ALBERT

Beschreibung: https://www.researchgate.net/publication/230891291_The_Orbital_Theory_of_Pleistocene_Climate_Support_frim_a_Revised_Chronology_of_the_Marine_d18O_Record

Beschreibung: Summary Holocene sediments of the North Lagoon, Bermuda, were studied with shallow seismic reflection profiles (200 km CSP-survey, UNIBOOM-system) and vibration coring (40 sediment cores, pneumatic vibration corer, Meischner et al., 1981). Seismic Stratigraphy Four seismic sequences are distinguishable by seismic stratigraphy. All seismic sequences correspond to depositional sequences built up during high sea levels in interglacial times. The seismic sequences are separated by unconformities which are often strongly reflective and correspond to emersion planes during glacial phases. The upper sequence (sequence 4) is related to Holocene sediments. The pre-Holocene bedrock is divided into three different seismic sequences (Kuhn et al., 1981): Sequence 1: oldest Pleistocene sequence (pre-Sangamon sea-level highstands), upper boundary with levelled relief (lower boundary not discernible), composed of strongly cemented carbonate sediments, forms the bedrock below Three Hill Shoals Sequence 2: Sangamon (125 ky sea-level highstand), distinct surface morphology, forms the bedrock of a large area below Holocene sediments, Holocene reefs grew up on elevations of the sequence 2 surface, the Holocene reef rim was developed on an elevated rim of sequence 2 Sequence 3: youngest Pleistocene sequence (Sangamon, 105 and 85 ky sealevel highstands lower than recent), deposited mainly in depressions of the bedrock deeper than -15 m below recent Mean Sea Level, levelling the older relief, peat sedimentation in places The distribution of recent reef areas and lagoonal basins is strongly controlled by pre-Holocene topography and geology of the bedrock. During the Holocene approx. 1050 x 106 m3 of carbonate sediments were deposited in the North Lagoon (290 km2) and approx. 1350 x 106 m3 in the reef rim area (170 km2). Sedimentology There are no larger oscillations of the Holocene sea level identifiable in the sedimentological record. The pre-Holocene topography was gradually drowned during the Holocene sea-level rise. At first, the depositional depressions were separated and landlocked. Fresh water peat marshes, fresh water ponds, marine ponds and bays were formed. With rising sea level, the land barriers were more and more eroded, drowned and lost their influence on the back-barrier sedimentation area. Autochthonous and allochthonous peat, lime gyttja and carbonate mud are a typical transgressive back-barrier sediment sequence. After destruction of the barrier, the depositional milieu changed from restricted marine to normal marine, open lagoonal. Sea-grass sediments and nearly mud-free carbonate sand were deposited in shallow water in an exposed environment. Hydrodynamic energy decreases with increasing water depth in the lagoonal basin. A more densely growing reef rim and intralagoonal reef growth added to the protection of the deeper lagoonal floors. Fine-grained sediments were deposited in this environment. They are distributed over a large area of the North Lagoon and form the top of the transgressive lagoonal sediment sequence. Holocene reefs mainly developed on rises of the pre-Holocene surface. In the early Holocene, solid reef build-ups were able to keep up with the rapid rise of sea level. Sand pockets in the reefs were left behind and filled up mainly in the later Holocene. The percentage of fine-grained sediments, produced and resuspended in the reef rim and deposited in the near lagoonal back-reef zone, increased during the Holocene. Two models of Holocene sedimentation in a depression and on an elevation of the pre-Holocene surface illustrate the dependence of vertical facies gradation on pre-Holocene topography. Trends of the mostly polymodal grain-size distributions of the Holocene sediments are a coarsening-upward in the back-barrier and a fining-upward in the lagoonal sediment sequences. Change in the composition of the molluscan fauna in the Holocene sediments (particle size 〉 2000 µm) is an Indication for fades changes. Gastropods are abundant in the basal backbarrier sediments. Bivalves are rare and their diversity 1s low. Sea-grass sediments contain Codakia orbicularis and Astraea phoebia shells. In the sheltered lagoonal environment shell fragments 〉 2000 µm become rare, common species are Gouldia cerina, Pitar fulminata and Finella sp. (approx. 1000 µm). Fine-grained reef-rim derived sediments differ from lagoonal sediments by a higher percentage of Homotrema rubrum fragments and Alcyonaria spicules.

Beschreibung: As an introduction to a number of researches of his own the author wishes to give the following data: „Veen” has two meanings in Dutch: 1. in a petrographic sense (peat) Von Büllow’s definition was accepted: „Torf” ist zu deflnieren als ein meist dunkles, kohlenstoffreiches und ± saures Gemenge unvollständig spezifisch-zersetzter Pflanzenteile, das erdgeschichtlich jüngste Glied der Verwantschaftsreihe der Kohlen, dessen Bildung noch heute andauert.” 2. in a plant-sociological-geographic sense (bog) the following definition has been suggested: a bog is a plot, the surface of which consists of a layer of peat, either covered or not with vegetation, with which that layer is genetically connected. The classification of bogs according to their position with regard to the water-level of the surroundings (Staring) and that of the geological chart were rejected on account of their ambiguous character. The classification suggested by Van Baren according to the environment in which the bogs have been formed, was likewise thought insufficient. Preference was given to the classification according to the plants which gave rise to the peat (eutrophic, mesotrophic and oligotrophic bogs) and according to the origin of the water needed for peat formation (topogenous, ombrogenous and soligenous bogs). The conditions of peat-formation are of a botanical (presence of a vegetation and micro-organisms), climatologic (presence of a certain temperature and moisture) and geological nature (presence of a basin, valley or dead river-branch, certain level of ground water, a possible impervious layer). With reference to a number of authors (Picardt; Van Lier; Grisebach, Venema and Staring; Weber) the alteration in conception as to peatformation from the 17th via the 18th and 19th to the 20th century has been given. The word „Peel” cannot be derived from „palus”. Nothing is certain about its origin. It may mean the low land, bog or marsh. The bogs of the Peel lie on the Brabant-Limburg border-plateau (fig. 2). Lorié and Pannekoek van Rheden have shown that the peatformation of the Peel is likely to have occurred in channels, which have been formed by the Meuse, in co-operation with wind and rain (fig. 4). The bogs were therefore in the first instance topogenous formations, which afterwards developed into ombrogenous bogs. For his own research the author collected peat in three ways: 1. by cutting lumps of peat from open profiles; 2. by boring with a simple peat-bore (photograph 1); 3. by boring with the Utrecht peat-bore, an improvement on Dachnowski’s (fig. 5). To assist in the pollen-analytic examination the samples were treated according to Erdtman’s method. The latter has the following advantages compared with the usual treatment with a 10% KOH-solution: 1. the surface-structures of the pollen-grains are more distinct and as a result the grains themselves can be recognized better; 2. the pollen is more concentrated, so that in spite of the method taking up much time, a saving of time is possible. How the method is applied may be found in the chapter concerned (p. 38 and following). For the stratigraphic examination the samples were broken apart in a glass-bowl of water and viewed with a binocular microscope. Dry sandy samples were broken in water, when seeds and other vegetative parts came floating to the top; next they were put with a brush on thick blotting paper and studied through the binocular microscope. The designations for the sediments and species of peat have been derived from Fægri & Gams. For Scheuchzeria peat a new designation has been added. A plea was made for replacing the word pollen-analysis by „palynology”. A survey of the observations and examinations up to abt. 1935 closes the introduction (see the diagrams of Weber, Erdtman and Duyfjes in the figs. 6, 7, 8 and 9). The author’s own research refers to the Southern and Astense Peel, as in the remaining grounds of the geological chart indicated I 4v (= raised bog) no samples could be taken owing to the digging off having progressed too far. 10 profiles were examined. The situation of the bore-sites has been given in the geological chart of the grounds (fig. 3). The result of the examination (figs. 10—27) and the discussion on it may be summerized as follows: Zoning of pollen-diagrams The sub-zoning of the late- and post-glacial periods according to Blytt & Sernander has proved useful as a zoning of pollen-diagrams, provided atlantic and sub-boreal are joined. It is desirable to replace Blytt & Sernander’s terminology by a different one, because the authors gave a climatologic connotation to their names of periods. The limit between pleistocene and holocene was drawn between preboreal and boreal as Florschütz did. As phases of the holocene the following names were suggested: young post-glacial = sub-atlantic mid post-glacial = sub-boreal and atlantic old post-glacial = boreal. Neither in the Peel nor elsewhere in Holland have Allerød-deposits been found. They are not likely to be found either, as on account of the long distance from the land-ice-margin the flora will have been hardly or not at all influenced by the Allerød interstadial period. For Holland therefore the zoning of the late-glacial according to Firbas (1935) may be considered sufficient. The names of the periods do not bear a climatologic connotation as those of the post-glacial phases do. For the sake of a unity the following names have been suggested: young late-glacial = pre-boreal mid late-glacial = sub-arctic period old late-glacial = arctic period. Forest-history In a table (p. 98), in which likewise the Peel diagrams of Weber, Erdtman and Duyfjes have been inserted, the examined profiles have been arranged from North to South. From each profile it has been stated whether it originated in a certain period (+) or not (—). The sub-arctic phase was characterized by forests of Betula and Pinus and was followed by the pre-boreal phase, in which Corylus and Alnus occurred. Also from the other Dutch diagrams (see list on p. 99) it appeared that in the Netherlands the Alnus pollen occurs with an equal frequency before, during and after that of the Quercetum mixtum. The old post-glacial zone of the diagrams shows a peak in the Pinusline. In contrast with the from Mid-Europe there is not always a maximum in the Corylus-curve after the Pinus-peak. In other Dutch diagrams this phenomenon is likewise found. Only in 28% of all Dutch profiles with a boreal zone does a hazel-maximum succeed a Pinus one. They often co-incide (16%), while in the remaining cases no hazelpeak has been established. There is no fixed order of sequence in the occurrence of the components of the Quercetum mixtum, either in the Peel or elsewhere in Holland. The mid post-glacial is the phase of culmination of warmth-loving forest elements: Alnus pollen shows the highest percentage in this zone. Quercus pollen also occurs in great quantities, while Ulmus and Tilia take up an important place up to the „Grenzhorizont”. The absolute and empiric Fagus pollen limits are found at different heights in the mid post-glacial zone of the diagrams, the rational limit lies somewhere near the „Grenzhorizont”. In the young post-glacial phase the Fagus pollen attains fairly high percentages (up to 30%). The maxima in the East and South-east of the Netherlands are between 20% and 38%; they decrease towards the coast and increase towards the South-east (Hautes Fagnes, Belgium) and East (Germany). It seems incorrect to class the Netherlands almost entirely among the oak-alderterritory poor in beeches, as Firbas did. An attempt has been made to fit the Peel-diagrams into Overbeck & Schneider’s zonation system. For the territory for which it has been made there are already difficulties (p. 104), for use in the Peel and other Dutch diagrams there are even more objections (p. 68, 104). Godwin’s zonation system appeared to be a little less forced, but not quite useful on account of too many details. From his horizons that of Ulmus proved useless for the continent. Neither for the Peel nor for the Netherlands and its surrounding territory can a detailed zonation system be designed. It has proved difficult to proceed any farther than Rudolph’s „Grundsukzession”: birch, pine-hazel-mixed oak-forest-beech, in which the alder generally joins the mixed oak-forest and the hornbeam the beech. Before drawing far-reaching conclusions from the course of the curves (as has been done by some authors) more palynological researches are needed in accordance with the actuality principle, known from geology. Pollen-grains from warmth-loving trees in seemingly sub-arctic spectra In profile 4 (Deurnse Peel II) pollen-grains of Abies, Alnus, Picea, Tilia, Ulmus and Corylus were found in the „late-glacial” zone (figs. 14, 15). Investigations were made as to which of the following possibilities would be the cause of their appearance: 1. in taking and preparing the samples pollution occurred; 2. pollen-transport over long distances has taken place; 3. the pollen-grains found have got secondarily into the deposit; 4. warmth-loving trees have occurred in favourable circumstances in the late-glacial phase or 5. in an interstadial period or in an interglacial phase. The said pollen-grains probably hail from a Würm interstadial or interglacial phase. Interglacial peat On the site of the bore-point 7 it was possible to collect samples from the layers under the peat. The upper 40 cm of the diagram Griendtsveen IX (fig. 27) of this profile proved a repetition of the lower 40 cm of the Griendtsveen I profile (fig. 18). The diagram shows that pollen of Carpinus, Picea and Abies occurs showing the deposit to be of interglacial age. The pollen-curves, however, pass unnoticed from an interglacial into a post-glacial portion. The limit is likely to be found between the two, about 30 cm below the mowing field. There is therefore a great stratigraphic hiatus. Pollen-analytically it could not be decided from which interglacial period the profile hails; on account of its situation on the middle terrace, it was deemed likely that it was an Eem sea deposit. The examined profile probably corresponds to Jessen & Milthers’ zone g; showing it to have been formed at the end of the Eem sea period. The Meuse therefore cannot have flowed through this part of the Astense Peel after the mid Eemean phase. Stratigraphy This is difficult to summarize. Compare various profiles. Individual mention may be made here of: 1. peat on a podsol layer; this was found in two places (Deurnse Peel I Kraaienhut and Griendtsveen VIII). Peat-formation may be thought to have occurred in the following way: heather started growing on drift-sand giving rise to a podsol layer. As the latter is impervious the vegetation surface became marshy. The heath was replaced by a Caricetum from which peat arose. Gradually more Eriophorum occurred, from which almost pure vaginatum peat arose. The bog-surface grew moister and moister, Sphagnum cuspidatum and Scheuchzeria could grow on it and formed a „Vorlaufstorf”. Only then could non-extremehydrophile Sphagna join in peat-formation. 2. the occurrence of Scheuchzeria-peat after the „Grenzhorizont” period. This species of peat, which is often found at the basis of the old Sphagnum-peat as a mesotrophic transition vegetation, has for the Netherlands only been found in the young post-glacial phase in the Peel (Deurnse Peel I Kraaienhut, Griendtsveen V and VIII and Nederweerd). At present the plant is very rare. The severe decline of this plant was also observed elsewhere. Probably it is caused by the gradual drying up or reclaiming of the raised bogs. Of the present station of Scheuchzeria near Ommen a short description has been given (p. 59 and photographs 2, 3, 4). 3. the „Grenzhorizont”. Where the young Sphagnum-peat has not been dug for the preparation of moss-litter, the Peel bogs show a clear „Grenzhorizont” (photograph 8). The conceptions about its origin have been discussed. The distinct separation between the old and the young Sphagnum-peat was not considered sufficiently explained. Though on the whole the „Grenzhorizont” is synchronous in the North-west European profiles, the point of transition from old to young Sphagnumpeat was fairly unstable and easily changeable as to time. Generally the date of the „Grenzhorizont” is fixed at about 500 A.D., though there are differences in opinion. There is a lack of archeological correlation which renders a correct dating impossible. Interference of man in the Peel Three ways of interference were stated: 1. peat has been dug off for the greater part in the territory of the Peel: young Sphagnum-peat for the preparation of moss-litter, old Sphagnum-peat for fuel. The trees which appeared when the bog was dug up in the „Veenderij der Maatschappij Griendtsveen” are sometimes in so good a condition, that they are used for building sheds. The 1 st, 2nd and 4th beam in the foreground of the shed in photo 5 has been sawn from a 30 m long subfossil pine. 2. in a native peat-digging it was possible to collect recent young Sphagnum-peat. 40 to 50 years ago the peasants living there had dug peat in holes, which were afterwards left to themselves. Sphagnum started growing again and the holes were filled in again. The diagram (fig. Griendtsveen VII) represents the surrounding heath with scattered pines and birches, sown by the wind, and a pine-plantation close by. 3. in the profiles Nieuwe Peel, Griendtsveen VI and VII it has been fixed by the indications given by Firbas, that only in the surface layers of the bog has corn-pollen occurred. So in these parts cultivation of cereals will be of recent date. This also appeared from the history of the reclamation of the said territory.

Beschreibung: Recently I got the opportunity of examining a specimen from the “Rijksherbarium”, Leiden, which was provided with a label on which ROTH had written in the middle the name of the plant, viz. “ Micranthus serpyllifol-Roth ” and in the lower right corner the name of the collector, viz. “Heyne”; in the lower left comer another hand had added “Ind. or. Hb. Roth”. As the specimen proved to answer the description of Micranthus serpyllifolius given on p. 282 of ROTH’s “Novae Plantarum Species, Halberstadt 1821,” there can be little doubt that it is either the type of this species or else a duplicate of the latter. This is the more important as none of the authors who in the past ventured an opinion with regard to the taxonomic position of ROTH’s species, apparently had seen the type. ROTH’s specimen was inserted in the Leiden Herbarium under the name Andrographis serpyllifolia R.W. (Acanthaceae), but this is obviously a misidentification. for Andrographis serpyllifolia does not fit ROTH’s description. The plant described by the latter has smaller and less numerous leaves and its flowers are arranged in terminal spikes instead of solitary or a few together in the axils of ordinary leaves.

Beschreibung: Nooit zal ik die Donderdagmorgen 10 Augustus 1944 vergeten, toen ik op het laboratorium hoorde dat in de krant — wie las dat vod nog in die tijd — stond dat UITTIEN gefusilleerd was. Het drong eerst niet goed tot mij door. Het kon niet waar zijn. De krant werd gehaald. Ja, daar stond zijn naam in een lange lijst van lotgenoten en het verschrikkelijke, het onherroepelijke, begon langzaam tot mij door te dringen. Koud en gevoelloos stond daar het bericht, van een leugenachtige argumentatie voorzien, dat men ook UITTIEN, die zachtmoedige, gevoelige, intelligente man, had vermoord. Woorden waren hiervoor op dat moment niet te vinden. Ik had alleen behoefte zijn oudste zuster, waaraan hij zeer gehecht was, op te zoeken. Door de slechte treinverbindingen kon ik eerst de volgende dag naar Brummen. Daar trof ik een diep verslagen kring van familie en vrienden van UITTIEN. Wij konden het ons nog zo moeilijk realiseren dat wij hem niet weer zouden zien. Eerst nu wij hem voor goed verloren hadden beseften wij in volle omvang hoe groot wel de plaats was die hij in ons aller leven innam. Van nature had UITTIEN weinig belangstelling voor politiek. Hij vond dat hij daar niets van wist en er dus ook niet aan mee behoefde te doen. Hij had dan ook de gewoonte zijn stembiljet blanco, ja zelfs zonder het open te vouwen, weer meteen in de bus te laten glijden, zeer tot ongenoegen van de partij-mannen die bij een dergelijke gelegenheid op het stembureau plegen te zitten. Wel was hij met hart en ziel het Koninklijk Huis toegedaan. Later heeft hij zijn blanco stemmerij opgegeven, daar het hem duidelijk was dat hij op die manier ongewild toch wel eens de door hem toen reeds verafschuwde N.S.B. zou kunnen steunen. De gang van zaken in Duitsland opende hem de ogen en reeds voor de oorlog liet hij zijn antinazi instelling duidelijk blijken. Zo zond hij na de overval van de Duitsers op Tsjecho-Slowakije een paar overdrukjes aan een botanicus in dat land met op het adres: .... Tsjecho-Slovakia, temporarily occupied by Germany. Dit had tot zijn intens genoegen een geheel onverwacht gevolg, n.1. een stroom van overdrukjes van allerlei Tsjechische botanici waarvan hij nog nooit gehoord had. Na de overval op ons land, het bombardement van Rotterdam, dat diepe indruk op hem maakte, en de daarop volgende bezetting, was UITTIEN dan ook een felle tegenstander van Duitsers en N.S.B.ers. Hij uitte dat waar hij kon in woord en daad. Op de Middelbare Koloniale Landbouwschool te Deventer waar hij leraar was, leidde dat tot wrijvingen met een N.S.B.-collega, die alles aan zijn Duitse meesters rapporteerde. Op 31 Aug. 1941, de verjaardag van H.M. de Koningin, kwam het tot een ernstige, maar niet onvermakelijke botsing met de Deventer zwarthemden, vanwege het feit dat hij binnenshuis met een oranjedas rondliep. Zijn huis aan de Dahliastraat werd door de N.S.B.ers belegerd, hetgeen een grote volksoploop en kloppartij tot gevolg had. Korte tijd daarna werd hij wegens dit feit en zijn „tartende” houding tegen de N.S.B.-collega ontslagen. Daar het departement een gunstige wachtgeldregeling maakte was dit geheel tot zijn genoegen. Sindsdien toch kon UITTIEN zich met nog meer energie wijden aan de taak, die hij zich ten bate van de oorlogvoering gesteld had, nl. het bijhouden van een uitvoerig dagboek en het verspreiden van door de radio opgevangen nieuwsberichten en van illegaal uitgegeven geschriften. Het is buitengewoon jammer dat dit dagboek in de laatste oorlogsmaanden door brand verloren is gegaan. Zijn folkloristische neigingen kwamen hem bij het samenstellen van dit dagboek goed van pas. Dagelijks tekende hij alles aan wat hij hoorde. Elk nieuwtje, elk gerucht, elke anecdote, met nauwkeurige opgave van plaats, tijd enz. Hoewel dus alles door elkaar kwam te staan, nl. alleen in de volgorde zoals hij de berichten kreeg, was het toch een verhaal dat men met spanning zat te lezen. Dat kwam natuurlijk ook vooral door de originele wijze waarop hij het gehoorde op schrift stelde. Zijn dagboek zou ongetwijfeld voor de geschiedschrijving van deze jaren van belang zijn geweest. Hoe ver zijn medewerking aan de illegale bladen zich uitstrekte, kan ik niet zeggen, daar hij dat begrijpelijk ook voor zijn familie en naaste vrienden verborgen hield. Wellicht heeft hij wel eens iets in deze bladen geschreven, maar zijn voornaamste medewerking was zeker de verspreiding. Op 29 Januari 1944 werd hij, op grond van verdenking van medewerking aan de verspreiding van „Trouw”, gearresteerd en naar het concentratiekamp Vught overgebracht. Voor zover wij wisten was er echter geen enkel positief bewijs tegen hem. Dat was dan ook waarschijnlijk de reden dat hij zelf dacht vrij te komen. De weinige brieven die hij uit zijn gevangenschap mocht schrijven waren merendeels opgewekt en getuigden van zijn onvergankelijke gevoel voor humor. Helaas werden zijn optimistische gedachten, geuit in zijn laatste brief, niet tot werkelijkheid. Hij schreef daarin dat hij nu wel spoedig dacht thuis te komen. In plaats daarvan werd echter zijn groep plotseling voor een standgerecht gebracht, en niet voor een gewone militaire rechtbank waarop zij recht hadden. De zaken gingen voor de Duitsers in die dagen slecht. De Amerikanen en Engelsen waren in het Westen doorgebroken. Vermoedelijk is er uit Berlijn een bericht gekomen, dat maar weer eens een voorbeeld moest worden gesteld om de schrik erin te houden. Zo werden deze mensen zonder dat iemand iets van de gang van zaken afwist ter dood veroordeeld en gefusilleerd. Weer was op een misdadige wijze met verkrachting van elk begrip van humaniteit en rechtsgevoel, aan 23 landgenoten het leven ontnomen, rouw en verbeten woede achterlatend.

Beschreibung: As has been stated in the introduction of the second part, this third part will include the remainder of the American part of the tribe Eupodostemeae of the subfamily Eupodostemoideae which was not treated in part I, viz. the genera Oserya, Devillea, Ceratolacis, Mniopsis, Podostemum and Castelnavia. Included are the dubious genera, and it also contains additions and corrections to part I, latin descriptions of new taxa, a list of collectors’ numbers in this part, new references to the literature, and a general index to the third part. The attention of the reader is drawn to a publication of SZAFER (1952) in which a fossil Podostemacea from Europe has been described. As I have not seen the material it is at present impossible to judge the value of the discovery though it seems highly improbable that Podostemaceae ever lived in Europe.

Beschreibung: De ontwikkeling van het vegetatiekundig onderzoek heeft in de laatste 10 jaar althans in ons land geen gelijke tred gehouden met de daarvoor in sommige gevallen noodzakelijke uitbouw van het terminologisch apparaat. Het mag sommige buitenstaanders misschien voorkomen, dat de terminologie van de vegetatiekunde reeds rijkelijk ingewikkeld is. Dit is echter slechts schijn. Weliswaar bestaat er een indrukwekkende reeks van termen, doch de meeste hiervan spelen in de practijk van het onderzoek geen enkele rol, en zijn slechts bedacht om er zich van te kunnen bedienen in extreme en vaak gezochte probleemstellingen. In de practijk van het onderzoek heeft men behoefte aan behoorlijk omschreven termen voor alle verschillende gevallen die zich kunnen voordoen bij het onderscheiden van vegetatie-eenheden. Daarbij moet bovenal zo nodig een scherp onderscheid gemaakt kunnen worden tussen concrete vegetaties, – die in het Duits “Bestand” genoemd worden, maar waarvoor geen Nederlands woord bestaat, – en abstracte eenheden. Wij hebben reeds herhaaldelijk betoogd, dat het veelal ontbreken van het besef van de noodzaak van dit onderscheid niet bevorderlijk is geweest voor de methodische ontwikkeling van de vegetatiekunde (zie bijv. Westhoff, 19501). Mat name bij de Zweedse, Engelse en Noordamerikaanse onderzoekers heeft dit besef veelal ontbroken. De zgn. Frans-Zwitserse school baseert haar methodiek wel op dit onderscheid, maar het wordt toch niet altijd scherp in het oog gehouden. Opvallend is bijv., dat in het laatst verschenen nummer (1957) van de “Mitteilungen der floristisch- soziologischen Arbeitsgemeinschaft” door prof. Tüxen een poging werd gedaan om het begrip plantengezelschap opnieuw te definiëren, en dat hij deze definitie, die overigens niet onverdienstelijk is, het concrete en het abstracte weer niet uit elkaar gehouden worden. Aan de andere kant is het juist een bezwaar van de Frans-Zwitserse school, dat men zich hier te veel heeft vastgelegd op de associatie als zgn. fundamentele eenheid, zonder er zich altijd voldoende rekenschap van te geven, dat het niet mogelijk is en ook niet de bedoeling van het Frans-Zwitserse systeem is om het gehele vegetatiedek in associaties op te delen. Wanneer men dus de associaties van een bepaald gebied heeft onderzocht en beschreven, blijven er een aantal vegetaties over, die niet of nauwelijks of slechts met gewrongen kunstgrepen tot deze associaties gebracht kunnen worden. Werden deze gevallen door vroegere onderzoekers min of meer gebagatelliseerd of eventueel genegeerd, dit is bij gedetailleerder en nauwkeuriger onderzoek niet aanvaardbaar en met name niet bij vegetatiekartering, waarbij men zich van elk stuk vegetatie methodisch rekenschap moet geven. Een derde moeilijkheid is hierin gelegen, dat vegetatie niet eendimensionaal, doch meerdimensionaal variëert, of om het wat beperkter en daardoor aanschouwelijker uit te drukken, dat een associatie niet alleen door de werking van locale edafische en biotische factoren variëert en dus in verschillende sub-associaties, varianten enz. verdeeld kan worden, doch ook over een grotere ruimte bezien een geografische differentiatie vertoont, zonder dat het nochtans altijd mogelijk is deze beide vormen van varianten scherp te scheiden. Dit probleem heeft in de laatste 20 jaar zeer zeker in de volle aandacht van de onderzoekers gestaan en het gevolg daarvan is eerder een verwarrend teveel dan een tekort aan terminologie geweest.

Beschreibung: Aan hen die nog ingevulde hoklijsten onder hun berusting hebben, wordt verzocht deze op te sturen aan het Rijksherharium, afd. Nederland. Er wordt op het ogenblik hard gewerkt aan het inboeken van alle gegevens in de albums.

Beschreibung: Door F. Drouet (Nat. Hist. Mus., Chicago) en mij werd thans een door J. Poolman op 27 aug. 1944 te Noorbeek (Zuid-Limburg) gevonden groenwier herkend als Chlorotylium cataractarum Kütz. Volgens W. Heering (in Pascher, Süsswasserfl. H.6, 1914), H. Printz (in Engl. -Prantl, Nat. Pfl. Fam. ed.2, Bd-3, 1927) en G.M. Smith (Freshw. Alg. U.S., Behoort deze soort tot de Chaetophoraceae, volgens F.E. Fritsch (Struct. and Reprod. I, 1935) tot de Trentepohliaceae. Zij groeit op hout en stenen in snel stromend water, zodat de soortnaam goed gekozen is. Uit Nederland was deze soort nog niet eerder Bekend, In Noorbeek groeide het wier in een drinkbak voor dieren, waar het water in stroomde uit een beek.

Beschreibung: Tragopogon dubius Scop. Het duinterrein te Nieuwe Sluis in de gemeente Groede, waar verleden jaar de planten van Tragopogon dubius Scop. groeiden, is de afgelopen winter in verband met herstellingen aan de zeewering met behulp van draglines en bulldozers geëgaliseerd, waardoor de Tragopogon naar ik dacht volkomen uitgeroeid zou zijn. Ik bemerkte dit pas dit voorjaar en kon dus geen maatregelen nemen om een gedeelte van het terrein te sparen. Bovendien vrees ik, dat men aan mijn verzoek toch geen gevolg had kunnen geven. Enkele weken geleden bezocht ik het terrein weer en tot mijn vreugde vond ik toch nog twee planten, die het overleefd hadden. Mij bleek echter, dat kneuen bijzonder verzot zijn op de onrijpe zaden van deze soort. Zij pikken de omwindsels stuk en halen zo de onrijpe zaden er uit, zodat het de vraag zal zijn of er nog iets voor het volgend jaar zal overblijven. Ook verleden jaar was mij dat opgevallen, doch bij de vele planten, die er toen groeiden, was dat niet zo’n bezwaar. Eigenaardig is, dat ik aan planten van Tragopogon pratensis iets dergelijks nimmer heb waargenomen. Hebben anderen dat wellicht wel gedaan?

Beschreibung: In 1957 waren wij weer herhaaldelijk in de gelegenheid het adventiefterrein “de Dwinger” aan de Langesloot tussen Wartena en Eernewoude te het laatst op 1 november, samen met M.T. Jansen. Hier volgt een opgave van de nieuw waargenomen planten (zie Corr.bl. no. 1 en 4). Buiten adventieven was het aantal verwilderde kultuurplanten vrij groot. De heren dr. S.J. van Ooststroom en Th.J. Reichgelt waren weer bereid het materiaal te controleren, terwijl de heer G. Bakker, direkteur der Gemeentereiniging Leeuwarden, opnieuw toestemming verleende het terrein te betreden.

Beschreibung: During 1954 the Gray Herbarium, the Orchid Herbarium of Oakes Ames, the paleobotanical collections of the Botanical Museum and a portion of the herbarium collections and the library of the Arnold Arboretum were moved into a new building in Cambridge, Massachusetts. This move was the culmination of a long period of planning to determine how the best interests of each institution as well as the field of systematic botany could be served best in this period of rapidly developing interrelationship of diverse scientific fields of knowledge. Additional considerations prior to the move were the isolation of the various taxonomic units at Harvard, the duplication of resources, efforts and goals, as well as the more mundane problems of increasing costs of labor, material and demands for additional storage facilities. In 1946 the President and Fellows of Harvard College appropriated from its unrestricted funds the sum of one million dollars to construct and equip a new and modern building to house the systematic work and collections of these institutions in Cambridge, to be in close proximity to the resources of the Department of Biology, the Museum of Comparative Zoology and the Farlow Reference Library and Herbarium of Cryptogamic Botany. The building, designed around the requirements established by the taxonomists of these institutions, was under construction during 1953 and was finished in the early months of 1954.

Beschreibung: Mr C.T. White is to be congratulated on being presented with, the Mueller Memorial Medal awarded by the Adelaide Meeting of the Australian and New Zealand Association for the Advancement of Science, Aug. 1946. This award is in recognition of his work on the systematic botany of Queensland. Dr Ir J.Ph. Pfeiffer, Director of Research, B.P.M.-lab., Amsterdam, died Nov. 18, 1947, at Amsterdam, 58 years old. He was formerly wood-technologist, and collected plants in Simaloer Island, NW Sumatra.

Beschreibung: Uit afschriften van door de heer W.W. Schipper te Winschoten in de jaren omstreeks 1904 ingevulde kwartierhokstaten blijkt, dat voor H8-61-42 (Bovenhuren bij Winschoten), H8-62-34 (bij Winschoten Oostereind) en H8-63-32 (bij Klein Ulsda) Agrimonia eupatoria L. opgegeven werd, doch nergens Agrimonia odorata (Gouan) Mill. Uit kwartierhok H8-61-41, eveneens Bovenburen, Winschoten, nl. langs de Kloosterweg, is mij Agrimonia odorata (Gouan) Mill. van 1917 tot ca. 1942 bekend geweest. De standplaats langs een meestal droge sloot op hoge fluvioglaciale zandgrond is ca. 1935 verwoest door verbetering van de weg en aanleg van lintbebouwing. Gelukkig bleek de plant toen een honderd meter verder aan de overkant van de weg een nieuwe groeiplaats te hebben verworven, die thans helaas niet meer bestaat. Ik heb er toen wat vruchten van verzameld, die zeven forse planten hebben opgeleverd, maar in de oorlogsjaren verloren zijn gegaan. Van de vruchten van deze exemplaren heb ik twee planten kunnen opkweken aan een veendijkje te Oude Pekela, doch ook die planten zijn vernietigd, toen in 1957 dat dijkje wegens een nieuwe waterschapsindeling werd afgegraven. Haar mijn weten komt Agrimonia odorata thans niet meer in Oostelijk Groningen voor. Ik vermoed evenwel, dat de bovenvermelde planten van A. eupatoria ook A. odorata geweest zullen zijn en dat het areaal in de Noordduitse laagvlakte zich over Groningen tot in Friesland uitstrekte.

Beschreibung: Index Kewensis. Suppl. 10. (1936-1940). Clarendon Press, Oxford, £4/4. (1947). Check List of British vascular plants (Journ. Ecol. 33 (1946) 308-347). Nomenclature accepted by the Brit. Ecol. Soc. to uniformize the binary names used for British plants.

Beschreibung: Besides the importance of correct identification the revision of a large genus should add considerably to knowledge of phytogeography and of infrageneric diversification. In all respects Ficus has much to contribute. It is a genus which the collector meets in abundance in all parts of tropical Asia and Australasia, whether in primary or secondary environments, and which he soon learns to recognise. It can be exploited, therefore, provided the species can be identified. The purpose of this note is to request intensified collection, because I believe it is possible to name satisfactorily sterile material. Only too often, valuable sterile material is left uncollected, as I know from my own experience, for sooner or later it can be recognised as a positive record from some locality. Some figs, too, fruit rarely and are in consequence ill-represented, though really frequent.

Beschreibung: Mr A.G.L. Adelbert of Herbarium Bogoriense was transferred to the new Garden Setia Mulya near Padang, Sumatra’s Westcoast, as leader of the staff, Dec. 1954. Cf. also chapter 6. Dr A.H.G. Alston was in Malaysia. Cf. chapter 5.

Beschreibung: ANDERSON, J.A.R., A checklist of the trees of Sarawak, 364 pp. (1983, Dewan Bahasa dan Pustaka Cawangan Sarawak, for Forest Department, Kuching, Sarawak). Cloth Mal$ 15.00. When Dr. Anderson retired from the Forest Department in 1973 he left the manuscript of this checklist for publication. Unfortunately publication was delayed for 10 years. It contains data on over 2500 arboreous plant species. The text consists mainly of two parts: the first is a list of vernacular names with their scientific equivalents, the second is a list of plant names alphabetically arranged by family. Each species is concisely annotated with its vernacular name(s), maximum diameter, ecology, frequency, soils, etc. Species names have been coded: the first two figures are for the family, the next two for the genus and the last two for the species. A list is given of the trees of the peat-swamp forests of which Anderson was a great expert. A small draw-back is that the literature of the last ten years has not been included. Nevertheless this is a most helpful book. — C.G.G.J. van Steenis.

Beschreibung: We are glad to be able to add to the list of herbaria which have agreed to send on loan herbarium specimens to collaborators of the Flora Malesiana: Herbarium of the Forestry Department, Sandakan, British North Borneo. Mr H.G. Keith, Conservator of Forests is in charge. Herbarium of the Forestry Department, Lae, Territory of New Guinea. Mr J.S. Womersley, Forest Botanist, is in charge (see p. 61).

Beschreibung: Dr C.A. Backer is now preparing the MS. on the Orchidaceae for the Flora of Java on the basis of a MS. by the late Dr J.J. Smith. Mr J. Monachino has finished his revision of the genus Alstonia (Apoc.); it is expected to be published early in 1949 in ”Pacific Science”, Hawaii.

Beschreibung: It is a great pleasure to announce that the technical difficulties delaying the printing of Flora Malesiana have now been overcome. The first part of volume 4 is in the press and, in all probability, will appear towards the end of this year. Sample sheets of volumes 1, 2, and 3 will be added to the initial instalment of volume 4. Owing to a generous grant by the Netherlands Indies Government of this first issue of the 4th vol. 2500 copies will be printed and distributed to all individual botanists and institutions which are believed to have an interest in the Flora, in order to enable them to form an idea of the scope, execution, and costs of subscription of the work. Those receiving this Bulletin will also receive the initial part. It is expected that volume 1 – which will be issued as one whole – will be in print at the end of this year.

Beschreibung: ( (Report in the ”Gardens’ Bulletin, Singapore”, vol. XI, pt 4, 1947). Prior to the Japanese attack on Malaya, most of the senior staff of the Gardens were seconded for other duties under the Department of Food Control and Information, for at least part of the time. The result was that botanical work was reduced, and considerable arrears of unnamed and undistributed specimens accumulated. The Gardens were maintained as usual, with the addition of demonstration plots of vegetables.

Beschreibung: Owing to shortage of time, this most precious element in the life of a taxonomist, nearly two years have elapsed since the last number (10) appeared (Febr. 1953). It should not be understood that our interest in editing this Bulletin has waned; we still regard it as a useful bond between Flora Malesiana Foundation, its collaborators, and its sympathizers. It also aims to chronicle some selected miscellaneous news to many people in Malaysia who are far from libraries or have only limited facilities to keep informed about progress. It tries to assemble data on activities of botanical work in the wide sense performed in the Malaysian area. Much work that is done in the field or is going on in establishments of forestry and botany in the Malaysian tropics is often locally known or published in technical reports which hardly reach the scientific botanic world. There is, hence, a field of mutual interest which this Bulletin tries to cover.

Beschreibung: In 1826 REINWARDT published in ”Sylloge Plantarum” &c, vol. 2, pp. 1-15 under the title ”Nova plantarum indicarum genera” an article containing descriptions of some Malaysian genera of phanerogams. Amongst them is described on pag 1: Angiopetalum punctatum Reinw. n.g.n.sp. from Java. Though assigned to the Myrsinaceae by DALIA TORRE & HARMS this genus has hitherto remained obscure, and has not even been mentioned by MIQUEL. However, there is a name Allopetalum punctatum REINW. mentioned by SCHEFFER (De Myrsin. 1967, 93) as a MS. name in the synonymy of Ardisia pumila BL., also mentioned by MEZ (Pfl. Reich 9 (1902) 171) for that plant, which is now commonly known as Labisia pumila (BL.) B. & H. The type specimens of Allopetalum punctatum REINW. at Leyden (sheets 908.133.- 614 and 903.255 – 190) are undoubtedly the type specimens of Angiopetalum punctatum REINW. The name under which this species was published differs from that found in REINWARDT’s handwriting hut this is of small significance. Many name-changes occur in the materials assembled by KUHL & VAN HASSELT, ZIPPEL, REINWAKDT (and BLUME) whose herbaria were left in BLUME’s care. On the type sheet of Orescia montana REINW. in the same paper of REINWARDT’s I found on the labels the following MS. names: Lysimachia montana BL., Phaemeria montana, Rumeria montana and Lysimachia cuspidata BL, an embarrassing choice from which only the last one has been validly published. In the case of Angiopetalum, REINWARDT who had probably the herbarium not at his disposal copied the name from MS. notes, the herbarium being with BLUME either in Java or at Brussels. Later he hardly paid any attention to phytography or nomenclature.

Beschreibung: As was pointed out in the first instalment, the management of Flora Malesiana acknowledge collaborating and co-operating institutions; for this purpose a distinction was made between collaborators and co-operators. The former take an active part in the composition of the work (by revising certain families or large groups), the latter give assistance through the loan of specimens, information about collections, biblographical assistance etc.

Beschreibung: Small evergreen trees, shrubs or lianas; two genera ( Cansjera and Opilia) are known to be root-parasites. Leaves distichous, simple, usually extremely variable in form and size, entire, exstipulate, pinnately veined; dried leaves mostly finely tubercled by cystoliths located in the mesophyll. Inflorescences axillary or cauliflorous, panicle-like, racemose, umbellate (in Africa) or spicate; bracts narrowly ovate or scale-like, in Opilia peltate, often early caducous. Flowers small, (3—) 4—5) (—6)-merous, mainly bisexual, sometimes unisexual and plants then dioecious ( Gjellerupia, Melientha, and Agonandra) or gynodioecious (Champereia). Perianth with valvate, free or sometimes partly united tepals (in ♀ flowers of Gjellerupia wanting). Stamens as many as and opposite to the tepals (in ♀ flowers only small staminodes); anthers introrse, 2-celled, longitudinally dehiscent. Disk intrastaminal, lobed (lobes alternating with the stamens), annular, or cupular. Ovary superior, 1-celled; style short or none, stigma entire or shallowly lobed. Ovule 1, pendulous from the apex of a central placenta, anatropous, unitegmic and tenuinucellar. Fruit drupaceous, pericarp rather thin, mesocarp ± fleshy-juicy, endocarp woody or crustaceous. Seed large, conform to the drupe, without testa; hilum basal, often in a funnel-shaped cavity. Embryo terete, embedded in rich, oily endosperm, nearly as long as the seed or shorter, with 3—4 linear cotyledons, radicle often very short. Distribution. There are 9 genera with about 30 spp., widespread in the tropics. Rhopalopilia is restricted to Africa and Madagascar, Agonandra to South and Central America. In Malesia: 7 genera, 5 of these only known from the eastern Old World (1 endemic: Gjellerupia in New Guinea); Opilia and Urobotrya occur also in tropical Africa.

Beschreibung: Monoecious, medium-sized to very large trees (rarely shrubby in very exposed situations). Either four independent cotyledons or two fused pairs (which may be retained in the seed after germination). The growing point of foliage shoots quite distinct between the two genera, being just a few highly reduced leaves in Araucaria and a highly organized bud formed of overlapping scales in Agathis. The leaves vary from scales or needles to broad leathery forms with many parallel veins sometimes on the same plant at different stages of growth. Pollen produced in cylindrical cones from one to as much as twenty cm long with numerous pedunculate spirally placed microsporophylls each with several to many pendent elongated pollen sacs attached to the lower side of an enlarged shieldlike apex which also projects apically more or less overlapping the adjacent microsporophylls. Pollen cones solitary, terminal or lateral, on branches separate from those bearing seed cones, subtended by a cluster of more or less modified leaves in the form of scales, deciduous when mature. Pollen globular, without ‘wings’. Seeds produced in large, well-formed cones which disintegrate when mature, dispensing the seeds in most cases with the help of wing-like structures; the seed cone terminal on a robust shoot or peduncle with more or less modified leaves that change in a brief transition zone at the base of the cone into cone bracts, formed of numerous spirally-placed bract complexes, usually maturing in the second year. Individual seed cone bract leathery or woody and fused with the fertile scale which bears one large inverted seed on its upper surface. Distribution. The 40 species in two genera are well represented in Malesia (13 spp.) and extend eastward and southward into Fiji, New Caledonia (18 spp.), Australia, and New Zealand, with 2 spp. also in the cooler parts of South America, giving the family a distinct Antarctic relationship. Only one species of Araucaria (in South America) occurs completely outside of the tropics, while the majority of the species in the family belong in the lowland tropics and others grow in the tropical highlands.

Beschreibung: Dioecious or monoecious small shrubs with thick woody roots. Leaves simple, opposite, sessile, fleshy, with a distinctly saccate, colourless base. Stipules minute. Flowers unisexual, either solitary and terminal or axillary, or in small axillary spikes. ♂ Flowers subtended by bracts, enclosed in a membranous spathella which opens with one or two transverse or radial slits giving rise to 2-4 lobes. Tepals 4, valvate. Stamens 4, alternitepalous; anthers dorsifixed, introrse, dehiscing lengthwise with 2 slits. Sometimes an abortive gynaecium present. ♀ Flowers merely consisting of a naked ovary, in the axil of leaves when solitary, in the axil of cordate bracts when growing in spikes, 2-carpellate, 4-celled by one true and one false septum; ovules 1 in each cell, basal, anatropous, with a long funicle. Stigmas 2, sessile, distinctly papillate. Fruit a septicidal berry dehiscing with 2 valves, either solitary or many united together with the bracts into a connate, spikelike whole. Seeds with a large, straight embryo, exalbuminous. Distr. The Batidaceae, consisting of one genus with two species, show a remarkably discontinuous area, viz B. maritima L. growing along the Atlantic and Pacific coasts of tropical America, the Hawaiian and Galapagos Islands, while B. argillicola has hitherto only been found in South New Guinea. As the distribution of the species is still rather insufficiently known and they are confined to littoral districts it has been found advisable to include both of them in the key given below.

Beschreibung: Small trees, mostly deciduous, bark gummy, wood soft, roots thickened, pungent; trunk often inflated. Leaves spread, imperfectly 2—4-imparipinnate; tissue with myrosin cells; pinnae opposite, provided with stipitate glands at the base of the petiolules and pinnae. Leaflets small, opposite, entire, all articulated. Stipules represented by blunt knobs. Flowers bisexual, zygomorphic, white (or yellow streaked red), in axillary panicles. Calyx tube short, as a hypanthium; lobes 5 imbricate, spreading or reflexed, separately dropping. Petals 5 free, anterior one largest and erect, others reflexed, posterior smallest. Disk lining the calyx tube, with a short free margin bearing the androecium. Perfect stamens 5 epipetalous; anthers dorsifixed, 1-celled, oblong, when lengthwise opened broader. Staminodes 5, subulate, with or without rudimentary anthers. Ovary superior, shortly stalked, 1-celled with 3 parietal placentas. Style filiform, stigma small. Ovules ~, in 2 series on each placenta. Capsule linear, beaked, 3—6-angled; valves thick, spongy, on the inside with pitted cavities in 1 row along the median line. Seeds 3-winged (or exalate), body roundish large. Embryo exalbuminous, straight, containing oil. Distr. Ca 10 spp., confined to the semi-arid countries of Somaliland, Madagascar, SW. Africa, NE. Africa, Asia Minor, 2 spp. in India.

Beschreibung: Small trees, shrubs or twining woody plants, rarely herbs; branches terete. Glands present in various parts. Indumentum consisting of simple hairs, or in Viburnum sometimes lepidote; glandular hairs mostly present. Stems often pithy. Leaves decussate, simple or deeply divided (Sambucus), sometimes provided with pitted or cup-shaped glands exuding resin. Stipules absent or very small. Flowers ♀, actinomorphic or zygomorphic, mostly cymosely arranged, 4—5-merous; outer flowers in an inflorescence sometimes differing from the normal ones, rarely ( Sambucus p.p.) some fls aborted into extra-floral nectaries. Calyx adnate to the ovary, (4—)5-fid or -toothed, mostly constricted below the limb; sepals often enlarged in fruit. Corolla epigynous, gamopetalous, sometimes 2-lipped, lobes mostly imbricate in bud. Stamens inserted on the corolla tube, alternating with the lobes, extrorse or introrse. Anthers free, 2-celled, dorsifixed, versatile, cells parallel, opening lengthwise, mostly introrse; filaments sometimes reflexed or curved in bud. Ovary inferior, 1-(2-)3-5(-8)-celled, in fruit cells sometimes partly abortive. Style terminal, often slender with one knoblike stigma, or 3 short partly connate styles. Ovules 1(-~), pendulous or axile. Fruit a drupe or berry, rarely a capsule. Seeds often only one per fruit, often with bony testa. Endosperm copious, sometimes ruminate; embryo straight, often small and linear, axial, cotyledons oval or oblong. Distr. Ca 10-14 genera, mainly distributed on the N. hemisphere, in the tropics mostly confined to the mountains, on the S. hemisphere only Viburnum and Sambucus, an endemic genus in New Zealand, two monotypic endemic genera in New Caledonia, in Australia only Sambucus in the eastern part.

Beschreibung: Annual or perennial herbs or shrubs, often fleshy, glabrous, papillate or hairy. Leaves opposite or alternate, exstipulate, sometimes seemingly wanting, stalked or sessile, entire, dentate-serrate-lobed or irregularly gashed. Flowers solitary, 2—3-nate or glomerate, usually sessile, either axillary or in terminal or axillary dense or interrupted spikes or panicles, ♀ or unisexual, monochlamydous, rarely achlamydous, small; bracts present or absent, usually small, rarely leafy. Perianth herbaceous or sometimes scarious, rarely (in ♀) absent, 3—5-partite with (in bud) imbricate segments, or sometimes almost entirely gamophyllous and then shortly lacerate-dentate or unilaterally cleft, persistent, after anthesis accrescent or not. Stamens often the same number as tepals and opposite to them, sometimes fewer, usually inserted on or near base of perianth; filaments free or shortly connate; anthers dorsifixed or inserted in a basal cleft, 2-celled (4-locellate); cells bursting longitudinally. Ovary free or at the base adnate to the perianth, 1-celled; ovule 1, basal, sessile and erect or suspended from a funicle; styles or stigmas 2-5, linear. Utricle either enclosed by the perianth or not, indehiscent or rarely operculate; seed erect, oblique or horizontal, usually compressed; endosperm mostly present, peripheral, surrounding the embryo; embryo annular or spirally twisted. Distr. Species numerous, inhabitants of the temperate and tropical zones of both hemispheres.

Beschreibung: Delicate, annual or perennial herbs, aquatic and then either entirely submersed, or floating in the upper part, or, in humid localities, not rarely terrestrial and creeping, with slender stems. Leaves opposite, at the summits of floating stems often spuriously rosulate, exstipulate, small, linear, elliptic, oblong or spathulate, entire, herbaceous, in the Mal. sp. triplenerved. Flowers minute, unisexual, axillary, solitary or rarely one ♂ and one ♀ flower from the same axil, often with 2 caducous, transversal, opposite, tender concave bracts. Calyx and corolla absent, ♂: Stamen 1; filament thin, anther 2-celled, cells bursting lengthwise, the slits becoming confluent at the top. ♀: Ovary sessile or subsessile, 4-lobed, 4-celled. Ovule solitary in each cell, pendulous from the top of the cavity. Styles 2, free, often long, papillose. Fruit 4-lobed, with longitudinally margined or winged lobes. Testa membranous; endosperm fleshy; embryo terete, straight. Distr. Only genus in the family, worldwide distributed, not yet known from S. Africa and in various regions scarce, in Malaysia apparently very rare, the only record proving its being indigenous is from the New Guinean highlands. Because of their small size terrestrial forms are easily overlooked.

Beschreibung: Adelbert, A.G.L.: Labiatae (in Backer, Beknopte Flora van Java (emergency edition) part 14, March 1954, 1-59, mimeograph). Full descr. and keys to genera and species; in Dutch.

Beschreibung: Annual or perennial aquatics and marsh plants, sometimes laticiferous. Leaves basal and erect, sometimes floating, rarely all submerged, sometimes some reduced to phyllodes, lanceolate to sagittate, rarely broad-elliptic to ovate, entire, with a hydathode on the apex, curvinerved, nerves more or less parallel and gradually joining the marginal nerve, connected by ascending cross-veins; petiole sheathing, mostly with air-channels, often septated. Inflorescences mostly erect, racemose or paniculate; peduncle sometimes hollow, mostly with air-channels. Bracts 3(-2) per whorl of flowers or branches. Flowers actinomorphous, bisexual or unisexual (and then rarely with rudiments of the other sex). Sepals 3, imbricate, green, parallel-nerved, convex, persistent. Petals 3, imbricate, white or faintly coloured, marcescent. Stamens 3-~, free, in a whorl; filaments filiform or dilated; anthers 2-celled, basifix, sometimes versatile, latrorsely lengthwise dehiscent. Carpels 2-~, free, in the Mal. spp. spirally arranged on the receptacle (in extra-Mal. Alisma in a whorl); style 1, ventrally or terminally inserted on each carpel, persistent. Ovule 1 (in extra-Mal. Damasonium 2 or more), basal, campylotropous, rarely anatropous (Damasonium), micropyle extrorse, rarely introrse ( Luronium). Achenes in a head (or whorl in Alisma), free, rarely connate at the base. Seeds oblong or horseshoeshaped; testa membranous; embryo horseshoe-shaped; albumen 0; radicula extrorse, rarely introrse (Luronium). Distribution. About 10 genera with c. 70 spp., all over the temperate and tropical zones except in the Pacific area (Micronesia, Melanesia, and Polynesia). The largest genera are Sagittaria and Echinodorus both centering in the New World.

Beschreibung: Many botanists must have wondered why as yet no volume of Flora Malesiana was dedicated to the outstanding botanist Carl Ludwig Blume, undisputed pioneer in planning the compilation of a ‘Flora Malesiana’. The writing of this Dedication would have been greatly facilitated if a full biography of BLUME had been existent, but none is available; there is not even a bibliography of his works. Only recently, in 1979, two biographical attempts were made, by J. MACLEAN and by A. DEN OUDEN, but only for the period 1820-1832; together with other biographical and obituary notes they are here assembled in Appendix B. I have also compiled a bibliography: Appendix A.²

Beschreibung: Annual or perennial, laticiferous herbs, rarely shrubs. Leaves (in Mal. spp.) spirally arranged, often incised. Stipules 0. Flowers mostly solitary, large, actinomorphic, ♀♂, 2—3-merous. Sepals caducous or calyptrate, free or united. Petals free, 4-6, rarely more or absent, biseriate, imbricate, often crumpled in bud; nectaries absent. Stamens ~, free; anthers 2-celled, dehiscing lengthwise. Ovary superior, 1-celled, with 2 or more parietal placentas (sometimes with protruding placentas or a spurious wall). Stigmas opposite or alternate with the placentas. Ovules 1 to numerous. Capsule opening by valves (or pores). Seeds small, with a crested or smooth raphe, or arillate; embryo minute; endosperm copious, fleshy or oily. Distr. About 23 genera, of which the bulk on the N. hemisphere, few in Central & S. America, almost absent from Africa and Australia, in Malaysia none native.

Beschreibung: Trees (or shrubs), often deciduous, producing gum and an orange juice. Leaves spread, palmatilobed, often with domatia in the axils of the main ribs; stipules caducous. Flowers actinomorphic, bisexual, showy, mostly golden-yellow, paniculate or racemose. Sepals 5 imbricate. Petals 5, imbricate or contorted, emarginate. Stamens ~, with free filaments, equal or subequal; anthers 2-celled, linear, basifixed, opening by introrse, short, often confluent pore-like slits. Ovary 1-celled with laminal placentas projecting into the cell, or perfectly or imperfectly 3-celled, the upper portion remaining 1-celled; ovules ~, style simple, stigma punctiform. Capsule 3—5-valved, valves of the endocarp separating from and alternating with those of the pericarp. Seeds covered by woolly hairs, mostly cochleate-reniform; endosperm copious, rich in oil; embryo large, conforming to the shape of the seed; cotyledons broad. Distr. Ca 15 spp., mostly in trop. and subtropical America, some in trop. Africa and SE. Asia, 3 species in N. Australia, rare in Malaysia; G. gillivrayi is possibly the only native Malaysian species. LAM assumed the genus to belong to the ‘antarctic’ type(Blumea 1 (1935) 135), but it is manifestly peri-tropical.

Beschreibung: Families and higher taxa have been entered under their name. Suprageneric epithets have been entered under the family name to which they belong preceded by the indication of their rank (tribes, e.g.).

Beschreibung: Evergreen, glabrous trees or shrubs, without resin-tubes. Leaves spread, simple, entire, more or less crowded towards the ends of the shoots, shining, exstipulate; midrib sulcate; shoots with perular terminal buds. Branches often in pseudowhorls. Inflorescences terminal, sometimes lateral, generally not exceeding the leaves. Flowers on the ultimate axis in fascicles of 3, towards the end solitary, pedicellate, bracteate. Calyx deeply 5-lobed, fleshy, persistent, petaloid, lobes inequal, concave, imbricate, 2 outermost smallest. Petals 5, thinner than the sepals, inserted at the margin of the disk-like receptacle. Stamens 5, attached to the base of the petals; filaments flattened or terete, slightly thickened towards the base; anthers dorsifixed, dehiscing lengthwise, intrors. Staminodes petaloid, dentate in the upper half, top mostly pointed, alternating with the petals. Disk glands 5, ovoid to ellipsoid, epistaminodial. Ovary ovoid, originally 2-celled, one cell soon abortive. Styles 1-2; stigma punctiform. Ovule 1, pendulous, anatropous. Fruit drupaceous, or a nut, with fibrous endocarp. Testa membranous; cotyledons planoconvex; albumen absent. Distr. Four spp., one each in New Zealand and adjacent islands, N. Caledonia, the New Hebrides, and N. Queensland & E. Malaysia.

Beschreibung: Mostly perennial, paludose, grass-like herbs with fibrous roots; stembase very rarely thickened, often profusely producing shoots. Leaves basal, distichous on each shoot, ensiform, linear or filiform, sometimes twisted; sheaths with a membranous margin (in Mal. spp.) producing mucilage (?always), with or without a short ligule; limb glabrous or with numerous, small hard papillae, sometimes with a stout nerve in either margin. Flowers ♀♂, in terminal, few- to many-flowered heads, 3-merous, yellow to white, ephemeral, each in the axil of a conspicuous bract; bracts conchate, imbricate, spirally arranged, lower ones sterile; one to few flowers simultaneously in anthesis. Peduncles scape-like, terete to compressed, sometimes winged or ribbed, glabrous or with numerous hard papillae, at the base with some sheaths provided with a short limb. Bracts entire, ciliate, fimbriate or lacerate, with one complete main nerve and some complete or incomplete longitudinal secondary (descending) nerves, in the apical part mostly with a small minutely-papillose field. Calyx zygomorphic; lateral sepals navicular, with entire, dentate or ciliate crest, wings membranous, entire, glabrous or ciliate; median sepal membranous, spathelliform or cap-shaped, enveloping the corolla, mostly obovate, 1-3(-5)-nerved, pushed out by the corolla in anthesis(?always). Corolla actinomorphic, ephemeral; petals with an orbicular to obovate limb and a long, narrow claw, free, cohering mutually or by the staminodes. Stamens mostly 3 fertile epipetalous inserted on the petals and 3 alternating staminodes, staminodes rarely absent, or all stamens fertile; filaments short; anthers basifix, dehiscing lengthwise extrorsely. Ovary superior, sessile to stipitate (in Australian spp. sometimes with 3 hard swellings at the top), 1- or 3-celled, or incompletely 3-celled. Placentas parietal, central, or basal, with ~ ovules; styles filiform, apex 3-fid, stigmas mostly capitate. Fruit shape similar to that of the ovary but larger, loculicidally 3-valved. Seeds ellipsoid to obovoid, often ribbed, with a long funicle. Distr. Xyridaceae are confined to the tropics throughout the world including the southern parts of North America; east of Malaysia and Australia hitherto only recorded from the Patau group (Korror) and New Caledonia.

Beschreibung: Monoecious, mostly deciduous shrubs or trees with perular buds. Pith triangular in section. Innovations often resiniferous. Leaves simple, (in Mal. spp.) spiral, penninerved, crenate or dentate, rarely entire, mostly with domatia in the nerveaxils beneath, in bud mostly folded along the midrib and plicate, often glandularlepidote beneath. Stipules caducous. Catkins unisexual, at least the pendent ♂ ones in terminal panicles above the ♀ ones, the latter mostly in stiff, axillary, poor racemes or terminal on short-shoots.—♂ Flowers in triads, each sustained by a bract. Perianth segments 4 (or less by abortion), mostly connate at the base. Stamens 4, epitepalous; filaments short; anthers glabrous, 2-celled; cells parallel, dehiscing lengthwise. No rudiment of ♀.—♀ Flowers in diads sustained by a bract concrescent with 4 bracteoles, accrescent and woody in fruit, densely packed and imbricate. Perianth 0. Ovary 2-celled, each cell with one anatropous, pendent ovule attached near the apex of the cell; styles 2, free, short, cylindric. Fruiting catkins cone-like. Nut small, compressed, 1-seeded, mostly winged and crowned by the styles. Seed without endosperm; embryo straight; cotyledons flat; testa membranous; embryo straight; endosperm 0; cotyledons flat. Distr. About 20 spp. mainly on the N. hemisphere except in the New World, mostly extra-tropical, in SE. Asia southward to Bengal, N. Assam, Tonkin, and Formosa, in Malaysia only cultivated.

Beschreibung: The Flora Malesiana is not preceded by a general key enabling one to identify any unknown native or wild plant to the family or genus to which it belongs. This is certainly a serious lack and presents a formidable handicap to inexperienced taxonomists in rapid naming current collections. However, there are several forcing arguments for omitting—at present—such an attempt which in itself would present no facile task, and could be accomplished only by a taxonomist thoroughly acquainted with the Malaysian flora. One could of course use some world key as a basis and cut out the entries leading to genera or families not represented in the Malaysian flora, but this procedure would be unsatisfactory, specially as these world keys make little use of vegetative characters; the latter appear to me very important specially in the earlier forks of the keys.

Beschreibung: Annual or perennial, unarmed or spinous, bitter herbs or undershrubs, often glandular-hairy. Stem terete, farctate, with a peripheral whorl of air-vessels. Leaves spread, simple, entire, exstipulate. Flowers ♀, actinomorphic, solitary, opposite or between the leaves, or by stunting of the leaves, more or less arranged in a racemiform or paniculiform inflorescence, distinctly pedicelled, lilac blue. Calyx persistent, 5-partite to near the base, segments lanceolate, imbricate in bud, after anthesis not or hardly accrescent. Corolla gamopetalous, deeply 5-partite; limb rotate; segments imbricate in bud, oval, obtuse. Stamens 5, free, inserted in the throat of the corolla, alternating with the segments; filaments filiform from a broadened base, glabrous or papillate; anthers 2-celled, bifid at the base and apex, opening lengthwise. Disk absent. Ovary superior, 2- (rarely 3-, very rarely more-) celled; placentas adnate to the dissepiment, spongy, entire or in cross-section bifid; styles 2 (rarely 3 or more), free; stigmas capitate-clavate. Ovules ~. Capsule globose or ellipsoid, loculicid, or both loculicid and septicid, 2(rarely more)-valved, or bursting irregularly. Seeds ~, very small, longitudinally ribbed; endosperm small, straight. Distr. Species ± 20, in the tropics of both hemispheres; in Malaysia 2, of which one indigenous, the other introduced and naturalized in Java.

Beschreibung: Rhizomes (rarely spiny) producing annual, mostly twining shoots, in Malaysia twining either to the right (fig. 4c) or the left (fig. 4a). Stems consisting of a main stem and sterile branches, both bearing leafless flowering axes. Leaves petiolate, generally cordate, simple and entire or palmately lobed, or palmately compound, except in the latter triplinerved; apex generally glandular, developed before the blade (forerunner tip); blade usually glandular on the lower side chiefly towards the base. Flowers hermaphrodite or dioecious, ♀ with staminodes, ♂ without even a rudimentary ovary, actinomorphic, 3-merous, mostly inconspicuous and greenish, ♂ often massed together and scented. Tepals in two whorls of 3. Stamens in 2 whorls of 3, the inner sometimes sterile; anthers usually introrse. Torus an urceolate, perianthoid chamber in Stenomeris, a saucer or cup in many spp. of Dioscorea, fleshy in Dioscorea § Enantiophyllum, in some spp. enlarged into a cone making the stamens appear to be connate. Style 1 with 3 bifid stigmas. Ovary 3-locular, inferior, sometimes separated from the perianth by a constriction. Ovules 2 in each cell or ~ (in Stenomeris), anatropous. Fruit a capsule, but it breaks up rather than dehisces in Trichopus. Seeds winged or wingless (in Trichopus); endosperm horny, embryo in a marginal pocket. Distr. Ca 9 genera and about 600 spp. (Dioscorea large, the other genera small or monotypic). Pantropic with considerable extensions into temperate regions. The Stenomerideae and Trichopodeae are restricted to the warm humid regions where Nepenthes grows and their geologic history must have been that of Nepenthes: they may be regarded as the survivors of the hermaphrodite ancestry of the Dioscoreeae.

Beschreibung: Herbs or shrubs, sometimes parasitic, usually with twining stems, occasionally prostrate or creeping, or erect, very rarely trees, often with milky juice. Leaves mostly spirally arranged, in parasitic species absent or nearly so, usually petioled; petiole sometimes with extra-floral nectaries. Stipules absent, pseudostipules (leaves of axillary shoot) rarely present. Inflorescences mostly cymose, one- to many-flowered, with mostly opposite or subopposite bracts at the base of the cymes or under the solitary flowers; rarely racemose. Flowers generally hermaphrodite, actinomorphic, rarely slightly zygomorphic, usually 5-merous, rarely 4-merous, various in size and colour, often showy. Sepals usually free, imbricate, with quincuncial aestivation, often persistent, sometimes accrescent in fruit. Corolla sympetalous, of various shapes, often funnel-shaped or campanulate, more rarely rotate, salver-shaped or urceolate; the limb nearly entire or more or less deeply lobed, often contorted-plicate in bud, or valvate or induplicate-valvate. Stamens isomerous, alternating with the corolla-lobes, adnate to the corolla, with usually slender, often filiform filaments and introrse or laterally and longitudinally dehiscing anthers. Pollen smooth or spinulose. Disk mostly present, annular or cupular. Ovary superior, mostly of 2 carpels, 2- or 1-celled, sometimes 4-celled by development of accessory partitions, rarely of 3 carpels and 3-celled; ovules 2 in each carpel, sessile, erect, anatropous. Style 1, often filiform, simple or forked, or 2 free styles, rarely very short or absent. Stigma entire or 2-lobed, rarely 3-lobed, or stigmas 2-4, of various shape, globular or ellipsoid to filiform, sometimes applanate, rarely peltate, kidney-shaped, conical or funnel-shaped. Fruit a capsule dehiscing by valves or circumscissile or irregularly dehiscing, rarely a berry or nut-like. Seeds as many as ovules or fewer; endosperm cartilaginous; cotyledons generally folded, sometimes obscure or absent. Distr. Ca 55 genera, with ca 1650 spp., widely distributed in the tropical, subtropical and temperate regions of both hemispheres; the greater part of the species in the tropics and subtropics of America and Asia. The larger genera Cuscuta (ca 165 spp.), Convolvulus (ca 250 spp.) and Ipomoea (ca 500 spp.) nearly throughout the range of the family but Convolvulus more in the temperate parts and Ipomoea more in the tropics and subtropics. Other large genera as Evolvulus (ca 100 spp.) and Jacquemontia (ca 120 spp.) nearly confined to America. Argyreia (ca 90 spp.) confined to tropical Asia. Malaysia, and a single sp. in Australia, and Merremia (ca 80 spp.) circumtropical. Several monotypic or small genera in E. Africa, Madagascar, and Australia.

Beschreibung: Erect or straggling herbs, shrubs or trees, sometimes monoecious or dioecious, the herbs sometimes rhizomatous; branches sometimes jointed at the nodes, sometimes without vessels ( Sarcandra). Leaves simple, decussate or sometimes whorled in fours, serrate, crenate or dentate, the teeth often thickened at the apex, penninerved, usually petiolate; petioles more or less connected at the base at least by a transverse line or connate into a distinct sheath; in Ascarina often alternating with leafless internodes which have the petiolar sheath; stipules minute to fairly conspicuous, subulate, borne on the petiole bases or sheath, occasionally pectinate. Flowers much reduced, without perianth, fully unisexual or essentially bisexual with the reduced anther-bearing organ adnate to the side of the ovary; arranged in spicate, paniculate, or capitate axillary or terminal inflorescences. — Male flowers bracteate or not, apparently consisting of 1—5 stamens, or in Hedyosmum consisting of numerous anthers in a cone-like structure; if 3 then the whole forming a fused 3-lobed organ sometimes enveloping the female flower by its edges, the central anther with 2 or aborted loculi and the laterals with single loculi, simply lobed or with connectives slightly to considerably produced so that the whole organ is 3-fingered; if with only 2 anther locelli then these on either side of a thickened filament plus connective. — Female flowers naked or enclosed by a cupular bract, the perianth adnate to the ovary, often minutely or shortly dentate at the apex and the ovary thus inferior; ovary 1-locular; stigma sessile or style short; truncate, 2-lipped, depressed or subcapitate (or horseshoe-shaped in one species), rarely linear or clavate. Ovule solitary, orthotropous, pendulous, bitegmic and crassinucellate. Drupes fleshy, small, ovoid or globose, sometimes more or less 3-sided in Hedyosmum, free or united into a mass by the bracts; endocarp hardened and crustaceous. Seeds subglobose, exarillate, with copious fleshy or oily endosperm and minute embryo, the cotyledons divaricate or scarcely formed. Distribution. Four genera with about 80 species. Since VESTER’S (1940) small-scale map the family (Ascarina) has been found in Madagascar. It is mainly tropical but Ascarina extends south to North Island of New Zealand (fig. 6) and Chloranthus and Sarcandra extend north to Japan, China, Korea and the eastern U.S.S.R. (Ussuri).

Beschreibung: It is not without some pride and much satisfaction that the present volume, fourth planned in the series, second in sequence of publication, is brought to a successful end. Satisfaction I feel through the fact that the scheme and aim of this work is not only understood by the scientific-botanical world, but has also been accepted in the administrative world: Notwithstanding the long term scope of the work, the High Government of the Republic of Indonesia, having realized the essential value of basic scientific work in the natural sciences for the welfare of the future generations of its young nation, has been instrumental in authorizing the Director of Kebun Raya Indonesia (Botanic Gardens of Indonesia, Bogor) to create a Flora Malesiana Foundation. Sponsored by the Indonesian Government, this Foundation knits together the work and interest of the Herbarium Bogoriense of Kebun Raya Indonesia and the Netherlands Rijksherbarium at Leyden, the direction of which have officially agreed to a long-range close co-operation.

Beschreibung: Halophobous, aquatic or palustrial perennial herbs, rooting in the mud or freefloating. Stem erect or floating, solid, with numerous air-chambers as are the petioles. Leaves rosulate or alternate, or solitary at the top of the stem, emersed, floating or submerged, broad or narrow, curvinerved (when emersed); petioles sheathing at the base. Flowers ♀, ephemerous, mostly in racemiform, spiciform, subumbelliform or paniculiform inflorescences which are subtended by 1-2 spathelike or tubular leaf-sheaths, rarely solitary or pairwise in the leaf-axils. Bracts minute or absent. Flowers often simultaneously or centrifugally expanding. Perianth choriphyllous or gamophyllous, 6-merous, actinomorphic or zygomorphic, blue or lilac, rarely yellow, after anthesis marcescent and tightly including the ovary or the fruit. Stamens 6 or 3, rarely 1, on the base, in the tube or in the throat of the perianth, often unequal; filaments free; anthers 2-celled, cells bursting lengthwise, rarely opening by pores. Ovary superior, sessile, 3-celled, with axile placentas or 1-celled with 3 parietal or with 1 apical placenta. Ovules numerous or 1 and then pendulous from the apex of the cell. Style 1; stigma entire or minutely 3-lobed. Fruit a 3-valved capsule or indehiscent. Seed(s) longitudinally ribbed. Embryo central, terete, straight, hardly shorter than the copious, mealy endosperm. Distr. About 8 small genera and ± 25 species, 6 genera confined to the New World, one in Madagascar, one widely distributed in the Old World; in Malaysia one native genus, one introduced and abundantly naturalized, and one occasionally cultivated as an ornamental.

Beschreibung: Trees or shrubs (or rarely suffrutices outside Malesia). Leaves simple, alternate, often coriaceous, glabrous or with an indumentum on undersurface, margin entire; petioles often with 2 lateral glands. Stipules 2, minute and caducous to large and persistent, usually linear-lanceolate. Inflorescence racemose, paniculate or cymose; flowers bracteate and usually bibracteolate; bracts and bracteoles small and caducous or larger and enclosing flower or groups of flowers and persistent. Flowers actinomorphic to zygomorphic, hermaphrodite or rarely polygamous, markedly perigynous. Receptacle campanulate to cylindrical or rarely flattened cupuliforum, often gibbous at base; calyx lobes 5, imbricate, often unequal, erect or reflexed. Petals 5 (absent in some Neotropical species), inserted on margin of disk, commonly unequal, imbricate, deciduous, rarely clawed. Stamens indefinite, 2—60 (to 300 in Neotropics), inserted on margin of the disk, in a complete circle or unilateral, all fertile or some without anthers and often reduced to small tooth-like staminodes; filaments filiform, free or ligulately connate, short and included to long and far exserted; anthers small, 2-locular, longitudinally dehiscent, glabrous or rarely pubescent. Ovary basically of three carpels but usually with only one developed, the other two aborted or vestigial, variously attached to (the base, middle or mouth of) receptacle, usually sessile or with short gynophore, pubescent or villous; ovary unilocular with two ovules or bilocular with one ovule in each locule. Ovules erect, with micropyle at base (epitropous). Style filiform, basally attached; stigma 3-lobed or truncate. Fruit a fleshy or dry drupe of varied size, interior often densely hairy; endocarp much varied, thick or thin, fibrous or bony, often with a special mechanism for seedling escape. Seed erect, exalbuminous, the testa membraneous; cotyledons amygdaloid, plano-convex, fleshy, sometimes ruminate. Germination hypogeal with the first leaves opposite or alternate or epigeal with opposite first leaves. An extensive review of the generic limits of the family has been published: G.T. PRANCE & F. WHITE, The genera of Chrysobalanaceae: a study in practical and theoretical taxonomy and its relevance to evolutionary biology, Phil. Trans. Roy. Soc. London 320 (1988) 1—184. This contains full details of taxonomic history, morphology, anatomy, pollen, ecology and distribution of the family. A condensed version of these subjects is given here. Details of the Neotropical members of the family are given in: G.T. PRANCE, Chrysobalanaceae, Flora Neotropica 9 (1972) 1—410. The African members of the family were treated in: F. WHITE, The taxonomy, ecology and chorology of African Chrysobalanaceae (excluding Acioa), Bull. Jard. Bot. Nat. Belg. 46 (1976) 265—350.

Beschreibung: Erect tall annual, usually branched. Leaves simple, with 2 free stipules, in the lower part of the stem opposite, in the higher part spirally arranged, long-petioled, palmate, 3—11-foliolate. Flowers (♂) (♀) or mostly (♂♀). Male flowers in short, dense cymes, which are united into lax, foliate, terminal panicles, very shortly pedicelled. Tepals 5, free, oblong, membranous, imbricate. Stamens 5, epitepalous; filaments erect and short in bud, linear, with a narrowed apex; anthers comparatively large, basifixed, 2-celled, cells opening longitudinally, rudimentary ovary absent. Female flowers solitary in the axil of a small, primary, membranous, entire bract closely enveloping the ovary, each enveloped by a spathaceous, conspicuous, acuminate, secondary bract. Perianth absent. Ovary sessile, 1-celled; style central; stigmas 2, sessile, long, filiform, caducous. Ovule solitary, pendulous. Achene closely enveloped by the much enlarged, secondary bract, broadly oval, with a concave rimmed base, much compressed, faintly keeled on the lateral margins; pericarp smooth, hard, crustaceous, easily splitting into two halves; albumen unilateral, scanty, fleshy; embryo large, horseshoe-shaped; cotyledons large; radicle long. Distr. Monotypic, native of Central Asia, cultivated in tropical Asia, naturalized in N. America.

Beschreibung: Trees or shrubs. Leaves spirally arranged or often distichous, sometimes crowded towards the top of the branchlets, simple, entire or crenate or serrate, crenations mostly glandular; petioles often thickened at the base and (or) the apex. Stipules small, rarely large and foliaceous, often early caducous, or wanting. Inflorescences subterminal or mostly axillary, sometimes on the old wood, in often spike-like racemes or in panicles or in short cymes, but sometimes condensed to glomerules or reduced to few-flowered fascicles or even to a solitary flower, apparently essentially cymose. Flowers bisexual or unisexual, in the latter case mostly dioecious, sometimes polygamous, actinomorphic, 3- to polymerous, cyclic (with sepals and petals) or rarely spiral (with perianth segments, in trib. Oncobeae). Pedicels often articulated near the base. Sepals 3-6, rarely more, mostly persistent, sometimes accrescent, imbricate or valvate, free or connate at the base into a calyx-tube, or calyx closed in bud and disrupting in different ways. Petals 3-8, rarely more, free, imbricate or valvate, mostly alternating with the sepals and caducous, sometimes persistent and accrescent, often inserted on the margin of a hypogynous or nearly perigynous disk, or absent. Receptacle often deepened in the centre, mostly with appendages such as an extrastaminal disk or disk-lobes, free glands between the stamens, or a corona of 5 phalanges, each of which is consisting of fine barbate threads, or staminode-like scales inserted on the inner side of the base of the petals, or with true, mostly barbate, staminodes. Stamens 5 to ~, hypogynous, mostly free, rarely the filaments connate into a column; anthers with 2, longitudinally dehiscent cells; connective sometimes with a short appendage. Ovary mostly free, rarely semi-inferior, unilocular with (2-)3-5(-8) parietal placentas, sometimes incompletely 2—8-celled by the deeply protruding placentas; ovules 2 to numerous, anatropous. Styles 1-10, free or connate; stigma sessile. Fruit a fleshy or dry berry or a capsule, rarely a drupe, 1- to many-seeded. Seeds sometimes arillate, with abundant endosperm; embryo straight; cotyledons mostly broad, foliaceous. Distr. About 84 genera with c. 1300 spp., nearly all woody, predominantly in the tropics, rapidly decreasing in number towards the subtropics, 2 genera with a total of 9 (mostly Chilean) spp. in the temperate zones of S. America.

Beschreibung: Small trees or erect shrubs. Leaves spirally arranged, simple, petioled, entire, palmatinerved, densely red-dotted. Stipules small, very caducous. Flowers in terminal corymbs or panicles, actinomorphic, ♀, rather large. Pedicel with 5-6 apical glands. Sepals 4-5, free, imbricate in bud, falling off as soon as the flower expands. Petals 4-7, free, imbricate in bud. Stamens numerous, inserted on an annular hypogynous disk; filaments thin, free; anthers horseshoe-shaped, passing over the top of the filament and with both ends closely applied to i , 2-celled; cells opening in the middle (on the top of the filament) by short slits which unite into a spuriously apical pore. Ovary superior, usually bristly, 1-celled, with 2 opposite parietal slightly intruding placentas. Style 1, sinuous, rather thick; stigma 2-dentate. Ovules very numerous. Capsule compressed contrary to the placentas, usually softly prickly, rarely smooth, loculicidally bivalved; endocarp membranous, separating from the valves. Seeds numerous, obovoid, angular; testa fleshy, very densely studded with small, round, red, sessile glands; albumen well-developed, not oil-containing; embryo rather large. Distr. Monotypic, native and cultivated in tropical America; cultivated in many other tropical countries.

Beschreibung: Annual (?)laticiferous herbs, with the habit of Phytolacca. Stem erect, somewhat succulent. Leaves spirally arranged, simple, entire, exstipulate. Inflorescences terminal, densely spicate, acropetal. Flowers subtended by a bract and two bracteoles, bisexual, actinomorphic. Calyx tube adnate to the ovary; segments 5, united below, imbricate, connivent, persistent. Corolla campanulate-urceolate, perigynous; lobes 5, imbricate. Stamens 5, epipetalous, alternating with the corolla lobes; filaments short; anthers rounded, 2-locular, dehiscing longitudinally. Ovary semi-inferior, 2-locular; style short, stigma capitate; ovules attached to large spongy stipitate axile placentas. Capsule cuneate-obconic, 2-locular, membranous, circumscissile; seeds ~, minute, oblong, rugose-costate, albumen very scanty or none (?); embryo axile, straight, subterete. Distr. Mono-generic, almost pantropical.

Beschreibung: Submerged, rootless, monoecious freshwater plants. Leaves verticillate, 2-4 times forked, segments linear dentate. Flowers actinomorphic, solitary, axillary, unisexual. Perianth valvate, segments 9-12, persistent, narrow. ♂: stamens 8-24; anthers nearly sessile rather broad, connective pointed, the 2 cells mostly crowned by a minute bristle; ovary rudiment absent. ♀: ovary superior, sessile, 1-celled with 1 ovule; style persistent, subulate, sulcate towards the apex; stamen rudiments absent. Fruit oblong, compressed, warty, not dehiscent, near the base with 2 straight or curved soft spines, or unarmed. Distr. Ca 2 spp., both ubiquitous.

Beschreibung: Annual or perennial, saprophytic or autotrophic herbs; the saprophytic species often colourless. Leaves usually spread or alternate, entire, simple, without stipules; non-saprophytic species with a radical rosette of linear leaves; stem leaves often reduced to small scales; sometimes the basal part of the stem provided with many decurrent, grass-like leaves. Flowers ♀♂, usually actinomorphic, solitary or in capitate or cymose inflorescences. Perianth corolline; limb consisting of 2 whorls; tube sometimes 3-winged. Anthers 3, subsessile in the perianth throat and dehiscing laterally with horizontal slits,or 6, hanging down in the perianth tube and dehiscing with longitudinal slits. Connective large, often appendiculate. Style filiform or shortly cylindrical or conical. Stigmas 3, sometimes connate. Ovary inferior, 1-celled with parietal placentation, or 3-celled with axile placentation. Ovules ~, anatropous, with 2 integuments; funicles often rather long. Fruit usually capsular, sometimes fleshy, crowned by the persistent perianth tube and the style, or by a thickened persistent basal ring of the perianth tube, dehiscing irregularly or with transverse slits at the top. Seeds ~, small, subglobose to linear, sometimes with loose, reticulate testa, with endosperm. Distr. About 125 species, widely distributed in the tropics of both hemispheres, also in subtropical America, Chicago area, Moçambique, Southern China, Japan, Southern Australia, New Zealand and Tasmania. As many species are rare, it is possible that only a part of their area is known. Most of them are found in moist regions. Among the autotrophic Malaysian Burmanniaceae there are 3 rather common species which are widely spread, viz Burmannia coelestis, B. disticha and B. longifolia. The latter two are absent from Java and the Lesser Sunda Islands, the former occurs in Java proper only in its western part. Of the saprophytic Malaysian species only 3 have been often collected, viz Burmannia championii, B. lutescens, and Gymnosiphon affinis.

Beschreibung: In the absence of a complete bibliography of the botanical literature of Malaysia, comparable to those on Eastern Asia and the Pacific by MERRILL & WALKER, and as the ‘Flora Malesiana’ will not to be completed within the near future, the need was felt to have at hand a concise, selected bibliography of existing revisions and other phytographical publications temporarily providing taxonomists with a reference to what is roughly available for the identification of Malaysian collections. When the ‘Flora Malesiana’ is completed, after some decades, this bibliography should no longer be required, as the references contained in it will all have been accounted for in the Flora itself in one way or another. In the meantime, however, a list arranged by families seems to serve a very practical purpose, as it gives access to the main body of accumulated knowledge precursory to the final revisions in the Flora.

Beschreibung: Trees, shrubs or lianas, rarely subherbaceous. Glands (in Mal. spp.) often present on the leaf-bases or petioles, and in lower marginal crenations. Indumentum of simple hairs, glandular hairs or multicellular hairs secreting calcium oxalate and forming scales, or present beneath the cuticle making the surface of the leaf minutely verruculose and sometimes pellucid-punctate. Leaves opposite, verticillate, spiral, or alternate, petioled (rarely sessile), exstipulate, simple, almost always entire. Flowers ♀♂ ♀♂ or ♀♂ and ♂ in the same inflorescences, usually protogynous, usually actinomorphic, rarely slightly zygomorphic, in axillary or extra-axillary elongated or subcapitate spikes or racemes or in terminal and sometimes axillary panicles. Receptacle (calyx-tube) usually in two distinct parts, the lower receptacle surrounding and adnate to the inferior ovary and the upper receptacle produced beyond to form a short or long tube terminating in the calyx-lobes, the latter sometimes poorly developed. Calyx-lobes 4 or 5 (rarely 6-8) or almost absent, sometimes accrescent ( Calycopteris). Petals 4 or 5 or absent, conspicuous or sometimes very small, inserted near the mouth of the upper receptacle. Stamens usually twice as many as the petals, borne inside the upper receptacle usually in two series, exserted or included; anthers dorsifixed, usually versatile (or rarely adnate to the filaments). Disk intrastaminal, usually present, hairy or glabrous. Style usually free (attached for part of its length to the upper receptacle in Quisqualis). Ovary inferior (semi-inferior in the West-African genus Strephonema), unilocular, with usually 2 (sometimes 2-6) pendulous, anatropous ovules of which only 1 usually developes. Fruit (botanically a pseudocarp) very variable in size and shape, fleshy or dry, usually indehiscent, often variously winged or ridged, 1-seeded. Albumen absent. Distr. 18 genera with c. 450 spp. in the tropics and subtropics: 2 are circumtropical ( Combretum and Terminalia), and are much the largest genera, 1 is confined to North Australia and Queensland (Macropteranthes), 2 confined to tropical Asia ( Finetia and Calycopteris) , 3 occur in Asia and Africa (Anogeissus, Lumnitzera, and Quisqualis), 1 is confined to Madagascar (Calopyxis), 3 are confined to tropical Africa (Guiera, Pteleopsis and Strephonema), 2 occur in tropical Africa and tropical America (Conocarpus and Laguncularia) and the remaining four ( Buchenavia, Bucida, Ramatuela and Thiloa) are confined to tropical and subtropical America.

Beschreibung: For various reasons the space occupied by pre-Linnean Malaysian phytography in this concise history seems too large and out of proportion in comparison to the survey of post-Linnean work. Modern plant description, though based on, and derived from, ancient beginnings and traditions, maintains but slender contacts with plant sciences earlier than the 18th century and it might claim to be allotted by far the larger space on account of its superior results, its greatly increased efficiency, its Consciousness of limitations and capabilities, its output, and its clearness of purpose. There exists, however, during the last decade, an increasing interest in the nearly forgotten botany of centuries long past, not only because of a certain taste for the quaint and attractive flavour of scientific efforts from minds so remote from our own, but also on account of a growing insight into the hidden springs of modern thought and method, which flow deeply, emerge unexpectedly, and appear to rise from distant roots. There is also, in connexion with this, the absorbing spectacle of discovery and of growth i.e. the development of a field of human culture that has bound devoted and excellent personalities in its service from the first glimmerings of our civilization.

Beschreibung: Perennial herbs, more commonly woody at the base, undershrubs or shrubs, erect, scrambling or scandent, sometimes high lianas. Rhizome not rarely tuberous. Branches often slightly swollen and jointed at nodes. Hairs simple, uni- or multicellular, short ones often with a hooked apex. Leaves simple, spiral or alternate, petioled (without an abscission zone), exstipulate; midrib usually prominent beneath, elevated or flat above; nervation commonly palmate, or pinnate, nerves often obliquely extending towards the margin. Flowers bisexual, actinomorphic or zygomorphic, solitary, fasciculate, or in axillary or cauligerous, racemose, paniculate or cymose inflorescences, usually only one or two flowers open at a time; bracts present and often persistent; pedicel often hardly distinct from the ovary. Calyx petaloid, gamosepalous, 3- (or 6-) lobed or 1-lipped; lobes valvate or induplicate. Petals (in Mal.) absent. Disk (?) 0, rarely present (e.g. a few Thottea spp.). Stamens 6 (4 or 5 in some extra-Mal. Aristolochia spp.) or 6—c. 36 (—46), in 1 whorl or in 2 (3 or 4) whorls (Thottea); filaments free or slightly mutually united at the base, and/or almost completely adnate to the style column to form a gynostemium; anthers free (Thottea) or dorsally united with the style column (Aristolochia), each consisting of 2 thecae with 4 pollen sacs, extrorse, rarely introrse (extra-Mal. spp.), dehiscing longitudinally. Ovary inferior (rarely half-inferior in extra-Mal. genera), 4—6-carpellate, 4—6-celled, syncarpous (or ± apocarpous in extra-Mal. Saruma); placentae parietal (distinct when young, then intruding and connivent axially, thus often seemingly axile); ovules usually many, anatropous, in 1 or 2 vertical rows in each locule of the ovary, horizontal or pendulous; style-column 3—many-lobed, sometimes some of the lobes redivided; stigmas or stigmatic tissue apical, lateral, or on the surface of style lobes. Fruits capsular or siliquiform (follicular or cocci in extra- Mal. genera), 4—6-celled; dehiscing apically towards the base (basipetal, e.g. Thottea) or basally towards the apex (acropetal, e.g. most Aristolochia); septicidal, rarely septifragal (some extra-Mai. Aristolochia) or bursting irregularly (extra-Mal. Asarum); rarely indehiscent (W. African Pararistolochia). Seeds many in each locule (1-seeded in extra-Mal. Euglypha), often coated with remains of placental tissue (membranous when dry), horizontal or pendulous, variously shaped; ovate, deltoid or triangular, flat, convex-concave, or longitudinally curved, or oblong (and triangular in cross-section), rugose, finely verrucose, or smooth, immarginate (Thottea; Aristolochia, p.p.) or winged (Aristolochia, p.p.); albumen fleshy, copious; embryo minute, cotyledons two, distinct. Distribution. There are 7 genera, Aristolochia worldwide, Asarum over the northern hemisphere, Thottea in continental Southeast Asia and Malesia, Pararistolochia in tropical Africa, and 3 monotypic genera, viz. Saruma in China, Holostylis and Euglypha in South America. As to number of species, Aristolochia is by far the largest with some 300 spp., largely concentrated in the New World, especially in Central and South America, in Malesia with 28 spp.; Asarum (incl Hexastylis and Heterotropa) with possibly some 70 spp. in northern temperate regions, Thottea with 26 spp., of which 22 in Malesia, and Pararistolochia with 12 spp. in West Africa.

Beschreibung: Scandent shrubs (often erect in youth), without resin; branches sympodial with a series of circinate woody hooks in one plane. Leaves spread, simple, entire, often rosette-crowded, cuneiform, penninervous, reticulate-veined, glabrous, both surfaces minutely pitted, each pit with a peltate small hair secreting a waxlike substance; petiole articulated, scar on the twigs often saddle-shaped; stipules absent. Flowers ♀♂, actinomorphic small. Inflor. few or several times dichotomous or spike-like, often provided with said hooks and single reduced bract-like leaves, branches often recurved. Pedicels articulated. Bracts with a glandular-thickened base, margin fimbriate-membranous. Calyx tube short, at length adnate to the base of the ovary; lobes 5 inequal imbricate, enlarged and wing-like in fruit. Petals 5, united at the base, slightly contorted in bud. Stamens mostly 10, rarely 5, the episepalous slightly longer. Filaments with broadened base; anthers basifixed, ± introrse to ± latrorse, 2-celled, opening lengthwise. Ovary for the greater part inferior, consisting of 3 carpels, 1-celled, protruding into a nippleshaped elongation bearing 3 articulated erect styles with a punctiform or horseshoe-shaped stigmatic apex; nipple enlarging in fruit. Ovule 1, basal, ascending, with 2 integuments. Nut not dehiscent, crowned by the enlarged calyx. Seed roundish with testa intruding between the cerebral-like folds of the endosperm. Exocarp leathery. Embryo straight, erect, obliquely placed; cotyledons diverging; hypocotyl rather thick. Distr. Disjunct, ca 3 spp. in trop. W. Africa, and 9 in SE. Asia, from the Deccan to Burma, Indochina, Hainan, S. China, the Malay Peninsula, Borneo and Sumatra (cf. fig. 2).

Beschreibung: Floating aquatic herbs with dimorphic leaves, submerged ones opposite pinnatifid rootlike, apical ones in a rosette, rhomboid, dentate, with spongy often inflated petiole, arranged in leaf-mosaic; stipules 4-8, minute. Flowers bisexual, small, solitary, axillary, short-pedicelled, 4-merous, white or lilac. Petals imbricate. Disk present. Ovary half-inferior with 1 style and 2-4 persistent sepals turning often to thorns or horns. Fruit mostly 1-celled, 1-seeded, shell bone-hard; thorns after withering often set with barbs at the apex. Seed often producing 2-5 free germ-stalks. Distr. Several species in the Old World, but not known from Australia.

Beschreibung: Priority of publication is internationally accepted as the basic principle of the ‘Rules of Botanical Nomenclature’. This has emphasized to a marked degree the importance of determining accurately the exact time when novelties are placed before the scientific public.

Beschreibung: Dioecious trees or shrubs. Leaves simple, scattered. Stipules O. Flowers unisexual, often in heads, in the axils of a bract and with 2 bracteoles. ♂: in axillary heads or short racemes; calyx entire or 5-toothed; petals 5, imbricate, often small, alternate with the calyx; stamens 8-16 in 2 alternating whorls; anthers small, dorsifixed with lateral lengthwise slits; disk pulvinate; style rudimentary. ♀: solitary, axillary or in 2-10-flowered heads; ovary inferior, 1-locular, connate with the 5-toothed or entire calyx; petals 5-8 often minute; stamens of inner whorl partly sterile, both petals and anthers soon dropping; style with 2 appressed later divergent often torulose branches stigmatose on their inside, brittle, often deficient in the herbarium. Ovule 1, hanging from the apex of the cell, anatropous with 2 integuments. Fruit drupaceous ovoid to oblong. Distr. Ca 6 spp., 4 in Atlantic N. America, 1 in China, 1 from India to W. Malaysia.

Beschreibung: Trees, rarely shrubs. Leaves simple, mostly glandular-punctate, exstipulate. Flowers ♀, actinomorphic, 5-merous. Calyx-tube short, tube (and usually segments) densely setulose-hairy within. Corolla represented by 7-40 deltoid to linear-subulate processes, rarely by a low entire annulus. Stamens 8-80; filaments free, short, slender; anthers hippocrepiform. Disk 0. Ovary (2-)3-5(-8)-locular; cells with one anatropous ovule pendulous from the apex. Style elongate, filiform, sometimes accompanied by ‘parastyles’ at the base; stigma small, capitate. Fruit a thick-walled, woody, dehiscent, 1—5-seeded capsule, or a thin-walled, (?) indehiscent, 1—2-seeded capsule. Seeds large, without chalazal fold, usually with aril. Endosperm 0. Distr. Almost confined to Malaysia, occurring in all parts of the archipelago except E. Java and the Lesser Sunda Isl.; found also in the Nicobar, Solomon and Fiji Islands. Genera 3. The greatest number of species is concentrated in Borneo, with apparently a marked inner centre of differentiation in the western part of the island. Fig. 1.

Beschreibung: Type material of Tulasnella cystidiophora Höhn. & Litsch. has been studied. The species is characterized by often moniliform gloeocystidia and clamp-less hyphae (at least in the subhymenium).

Beschreibung: The part certain lime-secreting marine algae play in the building of coral reefs and in the formation of banks was discussed chiefly at the end of the last and in the beginnig of this century. At that time it was already known that extensive parts of the sublittoral zone of the Arctic sea were covered by a luxuriant growth of Lithothamnion species. Kjellman states in 1883 (p. 96) that along the northern coast of Norway Lithothamnion soriferum “covers large spaces of the bottom in great masses”, and that off the shores of Spitsbergen and Nova Zembla in 10 to 20 fathoms of water Lithothamnion glaciale “covers the bottom in deep layers for several miles, and altogether determines the general aspect of the vegetation wherever it occurs”, whereas Lithothamnion norvegicum is said to form banks on the coasts of Iceland and of Greenland. Rosenvinge (1893, p. 772) reports that Lithothamnion ungeri forms banks on the coast of Iceland and of Greenland.

Beschreibung: There is much confusion about, the identity of the above mentioned aroid genera, the typification of which is still unsatisfactory.

Beschreibung: Most classifications of the genera of the Gramineae have been on the structure and arrangement of their spikelets, for these organs provide a far greater variety of readily distinguishing characters than do other parts of the grass plant. Nevertheless it has not always been possible to decide from morphological studies alone whether marked similarities in structure point to a close affinity or are merely examples of parallel development. The modern taxonomist, endeavouring to arrange the grass genera in as natural a sequence as possible in order to emphasise relationships and evolutionary trends, sooner or later meets with difficulties in this respect, for examples of parallelism are of common occurrence in this family. He is more fortunate, however, than his predecessors, in that his own intensive morphological studies, based on a wider range of specimens, may be supplemented by additional data gleaned from the ecological, anatomical and cytological researches of contemporary workers. Thus aided by the more complete information at his disposal, it has been possible for him to rearrange certain groups, particularly the Festuceae and Hordeeae, in which parallel development has occasionally led to unrelated genera such as Lolium, Agropyron and Nardus, being too closely associated. In the following account an attempt has been made to provide a more natural classification for about eighteen species frequently referred to the genus Lepturus R. Br. by reason of their similar spicate inflorescences.

Beschreibung: var. bullatus, nov. var. — Ramuli novelli ferrugineo-velutini. Folia 1—3-juga; foliola bullata. Folliculi inconspicue rostrati. Typus: Kostermans 4928 (fl., fr.), E. Borneo, Sangkulirang island, alt. 20 m (Holotype in L; Isotypes in BO, K).

Beschreibung: Urelytrum Henrardii Chippindall sp. nov.; ab U. agropyroidei Hack., cui e descriptione affine, culmis gracilibus, foliorum laminis non hirsutis, longe attenuatis, longioribus, racemis flavido-viridibus, spicularum sessilium gluma inferiore 5-nervi, arista breviore distinguendum — Fig. 1. Gramen perenne caespitosum, usque ad 92 cm altum. Culmi erecti, simplices, graciles, pauci-nodes, glabri, racemos versus asperuli. Folia plerumque basalia; vaginae internodiis longiores, sublaxae, striatae, apicem versus carinatae, basales glabrae laevesque, superiores pilis patulis laxe pilosae, ore villoso-barbatae; ligulae scariosae, rotundato-obtusae, 0.8—1.25 mm longae; laminae lineares, apice tenuiter setaceae, planae vel leviter conduplicatae, usque ad 38 cm longae, 3—3.8 mm latae, marginibus scabridis, costis asperulis, pone ligulam pilis longis exceptis glabrae. Racemi ad culmi apicem solitarii, stricti, fragiles, subcylindrici, fere glabri, flavidi vel pallide flavido-virides, saltem 16 cm longi; articuli rhacheos compressi, infimo usque ad 2 cm longo, scaberuli, margine uno superne rigide ciliati, appendice membranacea inaequaliter dentata ciliolata; pedicelli articulis similes, sed appendice minore. Spiculae sessiles biflorae, anguste lanceolato-oblongae, 7.5—8.2 mm longae (callo excluso); callus crassus, rotundato-obtusus, basi barbatus. Glumae subaequales, minute punctatae; inferior spiculam aequans, coriacea, marginibus hyalinis, explanata lanceolata, subconvexa, subacuta, 5-nervis, dorso apicem versus parce spinuloso-ciliata, superne bicarnata, carinis angustissime alatis, alis spinuloso-ciliatis; superior inferiore paulo brevior, firme membranacea, marginibus hyalinis apice minute ciliolata, lanceolata, acuta, 3-nervis, superne carinata, carina anguste alata, ala spinuloso-ciliata. Anthoecium inferum ♂: lemma tenuiter hyalinum, lanceolato-ovatum, 6—6.5 mm longum, 2-nerve, minute bidentatum, marginibus apicem versus minute ciliolatum; palea lemmati similis sed angustior et paulo longior; antherae 3 mm longae; lodiculae glabrae. Anthoecium superum ♀: lemma lemmati anthoecii inferi simile sed 3-nerve, apice latius; palea angustior. Spiculae pedicellatae illis sessilibus absimiles, neutrae, ad glumas lemmaque redactae, sine arista 2—2.75 mm longae. Glumae coriaceae, marginibus hyalinis superne ciliolatae, minute punctatae; inferior spiculae aequilonga, lanceolata, 5-nervis, ad carinam superne angustissime alata, ala spinulosociliata, in aristam scabridam 9—12.5 mm longam excurrente; superior inferiore paulo longior, apice integra, obtusa, superne carinata, carina anguste alata, ala spinuloso-ciliata, obscure 5-nervis. Lemma tenuiter hyalinum, parvum.

Beschreibung: According to general opinion the spikelets of Oryza consist, reckoned from their base upwards, of 2 sterile glumes, called hereafter I and II, one fertile glume (valvula inferior; lemma), called hereafter III, and the palea valvula superior) to this glume, called hereafter p3. The spikelets are placed singly on the very short ultimate branchlets, called hereafter pedicels, of a more or less strongly ramose panicle; the tips of the pedicels are broadened into a shallow infra-spicular cup, either distinctly 2-lobed or not; from the bottom of the cup arises a minute knob, on which the very distinct basal callus of the spikelet is jointed. When ripe, the spikelets of the wild species fall off as a whole, disarticulating at the joint (in dried specimens often long before maturity; hence in herbarium-specimens they are frequently lacking). In many cultivated forms they remain firmly attached to their pedicels, a property of very high economic value. The spikelets are strongly laterally compressed. I and II are either 1-nerved or nerveless; as a rule they are many times shorter than the spikelet, sometimes even very minute. Only in O. Ridleyi they are comparatively well-developed, reaching about half the length of the spikelet, but very narrow. III is very rigid, usually conspicuously granulate, boatshaped, keeled, either awned or not, 5-nerved, with a strong midrib; it has the ultimate lateral nerves along the margins. P3 is likewise boatshaped, shortly cuspidate or not, with a narrow, rather rounded, less often faintly keeled back, 3-nerved; it is about as long as III, awn disregarded, and has the same rigid granulate structure, excepted the narrowly incurved thinly membranaceous smooth marginal parts (hidden by III). It might be taken for a fertile glume, but this view is inadmissible because of the averted position of the lodicules. It has a rather thin mid-nerve and strong lateral nerves, separating the rigid central part from the membranaceous borders. The well-developed lodicules are glabrous; the six stamens are free; there are 2 free shortish styles with large plumose white or violet stigmas which, during anthesis, stick out from the sides of the spikelet in or below its middle. The ripe fruit is oblong or lanceolate, usually angular; it is free from glume and palea but remains firmly incarcerated between them.

Beschreibung: Dactyloctenium Henrardianum Bor spec. nov. quae ab omnibus aliis speciebus hujus generis inflorescentia racemosa haud digitata satis recedit. An annual grass. Culms slender, 10—30 cm tall, erect, smooth, glabrous, striate in robust specimens, terete, long-exserted from the uppermost leaf-sheath. Leaf-sheaths strongly keeled, loose, slipping from the culm, much shorter than the internode and leaf-blade, markedly striate, smooth and glabrous except for some bristles from bulbous bases sparsely arranged near the margins in the upper fourth; ligule a lacerate membrane not more than 2 mm long. Leaf-blades up to 10 cm long by 5 mm wide at the base, gradually narrowed into a fine point from the rounded base, very scabrid on the margins which also bear long bulbous-based bristles in the lower third; upper surface smooth; lower surface often with bulbous-based bristles; midrib strongly marked with 2—3 prominent parallel veins on either side.

Beschreibung: My colleague Lam, in whose honour this volume is composed, has made it very easy for contributors to find a subject in a field in which he has worked himself. His versatile interest nearly covered the whole of the taxonomy and phylogeny of vascular plants, subjects in theoretical biology, plant morphology, plant geography, and plant ecology. In the latter section his “Fragmenta papuana” contains an inspiring picture of tropical vegetation in correlation with environment. I have pleasure on this occasion in offering some considerations in the field of plant ecology. The subject which I have chosen deals with the way of reasoning when interpreting a correlation found to exist between vegetation and environment. I have not infrequently traced a deficiency in such interpretation and I feel a need of discussing this point which is, in my opinion, a matter of vital importance.

Beschreibung: Although Clarke saw the type of Scirpus erectus Poir. in the Paris Herbarium he misapplied the name to a quite different species occurring in Madagascar, S. and E. Asia, and tropical Australia. Herein he was followed by Ridley, Merrill, Backer, and others. It has now generally been accepted that the correct name of this species is Scirpus juncoides Roxb. and that the name Scirpus erectus Poir. does not belong to its synonymy. After having examined the type of S. erectus I am convinced that the question was admirably cleared up by Chermezon (see Arch. Bot. 4, 1931, 26, and also in Humbert, Fl. Madag., fam. 29, 1937, 149). Scirpus erectus is much nearer to the European S. supinus L. than to S. juncoides Roxb. It differs from S. supinus by the larger spikelets, the larger, more distinctly mucronate glumes, the bristly appendage of the connective, the bifid style, and the larger, biconvex, only faintly wavyridged, elliptic or suborbicular nuts. It is an African species extending from the Mediterranean region through tropical Africa to Madagascar and Mauritius. There can be no doubt that Isolepis uninodis Delile is conspecific with Scirpus erectus Poir. Delile’s description is very accurate: “épis cylindriques, ovoïdes-lanceolés ... écailles ovales, aiguës ... deux stigmates ... graine lenticulaire, transversalement rugueux vers les bords.” The differences with Scirpus supinus are clearly indicated: “ses graines [du S. supinus] sont ovoïdes-cunéiformes, trigones, ridées transversalement sur toute leur surface; ses styles sont trifides.” Moreover, Delile’s excellent figure leaves no doubt whatever on the identity of his species.

Beschreibung: In the development of the various scientific disciplines certain contacts have been established between neighbouring sciences, while other endeavour still proceeds almost on its own. The study of the influence of the environment on chemical reactions has given us a large part of physical chemistry, while the inverse, the study of the influence of chemical reactions on the (natural) environment has been, in the last decades, developed as geochemistry. Much of our physiology and ecology deals with the influence of the environment upon the organism, while the inverse, the influence of the organism upon the natural environment (geobiology) has hardly been studied systematically. This influence is great, as already realized by Pasteur a century ago. Moreover, this study completes the picture of the relations of “give and take” between the organism and its milieu and the author believes that a true ecology should concern itself with the mutual relations between organism and environment rather than view the landscape (or seascape) entirely from the biological point of view. Blumea twice before has been so kind as to accept ecological notes from the author of this paper. Because of the interest Professor Lam has always taken in this ecological approach, the author hopes that this short note will not be a complete dissonant in the, chiefly taxonomic, matter contained in this volume. By a study of the influence of various groups of organisms on the estuarine environment Baas Becking and Wood (1) arrived at the conclusion that, primarily, the following groups are of geochemical importance; sulphate-reducing bacteria, thiobacteria, purple bacteria, iron bacteria and algae. The limits of the potential algal environment surpass the regions of most other groups, the thiobacteria excepted. The range of conditions in which algae may occur, be it temperature, salinity, pH or electrode potential is very wide indeed. Algae occur in all aqueous environments; evaporate, geothermal, soft and hard freshwater, estuarine and marine water and also in buffered and non-buffered products of pyrite oxidation (Baas Becking, Wood and Kaplan, 4). The author recently isolated, from mirabilite at Mawson Base, Antarctica, a green polyblepharid while Kaplan (12) describes the occurrence of bluegreens at 86°C in the hotsprings of the Rotarua district, N—Z. From this same region Kaplan described a Navicula at pH 1.2, while several algae occur in a solution of trona (sodium carbonate-bicarbonate) at pH 10.55. The above anecdotical remarks only serve to illustrate the extent of the algal potential milieu.

Beschreibung: The name Arundo Bambos L. Sp. Pl. 81, 1753, is interpreted as properly belonging to the common thorny bamboo of India; therefore this species should be called Bambusa Bambos (L.) Voss. Arundo Bambos L. Sp. Pl. ed. 2, 120, 1762, insofar as it is represented by Linnaeus’ specimen labeled “1. Bambos” and by his description of this specimen, is based on a misidentification of a Chinese species: Bambusa flexuosa Munro (1868). Bambos arundinacea Retz. Obs. Bot. 5:24, 1789, is shown to have been based on the plant known today as Bambusa vulgaris Schrad. ex Wendl. (Coll. Pl. 2:26, pl. 47, 1810), and not on the common thorny bamboo of India, properly called Bambusa Bambos (L.) Voss. Bambusa arundinacea Willd. Sp. Pl. 2:245, 1799, is based on Bambos arundinacea Retz., but Willdenow is shown to have confused, in his text, as in his mind, at least two species under this name: 1. The plant which has since come to be known as Bambusa vulgaris Schrad. (of which he had a specimen labeled “B. arundinacea 1.”) and 2. The common thorny bamboo of India (properly called Bambusa Bambos [L.] Voss) of which he had no specimen. Traditional usage for 150 years has overlooked the facts in this case, and has erroneously applied Bambusa arundinacea Willd., and Bambusa arundinacea Retz. (as Bambos) to the common thorny bamboo of India. As a result of the long-continued misapplication of the name Bambos arundinacea Retz. and its variants, it will be exceedingly difficult to reïnvest the name with its original meaning. It may come to pass that consensus of leadership will be to avoid the use of the name Bambos arundinacea Retz and its variants altogether, at least for some time, because of the risk of being misunderstood, and to continue the use of the name Bambusa vulgaris Schrad., which is generally accepted in its proper sense. Those who use Bambusa arundinacea Retz. (as Bambos) or any of the other variants of the name, may be able to avoid being misunderstood by citing Bambusa vulgaris Schrad. as a synonym. Bambusa Schreb. Gen. Pl. 1:236, 1789, and Bambos Retz. Obs. Bot. 5:24, 1789, are synonymous, and are believed to have been based on the same species, namely the plant commonly known today as Bambusa vulgaris Schrad. Strict adherence to Recommendations IV and V of the fifth edition of the International Rules of Botanical Nomenclature, and probably the claims of priority, would indicate the replacement of Bambusa Schreb. by Bambos Retz. The continuation of the use of the generic name Bambusa Schreb., instead of Bambos Retz., has the sanction of tradition, and of contemporary preference; but in order to be fully justified and stabilized, this usage should be regularized and legalized by action of the International Botanical Congress, placing Bambusa Schreb. on the list of Nomina Conservanda. The genus Leleba Rumph. ex Nakai, Jour. Jap. Bot. 9: 9 et seq. 1933, is added to the recognized synonymy of Bambusa Schreb.

Beschreibung: On the 13th of October 1940 I found in the vicinity of a wool- and skinwork in Tilburg (The Netherlands, prov. N. Brabant) a sterile grasstuft, striking me by its peculiar habit. I transplanted it into my garden in Dordrecht and there it was flowering for the first time in June 1941, and in July it was collected to be dried. On the 4th of July 1941 I gathered one more fructifying specimen at the same locality in Tilburg. Doubtless the plant was a Deschampsia and my provisory identification was D. media R. et Sch.. Sending the material with this name to Dr P. Jansen in Amsterdam I got his reply: ”Certainly not D. media. It is a species, unknown to me or, more probably, a variety of D. flexuosa“. This conclusion, however, seemed unacceptable to me. The habit of the sterile as well as the fertile plant differs strongly from that of D. flexuosa. The tuft is denser and harder, with thicker and shorter leaves. The panicle is longer, wider and more diffuse, the branchlets less flexuous, the culms are relatively short, as long as the panicle or at most 1½—2 times the length of the panicle (in D. flexuosa 4—5 times). The characteristics of the flower are decisive. The lower glume is 5 mm long, the upper one 6 mm, both of them overtop the lemma and palea of the enclosed flower (in D. flexuosa the glumes are little different in length and equaling or overtopped by the flowers). The stipe of the upper flower, remaining attached to the lower one, when the spikelet falls asunder, is densily pencilshapedly hirsute and 1.5 mm long (in D. flexuosa 0.6—0.8 mm). The upper flower bears a similar stipe of a fully rudimental third flower, in other words: the rachilla is produced behind the upper palea as a hairy bristle. These properties sooner recall D. setacea than D. flexuosa, but the anthers are very small, 0.3—0.5 mm long, on much longer filaments (D. setacea has anthers, 1.5 mm long, filaments 0.5 mm, D. flexuosa: anthers 1.8 mm, filaments very short). All this: the habit, the pale green spikelets without any touch of purple, brown or blue, and the small anthers on long filaments justifies a specific differentiation of the Tilburgian wooladventive. I propose to name it, in honour of Dr J. Th. Henrard, whom I owe so much in the field of adventives in general and of Gramineae in particular: Deschampsia Henrardii nov. spec.

Beschreibung: A new species, Alstonia undulifolia Kochummen & Wong, is described from the Malay Peninsula. Two sections of the genus occur in the Malay Peninsula, Alstonia sect. Monuraspermum Mon. and Alstonia sect. Alstonia, the latter being the correct name for what was previously known as sect. Pala (Adr. Juss.) Benth. Various characteristics, including growth architecture, are examined for their usefulness in distinguishing these two sections of the genus. In comparing A. angustiloba Miq. and A. pneumatophora Berger, both of which have not been properly differentiated by characteristics of the reproductive organs, A. pneumatophora var. petiolata Mon. is reduced to synonymy under A. angustiloba. A key to the seven species of Alstonia native to the Malay Peninsula is provided.

Beschreibung: Revision of the Malesian species of the genus Steganthera, which centres in New Guinea; precursor to treatment in Flora Malesiana. There are 16 species accepted; 5 are described as new, 12 names are reduced, 3 are excluded and 9 are imperfectly known.

Beschreibung: In a recent thesis B.S. Fey (Zürich) has developed a new theory about the origin of the cupule in Fagaceae. He has concluded that the appendages (spines, lamellae, etc.) on the outside of the cupule are regularly arranged and that they reflect a condensation (concrescence) of a dichasial flower system, so that cupule and fruit(s) form together the representation of one ancestral inflorescence; the cupular appendages would then largely represent the bracts of the ancestral inflorescence. This stands in contrast with former opinions, in which the cupule was interpreted as of separate vegetative origin from the nut(s) which was (were) the remain (s) of the inflorescence.

Beschreibung: Recent studies in Sabah and Sarawak have demonstrated the presence of an undescribed species of Podocarpus.

Beschreibung: Pholidota kinabaluensis is transferred to the new monotypic genus Entomophobia. Coelogyne phaiostele, C. ridleyana, and Pholidota triloba are identical and transferred to the new genus Geesinkorchis, that also comprises the new species G. alaticallosa. The monotypic genus Sigmatochilus is reduced to Chelonistele, in which C. dentifera and C. lurida var. grandiflora are described as new. Chelonistele crassifolia is regarded as a variety of C. sulphurea.

Beschreibung: The genera Hunteria and Lepiniopsis of the family Apocynaceae are in Malesia represented by one species each. Distribution and ecology are cited in full.

Beschreibung: Five new species of Rafflesia (Rafflesiaceae) are described, while attention is drawn to a sixth, possibly also new one. A key to all recognized species is given.

Beschreibung: The morphology and leaf anatomy of Myxopyrum is described and a key to the species is given. Of the 15 species previously described four species and two subspecies are recognised: M. nervosum Bl. (synonyms M. horsfieldii, M. zippelii) with one subspecies coriaceum (Bl.) Kiew (synonym M. ellipticum), M. ovatum Hill (synonyms M. macrolobum, M. cordatum, M. philippinensis), M. pierrei Gagnep. (synonym M. hainanense) and M. smilacifolium Bl. (synonym M. serrulatum) with one subspecies confertum (Kerr) Kiew. Myxopyrum enerve Steen. is Chionanthus enerve (Steen.) Kiew. Descriptions for the extra-Malesian species, M. smilacifolium, is given.

Beschreibung: In subgenus Malachobatus twenty Malesian species are recognized, one of them ( Rubus moluccanus L.) with four varieties. Synonymy, descriptions, habitat notes, etc. are given. New names: R. moluccanus L. var. discolor (Bl.) Kalkm. and var. angulosus Kalkm. A key is given to the Malesian species.

Beschreibung: During a recent treatment of the Proteaceae for “Flora Malesiana” it has become evident that a revision of the generic status of all proteaceous taxa reported from S. Asia and Malaysia as well as from the adjacent regions of Micronesia, Melanesia, Polynesia and subtropical-tropical Australia had to be made to reach a satisfactory correlation of the genera and species concerned as a basis for the discussion of phytogeographical relations both within and outside the proper Malaysian area. During this work it appeared necessary to transfer some species to other genera. A revision of the genus Helicia showed that a group of species had to be segregated as a distinct new genus Heliciopsis. My studies are based on herbarium specimens borrowed from the following Institutions: Arnold Arboretum (A), Bot. Mus. Berlin-Dahlem (B, where the type-material of the family remained intact), Bogor (BO), Brisbane (BRI), Calcutta (CAL), Edinburgh (E), Florence (FT), Kepong (KEP), Lae (LAE), Leiden (L), Melbourne (MEL), Miinchen (M), New York (NY), Manila (PNH), Singapore (SING), Stockholm (S) and Utrecht (IT). The material preserved in the British Museum (BM), at Kew (K), and Paris (P) has been studied during a stay at London and Paris.

Beschreibung: For identifying the mosses collected in different localities of the Malaysian region, the need was felt for a key to the genera. In the preliminary one that I constructed to this end the genera were taken in the delimitation accepted in the second edition of Brotherus, Natürl. Pflanzf. In addition to the latter the genera published after 1925 and therefore not included in Brotherus 1. c. are taken into account. In revising the families for Flora Malesiana I will doubtless be compelled to alter the position of some of the species and the delimitation of some of the genera, and at the end of series III of Flora Malesiana, which will contain the Mosses, I therefore intend to give a final key. I sincerely hope that the preliminary key will in the meantime have been tested by different bryologists, and that they will let me profit by their remarks. For this reason it is published here. The analytical key is based as far as possible on vegetative characters, especially on the shape of the leaf cells. The principal features of the sporophyte are noted, but are not, as a rule, made use of as alternatives. This applies particularly to those alternatives that lead to the main groups. Only when no reliable vegetative characters could be found, have characters of the sporophyte, especially those of the peristome, been used. The habitat of each genus, not its distribution in the Malaysian region, is indicated in the key.

Beschreibung: In Southeast Asia (excluding India) 44 taxa are recognized, 39 species, of which four are newly described ( I. kerrii, I. luzoniensis, I. emmae, and one unnamed species A, which will be treated by Nguyen Van Thuan, Paris), four subspecies, one of which is new (I. sootepensis subsp. acutifolia) and three are new combinations ( (I. suffruticosa subsp. guatemalensis, I. trifoliata subsp. unifoliata, I. trita subsp. scabra) ), and one variety which is a new combination I. spicata var. siamensis). A key, descriptions and full synonymy are given as well as 4 distribution maps and 5 figures.

Beschreibung: Mapania holttumii Kern, nom. nov. — Mapania insignis Holttum, Gard. Bull. Sing. 11, 1947, 293, non Sandwith, Kew Bull. 1933, 496. When publishing the name Mapania insignis for a species occurring in the Malay Peninsula, Holttum overlooked the existence of the earlier homonym Mapania insignis Sandw. for a different species from British Guyana. I therefore propose to replace the illegitimate binomial by that of Mapania holttumii.

Beschreibung: Some time ago a paper was published by Brouwer (1954) on the geological interpretation of a vertical section through Pleistocene deposits near the village of Wezep in the northeastern corner of the province of Gelderland, in the central part of the Netherlands. The exact locality¹, known locally as the ""Moordenaarshoek"" (murderer’s corner), is situated on a heath along the edge of a pine forest, which grows on the lower slopes of the northeastern tip of the great Pleistocene pressure ridge or moraine along the western border of the valley of the river Yssel. The locality was described for the first time by Bursch, Florschütz and van der Vlerk (1938) who made an excavation there on the instigation of the late Colonel Mallinckrodt, an ardent amateur archaeologist. It was believed that the numerous flint objects which could be found in a peculiar kind of boulderclay there and at other localities in the neighbourhood, might be interpreted as artefacts of a primitive, somewhat Clactonian, type; a belief which was also accepted by the present author until some five years ago. With the object of collecting more data on these somewhat enigmatic phenomena², an excavation at approximately the same locality as that described by Bursch, Florschütz and van der Vlerk was made in the autumn of 1948 by the Biologisch-Archaeologisch Instituut of Groningen University (director Professor A. E. van Giffen) under supervision of Dr. A. Bohmers and the present author. Thanks to the energetic help of the late Colonel Malunckrodt two other localities, situated more to the West, on the Oldebroek artillery range, were also excavated. Here again the same flint objects, interspersed in either gravel and coarse sands, or in a loam comparable to that found at Wezep, were recovered. The layer of loam at one of the two Oldebroek localities was found to be much thinner than that encountered at Wezep, although not differing from it in any other aspect.