ALBERT

Beschreibung: Seedlings of Avicenniagerminans (L.) Stearn., Lagunculariaracemosa (L.) Gaertn., and Rhizophoramangle (L.) were subjected to flooding, signified by soil redox potentials around −92 mV, and salinity in the range of 342 mol•m−3. Leaf conductance and net carbon assimilation rates per unit area of leaf did not change significantly under flooding or salinity treatments compared with control plants. There was no significant interaction of flooding and salinity with leaf conductance and net carbon assimilation; however, significant reduction in total leaf area per plant in response to flooding (minus salinity) was found in L. racemosa and A. germinans compared with control plants, which would result in a substantial reduction of net carbon assimilation per plant. In R. mangle, total leaf area per plant did not change significantly in response to various treatments. Generally, salinity alone or combined with flooding enhanced dry weights, whereas flooding (minus salinity) resulted in reduced dry weights. The mean values of leaf conductance and net carbon assimilation differed significantly among the study species, with greatest values recorded in A. germinans. The differences in conductance in combination with changes in net carbon assimilation rates resulted in substantial differences in water-use efficiency among these species. Water-use efficiency was greatest in L. racemosa. The overall results showed that these species were tolerant of a wide range of salinity and waterlogging conditions, with differences in physiological responses being evident in changes in biomass partitioning.

Beschreibung: Morphological plasticity in terms of asymmetric display of crowns was compared between Piceaabies (L.) Karst. and Betulamaximowicziana Regel. To evaluate crown asymmetry in relation to local environment, a model that predicts crown asymmetry from topography and configuration of neighbors was applied to data derived from a mixed forest in Hokkaido, northern Japan. Betulamaximowicziana had greater crown asymmetry than P. abies in absolute value. However, observed crown asymmetry of both species was determined by local environment to similar degrees. Some other differences were found between the two species. Crowns of P. abies were more influenced by neighbors than topography, while crowns of B. maximowicziana were more influenced by topography than neighbors. Crowns of P. abies were influenced mainly by larger neighbors, while crowns of B. maximowicziana were influenced by large and by relatively small neighbors.

Beschreibung: Crown recession rates were estimated by branch mortality dating on 357 sectioned Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stems from temporary plots. Numerous nonlinear, logarithmic, and gamma-theory generalized linear models were developed for predicting 5-year crown recession across a range in tree, stand, and site conditions. Residual analyses and indices of fit demonstrated that a multiplicative model with lognormal errors was the most appropriate model form. The recommended logarithmic model predicts crown recession from current crown ratio, total height, breast height age, height growth, and crown competition factor. Data from southwestern Oregon indicate that within a given stand, trees with midsized crown ratios experience the most rapid crown recession.

Beschreibung: The rates of nutrient release were investigated during the Castanopsiskawakamii (Hayata) leaf, branch, and bark litter decomposition processes over a 360-day period within five different size classes of forest gaps that comprise a gap size gradient. Five different size classes of gaps were gap 1, under the closed canopy of pure C. kawakamii vegetation; gap 2, small gaps with a diameter of less than 5 m; gap 3, small to intermediate gaps with a diameter of 5–15 m; gap 4, intermediate to large gaps with a diameter of 15–30 m; gap 5, large gaps with a diameter of bigger than 30 m. After 360 days, decomposing plant litter under canopy condition or small gaps lost weight more rapidly than those in large gaps. Loss of K was highest among all nutrients measured for all three types of litter in all five size classes of gaps. Net immobilization of N and P occurred for all three types of litter in the first 2 or 4 months. Litter residue concentrations of N and P then declined until the end of this study, decreasing to 47% and 70%, respectively, of initial amounts. Loss of Ca and Mg averaged 60 and 40%, respectively, during the period of this study; loss of S averaged 50% and was continuous. Loss of nutrients measured were highest in leaves, intermediate in bark, and lowest in branches under the same size class of gaps, and were highest in gaps 1 and 2, intermediate in gap 3, and lowest in gaps 4 and 5 for the same type of litter. Cellulose and lignin components showed the similar patterns of mass loss as nutrients with highest in gaps 1 and 2, intermediate in gap 3, lowest in gaps 4 and 5 (P ≤ 0.001). Annual decomposition rate was correlated to the microclimatic factors for all types of litter within five different gap size conditions. Soil moisture content was the best predictor of annual decay rate (R2 = 0.922, P ≤ 0.001) among the microclimatic factors. The results indicated that small scale of natural disturbance do not influence the nutrient dynamics during plant litter decomposition; however, rates of nutrient release are strongly inhibited with the increase of scales of disturbance. In this study, gap size of 15 m in diameter is critical in determining the rates of nutrient release from plant litter decomposition processes.

Beschreibung: Six old-growth, late postfire Scots pine (Pinussylvestris L.) dominated forest stands of the Vaccinium–Cladina type were selected along a latitudinal gradient in northern Sweden. In two of the stands, Scots pine seedlings that had naturally regenerated during the last 40 years were surveyed in relation to field- and bottom-layer vegetation. The most abundant forest floor species, viz. Cladina spp., occupied 41% of the ground cover and dominated the microhabitat of Scots pine seedlings

Beschreibung: In northern Patagonia, Argentina, we examined the influences of climatic variation and inter-site variation in substrate stability on the dendroecological effects of earthquakes. In association with the great earthquake in 1960 centered off the coast of nearby Valdivia, Chile, extensive tree mortality occurred in northern Patagonia in Nothofagusdombeyi–Austrocedruschilensis stands on unstable debris fans. To examine the effects of the 1960 and earlier earthquakes on tree growth, we developed tree-ring chronologies from samples of the surviving A. chilensis on unstable debris fan sites and at adjacent nonfan sites of more stable substrates. For controlling the effects of regional climatic variation, we also produced a tree-ring chronology from this species in a more distant and undisturbed stand. Strong variations in tree-growth patterns on fan sites were associated with the historically documented major seismic events of south central Chile that occurred in 1737, 1751, 1837, and 1960. Tree-ring chronologies from nonfan sites (i.e., sites of greater substrate stability) showed much less response to these earthquakes. On the fan sites, strong growth suppressions were associated with the former three earthquakes, whereas strong releases followed the 1960 earthquake. The difference in response is explained by the occurrence of the 1960 earthquake during a period of drought, which in combination with the violent shaking of the ground, resulted in extensive tree mortality followed by growth releases of the survivors. However, severe droughts in the absence of earthquakes also can produce tree mortality and subsequent release of the survivors. Consequently, the synergistic effects of climatic variation and earthquake events must be carefully considered in developing records of both climatic variation and earthquakes.

Beschreibung: The relationships between site index and measures of soil, understory vegetation, and foliar nutrients were examined using data from 102 white spruce (Piceaglauca (Moench) Voss) stands in the Sub-boreal Spruce zone of British Columbia. Compared with soil physical properties (adjusted.R2 = 0.54), soil nutrient properties (adjusted.R2 = 0.29) were poor predictors of white spruce site index. The best soil model, using depth of major rooting zone, total nitrogen, and slope as predictors, explained 60% of the total variance in site index. The best understory vegetation model, using frequencies of soil-moisture and nitrogen indicator species groups as predictors, explained 53% of the total variance in site index. The best foliar nutrient model, using foliar phosphorus and sulphur as predictors, explained 64% of the total variance in site index. Adding variables from understory vegetation, foliar nutrients, or both into the best soil model significantly improved site index prediction. The resultant models explained up to 83% of the total variance in site index. When stands were stratified according to groundwater influence, regression models developed for three delineated strata were significantly improved compared with unstratified models. Based on practical consideration, four regression models were considered for predicting white spruce site index. Independent testing indicated that the stratified model, which used only soil variables as predictors, predicted white spruce site index with 79% of the estimate errors within 2 m in the study area.

Beschreibung: In view of the possible applications of ectomycorrhizae to forestry, this paper discusses the important functions of ectomycorrhizae, the conditions affecting their formation, and methods for the production and application of inoculum. A rationale for selecting appropriate ectomycorrhizal fungi and considerations in selecting sites where ectomycorrhizal seedlings should be planted are presented. Suggestions are also made on encouraging the use of ectomycorrhizal technology. A cost–benefit analysis of inoculation is done.

Beschreibung: Experiments were conducted using black spruce (Piceamariana (Mill.) B.S.P.) seedlings to evaluate diurnal variability in heat tolerance of roots and shoots and in levels of root heat shock proteins (HSPs). Shoot heat tolerance was significantly lower in the morning than in the afternoon. Similarly, root systems exposed to high temperatures in the morning suffered significantly more damage than those exposed in the afternoon. Root HSPs were identified using an antibody specific for the constitutive HSP73 and inducible HSP72. Both the constitutive levels of root HSPs and the levels of root HSPs following heat shock varied diurnally, and in different patterns for different protein fractions. The levels of soluble and nuclear root HSP72/73 were enhanced by heat shock, both in the morning and afternoon. In comparison, HSP levels in the root mitochondrial protein fractions increased after heat shock in the morning, but decreased after heat shock in the afternoon. HSPs in the root microsomal protein fraction declined following both morning and afternoon heat shock. The higher afternoon levels of root heat tolerance were associated with the greater constitutive levels of HSP73 in the mitochondrial and microsomal root protein fractions in the afternoon. To examine the effects of the shoot on diurnal variation in root system heat tolerance, root heat damage was compared between seedlings with intact shoots versus seedlings with shoots excised either 4–6 h prior to, or immediately before, heat treatment. Regardless of time of day, root systems died when shoots were excised and root systems were treated with high temperatures. In contrast, shoot excision or root heating alone resulted in no root mortality.

Beschreibung: The self-thinning rule describes in overcrowded even-aged plant monocultures a consistent relationship of mean mass to the approximately −3/2 power of plant density. Recent analyses revealed cases of statistically significant departures of the exponent from −3/2, confirming earlier suggestions. Shade tolerance, crown structure, and stand conditions have been suggested to influence variation in the self-thinning exponent; however, quantitative relationships of the species' traits to thinning exponent have been unclear. I examined patterns of crown fractal dimension, a quantitative measure of crown structure, and their relationship to the self-thinning exponent in four tree species, Nothofagussolandri var. cliffortioides (Hook. f.) Poole, Pinusbanksiana Lamb., Pinusdensiflora Sieb. et Zucc., and Cryptomeriajaponica (Linn. fil.) D. Don. Only stands at maximum crowding were used for analysis. Crown fractal dimension of monocultures is hypothesized to be a factor determining the species-specific variation in the self-thinning exponent of tree populations and is inversely proportional to the thinning exponent. Species-specific change in foliage packing through development of overcrowded populations is suggested to be closely related to variation in crown fractal dimension and the self-thinning exponent. Implications to possible effects of leaf shape and structural characteristics of crowns to crown fractal dimensions are discussed.

Beschreibung: Repeated measures data occur in a wide variety of experimental situations and are often analyzed without full consideration of the statistical issues involved. In this paper, a discussion of model construction, univariate versus multivariate solutions, and statistical assumptions is motivated by examples from a tree physiology experiment. In addition, several examples from the forestry literature are reviewed. It is hoped that this discussion will help scientists with little statistical training to become aware of the different analyses available and perhaps to recognize the associated models in their own research. The examples range from a simple repeated measures design with one within-subject factor and no between-subjects factors to a more complex design involving multiple within-subject and between-subjects factors. The modelling approach used here permits a straightforward comparison between the univariate and multivariate solutions. Although no single approach is consistently best, the multivariate approach is always appropriate and provides the same interpretations as the univariate approach. However, when appropriate assumptions such as sphericity are met, power considerations tend to favor the more traditional univariate analysis.

Beschreibung: In the assessment of S cycling in forest ecosystems, solutions passing through the forests are normally analyzed for inorganic SO4; other forms of S are rarely considered. In this study, organic S (estimated as the difference between total S and SO4-S) was measured in canopy and soil solutions from eight forest stands spanning a broad range of overstory and soil types. Organic-S concentrations varied among the different types of solutions and among the forests, with values ranging from 0 to 50 μmol S•L−1. Organic S was ≤10% of total S in precipitation, 5 to 54% in throughfall, 1 to 50% in stem flow, 16 to 46% in O-horizon solution, 11 to 21% in A- or E-horizon solutions, and 0 to 29% in B-horizon solutions. Organic S was positively correlated with organic C and organic N in Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and red alder (Alnusrubra Bong.) soil solutions and in Douglas-fir stem flow (r2 = 0.68 to 0.96, p 

Beschreibung: A probabilistic model predicts means and variances of the total number and volume of large woody debris pieces falling into a stream reach per unit time. The estimates of debris input are based on the density (trees/area), tree size distribution, and tree-fall probability of the riparian stand adjacent to the reach. Distributions of volume, length, and orientation of delivered debris pieces are also predicted. The model is applied to an old-growth coniferous stand in Oregon's Cascade Mountains. Observed debris inputs from the riparian stand exceeded the inputs predicted from tree mortality rates typical of similar nonriparian stands. Debris pieces observed in the stream were generally shorter, with less volume per piece, than those predicted by the model, probably because of bole breakage during tree fall. As a second application, predicted debris inputs from riparian management zones of various widths are compared with the input expected from an unharvested stand.

Beschreibung: Ten forest litters with decomposition state varying from 16.6 to 100% weight remaining were partitioned into sub-samples; each subsample was analyzed for proximate carbon fractions using one of two chemical analysis procedures (forage fiber and forest products analyses). Proximate carbon fractions from the simpler forage fiber techniques accurately estimated extractives, cellulose, lignin, and acid-hydrolyzed carbohydrates (R2 〉 0.83) determined by the more complex forest products analyses. Decomposition state accounted for most of the residual variance and significantly improved predictive equations for lignin and extractives. The relationship between proximate carbon fractions from the different techniques also varied somewhat among wood, hardwood leaves, and conifer leaves; however, variations were minor relative to the overall trend. Equations developed can be used to extend data availability for developing and validating decomposition models.

Beschreibung: Limits are frequently encountered in the range of values of independent variables included in data sets used to develop individual tree mortality models. If the resulting model is to be utilized, its ability to extrapolate to conditions outside these limits must be evaluated. This paper describes the development and evaluation of six assumptions required to extend the range of applicability of an individual tree mortality model previously described. The assumptions deal with mortality in very dense stands, mortality for very small trees, mortality on habitat types and regions poorly represented in the data, and mortality for species poorly represented in the data.

Beschreibung: The relevance of organosulfur formation and sulfate adsorption as S retention mechanisms in forest soils based upon the sulfur status of samples collected by horizon was investigated. Several forests of varying elevation, vegetation, location, and soil type were considered. Organic S was found to constitute over 78% of total S in the uppermost mineral (0–20 cm; A,E) horizons. This trend was also observed for both intermediate (20–40 cm; primarily A/B) and deeper (40+ cm; B,C) horizons, where organic S exceeded 65% of total S in all but one site examined. Adsorbed sulfate generally constituted only a minor component of the S pool in the uppermost mineral horizons (

Beschreibung: This paper reports results of a study designed to examine the control that soil temperature exerts on soil processes associated with nutrient flux, and in turn, on tree nutrition in interior Alaska black spruce ecosystems. Approximately 50 m2 of forest floor in a 140-year-old black spruce ecosystem, which had developed on permafrost, was heated to 8–10 °C above ambient temperature. This perturbation amounted to approximately a 1589 degree-day seasonal heat sum (above 0 °C), 1026 degree-days above the control total of 563 degree-days. The forest floor, surface 5 cm of mineral soil, and soil solution were compared with those of an adjacent control plot to evaluate the change in nutrient content and decomposition rate of the forest floor. The nutritional response to soil heating of current black spruce foliage also was evaluated. Soil heating significantly increased decomposition of the forest floor, principally because of an increase in biomass loss of the O21 layer. The increased decomposition resulted in greater extractable N and P concentrations in the forest floor, higher N concentrations in the soil solution, and elevated spruce needle N, P, and K concentrations for the experimental period. These results are discussed in light of the importance of soil temperature and other state factors that mediate ecosystem function.

Beschreibung: Previous research on chronosequences of even-aged northern hardwood stands has suggested that forest clearing is accompanied by large losses of nitrogen from the forest floor. The timing of the losses and the fate of a large fraction of the lost nitrogen are unclear. The purpose of this investigation was to study these questions through direct measurement of soil nitrogen concentrations and pools through time on an experimental catchment cleared in a whole-tree harvest in 1983–1984. Nitrogen losses from the forest floor at the site, the Hubbard Brook Experimental Forest, New Hampshire, were lower than predictions based on previous research. The mean forest floor nitrogen pool was 17% lower 8 years after clear-cutting of the site (P = 0.18). Predictions based on chronosequence studies suggest that 25–40% of the forest floor nitrogen would be lost after 8 years. Mechanical disturbance during logging may play a role in limiting short-term nitrogen losses. The steep midsection of the catchment experienced the greatest losses of nitrogen and carbon, while pools in the relatively flat spruce-fir zone at the upper elevations were unchanged. Carbon was preferentially lost from soil organic matter, relative to nitrogen, resulting in significant decreases in the C/N and C/organic matter ratios in the soil. The N/organic matter ratio was generally unchanged. Nitrogen losses can be limited after clear-cutting by minimizing organic matter losses and promoting rapid regrowth.

Beschreibung: Coarse woody debris from streamside forests plays important biological and physical roles in stream ecosystems. The distance from stream bank to rooting site was determined for at least 30 fallen trees at each study site on 39 streams in the Cascade and Coast ranges of Oregon and Washington. The study sites varied in channel size (first- through third-order), side-slope steepness (3 to 40°), and age of surrounding forest (mature or old-growth stands). The distribution of distances from rooting site to bank was similar among streams, with 11% of the total number of debris pieces originating within 1 m of the channel and over 70% originating within 20 m. Stands with taller trees (old-growth conifers) contributed coarse woody debris to streams from greater distances than did stands with shorter (mature) trees.

Beschreibung: Rooted cuttings of three clones of Sitka spruce (Piceasitchensis (Bong.) Carr.) were grown in 2 m tall transparent acrylic tubes of peat in insulated boxes out of doors. Roots of many of the trees formed mycorrhizas with Thelephoraterrestris Ehrh.:Fr.; the extra-matrical mycelium and associated strands were visible in the tubes, together with the roots. Some of the tubes were waterlogged to submerge the lower part of the root and mycelial systems. Waterlogging was carried out in October, when roots were growing slowly, or in November, when growth had stopped. The fungal mycelium was growing on both occasions. The tubes were drained in the following March, and survival of main roots and fungus was measured to the point of regeneration. There were large differences between the two waterlogging treatments in root survival. October waterlogging caused substantial dieback, and roots survived to a mean depth of only 122 mm below the water table. November waterlogging resulted in little dieback, and survival depth was 308 mm. Differences between clones in root survival were statistically significant but small. The extra-matrical hyphae of Tterrestris died in both treatments, but all of the strand systems survived and regenerated in the following spring. Factors affecting survival are discussed, including growth and oxygen transport in roots and fungal strands.

Beschreibung: We classified red spruce (Picearubens Sarg.) sites from northern Maine by radial growth release history. Two major releases were apparent for a majority of the sites. The first was a reduction and subsequent increase in radial increment in 1920. The second was an increase in radial increment from 1935 to 1955. Red spruce radial growth reduction in the 1960s is apparent only for sites released from 1935 to 1955 (approximately 54% of the sites in this study). These sites are now approaching the radial growth rates of the unreleased stands. Birch dieback is suggested as a probable contributor to the 1935–1955 red spruce growth increase and subsequent 1960s growth reduction.

Beschreibung: Effect of aluminum (Al) on the growth, anatomy, and nutrient content of eastern white pine (Pinusstrobus L.) was determined by growing newly germinated seedlings for 12 weeks in sand irrigated with nutrient solution (pH 3.8) containing 0, 12.5, 25, 50, 75, or 100 mg/L of Al. To determine the influence of mycorrhizal infection on Al toxicity, half of the seedlings were inoculated with the ectomycorrhizal fungus Pisolithustinctorius (Pers.) Coker & Couch. Across Al concentrations, shoot growth (height, needle length, biomass) but not root growth (lateral root length, biomass) was significantly greater in ectomycorrhizal (ECT) than nonmycorrhizal (NM) seedlings. Aluminum significantly decreased shoot and root growth in ECT and NM seedlings but had less effect on growth in ECT seedlings. Foliar symptoms of Al toxicity (chlorosis, tip necrosis) were more pronounced in NM than in ECT seedlings. Cellular and ultrastructural injury to needles and root meristematic tissues was observed at lower concentrations of Al in NM seedlings than in ECT seedlings. Amelioration of Al toxicity by mycorrhizal colonization appeared to result from enhanced uptake of nutrients, especially P, rather than to reduced uptake of Al.

Beschreibung: In a Rocky Mountain aspen forest, the detailed pattern of mass loss from decomposing leaf litter of trembling aspen (Populustremuloides Michx.) during the first 6 months of decay was compared with that from aspen leaves modified to produce a more recalcitrant litter type by removal of leachable material (31.7% of original mass). Leaching litter removed substantial quantities of N (24%) and P (54%), but did not change the litter's C/N ratio (77:1); and leached leaves still contained 33% labile (benzene alcohol soluble) material. Decomposition of intact aspen litter was best described by a double exponential model (k1 = −7.91/year, k2 = −0.21/year), except during the first 2 weeks, when an extremely rapid mass loss (14.2%) apparently resulted from leaching. Microbial metabolism was probably responsible for most of the subsequent decay (35% total in 6 months). In contrast, decomposition of leached aspen showed no exponential trend and was best described by a simple linear regression with a slope of −19.7%/year. Additional data from a 2nd year (12–15 months decay) reduced the regression estimates of decay rates but did not alter the best fit models. Fits were improved slightly if temperature sum replaced time in the regressions, especially if 2nd-year data were included.

Beschreibung: Ecotone vegetation may be especially sensitive to climate change. In particular, the invasion of subalpine meadows by tree seedlings has been well documented. However, there has been no systematic analysis of tree regeneration across the environmental heterogeneity of the alpine forest–tundra ecotone (FTE). Also, the position of the FTE may be relictual from more favorable climates of the past and therefore unresponsive to changing climate. To assess the environmental controls on FTE tree regeneration, to determine whether the ecotone might be relictual, and to determine whether tree invasion of nonforested FTE areas is occurring, we measured tree regeneration in various environments within the FTE of Rocky Mountain National Park, Colorado. Generally, seedling establishment appears to be controlled by patterns of soil moisture. Little seedling establishment was observed in krummholz openings, except for high seedling densities in willow wetlands. Tree seedling invasion of tundra is rare. Therefore, the upper limits of the FTE in Rocky Mountain National Park may be relictual from more favorable climates of the past. Abundant seedling establishment in patch forest openings suggests that patch forest may be poised to change to closed forest.

Beschreibung: Differences in N, P, and K status and N efficiency of some North American willow species and their clones were observed and evaluated. Salixeriocephala Michx., S. lucida Mühl., and S. exigua Nutt., represented by five clones each, were raised from cuttings in potted loamy sand for 113 days. Plants were subjected to three levels of fertilization (equivalent to 125, 250, or 500 kg N/ha) applied at exponential rates of addition (0.06, 0.071, and 0.082, respectively), using complete nutrient solutions and were compared with control plants (no fertilizer added). The experiment showed that nutrient concentration and nutrient uptake in the plants depended not only on treatments but also on species and clones. Thus, species and clones that removed smaller amounts of nutrients per unit of biomass could be selected. Species had a larger effect than clones on nutrient concentration and content. Total nutrient content, at the same treatment level, depended primarily on biomass production. Significant variation existed among species and clones in the ratio of biomass/total N. Salixeriocephala (the best species) produced 35.1% more than the other species, and the best experimental clones within species produced 27.8–41.6% more stem per unit of N than the poorest clones.

Beschreibung: An area-based forest plan is formulated and solved by mixed integer programming and a random search algorithm. This is a computationally difficult problem because operational and environmental constraints require that harvest units and road projects be defined as strict binary variables. It was found that the random search algorithm could easily identify several solutions with objective function values within 10% of the true optimum. The best solution found was within 3% of the optimum. The random search algorithm is simple and can be readily implemented on the microcomputer. It is concluded that the random search algorithm is an effective technique for generating acceptable alternatives to complex area-based planning problems.

Beschreibung: Current research suggests that projected climate change may influence the growth of individual trees. Therefore, growth and yield models that can respond to potential changes in climate must be developed, TREE-BGC, a variant of the ecosystem process model FOREST-BGC, calculates the cycling of carbon, water, and nitrogen in and through forested ecosystems. TREE-BGC allocates stand-level estimates of photosynthesis to "each tree using a competition algorithm that incorporates tree height, relative radiation-use efficiency, and absorbed photosynthetically active radiation, TREE-BGC simulated the growth of trees grown in a dense and an open stand of interior Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) near Kamloops, B.C. The competition algorithm dynamically allocated stand estimates of photosynthesis to individual trees, and the trees were grown using an allometric relationship between biomass increment and height and diameter increment. Asymptotic height growth and the changes in the height–diameter relationship with competition were also incorporated in the model algorithms. Sapwood and phloem volume were used to calculate maintenance respiration. Predicted reductions in diameter growth with stand density were similar to those observed in the study stands. Although the carbon balance of individual trees was not tested, simulated tree diameter increments and height increments were correlated with the actual measurements of tree diameter increment (r2 = 0.89) and tree height increment (r2 = 0.78) for the 5-year period (n = 352). Although the model did not work well with trees that had diameters

Beschreibung: This study related age structure and composition of forests in southeastern Pennsylvania to edaphic properties caused by the activities of a local charcoal iron furnace from the late 18th to the late 19th century. Forests surrounding charcoal hearths were harvested on 20- to 30-year rotations to supply wood for charcoal production in earthmound kilns. Charcoal hearths were created by a recurring cycle of disturbance, including the removal of established vegetation, intense heat, and the deposition of charcoal. Thirty-eight hearths of Hopewell Furnace National Historic Site were examined to describe differences in vegetation relative to surrounding forests that have developed since the cessation of charcoal production in 1883. Hearth forests were significantly younger than surrounding forests, with lower tree density and basal area, decreased sapling and shrub abundance, and higher herbaceous cover. Compositional differences on charcoal hearths included decreased importance of several Quercus species and ericaceous shrubs and increased importance of Liriodendrontulipifera L. Results of this study indicated that the initial effect of charcoal production was the prevention of stump sprouting and a delay in recolonization following disturbance due to extreme soil physical and chemical properties. Failure of hearth seedlings to recruit into larger size classes, age–diameter relationships of Liriodendrontulipifera L. trees, and the failure of calcifuge species to reestablish similar cover on charcoal hearths suggested that unfavorable properties of charcoal soils continue to negatively affect long-term productivity. Relative to nonhearth soils, large, significant differences in soil pH, cation exchange capacity, base saturation, and exchangeable Ca, Mg, and K indicated that continuing inhibition of woody plants on charcoal hearths may be related to alterations in soil chemistry that have persisted for 110 years.

Beschreibung: A model was developed to estimate total leaf dry weight of individual standing trees of Eucalyptusregnans F. Muell. in even-aged, monoculture stands aged 8–20 years. Tree biomass data were collected for a sample of 42 trees from four plots at three sites in Victoria and Tasmania. One plot had been heavily thinned 10 years prior to sampling. The model was based on the pipe model theory, L(x) = as(x)p(x), where L(x) is leaf dry weight above some height (x), s(x) and p(x) are, respectively, stem sapwood area and permeability at x, and a is a parameter. Permeability is known to vary with site and tree conditions, and an empirical model was developed that replaced ap(x) in the pipe model theory with a function in terms of tree age, diameter at breast height over bark, and distance along the stem from the tip of the tree. The resulting model to predict leaf weight appeared to be unbiased across sites, among trees within sites, and with respect to thinning treatment. It was found that given a prediction from the model of total leaf dry weight of one tree, it could be assumed with 95% confidence that the true value of leaf weight fell within an interval bounded by values of −60 to +76% of the estimate. When the model was used to predict stand leaf weight by summing estimates from many individual trees, the interval was bounded by minimum values of −3 to +19% of the stand estimate. These errors of estimate were probably conservative. It appeared that errors of similar magnitude may apply to other published models of this type.

Beschreibung: Thirty-six forest gaps were monitored for 12 years in an old-growth forest dominated by Acersaccharum Marsh. and Fagusgrandifolia Ehrh. Most gaps were formed by single tree falls, although some larger ones were created by the deaths of two to four canopy trees. Gap closure rates suggest that most saplings need the occurrence of more than one gap to reach the canopy. Newer, small-scale disturbances, because of their geometry, occur near older disturbances at high rates. After gaps form, woody vegetation increases in density for 5–10 years and then decreases in density. The four main species of this study show different patterns of response, with optima as follows: small gaps of all ages for Acer, old gaps of all sizes for Fagus, large young gaps for Fraxinusamericana L., and large gaps of all ages for Liriodendrontulipifera L. The presence, size distribution, and species composition of saplings immediately preceding gap formation help determine which species will dominate the gap. In general, the processes occurring in gaps are sufficient to maintain the stand at its current canopy composition. A map of disturbances found in different parts of the eastern deciduous forest suggests that many forests, particularly those located near the center of the biome, renew themselves primarily by way of gaps.

Beschreibung: Precise estimates of biomass are needed in productivity and nutrient cycling studies, and for improved estimates of potential productivity. Improvements in prediction of foliage and branch biomass were sought by comparing multiple regression models using stem diameter, sapwood radial thickness, and tree height as independent variables in stands of Sitka spruce (Piceasitchensis (Bong.) Carr.) in southeast Alaska. Five sites were sampled by stratifying trees into four diameter and three sapwood-thickness classes. Within stands, sample trees with thick sapwood consistently had 2–3 times more foliage and branch biomass than paired trees with thin sapwood but nearly equal diameter. Inclusion of both diameter and sapwood thickness in equations increased precision of foliage and branch biomass, leaf area, and net primary productivity by 15–31% and reduced standard error by 35–48% when compared with equations containing only diameter as an independent variable. Height growth over the last 30 years of intermediate and codominant trees with thick sapwood was 12–27% greater than that of paired trees with thin sapwood but nearly equal diameter at breast height. The addition of total tree height to multiple regression models, however, had little effect on their precision. Stem biomass equations were not improved by including tree height or sapwood thickness. The use of a diameter – sapwood thickness sampling matrix for construction of biomass equations may reduce the sample size needed and result in equations with wider application.

Beschreibung: Extensions of a basic allometric equation applied in predicting total foliage of individual trees were developed to estimate foliage and woody components of individual branches in western hemlock (Tsugaheterophylla (Raf.) Sarg.), Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco), and grand fir (Abiesgrandis (Dougl. ex D. Don) Lindl.). Nine initial equations were fitted on both nonlinear and log-transformed scales to data collected in the western Cascade Mountains of Washington. In all cases, the logarithmic form of each equation provided the best fit to the data based on a modified likelihood criterion and residual patterns. Branch diameter was the overall best predictor of branch foliage and woody mass; however, significant (p = 0.05) improvements in fit were obtained when other structural and positional variables were included in the equations (e.g., foliated branch length and depth into crown). The effects of fertilization and site (locality) differences were explored using indicator variables for western hemlock branches. Significant site effects were observed for intercept terms for all three branch components (total foliage area, total foliage mass, and total woody mass), while fertilization effects were observed in interaction with relative height above crown base for both foliage components.

Beschreibung: A field experiment was established between 1989 and 1993 to study the effects of (i) the age of clearcuts on damage by pine weevils (Hylobiusabietis L.) and (11) competing vegetation on the survival and growth of planted Norway spruce (Piceaabies (L.) Karst.). On each of four sites Norway spruce seedlings were planted on 0 to 4-year-old clearcuts. Effects of mounding, herbicide and mowing, removal of slash, and seedling stock type were also investigated. As many seedlings suffered severely from a drought that affected southern Sweden in 1992, this study reports a separate analysis of mortality due to drough. From the middle of May until the beginning of July 1992, only negligible precipitation was recorded on all four sites. The biomass of ground vegetation was correlated with the age of the clearcut. On fresh and 1-year-old clearcuts, only negligible amounts of vegetation were found, whereas about 2 Mg•ha−1 of ground vegetation were found on 2- and 3-year-old clearcuts. Low soil water potentials were recorded on 2- and 3-year-old clearcuts but not on fresh clearcuts. There was no effect of the slash removal treatment on soil water potential, but there was a significant effect of the soil and vegetation treatments. Mounding and herbicide treatments increased the soil water potentials compared with untreated controls and mowed plots. Mortality by drought was higher for seedlings planted on old clearcuts than for seedlings planted on fresh ones. Mortality was reduced by mounding, but late planting increased the number of dead seedlings significantly. The mortality of containerized seedlings was considerably less than that of bare-root seedlings. On old clearcuts growth of seedlings planted on plots with no vegetation control (untreated control and mown plots) was less than that of seedlings planted on plots where transpiration by ground vegetation was reduced (herbicide and mounding).

Beschreibung: Measurements of aboveground biomass and nitrogen (N) nutrition were made during July 1993 in 50-, 130-, and 380-year-old stands of Larixgmelinii (Rupr.) Rupr. in eastern Siberia. Constituting six forest types based on understorey plants, the stands were representative of vegetation throughout the Yakutsk region. Average tree height, diameter, and density ranged from 2 m, 23 mm, and 50 800 stems/ha in the 50-year-old stand to 11 m, 160 mm, and 600 stems/ha in the oldest stand. Aboveground biomass in the 50-year-old stand was 4.4 kg•m−2, and the aboveground N pool was 1.1 mol•m−2. This was slightly higher than the N pool in a 125-year-old stand with a Ledum understorey (1.0 mol•m−2), despite its higher biomass (7.2 kg•m−2). The highest observed aboveground biomass in a 125-year-old stand (characterized by the N2-fixing understorey plant Alnasterfruticosa) reached 12.0 kg•m−2, but the corresponding N pool was only 1.6 mol•m−2. In the oldest stand, aboveground biomass was 8.9 kg•m−2 and the N pool was 1.1 mol•m−2. There was thus a relatively constant quantity of N in the aboveground biomass of stands differing in age by almost 400 years. We postulate that N sets a limit on carbon accumulation in this boreal forest type. Trees were extremely slow growing, and there was essentially no aboveground biomass accumulation between the ages of 130 and 380 years because of a lack of available N. This conclusion was supported by graphical analysis indicating that the self-thinning process in our stands was not governed by the availability of radiation according to allometric theory. Much of the available N was used in the production of tree stems where 86% of the aboveground N (and 96% of aboveground biomass) was immobilized in the oldest stand. N in wood of the old stand exceeded the N pool in the litter layer and was 20% of the N pool in the Ah horizon. The processes of carbon and N partitioning were further explored by the estimation of carbon and N fluxes during three periods of forest development. We calculated a loss of ecosystem N during the period of self-thinning, while in the mature stands the N cycle appeared to be very tight. The immobilized N is returned from the wood into a plant-available form only by a recurrent fire cycle, which regenerates the N cycle. Thus fire is an essential component for the persistence of the L. gmelinii forest.

Beschreibung: Riparian forests are often identified as prime habitat for wildlife because of the presence of particular plant communities and edges creating a highly developed and diversified vegetation structure. However, in the northeastern boreal forests of Canada, where narrow land–water ecotones with abrupt edges are quite common, the relative habitat value of riparian forests remains to be demonstrated. We compared bird communities of eight pairs of riparian and nonriparian plots, similar in vegetation structure and composition, to verify the relative value for breeding birds of typical coniferous riparian forest stands of the southern boreal region of eastern Quebec. Bird abundance (P = 0.02), richness (P = 0.03), and diversity (P = 0.02) were significantly higher in the riparian stands, where the spatial sequence of three distinct habitats (a balsam fir (Abiesbalsamea (L.) Mill.)–northern white cedar (Thujaoccidentalis L.) forest, a narrow alder (Alnusrugosa (Du Roi) Spreng.) - grass wetland, and water) created high horizontal vegetation diversity. In riparian stands median bird richness and density were, respectively, 23.5 species and 437.5 territories/km2 compared with 19.0 species and 348.2 territories/km2 for interior stands. Nine species were observed exclusively in riparian plots. In addition to the species usually found in the studied nonriparian forests, the riparian plots were used by species typically related to the water edge such as the Northern Waterthrush (Seiurusnoveboracensis Gmelin) and Rusty Blackbird (Euphaguscarolinus Miiller) as well as species associated with the shrub and grass wetland such as the American Robin (Turdusmigratorius L.), the Veery (Catharusfuscescens Stephens), and the Common Yellowthroat (Geothlypistrichas L.). The shrub-foraging guild showed higher abundance in riparian stands. The tree-foraging and tree-bole foraging guilds, however, were comparable in both groups of stands. In this study, the natural conditions prevailing along he riparian sites appeared mostly positive for the breeding-bird community; it created what is perceived as being an "edge effect." The edge effect can be defined, in this case, as being the additional density and number of species induced by the added horizontal vegetation diversity created by the close association of three extremely different ecosystems: a forest stand, an aquatic ecosystem, and a narrow but distinct shrub-grass wetland. This also confirms the necessity of distinguishing natural edges that are permanent features of the landscape from induced edges created by human activity.

Beschreibung: A protocol is described for plantlet formation in juvenile tissues of Pinuscanariensis Sweet ex K. Spreng. (Canary Island pine). Adventitious buds were induced on 3-day-old cotyledonary explants cultured on Bornman's MCM medium supplemented with cytokinin. The concentration of benzylaminopurine, the use of other cytokinins alone or in combination with benzylaminopurine, and the concentration of mineral salts strongly affected the bud forming capacity of the cotyledonary explants. Also, the age of the explants significantly influenced the frequency of adventitious bud formation. Bud development was enhanced by the elimination of phytohormones, a reduction of mineral salts and sucrose, and the inclusion of activated charcoal in the medium. The conditions used during the induction phase strongly affected the ability of the induced buds to develop into vigorous rootable shoots. Vitrification problems were eliminated by transferring the shoots to the elongation medium solidified with Gelrite, and shoot remultiplication was enhanced by removing the apical bud. Maximum rooting was obtained by pulsing shoots with a high concentration of indolebutyric acid and by using peat–vermiculite or peat–vermiculite–perlite as substrates. Roots developed within 6–8 weeks, and the regenerated plantlets were transferred to soil under nonsterile conditions, where further development occurred.

Beschreibung: High cuticular water loss, desiccation, and leaf mortality during winter in conifers at alpine timberline represent especially dramatic examples of the ecophysiological importance of cuticular transpiration. However, little research has been conducted on the winter water relations of forest trees, even though recent evidence suggests a potentially high leaf mortality due, at least in part, to winter desiccation. Two factors that could have important influences on cuticular transpiration, leaf surface wax and the ratio of leaf area (A) to saturated water content (Mw), were compared for six conifer species common to the central Rocky Mountains, United States. Leaves were collected from forest trees at the end of winter (May) to compare the amount of leaf surface wax, cuticular conductance to water vapor (gc), and leaf water content. Decreases in relative leaf water content were much more linearly related to transpiration per unit saturated water content than either gc or A/Mw, separately. Only A/Mw appeared correlated with the rate of cuticular transpiration. Also, linear increases in gc occurred with increasing leaf water content. Estimated desiccation resistance during winter corresponded closely to the degree of drought tolerance characterized previously for each species based only on summer growth conditions. Thus, winter desiccation resistance may be an important, yet under-emphasized, ecological factor influencing conifer distribution patterns.

Beschreibung: The advance regeneration often present following clear-cutting in black spruce (Piceamariana (Mill.) B.S.P.) stands is generally composed of individuals of various heights. This initial height difference is hypothesized to affect the yield of the future stand. Height of the advance regeneration at time of release was determined on several black spruce trees within 33 stands distributed across the boreal forest of the province of Quebec, Canada. From these data, a regeneration structure index was developed and used to explain a part of the variation in subsequent stand yield. Dominant trees of stands developing for 38 to 65 years since clear-cutting generally originated from the tallest advance regeneration at time of release. Stands with the tallest regeneration at time of release produced the largest yields for a given site quality and number of years since harvesting. A 3-m increase in advance-regeneration height resulted in a similar yield gain as a 3-m increase in site index. Site indices calculated from age-height relationships of dominant trees originating from tall advance regeneration were less than those calculated from trees that have undergone a relatively short period of suppression. These results suggest that traditional management tools developed for fire-origin black spruce stands, i.e., stands mainly composed of seed-origin trees established after perturbation, should be adjusted for clearcut-origin stands that largely develop from advance regeneration.

Beschreibung: Nutrient movements in the senescing foliage of a Rhode Island Populustremuloides Michx. clone were measured during the years 1986–1988. Mean resorption of nitrogen, phosphorus, and copper was 43, 51, and 10%, respectively. Aluminum, boron, calcium, iron, magnesium, manganese, and zinc increased or remained unchanged in senescing foliage during 1986. Resorption of nitrogen and phosphorus decreased, respectively, from 56 and 64% in 1986 to 24 and 38% in 1988. Mean resorption differed among the 20 ramets studied. Older, larger ramets resorbed less nitrogen and copper than younger, smaller ramets. Timing of abscission strongly influenced resorption and may have been related to drought conditions in 1988 and to differential exposure to wind in all years. Resorption of nitrogen, phosphorus, and copper was lowest in those ramets that lost their leaves earliest and in leaves that senesced earliest on individual ramets. Because P. tremuloides ramets had the physiological potential to resorb more nitrogen and phosphorus than they actually did in 1988, we introduced the terms potential resorption and realized resorption to differentiate between physiological potential and ecological reality. Our data suggest that at least some portion of realized resorption is determined stochastically by environmental constraints.

Beschreibung: The ability of seedlings of Eucalyptustereticornis Smith to accumulate 137Cs and 90Sr, characteristic radioisotopes of nuclear fallout, from contaminated growth medium was tested. All seedlings were grown for 3 months in 165 cm3 of sphagnum peat moss–perlite (1:1 v/v) in a growth chamber before treatment with an isotope. After 1 month of exposure, seedlings had accumulated 31.0% of the 137Cs and 11.3% of the 90Sr originally present in the growth medium, with bioconcentration ratios of 54:1 for 137Cs and 13:1 for 90Sr. Accumulation of 137Cs and 90Sr in plant tissue was correlated curvilinearly with increasing time of exposure and with increasing concentration of radioisotope in the growth medium. Because seedlings of E. tereticornis accumulate these radioisotopes rapidly, they may be valuable in remediation of contaminated soils.

Beschreibung: This study characterized the height growth and crown form of 14-year-old balsam fir regenerated naturally under a light gradient varying from 3 to 83% of photosynthetic photon flux density (PPFD). After 14 years, total height growth was correlated with percent PPFD (R2 = 0.766; p 

Beschreibung: A linear mixed-effects model that accounts for the covariances among repeated measurements and for random plot effects is developed. A continuous-time autocorrelation error structure permits the model to be applied to irregularly spaced, unbalanced data. A strategy for fitting this model and diagnostic tools for assessing its goodness of fit are presented and demonstrated. The model is fitted to two permanent-plot data bases and shows marked improvement compared with models that do not account for the error structure.

Beschreibung: We studied the wood structure of eastern larch (Larixlaricina (Du Roi) K. Koch) affected by a larch sawfly (Pristiphoraerichsonii (Htg.)) infestation that occurred in 1939 in the Lac à l'Eau-Claire and Rivière-aux-Mélèzes areas, subarctic Quebec. The objective was to document variation in wood characteristics in relation to defoliation at three study sites. Lumen diameter of tracheids and cell wall thickness were measured from thin sections representing the 1931–1955 period. Significant changes in the wood structure of larch was noticed only at the Atkinson Island site, where the radial growth pattern was typical of the insect activity. A light ring was formed the 1st year of the infestation (1939). During the 2nd year (1940), the size of tracheids (lumen) decreased abruptly along with radial growth, while the proportion of latewood increased. The variation in wood anatomy among the three study sites paralleled that in radial growth. Wood structure appears to be a good indicator of the end of an infestation.

Beschreibung: A nonlinear model that yields logical predictions for tree crown ratio is presented. The model is based on data from permanent experimental plots located in even-aged Scots pine (Pinussylvestris L.) stands in southern and central Finland. Regressor variables in the model are stand dominant height, stand basal area, tree diameter, and tree height. The effect of thinning on tree crown ratio is modelled by incorporating a thinning response variable into the model. Thinning effect is dependent on thinning intensity and time elapsed from thinning, the latter of which is represented by the difference between current stand dominant height and dominant height at the time of thinning.