ALBERT

Description: The sedimentation regime off Northwest Africa is shaped by: (1) structur~al factors. which result in generallv low relief on land. shelf widths between 40 and more than 120 km. and av-erage sfope inclinations between 10 30' and 30; (2) land climates. which contral the delivery of terrigenous particles to the margin: (3) water movements including boundary currents and upwelling; and (4) the post- Pleistocene sea level rise. This chapter combines published and new results arising from research into the sedimentation processes off Northwest Africa. and emphasizes particularly the activities of the Kiel marine geological group during the past few years. Reviews of cruise activities and results were given in Closs et al. (1969) (Meteor cruise 8. 1967. off Morocco) . Seibold (1972) (Meteor cmise 25 . 1971. off Sahara to Central Senegal). Seibold and Hinz (1976) (Meteor cmise 39,1975 . and Valdivia cruise 10. 1975, from Morocco to South Senegal), and Waiden et al. (1974) (Meteor cmise 30, 1973, off Sierra Leone). Some of these cmises were used for pre- or post-site surveys for the Deep-Sea Drilling Project, or to add undisturbed Quaternary cores to the Glomar Challenger cores (leg 41, ] 975; Lancelot, et al .• 1978); leg 47 A, Arthur er al .• 1979; Lutze et al., 1979). We have concentrated our geological investigations on a number of standard profiles from the shelf to the upper continental rise as given in Figure 1. The manuscript was finished May 1979.

Description: https://www.researchgate.net/publication/230891291_The_Orbital_Theory_of_Pleistocene_Climate_Support_frim_a_Revised_Chronology_of_the_Marine_d18O_Record

Description: Summary Holocene sediments of the North Lagoon, Bermuda, were studied with shallow seismic reflection profiles (200 km CSP-survey, UNIBOOM-system) and vibration coring (40 sediment cores, pneumatic vibration corer, Meischner et al., 1981). Seismic Stratigraphy Four seismic sequences are distinguishable by seismic stratigraphy. All seismic sequences correspond to depositional sequences built up during high sea levels in interglacial times. The seismic sequences are separated by unconformities which are often strongly reflective and correspond to emersion planes during glacial phases. The upper sequence (sequence 4) is related to Holocene sediments. The pre-Holocene bedrock is divided into three different seismic sequences (Kuhn et al., 1981): Sequence 1: oldest Pleistocene sequence (pre-Sangamon sea-level highstands), upper boundary with levelled relief (lower boundary not discernible), composed of strongly cemented carbonate sediments, forms the bedrock below Three Hill Shoals Sequence 2: Sangamon (125 ky sea-level highstand), distinct surface morphology, forms the bedrock of a large area below Holocene sediments, Holocene reefs grew up on elevations of the sequence 2 surface, the Holocene reef rim was developed on an elevated rim of sequence 2 Sequence 3: youngest Pleistocene sequence (Sangamon, 105 and 85 ky sealevel highstands lower than recent), deposited mainly in depressions of the bedrock deeper than -15 m below recent Mean Sea Level, levelling the older relief, peat sedimentation in places The distribution of recent reef areas and lagoonal basins is strongly controlled by pre-Holocene topography and geology of the bedrock. During the Holocene approx. 1050 x 106 m3 of carbonate sediments were deposited in the North Lagoon (290 km2) and approx. 1350 x 106 m3 in the reef rim area (170 km2). Sedimentology There are no larger oscillations of the Holocene sea level identifiable in the sedimentological record. The pre-Holocene topography was gradually drowned during the Holocene sea-level rise. At first, the depositional depressions were separated and landlocked. Fresh water peat marshes, fresh water ponds, marine ponds and bays were formed. With rising sea level, the land barriers were more and more eroded, drowned and lost their influence on the back-barrier sedimentation area. Autochthonous and allochthonous peat, lime gyttja and carbonate mud are a typical transgressive back-barrier sediment sequence. After destruction of the barrier, the depositional milieu changed from restricted marine to normal marine, open lagoonal. Sea-grass sediments and nearly mud-free carbonate sand were deposited in shallow water in an exposed environment. Hydrodynamic energy decreases with increasing water depth in the lagoonal basin. A more densely growing reef rim and intralagoonal reef growth added to the protection of the deeper lagoonal floors. Fine-grained sediments were deposited in this environment. They are distributed over a large area of the North Lagoon and form the top of the transgressive lagoonal sediment sequence. Holocene reefs mainly developed on rises of the pre-Holocene surface. In the early Holocene, solid reef build-ups were able to keep up with the rapid rise of sea level. Sand pockets in the reefs were left behind and filled up mainly in the later Holocene. The percentage of fine-grained sediments, produced and resuspended in the reef rim and deposited in the near lagoonal back-reef zone, increased during the Holocene. Two models of Holocene sedimentation in a depression and on an elevation of the pre-Holocene surface illustrate the dependence of vertical facies gradation on pre-Holocene topography. Trends of the mostly polymodal grain-size distributions of the Holocene sediments are a coarsening-upward in the back-barrier and a fining-upward in the lagoonal sediment sequences. Change in the composition of the molluscan fauna in the Holocene sediments (particle size 〉 2000 µm) is an Indication for fades changes. Gastropods are abundant in the basal backbarrier sediments. Bivalves are rare and their diversity 1s low. Sea-grass sediments contain Codakia orbicularis and Astraea phoebia shells. In the sheltered lagoonal environment shell fragments 〉 2000 µm become rare, common species are Gouldia cerina, Pitar fulminata and Finella sp. (approx. 1000 µm). Fine-grained reef-rim derived sediments differ from lagoonal sediments by a higher percentage of Homotrema rubrum fragments and Alcyonaria spicules.

Description: Twelve species of terricolous microlichens from the Angmagssalik District, Southeast Greenland, are reported: Caloplaca friesii, C. livida, Lecanora boligera, Lecidea oligotropha and Leciophysma arctophila, which are new to the lichen flora of Greenland, Rinodina conradi, which is new to the eastcoast, and Baeomyces roseus, B. rufus, Buellia geophila, B. punctata, Caloplaca tornoensis and Mycoblastus tornoensis, new to Southeast Greenland. In a discussion of the greenlandic distribution, unpublished records from the herbarium of Copenhagen (C) are incorporated. Notes on the habitats are given and the pertinent phytosociological units indicated. Some morphological and anatomical characters are commented upon briefly.

Description: The 16 recognized species of Sorocea are listed with their synonyms and distribution. Two new taxa are described: S. steinbachii C.C. Berg and S. hirtella Mildbread ssp. oligotricha Akkermans & Berg. Three new combinations are made: S. muriculata Miquel ssp. uaupensis (Baillon) C.C. Berg, S. trophoides W. Burger ssp. rhodorachis (Cuatrecasas) C.C. Berg, and S. sprucei (Baillon) Macbride ssp. saxicola (Hassler) C.C. Berg. A key to the species is presented.

Description: From a bryological point of view Greenland is not well known in detail. Unlike Siberia or Arctic America, it is not known through a few large collections, but through multitudes of mostly relatively small gatherings. In this arctic island that is almost 2000 miles long and extends from below 60° N.lat. to nearly 84° N.lat., travel and logistics are difficult. Consequently no over-all study of the whole island has been made, although many collections have been achieved through casual or intensive studies of small areas. Like the other contributions to the knowledge of the bryoflora of Greenland this paper will only deal with a rather limited area; the Angmagssalik district on the East coast of Greenland, ranging from 65° N.lat. to 67°20' N.lat. In 1887 Lange and Jensen published the first and until now the only comprehensive review of specimens and publications on the Musci of Greenland. In their paper the first moss collection from the Angmagssalik area was reported, made in 1884-1885 in the course of ethnographical studies by Gustav Holm (Sphagnum girgensohnii and Polytrichum juniperinum), the first European to visit this part of East Greenland.

Description: The genus Stenandriopsis was created by S. Moore in Journ. of Bot. 44: 153. 1906 for a plant collected first by Vaughan Thompson and afterwards by Baron in an unspecified part of Madagascar. As the plate by which the description is accompanied depicts the specimen collected by Baron (n. 6708), the latter is to be regarded as the type. Stenandriopsis was referred by its author to the Justicieae, but this tribe is apparently accepted by him in the delimitation it received in BENTHAM and HOOKER’s “Genera Plantarum”, and as it is in this sense a most heterogeneous mixture, this does not greatly enlighten us. Of more importance is that Moore compares it with Crossandra Salisb. and Stenandrium Nees, i.e. with genera belonging to my subfamily Acanthoideae and referred by me respectively to the Acantheae and the Aphelandreae. However, in my paper on “The Acantheae of the Malesian Area. I. General Considerations” in Proc. Kon. Ned. Akad. v. Wetensch., Ser. c. 58: 166. 1955, I pointed out that it can not belong to the Acantheae as the corolla throat lacks the incision in the adaxial side which is characteristic for that tribe. It can not belong to the Aphelandreae either as the corolla limb is subactinomorphous instead of distinctly bilabiate. As I had to rely at that time entirely on Moore’s description and on the plate by which the latter is accompanied, I was unable to arrive at a conclusion, but I suggested that the genus might represent a new tribe of my Acanthoideae. Since then I have had the opportunity to inspect in the herbarium of the British Museum of Natural History the material on which the genus was based, for which I tender my best thanks to the Keeper, and now I am able to express a more definite opinion.

Description: Contrary to Europe, with only one Caltha species, North America has at least three species of this genus. These are the polymorphic C. palustris L., also widely distributed in Europe, the floating aquatic C. natans Pall, and the polymorphic C. leptosepala-biflora group. Two previous papers (Smit 1967, 1968) dealt with taxonomic aspects of C. palustris, that in North America were not essentially different from those of European material.

Description: The chromosome numbers of 16 species of Angiosperms, collected in Cameroun and the Ivory Coast, were determined. The numbers given for 14 species are new, in the remaining species the results of other authors could be confirmed.

Description: Neohattoria Kamim. is a monotypic genus of the Jubulaceae (= Frullaniaceae) with a single species, N. herzogii (Hatt.) Kamim., known from central to northern Japan and the southern part of the Kurile Islands. The present genus was segregated from Frullania by Kamimura (1961; sub. nom. Hattoria Kamim. nom. illeg., non Schust., 1961) on the basis of the branching type, the shape of the first leaf and underleaf on branch, the total lack of secondary pigmentation, the uniform cell structure of the stem in cross section, and the strongly toothed leaf lobes. The generic concept of Neohattoria was greatly expanded by Schuster (1970), who included eight species and classified them into two subgenera, subgen. Neohattoria (with a single species) and subgen. Microfrullania Schust. (with seven species); however, Hattori et al. (1972) transferred all species of subgen. Microfrullania to a newly segregated genus Schusterella Hatt. et al., thus retaining the monotypic status of Neohattoria. As already described and illustrated by Hattori (1955), Kamimura (1961), Mizutani (1961), Ladyzhenskaja (1963), Schuster (1970), and Hattori et al. (1972), Neohattoria herzogii is closely related to species of both Jubula and Frullania. Regarding the taxonomic desposition of Neohattoria, Mizutani (1961) and Mizutani & Hattori (1969) placed it with Jubula in a subfamily Jubuloideae of Lejeuneaceae and Hattori et al. placed it in Jubulaceae (s. lat.). But, Kamimura (1961), Schuster (1970, 1979), and Guercke (1978) placed it more close to Frullania, e.g. in a subfamily Frullanioideae of Jubulaceae (s. lat.); more recently, Asakawa et al. (1979b), admitting three distinct families, Jubulaceae, Frullaniaceae, and Lejeuneaceae, placed Neohattoria and Jubula in the Jubulaceae (s. str.) but Frullania and Schusterella in the Frullaniaceae.

Description: As an introduction to a number of researches of his own the author wishes to give the following data: „Veen” has two meanings in Dutch: 1. in a petrographic sense (peat) Von Büllow’s definition was accepted: „Torf” ist zu deflnieren als ein meist dunkles, kohlenstoffreiches und ± saures Gemenge unvollständig spezifisch-zersetzter Pflanzenteile, das erdgeschichtlich jüngste Glied der Verwantschaftsreihe der Kohlen, dessen Bildung noch heute andauert.” 2. in a plant-sociological-geographic sense (bog) the following definition has been suggested: a bog is a plot, the surface of which consists of a layer of peat, either covered or not with vegetation, with which that layer is genetically connected. The classification of bogs according to their position with regard to the water-level of the surroundings (Staring) and that of the geological chart were rejected on account of their ambiguous character. The classification suggested by Van Baren according to the environment in which the bogs have been formed, was likewise thought insufficient. Preference was given to the classification according to the plants which gave rise to the peat (eutrophic, mesotrophic and oligotrophic bogs) and according to the origin of the water needed for peat formation (topogenous, ombrogenous and soligenous bogs). The conditions of peat-formation are of a botanical (presence of a vegetation and micro-organisms), climatologic (presence of a certain temperature and moisture) and geological nature (presence of a basin, valley or dead river-branch, certain level of ground water, a possible impervious layer). With reference to a number of authors (Picardt; Van Lier; Grisebach, Venema and Staring; Weber) the alteration in conception as to peatformation from the 17th via the 18th and 19th to the 20th century has been given. The word „Peel” cannot be derived from „palus”. Nothing is certain about its origin. It may mean the low land, bog or marsh. The bogs of the Peel lie on the Brabant-Limburg border-plateau (fig. 2). Lorié and Pannekoek van Rheden have shown that the peatformation of the Peel is likely to have occurred in channels, which have been formed by the Meuse, in co-operation with wind and rain (fig. 4). The bogs were therefore in the first instance topogenous formations, which afterwards developed into ombrogenous bogs. For his own research the author collected peat in three ways: 1. by cutting lumps of peat from open profiles; 2. by boring with a simple peat-bore (photograph 1); 3. by boring with the Utrecht peat-bore, an improvement on Dachnowski’s (fig. 5). To assist in the pollen-analytic examination the samples were treated according to Erdtman’s method. The latter has the following advantages compared with the usual treatment with a 10% KOH-solution: 1. the surface-structures of the pollen-grains are more distinct and as a result the grains themselves can be recognized better; 2. the pollen is more concentrated, so that in spite of the method taking up much time, a saving of time is possible. How the method is applied may be found in the chapter concerned (p. 38 and following). For the stratigraphic examination the samples were broken apart in a glass-bowl of water and viewed with a binocular microscope. Dry sandy samples were broken in water, when seeds and other vegetative parts came floating to the top; next they were put with a brush on thick blotting paper and studied through the binocular microscope. The designations for the sediments and species of peat have been derived from Fægri & Gams. For Scheuchzeria peat a new designation has been added. A plea was made for replacing the word pollen-analysis by „palynology”. A survey of the observations and examinations up to abt. 1935 closes the introduction (see the diagrams of Weber, Erdtman and Duyfjes in the figs. 6, 7, 8 and 9). The author’s own research refers to the Southern and Astense Peel, as in the remaining grounds of the geological chart indicated I 4v (= raised bog) no samples could be taken owing to the digging off having progressed too far. 10 profiles were examined. The situation of the bore-sites has been given in the geological chart of the grounds (fig. 3). The result of the examination (figs. 10—27) and the discussion on it may be summerized as follows: Zoning of pollen-diagrams The sub-zoning of the late- and post-glacial periods according to Blytt & Sernander has proved useful as a zoning of pollen-diagrams, provided atlantic and sub-boreal are joined. It is desirable to replace Blytt & Sernander’s terminology by a different one, because the authors gave a climatologic connotation to their names of periods. The limit between pleistocene and holocene was drawn between preboreal and boreal as Florschütz did. As phases of the holocene the following names were suggested: young post-glacial = sub-atlantic mid post-glacial = sub-boreal and atlantic old post-glacial = boreal. Neither in the Peel nor elsewhere in Holland have Allerød-deposits been found. They are not likely to be found either, as on account of the long distance from the land-ice-margin the flora will have been hardly or not at all influenced by the Allerød interstadial period. For Holland therefore the zoning of the late-glacial according to Firbas (1935) may be considered sufficient. The names of the periods do not bear a climatologic connotation as those of the post-glacial phases do. For the sake of a unity the following names have been suggested: young late-glacial = pre-boreal mid late-glacial = sub-arctic period old late-glacial = arctic period. Forest-history In a table (p. 98), in which likewise the Peel diagrams of Weber, Erdtman and Duyfjes have been inserted, the examined profiles have been arranged from North to South. From each profile it has been stated whether it originated in a certain period (+) or not (—). The sub-arctic phase was characterized by forests of Betula and Pinus and was followed by the pre-boreal phase, in which Corylus and Alnus occurred. Also from the other Dutch diagrams (see list on p. 99) it appeared that in the Netherlands the Alnus pollen occurs with an equal frequency before, during and after that of the Quercetum mixtum. The old post-glacial zone of the diagrams shows a peak in the Pinusline. In contrast with the from Mid-Europe there is not always a maximum in the Corylus-curve after the Pinus-peak. In other Dutch diagrams this phenomenon is likewise found. Only in 28% of all Dutch profiles with a boreal zone does a hazel-maximum succeed a Pinus one. They often co-incide (16%), while in the remaining cases no hazelpeak has been established. There is no fixed order of sequence in the occurrence of the components of the Quercetum mixtum, either in the Peel or elsewhere in Holland. The mid post-glacial is the phase of culmination of warmth-loving forest elements: Alnus pollen shows the highest percentage in this zone. Quercus pollen also occurs in great quantities, while Ulmus and Tilia take up an important place up to the „Grenzhorizont”. The absolute and empiric Fagus pollen limits are found at different heights in the mid post-glacial zone of the diagrams, the rational limit lies somewhere near the „Grenzhorizont”. In the young post-glacial phase the Fagus pollen attains fairly high percentages (up to 30%). The maxima in the East and South-east of the Netherlands are between 20% and 38%; they decrease towards the coast and increase towards the South-east (Hautes Fagnes, Belgium) and East (Germany). It seems incorrect to class the Netherlands almost entirely among the oak-alderterritory poor in beeches, as Firbas did. An attempt has been made to fit the Peel-diagrams into Overbeck & Schneider’s zonation system. For the territory for which it has been made there are already difficulties (p. 104), for use in the Peel and other Dutch diagrams there are even more objections (p. 68, 104). Godwin’s zonation system appeared to be a little less forced, but not quite useful on account of too many details. From his horizons that of Ulmus proved useless for the continent. Neither for the Peel nor for the Netherlands and its surrounding territory can a detailed zonation system be designed. It has proved difficult to proceed any farther than Rudolph’s „Grundsukzession”: birch, pine-hazel-mixed oak-forest-beech, in which the alder generally joins the mixed oak-forest and the hornbeam the beech. Before drawing far-reaching conclusions from the course of the curves (as has been done by some authors) more palynological researches are needed in accordance with the actuality principle, known from geology. Pollen-grains from warmth-loving trees in seemingly sub-arctic spectra In profile 4 (Deurnse Peel II) pollen-grains of Abies, Alnus, Picea, Tilia, Ulmus and Corylus were found in the „late-glacial” zone (figs. 14, 15). Investigations were made as to which of the following possibilities would be the cause of their appearance: 1. in taking and preparing the samples pollution occurred; 2. pollen-transport over long distances has taken place; 3. the pollen-grains found have got secondarily into the deposit; 4. warmth-loving trees have occurred in favourable circumstances in the late-glacial phase or 5. in an interstadial period or in an interglacial phase. The said pollen-grains probably hail from a Würm interstadial or interglacial phase. Interglacial peat On the site of the bore-point 7 it was possible to collect samples from the layers under the peat. The upper 40 cm of the diagram Griendtsveen IX (fig. 27) of this profile proved a repetition of the lower 40 cm of the Griendtsveen I profile (fig. 18). The diagram shows that pollen of Carpinus, Picea and Abies occurs showing the deposit to be of interglacial age. The pollen-curves, however, pass unnoticed from an interglacial into a post-glacial portion. The limit is likely to be found between the two, about 30 cm below the mowing field. There is therefore a great stratigraphic hiatus. Pollen-analytically it could not be decided from which interglacial period the profile hails; on account of its situation on the middle terrace, it was deemed likely that it was an Eem sea deposit. The examined profile probably corresponds to Jessen & Milthers’ zone g; showing it to have been formed at the end of the Eem sea period. The Meuse therefore cannot have flowed through this part of the Astense Peel after the mid Eemean phase. Stratigraphy This is difficult to summarize. Compare various profiles. Individual mention may be made here of: 1. peat on a podsol layer; this was found in two places (Deurnse Peel I Kraaienhut and Griendtsveen VIII). Peat-formation may be thought to have occurred in the following way: heather started growing on drift-sand giving rise to a podsol layer. As the latter is impervious the vegetation surface became marshy. The heath was replaced by a Caricetum from which peat arose. Gradually more Eriophorum occurred, from which almost pure vaginatum peat arose. The bog-surface grew moister and moister, Sphagnum cuspidatum and Scheuchzeria could grow on it and formed a „Vorlaufstorf”. Only then could non-extremehydrophile Sphagna join in peat-formation. 2. the occurrence of Scheuchzeria-peat after the „Grenzhorizont” period. This species of peat, which is often found at the basis of the old Sphagnum-peat as a mesotrophic transition vegetation, has for the Netherlands only been found in the young post-glacial phase in the Peel (Deurnse Peel I Kraaienhut, Griendtsveen V and VIII and Nederweerd). At present the plant is very rare. The severe decline of this plant was also observed elsewhere. Probably it is caused by the gradual drying up or reclaiming of the raised bogs. Of the present station of Scheuchzeria near Ommen a short description has been given (p. 59 and photographs 2, 3, 4). 3. the „Grenzhorizont”. Where the young Sphagnum-peat has not been dug for the preparation of moss-litter, the Peel bogs show a clear „Grenzhorizont” (photograph 8). The conceptions about its origin have been discussed. The distinct separation between the old and the young Sphagnum-peat was not considered sufficiently explained. Though on the whole the „Grenzhorizont” is synchronous in the North-west European profiles, the point of transition from old to young Sphagnumpeat was fairly unstable and easily changeable as to time. Generally the date of the „Grenzhorizont” is fixed at about 500 A.D., though there are differences in opinion. There is a lack of archeological correlation which renders a correct dating impossible. Interference of man in the Peel Three ways of interference were stated: 1. peat has been dug off for the greater part in the territory of the Peel: young Sphagnum-peat for the preparation of moss-litter, old Sphagnum-peat for fuel. The trees which appeared when the bog was dug up in the „Veenderij der Maatschappij Griendtsveen” are sometimes in so good a condition, that they are used for building sheds. The 1 st, 2nd and 4th beam in the foreground of the shed in photo 5 has been sawn from a 30 m long subfossil pine. 2. in a native peat-digging it was possible to collect recent young Sphagnum-peat. 40 to 50 years ago the peasants living there had dug peat in holes, which were afterwards left to themselves. Sphagnum started growing again and the holes were filled in again. The diagram (fig. Griendtsveen VII) represents the surrounding heath with scattered pines and birches, sown by the wind, and a pine-plantation close by. 3. in the profiles Nieuwe Peel, Griendtsveen VI and VII it has been fixed by the indications given by Firbas, that only in the surface layers of the bog has corn-pollen occurred. So in these parts cultivation of cereals will be of recent date. This also appeared from the history of the reclamation of the said territory.

Description: The chromosome number of 157 species of Angiospermae occurring in the Netherlands is dealt with.

Description: It is commonly accepted that percentages of pollen in a pollen diagram do not express the exact composition of forests in earlier times. This inaccuracy is due to several factors, for instance the different quantities of pollen produced by plants, the distance of transport etc. A pollen diagram tells us only the change in pollen rain on the locality where we collected soil samples. In studying a pollen diagram we find a close relation between the variations in the percentages of a certain species and the area occupied by this species in the vegetation. When the percentage of pollen of a species increases, we conclude generally that the relative area occupied by this species in the vegetation increases too. However, such a connection might be doubted. The variety of factors controlling the dispersion of pollen is so great that the interpretation of a pollen diagram often meets with great difficulties. The connection between pollen rain and the composition of the vegetation is a simple one in the cases where we are dealing with a region of uniform vegetation. A diagram taken from a region in which the vegetation varies from place to place has to be regarded with some caution. Unfortunately such a heterogenity of the vegetation exists on the very place, where we want to compose a pollen diagram. The pollen rain which falls into a bog arises from two sources: a pollen rain from the local vegetation of the bog itself and one from the surrounding vegetation. When we are dealing with great bogs, the pollen produced by the vegetation of the bog itself will be mostly that of herbaceous plants, shrubs, and spores of the Bryophyta and the Pteridophyta. It is the rule rather than the exception that the bog will be treeless. The tree pollen in such a bog mostly takes its origin from the surrounding forests. It is a fortunate circumstance in a diagram that pollen of trees is separated from other pollen. However, one exception is seen in the way in which Iversen composes a diagram for late glacial times. This method, commonly used for late glacial times, embraces a pollen sum not only containing trees but also some herbaceous plants. The origin of the latter can, with some certainty, be accepted as from outside the bog. Therefore the local vegetation of the bog does not influence the percentages of tree pollen. The pollen sum thus comprises pollen of plants which grow under the same biotic conditions.

Description: Name: Wahlenbergia marginata (Thunb.) DC. Monogr. Camp. (1830) 143. Family: Campanulaceae.

Description: Alangium LAMK.—M. M. J. van Balgooy, Pac. Plant Areas 2: map 72. Complete; Old World, also incl. Indo-Malesia, E. Australia, Pacific (Solomons, New Caledonia, New Hebrides, Fiji); delineated except in Africa and Madagascar, localities indicated only in the Pacific, species density; monograph.

Description: In 1935 the present author reported the occurrence of this N. American species in the eastern part of Holland, province of Overijssel, in the vicinity of Almelo (JONKER, 1935). He found the species near the hamlet of Harbrinkhoek on a wet heath. The locality was also the only station of Wahlenbergia hederacea in the Netherlands, discovered a year before. Notwithstanding the extensive reclamations in that part of the country the species now still occurs in a number of localities around Almelo. The plants cannot be considered adventitious as they were found in places that are comparatively little influenced by human culture, judging from the occurrence, on the first-discovered locality, of e.g. Wahlenbergia hederacea. Gentiana pneumonanthe, Viola palustris, Radiola linoides, Linum catharticum, Scutellaria minor. The late Dr. Wachter discovered, in the herbarium of the Royal Botanical Society of the Netherlands, unidentified specimens of Hypericum canadense collected by Lako as early as 1909 in the same environment, perhaps even in the same station; and Dr. van Soest identified two specimens collected in 1918 by the late naturalist Bernink near Denekamp, about 20 km E of the above mentioned localities. Bouchard (1953, 1954, 1955) reported the discovery of the species in France, dept. Haute-Saône. The plants were found in large quantities, at the stony beach of oligotrophous lakes, together with Littorella uniflora. In his detailed publication of 1954 he discussed the possibilities of introduction. He concluded that the plants are not adventitious. They may be autochthonous or naturalized and then, when the latter is the fact, probably by U.S. army units that stayed in that area during world war I. He did not preclude, however, the possibility of a glacial relic. Bouchard overlooked the previous publication reporting the occurrence in Holland.

Description: The Upper Hunter palynological assemblage described includes 54 species, assigned to 29 genera. Seven genera (Scabratisporites, Guttatisporites, Lapposisporites, Pseudogravisporites, Paralundbladispora, Taeniaepollenites and Tubantiapollenites) and 39 species are new; three generic descriptions are emended (Apiculatasporites, Colpectopollis and Angustisulcites) and three new combinations are proposed. Bisaccate pollen grains average 84 %; within this group no Upper Permian elements could be demonstrated. The assemblage is compared with other European Lower and Middle Triassic assemblages; there are no close similarities to assemblages outside Europe.

Description: El género Plagiochila (hepatica) esta representada en las Islas Galapagos por ocho (8) especies diferentes: P. bursata (Desv.) Lindenbg., P. galapagona Inoue, P gradsteinii Inoue, P. guilleminiana Mont., P. inouei Grolle, P. scabrifolia Inoue, P. spinifera Ångstr. y P. subplana Lindenbg. El endemismo en este género es más alto que en otros géneros de las hepaticas, con cinco (5) especies que comienzan a conocerse solamente de los Galapagos ( P. galapagona, gradsteinii, scabrifolia, inouei, y spinifera). Las otras tres (3) son comunes y ampliamente distribuidas a lo largo de la America tropical. La mayoría de las especies estan restringidas a las zonas altas-húmedas de vegetación de las Islas Galapagos (matorrales de Zanthoxylum, Miconia y pampa) excepto P. guilleminiana muy común, la cual puede presentarse en la zona seca de transición de bosque. La más amplia variación de Plagiochila ha sido vista en Isabela (Cerro Azul), San Cristobal y Santa Cruz.

Description: The species Polypodium banaense C.Chr. is transferred to Crypsinus. The recognition of a genus Phymatopteris Pic. Ser. (= Phymatopsis J.Sm.) separate from Crypsinus is discussed.

Description: The six species of Curtia, including a hitherto undescribed species published here, as well as the monotypic genus Hockinia can be distinguished from each other by the seed coat structure. The anticlinal walls and the cuticle provide the most useful information. Curtia tenuifolia appears to be a complex species, but subsp. tenella can be readily separated from this complex by the seed coat structure. Heterostyly has been found in C. tenuifolia subsp. tenuifolia, C. obtusifolia, and Hockinia montana, but differences in seed coat structure can not be correlated with long-, short-, and equal-styled flowers. The differences in seed coat structure, the length of the seeds, and the number of cells per seed plead for maintaining Hockinia (closely related to Curtia) as a distinct genus. One new species of Curtia is described and a new combination is made.

Description: SETTEN, A. K. van & KOEK-NOORMAN, J.: Studies in Annonaceae. VI. A leafanatomical survey of genera of Annonaceae in the Neotropics. — Bot. Jahrb. Syst. 108: 17—50. 1986. — ISSN 0006-8152. Within the scope of the multidisciplinary research project on systematics of Annonaceae, a survey of the leafanatomical features and their distribution in the neotropical Annonaceae is presented. The studied specimens form a rather homogeneous group, as may appear from the family description given here. A detailed study of the leafanatomical features reveals, that differences are mainly found in the indument, the position and contents of the idioblasts, the structure of the primary vein, the type of crystals in the epidermal cells, and the type of sclereids. Based on character states, phenetic similarities and differences are discussed and compared with the classifications of FRIES (1959) and WALKER (1971).

Description: One new species of Dorstenia from Brazil is described: D. carautae C.C. Berg, and four new combinations are made: D. cayapia Vellozo subsp. asaroides (Hooker) C.C. Berg, D. cayapia Vellozo subsp. paraguariensis (Hassler) C.C. Berg, D. cayapia Vellozo subsp. vitifolia (Gardner) C.C. Berg, and D. ramosa (Desvaux) Carauta, Valente & Sucre subsp. dolichocaula (Pilger) C.C. Berg. A list of and a key to the 22 Dorstenia species distinguished in south-eastern tropical America are presented, together with synonymy and distributional data.

Description: In a previous palynological study in Cocoid palms the present authors (Punt and Wessels Boer, 1966) were able to demonstrate a rather strong but not absolute correlation between the pollen types and the staminate flower types within the genus Attalea in the broad sense. The existence of partly apparently primitive, partly very advanced flower types within the otherwise close related group of Geonomoid palms made it worthwhile to investigate the same feature in this group. The Geonomoid palms are usually considered to comprise 7-9 genera of monoecious Arecoid palms which share a large number of characteristics (Burret, 1930; Moore, 1966). The group is very obviously a most natural one. The generic distinctions are mainly based on the flower morphology, notably on differences found in the androeceum and pistil.

Description: Recently a multidisciplinary investigation program on the systematics of Annonaceae was started at Utrecht with special emphasis on the Neotropics. This project will be carried out largely within the framework of the UNESCO-project Flora Neotronica. The first goal is to provide a modern classification of the family as a whole, the second is the publication of a series of monographs for Flora Neotropica. The project has been planned and started in close consultation with leading botanists on the Neotropical flora. The Annonaceae are a family of pantropical distribution with between 2000 and 2500 species in ca. 130 genera as presently understood. In the Neotropics the family is represented by ca. 750 species and 35 genera. It is a family of trees, shrubs, and lianas. Its place is within the order of the Magnoliales and its supposedly closest relative is the family of the Myristicaceae. The Annonaceae, although generally considered primitive in many features, nevertheless offer a number of specialized features as well This makes it a promising object using various kinds of comparative morphological, karyological, and anatomical data. Besides, many species are of medicinal or commercial value, such as various species of Annona and Rollinia, the fruits of which are commonly eaten in most countries of Central America and South America; the Soursop (Annona muricata) is widely cultivated throughout the tropics.

Description: Vochysia sectio Ciliantha Stafleu, subsectio Ferrugineae Warming. A V. vismiifolia Spruce ex Warming stipulis incrassatis, foliis lanceolatis longe acuminatis, floribus calcari longo modice incurvo, petalo intermedio stamen aequante, stigmate terminali parvo instructis differt. Holotypus: “coll. unknown” (comm. D. Allen) in U, fl. 14 Nov. 1953. PERU, Nanay River near Iquitos, altitude 100 m., “quillo sisa”, tree more than 100 feet high, on clayey soil about 20 feet above river (Isotypes: US 2104976, Y 47782).

Description: Recently I got the opportunity of examining a specimen from the “Rijksherbarium”, Leiden, which was provided with a label on which ROTH had written in the middle the name of the plant, viz. “ Micranthus serpyllifol-Roth ” and in the lower right corner the name of the collector, viz. “Heyne”; in the lower left comer another hand had added “Ind. or. Hb. Roth”. As the specimen proved to answer the description of Micranthus serpyllifolius given on p. 282 of ROTH’s “Novae Plantarum Species, Halberstadt 1821,” there can be little doubt that it is either the type of this species or else a duplicate of the latter. This is the more important as none of the authors who in the past ventured an opinion with regard to the taxonomic position of ROTH’s species, apparently had seen the type. ROTH’s specimen was inserted in the Leiden Herbarium under the name Andrographis serpyllifolia R.W. (Acanthaceae), but this is obviously a misidentification. for Andrographis serpyllifolia does not fit ROTH’s description. The plant described by the latter has smaller and less numerous leaves and its flowers are arranged in terminal spikes instead of solitary or a few together in the axils of ordinary leaves.

Description: The chromosome numbers of 31 species of Angiospermae collected in S. Brazil were determined. Of these species 5 were studied before, the other numbers are new, 11 are first counts for genera and one even for a family. Some notes on the cytology and morphology are added.

Description: A new species of Asterophorum, A. mennegae, is described from the Sipaliwini Savanne (Suriname). The position of Asterophorum within the family is briefly reviewed, and a key to the 2 species is given.

Description: During my studies of the Surinam specimens belonging to this genus my attention was drawn to the often wrong interpretation of several old species. To avoid future misidentifications it seems useful to give a short review of the American species that are known up till now. It is emphasized, however, that this paper does not have the pretension to be a monograph of the American species. For the greater part my study of the species was confined to the type material and the variability therefore is not known. However, this contribution may serve as a base for a future monograph of this interesting group. Attention is drawn to the fact that only older leaves of the plants should be studied, because the leaf apex of the younger leaves is in all species acute and the lamina may not have reached its definite form.

Description: The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.

Description: In this preliminary investigation attention was paid to pollen morphology of West-European species of the Rosaceae. Some new terms were used like fastigium, endocingulus etc. The terminology of Iversen and Troels-Smith has been followed in addition to improvements by Erdtman. A key is given to the types and subtypes for the use of pollen analytical investigators. Sanguisorba officinalis appeared to be always 3-colporate and not 6-colporate.

Description: Nanocyperion communities (s.l.) are considered here as “warp-and-woof” communities; the Nanocyperion components are described separately as synusiae. On the Netherlands Frisian Islands, four main synusiae have been recognized. Raunkiaer life form spectra show few differences between the communities. Life strategy spectra of the Nanocyperion synusiae, based on systems for phanerogams (modified after Bakker 1966) and bryophytes, yield the clearest patterns. A comparison of the ecology of the communities and an interpretation of the spectra in terms of avoidance of stress or competition suggest that inundations and standing crop of the communities are the main factors determining the distribution of the synusiae. Winter inundations overrule the influence of differences in productivity level, which becomes prominent in drier situations.

Description: A palynological assemblage obtained from marly limestones of the Dutch Lower Muschelkalk is discussed. A qualitative analysis has disclosed its great resemblance to Upper Bunter (Röt) assemblages ; however, differences in quantitative composition were demonstrated. Utilizing palynological data new information can be added to the knowledge of the European Middle Triassic flora.

Description: The water economy, the mineral content of the soil, and human influence are the principal ecological factors governing the variation of the heath vegetation of a limited region. Sloping of the surface is also an important factor. In hilly country it is of a twofold nature: on the one hand the difference between high and low altitudes, based on the water economy, on the other hand differences in (micro-) climate. If the hills are higher, this results in greater climatic differences. In extremely oceanic and in boreal regions a rise in altitude of 100 m is sufficient for creating a noticeable decrease in temperature and an increase in precipitation, aerial moisture, and wind force. This results in the occurrence on the hills of heath communities that have their main distribution more to the North. The same observation was made by Gimingham (1961). On Slieve League on the Donegal coast (Ireland) Salix herbacea and Lycopodium selago occur in the heath at an altitude of 600 m, near Tongue on the Scottish north coast Dryas octopetala, Saxifraga oppositifolia, Alchemilla alpina and Thalictrum alpinum at an altitude of 60 m. West of Apeldoorn in the Netherlands are found extensive stretches of heath with abundant Vaccinium myrtillus and V. vitis-idaea at an elevation of 60-80 m, even on south-facing slopes. This is an area with high precipitation due to ascending air west of the hill ridge of the Eastern-Veluwe. Here the Vacciniums, elsewhere requiring the protection of the forest, can tolerate the habitat of the open heath (Stoutjesdijk, 1959; De Smidt, 1966). Higher elevation combined with north-facing slopes creates extreme conditions e.g. on Roc Trévézel (300—360 m) in Brittany, with Vaccinium myrtillus, Melampyrum pratense, Hymenophyllum wilsonii and Rhytidiadelphus loreus. These species are virtually lacking in the surrounding plains where the heath consists of such South Atlantic species as Erica cinerea, E. ciliaris, Ulex gallii, Lobelia urens, Lithospermum prostratum and Symethis planifolia.

Description: Frutex epiphyticus, ramis quadrangularibus, angulis lenticellatis. Folia ramorum fertilium petiolis 2-5 mm longis, 1.5-2 mm latis; lamina chartacea vel subcoriacea, oblanceolato-oblonga vel interdum elliptico-oblonga, 9-14 cm longa, 3-4.5 cm lata, ápice acuminata, acumen 1-2 cm longum, basi attenuata, costa subtus prominente, nervis lateralibus supra et subtus prominentibus vel prominulis, glandulis hypophyllis aliquot patelliformibus vel foveolatis, parvis, 3-5 in folii parte inferiore oblique seriatis, aliis minutis, punctiformibus, nigricantibus, aequaliter dispersis. Flores in racemis umbelliformibus (20-) 30-45-floris; rhachis ad circa 1 cm longa; nectaria clavato-cucullata, stipitata, stipes 5-8 mm longus, cucullus 1-1.5 cm longus, circa 4-5 mm diametro, ore late-rotundata, margine plerumque recurvo, apiculata; pedicelli 5-7 cm longi, lenticellati; bracteolae sepaloideae, circa 1 mm longae, 2-3 mm latae; sepala suborbicularia vel reniformia, circa 2-3 mm longa, circa 4-5 mm lata, margine glandulosa; corolla oblongo-subconoidea, circa 1 cm longa, circa 4-5 mm diametro; stamina 18-33, filamentis applanatis, liberis, inaequalibus, in alabastro 4-6 mm longis, antheris linearibus, 3-5 mm longis, circa 1 mm latis, basi subsagittatis; ovarium circa 2-3 mm diametro, 6-11-loculare. Fructus globosus, circa 9 mm diametro, stylo persistenti ornatus. Typus : Costa Rica, vicinity of Vara Blanca, North slope of Central Cordillera, between Poás and Barba Volcanoes, alt. 1700 m, April 1938, Skutch 3762 (holotype US; isotypes GH, MO, NY, S). Paratypes; Costa Rica: Heredia: Cerro de las Caricias, North of San Isidro. Standley & Valerio 52202, 52248,52375 (US); Panama:Chiriquí: Boquetedistrict, Bajo Chorro, Davidson 398 (GH, US).

Description: Although algology deals with a large group of plants, widespread and of a great morphological diversity, the history of this branch of botany is fairly young. Linnaeus (1753) listed in his Species Plantarum under the heading “Cryptogamia—Algae” only five genera of plants which are still accepted as algae at the present time. Under the same heading he also described a number of liverworts, lichens and sponges and a few other things.

Description: Joseph Gaertner (1732-1791) was the first to develop a carpological taxonomy in his book De fructibus et seminibus plantarum (1788-1791). The scope and background of this work are discussed; its history is sketched on the basis of the Banks correspondence at the British Museum; the main sources of material are listed. A brief outline of Gaertner’s life is given, also mainly based on letters from him and his contemporaries to Joseph Banks.

Description: This vegetation survey is the outcome of an investigation of the islands of the Netherlands Antilles carried out under the auspices of the Foundation for Scientific Research in Surinam and the Netherlands Antilles. The data on which the present study is based were obtained during a trip which lasted from September 1952 until October 1953. During this trip the following islands were visited: Curaςao, Bonaire, Aruba, St. Martin, Saba, and St. Eustatius. A short visit was also paid to the island of St. Kitts (B.W.I.). The present work gives an account of the actual vegetation of the Netherlands Antilles. Other studies, comprising the systematic results and conclusions of the survey, are being prepared, and will possibly be published in 1958.

Description: Previously, the chromosome numbers of some species of Loganiaceae were dealt with (Gadella, 1961, 1962, 1963). The chromosome numbers of 7 species are reported in this paper, of which 4 species, all belonging to the genus Strychnos, had not been investigated cytologically before. The materials, kindly supplied to me by Dr. A. J. M. Leeuwenberg and by Ir. F. Breteler, were collected in the form of seed-samples in the Ivory Coast and in Cameroun. The plants of 2 species originate from botanical gardens. Living material of all species (except for Strychnos lernata Gilg. ex Lwb.) is grown in the botanical garden of Wageningen (WAG). The determination of the chromosome numbers was based on the study of roottipmitoses. Roottips of the plants were fixed in Karpechenko, embedded in paraffin, sectioned at 15 micron and stained according to Heidenhain’s haematoxylin method. The results may be summarized as follows: 1. Nuxia floribunda Benth.: 2n = 38 Origin of the material: S. Africa, obtained from the University of Stellenbosch. Seeds probably collected in the wild. Herbarium material of the mother-plant: J. J. Bos, no. 310 (WAG). Herbarium material of seedling: A. J. M. Leeuwenberg no. 3665 (WAG). References: Gadella (1963): 2n = 38. 2. Strychnos dinklagei Gilg.: 2n = 44 Origin of the material: Ivory Coast, Forêt d’Abouabou, between Abidjan and Grand Bassam. Herbarium material of the mother-plant: R. A. A. Oldeman no. 845 (WAG). Herbarium material of the seedling: A. J. M. Leeuwenberg no. 3561 (WAG). References: Gadella (1963); 2n = 44. 3. Strychnos innocua Del. subsp. innocua: 2n = 44 Origin of the material: Ivory Coast, 16 km S. of Ferkéssédougou. Herbarium material of the mother-plant: A. J. M. Leeuwenberg no. 4435 (ABI, WAG). 4. Strychnos millepunctata Leeuwenberg: 2n= 441) Origin of the material: Ivory Coast, Forêt d’Abouabou, between Abidjan and Grand Bassam. Herbarium material of the mother-plant: J. J. F. E. de Wilde & A. J. M. Leeuwenberg no. 3447 (ABI, WAG). 5. Strychnos samba Duvign.: 2n = 44 Origin of the material: Cameroun, 4 km S. of Nguélémendouka. Herbarium material of the mother-plant: F. J. Breteler no. 2051 (WAG). 6. Strychnos spinosa Lam.: 2n = 44 Origin of the material: obtained from the botanical garden of Groningen, the Netherlands, origin unknown. Herbarium material of cutting: A. J. M. Leeuwenberg no. 3564 (WAG). References: Mangenot and Mangenot (1958): 2n = 44. Miège (1960): 2n = 44. Gadella (1962): 2n = 44. 7. Strychnos ternata Gilg. ex Leeuwenberg: 2n = 44 1) Origin of the material: Cameroun, 27 km from Bertoua on road to Bétaré Oya. Herbarium material of the mother-plant: F. J. Breteler no. 2196 (WAG). Herbarium material of the seedling: F. J. Breteler no. 2994 (WAG). Twenty-one species of the genus Strychnos have been studied up to the present. The following chromosome numbers have been counted: 2n = 24 (3 species, counted by Mohrbutter, 1936); 2n = 44 (16 species); 2n = 88 (2 species). From these data the conclusion may be drawn that the more common basic number of the genus Strychnos is X = 11 or 1 = 22.

Description: In my “Notes on the Acanthaceae of Java” (in Verh. Kon. Ned. Akad. v. Wetensch., Afd. Natuurk. 2nd Sect. 45, 2: 29,1948) I discussed the three epithets that had been applied to Rumph’s “Folium tinctorum” after the latter had been transferred to the genus Peristrophe, which, as is well known, was based on this species. Nees, the author of the genus, has used the name P. tinctoria, because he regarded Justicia tinctoria Roxb. as the oldest binomial that had been applied to it. This was contested both by Merrill and by Hochreutiner. Merrill was of opinion that Justicia bivalvis L (1759) was its oldest name, but as I pointed out l.c. this binomial must be regarded as a “nomen confusum”; the description indicates a Dicliptera species, whereas the plate in the “Hortus Malabaricus” and the specimina in Burman’s herbarium to which Linné referred, represent respectively Adhatoda vasica Nees and indeed “Folium tinctorum”. Hochreutiner, on the other hand, thought, that Justicia purpurea L (1753) was identical with Rumph’s plant, but this too proved to be a mistake. J. purpurea belongs, as R. Brown already had recognized, to Hypoëstes. As the binomials proposed by Merrill and Hochreutiner therefore had to be rejected, I accepted l.c. Peristrophe tinctoria (Roxb.) Nees as the correct name. This, however, is also erroneous, for Justicia tinctoria Roxb. itself is an illegitimate name, for which already long ago a legitimate one had been substituted. J. tinctoria Roxb. (1820) is a later homonym of J. tinctoria Lour. (1790). This was recognized already by Schultes (Mantissa 1: 140, 1822), who replaced Roxburgh’s epithet by roxburghiana quoting “ J. tinctoria Roxb., Fl. Ind. ed. Car. et Wall. I p. 124, n. 13 et hoc teste: Folium tinctorum Rumph. Amb. VI 51. t. XXII. f.l” adding “nomen mutandum erat ob tinctoriam antiquissimam Lour”. As Loureiro expressly stated that the plant described by him as J. tinctoria was not the same as “Folium tinctorum” of Rumph, it is clear that J. roxburghiana Schult. must be accepted as the oldest legitimate binomial for the latter. The correct name therefore becomes Peristrophe roxburghiana (Schult.) Brem. n. comb.

Description: A suitable subtitle for this paper would have been “The rise and fall of a family”. What is usually called the Cyphellaceae is an instructive example of a situation not uncommonly encountered in the current systematics of mycology: a family retained in a traditional sense by some mycologists and considered by them as good a family as any, while others are convinced that it is nothing but a handy bin from which part of the contents has already been taken out and disposed of by scattering it over various groups, but which is still needed for keeping what remains. We do not yet know what to do with this considerable remainder, mainly because the published accounts are inadequate and the species have not yet been scrutinized anew in the light of present-day taxonomic requirements. In order to understand the basic idea of the Cyphellaceae the type species may be briefly introduced. The fact that Cyphella digitalis was originally described as Peziza digitalis is telling, and one could not do better than characterize it as a ‘discomycete’ with basidia, viz. a cup-shaped fruit-body with the hymenium lining the smooth inside or ‘disk’. If one were pressed to form an opinion about its taxonomic position from a dried, not annotated collection and without the aid of the microscope, one would even now, very likely, dispose of it as a discomycete. However, there is little doubt that in nature the cup is directed downward at least when mature, in contradistinction to the average discomycete in which the hymenium containing the asci is directed upward. This difference is a reflection of the two modes of violent spore discharge inherent in the hymenomycetous basidium and ascus; it has been explained through Buller’s well-known researches. The cups in the various species are not always typically cup-shaped; in a number they are more or less tubular or else more flattened and even disk-like.

Description: La systématique scientifique, qui classe tous les êtres vivants, actuels et fossiles, d’après leurs affinités naturelles ou présumées telles, doit pouvoir donner à chaque taxon un nom stable, permettant de le désigner avec son rang taxonomique et de l’identifier facilement. La nomenclature botanique n’échappe pas à cette exigence et depuis que C. Linné généralisa, en 1753, la nomenclature binominale, le besoin s’est fait sentir de compléter et de préciser, sur le plan international, les principes et les règles établis par l’illustre botaniste suédois.

Description: From the general discussion of the effect of mass selection on the genotypic array of the next generation we have seen that under certain conditions mass selection can lead to homozygosity, but does not do so necessarily. Outcrossing, mutations and disadvantages of certain genotypes may prevent reaching equilibrium condition with complete homozygosity even in the simplest case of one locus with two alleles. It depends on the magnitude of these factors and the degree of heterozygosity of the variety how close to genetic uniformity we ultimately can come. The advancing of one generation has only a small effect, which becomes less as the frequency of one of the genotypes becomes less. Mass selection can reduce segregation in a variety, but only in ideal situations and only in small steps. Obviously the most effective way to promote genetic uniformity is to begin with non-segregating material obtained through careful inbreeding and within-family selection. Then, if this is available, mass selection is hardly necessary and seed collection should be done so as to prevent a return to a heterozygous condition. When only segregating populations are available, some changes can be expected if the selection intensity is low, but they will not be great and may not be noticed until after several generations. Returning to the original question of the stability of the tobacco variety, the conclusion can be drawn that when large numbers of plants from a field are selected as seedplants the changes in the next generation as a whole will be small, regardless of the selection procedure used. When 10-30 % of all plants are allowed to produce seed we cannot expect important changes. Also, when the environmental variations are as great as in shade tobacco, the possibilities of selecting against certain genotypes for a number of characteristics simultaneously becomes virtually impossible. Concerning whether or not mass selection can lead to improvements in the variety, it was explained that the selection intensity and the heritability of a character determine the selection response. Perhaps this response can be predicted in the case of one character, but it becomes difficult, if not impossible, to forecast the results when many characters are involved at the same time. Reference was also made to a selection index which has been effective in livestock breeding. Even if an index could be determined for shade tobacco, its use is not necessarily effective, as Kempthorne (1957) pointed out. The genotype-environment interaction makes questionable the choice of certain fields over others. Without experiments, such questions cannot be answered. It is likely that mass selection with low selection pressure changes a variety very little. Again, in a field where up to 30 % of the plants are selected for seed, the pressure cannot be very great and we should not expect great changes. Only when a few plants are carefully selected for certain characters should progress become noticeable, as is the experience of single plant selection in plant breeding. We conclude that mass selection for seed and plant breeding to improve a variety should not be confused with each other. Each has its own aims and methods, which are not interchangeable. Where uncertainty exists about seed, progeny tests usually are made to compare a seed with the parent seed. Bolsunov (1959) has described a number of such procedures for tobacco, though the more elaborate of them appear impractical.

Description: Eén van de grote giften van A. A. Pulle, Lanjouw’s voorganger als Hoogleraar in de Bijzondere Plantkunde en de Plantengeographie te Utrecht, was zijn goede keus van medewerkers. Hij kon die keus maken dank zij een andere gift, die van leermeester. Pulle’s stimulerende activiteit als hoogleraar luidde de wedergeboorte in van de Nederlandse plantensystematiek door een stroom van leerlingen waarvan er velen ook nu nog een belangrijke rol spelen in de botanische wereld. Vele van Pulle’s leerlingen uit de eerste tijd moesten Utrecht verlaten omdat er in de twintiger jaren nu eenmaal nog geen sprake was van een redelijk gesalariëerde wetenschappelijke staf. Eén leerling echter werd zo gefascineerd door de vele mogelijkheden die het Utrechtse instituut bood, dat hij bleef, niettegenstaande de soms zeer ongunstige economische en organisatorische omstandigheden. Deze leerling, Lanjouw, toonde reeds toen de taaie volhardendheid die zijn medewerkers en leerlingen in later jaren zo goed leerden kennen en waarderen. Reeds van de eerste jaren van zijn assistentschap aan stonden Lanjouw de idealen voor ogen die hij in de loop van zijn lange loopbaan van assistent tot directeur voor het grootste gedeelte zou weten te realiseren. Deze loopbaan begon toen hij op 1 januari 1926 benoemd werd tot assistent bij de Bijzondere Plantkunde. In het jaar waarin we het feit herdenken dat het veertig jaar geleden is dat Lanjouw verbonden werd aan het Botanisch Museum valt ook de honderdenvijftigste verjaardag van het herbarium. Plantensystematiek is te Utrecht in meerdere of mindere mate beoefend van de stichting van de Universiteit af. Op 30 oktober 1816 echter werd het eerste herbarium voor de Universiteit verworven. In 1966 herdenken we dus eigenlijk twee jubilea. De periode van werkelijk actief systematisch onderzoek is echter veel korter geweest: de korte jaren van Miquel’s werkzaamheid (1859-1871) en de periode die in 1906 begon met de aanstelling van Pulle tot Lector in de plantensystematiek. Pulle gaf, voor het eerst na Miquel, en mede dank zij de stimulerende invloed van Went, een nieuwe stoot aan het plantensystematisch onderzoek in Nederland en wel vooral aan het onderzoek van de tropische flora’s. Een reeks van leerlingen wijdde zich onder zijn leiding aan de studie van de flora’s van Suriname en Nederlands Indië. Het is voldoende hierbij de namen van van Sloten, Lam, van Steenis, en Uittien te noemen om deze ontwikkeling te illustreren. Lanjouw koos voor zijn proefschrift de Surinaamse Euphorbiaceae en het was uit dit werk dat de „Flora of Suriname” voortkwam. Zonder ook maar iets tekort te doen aan de rol van Pulle kan gezegd worden dat van het begin van zijn loopbaan af Lanjouw actief heeft medegewerkt aan de opbouw van het Botanisch Museum en dat hij bij de ontwikkeling van de zo belangrijke „Flora of Suriname” een doorslaggevende rol heeft gespeeld. Zijn dissertatie „The Euphorbiaceae of Suriname” vormde, ten dele, tegelijk de eerste aflevering van de Flora.

Description: The somewhat peculiar genus Vahlia Thunb. (1782) was formerly regarded as an anomalous member of the Saxifragaceae, but in 1959 it was segregated as the monotypic family Vahliaceae by Dandy (in Hutchinson, Fam. Fl. Pl. ed. 2, 1: 461). The genus includes about three species, all of them highly variable in stature, flower size, and indumentum; this has resulted in the description of more than twenty “species”, whose names now appear in synonymy. This note, however, is chiefly concerned with the generic name, since it appears that Vahlia Thunb. is not correct under the Code and must be replaced by Bistella Adans. (1763). In the synonymy under the new combinations, all the relevant names so far traced are included, but it may well be that an experimental approach to the problem of specific and infraspecific limits and the taxonomic value of the characters hitherto relied upon for differentiation would result in considerable changes. Bistella Adans. (1763) was cited in the form “ Bistella Lippi 243 – Ascyroides Lippi” and the name than fell into abeyance until it was resuscitated by Delile in 1826. He described plants collected by Cailliaud, and identified a number of them with those of Lippi. In particular, he identified Lippi 243 (and 244) with the Cailliaud plant which he named “ Bistella geminiflora Delil. (Descript, des plantes découv. par M. Cailliaud, pl. II. fig. 4). – Ascyroides Lippi (Manusc. nos. 243 et 244).” It must be emphasized that the taxonomic identification of “ Bistella Del.” with Vahlia Thunb. has never been questioned, but the identity of Adanson’s Bistella with Bistella geminiflora Del. has been overlooked; this species, as represented by Lippi’s plant, is the type of the generic name Bistella Adans.

Description: A survey is given of the types of the sclereids and the sclereid patterns occurring in the leaves of the Marcgraviaceae. Eight main categories of sclereids are distinguished on the base of the morphology of the sclereids. A comparison is made with the foliar sclereids found in some other families. The systematic value and the function of the sclereids are briefly discussed. Some new combinations of names are published.

Description: Among the endemic and sometimes localized genera of the Cruciferae occurring in South America, Mathewsia stands apart as an element of the distinctive desert flora of southwestern Peru and western Chile. As far as present records show, the genus is confined to a relatively narrow strip wholly west of the main Cordillera, not far from the Pacific Ocean. Some species are confined to washes or small arroyos known as quebrades. Others occur as part of the lomas vegetation of western Peru. Populations of the same species occupying different quebradas often differ from each other to a limited degree and in many instances probably little or no gene exchange occurs between them. The intervening dry areas, separating one quebrada from another, are completely unsuited to the growth of Mathewsia and form a natural barrier to the spread of any given population. Thus, with habitats only spottily available, the evolutionary divergence that has resulted in localized species of Mathewsia is readily understandable. One of the real difficulties in studying a group of species inhabiting areas that have been infrequently visited by botanists, is the paucity of available material. Furthermore, unlike some species of Cremolobus (Khanna and Rollins, 1965), the numbers of individuals in a given locality appear to be few and scattered. In some instances, only a single specimen of a given species has been found for study in all of the herbaria consulted. Under these circumstances, the bare essentials of the species are all that can be given and nothing can be said about their variation or distribution. The purpose of this paper is to present, as a first approximation, a taxonomic treatment of Mathewsia that will provide a sound basis for further research on the genus.

Description: A subdivision of pollen types based only on different dimensions is very dubious. An example is given, taken from the miocene browncoal in the Lower-Rhine area of Germany and the Netherlands.

Description: The Veluwe is a stretch of high ground in the central part of the Netherlands, north of the river Rhine and south of the IJssel Meer, i.e. the former Zuiderzee, and the polders reclaimed from the latter. Geologically the area consists of three formations: 1. ridges which owe their origin to the pressure of the land ise, and which consist of sands deposited as river sediments in preglacial times; 2. a fluvioglacial formation; on some of these plains small but steep hills are found; 3. aeolian sediments: löss and cover-sands (cf. VINK, 1949); they were deposited in the late-glacial period.

Description: Eight species of the genus Sanguisorba L. were studied. Two distinct types could be recognized, viz., the Sanguisorba minor and the Sanguisorba officinalis type. Sanguisorba filiformis (Hooker fil.) Handell-Mazzetti appeared to be a transition between these two types.

Description: The Colombian representatives of the lichen family Parmeliaceae with linear lobes and marginal cilia have been revised. A key is given and morphology, chemistry and distribution are treated of 12 species in three genera: Cetrariastrum Sipm. gen. nov, with C. andense (Kärnef.) Sipm. comb. nov., C. dubitans Sipm. spec. nov. and C. equadoriense (Sant.) Sipm. comb. nov., Everniastrum with E. catawbiense (Degel.) Hale, E. cirrhatum (Fr.) Hale, E. columbiense (Zahlbr.) Hale, E. fragile Sipm. spec. nov., E. planum Sipm. spec. nov., E. sorocheilum (Vain.) Hale and E. vexans (Zahlbr.) Hale, and Parmelina cleefii Sipm. spec. nov. and P. swinscowii (Hale) Hale.

Description: A phytosociological survey based on methods of the Zürich-Montpellier School was carried out in the páramo vegetation of the Cordillera Oriental, Colombia. The study area covers about 10,000 and comprises the páramo between the Nevado de Sumapaz (3°55'N, 4250 m), the Sierra Nevada del Cocuy (6°25'N, 5493 m) and the Páramo del Almorzadero (7°N, 4375 m). The páramo vegetation was studied along various altitudinal transects from the upper forest line (3000-3500 m) up to the lower limit of the snowcap (4800 m). A general description of the study area includes data on geology, geomorphology, soils, climate, flora, phytogeography, morphological characters of the vegetation, fauna and landuse. The evolution and Quaternary history of páramo vegetation and climate is reviewed, incorporating the first data from the Lateglacial and Holocene of the Páramo de Sumapaz. The general altitudinal zonation of the páramo vegetation was studied and is presented for both the dry and the humid side of the Cordillera. The zonal and azonal plant communities are described including their physiognomy, composition and syntaxonomy, habitat and distribution. Eighty five syntaxa from the rank of variant to that of the class are newly described, 17 of which are provisional. The vegetation is not ranked syntaxonomically yet, but described on the basis of preliminary tables. A number of azonal communities, part of them of lesser extent, are described in a similar way. The páramo vegetation is primarily determined by the tropical diurnal high mountain climate. The diversity of the páramo vegetation is related to temperature (altitudinal gradient) and to humidity (dry and wet climate). The presence of zonal bunchgrass páramo, bamboo-bunchgrass páramo or bamboo páramo mainly depends on the complex interrelation between these factors. Finally a synthesis is provided on ecology, morphology and phytogeography of the páramo vegetation of the study area.

Description: Studies in the Moraceae showed that Acanthinophyllum should be regarded as congeneric with Clarisia, and that Clarisia spruceana and Aliteria sagoti are conspecific with Clarisia ilicifolia. Some characters of the inflorescences and seeds and the position of Clarisia and its relationships with Trophis and Sorocea are discussed.

Description: Nineteen species of Stereocaulon are treated from the northern Andes, mainly from Colombia. Descriptions and keys are given, with notes on the north-Andean distribution and ecology. Seven species are new for the Colombian flora, viz. St. atlanticum, St. claviceps, St. corticatulum (chem. strain with atranorin and perlatolic acid), St. delisei, St. microcarpum, St. pachycephalum and St. pomiferum. St. crambidiocephalum is reported for the first time from Costa Rica, as is St. didymicum from Venezuela, and St. delisei is reported for the first time from the New World (Colombia and Costa Rica). St. cornutum Müll. Arg. is reduced to synonymy under St. pityrizans Nyl.

Description: The wood and leaf anatomy of representatives of the 9 genera of the Opiliaceae are described in detail. It is possible to separate the genera on the base of both wood- and leaf anatomical characters. Herein the presence of cystoliths of varying shape and size is important. Some comments on the taxonomy and possible phylogeny of the familiy are given.

Description: Isoëtes Palmeri with a distribution in the High Andes from the Páramo of Venezuela to the Páramo of Ecuador is described as a new taxon, and dedicated to the then American specialist of the genus, Thomas Chalkley Palmer (1860-1934). The new species belongs to the tropical-Andeanaustral-antarctic section Laeves, described as new here as well. The publication of the new species had to be anticipated to the projected monographic treatment of the South-American representatives of the genus Isoëtes, as A.M. Cleef, Utrecht intends to base a new association, the Isoëtetum Palmeri on this new taxon, observed and collected by him at many instances within the Colombian Páramo between 1971 and 1980 in the context of the preparation of his doctoral thesis now under way.

Description: The pollenflora of the browncoal mined in the quarry “Maria Theresia”, Herzogenrath, Western Germany has been investigated. Samples were taken along a section of 12.50 meters at intervals of 50 centimeters. Identifications were made on the basis of form-genera and form-species; this working method gives a broad information on pollen types, but does not relate the fossil pollen types primarily to recent plant taxa. Pollen types which are considered to belong to Betulaceae/Myricaceae make up the biggest part of the pollen assemblages. The construction of a “Composite diagram” for Betulaceae/Myricaceae proved to be useful and might give a better idea on paleoecologic conditions. The stratigraphic position of the browncoal examined is thought to be Lower “Hauptflöz”, possibly Morken or Frimmersdorf Horizont.

Description: Nooit zal ik die Donderdagmorgen 10 Augustus 1944 vergeten, toen ik op het laboratorium hoorde dat in de krant — wie las dat vod nog in die tijd — stond dat UITTIEN gefusilleerd was. Het drong eerst niet goed tot mij door. Het kon niet waar zijn. De krant werd gehaald. Ja, daar stond zijn naam in een lange lijst van lotgenoten en het verschrikkelijke, het onherroepelijke, begon langzaam tot mij door te dringen. Koud en gevoelloos stond daar het bericht, van een leugenachtige argumentatie voorzien, dat men ook UITTIEN, die zachtmoedige, gevoelige, intelligente man, had vermoord. Woorden waren hiervoor op dat moment niet te vinden. Ik had alleen behoefte zijn oudste zuster, waaraan hij zeer gehecht was, op te zoeken. Door de slechte treinverbindingen kon ik eerst de volgende dag naar Brummen. Daar trof ik een diep verslagen kring van familie en vrienden van UITTIEN. Wij konden het ons nog zo moeilijk realiseren dat wij hem niet weer zouden zien. Eerst nu wij hem voor goed verloren hadden beseften wij in volle omvang hoe groot wel de plaats was die hij in ons aller leven innam. Van nature had UITTIEN weinig belangstelling voor politiek. Hij vond dat hij daar niets van wist en er dus ook niet aan mee behoefde te doen. Hij had dan ook de gewoonte zijn stembiljet blanco, ja zelfs zonder het open te vouwen, weer meteen in de bus te laten glijden, zeer tot ongenoegen van de partij-mannen die bij een dergelijke gelegenheid op het stembureau plegen te zitten. Wel was hij met hart en ziel het Koninklijk Huis toegedaan. Later heeft hij zijn blanco stemmerij opgegeven, daar het hem duidelijk was dat hij op die manier ongewild toch wel eens de door hem toen reeds verafschuwde N.S.B. zou kunnen steunen. De gang van zaken in Duitsland opende hem de ogen en reeds voor de oorlog liet hij zijn antinazi instelling duidelijk blijken. Zo zond hij na de overval van de Duitsers op Tsjecho-Slowakije een paar overdrukjes aan een botanicus in dat land met op het adres: .... Tsjecho-Slovakia, temporarily occupied by Germany. Dit had tot zijn intens genoegen een geheel onverwacht gevolg, n.1. een stroom van overdrukjes van allerlei Tsjechische botanici waarvan hij nog nooit gehoord had. Na de overval op ons land, het bombardement van Rotterdam, dat diepe indruk op hem maakte, en de daarop volgende bezetting, was UITTIEN dan ook een felle tegenstander van Duitsers en N.S.B.ers. Hij uitte dat waar hij kon in woord en daad. Op de Middelbare Koloniale Landbouwschool te Deventer waar hij leraar was, leidde dat tot wrijvingen met een N.S.B.-collega, die alles aan zijn Duitse meesters rapporteerde. Op 31 Aug. 1941, de verjaardag van H.M. de Koningin, kwam het tot een ernstige, maar niet onvermakelijke botsing met de Deventer zwarthemden, vanwege het feit dat hij binnenshuis met een oranjedas rondliep. Zijn huis aan de Dahliastraat werd door de N.S.B.ers belegerd, hetgeen een grote volksoploop en kloppartij tot gevolg had. Korte tijd daarna werd hij wegens dit feit en zijn „tartende” houding tegen de N.S.B.-collega ontslagen. Daar het departement een gunstige wachtgeldregeling maakte was dit geheel tot zijn genoegen. Sindsdien toch kon UITTIEN zich met nog meer energie wijden aan de taak, die hij zich ten bate van de oorlogvoering gesteld had, nl. het bijhouden van een uitvoerig dagboek en het verspreiden van door de radio opgevangen nieuwsberichten en van illegaal uitgegeven geschriften. Het is buitengewoon jammer dat dit dagboek in de laatste oorlogsmaanden door brand verloren is gegaan. Zijn folkloristische neigingen kwamen hem bij het samenstellen van dit dagboek goed van pas. Dagelijks tekende hij alles aan wat hij hoorde. Elk nieuwtje, elk gerucht, elke anecdote, met nauwkeurige opgave van plaats, tijd enz. Hoewel dus alles door elkaar kwam te staan, nl. alleen in de volgorde zoals hij de berichten kreeg, was het toch een verhaal dat men met spanning zat te lezen. Dat kwam natuurlijk ook vooral door de originele wijze waarop hij het gehoorde op schrift stelde. Zijn dagboek zou ongetwijfeld voor de geschiedschrijving van deze jaren van belang zijn geweest. Hoe ver zijn medewerking aan de illegale bladen zich uitstrekte, kan ik niet zeggen, daar hij dat begrijpelijk ook voor zijn familie en naaste vrienden verborgen hield. Wellicht heeft hij wel eens iets in deze bladen geschreven, maar zijn voornaamste medewerking was zeker de verspreiding. Op 29 Januari 1944 werd hij, op grond van verdenking van medewerking aan de verspreiding van „Trouw”, gearresteerd en naar het concentratiekamp Vught overgebracht. Voor zover wij wisten was er echter geen enkel positief bewijs tegen hem. Dat was dan ook waarschijnlijk de reden dat hij zelf dacht vrij te komen. De weinige brieven die hij uit zijn gevangenschap mocht schrijven waren merendeels opgewekt en getuigden van zijn onvergankelijke gevoel voor humor. Helaas werden zijn optimistische gedachten, geuit in zijn laatste brief, niet tot werkelijkheid. Hij schreef daarin dat hij nu wel spoedig dacht thuis te komen. In plaats daarvan werd echter zijn groep plotseling voor een standgerecht gebracht, en niet voor een gewone militaire rechtbank waarop zij recht hadden. De zaken gingen voor de Duitsers in die dagen slecht. De Amerikanen en Engelsen waren in het Westen doorgebroken. Vermoedelijk is er uit Berlijn een bericht gekomen, dat maar weer eens een voorbeeld moest worden gesteld om de schrik erin te houden. Zo werden deze mensen zonder dat iemand iets van de gang van zaken afwist ter dood veroordeeld en gefusilleerd. Weer was op een misdadige wijze met verkrachting van elk begrip van humaniteit en rechtsgevoel, aan 23 landgenoten het leven ontnomen, rouw en verbeten woede achterlatend.

Description: In Sweden Erlandsson (1942) showed that the species Parnassia palustris L. has two chromosome numbers, 2n = 18 and 2n = 36. Rozanova (1940) found the same numbers in plants collected in the U.S.S.R. Some morphological differences could be demonstrated in the Swedish material (Erlandsson, 1942). This was confirmed by Löve and Löve (1944) and, therefore, these authors (1950) distinguish 2 species: the diploid Parnassia palustris L. em. Löve and the tetraploid Parnassia obtusiflora Rupr. em. Löve, also separated by sterility barriers and by their geographical distribution. In the Netherlands a tetraploid population was found by Gadella and Kliphuis (1963). As the tetraploid population occurs in an area situated far south of the circumpolar distribution area of the tetraploid plants, it seemed worthwhile to determine the chromosome numbers of other plants of Parnassia palustris L. growing in the Netherlands.

Description: Protium aracouchini (Aubl.) March. var. angustifolium Swart n. var. Folia 1- usque ad 5-juga; interjugum basale petiolo plerumque longius sed interjugis aliis brevius; petioluli breves; foliola oblongolanceolata 7.5-10 cm longa, 2-2.75 cm lata, apicem versus distincte angustata, apice gradatim in acumen lineare sexies usque ad nonies longius quam latius acuminato; nervi sec. utrinque usque ad 20. Holotypus: Lindeman 4504 (U, 078268B).

Description: Gradstein et al. (1982) propose to conserve four generic names of Lejeuneaceae: Lopholejeunea (Spruce) Schiffn., Acrolejeunea (Spruce) Schiffn., Trachylejeunea (Spruce) Schiffn. and Taxilejeunea (Spruce) Schiffn., each of which was introduced as a subgeneric name in Lejeunea by Spruce (1884), and subsequently raised to generic rank by Schiffner in his treatment of the Hepaticae in Engler-Prantl (preprint 1893) [see proposals to conserve 675-678 see p. 746]. Although Spruce (l.c.) used for his Lejeunea species a binary nomenclature by combining subgeneric names with specific epithets, it is clear (e.g. text, index) that the binomina are meant as Lejeunea combinations and they are considered as such by most authors (see Gradstein et al. for further details). Before 1893, however, the Sprucean subgeneric names were used in various papers by F. Stephani in a “seeming” generic rank; indeed Stephani now and then referred to them as “genus.” A chronological survey of a number of relevant papers by Stephani, mainly those published in Hedwigia, was given by Bonner et al. (1961), in conjunction with a brief discussion of the subject of this paper. These authors were the first to realize that on the basis of Art. 42 ICBN some generic names in Lejeuneaceae, e.g. Taxilejeunea and Trachylejeunea, can be considered as validly published by Stephani in Hedwigia 28, 1889. Later on Grolle (1979) demonstrated valid publication of monotypic new Lejeuneaceae genera by Stephani in the Bot. Gaz. 15, 1890, e.g. Lopho-Lejeunea and Acro-Lejeunea. For an evaluation of the status of Lopho- Lejeunea Steph., Acro-Lejeunea Steph., Trachylejeunea Steph. and Taxilejeunea Steph., one might consider these names against the background of the entire context of Stephani’s work on Lejeuneaceae until 1893. As the survey of Stephani’s papers in Bonner et al. is rather incomplete, and as there are several points of divergence in opinion, a new analysis of Stephani’s relevant papers (before Sep 1893) is presented below.

Description: Les systèmes radiculaires des espèces des prés salés du Juncion maritimi ont été étudiés. Nous avons distingué six types morphologiques principaux d’enracinement. Les particularités des différents types et espèces ont été décrites et dessinées. La répartition des différents types d’enracinement dans le Junco-Triglochinetum et le Caricetum divisae a été interprétée écologiquement. Les racines de nombreuses espèces du Juncion maritimi sont pourvues de lacunes aérifères. C’est une adaptation au niveau élevé de la nappe phréatique. La résistance mécanique du sol a également une influence sur le système radiculaire. La concurrence des systèmes radiculaires a été discutée. On a essayé de différencier les groupements végétaux par leurs types d’enracinement. Je tiens à remercier M. J. Braun-Blanquet, directeur de la Station Internationale de Géobotanique Méditerranéenne et Alpine à Montpellier, de son aide et de l’intérêt stimulant qu’il a porté à mon travail, et M. J.-M. Betsch de son conseil. Les subventions de la Koninklijke Nederlandse Akademie van Wetenschappen (Montpellier fonds) et du Utrechts Universiteitsfonds m’ont facilité le séjour à la Station et rendu possible ces recherches.

Description: In February and early March, 1961, the senior author spent three weeks on a small savanna in the approximate centre of Suriname, South of Tafelberg, (map 1). He was accompanied by Mr. W. H. A. Hekking. The time was spent in exploring the flora of the savanna and the adjacent forest. As a detailed study of the vegetation of the savannas of northern Suriname was then in progress, several extensive papers being in preparation (Heyligers, 1963; Van Donselaar, 1965; Van Donselaar-Ten Bokkel Huinink, 1966), it was felt that a more thorough inventory of the vegetation and the flora of the savanna might be rewarding. When a general impression of the plant-cover of the area had been obtained, eight representative sample-plots were selected, their vegetation was analyzed and described after the method of the French-Swiss school of phytosociology, and pits were dug in the soil down to bedrock, samples being taken in every distinctive-looking layer. This work was carried out jointly by the senior author and W. H. A. Hekking; part of the floristic exploration was also done by or with Dr. R. M. Tryon, Harvard Herbarium, Cambridge, Mass. The results are here presented. It was felt that in order to integrate them with those obtained elsewhere in Suriname, the collaboration of a specialist familiar with the Suriname savannas in general was required. This was the junior author’s task, who, after his prolonged work on the savannas of northern Suriname, later expanded his work to those of the southern part of the country. The preliminary results of the last-named study are in the press; more detailed field work is in progress as this paper goes to the press.

Description: As has been stated in the introduction of the second part, this third part will include the remainder of the American part of the tribe Eupodostemeae of the subfamily Eupodostemoideae which was not treated in part I, viz. the genera Oserya, Devillea, Ceratolacis, Mniopsis, Podostemum and Castelnavia. Included are the dubious genera, and it also contains additions and corrections to part I, latin descriptions of new taxa, a list of collectors’ numbers in this part, new references to the literature, and a general index to the third part. The attention of the reader is drawn to a publication of SZAFER (1952) in which a fossil Podostemacea from Europe has been described. As I have not seen the material it is at present impossible to judge the value of the discovery though it seems highly improbable that Podostemaceae ever lived in Europe.

Description: The pollen assemblages of a core in the coniferhardwood formation in northwestern Minnesota are compared with the floristics of the recent vegetation in the region. Percentage levels of the main tree components have been compared first with those from recent surface samples taken at the same short distance from various types of upland forests and second with the regional values of the pollen rain in this area (McAndrews 1966). To that end all the data were recalculated on the basis of special pollen sums. The regional diagram of Stevens Pond shows basically the same assemblage zones as established by McAndrews but without the late-glacial Picea-Populus assemblage zone. The pollen in the following Pinus-Pteridium assemblage zone has been interpreted as derived from a pine forest. During the midpostglacial expansion of the prairie eastwards the regional vegetation must have been a Quercus savanna, locally with prairie. Corylus reaches relatively high percentages in this zone. Among the prairie elements especially the occurrence of Lilium philadelphicum may be noted. In the next zone the pollen diagram shows a rise of the curves of mesic elements. In spite of this the comparison with recent surface samples indicates a xerophytic Quercus forest rather than a mesophytic deciduous forest. In the following Pinus assemblage zone pine was present along the margin of Stevens Pond and is therefore overrepresented in the diagram. In the uppermost zone the pollen curves show the effect of logging of the forest about 1900. Pollen of cultivated and introduced plants appear in this zone. Many local pollen types were found, on account of local overrepresentation. This made it possible to compare the local Stevens Pond sequence with the composition of recent lowland vegetation types. The pollen sequence was similar to a large extent to the recent pattern of lake filling, starting with a eutrophic vegetation of Typha latifolia and Salix in the prairie period and leading to a Larix forest and then to a mesotrophic Picea mariana forest, the present edaphic climax on peaty soils. There is a delay, however, in the introduction of acidophilous species, the Larix forest being without Sphagnum and Ericaceae. This is explained by assuming an influence of the vegetation of the surrounding slopes upon the local vegetation. About 1900 the bog forest was destroyed by logging operations and replaced by the present Typha latifolia mat.

Description: Arbor 8 m alta, altitudine pectoris 20 cm diam.; ramuli teretes, glabri. Folia coriacea, glabra, plana, late elliptica vel late obovata, apice obtusa, rotundata vel subacuminata, basi attenuata et in petiolum decurrentia, 9-12 cm longa, 5-7½ cm lata, statu sicco supra viridia, leviter nitida, infra olivacea, opaca, marginibus subrevolutis, integris; costa et nervi primarii utraque facie prominentes, nervi primarii infimi tenues, inconspicui, spatio brevi margine paralleli, ii paris secundi costae paralleli et valde proximi supra basin folii, tum divergentes sub angulo 30°, plus minusve recti inter costam et marginem et 1-1½ cm a margine remoti, nervis primariis paris tertii coniuncta in parte tertia superiore laminae; nervi primarii tenuiores singuli vel plures primariis maioribus intercalati; venae secundariae leves, paucae, ad marginem et apicem laminae arcuatim coniunctae, rete supra occulto, infra minime claro; petiolus 1-2 cm longus, 2 mm crassus. Inflorescentiae axillares, 10-14 cm longae; rhachis glabra; flores geminati; pedicellus communis perbrevis, pedicellus communis cum individuali 4-4½ mm longus, glaber; bracteae minimae, triangulares mox deciduae; alabastra cylindrica, apice clavata, lutea, 8-9 mm longa; lobi perianthii glabri, crassi, apice leviter excavati, 8-9 mm longi, 1 mm lati, anthesi recurvati, pallide flavo virides; filamenta brevissima, lata, parte superiore loborum perianthii affixa; antherae 2.6 mm longae; ovarium brevissime et adpresse fusco-pilosum, 2 mm altum, sensim in stylum protractum; stylus apice leviter clavatus, stigmate terminali; glandulae disci quaternae liberae, crassae, subglobosae, 0.4 mm altae et latae. Fructus ignotus.

Description: In his introductory statements to 'The Symposium on the Phylogeny and Classification of the Filicopsida' which was held in London, 1972, HOLTTUM, when dealing with 'dubious groups of relationships which would particularly repay investigation', mentioned the Polypodiaceae first (HOLTTUM, 1973: 6). Talking about Polypodiaceae the present authors deal with the Polypodiaceae sensu stricto only, thus excluding the Cheiropleuriaceae, Dipteridaceae, Grammitidaceae, and also the Loxogrammaceae, taxa which were formerly (or are still) included in the Polypodiaceae sensu lato. As delineated in this way, this almost exclusively pantropical family consists of about 600 species and an indefinite number of genera.

Description: Some of the basic concepts common to plant taxonomy, palaeobotany and palynology are discussed such as “taxon”, “taxonomic categories”, “genus and species concepts”, as well as “organ and form genera”. The development of the specifically palaeobotanical and palynological concepts of organ and form genera is briefly treated in the light of the shaping of palaeobotanical thought and methods since Adolphe Brongniart. The need for a single category of an artificial nature (i.e., form genus) is acknowledged; the need for a second category of such nature (i.e., organ genus) is questioned. The general guide lines for good palynological practice given by Faegri et al. (1950) are recommended for future use and are reprinted as an appendix to this paper.

Description: The new species Coussapoa manuënsis C.C. Berg is described.

Description: Three genera of the Proteaceae belonging to the tribe Grevilleeae occur in the Guianas and Brazil: Roupala, Panopsis, and Euplassa. The microscopical wood structure of one of these genera, Euplassa, has not been described before. Anatomically it proves to be nearly identical with Panopsis which it also resembles in general properties. Roupala differs in several respects. Within the genera differences are not fundamental and chiefly a matter of numbers and dimensions. A key to the genera is given. The relation of these three genera with the two other members of the tribe in South America, Orites and Gevuina, is discussed.

Description: Until recently relatively little attention has been paid to the study of chromosomes in liverworts. The first substantial contributions were made by Heitz (1927, 1928) and Lorbeer (1934). In the second half of this century chromosome studies on liverworts were mainly carried out in Europe (e.g. Fritsch 1972; Newton 1977, 1979) and Japan (e.g. Tatuno 1959; Segawa 1965a, b, c; Inoue 1968). Inoue (in Koponen 1979) reports that until now 28% of all bryophyte species in Japan have been investigated as to their chromosome complement. A comprehensive, but rather outdated, survey of chromosome numbers in Hepaticae and Anthocerotae was given by Berrie (1960). Work on a new, updated survey is now underway (Fritsch, in prep.). In the present article results are presented of a cytotaxonomic investigation of European species of the genera Aneura and Riccardia (Aneuraceae). Most specimens were gathered in the Netherlands, but some chromosome counts based on French and German plants are also included.