ALBERT

Beschreibung: Abschnitt 2.6 Bohrlochgeophysik, 10 S.

Beschreibung: The moult fast in emperor penguin (Aptenodytes forsteri) chicks was studied during January 1990 at Drescher Inlet, eastern Weddell Sea. In early January feeding of the chicks had stopped and about 4,000–5,000 chicks were in the inlet. The number of starving chicks decreased rapidly until 26 January when all chicks had either left the inlet or died. Mean body mass loss of starving chicks was 257 g/day and the evaluated specific daily mass loss was 25 g/kg body mass. The critical body mass, i.e. the mass below which chicks die, during moulting starvation was estimated to be 4 kg. Mean body mass was higher and mass loss lower in chicks at more advanced moult stages. Chicks left the inlet before moult was completed, although the sea-ice was still stable.

Beschreibung: The areas of marine pollen deposition are related to the pollen source areas by aeolian and fluvial transport regimes, whereas wind transport is much more important than river transport. Pollen distribution patterns ofPinus, Artemisia, Chenopodiaceae-Amaranthaceae, and Asteraceae Tubuliflorae trace atmospheric transport by the northeast trades. Pollen transport by the African Easterly Jet is reflected in the pollen distribution patterns of Chenopodiaceae-Amaranthaceae, Asteraceae Tubuliflorae, andMitracarpus. Grass pollen distribution registers the latitudinal extension of Sahel, savannas and dry open forests. Marine pollen distribution patterns of Combretaceae-Melastomataceae,Alchornea, andElaeis reflect the extension of wooded grasslands and transitional forests. Pollen from the Guinean-Congolian/Zambezian forest and from the Sudanian/Guinean vegetation zones mark the northernmost extension of the tropical rain forest.Rhizophora pollen in marine sediments traces the distribution of mangrove swamps. Only near the continent, pollen ofRhizophora, Mitracarpus, Chenopodiaceae-Amaranthaceae, and pollen from the Sudanian and Guinean vegetation zones are transported by the Upwelling Under Current and the Equatorial Under Current, where those currents act as bottom currents. The distribution of pollen in marine sediments, reflecting the position of major climatic zones (desert, dry tropics, humid tropics), can be used in tracing climatic changes in the past.

Beschreibung: In den 1980er Jahren wurden umfangreiche Bestandsaufnahmen zur Formenvielfalt der Fischkrankheiten sowie der saisonalen und regionalen Fluktuation ihrer Befallsraten in deutschen Küstengewässern angestellt. Sowohl Formenvielfalt als auch viele der Befallsraten scheinen im Wattenmeer und insbesondere in den dortigen Flußmündungen größer zu sein als in Vergleichsgewässern. Drei “neue” Krankheitsformen werden vorgestellt: die nahrungsinduzierte Maulgranulomatose des Stints, die vermutlich durch Flexibacter-Bakterien verursachte Gelbe Pest des Kabeljaus und eine Fibrom des Steinpickers, in dem lentivirusähnliche Partikel nachgewiesen wurden. Das Schwergewicht der bisherigen Forschungsvorhaben lag auf dem Sammeln von Felddaten. Mangels experimenteller Arbeitsansätze müssen viele wichtige Fragen zur Bedeutung dieser Krankheiten daher heute noch unbeantwortet bleiben. So bleibt die Vermutung unbewiesen, daß ein Reihe von parasitenbedingten Krankheiten einen entscheidenden Beitrag zur natürlichen Sterblichkeit vor allem von jungen Nutzfischen im Wattenmeer liefert. Auch das komplizierte Wirkungsgefüge anthropogener und natürlicher Faktoren bei der Entstehung von Fischkrankheiten im Gebiet wird ohne gezielte experimentelle Arbeiten nicht aufzudecken sein. Das Auftreten von Gelber Pest, Ichthyophonus-Krankheit und lentivirusähnlichen Partikeln in Fischtumoren stellt ein beträchtliches Gefährungspotential für die deutsche Fischwirtschaft dar. Allgemein ist festzustellen, daß in Deutschland eine große Empfänglichkeit gegenüber Umweltproblemen auf dem Gebiet der Marinen Pathologie besteht. Die nationalen Forschungskapazitäten sind aufgrund anderer, meist kurzfristig angelegter Arbeitsschwerpunkte jedoch nicht ausreichend darauf vorbereitet, diese Probleme auch erfolgreich zu analysieren und zu bewerten. Als Konsequenz wird gefordert, der bislang vernachlässigten Grundlagenforschung auf dem Gebiet der Marinen Pathologie neben der reinen Datenerhebung auf See künftig eine größere Bedeutung zuzumessen. Das gilt insbesondere für experimentelle und für biochemisch- und mikroskopisch-diagnostische Arbeiten. Künftig auftretende Probleme werden nur dann rechzeitig zu bewerten und zu lösen sein, wenn sich ein ausreichend breit angelegtes und fundiertes Wissenspotential kontinuierlich weiterentwickeln kann.

Beschreibung: The present study demonstrates that a distinct land-associated community of mesopelagic micronekton exists around the Hawaiian Islands. This “mesopelagic-boundary community” replaces the oceanic mesopelagic community over bottom depths of approx 400 to 700 m and includes about 14 species of fishes, 5 of shrimps and 4 of squids. Similar species of the mesopelagic micronekton have been reported in association with other landmasses at the boundary between the oceanic mesopelagic realm and upper continental or island slopes. These species may form a cosmopolitan “mesopelagic-boundary community” which shows regional differences in taxonomic composition, abundance and diversity. Boundary communities, with populations which are both tightly constrained geographically and relatively accessible to shore-based research programs, offer unique opportunities for studying biological processes of the mesopelagic realm and the interactions between neritic and oceanic populations. Data is presented from three midwater and two neuston sampling projects undertaken around the main Hawaiian Islands between 1987 and 1989; additional evidence from the literature is also discussed.

Beschreibung: A seabird and mammal census was carried out in the north-eastern Weddell Sea during the austral winter of 1986. The German research icebreaker Polarstern operated in heavy pack ice along the Greenwich Meridian between the northern sea ice boundary and the Antarctic coast. Crabeater seals (Lobodon carcinophagus), minke whales (Balaenoptera acutorostrata), Adélie penguins (Pygoscelis adeliae), Antarctic petrels (Thalassoica antarctica) and snow petrels (Pagodroma nivea) were found to be more abundant in the vicinity of the submarine Maud Rise, about 700 km north of the continental margin, than in other areas of substantial ice cover traversed during that cruise. The aggregations of birds and mammals are expected to reflect aggregations of their principal food, krill (Euphausia superba) wintering underneath the ice cover. The distribution pattern of krill predators coincides with the course of a warm water belt upwelling near Maud Rise. This upwelling could induce local ice melting which in turn may result in an increased release of sea ice algae.

Beschreibung: In 1987 and 1989 coelacanths were observed for the first time in their natural habitat with the help of submersibles. Coelacanths were found between 150–253 m depth, their preferential depth seems to be around 200 m; the water temperature ranged between 16.5–22.8° C. During the day coelacanths aggregate in small non-aggressive groups in sheltered lava-caves. Caves might be a limiting factor for distribution. At night they leave the caves for hunting by drifting singly along the steep lava slopes. They migrate between different caves located within a large home range covering more than 8 km coastline. Coelacanths are site-attached, some for a period of at least 2 years. Our own observations and earlier catch records show that only the west coast of Grand Comoro is a suitable coelacanth habitat with more structural complexity and prey fish abundance than other coastlines of the island. From our survey we estimated a total coelacanth population off Grand Comoro to be 150–210 individuals; a saturated population would be 370–510 individuals. This small relict population seems to be stable. International protection of coelacanths against commercial interests is needed.

Beschreibung: We report first observations on social behavior ofLatimeria chalumnae in its natural environment at around 200 m depth in the Comoro Archipelago, Western Indian Ocean. Coelacanths aggregate in small nonaggressive groups in sheltered lava-caves. They live in open site-attached social systems with individuals occupying large overlapping home ranges of more than 8 km coastline, some for a period of at least 2 years.

Beschreibung: This paper describes (i) the Recent distribution pattern of various types of carbonate sediments in Troms county (northern Norway), and (ii) the onset of carbonate sedimentation after the final deglaciation of the Fennoscandian ice-shield. The distribution of major facies belts is strongly dependent on hydrographic and topographic constraints. The main bulk of carbonate deposits is derived from maerl-producing coralline algal biotopes that are restricted to the photic zone of waveprotected areas and influenced by tidal currents. Furthermore, extended mollusc and echinoderm-rich arenites are present in the area investigated. In deeper subtidal areas, terrigenous sediments of Late Weichselian to Early Holocene age are preserved. However, these are strongly influenced by later winnowing processes, generating a coquina lag deposit that serves as a secondary hardground for a diverse fouling community. The Holocene facies successions can be seen in several raised outcrops, containing the transition from glacigenic to Recent non-glacigenic depositional conditions. Autochthonous radiocarbon dated rhodolith banks, which mark the onset of carbonate sedimentation, yielded surprisingly young ages of 5,500 YBP. Around 5,500 YBP, present-day oceanographic and climatic conditions had already been firmly established. This time-lag can be explained with the behaviour of postglacial sea level fluctuations in the area. From 10,000 their cooperation in this project. We address our sincere thanks specifically to Prof. T.O. Vorren, Dr. M. Hald, Dr. B. Gulliksen, Dr. J. Hansen, Dr. J.J. Møller, Dr. E. Thomsen, and Dr. G. Corner for their dedicate help in organizing our field trips in Troms and for extensive scientific discussions. Special thanks to capitain K. Bendiksen for his patience and professional skill while operating the vessel ‘OTTAR’ in the skerry area even under bad weather conditions. S. Rasmussen and G. Ellingsen assisted during the cruises and very kindly introduced us to the Norwegian way of life. Special thanks to the two reviewers (Prof. Dr. W.-Ch. Dullo and anonymus) for their critical but constructive remarks. J. Welling and Dr. M. Kaminski improved the language of the final version of the manuscript. This project is part of the German Science Foundation’s main research program ‘Globale und regionale Steuerungsprozesse biogener Sedimentation”. Financial support was kindly provided by the Deutsche Forschungs-gemeinschaft (He 1671/1-1, 2).

Beschreibung: In the Atlantic sector of the Southern Ocean, data from vertebrate predators and commercial fisheries suggests that the distribution of the ommast rephid squid Martialia hyadesi is related to the Antarctic Polar Frontal Zone, but it spreads further to the north in some years (Rodhouse, in press). A mass stranding of M. hyadesi occurred on Macquarie Island in 1971 (O'Sullivan et al. 1983) suggesting that its distribution is c ircumpolar (Rodhouse and Yeatman 1990). However, apart from a single beak collected from the s tomach of a wandering albatross at Mar ion Island (Imber and Berruti 1981) its presence has not, until now, been confirmed in the Indian Ocean sector and in particular it is not included in the list of cephalopods from the Kerguelenian Province (Lu and Mangold 1978). M. hyadesi is a major prey item of the grey-headed albatross, Diomedea chrysostoma, and the southern elephant seal, Mirounoa leonina, at South Georgia (Rodhouse et al. 1990; Rodhouse et al., unpublished data) and is present in the diet o f several other predators in the Scotia Sea area including the wandering albatross, Diomedea exulans (Rodhouse et al. 1987) and the giant petrels, Macronectes halli and M. gioanteus (Hunter 1983). It occasionally occurs as a significant by-catch in the lllex argentinus fishery on the Pa tagonian Shelf and has been taken during commercial squid jigging trials in the Scotia Sea at the Antarctic Polar Frontal Zone (Rodhouse, in press). It appears to have potential for commercial exploitation in the sub-Antarctic waters of the Atlantic sector of the Southern Ocean (Rodhouse 1990). In view of the ecological importance of M. hyadesi to Antarctic predators, and the likelihood that it will be commercially exploited in the future, it is important to thoroughly establish its geographical range, and in particular to confirm its circumpolar distribution.

Beschreibung: Chemical elements in calcitic foraminiferal tests may serve to reconstruct the spatial and temporal variability as weil as the thermal history of deep and surface water masses. Multi-element analyses of benthic (Cassidulina spp., C. wuellerstorfi, 0. umbonatus, H. elegans, Uvigerina spp.) and planklonie foraminiferal tests (N. pachyderma sin.) performed by miereprobe reveal relations between the geochemical composition of foraminiferal tests and oceanographic and climatic parameters: the analysis of magnesium in calcitic foraminiferal tests makes the thermal reconstruction of surtace water masses possible. Cadmium and barium as paleoproductivity indicators in boliom and surface water masses may reveal oceanographic changes Sediment cores along a transect reaching from the Rockall Plateau in the northeastern North Atlantic to the Fram Strait and lying under the influence of the North Atlantic Drift and the Norwegian Current were used to systematically investigate geochemical variations for several glacial/interglacial changes down to oxygen isotope stage 6 (approximately the last 180,000 years).

Beschreibung: Dinoflagellate cysts have been investigated in surface from the Norwegian-Greenland Sea and short sediment cores Norwegian Sea spanning the last 15,000 years. sediments from the The distribution of single species and assemblages is related to the bathymetry and oceanography of the Norwegian-Greenland Sea. Oceanographic fronts can be recognized in the distribution of species and assemblages. Round protoperidinoid cysts, MuZtispinuZa minuta s.l. and HaZodinium spp., characterize the assemblages from the East Greenland Shelf. Nematosphaeropsis Zabyrinthus and ?Impagidinium paZZidum dominate the assemblages in the central leeland and Greenland Seas. Operculodinium centrocarpum is important in the marginal area of the Arctic domain and dominates clearly the assemblages from the Norwegian Sea. Assernblages from the shelf of north leeland are marked by cysts of Peridinium faeroense. The distribution pattern of single species and assemblages reveals that relatively warm north Atlantic waters only pass through the Faeroe Shetland Canal into the Norwegian-Greenland Sea and further · up into the eastern Arctic Ocean and the Barents Sea. In the southern Norwegian Sea the Atlantic water masses are already modified by advection of surface waters from the North Sea. The development of the Norwegian current has been reconstructed by means of dinoflagellate cysts since termination r •. North Atlantic water masses have almost always influenced the surface water masses in the last 15,000 years. Since ca. 12,000 to 13,000 BP, the influence of warmer North Atlantic waters increased significantly in the NorwegianGreenland Sea. The modern circulation system was established around 10,000 BP. First, the Norwegian current was cooler and less saline, and then obtained its modern hydrographic properties around 6,000 to 7,000 BP. A slight change towards cooler conditions is documented in the western marginal areas in the last 2,000 to 3,000 years. Changes in the oceanography of the Norwegian-Greenland Sea must be related to modifications in the water masses and the circulation system of the entire North Atlantic. The discontinous influx of meltwater may have been an important factor for changes in the oceanography and the ecological conditions.

Beschreibung: Sediment samples from the Mid-Atlantic Reykjanes Ridge (59°N) were taken to get information about sediment genesis and to identify different sources during the late Quaternary. Samples were investigated by X-ray diffraction and grain-size analyses. The clay mineral assemblages in sediments of the Reykjanes Ridge reflect paleoceanographic changes during the late Quaternary. Holocene sediments are characterized by high contents of smectite, mainly of less developed crystallinity. In the spatial distribution of clay minerals high smectite concentrations on the eastern flank and slightly decreasing concentrations on the western flank of the Reykjanes Ridge indicate the action of bottom-water transport. The smectite originates mainly from the volcanogenous Icelandic shelf and reflects the influence of Iceland-Scotland Overflow Water (ISOW). Stratigraphic variability in the clay mineral composition reflects predominantly the influence of different sources, resulting from oceanographic and glacial transport processes. During glacial time sediment transport is due mainly to input by icebergs. Increasing amounts of illite, chlorite, and kaolinite characterize ice-rafted sediments of the “Heinrich layers”. In these sediments smectite crystallinity is well developed. In contrast, several other ice-rafted layers contain smectite with low crystallographic order, similar to smectites of Holocene age. The icelandic source was proved by distinct amounts of basaltic glass in the coarse-grained sediment. At approximately 55 ka increasing amounts of chlorite and kaolinite suggest an enhanced influx of warm North Atlantic surface waters. This hypothesis is supported by a high carbonate shell production at this time. Relative low concentrations and the well-developed crystallinity of smectite minerals characterize the Last Glacial Maximum (LGM; 18–16 ka), indicating a reduced supply of fine icelandic material. Shortly after the LGM, at the beginning of termination IA, a distinct increase in fine-grained quartz (〈2µm) and smectite are visible, which are proposed to reflect a supply of fine-grained ice-rafted material. At 13 ka linear increasing smectite concentrations of lower crystallographic order indicate increasing supply of fine-grained material from Iceland, linked to reinitiation of bottom currents of the ISOW. Full reinitiation is indicated at around 10 ka, where a strong increase in smectite of low crystallographic order is detected.

Beschreibung: Seit jeher will der Geologe wissen, aus welchem Material die Erde besteht, wie ihre Struktur aussieht und welche physikalischen, chemischen und biologischen Vorgänge dabei im Laufe der Erdgeschichte zusammengewirkt haben. Indessen muß er sich, wie alle Historiker, auch heute noch weitgehend damit bescheiden zu ermitteln, wie und wann etwas geschehen ist, und nicht, warum. Daran ist vor allem die Komplexität der Fragen schuld und die Schwierigkeit, einschlägig zu experimentieren.

Beschreibung: Seismic profiles from a venting area on the western margin of Paramushir Island (Sea of Okhotsk) reveal a local complex structure and an interesting, unusual pattern of the bottom simulating reflector (BSR). The BSR is gradual rising towards the venting area. The geothermal gradient and the bottom temperature confirmed the methane hydrate. The temperature appears to be the most important factor controlling the hydrate stability. A locally higher heat flow caused the upward migration of the hydrate stability field and the subsequent degradation of the hydrated sediments, causing gas vent formation and the flux of methane gas into the water column.

Beschreibung: Picritic units of the Miocene shield volcanics on Gran Canaria, Canary Islands, contain olivine and clinopyroxene phenocrysts with abundant primary melt, crystal and fluid inclusions. Composition and crystallization conditions of primary magmas in equilibrium with olivine Fo90-92 were inferred from high-temperature microthermometric quench experiments, low-temperature microthermometry of fluid inclusions and simulation of the reverse path of olivine fractional crystallization based on major element composition of melt inclusions. Primary magmas parental for the Miocene shield basalts range from transitional to alkaline picrites (14.7–19.3 wt% MgO, 43.2–45.7 wt% SiO2). Crystallization of these primary magmas is believed to have occurred over the temperature range 1490–1150° C at pressures ≈5 kbar producing olivine of Fo80.6-90.2, high-Ti chrome spinel [Mg/ (Mg+Fe2+)=0.32–0.56, Cr/(Cr+Al)=0.50–0.78, 2.52–8.58 wt% TiO2], and clinopyroxene [Mg/(Mg+Fe)=0.79–0.88, Wo44.1-45.3, En43.9-48.0, Fs6.8-11.0] which appeared on the liquidus together with olivine≈Fo86. Redox conditions evolved from intermediate between the QFM and WM buffers to late-stage conditions of NNO+1 to NNO+2. The primary magmas crystallized in the presence of an essentially pure CO2 fluid. The primary magmas originated at pressures 〉30 kbar and temperatures of 1500–1600° C, assuming equilibrium with mantle peridotite. This implies melting of the mantle source at a depth of ≈100 km within the garnet stability field followed by migration of melts into magma reservoirs located at the boundary between the upper mantle and lower crust. The temperatures and pressures of primary magma generation suggest that the Canarian plume originated in the lower mantle at depth ≈900 km that supports the plume concept of origin of the Canary Islands.

Beschreibung: Tectonics and climate are both directly and indirectly related. The direct connection is between uplift, atmospheric circulation, and the hydrologic cycle. The indirect links are via subduction, volcanism, the introduction of gasses into the atmosphere, and through erosion and consumption of atmospheric gases by chemical weathering. Rifting of continental blocks involves broad upwarping followed by subsidence of a central valley and uplift of marginal shoulders. The result is an evolving regional climate which has been repeated many times in the Phanerozoic: first a vapor-trapping arch, followed by a rift valley with fresh-water lakes, culminating in an arid rift bordered by mountains intercepting incoming precipitation. Convergence tectonics affects climate on a larger scale. A mountain range is a barrier to atmospheric circulation, especially if perpendicular to the circulation. It also traps water vapor converting latent to sensible heat. Broad uplift results in a shorter path for both incoming and outgoing radiation resulting in seasonal climate extremes with reversals of atmospheric pressure and enhanced monsoonal circulation. Volcanism affects climate by introducing ash and aerosols into the atmosphere, but unless these are injected into the stratosphere, they have little effect. Stratospheric injection is most likely to occur at high latitudes, where the thickness of the troposphere is minimal. Volcanoes introduce CO2, a greenhouse gas, into the atmosphere. Geochemical effects of tectonic uplift and unroofing relate to the weathering of silicate rocks, the means by which CO2 is removed from the atmosphere-ocean system on long-term time scales.

Beschreibung: Factors influencing the fate of ice algae released from melting sea ice were studied during a R V Polarstern cruise (EPOS Leg 2) to the northwestern Weddell Sea. The large-scale phytoplankton distribution patterns across the receding ice edge and small-scale profiling of the water column adjacent to melting ice floes indicated marked patchiness on both scales. The contribution of typical ice algae to the phytoplankton was not significant. In experiments simulating the conditions during sea ice melting, ice algae revealed a strong propensity to form aggregates. Differences in the aggregation potential were found for algal assemblages collected from the ice interior and the infiltration layer. Although all algal species collected from the ice were also found in aggregates, the species composition of dispersed and aggregated algae differed significantly. Aggregates were of a characteristic structure consisting of monospecific microaggregates which are likely to have formed in the minute brine pockets and channels within the ice. Sinking rates of aggregates were three orders of magnitude higher than those of dispersed ice algae. These observations, combined with the negligible seeding effect of ice algae found during this study, suggest that ice algae released from the melting sea ice are subject to rapid sedimentation. High grazing pressure at the ice edge of the investigation area is another factor eliminating ice algae released during melting. Data presented here were collected during the European Polarstern Study (EPOS) sponsored by the European Science Foundation

Beschreibung: High-resolution records of the natural radionuclide230Th were measured in sediments from the eastern Atlantic sector of the Antarctic circumpolar current to obtain a detailed reconstruction of the sedimentation history of this key area for global climate change during the late Quaternary. High-resolution dating rests on the assumption that the230Thex flux to the sediments is constant. Short periods of drastically increased sediment accumulation rates (up to a factor of 8) were determined in the sediments of the Antarctic zone during the climate optima at the beginning of the Holocene and the isotope stage 5e. By comparing expected and measured accumulation rate of230Thex, lateral sediment redistribution was quantified and vertical particle rain rates originating from the surface water above were calculated. We show that lateral contributions locally were up to 6.5 times higher than the vertical particle rain rates. At other locations only 15% of the expected vertical particle rain rate were deposited.

Beschreibung: Hydrographic data along 11°S occupied in 1983 by the R.V. OCEANUS are used together with various wind climatologies to estimate the annual average transport of heat at this latitude. Some motivation for expecting fairly well-defined estimates at this latitude compared to others comes from the absence of a strong western boundary current. Results include flow in four layers representing the thermocline, Antarctic Intermediate Water, North Atlantic Deep Water, and Antarctic Bottom Water, using zero velocity reference level choices based on property distributions. The annual average heat transport is estimated to be 0.6 ± 0.17 x 1015 W. Previous estimates of the transport at 8–16°S range from 0.2 PW to greater than 1 PW. Interannual variability from the wind field alone leads to interannual heat transport variability of about 0.05 PW. Comparisons with other recent studies at 45–30°S and 11°N are made.

Beschreibung: The data from six zonal sections in the World Ocean Circulation Experiment (WOCE) in the tropical and southem Atlantic are used to describe the distribution of water masses. Due to the high spatial resolution, the structure oftemperature, salinity, oxygen, silicate and nitrate displays details related to transport and mixing in this region. Temperature-salinity diagrams are also presented which indicate the effects ofbranching and recirculation loops in the water mass flow.

Beschreibung: During the past quarter century the study of air-sea interaction has evolved from a small branch of marine climatology to play a key role in the modelling of the coupled system of ocean and atmosphere. Knowledge of air- sea fluxes has grown, based on Monin-Obukhov similarity theory for surface boundary layers and on direct and indirect techniques of measuring the fluxes. This has been the basis for providing boundary conditions needed to couple atmospheric and oceanic circulation models that are used to forecast weather and climate. An overview of current understanding is followed by a discussion of parameterisation schemes and a chronicle of some of the experimental work that has tested theories and quantified their conclusions.

Beschreibung: Zooplankton was sampled by a Rectangular Midwater Trawl (RMT 1 + 8) in Weddell Sea surface waters (0 to 300 m) between 66 and 78°S during austral summer (February – March 1983). Sixty-nine taxa including different developmental stages were considered and divided into 16 size classes between 〈1 and 〉39.5 mm length. Biomass was determined by taxon and size class for three different meso- and macroplankton communities in the oceanic region, on the northeastern shelf and on the southern shelf of the Weddell Sea. The highest biomass of 11.2 mg DW m−3 (3.4 g DW m−2) was found in the northeastern shelf community (70 to 74°S), where juvenile and adultEuphausia crystallorophias accounted for 3.7 mg DW m−3 (1.1 g DW m−2). Although not quantitatively sampled, early copepodite stages (CI to CIII) ofCalanoides acutus andCalanus propinquus ranked second with 2.7 mg DW m−3 (0.8 g DW m−2). Biomass in the northeastern shelf community was concentrated in the size ranges 1 to 4 mm and 19.5 to 39.5 mm. The oceanic community of the central Weddell Sea was dominated by copepods smaller than 5 mm, which made up half of the total oceanic biomass. The tunicateSalpa thompsoni (7.0 to 8.5 mm) was the dominant single species with 1.6 mg DW m−3 (0.5 g DW m−2). Euphausiids, mainly juvenile and adult krillEuphausia superba, comprised 1.2 mg DW m−3 (0.4 g DW m−2). Total standing stock in the oceanic community was 9.4 mg DWm−3 (2.8 g DW m−2). Lowest biomass values were found in the southern shelf community (south of 75°S) with 4.0 mg DW m−3 (1.2 g DW m−2), concentrated in the 1 to 4 mm and 14.5 to 34.5 mm size classes. Abundant species were the pteropodLimacina helicina (1 to 2 mm; 0.7 mg DW m−3; 0.2 g DW m−2) andE. crystallorophias (24.5 to 39.5 mm; 0.9 mg DW m−3; 0.3 g DW m−2). The data reveal that it is essential to distinguish among subsystems in the Southern Ocean. This leads to a better understanding of the structure and function of those pelagic food webs which represent alternatives to the paradigmatic krill-centered system.

Beschreibung: The Deep Basin Experiment (DBE), a part of the World Ocean Circulation Experiment (WOCE), is presently underway in the Brazil Basin of the South Atlantic. The program objectives and design philosophy are reviewed and early results are presented. Observations from a moored array along the southern boundary and neutrally buoyant float trajectories in the North Atlantic Deep Water and Antarctic Bottom Water are described with emphasis on their relationship to the recent flow schemes offered by Reid (1989). Also discussed are the process of cross isotherm mixing within the intense flow regime of the Verna Channel and observations of long period warming of the bottom water.

Beschreibung: This is a selected account of recent developments in plankton ecology. The examples have been chosen for their degree of innovation during the past two decades and for their general ecological importance. They range from plankton autecology over interactions between populations to community ecology. The autecology of plankton is represented by the hydromechanics of plankton (the problem of life in a viscous environment) and by the nutritional ecology of phyto- and zooplankton. Population level studies are represented by competition, herbivory (grazing), and zooplankton responses to predation. Community ecology is represented by the debate about bottom- up vs. top-down control of community organization, by the PEG model of seasonal plankton succession, and by the recent discovery of the microbial food web.

Beschreibung: The dependence of results from coarse-resolution models of the North Atlantic circulation on the numerical advection algorithm is studied. In particular, the sensitivity of parameters relevant for climate simulations as e.g., meridional transport of mass and heat and main thermocline thickness is investigated. Three algorithms were considered: (a) a central difference scheme with constant values for horizontal and vertical diffusion, (b) an upstream scheme with no explicit diffusion, and (c) a flux-corrected transport (FCT) scheme with constant and strictly isopycnal diffusion. The temporal evolution of the three models on time scales of centuries is markedly different, the upstream scheme resulting in much shorter adjustment time whereas the central difference scheme is slower and controlled by vertical diffusion rather than advection. In the steady state, the main thermocline structure is much less diffusive in the FCT calculation which also has much lower heat transport. Both horizontal circulation and overturning in the meridional-vertical plane are strongest in the upstream-model. The results are discussed in terms of the effective vertical (diapycnal) mixing in the different models. A significant increase in vertical resolution would be required to eliminate the high sensitivity due to the numerical algorithms, and allow physically motivated mixing formulations to become effective.

Beschreibung: U-series ages measured by thermal ionisation mass spectrometry (TIMS) are reported for a Last Interglacial (LI) fossil coral core from the Turtle Bay, Houtman Abrolhos islands, western Australia. The core is 33.4 m long the top of which is approximately 5 m a.p.s.l. (above present sea level). From the232Th concentrations and the reliability of the U-series ages, two sections in the core can be distinguished. Calculated U/Th ages in core section I (3.3 m a.p.s.l to 11 m b.p.s.l) vary between 124±1.7 ka BP (3.3 m a.p.s.l.) and 132.5±1.8 ka (4 m b.p.s.l., i.e. below present sea level), and those of section II (11–23 m b.p.s.l.) between 140±3 and 214±5 ka BP, respectively. The ages of core section I are in almost perfect chronological order, whereas for section II no clear age-depth relationship of the samples can be recognised. Further assessments based on the ϖ234U(T) criteria reveal that none of the samples of core section II give reliable ages, whereas for core section I several samples can be considered to be moderately reliable within 2 ka. The data of the Turtle Bay core complement and extend our previous work from the Houtman Abrolhos showing that the sea level reached a height of approximately 4 m b.p.s.l at approximately 134 ka BP and a sea level highstand of at least 3.3 m a.p.s.l. at approximately 124 ka BP. Sea level dropped below its present position at approximately 116 ka BP. Although the new data are in general accord with the Milankovitch theory of climate change, a detailed comparison reveals considerable differences between the Holocene and LI sea level rise as monitored relative to the Houtman Abrolhos islands. These observation apparently add further evidence to the growing set of data that the LI sea level rise started earlier than recognised by SPECMAP chronology. A reconciliation of these contradictionary observations following the line of arguments presented by Crowley (1994) are discussed with respect to the Milankovitch theory.

Beschreibung: Planktonic foraminifera first appeared in the Early Jurassic, approx. 200 million years ago. They occurred in great abundance in the oceans only after an important marker Ievel of 130 million years (Early Cretaceous). Since that time, planktonic foraminifera are the most important first ranking Stratigraphie markers and indicators forachanging world's ocean. Knowledge of the ecological demands of living planktonic foraminifera help one to understand their environment in the past because of their high sensitivity to environmental changes; thus, they can be used to reconstruct the oceanic paleoclimate by means of their morphology, faunal assemblage and chemical composition.

Beschreibung: Predation among pelagic cnidarians and ctenophores is reviewed. The diets of semaeostome scyphomedusae and hydromedusae commonly include other gelatinous zooplanktivores. However, few species of siphonophores and ctenophores are known to consume other gelatinous species. Most of these species can be said to exhibit intraguild predation, since they consume species that potentially compete with them for food. In addition, some hydromedusan and ctenophore species may consume other gelatinous zooplanktivores exclusively. Characteristics of cnidarians and ctenophores as predators and as prey of other gelatinous species are discussed.

Beschreibung: Using bacteria that tolerate fairly high sodium sulphate concentrations and reduce sulphate and a coarse clay with buffering properties as solid electrolytes, biological fuel cells have been developed for providing electrical energy discontinuously. The biological active anodes, which accumulate fuel, contained a metal hydroxide catalyst that was converted during the biological process into a metal sulphide catalyst and functioned in a solidified aqueous electrolyte that contained sodium sulphate. The active carbon or titanium cathodes employed were activated with cobalt oxide spinels, molybdenum/vanadium compounds or metal chelates. When employed in continuous service for discontinuous energy consumption, fuel cells were capable of being operated with humus constituents or sugar waste as fuel over a period of 5 years without malfunction and maintenance. Positive results were achieved in the purification of waste-water with a modified type of fuel cell in which the production of energy was not dominant.

Beschreibung: A large-scale trawl survey in August 1989 and data from other surveys indicate that the mesopelagic zone (depth 200-500m) of the Sea of Okhotsk is most densely populated with squids, wit maximum catches of more than 500kg/h. The greatest concentrations occur above the continental slope, particularly in the northern TINRO basin and off the southwestern coast of Kamchatka, adjoining the most productive regions of the sea. Five squid associations are distinguished in terms of the makeup of the catches.

Beschreibung: The systematics of the genus Sepia is not yet clear. Morphological evidence has led to S. officinalis Linnaeus, 1758 being considered as belonging to the subgenus Sepia sensu stricto, and S. obignyana Férusac, 1826 and S. elegans Blainville, 1827 as belonging to the subgenus Rhombosepion. Samples of 30 individuals ocenographic boundary off the north-west Iberian Peninsula, and a sample of S. elegans from the northern side, were collected in 1993-1994. Allozyme electrophoresis for 32 presumptive loci revealed low levels of genetic variability for the three Sepia species (mean expected heterozygosity estimates were 〈0.052). No significant differences in allozyme frequencies were detected among populations of either S. officinalis and S. orbignyana (I=0.12) and of S. elegans (I=0.13) were significantly different from that of S. orbignyana and S. elegans (I=0.49). The former are typical of values for confamiliar genera, and a new generic status is proposed for the latter two species, which become Rhombosepion orbignyana (Férussac, 1826) and R. elegans (Blainville, 1827).

Beschreibung: The energy consumption of Adélie penguins while at rest in water (8.4 W·kg-1 at 4°C) or swimming below the surface was determined using a 21 m long canal fitted with respiration chambers at each end. Penguins chose to swim 86% of the time at speeds recorded in nature. Cost of transport was lowest (7.9 J·kg-1·m-1) at 1.7–2.3 m·s-1, corresponding to a power input of 15.8 W·kg-1, and only 50% as high as previously reported. Assuming a muscle efficiency of 0.25, propulsion efficiency is 0.4 and overall efficiency is 0.1. Calculated food requirements vary between 1060 g krill per adult and foraging trip at the beginning of the breeding season and 2500 g at the period of highest demand, prior to crèching of the chicks.

Beschreibung: To investigate the role of sea ice cover on penguin populations we used principal component analysis to compare population variables of Adélie (Pygoscelis adeliae) and chinstrap (Pygoscelis antarctica) penguins breeding on Signy Island, South Orkney Islands with local (from direct observations) and regional (from remote sensing data) sea ice variables. Throughout the study period, the Adélie penguin population size remained stable, whereas that of chinstrap penguins decreased slightly. For neither species were there significant relationships between population size and breeding success, except for an apparent inverse density-dependent relationship between the number of Adélie breeding pairs and the number of eggs hatching. For both species, no general relationship was found between either population size or breeding success and the local sea ice conditions. However, the regional sea ice extent at a particular time prior to the start of the breeding season was related to the number of birds that arrived to breed. For both species, this period occurred before the sea ice reached its maximum extent and was slightly earlier for Adélie than for chinstrap penguins. These results suggest that sea ice conditions outside the breeding season may play an important role in penguin population processes.

Beschreibung: Major outer membrane proteins with porin activity were isolated from cell envelopes of the halophilic strains Ectothiorhodospira shaposhnikovii N1 and Ectothiorhodospira vacuolataβ1. The porins were obtained as oligomers. They dissociated into monomers by heat or EDTA treatment. The molecular masses of the monomers were determined by mass spectrometry to be 39,285 and 37,160 Da for E. shaposhnikovii N1 and E. vacuolataβ1, respectively. Both were shown by analytical ultracentrifugation to be trimers of about 112,000 Da. Circular dichroism spectra indicated predominantly β-sheet structure. The 18 N-terminal amino acid sequences of the two porins were identical except for the amino acids in positions 12 and 14. No sequence similarity with the primary structure of known porins was found. In reconstitution experiments with lipid bilayers, the porins of E. shaposhnikovii N1 and E. vacuolataβ1 formed channels with a single-channel conductance of 1.5 and 0.7 nS, respectively, in 1 M KCl. The single-channel conductance saturated with increasing salt concentration, indicating a putative binding-site for anions in the channel since both porins exhibited anion-selectivity. For the porin of E. vacuolataβ1, but not for that of E. shaposhnikovii N1, an influence of detergent concentration on the single-channel conductance was observed.

Beschreibung: Direct measurements of magnitude of the northward flow of the Malvinas (Falkland) Current have recently been made with two types of Lagrangian platforms: ALACE floats which cycled between 750-m depth and the sea surface, and 100-m drogued surface drifters. Each data set clearly delineates the path of the Malvinas Current, and the vertical shears inferred from them are commensurate with historical geostrophic shears. Velocities from the surface drifters are used here to adjust geostrophic shears from historical measurements, and the results confirm a large transport of the current, as previously implied by numerical models and a regional inverse calculation. At 42°S, the northward transport of the Malvinas Current in the upper 3000 m appears to be about 70 Sv, several times larger than estimates obtained by adjusting geostrophic shears to assumed levels of no motion. This large barotropie component may have significance in the cross-frontal transfer of intermediate and deep waters from the circumpolar current to the adjacent flow regimes in the South Atlantic, and thus on the inter-basin exchange of water masses.

Beschreibung: Mohns Ridge lavas between 71 and 72°30′N (∼360 km) have heterogeneous compositions varying between alkali basalts and incompatible-element-depleted tholeiites. On a large scale there is a continuity of incompatible element and isotopic compositions between the alkali basalts from the island Jan Mayen and Mohns Ridge tholeiites. The variation in isotopes suggests a heterogeneous mantle which appears to be tapped preferentially by low degree melts (∼5%) close to Jan Mayen but also shows its signature much further north on Mohns Ridge. Three lava types with different incompatible element compositions [e.g. chondrite-normalized (La/Sm)N〈1 to 〉2] occur in the area at 72°N and were generated from this heterogeneous mantle. The relatively depleted tholeiitic melts were mixed with a small degree melt from an enriched source. The elements Ba, Rb and K of the enriched melt were probably buffered in the mantle by residual amphibole or phlogopite. That such a residual phase is stable in this region of oceanic mantle suggests both high water contents and low mantle temperatures, at odds with a hotspot origin for Jan Mayen. Instead we suggest that the melting may be induced by the lowered solidus temperature of a “wet” mantle. Mohns MORB (mid ocean ridge basalt) and Jan Mayen area alkali basalts have high contents of Ba and Rb compared to other incompatible elements (e.g. Ba/La 〉10). These ratios reflect the signature of the mantle source. Ratios of Ce/Pb and Rb/Cs are normal MORB mantle ratios of 25 and 80, respectively, thus the enrichments of Ba and Rb are not indicative of a sedimentary component added to the mantle source but were probably generated by the influence of a metasomatizing fluid, as supported by the presence of hydrous phases during the petrogenesis of the alkali basalts. Geophysical and petrological models suggest that Jan Mayen is not the product of hotspot activity above a mantle plume, and suggest instead that it owes its existence to the unique juxtaposition of a continental fragment, a fracture zone and a spreading axis in this part of the North Atlantic.

Beschreibung: In February 1992, 34 faecal samples from non-breeding male Antarctic fur seals, Arctophalus gazella, were collected at Stranger Point, King George Island, South Shetlands. Fish constituted an important part of the diet, occuring in 90% of those scats containing prey remains. From 1162 otoliths found in the Myctophids and nototheniids represented together almost 90% of the fish eaten. The dominant species were Gymnoscopelus nicholsi, Pleuragramma antarcticum and Electrona antarctica, contributing 33.3%, 30.8% and 12.0% of the otoliths respectively. The standard length of these three species was estimated from otoliths with little or no signs of erosion. This study showed that fur seals fed mainly on pelagic fish species that are often associated with krill. These findings are corroborated by fur seal diving patterns.

Beschreibung: The diet of lactating female Antarctic fur seals Arctocephalus gazella at South Georgia was investigated during the pup-rearing period (January/March) of 1991-1994. Antarctic krill Euphausia superba was the main prey item, occuring in 88% of all scats (n=497), whereas fish occured in 47% and squid in 5% of all scats. There was a considerable intra- and inter-annual variation in the characteristics of krill taken by fur seals. The distribution of krill sizes takes suggests that fur seals are not actively selecting particular sizes of krill available around South Georgia. The absence of group 3 krill (44-48mm in length) in the South Georgia area, as indicated by their absence in the diet of seals, is suggested as a possible reason for low availability of krill and the subsequent reproductive failure among krill predators. The frequency of occurence of fish was much higher than in previous studies; the pattern of fish consumption showed a consistent seasonal pattern in 3 of the 4 years studied. Of the total number of the myctophid Protomyctophum choriodon, the most numerous fish taxon, 98% were taken between early February and the middle of March. Champsocephalus gunnari and Lepidonothen larseni agg., which both feed on krill, dominated the fish component of diet outsided this period and together constituted 94% of the total estimated biomass of fish consumed. The intra- and inter-annual variability in the diet of Antarctic fur seals emphasise the need for diet studies to be conducted during the entire pup-rearing periods of several years.

Beschreibung: The diet of king penguins, Aptendonytes patagonicus, rearing chicks was studied during three consecutive austral winters (1990, 1991 and 1992) at Crozet Islands. The mean stomach content mass of the 47 samples was 503g. Percentages of wet and reconstituted masses showed that both fishes (66 and 36% respectively) and squid (34 and 64%) are important components of the winter diet. Juveniles of the demersal onychoteuthid squid Moroteuthis ingens from the bulk of the cephalopod diet, and this was the main species of mycophids eaten in summer by king penguins were either very rare in winter (Electrona carlsbergi) or accounted for a smaller proportion of the diet (Krefftichthys anderssoni = 1.5% by mass and Protomyctophum tenisoni = 4.6%). Five other myctophids, which are rarely consumed in summer, contributed 24% of the diet by mass in winter (Gymnoscopelus piabilis = 18.1%, Lampichthys procerus = 2.4%, G. nicholsi = 1.3%, and Metelectrona ventralis and Electrona subaspera = 1.0%). The greater diversity of prey in winter suggests a more opportunistic feefing behaviour at a time probably marked by a change in prey availability. Both the known ecology of the fish and squid prey and the barely digested state of some items suggest that in winter breeding adults forage in the outer shelf, upper slope and oceanic areas in the close vicinity of the Crozet Islands to feed their chicks. Finally, using kind penguins as biological samplers, the present work provides novel data on the previously unstudied mesopelagic/epibenthic marine community in waters surrounding the Crozet Islands. Seventeen myctophid fish have been identified to species level. These include several poorly known species in the southern Indian Ocean. The occurence of small, nearly intact, cephalopods in the diet of king penguins suggests that spwaning grounds of four squid species may be located near the Crozet Islands. Seventeen myctophid fish have been identified to species level. These include several poorly known species in the southern Indian Ocean. The occurence of small, nearly intact, cephalopods in the diet of king penguins suggest that spawning ground for four squid species may be located near the Crozet Archipelago.

Beschreibung: Confusion abounds regarding the relative generic status of many member species of the squid family Loliginidae. A taxonomic reorganisation within the family has been proposed in which Loligo species from the Indo-Pacific possessing photophores on the ink sac, including Loligo edulis and Loligo chinensis, are removed to the newly created genus Photololigo. This system of classification has not however gauined general acceptance, and some authors have continued to refer to these species as Loligo. Here biochemical genetic data gathererd using allozyme electrophoresis are presented supporting the assertion that L. edulis and L. chinensis should indeed be positioned in a genus distinct from that characterised by the type species Loligo vulgaris vulgaris. Cluster analysis of allele frequency data from 22 putative enzyme-coding loci suggests that L. edulis and L. vulgaris vulgaris as are members of the confamilial genera Alloteuthis, Uroteuthis and Sepioteuthis, and as such warrant separate generic status. We conclude that the genus Photololigo is valid.

Beschreibung: Paralarval behaviour of eight species of the family Gonatidae (Teuthodiea, Cephalopoda) was examined in small 3-1 aquaria on board ship during planktonic surveys, which were carried out above and off the continental slope of the western part of the Bering Sea. Undisturbed paralarvae moved in aquaria with an average frequency of 15-20 mantle contractions per minute. In response to a sudden disturbance (flash of light, impact to the aquarium wall) squids exhibited a defensive body posture, relaxing the mantle and pulling the head with tentacles and arms into the mantle cavity, thereby becoming similar in appearance, size and colour to small jellyfishes Aglantha digitalis (Hydromedusae)

Beschreibung: Climatologists generally expect an anthropogenic global warming that could raise sea level 50-150 centimeters in the next century and more thereafter. One of the impacts would be the loss of coastal wetlands. Although the inundation of adjacent dry land would enable new wetlands to form, much of this land is or will soon be developed. If developed areas are protected, wetlands will be squeezed between an advancing sea and the land being protected, which has already happened in China and the Netherlands, where people have built dikes for centuries. Unlike those countries, the United States has enough land to accommodate the landward migration of wetlands; but governments lack the funds to purchase all the coastal lowlands that might be inundated and the legal authority to prohibit their development. We propose a third approach: allowing property owners to use coastal lowlands today as they choose, but setting up a legal mechanism to ensure that the land is abandoned if and when sea level rises enough to inundate it. Although compensation may be required, this approach would cost less than 1 percent as much as purchasing the land, and would be (1) economically efficient by enabling real estate markets to incorporate expectations of future sea level rise; (2) constitutional by compensating property owners; and (3) politically feasible by pleasing people who care about the long-term fate of the coastal environment without disturbing people who either are unconcerned about the distant future or do not believe sea level will rise. Because planning today would be more effective than reacting later, the U.S. government should develop a long- term strategy in the next three years.

Beschreibung: Beak length analysis was undertaken for the arrow squid Nototodarus sloanii in the New Zealand sector of the Southern Ocean between Stewart Island and the Auckland Islands. N. sloanii has a restricted range and is endemic to the waters around New Zealand and its associated southern islands, where it supports a fishery and is preyed upon by a number of fishes, marine mammals and birds. Lower rostral length (LRL) and upper rostral length (URL) were analysed to see how these beak measurements related to both mantle lentgh (ML) and wet weight (W). Both lower and upper rostral lengths could be used as useful predictors of ML and W in N. sloanii, as relationships had limited scatter and high correlation coefficients. The relationships between LRL and ML, and URL and ML were only linear after regressing log-transformed values of beak length against ML. However, the relationships between LRL and W, and URL and W were linear without transforming either the x or y values. These results are different from previously published beak length analysis of Nototodarus in New Zealand waters.

Beschreibung: Different initial mixtures of phyto-and zooplankton from different lakes were grown under identical chemical and physical conditions in medium size (8-and 12–1) laboratory microcosm cultures until convergence of phytoplankton species composition was attained. Five such experiments with four (four experiments) or three (one experiment) microcosm cultures were run. Three experiments were performed with weak stirring which permitted sedimentary elimination of the diatoms. Two experiments were conducted with stronger stirring to prevent sedimentation. In the three ldquosedimentation intensiverdquo experiments, the final phytoplankton community was composed of the filamentous chlorophyte Mougeotia thylespora together with a smaller biomass of nanoplanktic algae. In the two ldquosedimentation freerdquo experiments the final phytoplankton community consisted of pennate diatoms. Both dissolved nutrient concentrations and the chemical composition of biomass suggested strong nutrient limitation of algal growth rates in the final phase of the experiments. The zooplankton communities at the end of the experiments were composed of species that were apparently unable to ingest the large, dominant algae and that presumably fed on the nanoplanktic ldquoundergrowthrdquo and the bacteria. There was a distinct sequence of events in all experiments: first, the large zooplankton species (Daphnia and Copepoda) were replaced by smaller ones (Chydorus, Bosmina, rotifers); second, all cultures within one experiment developed the same nutritional status (limitation by the same nutrient); and third, the taxonomic composition of phytoplankton of the different cultures within one experiment converged. The last took 7–9 weeks, with is about 2–3 times as long as the time needed in a phytoplankton competition experiment to reach the final outcome.

Beschreibung: Thorium- and uranium isotopes were measured in a diagenetic manganese nodule from the Peru basin applying alpha- and thermal ionization mass spectrometry (TIMS). Alpha-counting of 62 samples was carried out with a depth resolution of 0.4 mm to gain a high-resolution230Thexcess profile. In addition, 17 samples were measured with TIMS to obtain precise isotope concentrations and isotope ratios. We got values of 0.06–0.59 ppb (230Th), 0.43–1.40 ppm (232Th), 0.09–0.49 ppb (234U) and 1.66–8.24 ppm (238U). The uranium activity ratio in the uppermost samples (1–6 mm) and in two further sections in the nodule at 12.5±1.0 mm and 27.3–33.5 mm comes close to the present ocean water value of 1.144±0.004. In two other sections of the nodule, this ratio is significantly higher, probably reflecting incorporation of diagenetic uranium. The upper 25 mm section of the Mn nodule shows a relatively smooth exponential decrease in the230Thexcess concentration (TIMS). The slope of the best fit yields a growth rate of 110 mm/Ma up to 24.5 mm depth. The section from 25 to 30.3 mm depth shows constant230Thexcess concentrations probably due to growth rates even faster than those in the top section of the nodule. From 33 to 50 mm depth, the growth rate is approximately 60 mm/Ma. Two layers in the nodule with distinct laminations (11–15 and 28–33 mm depth) probably formed during the transition from isotopic stage 8 to 7 and in stage 5e, respectively. The Mn/Fe ratio shows higher values during interglacials 5 and 7, and lower ones during glacials 4 and 6. A comparison of our data with data from adjacent sediment cores suggests (a) a variable supply of hydrothermal Mn to sediments and Mn nodules of the Peru basin or (b) suboxic conditions at the water sediment interface during periods with lower Mn/Fe ratios.

Beschreibung: The Jurassic carbonate series of the Lechtal and Allgäu Nappes in the central part of the Northern Calcareous Alps reflect formation of orogen-parallel structures with swells and basins. Regional facies patterns display the morphologies of the various depositional environments. During the Middle Jurassic, an elongated swell evolved parallel to the overall structural strike in the central part of Lechtal Nappe, while in the southern part a basin started to subside. This configuration reflects the initial stage of rotational block faulting on the southern continental margin of the Tethys. Similar structural and facies settings were also established in the northern part of the Lechtal Nappe and in the southern Allgäu Nappe. Synsedimentary tectonics induced a variety of downslope sediment mass movements and increased facies differentiation on the slopes. In the upper section of the middle Jurassic sequences red nodular limestones with frequent intercalations of intraformational breccias and conglomerates indicate downslope sediment movements. During the Oxfordian, the Tethyan-wide deposition of radiolarites also covered the basin in the southern Lechtal Nappe. Contemporaneous deposition of pelagic radiolarian-bearing limestones dominated on the slope of the surrounding northern swell, while its peak was covered by a shallow water carbonate facies, e.g. a specific pseudopeloid and oolithic facies, which was also injected downslope into the pelagic facies. The Oxfordian to Tithonian section reveals a characteristic pelagic carbonate facies succession, e.g. with Protoglobigerina facies at the base, followed by aSaccocoma facies and a calpionellids facies on top. In the northern Lechtal Nappe and in the Allgäu Nappe various similar radiolarite basins with intersected swells were discovered.