ALBERT

Description: Dark limestones in the old quarries at Leny, Perthshire contain sparse beds with tiny fossils. They are poorly preserved and, though barely affected by the Ordovician Grampian Event tectonism, there is some taphonomic distortion and many are corroded along stylolitised horizons. The fauna mainly comprises trilobites of two types, open-ocean miomerids and polymerid shelf dwellers. MiomeridsCondylopygecf.eliandKiskinella cristataindicate a stratigraphical position equivalent to the base of the paradoxidid Amgan Stage of Siberia; traditionally regarded as ‘Middle Cambrian’. However, the bulk of the Leny miomerids, notably species ofPagetides, are forms described from the outer edge of Laurentia, within theBonnia–OlenellusZone, where it is considered to be ‘Lower Cambrian’. The Leny polymerids were likely transported off-shelf and some are conspecific with taxa in the Laurentian allochthonous Quebec and New York successions of the Early–Middle Ordovician (Taconic) Appalachian Orogen. The Leny Limestone and Shale Member of the Keltie Water Grit Formation is part of the Dalradian Supergroup deposited in an off-shelf Caledonide Grampian Terrane of the Humber Tectonostratigraphical Zone, midway between the North American successions and the Greenland Caledonides.Additional to the trilobites, brachiopods, sponges, hyoliths, bradoriids and a selection of indeterminable organic fragments occur; none of which has any particular age significance.

Description: Holocene sea-level changes affected people living in the Pacific Islands and their ancestors along the western Pacific Rim. Sea-level changes, particularly those that were rapid, may have led to profound and enduring societal/lifestyle changes. Examples are given of (1) how a rapid sea-level rise (CRE-3) about 7600 BP could ultimately have led to the earliest significant cross-ocean movements of people from the western Pacific Rim into the islands; (2) how mid to late Holocene sea-level changes gradually created coastal environments on Pacific Islands that were highly attractive to human settlers; (3) a hypothesis that rapid sea-level fall during the ‘AD 1300 Event' brought about widespread disruption to trajectories of cultural evolution throughout the Pacific Islands; and (4) the effects of recent and likely future sea-level rise on Pacific Island peoples.

Description: The potential of igneous quartz for providing a better understanding of magmatic processes is demonstrated by studying late-Hercynian rhyolites and granites from central and western Europe. Cathodoluminescence (CL) reveals growth patterns and alteration structures within igneous quartz reflecting the magma crystallisation history. The relatively stable and blue-dominant CL of zoned phenocrysts is principally related to variations in the Ti concentration, which is a function of the crystallisation temperature. The Al/Ti ratio of igneous quartz increases with progressive magma differentiation, as Ti is more compatible, compared to Al, Li, K, Ge, B, Fe, P during magma evolution. The red-dominant CL of the anhedral groundmass quartz in granite is unstable during electron bombardment and associated with OH- and H2O-bearing lattice defects. Thus, CL properties of quartz are different for rocks formed from H2O-poor and H2O-rich melts. Both groundmass and phenocrysts in granites are rich in alteration structures as a result of interaction with deuteric fluids during cooling, whereas phenocrysts in extrusive rocks do not usually contain such structures. The combined study of trace elements along with the analysis of quartz textures and melt inclusion inventories may reveal detailed PTX-paths of granite magmas. This study shows that quartz is a sensitive indicator for physico-chemical changes during the evolution of silicarich magmas. Common growth textures show a wide variety in quartz phenocrysts in rhyolites and some granites. This paper presents a classification of textures, which formed as a result of heterogeneous intra-granular lattice defects and impurities. The alternation of growth and resorption microtextures reflects stepwise adiabatic and non-adiabatic magma ascent, temporary storage of magma in reservoirs and mixing with more mafic, hotter magma. The anhedral groundmass quartz overgrowing early-magmatic phenocrysts in granites is free of growth zoning.

Description: This paper describes three mid-Tertiary intrusions from the Henry Mountains (Utah, USA) that were assembled from amalgamation of multiple horizontal sheet-like magma pulses in the absence of regional deformation. The three-dimensional intrusion geometries are exceptionally well preserved and include: (1) a highly lobate sill; (2) a laccolith; and (3) a bysmalith (a cylindrical, fault-bounded, piston-like laccolith). Individual intrusive sheets are recognised on the margins of the bodies by stacked lobate contacts, and within the intrusions by both intercalated sedimentary wallrock and formation of solid-state fabrics. Finally, conduits feeding these intrusions were mostly sub-horizontal and pipe-like, as determined by both direct observation and modelling of geophysical data.%The intrusion geometries, in aggregate, are interpreted to reflect the time evolution of an idealised upper crustal pluton. These intrusions initiate as sills, evolve into laccoliths, and eventually become piston-like bysmaliths. The emplacement of multiple magma sheets was rapid and pulsed; the largest intrusion was assembled in less than 100 years. The magmatic fabrics are interpreted as recording the internal flow of the sheets preserved by fast cooling rates in the upper crust. Because there are multiple magma sheets, fabrics may vary vertically as different sheets are traversed. These bodies provide unambiguous evidence that some intrusions are emplaced in multiple pulses, and that igneous assembly can be highly heterogeneous in both space and time. The features diagnostic of pulsed assembly observed in these small intrusions can be easily destroyed in larger plutons, particularly in tectonically active regions.

Description: The anisotropy of magnetic susceptibility (AMS) is widely and routinely used to measure the preferred orientations of Fe-rich minerals in undeformed and weakly deformed granite plutons. The interpretation of the mapped AMS fabrics depends on rock-textural observations, on the map patterns of the fabrics in plutons, and on comparisons of the pluton fabrics to tectonic structures in the country rocks. The AMS may document emplacement-flow related fabrics, but the emplacement fabrics may be reworked or completely overprinted by rather weak tectonic strains of the magma mush or the cooling pluton, especially in syntectonic intrusions. The Late Devonian Canso granite pluton is an excellent example of overprinting of emplacement fabrics by weak tectonic strains. The Canso pluton was emplaced ca. 370 Ma along the boundary between the Meguma and Avalon tectonic terranes, in the northern Appalachian orogen. The AMS was mapped along two traverses that cross the pluton and that are perpendicular to the terrane boundary. Textural evidence suggests the rocks underwent very modest post-full crystallisation strains. The AMS records the dextral transcurrent shearing that occurred on the terrane boundary during emplacement and cooling of the Canso pluton, supporting interpretations that weakly deformed syntectonic granites can be used as indicators of regional bulk kinematics. AMS fabrics in Late Devonian granites of the Meguma Terrane suggest partitioning of the non-coaxial shearing into the terrane bounding fault, with pure-shear dominated deformation further from the fault. Numerical simulations suggest that the kinematics recorded by the fabrics in the Canso pluton was simple-shear, or transpression or transpression with small components of pure shear oriented perpendicular to the bounding shear zone.

Description: A comparative study of three main igneous rock associations that plot in the K2O–SiO2 diagram shoshonite field: shoshonite series absarokites–shoshonites–banakites (henceforth referred to as shoshonites s.l.), vaugnerites, and potassic lamprophyres, reveals that similarities between the associations are superficial. Vaugnerites and lamprophyres are more magnesian, richer in large ion lithophile and high field strength elements and have higher light rare earth/heavy rare earth ratios than shoshonites. Furthermore, shoshonites have low radiogenic heat production, typical of subduction-related rocks, but most vaugnerites and some lamprophyres are highly radioactive. Relative to bulk-Earth, shoshonites have depleted, asthenospheric mantle-like Sr and Nd isotope signatures, whereas vaugnerites and potassic lamprophyres have enriched, crust or lithospheric mantle-like compositions. Though vaugnerites and some lamprophyres show evidence of crustal contamination, the contaminated magma was not originally shoshonitic. Their composition is consistent with derivation from a metasomatised upper mantle source enriched long before melting, thus precluding an active subduction setting. In conclusion, the term shoshonite, implying late-stage arc magmas, cannot be applied to a rock series simply because it plots into the K2O–SiO2 diagram shoshonite field. Shoshonites with a subduction-related source may, however, be identified by discriminant function analysis.

Description: India–Asia collision resulted in crustal thickening and shortening, metamorphism and partial melting along the 2200 km-long Himalayan range. In the core of the Greater Himalaya, widespread in situ partial melting in sillimanite+K-feldspar gneisses resulted in formation of migmatites and Ms+Bt+Grt+Tur±Crd±Sil leucogranites, mainly by muscovite dehydration melting. Melting occurred at shallow depths (4–6 kbar; 15–20 km depth) in the middle crust, but not in the lower crust. 87Sr/86Sr ratios of leucogranites are very high (0·74–0·79) and heterogeneous, indicating a 100 crustal protolith. Melts were sourced from fertile muscovite-bearing pelites and quartzo-feldspathic gneisses of the Neo-Proterozoic Haimanta–Cheka Formations. Melting was induced through a combination of thermal relaxation due to crustal thickening and from high internal heat production rates within the Proterozoic source rocks in the middle crust. Himalayan granites have highly radiogenic Pb isotopes and extremely high uranium concentrations. Little or no heat was derived either from the mantle or from shear heating along thrust faults. Mid-crustal melting triggered southward ductile extrusion (channel flow) of a mid-crustal layer bounded by a crustal-scale thrust fault and shear zone (Main Central Thrust; MCT) along the base, and a low-angle ductile shear zone and normal fault (South Tibetan Detachment; STD) along the top. Multi-system thermochronology (U–Pb, Sm–Nd, 40Ar–39Ar and fission track dating) show that partial melting spanned ̃24–15 Ma and triggered mid-crustal flow between the simultaneously active shear zones of the MCT and STD. Granite melting was restricted in both time (Early Miocene) and space (middle crust) along the entire length of the Himalaya. Melts were channelled up via hydraulic fracturing into sheeted sill complexes from the underthrust Indian plate source beneath southern Tibet, and intruded for up to 100 km parallel to the foliation in the host sillimanite gneisses. Crystallisation of the leucogranites was immediately followed by rapid exhumation, cooling and enhanced erosion during the Early–Middle Miocene.

Description: :Finite difference numerical simulations were used to characterise the rates of diffusion-controlled dissolution and growth of zircon in melts of granitic composition under geologically realistic conditions. The simulations incorporated known solubility and Zr diffusivity relationships for melts containing 3 wt% dissolved H2O and were carried out in both one and thre dimensions under conditions of constant temperature, linearly time-dependent temperature and for a variety of host system thermal histories. The rate of zircon dissolution at constant temperature depends systematically on time (t½−12;), temperature (exp T−1) and degree of undersaturation of the melt with respect to zircon (in ppm Zr). Linear dissolution and growth rates fall in the range 10−19 10−15 cm s−1 at temperatures of 650-850°C. Radial rates are strongly dependent on crystal size (varying in inverse proportion to the radius, r): for r〉30 μm, dissolution and growth rates fall between 10−17 and 10−13 cm s−1. During crustal magmatism, the chances of survival for relict cores of protolith zircons depend on several factors, the most important of which are: the initial radius of the zircon; the intensity and duration of the magmatic event; and the volume of the local melt reservoir with which the zircon interacts. In general, only the largest protolith zircons (〉120 μm radius) are likely to survive magmatic events exceeding 850°C. Conversely, only the smallest zircons (

Description: :Ore element ratios in intrusion-related mineralisation are in part a function of the relative oxidation state and degree of fractionation of the associated granite suite. A continuum from Cu-Au through W to Mo dominated mineralisation related to progressively more fractionated, oxidised I-type magmas can be traced within single suites and supersuites. Such systematic relationships provide strong evidence for the magmatic source of ore elements in granite-related mineral deposits and for the production of the observed ore element ratios dominantly through magmatic processes. The distribution of mineralised intrusive suites can be used to define a series of igneous metallogenic provinces in eastern Australia. In general, there is a correlated evolution in the observed metallogeny (as modelled based on the compatibility of ore elements during fractionation) with increasing degree of chemical evolution of the associated magmatic suite. This is from Cu-Au associated with chemically relatively unevolved magmas, through to Sn and Mo-rich mineralisation associated with highly evolved magmas that had undergone fractional crystallisation. Provinces recognised in that way do not necessarily correlate with the tectonostratigraphic boundaries defined by the near-surface geology, indicating that the areal distribution of some granite source regions in the deep crust is unrelated to upper crustal geology.

Description: :Most granitic batholiths contain plutons which are composed of low-variance mineral assemblages amenable to quantification of the P– conditions that characterise emplacement. Some mineral thermometers, such as those based on two feldspars or two Fe–Ti oxides, commonly undergo subsolidus re-equilibration. Others are more robust, including hornblende–plagioclase, hornblende–clinopyroxene, pyroxene–ilmenite, pyroxene–biotite, garnet–hornblende, muscovite-biotite and garnet–biotite. The quality of their calibration is variable and a major challenge resides in the large range of liquidus to solidus crystallisation temperatures that are incompletely preserved in mineral profiles. Further, the addition of components that affect Kd relations between non-ideal solutions remains inadequately understood. Estimation of solidus and near-solidus conditions derived from exchange thermometry often yield results 〉700°C and above that expected for crystallisation in the presence of an H2O-rich volatile phase. These results suggest that the assumption of crystallisation on an H2O-saturated solidus may not be an accurate characterisation of some granitic rocks.Vapour undersaturation and volatile phase composition dramatically affect solidus temperatures. Equilibria including hypersthene–biotite–sanidine–quartz, fayalite–sanidine–biotite, and annite–sanidine–magnetite (ASM) allow estimation of Estimates by the latter assemblage, however, are highly dependent on . Oxygen fugacity varies widely (from two or more log units below the QFM buffer to a few log units below the HM buffer) and can have a strong affect on mafic phase composition. Ilmenite–magnetite, quartz–ulvospinel–ilmenite–fayalite (QUILF), annite–sanidine–magnetite, biotite–almandine–muscovite–magnetite (BAMM), and titanite–magnetite–quartz (TMQ) are equilibria providing a basis for the calculation of .Granite barometry plays a critical part in constraining tectonic history. Metaluminous granites offer a range of barometers including ferrosilite–fayalite–quartz, garnet–plagioclase–hornblende–quartz and Al-in-hornblende. The latter barometer remains at the developmental stage, but has potential when the effects of temperature are considered. Likewise, peraluminous granites often contain mineral assemblages that enable pressure determinations, including garnet–biotite–muscovite–plagioclase and muscovite–biotite–alkali–feldspar–quartz. Limiting pressures can be obtained from the presence of magmatic epidote and, for low-Ca pegmatites or aplites, the presence of subsolvus versus hypersolvus alkali feldspars.As with all barometers, the influence of temperature, , and choice of activity model are critical factors. Foremost is the fact that batholiths are not static features. Mineral compositions imperfectly record conditions acquired during ascent and over a range of temperature and pressure and great care must be taken in properly quantifying intensive parameters.

Description: :The Cretaceous granitic rocks and associated regional metamorphic rocks in SW Japan were formed by a Cordilleran-type orogeny. Southwest Japan is regarded as a hypothetical cross-section of the upper to middle crust of the Eurasian continental margin in the Cretaceous, comprising (1) high-level granitoids (called San-yo type) and weakly to unmetamorphosed accretionary complexes that are exposed on the back-arc side and (2) low-level (Ryoke type) granitoids with high-grade metamorphites up to migmatitic gneisses on the forearc side. All these granitoids are of the ilmenite series, and predominantly I-type, with a subordinate amount of garnet- or muscovite-bearing varieties in the Ryoke zone, but none of these contains cordierite. These mineralogical variations are likely to depend more on their slightly peraluminous chemistry rather than the pressure differences during crystallisation.In the eastern part of SW Japan, the granitoids of both levels give K–Ar biotite ages of approximately 65 Ma, whereas the magmatic age of high-level granitoids is approximately 70 Ma, 15 Ma younger than the nearly 85 Ma old lower level granitoids. This implies that the formation of the middle crust started approximately 15 Ma before that of the upper crust. The middle crust material was kept over 500°C for 15–20 Ma after solidification, then it cooled together with the upper crust to 300°C, 6–7 Ma after the formation of the upper crust. The coincidence of cooling history below 500°C of the upper and middle crust may reflect the regional uplift of the crust.The low-level granitoids have higher 87Sr/86Sr initial ratios than those of high-level granitoids in the middle-western part (Chugoku district), but the relationship appears to be opposite in the eastern part. This may imply that the two plutonic series formed by separate magmatic pulses at an interval of c. 15 Ma, even though they are not independent, but rather part of a larger episode of crustal growth.

Description: :Granite is the final product of the high-temperature, magmatic, predominantly endogenic, chemical differentiation of the earth. Our understanding of the origin and evolution of granitoid rocks comes from a combination of direct observation, analogue experimentation and numerical modelling. A brief historical overview shows an exceptional level of such research activity over the last 50 years. The number and complexity of questions have resulted in both an absolute and a relative growth of the science since the plate tectonic revolution, largely consisting of refining the current magmatic paradigm within its overarching context. Current research activity involves large components of mineralogical–petrological–geochemical and structural–tectonic work, with much lower levels of experimental, geophysical and geochronological investigations. Many important questions concerning the thermal, physical and chemical aspects of the origin and evolution of granites remain. In keeping with the general progress of science, the complexity of the questions, the declining financial support and the revolution in information technology, directions of granite research in the foreseeable future will change from concrete and qualitative to abstract and quantitative, from expensive and active to cheap and armchair, from reductionist to holistic, and from periodic communication to continuous communication.

Description: :Anhydrite has been identified as a phenocrystic phase in some silicic volcanic magmas, but it is not commonly described in plutonic rocks. Anhydrite-bearing magmas tend to form in arc environments and to contain hydrous, low-temperature, oxidised mineral assemblages. Phenocrystic anhydrite coexists with sulphur-enriched apatite and sometimes with pyrrhotite, in silicate melt that contains from 50 ppm to 1 wt% S, depending on temperature and conditions. Vapour coexisting with anhydrite- and water-saturated magma may contain from a few tenths of a mole per cent to a few mole per cent sulphur gases (SO2 and H2S), with the exact composition and gas speciation depending on temperature and oxygen fugacity. Samples of one anhydrite-bearing magma, the 1991 Pinatubo dacite, have been experimentally crystallised to determine whether the magma retains its characteristic sulphur-rich mineral phases during solidification. Results show that anhydrite and sulphur-rich apatite are retained throughout crystallisation and vapour phase evolution. This suggests that anhydrite-bearing intrusive equivalents of the Pinatubo dacite should be present in arc plutonic complexes.

Description: :In Patagonia a Triassic-Early Jurassic Cordilleran interior magmatic belt preceded the widespread eruption of Middle Jurassic syn-extensional rhyolites. Two plutons (La Calandria and La Leona) represent the easternmost plutonic rocks of this belt, 〉 750 km east of the present oceanic trench. They define a high-K calc-alkaline monzonite series in contrast with the main Andinotype arc magmatism of the Pacific margin: they are enriched in large ion lithophile elements (K, Rb, Ba, Sr and Th), LREE and P2O5and depleted in HREE and Y, with low FeO*/MgO ratio. The range of observed compositions (56-76% SiO2) resulted from high-level fractionation of plagioclase, hornbleńde, biotite, K-feldspar and accessories (sphene, apatite and zircon).Initial87Sr/86Sr ratios, average εNdtand mean depleted-mantle Nd model ages of the two plutons are 0·70487, -0·5 and 1050 Ma for La Calandria and 0·70509, -1·4 and 1125 Ma for La Leona, respectively. They are thus isotopically more primitive than the Middle Jurassic rhyolites, previously attributed to partial melting of Mesoproterozoic mafic lower crust. The preferred model for the origin of the monzonites is remelting of an amphibole- + garnet-bearing, plagioclase-poor, high-K mafic source (?underplating). This occurred in a distal sector of a dying oblique subduction regime, immediately preceding the extensional silicic volcanism.

Description: :Petrological investigations of granite commonly reveal multiple periods of growth punctuated by resorption for many of the constituent minerals. Complementary to such textures are mineral compositional heterogeneity manifested by zoning or grain to grain variability. These features ultimately reflect changes in the intensive parameters or activities of components during melt solidification. Such complexities of granite crystallisation can be simultaneously modelled in a reaction space constructed from the set of linearly independent reactions describing the equilibria among all phases and components in the system of interest.The topology of the linearly independent reactions that define the reaction space for garnetmuscovite-biotite granites yields the following insights: (1) there is no one unique reaction that produces or consumes aluminous minerals (e.g. garnet); (2) minerals can alternate as reactants or products in different reactions accounting for textures indicating multiple periods of crystallisation separated by resorption; (3) mineral compositions are regulated by the reaction(s) producing them and vary as the stoichiometry of the reaction(s) producing them varies; (4) resorption of early crystallising garnet is likely to reflect decreasing pressure, presumably during magma ascent; (5) late crystallisation of garnet, at the expense of biotite, reflects an increase in melt aluminosity and does not necessarily require high Mn activities for the melt and (6) increasing melt H2O, at H2Oundersaturated conditions, favours the formation of biotite–muscovite granite.Application of the reaction space method to other granite types holds considerable promise for elucidating reactions that regulate mineral assemblages and compositions during crystallisation.

Description: :Dehydration melting of crustal rocks may commonly occur in response to the intrusion of mafic magma in the mid- or lower crust. However, the relative importance of melt buoyancy, shear or dyking in melt generation and extraction under geologically relevant conditions is not well understood. A numerical model of the partial melting of a metapelite is presented and the model results are compared with the Ivrea-Verbano Zone in northern Italy. The numerical model uses the mixture theory approach to modelling simultaneous convection and phase change and includes special ramping and switching functions to accommodate the rheology of crystal-melt mixtures in accordance with the results of deformation experiments. The model explicitly includes both porous media flow and thermally and compositionally driven bulk convection of a restitecharged melt mass. A range of melt viscosity and critical melt fraction models is considered. General agreement was found between predicted positions of isopleths and those from the Ivrea-Verbano Zone. Maximum melt velocities in the region of porous flow are found to be 1 × 10−7 and 1 × 10−1m per year in the region of viscous flow. The results indicate that melt buoyancy alone may not be a sufficient agent for melt extraction and that extensive, vigorous convection of partially molten rocks above mafic bodies is unlikely, in accord with direct geological examples.

Description: :The behaviour of trace elements during partial melting depends primarily on their mode of occurrence. For elements occurring as trace constituents of major phases (e.g. Li, Rb, Cs, Eu, Sr, Ba, Ga, etc.), slow intracrystalline diffusion (D ≍ 10−16 cm2 s−1) at the temperature range of crustal anatexis causes all effective crystal-melt partition coefficients to have a value close to unity and impedes further melt-restite re-equilibration. Usually, therefore, the trace element composition of crustal melts simply depends on the mass balance between the proportion and composition of phases that melt and the proportion and composition of newly formed phases. The behaviour of trace elements occurring as essential structural components in accessory phases (e.g. P, La-Sm, Gd-Lu, Y, Th, U, Zr, Hf, etc.) depends on the solubility, solution kinetics, grain size and the textural position of accessory phases. In common crustal protoliths a significant mass fraction of monazite, zircon, xenotime, Th-orthosilicates, uraninite; etc.—but not apatite—is included within other major and accessory phases. During low melt fraction anatexis the amount of accessory phases available for the melt is not sufficient for saturation, thus producing leucosomes with concentrations of La-Sm, Gd-Lu, Y, Th, U and Zr lower than expected from solubility equations. Low concentrations of these elements may also occur if the melt is prevented from reaching equilibrium with the accessories due to fast segregation. However, the first mechanism seems more feasible as leucosomes that are undersaturated with respect to monazite and zircon are frequently saturated, even oversaturated, with respect to apatite.

Description: :Until the last few years, diapirism reigned supreme among granitoid ascent mechanisms. Granitoid masses in a variety of material states, from pure melt through semi-molten crystal mushes to solid rock, were believed to have risen forcefully through the continental crust to their final emplacement levels in a way analogous to salt domes. The structural analogy between granite plutons and salt diapirs, which gained acceptance in the 1930s, has clearly been attractive despite the pessimistic outcomes of thermal models and, at best, ambiguous field evidence.In contrast with traditional diapiric ascent, dyke transport of granitoid magmas has a number of important implications for the emplacement and geochemistry of granites that have yet to be fully explored. Rapid ascent rates of ≍ 10 2m/s predicted for granite melts in dykes (cf. m/a for diapirs) mean that felsic magmas can be transported through the continental crust in months rather than thousands (or even millions) of years, and that large plutons can in principle be filled in

Description: :Diapirism has been discredited as a transport mechanism for magmas partly because diapirs seem to be unable to bring magmas to shallow crustal levels (

Description: :After many years of systematic experimental investigations, it is now possible to quantify the conditions for optimum fertility to melt production of most common crustal rock types as functions of temperature and to about 30 kbar pressure. Quartzo-feldspathic melting produces steady increases in melt proportion with increasing temperature. The exact melt fraction depends on the mineral mode relative to quartz-feldspar eutectics and the temperatures of mica dehydration melting reactions. Mica melting consumes SiO2 from residual quartz during the formation of refractory Al2SiO5, orthopyroxene, garnet or cordierite.A simple graphical interpretation of experimental results allows a deduction of the proportions of mica and feldspar leading to optimum fertility. In effect, the mica dehydration melting reactions, at specific pressure and are superimposed on quartz-feldspar melting relations projected onto Ab-An-Or. Fertility to melt production varies with the mica to feldspar ratio and pressure. Pelites are more fertile than psammites at low pressures (e.g. 5 kbar), especially if they contain An40 to An50 plagioclase. At higher pressure (e.g. 10-20 kbar) and for rocks containing albitic plagioclase, psammites are more fertile than pelites. For a typical pelite (e.g. with An25 at 20 kbar), the cotectic with muscovite lies at higher (≍·) and XAb (≍0·42) than with biotite :≍0·35; XAb(≍·), thus dehydration melting of muscovite requires 10% more plagioclase for fertility than does biotite.The first melts from dehydration melting of muscovite (with Plg + Qtz) are more sodic and form at lower temperatures than the first melts from Bio + Plg + Qtz. With increasing pressure, to at least 30 kbar, granite minimum and mica dehydration melts become more sodic. This indicates that of such melts is greater than 0·3.

Description: :Melting experiments with and without added H2O on a model metagreywacke and a natural metapelite demonstrate how pressure and H2O content control the compositions of melts and residual assemblages. Several effects are observed under isothermal conditions. Firstly, the stability field of biotite shrinks with decreasing pressure and with increasing H2O content, whereas that of plagioclase shrinks with increasing pressure and H2O content. Secondly, the ferromagnesian content of melts at the source (i.e. coexisting with their residual assemblages) decreases with decreasing H2O activity. Thirdly, with increasing pressure the Ca/Mg and Ca/Fe ratios of melts decrease relative to those of coexisting garnet. As a consequence, a wide spectrum of melts and crystalline residues can be generated from the same source material. For example, H2O-starved dehydration melting of metagreywacke at low pressure (≤10 kbar) generates K-rich (granitic) melts that coexist with pyroxene- and plagioclase-rich residues, whereas melting of the same material at high pressure (≍15 kbar) and with minor H2O infiltration can generate leucocratic Na-rich and Ca-poor (trondhjemitic) melts that coexist with biotite- and garnet-rich residues. An increased H2O content stabilises orthopyroxene at the expense of garnet + biotite + plagioclase, causing melts to shift towards granodioritic or perhaps tonalitic compositions.

Description: Foraminiferal, algal and problematica assemblages from the Mississippian (late Viséan and early Serpukhovian) Lower Limestone Formation have been studied in order to validate lithostratigraphical correlations of limestones within the central and western parts of the Midland Valley of Scotland. Analysis of more than 100 outcrops allows recognition of four calcareous microfossil assemblages, which span the late Brigantian and early Pendleian, and enables a detailed correlation to be made within the Central Coalfield (north Lanarkshire) and with the thinner sequences to the west (north Ayrshire), to the south (Douglas area, south Lanarkshire), and to the east (Bathgate area, West Lothian). The age of the Lower Limestone Formation is modified because the upper part of this formation is now assigned to the Pendleian (due to the first occurrences of new foraminiferans and the co-occurrence with the Namurian goniatites), and some individual limestone horizons within the formation are repositioned, or their precise correlation with other limestones is established. A refined stratigraphical framework is proposed for the above noted areas, and a correlation between them and the Pennine region in northern England is proposed, passing through the Archerbeck Borehole sequence in the Scottish Borders.

Description: The Cape Granites are a granitic suite intruded into Neoproterozoic greywackes and slates, and unconformably overlain by early Palaeozoic Table Mountain Group orthoquartzites. They were first recognised at Paarl in 1776 by Francis Masson, and by William Anderson and William Hamilton in 1778. Studies of the Cape Granites were central to some of the early debates between the Wernerian Neptunists (Robert Jameson and his former pupils) and the Huttonian Plutonists (John Playfair, Basil Hall, Charles Darwin), in the first decades of the 19th Century, since it is at the foot of Table Mountain that the first intrusive granites outside of Scotland were described by Hall in 1812. The Neptunists believed that all rocks, including granite and basalt, were precipitated from the primordial oceans, whereas the Plutonists believed in the intrusive origin of some igneous rocks, such as granite. In this paper, some of the early descriptions and debates concerning the Cape Granites are reviewed, and the history of the development of ideas on granites (as well as on contact metamorphism and sea level changes) at the Cape in the late 18th Century and early to mid 19th Century, during the emerging years of the discipline of geology, is presented for the first time.

Description: The Dalradian Supergroup contains three distinct glacigenic units, formerly termed ‘Boulder Beds’, which are correlated with widespread Neoproterozoic glaciations. The oldest and thickest unit, the Port Askaig Formation, marks the Appin–Argyll group boundary of the Dalradian Supergroup and has been correlated with the Middle Cryogenian (Sturtian) glaciation. The Auchnahyle Formation, a diamictite-bearing sequence near Tomintoul in NE Scotland, exhibits strong lithological similarities to the Port Askaig Formation. Both these glacigenic ‘Boulder Bed’ units contain abundant dolomite clasts in their lower parts and more granitic material at higher levels. Both metadiamictite units are overlain by thick shallow-marine quartzite units. C isotope data from Appin Group carbonate strata below the Auchnahyle Formation support this correlation. U–Pb laser ablation inductively coupled plasma mass spectrometry (LA-ICP-MS) detrital zircon data from the Auchnahyle Formation metadiamictite differ slightly from the Port Askaig Formation, but are similar to detrital zircon spectra obtained from the Macduff Formation, a diamictite unit in the younger Southern Highland Group of the Dalradian Supergroup; both apparently reflect derivation from local basement rocks. No detritus younger than 0·9 Ga is observed, so the data do not constrain significantly the depositional age of the glacial strata. A thin tholeiitic pillow basalt unit in the lower part of the Auchnahyle Formation is geochemically distinct from pre-tectonic metadolerite sills and from basic metavolcanic rocks up-section. A Sturtian (c. 720–700 Ma) age for the Auchnahyle Formation metadiamictite would imply that this basaltic volcanism represents the oldest recorded volcanic activity in the Dalradian Supergroup and is inferred to represent an early, local phase of proto-Iapetan rifting within the Rodinian supercontinent.

Description: Conodont faunal dynamics and high-resolution biostratigraphy in the lithologically and faunally anomalous ‘Täljsten’ succession, which spans the DarriwilianLenodus variabilis–Yangtzeplacognathus crassusZone boundary, were investigated in a 2·5 m-thick section on Mt Kinnekulle that includes an interval yielding fossil meteorites and extraterrestrial chromite. The previous interpretation that this interval reflects a regression is consistent with the occurrence and abundance patterns of some conodont taxa. The disappearance of e.g.,Periodon, suggests that the regression began prior to the deposition of the grey ‘Täljsten’. The transition from red to grey limestone coincides with a conspicuous faunal re-arrangement. The lower half of the ‘Täljsten’ reflects a gradual shallowing favourable for some taxa, such asLenodus, and the immigration ofMicrozarkodinacf.ozarkodellaandHistiodella holodentata. In the middle of the ‘Täljsten’ interval, coinciding with the appearance of abundant cystoids, conditions became less hospitable for conodonts, resulting in a low diversity and low abundance fauna, which occurs to the top of the interval. The overlying red limestone, apparently deposited during a deepening event, marks a return to pre-‘Täljsten’ conditions with a re-organised fauna. The close correlation between the lithologic shifts and conodont faunal changes demonstrates the usefulness of conodonts as environmental indicators.

Description: Two regional-scale, recumbent folds control the structure of the Beinn Udlaidh area, Tyndrum, Perthshire. They reached their maximum development during D2, following the regional metamorphic peak, and are part of a stack of larger SE-facing recumbent folds formed during the ∼470 Ma Grampian Orogeny. The rocks belong to the Neoproterozoic–Lower Ordovician Dalradian Supergroup, and preserve a sedimentary transition between the Grampian Group and the overlying Appin Group. The latter occupies the core of the S-facing, recumbent Beinn Udlaidh Syncline (D2) which, with the underlying complementary Glen Lochy Anticline, is gently folded by a regional-scale structure, the Orchy Dome. The recumbent folds postdate an early fabric (S1), which is generally obliterated by the D2 imprint, but preserved as inclusion trails in regional metamorphic garnets, that are highly oblique to, and wrapped by, S2. It is concluded that the Dalradian rocks described here from below the Iltay Boundary Slide are in structural continuity with those of the Tay Nappe above, and that the Slide represents a structurally-modified disconformity between the Leven Schist (Appin Group) and the overlying Ben Lui Schist (Argyll Group). The Orchy Dome probably influenced the spatial distribution of minor intrusions and explosion vents of the lamprophyre suite.

Description: Dispersed spore assemblages have been recovered from the Am Binnein Sandstones from the upper part of the ‘Lower Old Red Sandstone’ sequence on the island of Arran, Scotland. The spore assemblages belong with theEmphanisporites annulatus–Camarozonotriletes sextantii(AS) Spore Assemblage Biozone (SAB), indicating an Early Devonian, Emsian (but not earliest Emsian or latest Emsian) age. This is the first reliable age constraint for the ‘Lower Old Red Sandstone’ of Arran, and enables correlation with the more extensive sequence developed on the mainland in the Midland Valley of Scotland. The Am Binnein Sandstones are confirmed as correlatives of the Strathmore Group.

Description: The authors revise the occurrences of burrow networks with striated walls having dominantly transverse to oblique striae, which have been assigned to the ichnogeneraSpongeliomorphaSaporta, 1887, andSteinichnusBromley & Asgaard, 1979. The taxonomic status of the ichnogenusSteinichnusBromley & Asgaard, 1979 is examined and it is suggested that this ichnogenus is a subjective junior synonym ofSpongeliomorphaSaporta, 1887.Spongeliomorphais best reserved for an unlined network of burrows having distinct surface ridges or grooves of different orientation and massive filling. The diagnosis ofSpongeliomorphais emended accordingly and the proposed ichnospecies revised for consistency with the diagnostic features of the ichnogenus.Spongeliomorpha milfordensisMetz, 1993a is considered a subjective junior synonym ofSpongeliomorpha carlsbergi(Bromley & Asgaard, 1979) after a visual comparison and statistical analysis of the angle of striation with respect to the burrow midline in the type material. Nevertheless, the use of statistical techniques is not advocated for distinction of ichnotaxa, but may support observations.Spongeliomorpha carlsbergiis considered as an indicator of nonmarine settings and was probably produced by burrowing insects. Proposed ichnospecies ofSpongeliomorphathat fit the emended diagnosis includeS. sudolica(Zaręczny, 1878);S. ibericaSaporta, 1887;S. siculaD’Alessandro & Bromley, 1995;S. chevronensisMuñiz & Mayoral, 2001; andSpongeliomorphaisp. nov. aff.siculaLewy & Goldring, 2006.

Description: The morphology of the amphibamid temnospondyl amphibian Platyrhinops lyelli from the Middle Pennsylvanian of Linton, Ohio is reassessed from previously described and undescribed specimens. Newly reported or newly significant features include the presence of bicuspid marginal teeth, anteriorly widened frontals, elongate choanae, a broad rhomboidal sphenethmoid and a pair of flattened blade-like ceratohyals. One specimen shows an unusual distribution of tooth sizes along the premaxillaries. No derived characters can be found to justify reference of the species lyelli to the genus Amphibamus as represented by its type species A. grandiceps, but Platyrhinops does belong to the ‘Amphibamus branch’ of the Amphibamidae.

Description: This paper draws on epistemology, theory, published data, hypothesis testing and synthesis to explore the predictive and explanatory power of a far-from-equilibrium interpretation of the history of human settlement on Rapa Nui. Our interpretation of the last 1500 years of human settlement on Rapa Nui provides an important competing hypothesis by which to test the explanatory power of earlier equilibrium interpretations of this history. Our evaluation of this competing hypothesis suggests that it provides, overall, a more satisfactory theory that has interesting implications for the goal of sustaining human civilisation.

Description: As an isolated island group lying off the NW European mainland which was uninhabited until the mid-first millennium AD, the Faroes offer a unique opportunity to study natural processes of Holocene ecosystem development in a region where anthropogenic activity is usually a complicating factor. In this paper new radiocarbon dates and pollen-analytical data from the island of Sandoy, in the centre of the Faroes archipelago, are presented. Together with existing pollen and plant macrofossil records, these data allow a reconstruction of patterns of Holocene vegetational and edaphic change. Basal peat dates indicate that large areas of blanket mire were established long before the first human settlement, demonstrating conclusively that human impact is not necessary for the development of such ecosystems. The timing of the initiation of the blanket peats varies markedly, both across the Faroes as a whole and at a landscape scale, with dates distributed evenly over 9000 years. This suggests that, in the Faroes at least, pedogenesis was more important than climatic change in determining the timing of the spread of blanket peat systems.

Description: Though typically exhibiting considerable scatter, geochemical variations in granitic plutons and silicic volcanic deposits are commonly modelled as products of differentiation of originally homogeneous magmas. However, many silicic igneous bodies, particularly those classified as S-types, are internally heterogeneous in their mineralogy, geochemistry and isotope ratios, on scales from hundreds of metres down to one metre or less. The preservation of these heterogeneities supports recent models for the construction of granitic magma bodies through incremental additions of numerous batches (pulses) of magma derived from contrasting sources. Such pulses result from the sequential nature of the melting reactions and the commonly layered structure of crustal magma sources. Internal differentiation of these batches occurs, but not generally on the scales of whole magma chambers. Rather than being created through differentiation or hybridisation processes, at or near emplacement levels, much of the variation within such bodies (e.g. trace-element or Mg# variation with SiO2 or isotope ratios) is a primary or near-source feature. At emplacement levels, the relatively high magma viscosities and slow diffusion rates of many chemical components in silicic melts probably inhibit processes that would lead to homogenisation. This permits at least partial preservation of the primary heterogeneities.

Description: Trace element abundances in garnet from a polyphase migmatite were measured by secondary ion mass spectrometry (SIMS) in order to identify some of the effective variables on the trace element distribution between garnet and melanosome or leucosome. In general, garnet is zoned with respect to REE, in which garnet cores are enriched by a factor of 2–3 relative to the rims. For an inclusion-rich garnet from the melanosome, equilibrium distribution following a simple Rayleigh fractionation is responsible for the decreasing concentrations in REE from core to rim. Inclusion-poor garnet from the same melanosome located in the vicinity of the leucosomes shows distinct enrichment and depletion patterns for REE from core to rim. These features suggest disequilibrium between garnet and the host rock which, in this case, could have been an in-situ derived melt. This would probably indicate a period of open-system behaviour at a time when the garnet, originally nucleated in the metamorphic environment reacted with the melt. In addition, non-gradual variation in trace element abundances between core and rim may suggest variable garnet growth rates. Inclusion-free garnet from the leucosome, interpreted to have crystallised in the presence of a melt, has a small core with high REE abundances and a broad rim with lower REE abundances. Here, crystal-liquid diffusion-controlled partitioning is a likely process to explain the trace element variation.

Description: Like most metazoan biomineralisations, the brachiopod shell is the end product of a biologically controlled calcification process. The main agent of the control is the extracellular matrix, which is secreted by the outer mantle epithelium. This matrix mediates the calcification process by allowing crystal nucleation and elongation in specific orientations and finally, by stopping crystal growth. The proteinaceous moiety of brachiopod shell matrices has been extensively studied. Less known are the post-translational modifications that occur in these matrices, in particular glycosylations. In this comparison of five species of Recent articulated brachiopods, the ratio of soluble to insoluble organic matrix varies between the species. Polydisperse macromolecular materials occur in each of these species with discrete proteins of 50 kDa in Notosaria nigricans, Calloria inconspicua and Neothyris lenticularis, 37 kDa in Terebratulina retusa and Gryphus vitreus and 20–25 kDa in N. nigricans. Protein mixtures from all five species respond differently to anionic stains (Stains-All and Alcian Blue). PAS staining results in a positive smear in C. inconspicua and T. retusa and highlights low molecular weight glycoproteins in C. inconspicua. The polysaccharide composition of the soluble matrix of T. retusa is different from the others due to high proportions of arabinose and low proportions of fucose. In all cases, polysaccharide composition of the insoluble matrix is dominated by glucose and glucosamine. Insoluble matrices have more glucose and xylose and less galactosamine and glucosamine than the corresponding soluble matrix. Relatively high amounts of glucosamine may suggest the presence of chitin in the shell matrix of rhynchonelliform brachiopods.

Description: The causes of compositional diversity in the Tuolumne Batholith, whether source heterogeneity, magma mixing, or fractional crystallisation, is a matter of longstanding debate. This paper presents data from detailed mapping and a microstructural and major element, trace element and isotopic study of an elongate lobe of the Half Dome granodiorite that protrudes from the southern end of the batholith. The lobe is normally zoned from quartz diorite along the outer margin to high-silica leucogranite in the core. Contacts are steep and gradational, except for the central leucogranite contact, which is locally sharp: magmatic fabrics overprint contacts. A striking feature of the lobe is the 18 wt SiO2 range comparable to that observed for the entire Tuolumne Batholith. Feldspar-compatible elements (Sr and Ba) decrease towards the centre, while Rb increases. Light and middle REEs show a smooth decrease towards the centre of the lobe. Calculated initial isotopic ratios of 87Sr/86Sr(i) and εNd(t) have identical values within error across the lobe, except in the central leucogranite, the most silica rich phase, which shows a slightly more crustal signature. Field, structural, geochemical and isotopic data suggest that fractionation was the dominant process causing compositional variation in this lobe. It is envisioned that this fractionation/crystal sorting occurred in a vertically flowing and evolving magma column with the present map pattern representing a cross-section of this column. Thus the areal extent of the lobe represents a minimum size of interconnected melt at the emplacement level of the Tuolumne Batholith and, given its marginal position, limited width and proximity to colder host rocks, implies that fractionation in larger chambers likely occurred in the main Tuolumne Batholith magma chamber(s).

Description: In the recently completed and formally ratified new series and stage classification of the Ordovician System, the base of the Middle Ordovician Series coincides with the base of the global Dapingian Stage. In the Global Stratotype Section and Point (GSSP) of this stage, which is located at Huanghuachang in southern China, the base of the Dapingian Stage is defined as the level of first appearance of the conodont Baltoniodus triangularis. The fact that this species, along with some other taxa present at the Dapingian GSSP, occurs in many sections in Baltoscandia makes it possible to recognise with considerable precision the level of this global stage boundary in Sweden, Estonia, northwestern Russia, and Denmark. In several, but not all, regions, especially in the East Baltic, the global stage boundary coincides with the base of the regional Volkhov Stage and can be tied to the base of the Megistaspis polyphemus Trilobite Zone. The regionally somewhat different relationships between the position of the global stage boundary and a very widespread hardground complex are probably due to the occurrence of local and/or regional unconformities in the upper Floian–lower Dapingian interval. Although biostratigraphically important graptolites are present in the study interval in some Baltoscandic sections, the precise graptolite correlation of the base of the Dapingian Stage remains somewhat unclear, although it appears to be near, or at, the base of the Isograptus victoriae victoriae Zone (Ca 2).

Description: New three-dimensionally preserved specimens of two euthycarcinoid arthropods, namely Schramixerxes gerem and Sottyxerxes multiplex, allow complete description of both the dorsal and ventral sides of the exoskeleton. The functional morphology is tentatively interpreted for the first time. In S. gerem, the ‘thirteenth somite’, or ‘monosomite’ is fully described and re-interpreted as the main articulatory process of the body, between the cephalic region and the preabdomen. The morphology and arrangement of the two parts of the process clearly indicate that the anterior cephalic region of the body could move laterally and bend ventrally, while posterior somites could only move ventrally. Unlike several other euthycarcinoid species, the ventral side of the head area exhibits one or two plates instead of mandibles; such distinct morphologies are indicative of different feeding mechanisms and behaviours among euthycarcinoid arthropods. Possible homologies with the labrum of Hexapoda support the hypothesis that euthycarcinoids have hexapod affinities.

Description: This study explores the implications of different decisions about taxon sampling in studies of phylogeny and macroevolution – how would phylogenetic results differ as decisions about inclusion of taxa differed? The focus is on investigating phylogenetic relationships among families of Devonian terebratulides, and include all 71 named genera in our analyses. Subsets of taxa were experimented with the most complete morphologically from fossil specimens; those that occur earliest in the stratigraphic record; those that include only the name-bearing genera from each family. Including only the most completely known genera produces a result essentially similar to one including all genera, even those for which less than half the characters can be coded. Including only the stratigraphically earliest genera produces a result at odds with the other analyses. Including only name-bearers, representing 13% of all genera, produces a result generally similar to the analysis including all taxa. None of the results of these phylogenetic experiments involving subsets of genera corresponds strongly with the recently revised classification in the Treatise on Invertebrate Paleontology, but general similarities can be discerned. The lack of strong correspondence between classification and several different experimental phylogenetic hypotheses could be ascribed to their different overall goals, and highlights the potential dangers of ascribing evolutionary significance to simple counts of taxa, particularly families, as warned by Alwyn Williams 50 years ago.

Description: The records of columnar shell structures of linguloid brachiopods (Class Lingulata, Order Lingulida, Superfamily Linguloidea) are reviewed in the light of the discovery of two new taxa from the Middle Cambrian Forsemölla Limestone Bed of southern Sweden. The linguloid taxa, described here as Eoobolus? sp. aff. E. priscus (Poulsen) and Canalilatus? simplex sp. nov., are both characterised by a columnar shell structure, a structural type that is representative for acrotretoid brachiopods and that has previously only rarely been reported from the linguloids. Though the two taxa are superficially similar to known genera, i.e., Eoobolus and Canalilatus, their shell structure challenges such affiliations, as the shell structure of the type species of these genera is previously unknown. Linguloid families whose morphological characteristics agree the most with those of the new taxa, i.e., the Zhanatellidae and the Eoobolidae, and from which columnar shell structures have been reported, i.e., the Lingulellotretidae and the Kyrshabaktellidae, are reviewed briefly. Many taxa assigned to these families completely lack shell structure data and are in need of restudy in order to elucidate their systematic position. Knowledge of the representative type of shell structure of the various suprageneric taxa within the Linguloidea is considered crucial, in order to unravel their currently poorly resolved phylogenetic relationships.

Description: Earliest Silurian (basal Llandovery) brachiopod faunas are surveyed and listed from around the globe, and divided between Lower Rhuddanian and Upper Rhuddanian occurrences. 60 genera are known from the Lower Rhuddanian within 20 superfamilies and there are 87 genera in 25 superfamilies in the Upper Rhuddanian. The 29 areas surveyed span the globe, both latitudinally and longitudinally. Only six superfamilies are Lazarus taxa which are known both from the Ordovician and Middle Llandovery (Aeronian) and later rocks but have not been recorded from the Rhuddanian. These are surprising results, since many previous studies have inferred that the Rhuddanian was a time of very sparse faunas. The global warming that followed the latest Ordovician (Hirnantian) ice age did not proceed quickly, with an ice-cap probably present through at least the Llandovery. There is a marked absence of Lower Rhuddanian bioherms even at low palaeolatitudes; however, the ecological recovery rate was far faster than that following the end-Permian mass extinction event. The partitioning of the Rhuddanian shelf faunas into well-defined benthic assemblages progressed slowly over the interval.

Description: The Lower Permian of the Haushi basin, Interior Oman (Al Khlata Formation to Saiwan Formation/lower Gharif member) records climate change from glaciation, through marine sedimentation in the Haushi sea, to subtropical desert. To investigate the palaeoclimatic evolution of the Haushi Sea we used O, C, and Sr isotopes from 31 brachiopod shells of eight species collected bed by bed within the type-section of the Saiwan Formation. We assessed diagenesis by scanning electron microscopy of ultrastructure, cathodoluminescence, and geochemistry, and rejected fifteen shells not meeting specific preservation criteria. Spiriferids and spiriferinids show better preservation of the fibrous secondary layer than do orthotetids and productids and are therefore more suitable for isotopic analysis. δ18O of −3·7 to −3·1℅ from brachiopods at the base of the Saiwan Formation are probably related to glacial meltwater. Above this, an increase in δ18O may indicate ice accumulation elsewhere in Gondwana or more probably that the Haushi sea was an evaporating embayment of the Neotethys Ocean. δ13C varies little and is within the range of published data: its trend towards heavier values is consistent with increasing aridity and oligotrophy. Saiwan Sr isotope signatures are less radiogenic than those of the Sakmarian LOWESS seawater curve, which is based on extrapolation between few data points. In the scenario of evaporation in a restricted Haushi basin, the variation in Sr isotope composition may reflect a fluvial component.

Description: A diverse, previously undescribed trilobite fauna from northwest Iran sheds new light on the Early Palaeozoic geography of the Alborz terrane. The structural history of the Alborz region is well known, though faunal data for the area are sparse, especially for the Ordovician. Trilobites, brachiopods, conodonts, bryozoans and echinoderms occur in a red grainstone/packstone near the village of Tatavrud, 35 km southwest of Bandar-e-Anzali. The fossils indicate a Late Ordovician age, although lithologically similar outcrops in northern Iran have previously been considered Silurian. Trilobite genera occurring include Trinodus, Geragnostus, Illaenus, Panderia, Phorocephala, Ovalocephalus, Dicranopeltis, Symphysops, Cyclopyge, and Sphaerexochus. Similar assemblages have been reported from the Late Ordovician of Ireland, Spain, Poland, Norway, northwest China, Kazakhstan and the Turkistan–Alai Ridge (Uzbekistan–Kyrgyzstan border). Reconstructions of Late Ordovician geography place the Alborz terrane near the eastern margin of Gondwana, but there is doubt as to its precise location. Some workers have considered it part of the margin while others have considered it a separate terrane. Several species present, including Mezzaluna tatavrudensis n. sp. and Phorocephala cf. ulugtana (Petrunina, 1975) are very similar to taxa described from the Turkistan–Alai Ridge. In addition, the isocolid genus Paratiresias has only been reported from Iran, the Turkistan–Alai Ridge and northwest China. The presence of these taxa indicates that the Alborz terrane was in close proximity to the eastern margin of Gondwana at this time.

Description: We consider the reinterpretation of Bethia by Bassett et al. (2008) to be flawed in several regards. Details of the pedicle rootlets and wrinkles strongly imply a soft-tissue structure rather than calcified sheath. The ornament is unusual, but equally so under either an orthide or a plectambonitoidean model. The reinterpretation of the ‘deltial plates’ as ‘chilidial plates’ is likely correct, but argues against a plectambonitoidean affinity by implying that the pedicle emerges between the valves. Other arguments presented in favour of a plectambonitoidean affinity are also discussed; we consider them unpersuasive. Finally, we contend that Bethia is sufficiently well characterised to deserve a taxonomic name.

Description: The Palaeogene volcanic succession of the Faroe Islands in the NE Atlantic Ocean is formalised using a purely lithostratigraphic approach and following international guidelines. The Faroe Islands Basalt Group (FIBG) has a gross stratigraphic thickness of ∼6·6 km, dominated by subaerial basalt lava flows, and is subdivided into seven formations. The Lopra Formation forms the basal ∼1·1 km of the Lopra-1/1A borehole, dominated by hyaloclastites, volcaniclastic sandstones and invasive basaltic lavas/sills. It is overlain by the ∼3·25 km-thick Beinisvørð Formation, dominated by laterally extensive basalt sheet lobes separated by minor volcaniclastic lithologies. The Beinisvørð Formation is overlain by the

Description: The exposed nature of the northern Faroe Islands high relief landscape enabled widespread Holocene slope process activity and deposition of related landforms, which seem largely controlled by extreme meteorological conditions. Three different slope landforms – the large colluvial Glyvurs fan, the lower Marknastiggjur mountainside debris-flow deposits in the town of Klaksvik, and the mountain top aeolian sediment cover on Eidiskollur – were investigated by a combination of geomorphological, stratigraphic, sedimentological, chronological and modern process studies.Sporadic Holocene snow-avalanche and debris-flow activity were documented, with sedimentation starting significantly before 8000 cal yr BP in the Glyvurs fan, which still sporadically experiences activity. The largest amounts of Holocene slope sedimentation seem to occur in colluvial fans, such as the Glyvurs fan, which are located below large dyke canyons, called gjogvs. The lower Marknastiggjur mountainside consists of mainly debris-flow deposits, which started before 7800 cal yr BP. A relatively small amount of precipitation, but with high precipitation intensity after a dry summer, triggered modern small-scale debris-flows in the northern islands, also at the Marknastiggjur mountainside, early in Autumn 2000. Extensive continuous mountaintop aeolian sedimentation from cliff weathering started around 6900 cal yr BP on the Eidiskollur peninsula.No direct influence of settlement on slope process activity was found at the different investigated slope landforms in the northern Faroe Islands.

Description: Campbell Island is a small, uninhabited peat-covered island lying in the cool southern ocean 600 km south of the New Zealand mainland. Dracophyllum scrub is the main cover from sea level to 200 m, above which tussock grassland, macrophyllous forbs and tundra dominate. Seven peat profiles from sea level to the tundra zone provide an elevational transect for pollen and charcoal records spanning the last 500 years. Scrub density was relatively low between 200 and 400 cal yrs BP, possibly due to Little Ice Age cooling, but had recovered by the time Europeans discovered the island in AD 1810. Burning and grazing during a brief farming episode (AD 1895–1931) severely reduced scrub and palatable grasses and forbs. Vegetation recovery is now well advanced following cessation of farming and the later elimination of all feral grazing animals, cats and rats. Climates were cool in the southwest Pacific during the farming period, and since AD 1970 the island has warmed by c. 0·5°C. However, there has been no upwards movement of the scrubline despite vigorous regeneration of scrub at lower altitudes. The island's cloudy, highly oceanic climate appears to offset increasing summer warmth, and scrubline is likely to rise only if clearer and less windy, as well as warmer, summers eventuate.

Description: A study of type and newly collected material of the Silurian ostracodRichteria migrans(Barrande, 1872) demonstrates that it had wide distribution, occurring in at least the Czech Republic, France, Sardinia, Wales, central Asia and probably Poland.R. migranshas biostratigraphic value, as an indicator for the mid to late part of the Ludlow Series. It was almost certainly pelagic, living predominantly in probable shelf topographic lows to marginal/off-shelf environments, characteristically with cephalopod-graptolite-bivalve-dominant associates. It had at least five to six growth stages, exhibits polymorphic variation, and its morphology provides evidence to endorse the notion that ‘entomozoaceans’ are myodocopes.

Description: :Granites within suites share compositional properties that reflect features of their source rocks. Variation within suites results dominantly from crystal fractionation, either of restite crystals entrained from the source, or by the fractional crystallisation of precipitated crystals. At least in the Lachlan Fold Belt, the processes of magma mixing, assimilation or hydrothermal alteration were insignificant in producing the major compositional variations within suites. Fractional crystallisation produced the complete variation in only one significant group of rocks of that area, the relatively high temperature Boggy Plain Supersuite. Modelling of Sr, Ba and Rb variations in the I-type Glenbog and Moruya suites and the S-type Bullenbalong Suite shows that variation within those suites cannot be the result of fractional crystallisation, but can be readily accounted for by restite fractionation. Direct evidence for the dominance of restite fractionation includes the close chemical equivalence of some plutonic and volcanic rocks, the presence of plagioclase cores that were not derived from a mingled mafic component, and the occurrence of older cores in many zircon crystals. In the Lachlan Fold Belt, granite suites typically evolved through a protracted phase of restite fractionation, with a brief episode of fractional crystallisation sometimes evident in the most felsic rocks. Evolution of the S-type Koetong Suite passed at about 69% SiO2 from a stage dominated by restite separation to one of fractional crystallisation. Other suites exist where felsic rocks evolved in the same way, but the more mafic rocks are absent. In terranes in which tonalitic rocks formed at high temperatures are more common, fractional crystallisation would be a more important process than was the case for the Lachlan Fold Belt.

Description: :To form a granite pluton, the felsic melt produced by partial melting of the middle and lower continental crust must separate from its source and residuum. This can happen in three ways: (1) simple melt segregation, where only the melt fraction moves; (2) magma mobility, in which all the melt and residuum move together; and (3) magma mobility with melt segregation, in which the melt and residuum move together as a magma, but become separated during flow. The first mechanism applies to metatexite migmatites and the other two to diatexite migmatites, but the primary driving forces for each are deviatoric stresses related to regional-scale deformation. Neither of the first two mechanisms generates parental granite magmas. In the first mechanism segregation is so effective that the resulting magmas are too depleted in FeOT, MgO, Rb, Zr, Th and the REEs, and in the second no segregation occurs. Only the third mechanism produces magmas with compositions comparable with parental granites, and occurs at a large enough scale in the highest grade parts of migmatite terranes, to be considered representative of the segregation processes occurring in the source regions of granites.

Description: :Buddington (1959) pointed out that the construction of large crustal magma chambers involves complex internal processes as well as multiple country rock material transfer processes (MTPs), which reflect large horizontal, vertical and temporal gradients in physical conditions. Thus, we have attempted to determine the relative importance of different magmatic and country rock MTPs at various crustal depths, and whether country rock MTPs largely transport material vertically or horizontally, rather than seeking a single model of magma ascent and emplacement.Partially preserved roofs of nine plutons and in some cases roof–wall transitions with roof emplacement depths of 1·5–11 km were mapped. During emplacement, these roofs were not deformed in a ductile manner, detached or extended by faults, or significantly uplifted. Instead, sharp, irregular, discordant contacts are the rule with stoped blocks often preserved immediately below the roof, even at depths of 10 km. The upper portions of these magma chambers are varied, sometimes preserving the crests of more evolved magmas or local zones of volatile-rich phases and complex zones of dyking and magma mingling. Magmatic structures near roofs display a wide variety of patterns and generally formed after emplacement. Transitions from gently dipping roofs to steep walls are abrupt. At shallow crustal levels, steep wall contacts have sharp, discordant, stepped patterns with locally preserved stoped blocks indicating that the chamber grew sideways in part by stoping. Around deeper plutons, an abrupt transition (sometimes within hundreds of metres) occurs in the country rock from discordant, brittle roofs to moderately concordant, walls deformed in a ductile manner defining narrow structural aureoles. Brittle or ductile faults are not present at roof–wall joins.Near steep wall contacts at shallow to mid-crustal depths (5–15 km), vertical and horizontal deflections of pre-emplacement markers (e.g. bedding, faults, dykes), and ductile strains in narrow aureoles (0·1–0·3 body radii) give a complete range of bulk strain values that account for 0–100% of the needed space, but average around 30%, or less, particularly for larger batholiths. A lack of far-field deflection of these same markers rules out significant horizontal displacement outside the aureoles and requires that any near-field lateral shortening is accommodated by vertical flow. Lateral variations from ductile (inner aureole) to brittle (outer aureole) MTPs are typically observed. Compositional zoning is widespread within these magma bodies and is thought to represent separately evolved pulses that travelled up the same magma plumbing system. Magmatic foliations and lineations commonly cross-cut contacts between pulses and reflect the strain caused either by the late flow of melt or regional deformation.Country rocks near the few examined mid- to deep crustal walls (10–30 km) are extensively deformed, with both discordant and concordant contacts present; however, the distinction between regional and emplacement-related deformation is less clear than for shallower plutons. Internal sheeting is more common, although elliptical masses are present. Lateral compositional variations are as large as vertical variations at shallower depths and occur over shorter distances. Magmatic foliations and lineations often reflect regional deformation rather than emplacement processes.The lack of evidence for horizontal displacement outside the narrow, shallow to mid-crustal aureoles and the lack of lateral or upwards displacement of pluton roofs indicate that during emplacement most country rock is transported downwards in the region now occupied by the magma body and its aureole. The internal sheeting and zoning indicate that during the downwards flow of country rock, multiple pulses of magma travelled up the same magma system. If these relationships are widespread in arcs, magma emplacement is the driving mechanism for a huge crustal-scale exchange process.

Description: :Comb-layered quartz is a type of unidirectional solidification texture found at the roofs of shallow silicic intrusions that are often associated spatially with Mo and W mineralisation. The texture consists of multiple layers of euhedral, prismatic quartz crystals (Type I) that have grown on subplanar aplite substrates. The layers are separated by porphyritic aplite containing equant phenocrysts of quartz (Type II), which resemble quartz typical of volcanic rocks and porphyry intrusions. At Logtung, Type I quartz within comb layers is zoned with respect to a number of trace elements, including Al and K. Concentrations of these elements as well as Mn, Ti, Ge, Rb and H are anomalous and much higher than found in Type II quartz from Logtung or in igneous quartz reported elsewhere. The two populations appear to have formed under different conditions. The Type II quartz phenocrysts almost certainly grew from a high-silica melt between 600 and 800°C (as β-quartz); in contrast, the morphology of Type I quartz is consistent with precipitation from a hydrothermal solution, possibly as α-quartz grown below 600°C. The bulk compositions of comb-layered rocks, as well as the aplite interlayers, are consistent with the hypothesis that these textures did not precipitate solely from a crystallising silicate melt. Instead, Type I quartz may have grown from pockets of exsolved magmatic fluid located between the magma and its crystallised border. The Type II quartz represents pre-existing phenocrysts in the underlying magma; this magma was quenched to aplite during fracturing/degassing events. Renewed and repeated formation and disruption of the pockets of exsolved aqueous fluid accounts for the rhythmic banding of the rocks.

Description: :Interaction processes between acid and basic magmas are widespread in the Sardinia–Corsica Batholith. The resulting hybrid magmas are extremely variable and can be broadly divided into: (i) microgranular mafic enclaves with geochemical characteristics of both magmatic liquids and cumulates; (ii) basic gabbroic complexes with internal parts mainly formed by cumulates and with interaction zones developing only in the marginal parts; and (iii) basic septa with the form of discrete, lenticular-like bodies often mechanically fragmented in the host rock. Different styles of interaction, ranging from mixing to mingling, have been related to variations in several physicochemical parameterś, such as: (i) the initial contrast in chemical composition, temperature and viscosity; (ii) the relative mass fractions and the physical state of interacting magmas; and (iii) the static versus dynamic environment of interaction.A model is presented for the origin and history of interaction processes between basic and acid magmas based on the geochemical characteristics of hybrid magmas. Physico-chemical processes responsible for the formation of hybrid magmas can be attributed to: (i) fractional crystallisation of basic magma and contamination by acid magma; (ii) loss of the liquid phase from the evolving basic magma by filter pressing processes; (iii) mechanical mixing between basic and acid magmas; and (iv) liquid state isotopic diffusion during the attainment of thermal equilibrium.

Description: :The primary focus of this review is on P-T conditions, mineralogical indicators of melt or fluid composition and textural evolution; lesser treatment is given to pegmatite sources or to pegmatite-wallrock interactions. Investigations of stable and radiogenic isotopes have revealed that the source materials for pegmatites are likely to be more heterogeneous or varied than previously thought, especially for peraluminous pegmatites, but that overall pegmatites bear a clear intrusive relationship with their hosts, as opposed to an origin in situ. The P-T conditions of crystallisation of some lithium-rich pegmatites have been constrained by lithium aluminosilicate stability relations in combination with stable isotope or fluid inclusion methods. Experimental studies have elucidated the effects of components such as Li, B, P and F, which are common in some classes of pegmatites, to liquidus relations in the hydrous haplogranite system. Experimentation has also provided corroboration of an old concept of pegmatite crystallisation—that pegmatites owe their distinctive textures and mineral/chemical zonation to relatively rapid crystallisation of melt from the margins inwards at conditions far from the equilibrium (i.e. from supercooled liquids). The origin of aplites, whether alone, layered, or paired with pegmatites, remains an active area of research. Studies of fluid inclusions, crystal-vapour equilibria and wallrock alteration have helped to define the timing and compositions of vapour phases in pegmatites and to aid in the economic evaluation of deposits.

Description: :The current underlying assumption in most geochemical studies of granitic rocks is that granitic magmas reflect their source regions. However, the mechanisms by which source rocks control the intensive and compositional parameters of the magmas remain poorly known. Recent experimental data are used to evaluate the ‘source rock model’ and to discuss controls of (1) redox states and (2) the Sr isotopic compositions of granitic magmas.Experimental studies have been performed in parallel on biotite-muscovite and tourmaline-muscovite leucogranites from the High Himalayas. Results under reducing conditions ( = FMQ – 0·5) at 4 kbar and variable suggest that the tourmaline-muscovite granite evolved under progressively more oxidising conditions during crystallisation, up to values more than four log units above the FMQ buffer. Leucogranite magmas thus provide an example of the control of redox conditions by post-segregation rather than by partial melting processes.Other experiments designed to test the mechanisms of isotopic equilibration of Sr during partial melting of a model crustal assemblage show that kinetic factors can dominate the isotopic signature in the case of source rocks not previously homogenised during an earlier metamorphic event. The possibility is therefore raised that partial melts may not necessarily reflect the Sr isotopic composition of their sources, weakening in a fundamental way the source rock model.

Description: Zavitokichnus fusiformisn. igen. et n. isp. occurs in Lower Cretaceous (Valanginian to Hauterivian) limestones of the Fatric Superunit in the Western Carpathians. Typical cross sections of this more or less spiral trace fossil are sometimes U–O–C–S-shaped. In cross-section the trace fossil passes from a simple linear form, and spreads to a wider rolled-up or rolled-out form and then it returns to a linear trace. Spreite-like lamellae are distinguishable on several cross-section examples. The trace fossil was produced by a deposit feeder and it might be classified as a fodinichnion.Z. fusiformisco-occurs with trace fossil associations ofZoophycos,Chondrites,Planolites,HormosiroideaandPalaeophycusin carbonate sediments of a deep-seated ramp along the margin of the Fatric intrashelf basin.

Description: A specific type of granitoid, referred to as sanukitoid (Shirey & Hanson 1984), was emplaced mainly across the Archaean–Proterozoic transition. The major and trace element composition of sanukitoids is intermediate between typical Archaean TTG and modern arc granitoids. However, among sanukitoids, two groups can be distinguished on the basis of the Ti content of the less differentiated rocks of the suite: high- and low-Ti sanukitoids. Melting experiments and petrogenetic modelling show that they may have formed by either (1) melting of mantle peridotite previously metasomatised by felsic melts of TTG composition, or (2) by reaction between TTG melts and mantle peridotite (assimilation). Rocks of the sanukitoid suite were emplaced at the Archaean–Proterozoic boundary, possibly marking the time when TTG-dominated granitoid magmatism changed to a more modern-style, arc-dominated magmatism. Consequently, the intermediate character of sanukitoids is not only compositional but chronological. The succession of granitoid magmatism with time is integrated in a plate tectonic model where it is linked to the thermal evolution of subduction zones, reflecting the progressive cooling of Earth: (1) the Archaean Earth’s heat production was high enough to allow the production of large amounts of TTG granitoids formed by partial melting of recycled basaltic crust (‘slab melting’); (2) at the end of the Archaean, due to the progressive cooling of the Earth, the extent of slab melting was reduced, resulting in lower melt:rock ratios. In such conditions the slab melts can be strongly contaminated by assimilation of mantle peridotite, thus giving rise to low-Ti sanukitoids. It is also possible that the slab melts were totally consumed in reactions with mantle peridotite, subsequent melting of this ‘melt-metasomatised mantle’ producing the high-Ti sanukitoid magmas; (3) after 2·5 Ga, Earth heat production was too low to allow slab melting, except in relatively rare geodynamic circumstances, and most modern arc magmas are produced by melting of the mantle wedge peridotite metasomatised by fluids from dehydration of the subducted slab. Of course, such changes did not take place exactly at the same time all over the world. The Archaean mechanisms coexisted with new processes over a relatively long time period, even if they were subordinate to the more modern processes.

Description: Electron backscatter diffraction (EBSD) is used to determine the detailed crystallographic orientation of calcite crystals of craniid brachiopods in the context of shell ultrastructure. Sections of shells of two Recent species, Novocrania anomala and Novocrania huttoni, are analysed to provide 3D crystallographic patterns at high spatial resolution. The c-axis of semi-nacre calcite crystals is oriented parallel to the laminae that define the ultrastructure of the secondary layer. This orientation differs from that of rhynchonelliform calcitic brachiopods where the c-axis is perpendicular to the length of morphological fibres and to the shell exterior.

Description: This study describes bioerosion traces ascribed to either predation or endo- and epibiont activity in twenty assemblages from the Mediterranean region and Paratethys, spanning in age from Eocene to Recent. Statistical analysis of the distribution of bioerosion traces among genera and assemblages revealed that there is higher drilling predation intensity on smaller species. Larger species seem to be primarily affected by non-drilling predators. Greatest variety in types of bioerosion could be related to species’ ecology and body size. Both major categories of bioerosion (etchings and traces of predatory activity) vary considerably among samples. Different genera show significant differences in the frequency of different bioerosion types. Shell size seems a major factor contributing to these differences.

Description: The lower Meitan Formation (Floian, upper Lower Ordovician) at the Dajiaosi section, Zunyi District, northern Guizhou Province of South China, contains a moderately rich and diverse Sinorthis Fauna, with 22 species attributable separately to 15 families and seven orders of brachiopods. The fauna can be differentiated into three associations: the Paralenorthis serica, the Sinorthis typica, and the Tarfaya intercalare associations. These occupied a relatively wide palaeoecological range from lower BA3 to upper BA2 settings and from silty to clay substrate conditions. In South China, the Early Ordovician brachiopod radiation was marked by the diversification of the orthide-dominated Sinorthis Fauna, which first appeared and diversified in the middle part (relatively deep water) of the Upper Yangtze Platform in the Didymograptellus eobifidus Biozone, but rapidly declined in the succeeding Corymbograptus deflexus and basal Azygograptus suecicus biozones. During the latest Floian and early Dapingian, it expanded into comparatively shallow-water settings in the onshore direction (Changning area, southern Sichuan), offshore carbonate platform (Yichang, Hubei), and areas adjacent to the submergent Qianzhong Arch (such as the Zunyi area). The first appearance datum of the Sinorthis Fauna in shallower-water settings generally postdates that in the deeper-water environment in the central Upper Yangtze Platform, probably as a result of the fauna tracking a favoured BA3 setting during a gradual marine transgression.

Description: Siphonotretid brachiopod distribution in the Lower Ordovician of the Prague Basin is described and discussed; the new species Alichovia cometa is established. The preservation and distribution of the group indicate a preference for shallow-water offshore, high-energy environments: siphonotretids are abundant shelly fossils in the shallower-water, mostly marginal sites of the basin but are absent in the deeper parts. Their environmental limits were complementary to those of the graptolites; siphonotretids co-occur with epiplanktonic graptolites at only a few localities. Deeper offshore deposits with biramose dichograptids and diverse, generally delicate benthic dendroids lack siphonotretids. Indeed, siphonotretids were intolerant to dysoxia and preferred a well-aerated siliciclastic, firm sea floor in the shallower parts of the basin. They were tolerant to suspended coarser detritus, using their spines which functioned as an effective sieve. The density and ramification of spines indicate three sifting strategies. Suggested life styles above the seabed on elevated surfaces (algae, sponges, dendroids) are not supported by direct evidence. They more likely lived within more supple and soft matrices (sponges, tufts of algae) stabilised by their spines, with the spinose basket maintaining free space for inhalant and exhalant currents.

Description: This paper describes a terebratelloid articulate brachiopod, Gyrothyris williamsi sp. nov., based on 95 specimens from seamounts on the Lord Howe Rise, Coral Sea, SW Pacific Ocean. The new species is attributed to Gyrothyris on the basis of (a) morphological and growth trajectory similarities; (b) phylogenetic analyses of an alignment of DNA sequence (∼2900-sites) obtained from nuclear-encoded small- and large-subunit ribosomal RNA genes (SSU and LSU); and (c) the presence of a distinctive, two-part deletion in the LSU gene. It is distinguished morphologically from Gyrothyris mawsoni and its subspecies by both internal and external morphology and by its isolated geographical distribution, which extends the patchy, known range of this genus to an area some 2000 km north of its previous northern limit around New Zealand. Phylogenetic analyses of the rDNAs and of mitochondrial cox1 gene sequences (663 sites) confirm previous indications that the New Zealand endemic terebratelloid genera form a clade (Neothyris (Calloria, Gyrothyris, Terebratella), but the position of Terebratella with respect to Calloria and Gyrothyris remains weakly established. These sequences disagree inexplicably about the closeness of the relationship between Neothyris parva and N. lenticularis. Analyses of the first sequences from Calloria variegata, a species restricted to the Hauraki Gulf, New Zealand, are consistent with the possibility that it originated locally, and recently, from C. inconspicua. Magellania venosa from S. America/Falklands joins with Antarctic Magellania fragilis and M. joubini to form an rDNA clade that excludes Terebratalia as the putative sister-group of the New Zealand terebratelloid clade. The cox1 (but not the rDNA) sequences of the New Zealand clade pass a test for clock-like rates of evolution, and maximum likelihood pairwise distances suggest that if genetic isolation between the ancestor of Antarctic Magellania and the last common ancestor of the New Zealand terebratelloid clade was initiated by separation of the Antarctic and New Zealand plates ∼90 Mya, isolation from M. venosa was initiated earlier, perhaps ∼145 Mya. However, in the simple phylogenetic reconstruction presented here from cox1 sequences, S. American and Antarctic Magellania spp. do not yield a well-supported clade, perhaps because of differences in base composition.

Description: Though little is known of the graptoloid reproductive mechanism, graptolites with putatively sac-like appendages, supposedly ovarian vesicles, have been known from the Moffat Shales Group, Southern Uplands, Scotland, for over 150 years. Locally, these co-occur with isolated, two-dimensional, discoidal or ovato-triangular fossils. In the 1870s, Nicholson interpreted these isolated fossils as being graptoloid ‘egg-sacs’ detached from their parent and existing as free-swimming bodies. He assigned them to his genus Dawsonia, though the name was pre-occupied by a trilobite, and named four species: D. campanulata, D. acuminata, D. rotunda and D. tenuistriata. A reassessment of Nicholson’s type material from the Silurian of Moffatdale, Scotland, and from the Ordovician Lévis Formation of Quebec, Canada, shows that Dawsonia Nicholson comprises the inarticulate brachiopods Acrosaccus? rotundus, Paterula? tenuistriata and Discotreta cf. levisensis, the tail-piece of the crustacean Caryocaris acuminata and the problematic fossil D. campanulata. Though D. campanulata resembles sac-like graptolite appendages, morphometric analysis reveals the similarity to be superficial and the systematic position of this taxon remains uncertain. There is no definite evidence of either D. campanulata or sac-like graptoloid appendages having had a reproductive function.

Description: A number of approaches to interpreting geochemical data for categorising and correlating metabentonites are presented. Bulk rock geochemistry, combined with data from cognate minerals, provides the most robust means of characterising an altered ash bed. While it is not possible at present to identify one single parameter which can uniquely fingerprint an altered ash bed, it is possible to come close to an unique fingerprint, or ‘Golden Spike’, by utilising an array of geochemical parameters comprising bulk rock element ratios, Rare Earth Elements, Sr and Nd isotopic ratios in apatite, composition of sandine phenocrysts, and the chemical composition of melt inclusions.

Description: An alteration zone at the base of a Carboniferous lava flow on the island of Bute resembles modern and ancient weathering profiles, but it is ‘upside down’ in the sense that alteration is most intense at the base and decreases upwards. Values for a Chemical Index of Alteration (CIA) increase from fresh lava and corestones downwards to fine-grained structureless material. The altered material is Mg- and Cu-rich, possibly as a result of migration of these elements from underlying sediments. Rare earth elements (REE) display considerable and systematic mobility and fractionation. In general heavy rare earth elements (HREE) are concentrated during alteration, whereas the light rare earth elements (LREE) are lost. Mobility of the REE appears to be related to atomic weight, with La (the lightest REE) being the most depleted through to Lu, which is the most concentrated REE in the highly altered material. Similar systematic fractionation is shown by some weathering profiles developed on mafic igneous rocks. Movement of water into the volcanic rocks was probably driven by a steep thermal gradient between the hot lava and its sedimentary substrate.

Description: A review is presented of the genera and species, both living and fossil, comprising the Superfamily Kraussinoidea and of their geographic and stratigraphic distributions. Two new fossil species are described, Kraussina chilensis and Megerlina miracula, from the Miocene of Chile and New Zealand respectively. The ontogenetic development of the brachidium in each of the four kraussinid genera is described and on the basis of recognised differences it is proposed that a new taxonomic subdivision is made that places Megerlia in a new subfamily Megerliinae, distinct from the other three genera in the existing subfamily Kraussininae Dall. The loop ontogeny of kraussinoids is compared with that of other terebratellidines and a phylogenetic linkage between kraussinoids and laqueoids is suggested.

Description: The size of the protegulum on the valves of Early Palaeozoic rhynchonelliform brachiopods has previously been used as a criterion to assign various taxa to either a planktotrophic or a lecithotrophic larval mode. A more reliable quantitative character is evaluated in juvenile samples of two genera from the Ordovician Mifflin Formation, the presumed planktotrophic orthid Doleroides pervetus and the presumed lecithotrophic atrypid Protozyga nicolleti, using the coefficient of variation in measurements of protegular size. The Doleroides sample has a significantly higher coefficient of variation than does the Protozyga sample. The results are in accord with observations on living marine invertebrates which indicate that at settlement (when mineralisation of the protegular mantle in brachiopods occurs), planktotrophic larval size is more variable than larval size in lecithotrophs because of fluctuations in food availability and duration of the planktonic phase. This result confirms that protegular size variance can be used to make inferences about larval trophic mode.

Description: A revision of the Lower Devonian dalmanitid trilobites of the Prague Basin (Czech Republic) is presented. The subfamily Odontochilinae Šnajdr is considered a synonym of Dalmanitidae Vogdes. Twenty-one previously and five newly described (three in open nomenclature) species and subspecies occur in the Prague Basin from the lowermost Pragian (one problematic specimen possibly comes from the uppermost Lochkovian) to the Lower Emsian; the last questionable record is from the Upper Emsian. The species have been assigned to four genera and subgenera: Odontochile Hawle & Corda, Reussiana Šnajdr, Zlichovaspis (Zlichovaspis) Přibyl & Vaněk and Zlichovaspis (Devonodontochile) Šnajdr. These trilobites are considered as scavengers or opportunistic predators, living most of their lives as vagrant benthos burying in the muddy substrate to find organic remains. The first undoubted adult-like meraspid specimens are described.

Description: With their extensive fossil record and shells of stable low-Mg calcite, rhynchonelliform brachiopods are attractive sources of climate information via seawater temperature proxies such as stable oxygen isotope composition. In Terebratalia transversa (Sowerby) there is a progression towards oxygen isotope equilibrium in the calcite of the innermost secondary layer. This study confirms the lack of any vital effects influencing oxygen isotope composition of T. transversa, even in specialised areas of the innermost secondary layer. Calcite Mg/Ca ratio is another potential seawater temperature proxy, that has the advantage of not being influenced by salinity. Mg concentrations measured by electron microprobe analyses indicate that there is no concomitant decrease in Mg concentration towards the inner secondary layer, associated with the progressive shift towards oxygen isotope equilibrium. Mg distribution is heterogeneous throughout the shell and correlates with that of sulphur, which may be a proxy for organic components, suggesting that some of the Mg may not be in the calcite lattice. It is essential therefore, to determine the chemical environment of the magnesium ions to avoid any erroneous temperature extrapolations in brachiopods or any other calcite biomineral.

Description: In geochemical diagrams, granitoids define ‘trends’ that reflect increasing differentiation or melting degree. The position of an individual sample in such a trend, whilst linked to the temperature of equilibration, is difficult to interpret. On the other hand, the positions of the trends within the geochemical space (and not the position of a sample within a trend) carry important genetic information, as they reflect the nature of the source (degree of enrichment) and the depth of melting. This paper discusses the interpretation of geochemical trends, to extract information relating to the sources of granitoid magmas and the depth of melting.%Applying this approach to mid-Archaean granitoids from both the Barberton granite–greenstone terrane (South Africa) and the Pilbara Craton (Australia) reveals two features. The first is the diversity of the group generally referred to as ‘TTGs’ (tonalites, trondhjemites and granodiorites). These appear to be composed of at least three distinct sub-series, one resulting from deep melting of relatively depleted sources, the second from shallower melting of depleted sources, and the third from shallow melting of enriched sources. The second feature is the contrast between the (spatial as well as temporal) distributions and associations of the granites in both cratons.

Description: A detailed account of the morphology and ontogeny of the late Middle Cambrian crustacean †Henningsmoenicaris scutula is presented. Ten successive ontogenetic stages could be recognised in the material collected from various localities in Sweden. Morphogenetic changes include the development of a pair of stalked lateral eyes and the increase in the number and size of appendages and their setal armature. Notably, early stages lack ‘proximal endites’ on all post-antennular appendages; such a spine-bearing endite has previously been thought to appear simultaneously on these limbs. In †H. scutula a single functional endite appears on the third limb in an advanced stage; an additional endite appears on the second limb and, subsequently, further endites appear on more posterior limbs. Furthermore, a single specimen of †Sandtorpia vestrogothiensis gen. et sp. nov. is described. Based on this new information and data of other ‘Orsten’ taxa, particularly those assigned already to the early evolutionary lineage of Crustacea, a small-scale computer-based phylogenetic analysis was performed. This resolved the basal branchings of Crustacea s. l. as follows: †Oelandocarididae (=†Oelandocaris oelandica+†H. scutula+†S. vestrogothiensis)+(†Cambropachycopidae (=†Goticaris longispinosa+†Cambropachycope clarksoni)+ (†Martinssonia elongata+Labrophora (=†Phosphatocopina+Eucrustacea))). Plotting ontogenetic data on the phylogram and comparing the ground pattern at every node led to the detection of three peramorphic heterochronic events in the evolutionary lineage towards Eucrustacea.

Description: The Sixth Hutton Symposium on the Origin of Granites and Related Rocks was held on July 2–6, 2007 at the University of Stellenbosch, South Africa, founded on granite, nestled at the feet of towering mountains and fringed by the rolling winelands of the Western Cape. This Special Issue opens with Master’s historical account of how the Cape granites influenced 18th and early 19th century thinking on the origins of these rocks. The fascinating fact is that the granites of the Western Cape were apparently the first intrusive granites recognised outside Britain. The balance of the volume contains a collection of research papers derived from the meeting and illustrates some of the important directions in which granite research may be evolving. One of the characteristics of the papers and talks presented at the meeting was that there seemed to be some shift in interest, away from the crust as a source of granitic magmas and towards mantle rocks that have been metasomatised by subduction-zone fluids or melts. Nevertheless, the crust still holds pride of place as the cradle of granite genesis.

Description: Several authors have proposed that granitic melt accumulation and transport from the source region occurs in networks of connected melt-filled veins and dykes. These models envisage the smallest leucosomes as ‘rivulets’ that connect to feed larger dykes that form the ‘rivers’ through which magma ascends through the sub-solidus crust. This paper critically reviews this ‘rivulets-feeding-rivers’ model. It is argued that such melt-filled networks are unlikely to develop in nature, because melt flows and accumulates well before a fully connected network can be established. In the alternative stepwise accumulation model, flow and accumulation is transient in both space and time. Observations on migmatites at Port Navalo, France, that were used to support the existence of melt-filled networks are discussed and reinterpreted. In this interpretation, the structures in these migmatites are consistent with the collapse and draining of individual melt batches, supporting the stepwise accumulation model.

Description: The Murotomisaki Gabbroic Intrusion is a sill-like layered gabbro emplaced in sedimentary strata of Tertiary age in southwest Japan. The zoning (including resorption structures) and the compositional variations of plagioclase from throughout the intrusion were studied, and it was found that the zoning pattern may be classified into four types, which may well correlated with the hosting rock types, the mode of occurrences and their stratigraphic positions in the intrusion. The plagioclase zoning was successfully decoded, and the sequence of events that took place during the magmatic differentiation was deduced and further interpreted in the context of a stratified basal boundary layer slowly ascending in a solidifying magma body. It was shown that various layered structures – modal layering, podiform gabbroic pegmatites and anorthositic layers – observed in the Murotomisaki Gabbro were formed within the moving basal boundary layer by flushing of H2O-rich fluid and fractionated silicate melts from below. By the fluxing of hydrous fluids, plagioclase crystals preferentially dissolved and then melt fraction increased in the basal boundary layer. Under these circumstances, plagioclase-rich fractionated melts diapirically segregated from the crystal pile. Calcic plagioclases, which are out of equilibrium in the central part of the intrusion, may have originated from the basal boundary layer in this manner.

Description: Regional scale biotite and cordierite-bearing granites (s.l.) in the Variscan of the Central Iberian Zone (CIZ) are spatially closely associated with cordierite-rich nebulites and cordierite-bearing two-mica granites, and with cordierite-rich high grade hornfelses and cordieritites (〉60% cordierite) that are relatively common in the aureoles of these granites. Building on published field evidence, petrological data are presented which, combined with new chemical and isotopic (Sr–Nd) modelling, indicate that the cordierite-bearing granites cannot be derived by simple anatexis of regional sedimentary protoliths; but the data are consistent with a process of reactive assimilation that involves the interaction of biotite granite magma with high-grade host rocks ranging from cordierite nebulites to andalusite-bearing cordieritites. The contribution of the postulated cordierite-rich contaminants to the diversity of cordierite granite compositions is modelled using the compositions of regional Lower Cambrian–Upper Neoproterozoic metasedimentary rocks that are generally chemically mature (CaO very rarely exceeds 1·4%). These rocks include specific horizons in which extreme chemical alteration is attributable to sediment reworking during eustatic falls in sea level. Such compositions may account for the presence of the high concentrations in Al that later produced cordieritites. Fractional crystallisation is also important, particularly in generating the more evolved cordierite granite and cordierite biotite muscovite granite compositions. Although assimilation in situ is normally regarded as a minor contributor volumetrically to evolving plutons, in this instance the emplacement of large volumes of granite magma into a high-T–low-P environment significantly increased the potential for reactive assimilation.

Description: Microzarkodina is a genus of mainly Middle Ordovician conodonts that has its centre of distribution in Baltoscandia, and much less commonly occurs in southern China, Australia, Argentina and Laurentia. In Baltica a series of species, Microzarkodina russica n. sp., M. flabellum, M. parva, M. bella, M. hagetiana and M. ozarkodella, established themselves successfully. The succession of species ranges from just below the base of the Middle Ordovician (M. russica) to the upper part of the Middle Ordovician (M. ozarkodella). The species are frequently used for biostratigraphical purposes. The largely contemporaneous species Microzarkodina bella and M. hagetiana probably both evolved from M. parva and mostly occurred in separate areas. Microzarkodina ozarkodella probably evolved from M. hagetiana. This present investigation is based on a total of 94,208 elements, collected from 20 sections and one drill-core site in Sweden, one drill-core site and one outcrop in Estonia and two sections in the St Petersburg area in Russia. The Microzarkodina apparatus probably consisted of 15 or 17 elements: four P, two or four M and nine S elements. The S elements include different Sa, Sb1, Sb2, and Sc element types.

Description: The regionally metamorphosed, Riphean–Cambrian Argyll Group Dalradian rocks of NW Achill Island, western Ireland are disposed in a large-scale, regionally west-facing, tight, recumbent F2 curvilinear fold, with which two ductile shear zones are associated. Clasts in conglomerates within the Dalradian sequence that are deformed by the shear zones preserve evidence for a constrictional overprint of earlier plane strain as the fold became curvilinear, while stretched clasts maintained a constant orientation as the hinge curvilinearity developed. During the constrictional overprint a crenulation fabric, S2b, overprinted a penetrative foliation, S2a, in the shear zones. The S2b has an orientation that varies systematically with that of the fold hinge. It is inferred that, although the S2b surfaces initiated as a dip-slip fabric, there was an increasing degree of strike slip on these surfaces as the fold hinge approached parallelism with the direction of tectonic transport. It is possible that many curvilinear folds have an early history involving plane strain, but that increasing constrictional strain is intrinsic to the later stages of their development.

Description: The early Cambrian Chengjiang biota in Yunnan, southern China has yielded many articulated exoskeletons of the spiny redlichiid trilobite Eoredlichia intermedia, of which some have their appendages exceptionally well preserved. Both of the paired uniramous antennae of a medium-sized holaspis consist of 46–50 short segments (articles), each of which bears a fine spine near its inner edge. Behind the antennae there are twenty-one pairs of biramous limbs: three pairs are situated underneath the cephalon, one pair underneath each of the fifteen thoracic segments, and probably three pairs underneath the small pygidium. The endopod consists of a broad basis and seven podomeres, of which the last is divided into three terminal spines. The exopod is blade-like, and according to one interpretation, is dorsally hinged to the basis of the endopod; an alternative suggestion being that both the endopod and exopod are split from the basis, the latter being independent and not forming part of the endopod. The exopod has a prominent anterior rim, and possesses about forty long filaments along the posterior margin, and short setae along the rounded distal lobe. The basic appendage features of the redlichiid trilobites, and likewise the gut, are comparable to those of other known Cambrian polymerid trilobites that belong to more distantly related clades.

Description: Animals can respond in three main ways to environments that change and deteriorate. They can cope with their inherent physiological flexibility, evolve or adapt to new conditions, or migrate away to areas consistent with survival. The main factors dictating survival are the rates of change and capacities of species to cope. Articulated brachiopods are known as a group that employ low energy solutions and exhibit slow rates in life histories characters, especially in growth and metabolic rates. This way of life carries with it a range of consequences, including poor abilities to raise metabolic rates to perform work. This brings with it poor abilities to cope with elevated temperature. Slow growth, deferred maturity and low feeding rates make them less likely to produce novel adaptations compared with other marine invertebrate taxa. Being largely restricted to hard substrata and low energy supply environments also makes it harder for them to migrate to new sites compared to other groups, because of the large geographic distances between colonisable areas. The only clear area they may have that gives them advantages is in the flexibility of their larval release strategies, where settlement can be from very short, millimetre-scale distances to hundreds of kilometres.

Description: The loop configuration, structure and growth are studied in a series of megathiridoid dorsal valves ranging in length from 0·7 to 6 mm. The structure of the posterior and anterior sectors of the descending lamellae and their relationships with the socket ridges, dorsal septa and dorsal valve floor, are studied, illustrated and described. Detailed studies show that internal structures display great variability. In the megathiridoids the lamellae of more typical terebratulide loops are replaced by partially developed loops in early developmental stages, resulting in a mix of free sections of loop together with sections where the lophophore rests directly on the epithelium of the valve floor. Comments on the ecology of the megathiridoid species are included and relate their unusually wide gapes, which position their lophophores fully accessible to the open sea, with their strong lophophore attachments and loss of typical free loops. The new genus Joania (type species Terebratula cordata Risso, 1826) is erected for those Argyrotheca which, although having a typical megathiridoid brachidium, differ in their adult crural development, their narrow hinge line, their prominent cardinal process, their characteristic dorsal median septum and their tuberculate radial ridges terminating anteriorly in tubercles.

Description: Spire-bearing brachiopods formally comprise four different rhynchonelliform orders. A calcified spiral brachidium (presumably supporting a spirolophe when alive) and variable median fold and sulcus (probably aiding separation of incurrent from excurrent flows) are peculiar characteristics they all share. Inferences regarding feeding current systems for these extinct taxa have long remained controversial. Two rival models (the Williams–Ager model and the Rudwick–Vogel model) have been developed, each of which has gained supporters as well as critics over the years. In this present paper they are both contrasted and reassessed on the basis of available evidence, together with a new approach that combines: (a) a morpho-functional analysis applying the plankton net as a suitable seston-collecting paradigm; (b) a review of actualistic data showing that all extant spirolophes are functionally inhalant (irrespective of water entering the valves laterally or not); (c) an evaluation of known outcomes from flume experiments yielding consistent empirical results where gaping shells are oriented transversally and dorsally upcurrent; and (d) a reappraisal of the distributions of certain epizoobionts and endosymbionts revealing compatible patterns. The evidence thus accumulated supports the main conclusion that, in most groups (with laterally tapering spiralia), the inhalant current was located medially with the exhalant currents on either side; only in atrypides (with centrally to dorsally tapering spiralia) does the reverse situation appear to have occurred.

Description: The interpretation of pedicle soft tissue preservation in a unique brachiopod specimen of Wenlock (Silurian) age from Herefordshire, western England, is re-assessed. Bethia serraticulma, assigned originally to the Orthida, is more probably a member of the Strophomenida (Plectambonitoidea). The supposed pedicle structure is more plausibly a weakly mineralised pedicle sheath, which is a common morphological and functional development in the early ontogeny of a number of Palaeozoic brachiopod lineages.

Description: Two Ordovician plectambonitoid genera, Alwynella and Grorudia, occur in drill core sections of Latvia in the East Baltic, and in exposures and loose blocks on the Swedish Island of Öland in the Baltic Sea. The new material confirms differences between the two taxa that are assigned herein to separate families, Alwynellidae fam. nov. and Grorudiidae Cocks & Rong, 1989. In particular, the undercut cardinalia separates Alwynella from Grorudia and indicates its proximity to the sowerbyellids. The genus Grorudia, which is externally similar to Alwynella, is more closely related to the palaeostrophomenines. A new species Grorudia morrisoni sp. nov. is established in the East Baltic. The specimens from Öland are included tentatively within the genus Grorudia due to lack of interiors. Both Alwynella and Grorudia were confined to deeper-water facies in the Baltic palaeobasin, within successions ranging in age from latest Mid (late Llanvirn) to earliest Late Ordovician (mid Caradoc).

Description: The Late Carboniferous whip spiderGraeophonus anglicusPocock, 1911 (Arachnida: Amblypygi), is redescribed on the basis of the holotype and nine other specimens all preserved in sideritic nodules from the British Middle Coal Measures of Coseley, Staffordshire, UK. This species is clearly basal with respect to most living whip spiders, expressing numerous plesiomorphic character states and can be referred to both the suborder Paleoamblypygi and the ‘living fossil’ family Paracharontidae (with one Recent species), the latter based on an explict character of dorsal spination on the pedipalp femur. This suggests that crown-group Amblypygi originated by at least the mid-Palaeozoic.

Description: Oceanic islands offer outstanding ‘model systems' for investigating long-term dynamics between human populations and their ecosystems. Whilst the state factors involved in human-environment dynamics on islands are often simpler than on continents, the same essential processes are involved. This paper applies a comparative approach to understanding the reciprocal interactions between a set of four Polynesian island cases (Tikopia, Mangaia, Mangareva, Hawaiian Islands), over time scales of between one and three thousand years (kyr). In all cases, the island ecosystems were colonised by Polynesian populations derived from the same ancestral culture, with similar socioeconomic patterns. However, the ecosystems vary significantly in scale, geologic age, and other characteristics. Comparing the historical trajectories of these human–environment dynamics, as revealed by archaeological and palaeoecological study, provides insights into the relative impact of humans on pristine island ecosystems, the influence of environment on ecosystem vulnerability, and the ways in which societies have modified their economies, sociopolitical structures, and other aspects of culture in response to long-term environmental changes.

Description: Archaeological investigations demonstrate that peoples first settled the Caribbean islands approximately 6000–7000 years ago. At least four major, and multiple minor, migrations took place over the next millennia by peoples from Mesoamerica and South America who practised various subsistence strategies and had different levels of technology. For decades, researchers have been interested in investigating how these groups adapted to and impacted insular environments through time. This paper combines archaeological, palaeoecological, historical, and modern biological data to examine the effects of humans on Caribbean island ecosystems using a historical ecology approach. By synthesising a wide range of data sources, we take a human/nature dialectical perspective to understanding how peoples adapted to and modified their environments. The data suggest that earlier foraging/fishing Archaic groups (ca. 6000–3000 BP), who used a stone tool and shell technology and transported few, if any non-indigenous plants or animals, still impacted island landscapes as evidenced by bird and sloth extinctions. As more advanced ceramic making horticulturalists entered the Antillean chain around 2500 BP, there is an observable change to island environments as a result of forest clearance, overexploitation of both terrestrial and marine resources, and growing populations. Palaeoecological and palaeoenvironmental records also suggest, however, that an increased moisture regime during the late Holocene probably led to a decrease in near-shore salinity and heavier sediment and nutrient loads in rivers. These conditions would have been exacerbated by land clearance for agriculture, leading to coastline progradation, increased turbidity, and mangrove development resulting in changes to the availability of resources for humans on some islands. Although prehistoric peoples in the Caribbean were certainly impacting their environments, it was not until Europeans arrived and population centres grew that intensive and widespread degradation of island landscapes and resources occurred. Modern ecological studies, along with historical and archaeological data, indicate that hundreds of species have been driven to extinction or extirpation – many others have significantly diminished in number, especially within the last two millennia.

Description: This two-part special issue, Brachiopod Research into the Third Millennium, is published in honour of Sir Alwyn Williams (8 June 1921–4 April 2004). One section deals with fossil brachiopods while the other concentrates on aspects of modern material. The special issue comprises 23 invited papers, all dealing with current brachiopod research. In producing this special issue we honour the immense contribution made by the late Alwyn Williams to this field of research.

Description: A new genus of pentameride brachiopod, Costilamnulella, described herein from the middle Ashgill of Sweden, is particularly interesting in that the ventral valve possesses both an interarea and a palintrope. Brachiopods normally have one or the other, but both structures do occur amongst the pentamerides, e.g. in Clorinda, Anastrophia and Brevilamnulella. The presence of these structures in three separate pentameride families indicates the persistence of the more primitive condition of an interarea whilst the more advanced palintrope is becoming established. Costilamnulella is interpreted as a restricted side line of the main development of the more widely occurring Brevilamnulella, which survived the end-Ordovician event and may have shared its ancestry with the ubiquitous Clorinda.

Description: New cranial material of the ‘rhynchodipterid’ lungfishSoederberghia groenlandicafrom Upper Devonian (Famennian) strata in East Greenland is described. Previously unknown structures identified here include components of the hyoid arch (ceratohyal, hypohyal) and the lower jaw (prearticular, dentary). Earlier interpretations of the cheek and mandible ofSoederberghiaare reconsidered in the light of new fossil specimens. Some of the difficulties in assessing the homologies between cheek bones inSoederberghiaand those of other lungfishes stem from confusion over the arrangement inRhynchodipterus, and a revised interpretation of this genus is proposed. The single infraorbital bar found inSoederberghiaprobably originates, in part, from an expanded bone 10 (quadratojugal) of the kind found inGriphognathus. Hypotheses that posit ‘rhynchodipterid’ polyphyly seem unlikely in light of a set of derived cranial characters that define a coherent radiation of long-snouted, denticle-bearing lungfishes known from the Late Devonian. The hypothesis presented here places ‘rhynchodipterids’ as a paraphyletic grade with respect to fleurantiids.Rhynchodipterus,Soederberghia, and fleurantiids form a clade to the exclusion of the species ofGriphognathus.G. minutidensis the sister taxon to this apical group, whileG. sculptaandG. whiteiare more remote from it.

Description: We present the first record of Holocene and Pleistocene environmental change derived from the chemical and stable-isotope composition of a tropical cave guano sequence from Makangit Cave in northern Palawan (Philippines). The 180 cm sequence of guano, derived predominantly from insectivorous bats and birds, consists of two distinct units. An upper section of reddish-brown oxidised guano to 110 cm was deposited since the mid-Holocene while a lower section of black, reduced guano was deposited through the Last Glacial Maximum (LGM) to 〉30 000 BP. Carbon-isotope (δ13C) values in guano deposited during the LGM are as high as −13·5‰ indicating that a C4-dominated grassland existed in the area around the cave at this time. Guano δ13C values of − 25‰ to − 28‰ suggest that this open vegetation was replaced by C3-dominated closed tropical forest, similar to that of the present, by the mid-Holocene. The results suggest that the climate of northern Palawan was substantially drier at the LGM than is currently the case.

Description: A total of 47 genera, in 80 species, of Mississippian, or Lower Carboniferous, crinoids are evaluated from 61 localities in Scotland based on a modern update of the literature, study of museum collections, and new field work. Among the 80 species, 76 are considered valid, with eight requiring new combinations of genus and species names. In addition, one species is considered anomen dubium, and three can only be assigned at the generic level. Asbian faunas have a moderate generic richness of 13. Brigantian faunas have the highest generic richness at 40. Arnsbergian faunas have the lowest generic richness at three. There are no Mississippian crinoid faunas in Scotland older than Asbian. The dominant Brigantian crinoid faunas occur in limestones and shales in the shallow shelf marine intervals of Yoredale-style cyclothems. The Asbian faunas occur in shallower, nearshore mudstones and limestones of Yoredale-style cyclothems. The late Viséan (Brigantian) fauna is more similar in overall taxonomic composition to the global record for the Serpukhovian rather than the Viséan. This reflects the establishment of the Late Palaeozoic Crinoid Macroevolutionary Fauna, dominated by advanced cladid crinoids, by the end of the Viséan in Scotland.

Description: The 1800 km-long Aleutian archipelago represents a model ecosystem to track human–environmental interactions across space and through time. Defining the southern margin of Beringia across which much of the early peopling of the Americas occurred, the Aleutians present a 9000 year record of human occupation in the eastern part of the island chain, and more than 3000 years in the west. Molecular evidence demonstrates: (1) that Aleuts shared common ancestry with Chukchi and Siberian Eskimos of Chukotka; (2) the original patterns of migration into the Aleutian islands were from the Alaskan peninsula in a westward direction with no evidence for island-hopping from Kamchatka; and (3) a highly significant statistical relationship between geography and genetics, based on mtDNA sequences, was observed despite previous population disruption. Historically, the Aleutian region is a rich ecotone, with ocean fisheries, abundant populations of large marine mammals, thick kelp forests, complex near-shore ecosystems and intertidal zones, spawning streams, and a highly diverse avian fauna. Each of these environments and resources has been pivotal in shaping the adaptive strategies of human occupants of the island chain since the initial colonisation of the Aleutians from the Alaskan Peninsula. In turn, Holocene human immigration, prehistoric cultural adaptations and subsequent historic events have had reciprocal impacts on the natural systems of the Aleutians.