly species dependent, it did not differ between the two sites. The decomposition constant was weakly related to lignin and phosphorus concentration in the original stems (r2 = 0.44 and 0.49, respectively) and not significantly related to original nitrogen concentrations. Nutrient concentration increased during decomposition, particularly for nitrogen and phosphorus. Nitrogen content increased during decomposition, and potassium content decreased. Calcium and magnesium content changed little during decomposition, whereas phosphorus content increased or decreased, depending on species. In general, species with the lowest initial nutrient contents had the greatest nutrient increases during decomposition. Nutrient content of boles of all species became similar during decomposition.

Description: Tree-ring analysis was used to study historical patterns of basal area increment (BAI) by healthy (0–5% dieback) and declined (greater than 30% dieback) overstory sugar maples (Acersaccharum Marsh.) in four stands in Pennsylvania. The objectives were to establish if and when BAI decreased in healthy and declined trees and to identify causal factors associated with decreased BAI. Reduction in BAI of declined compared with healthy trees was first evident following a series of defoliations and summer droughts in the mid-1960s and early 1970s. These stresses were followed by repeated damage by pear thrips (Taeniothripsinconsequens Uzel) in the 1980s and a drought in 1988, during which large reductions in BAI occurred for both declined and healthy trees in all stands. Foliar nutrient analysis of two stands suggested deficiency of Mg and Ca. Other factors hypothesized to be associated with this sugar maple decline as either predisposing or inciting factors include unusually warm winter temperatures in the 1980s and air pollution.

Description: On the rocky shores and islands of Lake Duparquet, in the southwestern Quebec boreal forest, Thujaoccidentalis L. reaches ages in excess of 800 years. Annual ring widths from 38 trees were used to develop an 802-year chronology (1186–1987) standardized by polynomial regressions. Excellent cross dating, correlation with a shorter chronology located 14 km inland, and 33.6% common variance in a chronology subsample all point to the existence of a climatic signal. After autoregressive modeling to obtain a serially random residual chronology, correlation and response functions were used to identify the growth–climate relationship. The resulting model reduced 19.2% of the chronology variance. Precipitation in June as well as low temperature in June or July seemed to have a positive influence on growth. Likewise, a drought index was closely related to growth, indicating that the chronology could be used to estimate past drought conditions. Moisture deficits are thus inferred for the 13th century as well as during the Little Ice Age (17th century to late 19th century). Since the end of the latter period, precipitation seems to have followed an upward trend.

Description: Three southern Appalachian stands with sparse and unproductive pine–hardwood overstories and dense Kalmialatifolia L. understories were treated to restore productivity and diversity on steep slopes. An adaptation of the fell and burn practice was applied in summer and fall 1990. About one-half of the woody fuels were consumed at each site. A range of fire intensities was observed. Flame temperatures approached 800 °C, but the heat pulse into the forest floor only reached 60 °C at 5 cm. Humus and charred leaf litter remained on most of the surface after burning. Evidence of soil erosion was spotty and related to points of local soil disturbance. No soil left the sites. At the end of the first growing season, 23% of the burned surfaces were covered by growing plants and 62% by residual forest floor and woody debris. Felling and burning reduced evapotranspiration so that soil in the treated areas remained moister than under adjacent uncut stands. Opening the sites increased soil temperatures 2 to 5 °C at 10 cm during the first 16 months after treatment.

Description: Three paired watersheds treated with a fell and burn prescription were studied to determine the effects on soil, soil water, and stream water. Soil nitrification and mineralization were measured by in situ closed-core incubation. Soil water was collected with porous cup lysimeters placed at 30 and 60 cm depths, and water samples were collected from streams draining control and burned areas on one of the three sites. All data were collected for 6 months prior to and 12 months after treatment. Soil ammonium (NH4+) content increased significantly in all three sites after burning, but the magnitude differed greatly among sites. However, there was no change in soil nitrate (NO3−) content. In situ measurements of net mineralization showed increased rates with increasing burn severity. Net nitrification displayed no treatment response. Slight and nonsignificant increases in soil water NO3− concentration occurred after burning in two of the three sites. Stream water NO3− concentrations increased in the one stream sampled. Thus, while prescribed burning increased available soil N, there was little change in N transformation rates or movement of dissolved inorganic N off site during the first year after burning.

Description: Change in the health of sugar maple (Acersaccharum Marsh.) and associated northern hardwoods was evaluated for 3 years (1988–1990) in seven states and four provinces. Generally, levels of crown dieback and crown transparency (a measure of foliage density) in 165 stands decreased during this period. In 1990, less than 7% of all dominant–codominant sugar maples (n = 7317) exhibited crown dieback ≥ 20%. Significantly (p = 0.05) fewer of these maples were classified as having high crown transparency (≥ 30%) in 1990 compared with 1988. Crowns of maples that received moderate (31–60%) or heavy (〉 60%) pear thrips (Taeniothripsinconsequens (Uzel)) damage for 1 year recovered the following year. Crowns of maples exposed to severe drought in 1988 (Wisconsin) continued to show the effects (high transparency) of this stress in 1990. A majority (69–71%) of the dominant–codominant sugar maples with high (≥ 20%) crown dieback had bole and (or) root damage. Of those maples with crown dieback ≥ 50%, 86% had bole and (or) root damage. The condition of sugar maple in operating sugar bushes and undisturbed stands was similar. The condition of sugar maple crowns was similar in locations presumably exposed to low, medium, and high levels of sulfate deposition.

Description: A series of experiments on field-grown seed-derived trees between 2 and 17 years old demonstrated that the growth regulator paclobutrazol could be used to reduce vegetative growth and enhance flower-bud production in Eucalyptusglobulus Labill. and Eucalyptusnitens (Dean & Maid.) ex Maid. Responses to high levels of trunk injection and collar drenching persisted for up to six growing seasons, yielding both increases in frequency of flowering and heaviness of bud crop. Growth responses were expressed in the immediate growing season, but flowering responses were not evident for another year. Foliar spray treatments reduced vegetative growth in young trees of both species for one growing season, but only the E. globulus showed an associated flowering response. Assessment of seed yield per capsule and subsequent germination tests showed no deleterious effects on seed development or quality. Choice of application method will vary with objective and size of tree. Collar drenching shows the most promise for treating large numbers of seed orchard trees because application time is substantially independent of tree size and weather conditions.

Description: This paper describes a decision support system that forest managers can use to help evaluate short-term, site-specific silvicultural operating plans in terms of their potential impact on long-term, forest-level strategic objectives. The system is based upon strategic and tactical forest-level silvicultural planning models that are linked with each other and with a geographical information system. Managers can first use the strategic mathematical programming model to develop broad silvicultural strategies based on aggregate timber strata. These strategies help them to subjectively delineate specific candidate sites that might be treated during the first 10 years of a much longer planning horizon using a geographical information system and to describe potential silvicultural prescriptions for each candidate site. The tactical model identifies an annual silvicultural schedule for these candidate sites in the first 10 years, and a harvesting and regeneration schedule by 10-year periods for aggregate timber strata for the remainder of the planning horizon, that will maximize the sustainable yield of one or more timber species in the whole forest, given the candidate sites and treatments specified by the managers. The system is demonstrated on a 90 000 - ha area in northeastern Ontario.

Description: A simple conceptual model is proposed concerning how leaf area efficiency (stemwood growth per unit leaf area) changes with leaf area for trees within a stand. Greater leaf area is generally associated with (i) improved light environment due to greater height and (ii) a lower ratio of photosynthetic to nonphotosynthetic tissue. Greater height and improved light environment result in higher photosynthetic production, which should increase leaf area efficiency. A lower ratio of photosynthetic to nonphotosynthetic tissue suggests that the ratio of respiration to photosynthesis increases, which should decrease leaf area efficiency. In relatively small trees, the influence of increased height (associated with greater leaf area) should more than offset the influence of the increased respiration:photosynthesis ratio; as a result, leaf area efficiency should increase with leaf area. In large trees, further increases in leaf area are associated with minimal increases in height, and leaf area efficiency should decline as the respiration:photosynthesis ratio increases. Predictions from this conceptual model were examined with data from stands of subalpine fir (Abieslasiocarpa (Hook.) Nutt.).

Description: We studied radial growth reduction in American beech (Fagusgrandifolia Ehrh.) in relation to the level of defect induced by beech bark disease, in second-growth and old-growth northern hardwoods stands in New Hampshire. In the second-growth stand at Moose Mountain (n = 243 trees), 1989–1990 radial growth declined significantly with increasing severity of external symptoms. The severity of external symptoms increased significantly with DBH. To examine temporal trends, internal defect induced by the disease was quantified as the percentage of growth sheath cankered in each year, by cross-sectioning a subsample of 40 trees. Internal defect first appeared in 1950, increased through 1969, then declined until a major pulse of infection in the period 1983–1987. Sectioned trees were divided into infection classes based on a cumulative measure of internal defect. Growth of severely infected trees first fell below that of uninfected trees in 1965, and was consistently lower after 1972. The growth ratio of severely infected to uninfected trees generally declined from 1960 to 1990; by 1990, growth of severely infected trees was reduced by more than 40% relative to healthy trees. This decline in the growth ratio corresponded well to the increase in cumulative internal defect in the stand, suggesting that disease stress had cumulative effects on tree vigor. The relation between beech bark disease and growth was also examined on an individual-tree basis; recent growth decline was significantly greater for trees with higher levels of internal defect. Internal defect was a better predictor of growth trends than was external defect. External defect was only moderately correlated with internal defect (r2 = 0.503). In the old-growth stand at Bartlett, N.H. (n = 40 trees) infection was quantified from external symptoms only. As in the second-growth stand, the growth of severely infected trees in the old-growth stand fell significantly below that of uninfected trees. However, significant differences in growth between uninfected and severely infected trees occurred earlier in the old-growth stand, first appearing in 1949. Delayed growth reductions in the second-growth stand may be associated with changes in shade and moisture affecting the beech scale, changes in tree physiological stress after selective logging, or changes in the density of large trees. Beech may survive long periods of infection by beech bark disease. However, our results demonstrate clearly that beech bark disease has reduced the growth of American beech in both second-growth and old-growth northern hardwoods stands for several decades.

Description: Stratum-based timber harvest schedules must be disaggregated into operational plans prior to implementation. In most cases this is an expensive and time-consuming manual task that does not ensure consistency between the long-term harvest schedule and short-term operational activities. This paper presents the results of applying the CRYSTAL algorithm, which automates the disaggregation and allocation of a stratum-based harvest schedule into harvest blocks, to a small forest in New Brunswick. The results indicate that it is possible to use a set of allocation guidelines to quickly delineate harvest blocks in a consistent, reproducible manner. We also discuss how the algorithm is used in conjunction with a Monte Carlo integer programming model to estimate the potential losses in timber harvest volumes attributable to deviations from the stratum-based schedule and the addition of adjacency constraints.

Description: Identifying the factors that influence foresters to adopt management innovations is essential for developing models that accurately forecast the rate at which new technology will be adopted. Models were developed to evaluate the impact of sociodemographic characteristics, attitudes, and structural–external variables on New Hampshire foresters' decisions to adopt three distinct soil information sources. The soil information sources studied were County Soil Surveys, Important Forest Soils Groups, and Leak's Habitat Classification System. Discriminant analytic models were developed and evaluated for each information source. Information on use and other variables were collected by a mail survey with a response rate of approximately 60%. The results reveal that structural variables exerted the greatest influence on adoption decisions, while sociodemographic characteristics were the least important. College and noncollege sources of information about the innovations were the most important variables in the discriminant models. Other significant structural variables included employer, prior use of soils information, and participation in continuing education. All three models correctly classified more than 75% of the respondents, with one model classifying more than 90%. The difference in accuracy may be attributed to differences in information access among the three soils information sources.

Description: The reactions of sessile oak (Quercuspetraea (Mattuschka) Liebl.) to drought were studied under natural conditions in a 32-year-old stand near Nancy (northeastern France) during the summers of 1989 (strongly rain deficient) and 1990. A plot of five trees was subjected to imposed water shortage, while a group of irrigated trees was used as a control. Measurements of xylem sap flows and water potential enabled the computation of plot transpiration, canopy conductance, and specific hydraulic conductance in the soil–tree continuum. Stomatal conductance was measured directly with a porometer. Specific hydraulic conductance of our oaks was of the same order of magnitude as that reported for other species. It decreased significantly during spring because of a time lag between cambial growth and leaf area expansion. Measured transpiration was close to potential evapotranspiration, except during days with high vapor pressure deficits, which promoted stomatal closure in the absence of soil water deficits. Imposed drought caused predawn leaf water potentials to reach values as low as −2.0 MPa and a progressive decline in hydraulic conductance, which was probably attributable to modifications in hydraulic properties at the soil–root interface. This gradual decline in conductance was attributed to their deep rooting (1.40 m). This study revealed that Q. petraea may be considered as drought tolerant because of adaptations like deep rooting, efficient and safe xylem sap transport, maintenance of significant stomatal conductance, and significant transpiration, even during strong drought stress.

Description: This review presents information on the occurrence, persistence and nontarget effects of Bacillusthuringiensis Berliner (B.t.) in soil, with particular emphasis on forest ecosystems. Both field and laboratory studies have confirmed that B.t. is able to survive for several years after spraying, but studies on long-term persistence and possible accumulation of spores in soils are inadequate. The ecological role of B.t. in the soil ecosystem is poorly understood; we do not even know where and under what conditions B.t. multiplies in nature. Information on the effects of B.t. on soil microflora is contradictory, with studies variously suggesting that B.t. caused an increase, a decrease, or did not affect indigenous bacterial populations. Toxicity of B.t. to species of several invertebrate taxa (Acarina, Nematoda, Collembola, Annelida, Hymenoptera) inhabiting the soil has been demonstrated, but only rarely is it possible to relate dosage information to field situations, and in many cases the B.t. subspecies tested are not currently used for pest control in North America. There is an urgent need for further research to elucidate the relationships between B.t. and the natural soil microflora and fauna.

Description: Development of humus layer soil microbial biomass C (Cmic) and N (Nmic), fungal biomass (as soil ergosterol content), microbial respiration activity, and the soil organic C (Corg) and N (Ntot) were determined in coniferous forest soils that had received a single prescribed fire treatment at different times over a period of 45 years. The ratio of soil respiration rate to microbial biomass C (qCO2) and the Cmic/Corg and Nmic/Ntot percentages were derived from the measurements taken. All the measured biomass indicators reacted identically to show recovery from prescribed burning within 12 years. A raised metabolic quotient (qCO2) was detected in soils over the first 2 years following the fire treatment, but after the third year it had decreased to a stable level. These observations suggest that during the first few years after fire the soil microflora can be characterized on the basis of simple substrate–decomposer relationships. The first 12 years were characterized by increasing Cmic/Corg and Nmic/Ntot percentages, which then stabilized at mean values of 1.3 and 5.5%, respectively. The observed rise in the Cmic within a large pool of Corg suggested increasing availability of energy-rich C sources. These C sources are probably derived from the organic C input resulting from postfire plant succession.

Description: During a 3-year study, soil microbial biomass C and N, length of the fungal hyphae, soil respiration, and the percent mass loss of needle litter were recorded in coniferous forest soil humus layers following a prescribed burning (PB) treatment or a forest fire simulation (FF) treatment (five plots per treatment). Unburned humus from adjacent plots served as controls (PC and FC, respectively). Prescribed burning was more intensive than the forest fire, and this was reflected in all the measurements taken. The amounts of microbial biomass C and N, length of fungal hyphae, and soil respiration in the PB area did not recover to their controls levels, whereas unchanged microbial biomass N and recovery of the length of the fungal hyphae to control levels were observed in the FF area. The mean microbial C/N ratio was approximately 7 in all the areas, which reflected the C/N ratio of the soil microbial community. Deviation from this mean value, as observed during the first three samplings from the PB area (3, 18, and 35 days after fire treatment), suggested a change in the composition of the microbial community. Of the two treated areas, the decrease in soil respiration (laboratory measurements) was much more pronounced in the PB area. However, when the humus samples from both areas were adjusted to 60% water holding capacity, no differences in respiration capacity were observed. The drier humus, due to higher soil temperatures, of the PB area is a likely explanation for the low soil respiration. Lower soil respiration was not reflected in lower litter decomposition rates of the PB area, since there was a significantly higher needle litter mass loss during the first year in the PB area followed by a decline to the control level during the second year. Consistently higher mass losses were recorded in the FC area than in the FF area.

Description: Red spruce (Picearubens Sarg.), black spruce (Piceamariana (Mill.) B.S.P.), and balsam fir (Abiesbalsamea (L.) Mill.) seedlings were root inoculated with tomato mosaic virus, potted, and maintained in a cold frame. The virus was detected by enzyme-linked immunosorbent assay in root extracts of several inoculated seedlings of each species 6–12 months postinoculation, and in root extracts of approximately 35–40% of both inoculated and noninoculated seedlings of each species 12–18 months postinoculation. Virus spread apparently occurred from the roots of infected to noninfected seedlings within the cold frame. The presence of virus in root extracts was confirmed by immunoelectron microscopy. The virus was not detected in the needles of any seedling at any time by either enzyme-linked immunosorbent assay or immunoelectron microscopy. A transitory needle chlorosis was observed in approximately 14% of the inoculated black spruce and 30% of the inoculated red spruce seedlings, but tomato mosaic virus was not detected in all plants with symptoms.

Description: Rising atmospheric carbon dioxide concentrations may have important consequences for forest ecosystems. We studied above- and below-ground growth and leaf gas exchange responses of Populusgrandidentata Michx. to elevated CO2 under natural forest conditions over the course of a growing season. Recently emerged P. grandidentata seedlings were grown in native, nutrient-poor soils at ambient and twice ambient (707 μbar (1 bar = 100 kPa)) CO2 partial pressure for 70 days in open-top chambers in northern lower Michigan. Total leaf area and shoot and root dry weight all increased in high CO2 grown plants. Photosynthetic light and CO2 response characteristics were measured 28, 45, and 68 days after exposure to elevated CO2. In ambient grown plants, light saturated assimilation rates increased from day 28 to day 45 and then declined at day 68 (15 September). This late-season decline, typical of senescing Populus leaves, was due both to a decrease in the initial slope of the net CO2 assimilation versus intercellular CO2 partial pressure relationship and to decreased CO2 saturated assimilation rates. Specific leaf nitrogen (mg N•(cm2 leaf area)−1) did not change during this period, although leaf carbon content and leaf weight (mg•cm−2) both increased. In ambient grown plants stomatal conductance also declined at day 68. In contrast, plants grown at elevated CO2 showed no late-season decline in photosynthetic capacity or changes in leaf weight, suggesting a delay in senescence with long-term exposure to high CO2. High CO2 grown plants also maintained photosynthetic sensitivity to increasing Ci throughout the exposure period, while ambient CO2 grown plants were insensitive to Ci above 400 μbar on day 68. These results indicate the potential for direct CO2 fertilization of P. grandidentata in the field and provide evidence for a new mechanism by which elevated atmospheric CO2 could influence seasonal carbon gain.

Description: Nitrogen (N) mineralization and nitrification were compared among ecosystems representing a primary successional sequence on the Tanana River floodplain of interior Alaska. These processes displayed marked seasonality, were closely related to substrate chemistry, and reflected the impact of vegetation clearing. The highest rates of N mineralization were encountered in the June to July incubation periods, and rates generally declined during the remainder of the summer. The early season period (June to July) was the interval of most favorable litter and mineral soil temperature and most available energy supply for microbial mineralization of detrital materials. Minimal rates were encountered during the winter. Litter layer N mineralization rates were highest in the early-successional poplar–alder (Populusbalsamifera–Alnustenuifolia (Nutt.) stage and declined with advancing succession in poplar (Populusbalsamifera) and mature white spruce (Piceaglauca) (Moench) Voss) stands. The poplar–alder stage displayed the highest rate of nitrification. Nitrate constituted 98% of the mineralized N in early-successional poplar–alder forest floors but fell to 4 and 0% in poplar and white spruce forest floors, respectively. Nitrogen mineralization was closely related to significant increases in the lignin/N ratio across the sequence of vegetation types. The rate of surface mineral soil net N mineralization increased with succession in response to higher soil organic matter content. The range of average total seasonal net N mineralization (260–1600 mg N•m−2) for litter layer plus mineral soil among successional stages in this study was generally lower than the 1200–8400 mg N•m−2 reported by investigators for other studies in temperate latitudes. Vegetation clearing increased the magnitude of temporal fluxes as well as total annual mineral N production. The most consistent increases were encountered in the poplar–alder vegetation type. The average seasonal total net N mineralization for forest floor plus mineral soil in this vegetation type increased from 1500 to 3264 mg N•m−2 as a result of clearing. Soil temperature declined with advancing succession and generally increased as a consequence of clearing. However, these changes were not as closely correlated with N mineralization as were the changes in substrate chemistry encountered across this successional sequence.

Description: Variation in diameter, height, and stem volume of 57-year-old Douglas-fir (Pseudotsugamenziesii var. menziesii (Mirb.) Franco) was related to distance of these trees from a 27 m wide strip in the same Douglas-fir plantation that had been interplanted with red alder (Alnusrubra Bong.). Within the interplanted strip and despite its greater total stand density, bole volume of dominant and codominant Douglas-fir averaged 1.27 m3 compared with 0.55, 0.45, 0.46, or 0.49 m3 in trees 15, 30,45, or 60 m, respectively, from the edge of the mixed stand. Some positive influence of nitrogen-fixing red alder apparently extended about 15 m beyond the edge of the mixed stand at this poor quality site in southwest Washington. We infer that similar ribbonlike distributions of naturally regenerated red alder could be retained to improve growth of nearby conifers on nitrogen-deficient sites.

Description: Attraction of scolytids and other bark- and wood-dwelling beetles to volatile constituents of Norway spruce (Piceaabies (L.) Karst.) was studied in field experiments in central Sweden. The volatiles were released from chips of newly cut spruce stems as well as from stems stored over winter. These host materials were kept in cylinders covered on both ends with fine nylon mesh. Attracted insects were caught in barrier traps. The composition of volatile constituents of wood samples was determined using gas chromatography. Hylurgopspalliatus (Gyll.), Hylastescunicularius Er., Ipstypographus (L.) (Scolytidae), and Glischrochilusquadripunctatus (L.) (Nitidulidae) were particularly attracted to stored spruce wood. Trypodendronlineatum (Oliv.) (Scolytidae) and Pityophagusferrugineus (L.) (Nitidulidae) were only attracted to stored wood. There were indications that Tomicuspiniperda (L.) and Hylastesbrunneus Er. (Scolytidae) preferred fresh wood. Chemical analyses revealed that during storage ethanol and acetaldehyde increased considerably in two out of the five examined tree stems.

Description: One-year-old Cornusflorida L. seedlings were randomly assigned to four treatments of simulated rain (pH 2.5, 3.5, 4.5, and 5.5) in two greenhouse–laboratory studies in 1989 and 1990. After 10 applications over a 42-day period, the seedlings were moved to a temperature-controlled laboratory, placed in plastic bags, humidified, and sprayed with a spore suspension of five Disculadestructiva isolates. About 30 days later, the seedlings were examined for percentage of leaves exhibiting anthracnose symptoms and disease severity on affected leaves. Both trials showed that as the acidity of the simulated rain increased, the incidence and severity of anthracnose leaf symptoms increased. The 1989 study included a soil lime treatment that showed the same trend but the overall occurrence and severity of symptoms was higher.

Description: The prevalence of individual-tree growth decline was determined for red spruce (Picearubens Sarg.) populations at three locations in the southern Appalachians: Mount Rogers National Recreation Area, the Black Mountains, and Great Smoky Mountain National Park. An index of annual stemwood volume increment (AVI) was computed from dendrochronological data and a site-specific DBH–height regression equation. Individual-tree AVI time series were analyzed to identify changes in 20-year periodic mean AVI and AVI trend. The proportion of red spruce that exhibited decreasing mean AVI or negative AVI trend was determined for the most recent 20-year period, and this was compared with the estimated historical prevalence of these indications of growth decline. Also, the prevalence of growth decline was compared among subpopulations that differed with regard to various tree, stand, and site characteristics. Of 263 red spruce sampled, 25% exhibited a decrease in mean AVI during the period 1967–1986, 8% exhibited a negative AVI trend without a reduction in mean AVI, and 17% exhibited a reduction in the slope of the AVI curve. The proportion of trees that exhibited decreasing or slowed growth after 1967 was substantially greater among trees growing at 1980 m than in populations at lower elevations; no relationship was found between elevation and growth decline below 1980 m. No difference was found in prevalence of growth decline between subpopulations that differed with regard to age, DBH, competitive status, stand density, slope aspect, or site exposure. The prevalence of individual-tree growth decline for the most recent 20-year period did not exceed estimated levels for historical periods of decline in the Great Smoky Mountains population.

Description: Phenotypic variation in defoliation of ponderosa pine, Pinusponderosa Laws., by the pine sawfly, Neodiprionfulviceps, is reported. Two field and one laboratory bioassays of paired phenotypically resistant and susceptible trees consistently indicated that sawfly egg, larval, and total survival were 5–14% lower on resistant trees. A suite of host-plant traits were assayed on resistant and susceptible trees including anatomical characteristics of the needles and concentration of terpene and nitrogen. Resistant trees had thicker outer cell layers, more fiber cells, higher toughness, and lower soluble nitrogen than susceptible trees. Resistant and susceptible trees were not distinguished by differences in terpenes. Generational survival of N. fulviceps was inversely related to foliage toughness and number of fiber cells.

Description: In a routine determination of jack pine (Pinusbanksiana Lamb.) tree ages, several rings were overlooked or were difficult to count. Sections of the wood, when examined at higher magnification, revealed an unusually high proportion of "light rings." Light rings are characterized by having a small number of latewood cells whose walls are not as thick as those of latewood cells found in normal rings. Under low magnification, bands of these light rings may be interpreted as a single annual ring. Light rings may thus be a source of error in determining tree ages for forest productivity studies, particularly in older stands that have been affected by defoliators and root disease.

Description: A base-age invariant site index equation for jack pine based on the Chapman–Richards function was produced that satisfied nine criteria of preferred behavior for site index equations. A difference form of the Chapman–Richards equation produced the best behavior; height equalled site index at base age, and the shape of the curves reflected the data. The data structure used to fit the difference equation was all possible differences rather than the conventional nonoverlapping sequential intervals because this improved the behavior of the model. Height-prediction equations typically use height at base age (site index) as a predictor variable. As site index is measured with error, the equation will be biased. This bias will be evident in the predicted height at base age and in the shape of the curves. Base-age invariant equations predict height and site index with the same equation and thus diminish the effect of stochastic predictor variables. The equation performed comparably to a previously published equation with a specific base age of 50 years.

Description: Height and diameter growth, stem volume production, leaf phenology and leaf number, and number of branches of Populustrichocarpa Torr. & Gray, Populusdeltoides Bartr., and their F1 hybrids (P. trichocarpa × P. deltoides) were studied for 4 years in a research plantation in western Washington, United States. Twelve clones (three of each species and six of the hybrids) grew under a short-rotation silviculture regime in monoclonal plots at spacings of 1 × 1 m (10 000 stems/ha). Clones represented a north-south gradient within the geographic distribution of both the two North American poplar species and the parentage of the hybrid material. The results support earlier work by contributing additional evidence for the superiority of the hybrids. However, the relative hybrid superiority in these monoclonal plots was less pronounced than that found earlier in field trials with single-tree plots because of heightened intraclonal competition. After 4 years, mean estimated stem volume of the hybrids was 1.5 times that of P. trichocarpa and 2.3 times that of P. deltoides. Total tree height of the hybrids was 1.1 times that off. trichocarpa and 1.3 times that off. deltoides. Clonal variation was the dominant theme in height and diameter growth, stem volume productivity, time of bud break and bud set, tree mortality, and number of branches. Populustrichocarpa had the highest number of sylleptic branches, P. deltoides had the lowest, and hybrids were intermediate. Significant clone by replicate interactions were observed in height, diameter, and volume growth. Phenological traits, such as the dates of bud break and bud set, and the length of growing period only partly explained the observed differences in growth between the P. trichocarpa × P. deltoides hybrids and the parental species.

Description: Nitrogen transformations were examined in two pairs of adjacent, 55-year-old forests dominated by conifers (primarily Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco)) and by conifers and nitrogen-fixing red alder (Alnusrubra Bong.). Nitrogen availability was examined in aerobic and anaerobic incubations in the laboratory, and with resin-core and buried-bag incubations in the field. Rates of nitrogen mineralization and immobilization were examined in the field incubations using [15N]ammonium. Net nitrogen mineralization in a year-long series of resin-core incubations of forest floor plus 0–0.15 m depth soil was about 9 kmol•ha−1•year−1 for the two alder–conifer stands, but almost nil in the conifer stand at the low-productivity Wind River site, and 2.1 kmol•ha−1•year−1 in the conifer stand at the high-productivity Cascade Head site. The 15N pool dilution experiment showed that buried-bag incubations demonstrated more differences among stands than did resin cores; resin cores typically gave greater rates than buried bags. Previous estimates of nitrogen budgets were coupled with net mineralization estimates to examine how well the estimated fluxes balanced at an ecosystem scale. This tabulation of the complete nitrogen cycles showed substantial discrepancies, prompting caution in interpretation of some of the differences among the stands.

Description: Nitrification and denitrification were measured in a 120-year-old Piceaglauca (Moench) Voss forest in southwestern Alberta. Nitrifying activity could not be detected using short-term incubations of F–H and upper mineral soil horizons. Long-term incubations for nitrifying potential indicated that nitrifying organisms were present. The results suggest that the population of nitrifying organisms must have been small, their activity was limited, or the microhabitat necessary for their activity was not adequately simulated by the soil slurries in the short-term incubations. Low rates of denitrification were detected but probably did not represent a substantial loss of N from the soil–plant system. Low rates of denitrification may have been due to a lack of substrate (NO3−), reflecting the low rates of nitrification.

Description: The nutritional requirements of Eucalyptusgrandis Hill ex Maiden seedlings were studied in glasshouse pot experiments. Nitrogen (N), phosphorus (P), and potassium (K) demand was assessed by bioassays, in which the uptake of tracers (15N, 32P, or 86Rb) by roots excised from the seedlings grown at different levels of nutrient supply was measured. A pilot study showed that there was an inverse relationship between nutrient supply and influx. The measured influx was affected by the length of time that the roots were stored before the bioassay was applied. In a 33 N, P, and K factorial nutrient experiment, growth was primarily influenced by N. Maximum growth occurred at the highest level of supply of each of the three nutrients. N and K influx in the root bioassay was inversely related to foliar N and K concentration, but P uptake was not related to foliar P concentration. These results indicate that the root bioassays are more sensitive determinants of nutrient limitation than. foliar analysis. Interactions between N and P and between N and K alter the stem:leaf ratio, which may be of importance in optimizing wood production.

Description: Forest floors and mineral soils from early (open willow), middle (poplar–alder), and late (white spruce) floodplain primary successional stages were examined for nitrogen fixation and denitrification. The acetylene-reduction and acetylene-inhibition techniques were used separately and in combination to measure nitrogenase and denitrification activities, both in laboratory and field studies. In situ N2O production was undetectable at all sites and during all sampling periods. Denitrifying activity measured in the field with acetylene amendments was low to undetectable, except after a brief flood in the open willow stand when N2O production ranged from undetectable to 34 ng N•cm−2•h−1 within the newly deposited alluvium–old mineral soil interface. Intact core assays also had low to undetectable denitrification activities; the highest activities (259 ng N•g−1 h−1) were measured in the poplar–alder forest floor in the fall. Laboratory studies showed that potential denitrification enzyme activity (DEA) was also greatest in the poplar–alder forest floor (4332 ng N•g−1•h−1), once again occurring in the fall. In early and midsuccessional stages, the interactive effects of temperature, carbon, and NO3− limited denitrification, yet even with the addition of the limiting amendments, low to undetectable DEA was observed in mineral soils. The later white spruce successional stage also had low to undetectable DEA, increasing only with the addition of the full DEA media and independent of temperature changes. Nonsymbiotic nitrogenase activities were highly variable, ranging from undetectable to 30 ng N•cm−2•h−1. Highest activities were seen in the open willow, newly deposited alluvium–old mineral soil interface immediately after a flood and approximately 1 month after the flood on the newly deposited silt surface. Only the white spruce forest floor had measurable nonsymbiotic nitrogenase activity at all sampling times. Alder root nodule nitrogenase activity showed no significant differences between sampling periods. The estimated annual nitrogen fixation rate of 164 kg N•ha−1 for alder root nodules is a substantial N contribution to the alder stand and to the floodplain ecosystem in general.

Description: The variable-form taper function was tested on a data set of 5074 trees, consisting of jack pine (Pinusbanksiana Lamb.), lodgepole pine {Pinuscontorta Dougl.), white spruce (Piceaglauca (Moench) Voss), and trembling aspen (Populustremuloides Michx.), from Alberta. When compared with the results of tests performed on the same data using the variable-exponent function and the segmented polynomial model, the variable-form model was found to be superior (based on residual mean squares) for estimating both upper stem diameter ratios and total stem volumes. The two pine species could be combined and a single taper function used without significant loss of accuracy. Stem form in two of the volume sampling regions was significantly different from that in other regions of the province, but this may have been because the samples were not representative of the two regions. The variable-form function gave accurate and unbiased estimates of both merchantable height and merchantable volume.

Description: Eighteen natural second-growth stands of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) from a wide range of stand densities and stocking levels were measured for stand leaf area index (LAI) from allometric relationships (direct LAI) and the LI-COR LAI-2000 plant canopy analyzer (indirect LAI). LAI was based on one-half total leaf area (m2) per square metre of ground surface area, normal to any slope. The LAI-2000 underestimated direct LAI by 62%. The underestimate of LAI was due to the confinement of needles within branches (74%) and nonrandom branch distribution (26%). In effect, the branches behaved like big leaves. The LAI-2000 can be used to estimate LAI across a variety of stand density and stocking levels by using a single stand clumping factor of 2.63.

Description: Boreal forests contain large quantities of soil carbon, prompting concern that climatic warming may stimulate decomposition and accentuate increasing atmospheric CO2 concentrations. While soil warming increases decomposition rates, the accompanying increase in nutrient mineralization may promote tree growth in these nutrient-poor soils and thereby compensate for the increased carbon loss during decomposition. We used a model of production and decomposition to test this hypothesis. In black spruce (Piceamariana (Mill.) B.S.P.), white spruce (Piceaglauca (Moench) Voss), and paper birch (Betulapapyrifera Marsh.) forests, decomposition increased with the soil warming caused by a 5 °C increase in air temperature. However, increased nitrogen mineralization promoted tree growth, offsetting the increased carbon loss during decomposition. In the black spruce forest, increased tree production was maintained for the 25 years of simulation. Whether this can be maintained indefinitely is unknown. In the birch forest, tree production decreased to prewarming levels after about 10 years. Our analyses examined only the consequences of belowground feedbacks that affect ecosystem carbon uptake with climatic warming. These analyses highlight the importance of interactions among net primary production, decomposition, and nitrogen mineralization in determining the response of forest ecosystems to climatic change.

Description: The effect of convection column air temperature and live needle moisture content on ignitability of tree branches was verified and quantified by exposing branches of three conifer species to a hot-air convection column, at temperatures between 400 and 640 °C, and measuring time to ignition. The three species were ponderosa pine (Pinusponderosa Laws.), Douglas-fir (Pseudotsugamenziesii var glauca (Beissn.) Franco), and lodgepole pine (Pinuscontorta Dougl.). The experiment was repeated monthly over the course of a year, taking advantage of the natural fluctuation of live needle moisture content. Three multiple regression equations for the prediction of time to ignition with air temperature and needle moisture as the independent variables were developed.

Description: Concepts and procedures are presented for the analysis of progeny trials that incorporate clonal replication as a means to resolve variance arising from nonadditive gene effects. Components of variance from the linear model may be expressed in terms of expected covariances among relatives, and these, in turn, may be used to derive approximations of additive, dominance, and epistatic components of genetic variance. In addition to the usual assumptions applied to conventional progeny trials, the use of this expanded genetic model in the analysis of tests with clonal replicates assumes that the greatest portion of the total epistasis is due to interactions involving groups of more than two or three loci. If this assumption is not satisfied, estimates of additive and dominance variance, including those from trials without clonal replicates, will be contaminated by a large fraction of epistasis, and total epistasis will be underestimated by a corresponding amount. Heritability and gain formulae for alternative selection and deployment schemes are developed and illustrate the use of genetic parameters in the comparison of seedling and clonal reforestation strategies.

Description: Identical pair crosses, including reciprocals, in Pinussylvestris L. (Scots pine) were made on ramets of the same clones in three clonal archives (seed orchards) in Sweden: Sävar (64°N), Röskär (59.5°N), and Degeberga (56°N). The offspring were used to test the hypothesis that the parental environment could affect the performance of the progeny (aftereffects). Growth and freezing tests were performed in the controlled conditions of the Stockholm Phytotron. Parental environment affected seed weight: the heaviest seeds came from Röskär and the lightest seeds, from Degeberga. Height development was affected in the two growth periods tested: seeds from Sävar produced the shortest plants and seeds from Röskär, the tallest plants. There was an effect on the autumn frost hardiness in the first growth period that disappeared after the second growth period. The most hardy progenies came from Sävar. The aftereffects of the parental environment were less than the maternal effects on seed weight and also less than the effects of full-sib families on growth and autumn frost hardiness. Small but mostly significant reciprocal effects were found for height and height increment during the second growth period. There was a significant reciprocal effect for seed weight. Seed weight differences could explain only a small part of the effects on growth and none of the effects on hardiness.

Description: The frost hardiness of 15- to 25-year-old Scots pine (Pinussylvestris L.) and Norway spruce (Piceaabies (L.) Karst.) growing under central Finnish conditions was followed during 1985–1987. Shoots were subjected to artificial frost in the laboratory. Frost hardiness was assessed by the impedance method and by visual scoring. Frost hardiness varied during the years from −3.5 °C to lower than −40 °C. The rate of dehardening increased after about mid-April in both tree species when the daily mean temperature increased by several degrees above 0 °C. The maximum rate of dehardening varied slightly from year to year. In both species the frost hardiness of the previous year's shoot decreased during shoot elongation. This phenomenon was more prominent in pine than in spruce. Shoots were most susceptible to frost damage at the time when shoot elongation was ceasing. The onset and development of hardiness in autumn varied from year to year, especially in spruce. Some difference in hardening was found between the current and the previous year's shoots. The rate of hardening increased typically around mid-September in both species, when the mean daily temperatures decreased to within the range of 5–10 °C.

Description: Sampling of 24-, 34-, and 44-year-old patch cuts (324–2400 m2) in the Bartlett Experimental Forest, New Hampshire, was undertaken to assess the effect of gap size and location within a gap on tree species abundance (relative basal area and relative density). Shade-tolerant species, especially eastern hemlock (Tsugacanadensis (L.) Carr.) and American beech (Fagusgrandifolia Ehrh.), were relatively more abundant in small gaps and gap edges and generally decreased with increasing gap size. Shade-intolerant species, including paper birch (Betulapapyrifera Marsh.) and pin cherry (Prunuspensylvanica L.f.), were relatively more abundant in large gaps and gap centers and increased with increasing gap size. Intermediately shade-tolerant species, especially yellow birch (Betulaalleghaniensis Britt.) and red maple (Acerrubrum L.), were relatively more abundant in gap centers. Striped maple (Acerpensylvanicum L.) was relatively more abundant in gap edges. Many of these relationships were complex due to interactions with gap age and slope. Sugar maple (Acersaccharum Marsh.) relative abundance was not associated with gap size or location within a gap. Analyses isolating irradiance as a factor influencing species composition were inconclusive. Instead, other effects of gap disturbance and characteristics associated with different locations in the gap, such as soil conditions and root competition, may play an important role in the gap dynamics of this northern hardwoods forest. Gap age had a strong effect on species relative abundances and these patterns reflected typical successional sequences in northern hardwood forests. The gap disturbances increased species richness and diversity in this forest. Gaps contained species not present in the old-growth forest, and the species compositional variations among different gap sizes suggest that a forest with a range of gap sizes will have high diversity. Competitive exclusion appeared to be prevented by the gap disturbances, a likely consequence of the release of previously unavailable resources.

Description: Biomass and nutrient dynamics were examined in a subalpine Piceaengelmannii Parry–Abieslasiocarpa (Hook.) Nutt. forest within Loch Vale watershed in north central Colorado by quantifying annual above- and below-ground production, biomass, nutrient pools, and internal nutrient transfers. Subalpine forest covers only 6% of the watershed, and the values reported here refer only to the forested area. Total ecosystem biomass was 42 kg•m−2, of which 30% was soil organic matter, 33% was detrital biomass (including deadwood and forest floor), and 36% was living biomass. Total forest biomass (not including soil organic matter) was 28.9 kg•m−2, of which root biomass was 11%. Net primary production was 520 g•m−2•year−1, of which fine root production was about 27% and foliar production was 30%. Much more N was recycled via fine root turnover than via aboveground litter fall (1.6 versus 0.9 g•m−2•year−1), whereas four times more Ca was returned via litter fall than via fine roots. Compared with other temperate coniferous forests, this subalpine forest had low production. Nutrient resorption contributed between 35 and 38% of the annual requirements of N, P, and K, but only 9% of Ca and 12% of Mg. Although a higher percentage of annual N requirement was met by resorption, this forest used N less efficiently than a similar forest in southwestern Alberta.

Description: Height-growth patterns of black spruce (Piceamariana (Mill.) B.S.P.) in the boreal forest were studied in layer-origin stands released by clear-cutting in the first half of this century. Most stems responded to release brought about by clear-cutting of the main stand. Initially, small stems responded more rapidly and more strongly, but their height-growth rate culminated between 20 and 30 years following release, the same period when height-growth rate culminated for the taller, older stems. After the culmination period, growth rates were independent of height at the time of release. Consequently, the taller stems at time of release were able to maintain their dominant position. Height growth over the recent years indicates that this position could be maintained in the years to come, and even indefinitely. It seems that second-growth black spruce stands behave like even-aged stands despite their uneven-aged structure. Consequently, years since release and stem height are more accurate measures of stem effective age than total age. Moreover, the taller stems in the understory, which are usually considerably older, should be considered to be as valuable as smaller regeneration when a stand is cut.

Description: Analysis was performed on 112 stems of black spruce (Piceamariana (Mill.) BSP) from the Réserve faunique des Laurentides to compare the growth in height, DBH, and volume of layers released by clear-cutting to the growth of individuals of comparable age originating from seeds after fire. The sampled stands originated from fires and clear-cuttings that occurred between 1894 and 1941. Stem analysis also permitted the calculation of specific volume increment, which corresponds to the annual volume increment divided by the surface of the cambium. Results show that height, DBH, and volume measured 60 years after clear-cutting were better correlated to the height of the advanced growth at the moment of release (Ho) than to the number of years of suppression. The mean annual increments in height and DBH of released layers were positively related to height at the year of logging when the height was less than 2 m. When layers were taller, mean annual increments in height and DBH were negatively related to initial height. These relationships were however variable, since for two-thirds of the stand's life, the periodic annual increments in height and diameter did not differ significantly (α = 0.05) between small (Ho  2 m). The superiority of taller layers is consequently due to greater heights before release. Volume growth rate of layers was positively related to initial height until approximately 60 years after clear-cutting. However, no differences in specific volume increment could be associated with initial height. Therefore, the relationship between initial height and volume increment can be attributed to the difference of cambial area between small, medium, and large second-growth spruces. Black spruce originating from seeds had greater height, DBH, and specific volume increments than second-growth trees, until about 50 years after stand origin. Thereafter, growth rates are comparable. Consequently, after 40 years, stands originating from seeds are comparable in height, volume, and DBH to layers that reached between 1 and 2 m in height after clear-cutting. Neither drainage class nor point density had a significant effect on the sampled black spruces. Stem analysis also revealed an important growth reduction that can be associated with the last spruce budworm (Choristoneurafumiferana Clem.) outbreak in the Réserve faunique des Laurentides. We can deduce from the results of this study that the performance of second-growth stands compared with fire-origin stands will mainly depend on the density of the advanced growth and its height structure.

Description: Two-year-old Sitka spruce (Piceasitchensis (Bong.) Cam), Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco), and Japanese larch (Larixleptolepis (Sieb. & Zucc.) Gord.) were cold-stored at 1 °C for 1 to 6 months. In April at the end of the cold storage period, root growth potential and electrolyte leakage from the fine roots were assessed and related to plant survival and height growth. After cold storage, seedlings were planted on a second-rotation, cultivated site. In two experiments planted in 1989 and 1990, fine-root electrolyte leakage was closely correlated with survival and height growth. Fine-root leakage has also practical advantages over other available methods of assessing plant vitality after cold storage.

Description: A system of three conditioned equations is used to describe whole-tree taper for western hemlock (Tsugaheterophylla (Raf.) Sarg.). Geometric attributes of the system are predicted as empirical functions of DBH and total height. This allows for a wide range of predicted stem profiles without risk of illogical profile predictions for any tree size. Fitting is by multivariate nonlinear regression, where the dependent variables are inside-bark diameters at 12 points on each sample tree. Residual analysis shows that the system does mimic the data in the prediction of different profiles for various combinations of DBH and height. The resultant system can produce analytically tractable estimates of volume. The goodness of fit of the compatible volume estimator is comparable with that of the best whole-tree volume equations. A product of the analysis is a multivariate description of the error distribution; this could have application in the development of profile prediction systems that utilize diameter measurements at multiple heights.

Description: Increasing needle chlorosis at a declining Piceaabies (L.) Karst. site in northeast Bavaria, Germany, as well as in other declining P. abies stands in southern Germany, was associated with a decrease in the concentrations of both chlorophyll a and b, and, to a lesser extent, of carotenoids. Needle chlorosis related more to a decrease in the chlorophyll concentration than to a decrease in the ratio of total chlorophyll to carotenoids. In forests growing on relatively acidified soils, concentrations of both chlorophyll a and b, and of carotenoids in chlorotic needles were related only to foliar magnesium concentration. In forests growing on more alkaline soils, chlorophyll concentration increased with potassium and phosphorus concentrations and decreased with calcium and magnesium concentrations. Green needles from a declining site in northeast Bavaria were also compared with those from a nearby healthy site. The chlorophyll concentrations in needles increased more at the healthy site than at the declining site during the growing season and with increasing needle age as predicted by a nutritional disharmony model. The use of foliar chlorophyll and nutrient analyses of needles representing a range in severity of chlorosis is proposed as a quick method of identifying the soil processes which, in some areas, cause forest decline.

Description: Simulated annealing is a stochastic approach to solving large combinatorial problems. This approach was used to model a harvest scheduling problem having block size constraints (no limit, 100–200, and 200–400 ha), a 20-year adjacency delay, and objectives to meet harvest volume targets on the minimum area possible. Spatially explicit harvest schedules complying with the constraints were successfully generated on test data sets of 6148 and 27 548 forest stands.

Description: Nitrogen (N) mineralization potential and net N mineralization insitu were measured monthly over 7 months for the forest floor horizons (Oi, Oe, Oa) and mineral soil (0–15 cm) of a pine stand and the mineral soil (0–15 cm) of a maple stand in Massachusetts, United States. In all cases, N mineralization potential per unit organic matter (anaerobic laboratory incubation) varied significantly by sampling month but was unrelated to the seasonal pattern for net N mineralization (buried-bag method). The organic horizons in the pine stand exhibited the most variable N mineralization potential, with the Oe horizon having more than a fourfold seasonal range. For the pine stand the Oe horizon also had the highest N mineralization potential (per unit organic matter) and the highest net N mineralization insitu (per unit area). In general, temporal and depth-wise variability should be considered when sites are assessed with respect to the pool of mineralizable N.

Description: Herbaceous weed control studies installed by the Auburn University Silvicultural Herbicide Cooperative to examine response to methods and duration of herbaceous weed control in eight loblolly pine (Pinustaeda L.) plantations were analyzed to determine stand response through age 9. Studies were designed to compare weed control treatments with an untreated check, weed control methods (band vs. broadcast), and weed control duration (first year vs. first 2 years). Pine growth was increased by weed control on all sites. Growth was increased by an additional year of weed control (duration) on about one-half of the sites, but did not differ between band and broadcast treatments (method). Age 9 volume response above the check averaged 27.3 m3/ha for first-year weed control and 42.9 m3/ha for the first 2 years of weed control. Individual-tree height growth between ages 7 and 9 did not differ by treatment at most sites, but stand volume growth was higher with weed control at six of the eight sites. Uniformity of individual tree size, as represented by the standard deviation of DBH adjusted for dominant height, was more dependent on survival, hardwood encroachment, and level of fusiform rust stem infection, which varied by treatment and site, than on the result of herbaceous weed control per se. Growth projections made with the least intensive weed control treatment at each site indicated that on average, merchantable volume at age 22 with weed control will equal that of an age 25 stand without weed control. Largest gains were on sites where weed control increased survival.

Description: Soil nutrient availability was assessed on unharvested white spruce (Piceaglauca (Moench) Voss) sites, on a recently harvested site and on 14-year-old postharvested sites stratified into four different regeneration types defined by surface soil conditions and colonizing species. These values were compared with field aboveground biomass production that had been estimated in a previous study and with biomass production of bioassay seedlings. All sites were upland and south facing. On this range of sites, laboratory net N mineralization was the soil characteristic that was the most strongly associated with plant growth in the field as well as in the greenhouse bioassay. The significance of this relationship was mainly caused by the presence of sites regenerating to aspen (Populustremuloides Michx.) which showed high plant biomass production and high soil N availability. Total soil N content, cumulative field soil temperature and soil moisture content were poorly related to N mineralization estimates and to plant biomass production. Soil temperature had an effect on N mineralization and plant growth only when sites where the forest floor had been scraped during the harvesting operations, were included in the computations. Despite a higher soil temperature, these sites showed decreased N mineralization rates and decreased plant biomass production. These results suggest that on south facing postharvested white spruce sites (i) soil temperature does not show enough variability to be an important factor controlling nutrient availability and plant growth unless the soil is severely disturbed, (ii) the rate of N mineralization is controlled by a small pool of rapidly cycling N which is poorly related to forest floor total N concentrations, and (iii) N availability and vegetation production vary with regeneration type.

Description: Competition for water and nutrients between trees and other vegetation is discussed using examples from the interactions between tree and weeds in production forests, and trees and pasture in agroforestry systems. Production and economic viability of plantation forests are dependent upon sound weed management practices. Competition for water and nutrients by a plant is registered as a water deficit or a nutrient deficit. Plant responses to competition are similar to those for coping with water and nutrient deficiency in the soil. One species may have a competitive advantage over another for water and nutrients by (i) acquiring a greater proportion of available soil water and (or) nutrients, (ii) using water and nutrients more efficiently in producing biomass, and (or) (iii) allocating assimilate in ways that maximize survival and growth. The benefits from managing weeds during establishment of a stand have been demonstrated, but the value of managing understory in older stands is unclear. It is not possible to have water stress (through competition) without some degree of nutrient stress, but the opposite may not be the case in some environments. Managing competition effectively requires a clearer understanding of the dynamics of water × nutrient interactions, as well as the dynamics of the interactions between trees and associated vegetation and how this is modified by silviculture. Experiments where variables are well controlled and supported by simultaneous and regular measurements of both water and nutrients are required.

Description: Glyphosate herbicide was applied by helicopter to release small conifer plants from the effects of competition with silvicultural "weeds" on regenerating clearcuts at four study sites. Prior to clear-cutting the study sites were (i) a mixed conifer-hardwood stand in which the regeneration was dominated by red maple (Acerrubrum L.), raspberries, and dicotyledonous herbs; (ii) a conifer stand, with regeneration dominated by heath shrubs, bracken, and birches; (iii) an old-field stand of white spruce (Piceaglauca (Moench) Voss), with regeneration dominated by alder, Asteraceae, and graminoids; and (iv) a mixed conifer–hardwood stand, with regeneration dominated by red maple, raspberries, Asteraceae, and graminoids. The study design included spray and reference (unsprayed) plots at each study site, and prespray surveys of vegetation in permanent plots, followed by as many as 6 years of measurement of regeneration after herbicide treatment. Aerial glyphosate treatment caused large decreases in the abundance of vegetation, especially of pteridophytes and herbaceous and woody angiosperms. There was a substantial recovery of some plant taxa by the end of the first and second postspray growing seasons, especially raspberry and various herbaceous angiosperms. By the end of the study period, no plant taxa had been eliminated, but there were substantial changes in relative abundance. In particular, conifers had a relatively greater abundance on sprayed plots than on unsprayed plots, suggesting that sprayed clearcuts will more rapidly develop into conifer-dominated stands.

Description: The forces and mechanics related to the wind firmness of 30-year-old loblolly pine (Pinustaeda L.) trees were tested by toppling them with a winch and cable system. The ability of trees to resist toppling, expressed as the critical turning moment, was determined by measuring the force exerted by the winch and the height on the tree where the center of force was exerted. Critical turning moments were closely and positively related to stem taper (R2 = 0.91) and various measures of tree size, including tree weight (R2 = 0.96), stem volume (R2 = 0.94), and cubic diameter at breast height (R2 = 0.93). The flexibility of tree stems, measured by the angle of stem deflection during tree pulling, was negatively related to tree size. Measures of center of gravity, crown:stem ratio, and stem moisture content were not significantly related to critical turning moment. Soil moisture content was only weakly significant and negatively related to critical turning moment. With few exceptions, trees subjected to simulated wind stress treatment resembled southern pines subjected to natural acute wind stress. In both cases, root systems were rarely damaged and stem failure occurred instead of uprooting. However, pulled trees tended to break lower on the stem than wind-broken trees.

Description: Torreyataxifolia Arn. is an endangered conifer restricted to ravine slopes along the Apalachicola River in northern Florida. The species began a decline in the 1950s that has resulted in fewer than 1500 trees remaining extant in the wild. Allelic variation was examined for 189 trees throughout its range. The species is characterized by low overall levels of genetic variation: seven of 20 loci were variable; each polymorphic locus had no more than two alleles; mean heterozygosity (He) was 0.064. Over 78% of this measured genetic variation was found within populations; no spatial pattern was detected in the remaining variation between populations within drainage regions, across drainage regions, or across soil types. This genetic information was used to plan tissue collection for a permanent offsite population.

Description: Results of the study of salt-affected soils on the Tanana River floodplain in interior Alaska raised a number of key issues that are important to resolve with future research in this fluvial environment. The results emphasized the prominent role of plant succession in ecosystem structure and function in northern boreal forests. For example, alder plays a crucial role in organic matter and nitrogen dynamics in floodplain ecosystems. This plant species has an impact on ecosystem processes in successional stages beyond its period of dominance on the floodplain. The potential facilitative nature of nitrogen accumulation, mediated by alder, to plant community development during and subsequent to the alder phase of succession should be examined in future studies. Organic matter chemistry and soil temperature appeared to be more important controls of soil processes than high soil salt content. Moreover, secondary plant chemicals may play a role in determining rates of soil nitrification in these floodplain ecosystems. The importance of ion balance to plant nutrition and primary production in salt-affected soils is an important topic for future research. Groundwater may be a unique source of water and nutrients to floodplain plant communities. The magnitude of water flux to the rooting zone in relation to terrace elevation and river level and its importance to plant growth is a significant consideration in the semiarid environment of interior Alaska. The established plant community indicates which species are successful in this environment. Optimum species-terrace combinations may exist that maximize productivity through use of unique moisture and nutrient supplies associated with a shallow water table. Results of this work suggested that new research initiatives are crucial to advance the fundamental understanding of controls of ecosystem processes and as a base of information to support forest resource management.

Description: Seedlings of Alaskan floodplain species (Populusbalsamifera L. (balsam poplar), Alnustenuifolia Nutt. (thinleaf alder), and Piceaglauca (Moench) Voss (white spruce)) and an upland species (Populustremuloides Michx. (trembling aspen)) were grown in early-successional floodplain soils treated with a floodplain salt (calcium sulfate, CaSO4), an osmoticant (polyethylene glycol), and nitrogen. CaSO4 reduced the growth of aspen relative to controls but also reduced the growth of some typical floodplain colonizers (alder at low nitrogen and poplar at high nitrogen). Aspen and poplar were the most rapidly growing species, even when grown with salt or polyethylene glycol. Effects of CaSO4 on growth, therefore, do not explain why aspen is less abundant on the floodplain than are typical floodplain colonizers. CaSO4 reduced growth directly in salt-sensitive species, judging from the insensitivity of water potential, transpiration, and photosynthesis to CaSO4 addition. Tissue concentrations of nitrogen and phosphorus were unaffected by CaSO4, suggesting that the declines in nutrient accumulation by salt-sensitive species in response to CaSO4 addition reflected a decline in nutrient demands for growth rather than being the cause of the reduction in growth. Growth and nutrient accumulation were stimulated by nitrogen addition in all species. We suggest that floodplain salts may be important in succession by slowing the establishment and growth of alder, which is responsible for most of the nitrogen acquired by plants during succession.

Description: Ninety hectares in a treed fen in north central Alberta were drained to improve growth of stagnant black spruce (Piceamariana (Mill.) B.S.P.) and tamarack (Larixlaricina (Du Roi) K. Koch) stands. Installation of 30-, 40-, 50-, and 60-m ditch spacings resulted in a lowering of the average water table by 79, 66, 56, and 73 cm, respectively. The results and the groundwater level criteria used (drainage norm, 40 cm; flood duration limit, 14 days) indicated that the 50-m ditch spacing was hydrologically the most appropriate one for this area. Given the relatively high hydraulic conductivity of the area, it is believed the 30-m spacing was too narrow and resulted in an excessively low average water table. The 60-m spacing was also overeffective, but in this case, overeffectiveness was attributed more to "edge effects" i.e., to site factors such as the proximity to uplands and the small size of upstream source areas, than to the distance between ditches. The results illustrate the importance, for ditch network design purposes, of taking into account hydrologic conditions both within and well beyond the boundaries of an area proposed to be drained. Peat subsidence after drainage appeared to be related to the average drop in water table level and amounted to about 5 cm•a−1.

Description: One possible consequence of predicted increases in global temperature is an increase in soil organic matter decomposition rates and (or) nitrification rates. In nitrogen-saturated ecosystems, such a change could lead to an increase in nitrate production, since much of the ammonium released from decomposition of organic matter would be converted to nitrate rather than taken up by plants or microorganisms. In a high-elevation red spruce (Picearubens Sarg.) stand, soil at 15 cm below the surface of the sunny and shaded sides of a recent clearing were found to exhibit a significant temperature differential (Δ = 1.2 °C, p 

Description: Availability of N and P was compared in the forest floors of old-growth forests of western red cedar (Thujaplicata Donn)and western hemlock (Tsugaheterophylla (Raf.) Sarg.) (CH forests), and second-growth forests of western hemlock and amabilis fir (Abiesamabilis (Dougl.) Forbes) (HA forests) of windthrow origin. Five samples of each forest floor layer (litter, fermentation (woody and nonwoody), and humus (woody and nonwoody)) were collected from three forests of each type (CH and HA). All layers of CH forest floors had smaller concentrations of total and extractable N and mineralized less N during 40-day aerobic incubations in the laboratory. Total and extractable P was lower in the litter layer of CH forest floors. Seedlings of western red cedar, Sitka spruce (Piceasitchensis (Bong.) Carr.), western hemlock, and amabilis fir grown from seed in forest floor material from CH forests grew more slowly and took up less N and P than did seedlings grown in HA forest floor material. The low supply of N and P in CH forest floors may contribute to the nutrient supply problems encountered by regenerating trees on cutovers of this forest type.

Description: Both photosynthetically active radiation penetrating the overstory canopy and overstory leaf area index were determined in forty-three 12 × 12 m plots in even-aged Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stands. Stands were located on the east side of Vancouver Island, British Columbia, between 300 and 800 m on flat to south-facing slopes and were from a wide range of stem densities and stocking levels. In nine core plots total tree leaf area index was estimated using regression equations from individual-tree stem diameter and stand relative density. A single extinction coefficient did not govern the attenuation of photosynthetically active radiation with respect to leaf area index. For a given leaf area index, the extinction coefficient was smaller at low relative densities because of the presence of canopy gaps. Light attenuation models assuming a single extinction coefficient seriously underpredict stand leaf area index, especially at low stand densities. A modified Beer's Law model was used to predict light penetration, which implicitly accounted for foliage clumping.

Description: Rates of net photosynthesis and stomatal conductance of red spruce (Picearubens Sarg.) twigs were measured in field conditions before and after twig detachment on clear midsummer days in New Brunswick. Although these variables did not change significantly within about 6 min after twig detachment, they did drop continually after detachment. Regression models for the rates of net photosynthesis and stomatal conductance were developed that can be used to simulate this drop. Photosynthetically active radiation, leaf temperature, relative humidity, and CO2 concentration were the principal covariates in these models. The regressions improved by including simulated leaf moisture values in the analysis of data obtained after twig detachment.

Description: A litter-bag technique was used to measure decay rates and assess changes in organic and inorganic constituents of ponderosa pine (Pinusponderosa Laws.) needle litter during decomposition over a 2-year period in old- and young-growth forests in the Sierra Nevada of California. Rates of mass loss were among the lowest reported for temperate and boreal forests, with annual decomposition constants of about 0.08 and 0.18 year−1 for the old- and young-growth forests, respectively. Apparently, the temporal separation of warm temperatures and moist conditions found in Mediterranean-type climates severely limits decomposition in these coniferous forests. In the old-growth forest, comparison of estimates of tree nutrient uptake with net releases of nutrients from fine litter during their 1st year of decomposition suggests that recent litter fall potentially acts as a significant source of P, Mg, and K for tree uptake in this forest; in contrast, recently fallen litter acts as a net sink for N, S, and Ca. Despite initially lower indices of litter quality for litter originating from the old–growth relative to the young–growth forest, no significant difference in decomposition rates of these two litter age-classes was found when placed at either site. This result does not support the hypothesis that decreases in decomposition rates during forest development are driven by decreases in the quality of litter fall.

Description: Stem analysis was used to compare the height, diameter at breast height, and volume growth of seven merchantable black spruce (Piceamariana (Mill.) B.S.P.) stands regenerated after harvesting from advance growth of layer origin with the growth of three merchantable black spruce stands regenerated after fire from seed. The year of harvesting in the second-growth stands was precisely determined using synchronous growth release after logging, scars left by the logging operation, and historical records. The year of the fires in seed-origin stands was determined using fire scars and historical records. Fire-origin stands showed typical even-aged structure, and logged, second-growth stands showed an uneven-aged structure associated with an asymmetric curve. When compared with seed-origin stands, layer-origin stands showed a significantly greater total height 30 years after the stand origin because of the initial height of the layers. However, annual height increments were similar between the two origin types at 30 years. The mean diameter increment at 30 years was significantly higher in the second-growth stands. The mean annual specific volume increment values for the entire period of growth were slightly higher for the fire-origin stands. Layers that were small at the time of logging (2 m). There was a significant negative correlation between the height, diameter, and age of the layers at the time of logging and both the mean specific volume increment and the mean annual height increment 30 years after logging. In the second-growth stands, the number of merchantable trees and volume increment increased gradually because of the uneven structure of the stands. In contrast, in the seed-origin stands, the trees attained merchantable size at around 30 years after the fire, and the merchantable volume rose rapidly after this. The layer-origin populations had a significant advantage over the seed-origin populations because of the initial height and diameter of the layers at the time of logging. All seven layer-origin stands achieved, or were predicted to achieve, higher merchantable volumes than the seed-origin stands at 40 years. Our results indicate that the second-growth stands growing on mesic sites have the potential to produce merchantable forests comparable to the yield tables available for black spruce provided that the number of stems per hectare is adequate.

Description: Alkaline soils exist under forest vegetation on the Tanana River floodplain of interior Alaska. The objectives of this study were to describe the soil chemical properties and to examine controls on these chemical properties along a forest primary-successional sequence. Soil saturation pastes were prepared from duplicate sites along the successional sequence representing bare alluvium (stage I), open willow (stage III), poplar–alder (stage V), and white spruce (stage VIII). Calcium, Mg, SO4, and ions responsible for alkalinity were the dominant solutes in the saturation extracts. Soil horizons were generally calcareous (CaCO3) and therefore alkaline (pH 〉 7.0) across the successional sequence. The CaCO3-containing soil horizons were saturated to supersaturated with respect to calcite. Many soil horizons in the plots of stages III and V from one site were saturated with respect to gypsum (CaSO4•2H2O), whereas other stages and sites were generally undersaturated. Phosphate availability in CaCO3-containing soil horizons was apparently constrained by CaCO3 solubility and the solubility products of β-tricalcium phosphate and hydroxyapatite. There was a strong positive relationship between soluble salt content and silt concentration in the early stages, suggesting that initial salt content is controlled by the texture of the alluvial material; the variation in initial texture largely accounts for the site to site variation in salts. Evaporation apparently plays a role in concentrating soluble salts at the surface during the early stages (III and V), but by the late stages (VIII), the biotic and topographic factors apparently reduce surficial evaporation. This reduction in evaporation, coupled with the production of organic acids, leads to a 42% loss of soluble salts from the soils. This study supports our original hypotheses that physical factors (texture, evaporation) are most important early in the successional sequence and that biotic factors (transpiration, acid leaching) are most important late in the successional sequence in controlling salt distribution in soils on the Tanana River floodplain.

Description: Cold climatic conditions govern the productivity of taiga forests, yet within a successional sequence the microclimatic and biogeochemical variations also have a major effect on soil microbial activities, thus affecting plant productivity through nutrient availability. Nitrogen mineralization and nitrification were measured in primary-successional floodplain forests of interior Alaska. Forest floor and mineral soils from an early (open willow), middle (poplar–alder), and late (white spruce) successional stage were used. The effects of temperature, moisture, and NH4+ were tested in the laboratory for each of the successional stages. Potential nitrification was estimated using the chlorate-inhibition technique. Surface mineral soils and white spruce forest floor had low to undetectable rates of nitrogen mineralization and nitrification (

Description: The increasing commercial demand for quaking aspen (Populustremuloides Michx.) has sparked renewed interest in aspen silvics and management. While aspen generally regenerates vigorously by root suckering following clear-cutting, recent reports of unsuccessful regeneration suggest a linkage to harvesting activities. Accordingly we evaluated the effects of harvest season and equipment trafficking on aspen regeneration in a glaciolacustrine landform in northern Minnesota. Regeneration vigor, as measured by stem density, stem height, and crown closure was greater in winter-harvested areas. Increased equipment trafficking significantly reduced regeneration in stands harvested during the summer. We detected no significant increases in soil density associated with equipment trafficking; however, reduced regeneration was correlated with increased amounts of rutting and soil scarification. For a given level of trafficking, aspen regeneration increased along the annual chronosequence of harvesting later and later in the summer.

Description: In remote subarctic North America, instrumental records are very short and sparsely distributed. Yet a long-term understanding of subarctic climate is critical to studies of global change. Annual tree-ring width and maximum latewood density are complementary, high-resolution parameters with different environmental and physiological controls that can be used to assess recent centuries of climatic change. In this paper we present a comparison of the different temperature information inferred from these parameters for white spruce (Piceaglauca (Moench) Voss), a dominant North American latitudinal tree line species. Ring-width and maximum latewood density chronologies (with a common period from 1720–1977) are shown for five sites along a widely spaced transect of the forest–tundra transition in northern Canada. The positive temperature response of maximum latewood density to year to year local temperatures is more consistent and covers a longer portion of the growing season than does that of ring width. Unlike density, the ring-width data show a preference for cold spring conditions. Some, but not all, of the ring-width and density series display increases during the recent century's large-scale climatic warming trend. It is concluded that both types of parameters are necessary for understanding changes in climate and forest dynamics at the northern tree line.

Description: The dynamics of fine (

Description: A process-oriented forest soil temperature model, FORSTEM, is presented. FORSTEM considers vertical heat conduction as well as freezing and thawing, and it lumps the effects of forest canopies on soil surface temperature with the surface heat transfer coefficient. It runs in conjunction with the forest hydrologic model, FORHYM. FORSTEM and FORHYM input is limited to (i) air temperature; (ii) precipitation and its snow fraction; and (iii) descriptive site information (latitude, elevation, slope, aspect, forest coverage, and soil layer thickness and texture). FORSTEM uses generalized parameters derived from existing empirical information. The model was applied to 10 different cover type–site conditions, including lawns, deciduous forests, and coniferous forests before and after clear-cutting in Ontario, Quebec, New Brunswick, and Colorado. The only model parameter we calibrated for different sites was the effective ground/air conductance ratio. The ratio was found to be a function of incoming solar radiation and vegetative area index. Differences between monthly simulations and field measurements fell within ± 1.5 °C for at least about three-quarters of the data cases at individual sites. Major exceptions occurred when temperature measurements showed no damping down the soil profile or with soils containing large air gaps between coarse rock fragments.

Description: A parameter recovery procedure for the Weibull distribution function, based on diameter percentiles, was modified to incorporate the effects of interfering vegetation in young Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco var. menziesii) plantations. The applicability of the system was tested by using data from sites in the Coast Ranges of Oregon and Washington and in the Siskiyou Mountains of southwestern Oregon. Four percentiles (0, 25th, 50th, 95th) of the cumulative probability distribution were predicted as functions of quadratic mean diameter and age. In the Siskiyou study, cover and total vegetation control affected quadratic mean diameter and all four percentiles; intensity of the vegetation treatments affected the 0 and 25th percentiles, and the interaction between intensity and timing of treatment affected quadratic mean diameter. In the Coast Ranges study, only quadratic mean diameter was affected by cover of woody vegetation, while quadratic mean diameter and the 25th percentile were significantly affected by total vegetation control. The predicted distributions showed decreasing variance with increasing cover, particularly in the Siskiyou Mountains. In the Coast Ranges study, the coefficient of variation increased with increasing cover, indicating that the variance of stem diameters was affected by average size. On xeric sites in the Siskiyou Mountains, high diameter variability in plots with total vegetation control suggests that interspecific competition may inhibit the expression of microsite variation.

Description: Landscape ecosystems of a 60-ha area, representative of the pine–mixed hardwood forest of the Changbai Shan Preserve in Jilin Province of northeastern China, were identified, described, and contrasted. Site–species relationships and successional trends were examined together with a comparison of these ecosystems and species with those of northern hardwood forests of eastern North America. Ecosystem components of physiography, soil, and vegetation were used to distinguish two major ecosystem types. The more widespread ecosystem 1 differed from ecosystem 2 in having a flatter topography and more moist and nutrient-rich soil. The overstory of ecosystem 1 was dominated by Tiliaamurensis Rupr., Pinuskoraiensis Sieb. & Zucc, Quercusmongolica Fisch. & Turcz., and Fraxinusmandshurica L., whereas that of ecosystem 2 was dominated primarily by Pinuskoraiensis and Quercusmongolica. Understory species and ground-cover vegetation also reflected the difference in physiography and soil between the two ecosystem types. Six Acer species were recorded; they occurred primarily in the subdominant overstory and the understory of both ecosystems. Without catastrophic disturbance, succession favors the more shade tolerant species in all layers. Pinus and Quercus are rare in the ground cover and understory. Acermono Maxim, is much less dominant than its North American counterpart, Acersaccharum Marsh., in their respective mesic ecosystems in the Changbai Shan forest and forests of western upper Michigan. Fagus and Tsuga, characteristic dominants of northern hardwood forests of eastern North America, are absent. The establishment ecology of Pinuskoraiensis, a five-needled pine with wingless seeds, in the mixed hardwood forest is discussed.

Description: Landform is related to environmental factors that affect site productivity in mountainous areas. I devised a simple index of landform and tested this index as a predictor of site index in the Blue Ridge physiographic province. The landform index is the mean of eight slope gradients from plot center to skyline. A preliminary test indicated that the index was significantly associated with slope position and three classes of landform (ridge, slope, and cove). In a test with data from four locations, site index of yellow-poplar (Liriodendrontulipifera L.) was significantly correlated with landform index for each location (r = 0.45–0.65). Landform index and two other topographic variables together accounted for 31 percent of the variation in yellow-poplar site index throughout the Blue Ridge province. Landform index is a conveniently measured site variable that may be useful in various forestry-related applications, including multivariate analysis of the distribution and composition of forest vegetation.

Description: A stem map showing the symptom status of 110-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) trees along the edge of a large Armillaria center was prepared in 1989 and the area was resurveyed in 1992. The rate of spread of Armillariaostoyae (Romagnesi) Herink during this time was 0.22 m/year. The average time from first appearance of aboveground symptoms (mainly basal resinosis) to tree death was 6 years. Small trees died more quickly than large ones.

Description: This study examines the condition of fine-root systems of healthy and declining sugar maples (Acersaccharum Marsh.) at two sites in Central Ontario, one moderately declining and the other severely declining. Roots are frequently sensitive indicators of soil nutrition, thus ingrowth cores (soil transplants) were used to assess the effect of bulk surface (F-layered Ah horizon) soils collected from beneath healthy or declining trees on fine-root chemistry and growth. Soil at each site was collected from healthy and declining trees and reciprocally transplanted (buried) in mesh bags, and roots were allowed to grow into the soil cores for 8 weeks. In addition to transplants of untreated soils, additional transplants of "healthy" and "decline" soils treated by either (i) steam sterilization to remove pathogens or (ii) fertilization with superphosphate were made. Root dry weight of healthy and declining trees was reduced 25–70% in untreated decline soils compared with healthy soils (p ≤ 0.007). This was consistent with lower Ca, Mg, Mn, and Fe in the soil solution of declining trees at the severely declining site and a lower Mn concentration and Ca/Al and Mg/Al ratios in the soil solution at the moderately declining site. Compared with roots of healthy trees, roots of declining trees had significantly lower Ca concentrations for both stands, and in addition, significantly lower concentrations of P, Mg, S, K, Mn, Al, Fe, and Zn in the more severely declining stand. The experimental soil transplants established that lower nutrient concentrations in soils from declining trees are reflected in the chemistry of roots growing in them. On the whole, the fertilizer treatment significantly increased root growth and the steam sterilization treatment had no effect on root growth. While this study shows that reduced nutrient availability in the rooting substrate is an important factor in decline, it is not known how recently this nutrient depletion has occurred, nor what is the cause. Although other mechanisms may be involved, reduced nutrient availability in the rooting substrate is consistent with the speculation that deposition of acidic salt solutions of sulphate and nitrate to these forest soils has caused accelerated soil base cation leaching losses that are reflected in nutritional deficiencies and growth decline.

Description: The soils and vegetation of 12 stages of forest succession on the floodplain of the Tanana River are described. Succession begins with the invasion of newly deposited alluvium by willows (Salix spp.) and develops through a willow–alder (Alnustenuifolia Nutt.) stage to forest stands of balsam poplar (Populusbalsamifera L.), followed by white spruce (Piceaglauca (Moench) Voss), and finally black spruce (Piceamariana (Mill.) B.S.P.). The principal changes in substrate characteristics during the successional sequence are (i) change from sand to silt loam, (ii) increase in terrace height and distance from the water table, (iii) development of a forest floor, first of leaf litter and then live and dead feather mosses, (iv) burial of organic layers by flooding, and (v) the development of permafrost as soils are insulated by a thick organic layer. Soils and vegetation of six stands occurring in three successional stages used in the salt-affected soils study are described in detail: open willow stands (stage III), balsam poplar–alder stands (stage VI), and a mature white spruce stand (stage VIII). There is a general progression of plant species resulting from the modification of the environment by the developing vegetation and changes in soil characteristics. Life history and stochastic events are important in the early stages of succession, and biological controls such as facilitation and competition become more important in middle and late stages of succession.

Description: Wood density in relation to cambial age and ring width was studied in 23 trees of European oak (Quercuspetraea (Matt) Liebl. and Quercusrobur L.) from northeast France, and based on these relationships, a model predicting intratree wood density through cambial age and ring width was developed. Wood density is closely correlated with both cambial age and ring width, but its relationships with cambial age and ring width, to some extent, vary from tree to tree. The variation, however, does not show an appreciable pattern with tree height. The model developed is able to predict satisfactorily the average wood density of growth rings in European oak, and to explain the main part of the variation in wood density of individual trees. Furthermore, the model appears also possible to predict the average density of a given small piece of wood from any part of the merchantable bole of the trees. A general model, however, accounts for a relatively low percentage of the variation because there is large intertree variability of wood density in the species. The simulation results reveal that the dynamic silviculture applied to European oak might result in the production of denser and more uniform (in the sense of the radial variation of wood density) wood, compared with classical silviculture.

Description: We introduce a new spatially explicit model of forest dynamics. The model is constructed from submodels that predict an individual tree's growth, survival, dispersal, and recruitment, and submodels that predict the local availability of resources. Competition is entirely mechanistic; plants interfere with one another only by depleting resources. We also describe maximum likelihood methods for estimating each of the submodels from data collected in the field. Over the past two years, we collected the necessary data for the dominant tree species in the Great Mountain Forest (Norfolk, Conn.). We report estimates of submodels for each species, and show that the calibrated population dynamic model predicts the structure and dynamics of natural forests. Finally, we contrast our model with the JABOWA–FORET family of forest models.

Description: Wet deposition of nitrogen compounds was measured and dry and cloud water deposition were estimated at 11 forested sites in North America and one site in Europe. Dry deposition was a significant pathway of N input to all the forests, averaging 46% of the total deposition. At most of these sites, NH4+ was the dominant form of fixed N in the air, but HNO3 vapor dominated the dry deposition of N. Coarse-particle deposition was often important, but fine-particle deposition usually contributed only a small amount of the dry-deposited N. The deposition rates of inorganic N, which ranged between 4.8 and 27 kg N•ha−1•year−1, were generally much higher than has been reported by other studies measuring only wet or bulk deposition. The highest deposition rates were at the high-elevation sites in the southeastern and northeastern United States and much of the deposition at these sites was attributed to cloud water. Throughfall and stemflow (TF + SF) flux was also measured at all sites, and the net canopy exchange (NCE = (TF + SF)–total deposition) was found to be negative (indicating consumption of N in the canopy) for NH4+ and NO3−, and positive (indicating canopy release) for organic N. Past reports of canopy release of NO3− can probably be attributed to washoff of dry-deposited NO3− species. Consumption of inorganic N in the canopy ranged from 1 to 12 kg N•ha−1•year−1, and was highest in the spruce and spruce–fir stands. When organic N was included in the canopy N balance, the net canopy uptake of N was generally

Description: Tree-ring data were used to define regional tree-growth anomalies (i.e., recurring spatial patterns of growth that differ from long-term averages) for 11 species growing in a network of sites spanning the deciduous and mixed hardwood–conifer forest boundary in the upper Great Lakes region. Tree-ring samples were collected at 11 sites that are classified as mesic to dry–mesic based on species composition. At each stand at least 20 trees were sampled of each species dominant in the canopy, resulting in one to five species collections per stand and 31 chronologies in total. Principal component analysis was used to define the common variance among the chronologies. Three components explain 57.4% of the variation among the chronologies, indicating that common patterns of tree growth exist within the multispecies network. Component loadings indicate that (i) species to species variation is more important than site to site variation and (ii) species can be segregated into distinct groups based on their common patterns of growth through time. Correlations between the three-component-score time series and climatic data indicate that growth anomaly patterns are weakly, but significantly, correlated with growing-season temperature and precipitation variables. Extreme climatic events (i.e., greater than 1 SD above or below the mean) are very important in generating differential growth rates among the species sampled.

Description: Black birch (Betulalenta L.) and red maple (Acerrubrum L.) trees 1.5–2 m tall responded by compensatory (increased) shoot growth following a single treatment where shoots on the whole tree were clipped. Total shoot length of clipped trees recovered to control levels after 1 year's growth. The remaining buds on clipped shoots grew more than they would have without clipping. Frequency distributions of shoot lengths shifted toward longer lengths in clipped trees in the 1st year, but were similar to controls 2 years after clipping. The longest shoots were as long, or longer, on clipped shoots as on unclipped shoots. The major compensatory growth mechanism was increased growth of lateral buds. Clipping induced a few preventi-tious buds to form shoots and a few unclipped short shoots to grow as long shoots.

Description: Piceamariana (Mill.) B.S.P. seedlings were grown in containers under three levels of N–P–K fertilization. They were inoculated with one of three levels of mycelial slurry of either an isolate of Laccariabicolor (Maire) Orton or of Laccarialongipes G.M. Mueller. Seedlings were grown for 16 weeks in a glasshouse before planting on two different sites (organic and mineral soil) in northern Minnesota. Persistence of the two fungi was monitored over a 2.5-year period. Dikaryotic-monokaryotic pairings indicated that trees on the mineral soil site, inoculated with L. bicolor, remained colonized by that isolate for the entire test period. Controls and L. longipes treated seedlings on the same mineral soil site became colonized by indigenous isolates of L. bicolor. The half-strength fertilizer treatment produced significantly taller seedlings.

Description: Protoplasts were isolated from developing xylem of Pinusbanksiana Lamb, and Pinusstrobus L. by incubating freshly harvested tissue in a cellulose–pectinase mixture having mannitol as osmoticum. Protoplasts were then purified using a discontinuous sucrose–mannitol gradient. More than 70% of the isolated protoplasts were of small diameter (12–27 μm) and had dense cytoplasm and many small vacuoles, suggesting that they originated from ray cells. Larger protoplasts constituted about 25% of the protoplast population; these contained single large vacuoles and only parietal cytoplasm, suggesting that they originated from fusiform cells. Using combined gas chromatography–mass spectrometry, coniferin was confirmed to be present in protoplast preparations. By high-performance liquid chromatography (258 nm UV detection), coniferin was readily detected in protoplasts and in extracts of developing xylem from both species. On a fresh-weight basis, coniferin occurred at 1.0–1.6 mM in protoplasts. In late June, coniferin in developing xylem could be accounted for totally by protoplast coniferin content. In late July, protoplasts contained 93 and 61% of the coniferin content in developing xylem of P. strobus and P. banksiana, respectively.

Description: The effects of acidic atmospheric deposition on leaching of base cations from the canopy and the origin of the major ions in throughfall and stemflow were evaluated in a 2-year study of adjacent deciduous and coniferous forests at Panola Mountain Research Watershed in the Georgia Piedmont. In each forest, the NO3− and SO42− in throughfall and stemflow were derived primarily from atmospheric deposition, whereas the base cations Ca2+, Mg2+, and K+ were derived primarily from canopy leaching. Acidic atmospheric deposition was partially neutralized in each forest. Exchange of H+ with base cations appeared to be the major mechanism for the neutralization of atmospheric acidity by the deciduous canopy. Major neutralization mechanisms could not be differentiated in the coniferous canopy. Base-cation leaching accounted for 86% of the base cations in throughfall and stemflow in the deciduous forest and 69% in the coniferous forest. Exchange with H+ accounted for about 30% of base cations in throughfall in the deciduous forest, whereas it could not be clearly estimated in coniferous throughfall. The current level of acidic atmospheric deposition is hypothesized to have caused an increased leaching of base cations of the deciduous canopy, but methods were insufficient to determine its effect on the coniferous canopy.

Description: Indoor and outdoor experiments demonstrated that allelopathy is an important factor explaining seed regeneration failures of Scots pine (Pinussilvestris L.) in forest floor vegetation dominated by the dwarf shrub Empetrumhermaphroditum Hagerup. Scanning electron micrograph views of the leaf surfaces of E. hermaphroditum reveal secretory glands that are shown to be involved in the release of water-soluble phytotoxic substances. Bioassays indicate that low doses and short exposure times of seeds to leachates have strong negative effects on germination and early root development. Activated carbon can eliminate the inhibitory effects of leachates and litter. This technique demonstrates the occurrence of allelopathic interference by E. hermaphroditum on seed germination of both Scots pine and aspen (Populustremula L.). In a field experiment the allelopathic effects by E. hermaphroditum are strong during early spring when germination and growth initiate and ground ice still is present. Extracts passed through soils collected from an E. hermaphroditum site were detoxified, while those passed through sterilized soil were not. Therefore, microorganisms may detoxify the allelochemicals under some environmental conditions.

Description: Insitu snow evaporation was measured in circular openings, from 0H (full forest) to 5H (where H is the height of the surrounding trees; approximately 20 m), cut in the lodgepole pine (Pinuscontorta Dougl.) forest of the Alberta Foothills. Additional measurements were also made in a large 40H irregular clearing. The ratios of snow evaporation measured in the circular clearings to that measured in the large irregular opening were 0.24, 0.23, 0.28, and 0.41 for the 0H, 1H, 3H, and 5H, respectively. The aerodynamic approach yielded a good estimate of average daily snow evaporation in the large opening, with a computed value of 1.01 mm versus a measured one of 1.07 mm. Snow evaporation in the 0H opening was enhanced over that attributable to turbulent transport by nighttime radiative transfer from the canopy, which kept the snow warmer than in the other openings and prevented condensation. Vertical transfer efficiencies, the slope of the curve relating snow evaporation to the product of horizontal wind speed and vertical humidity gradient, were similar in all clearing sizes in spite of possible fundamental differences in wind fields created by the various canopy–opening interactions. Comparison of data from smaller openings with data from a previous study by A.J. West suggests that differences in snow evaporation between openings can be indexed to opening size.

Description: Assessment of tree health requires accurate estimates of crown condition and identification of specific biotic and abiotic agents that may affect crowns, stems, or roots. ForestHealth, an expert advisory system written in C language for the Macintosh™ platform, provides a diagnostic module and two training modules. The diagnostic module provides guidance in identifying foliar symptoms and the common insects, diseases, and abiotic disorders found on leaves of: sugar maple (Acersaccharum Marsh.), red maple (Acerrubrum L.), black cherry (Prunusserotina Ehrh.), northern red oak (Quercusrubra L.), and white oak (Quercusalba L.). Graphical user interfaces are central to both training modules. The Foliar Severity routine trains users to categorize injury (percent leaf area) on single leaves using a modified Horsfall–Barratt scale. The Crown Dieback routine displays different types and degrees of crown injury that users must identify and classify. Each crown is unique, generated in "real time" using tree–branch parameters selected by the user. Since the crowns are created on a three-dimensional basis, multiple views are possible. Written output from training sessions or periodic checks provide quantitative information for quality assurance and quality control. ForestHealth provides diagnostic assistance, training, quality assurance and quality control data, and standardization for research or survey projects involving forest health.

Description: Four species of chestnut trees and four kinds of hybrids were inoculated with two virulent strains of the chestnut blight fungus (Cryphonectriaparasitica (Murr.) Barr). Rate of canker expansion was measured over 114 days (June–September), and rate change was used as a quantitative assessment of individual tree resistance. Cankers on two typical American chestnut trees (Castaneadentata (Marsh.) Borkh.) expanded at a rate of 1 mm/day. Expansion rates on Chinese chestnut (Castaneamollissima Blume) ranged from 0.2 to 0.6 mm/day, and the rates on two Japanese chestnut trees (Castaneacrenata Siebold & Zucc.) were 0.08 to 0.2 mm/day. Our single Henry chestnut (Castaneahenryi (Skan) Reh. & Wilson) tree allowed rates of 0.3 and 0.8 mm/day. Most rates on American–Chinese and Japanese–American hybrids were from intermediate between rates on Asian and American trees to near the rates on American trees. However, a first-generation backcross of Chinese–American × American only allowed expansion rates of 0.3 and 0.4 mm/day, similar to the lowest rates on Asian trees.

Description: Since 1977, the extent of forest wildfires in the boreal and western regions of North America increased 6- to 9-fold over long-term trends, and an estimated 132 × 106 ha of temperate and boreal forest burned across the northern hemisphere. Emissions during and after burning may have been a significant feedback to global warming. Simulated carbon budgets indicated a hemispheric release of 1.4 Pg C during burning and 4.1 Pg C gross from CO2 fluxes postfire. The total release (5.5 Pg C) was 43% of the biospheric CO2 release to the atmosphere, 1977–1990. Over the next century (1991–2090), continuing emissions from wood and soil decomposition will release an additional 6.9 Pg C gross. A large CO2 release was contrary to assumptions of little net carbon flux in the temperate and boreal forests. The pattern of attenuated CO2 release in northern forests also contrasted with sharp emission peaks in tropical deforestation. A simulation experiment indicated that the CO2 pulse from direct emissions per unit area was 10-fold larger in tropical deforestation than in northern forest wildfires on average; postfire release in the northern systems, however, was about 10 times longer in duration and only slightly less overall than in tropical deforestation fires.

Description: Correlations and functional relationships of sapwood and annual ring cross-sectional areas versus the number of leaves supported inside the live crowns of red oak (Quercusrubra L.) were found, based on analyses of six 20- to 25-year-old oaks from monocultures in southern Ontario. Differences in numbers of leaves supported across intervals inside the crowns were compared, using correlation analyses, with the corresponding differences in conductive sapwood (outer two annual rings) cross-sectional areas, and the current and previous year's annual ring's cross-sectional areas. These analyses showed that the current year's annual ring area had a lower correlation with number of current leaves supported (R = 0.918, P 

Description: The objective of this field study was to test whether Ca and (or) Mg was deficient in two red spruce (Picearubens Sarg.) sites located at 1720 and 1950 m in elevation in the Great Smoky Mountains National Park near Clingmans Dome, North Carolina. Initial current needle Ca and Mg concentrations were, respectively, 1700 and 620 μg/g at the upper site, and 1940 and 670 μg/g at the lower site, suggesting poorer nutrient conditions at the upper site. Twenty-eight saplings at each site stratified by height were involved in an individual-tree fertilization trial. Within each stratum four treatments were applied randomly: (i) no fertilization, (ii) 200 kg/ha Ca as CaCl2, (iii) 100 kg/ha Mg as MgCl2, and (iv) 200 kg/ha Ca as CaCl2 plus 100 kg/ha Mg as MgCl2. Fertilizer was applied in April 1989 and 1990, and needles that subsequently formed in the 1989 and 1990 growing seasons were sampled in November 1989 and 1990, respectively. Post-fertilization nutrient concentrations, needle weights, and nutrient contents were compared through analysis of covariance with the pre-treatment needle weight as covariate. Vector analysis suggested an improvement in Ca nutrition and potential growth response with Ca and Ca + Mg fertilization at the upper site in the 1st year but not at the lower site. Neither site appeared to be Mg deficient. Magnesium fertilization had an antagonistic effect on Ca uptake at both sites, whereas Ca addition seemed to improve Mg uptake. Our study suggests that foliar Mg concentrations of 600 μg/g are well within the sufficiency range, but that red spruce saplings may experience incipient Ca deficiency in the field when Ca concentrations in the current needles are

Description: We studied post-fire Pinusalbicaulis Engelm. seedling establishment in the Sleeping Child and Saddle Mountain burns of western Montana 26 and 28 years, respectively, after fire. Pinusalbicaulis seeds are dispersed primarily by Clark's nutcracker (Nucifragacolumbiana Wilson) in caches of 1–15 or more seeds. Consequently, P. albicaulis seedlings and trees often occur in clusters. In the Sleeping Child burn we collected regeneration data from 63 quadrats along a 3.6-km transect and from 14 quadrats along a 20.6-km road transect; in the Saddle Mountain buhn we collected data from 41 quadrats along a 3.2-km transect. We determined regression models for height versus age and age versus height relationships for P. albicaulis and Abieslasiocarpa (Hook.) Nutt. and used the latter regression models to construct regeneration age structures for both bums. For both conifers a disproportionately high percentage of regeneration was established between 1977 and 1985, about 17–25 years after fire. Thus, major regeneration was episodic. A small increase in mean seasonal rainfall between 1977 and 1985 possibly contributed to this recruitment surge. Age differences of 1–8 years occurred among seedlings in 65–86% of the seedling and tree clusters sampled, suggesting delayed germination of some cached seeds. Pinusalbicaulis regeneration was more frequently associated with Vacciniumscoparium Leiberg and less frequently associated with Xerophyllumtenax (Pursh) Nutt. and Luzulahitchcockii Hamet-Ahti relative to the overall frequency of occurrence of these plants among quadrats.

Description: The effects of thinning and nitrogen fertilization, singly and in combination, on growth of 45-year-old Scots pine (Pinussylvestris L.) trees in northern Sweden were examined for 5 years after treatment. Annual examination of trees determined height growth, diameter growth, crown development, annual biomass increment, aboveground biomass allocation, and needle efficiency in relation to treatments. Nitrogen fertilization increased total dry matter production. The combination of thinning and nitrogen fertilization increased total dry matter production per tree more than the additive effects of thinning and nitrogen fertilization singly. Dry matter production per hectare was increased by nitrogen fertilization. Needle efficiency, i.e., total annual aboveground biomass production per unit of needle mass, was also increased by nitrogen fertilization. The increased aboveground production after nitrogen fertilization resulted from both an increase in biomass of needles and from the increased needle efficiency. The allocation pattern indicated that an increased production of stemwood was a result of a larger biomass of needles. An increase in cone production of the control trees was associated with a decrease in needle efficiency.

Description: Various measurements of the abundance, proximity, and stature of neighbouring plants are utilized as indices of competitive intensity experienced by crop trees. These indices can be useful in assessing the desirability of vegetation control and stand thinnings, and in simulating stand development. Static competition indices, however, have fundamental limitations which should be more widely considered. Competition is usually a constraint to growth, not a determinant of growth, so any measurement of competition alone is fundamentally limited in its ability to predict individual crop tree performance. Differences in the microsite effects of competitors and the autecological responses of crop species result in wide variation in the size of the functional competitive arena, and suggest the need for distinct weightings of competitive intensity for different combinations of species and site. Site differences and weather variation, when coupled with species differences in above- and below-ground allocation, will alter the mode and intensity of competition within a stand and from year to year. The phenology of competitors and focal species will likewise alter interference patterns within a year, making one-time measurements sometimes misleading. Suggested alternatives for determining the importance and intensity of competition include the maintenance of competition-free seedlings ("phytometers") in order to gauge the competitive constraints experienced elsewhere in a plantation. Simulation models of tree growth and stand development also show promise, but they must incorporate the dynamics of noncrop vegetation development. In developing guidelines for the assessment of noncrop interference and the advisability of vegetation control, there is still a need to determine, in a generalizable manner, appropriate survey plot sizes and critical competition thresholds for different combinations of site type, crop species, and vegetation complex.

Description: Alluvial soils on the Tanana River floodplain near Fairbanks, Alaska, were examined for development of physical and chemical properties in relation to soil depth and across a 200-year vegetation development sequence. Development was mediated by ecosystem controls including successional time, vegetation, terrace height, soil physical and chemical properties, and microclimate. These controls interact and are conditioned by the state factors time, flora, topography, parent material, and climate, respectively. On early-successional (

Description: Fifty-nine mixed oak stands in the Ridge and Valley Province of central Pennsylvania were sampled to evaluate the potential for oak replacement by later successional species or oak stability across a diverse landscape. Using species importance, stands were separated into four groups along detrended correspondence analysis axes 1 and 2. Betulalenta L.–Quercusprinus L.–Quercusrubra L. and Q. prinus–Q. rubra groups occurred on sandstone-based Inceptisols and Ultisols on ridges, whereas mixed–Quercus and Quercusalba L. groups were representative of limestone-based Alfisols on valley floors. Quercus species dominated the overstory of all groups; however, the understories mainly comprised shade-adapted Acerrubrum L., Acerpensylvanicum L., and (or) B. lenta on the xeric ridges and A. rubrum and Prunusserotina Ehrh. in the mesic valleys. The presettlement forests consisted of Quercus, Pinus, and Castaneadentata (Marsh.) Borkh. on the ridges and Quercus, Pinus, and Carya in the valleys. Comparisons between presettlement and present-day forests indicate that Pinusstrobus L. and C. dentata have declined dramatically, whereas Quercus species have increased. Changes in disturbance patterns following European settlement (e.g., the charcoal iron industry) initially favored Quercus expansion, followed later by increases in Acer, Prunus, and Betula. Under existing conditions where large-scale disturbances from fire or logging are minimal, Quercus species are expected to decline slowly in importance, being replaced by Acer, Prunus, and (or) Betula species regardless of site moisture relations. Thus, mixed oak forests do not appear to represent a prominent edaphic climax in the region, despite the long history of oak domination.