ALBERT

Description: Estimating site productivity in irregular structures is complicated by variations in stand density, structure, composition in mixed stands, and suppression experienced by subordinate trees. Our objective was to develop an alternate to site index (SI) and demonstrate its application in models of individual-tree and stand growth. We analyzed coast redwood (Sequoia sempervirens (Lamb. ex D. Don) Endl.) tree and stand growth in a grid of 234 permanent sample plots covering a 110 ha study area in north coastal California. Partial harvesting created a mosaic of densities and openings throughout the 60-year-old redwood-dominated forest. Redwood SI was a poor predictor of volume increment (VI) per hectare among redwood in each plot over two decades after harvest. A new index of redwood basal area increment (BAI) productivity, calculated using inventory data for all stems in even-aged stands and the oldest cohort of multiaged stands, was a stronger predictor of VI. Diameter increment of individual redwood trees correlated strongly with stand density and the new BAI index. Forest managers should expect widely divergent responses following partial harvesting in crowded even-aged stands, with the greatest response coming from dominant redwoods with long crowns retained in areas with low residual stand density and high BAI index.

Description: Stand development of a subalpine forest in the Colorado Front Range following a ca. 15-ha blowdown was examined by analyzing tree population age structures and radial growth patterns. The stand studied was initiated by a fire at the start of the 18th century and was dominated by a dense population of lodgepole pine (Pinuscontorta Dougl. var. latifolia Engelm.) at the time of blowdown in 1973. Before the blowdown, the subcanopy was characterized by abundant subalpine fir (Abieslasiocarpa (Hook.) Nutt.) and scarce Engelmann spruce (Piceaengelmannii (Parry) Engelm.). Comparison with an adjacent control stand, affected only slightly by the blowdown, indicates that new seedling establishment following the blowdown was slight. Instead, the response was dominated by the release of the subcanopy fir and spruce, resulting in acceleration of the successional replacement of lodgepole pine by these shade-tolerant species. Given the 〉300 years required for an old-growth fir and spruce stand to develop following catastrophic fire, the likelihood of a major canopy disturbance in the form of blowdown and (or) lethal insect attack is high and should be explicitly incorporated into general explanations of stand development of subalpine forests in the southern Rocky Mountains.

Description: A synthesis was carried out to examine Alaska’s boreal forest fire regime. During the 2000s, an average of 767 000 ha·year–1 burned, 50% higher than in any previous decade since the 1940s. Over the past 60 years, there was a decrease in the number of lightning-ignited fires, an increase in extreme lightning-ignited fire events, an increase in human-ignited fires, and a decrease in the number of extreme human-ignited fire events. The fraction of area burned from human-ignited fires fell from 26% for the 1950s and 1960s to 5% for the 1990s and 2000s, a result from the change in fire policy that gave the highest suppression priorities to fire events that occurred near human settlements. The amount of area burned during late-season fires increased over the past two decades. Deeper burning of surface organic layers in black spruce ( Picea mariana (Mill.) BSP) forests occurred during late-growing-season fires and on more well-drained sites. These trends all point to black spruce forests becoming increasingly vulnerable to the combined changes of key characteristics of Alaska’s fire regime, except on poorly drained sites, which are resistant to deep burning. The implications of these fire regime changes to the vulnerability and resilience of Alaska’s boreal forests and land and fire management are discussed.

Description: Broad-scale fire regime modelling is frequently based on large ecological and (or) administrative units. However, these units may not capture spatial heterogeneity in fire regimes and may thus lead to spatially inaccurate estimates of future fire activity. In this study, we defined homogeneous fire regime (HFR) zones for Canada based on annual area burned (AAB) and fire occurrence (FireOcc), and we used them to model future (2011–2040, 2041–2070, and 2071–2100) fire activity using multivariate adaptive regression splines (MARS). We identified a total of 16 HFR zones explaining 47.7% of the heterogeneity in AAB and FireOcc for the 1959–1999 period. MARS models based on HFR zones projected a 3.7-fold increase in AAB and a 3.0-fold increase in FireOcc by 2100 when compared with 1961–1990, with great interzone heterogeneity. The greatest increases would occur in zones located in central and northwestern Canada. Much of the increase in AAB would result from a sharp increase in fire activity during July and August. Ecozone- and HFR-based models projected relatively similar nationwide FireOcc and AAB. However, very high spatial discrepancies were noted between zonations over extensive areas. The proposed HFR zonation should help providing more spatially accurate estimates of future ecological patterns largely driven by fire in the boreal forest such as biodiversity patterns, energy flows, and carbon storage than those obtained from large-scale multipurpose classification units.

Description: Analysis of the frequency of past moderate and high-intensity disturbances has been hindered in forests of complex age structure by methodological problems. A methodology is proposed for developing a disturbance chronology in such stands by identifying the probable date of canopy accession for each sample tree. Canopy accession dates are based on an evaluation of radial growth pattern and early growth rates of existing canopy trees. Canopy disturbance intensity is defined as the percentage of sample trees with canopy accession events in each decade. Rotation periods for disturbances of various intensities are calculated from the chronology. The method was evaluated using 893 increment cores from 70 plots in northern hardwood stands of western Upper Michigan. The estimated average disturbance rate for all plots and decades was 5.7–6.9% of land area per decade, with an implied average canopy tree residence time of 145–175 years. These estimates are similar to those obtained by on-site estimates of canopy tree residence time and studies in the literature on the rate of gap formation. Problems in radial increment analysis and possible solutions are discussed.

Description: This study investigates geographic patterns of genetic variation in aspen (Populus tremuloides Michaux.) spring phenology with the aim of understanding adaptation of populations to climatic risk environments and the practical application of guiding seed transfer. We use a classical common garden experiment to reveal genetic differences among populations from western Canada and Minnesota, and we present a novel method to seamlessly map heat-sum requirements from remotely sensed green-up dates. Both approaches reveal similar geographic patterns: we find low heat-sum requirements in northern and high-elevation aspen populations, allowing them to take full advantage of a short growing season. High heat-sum requirements were found in populations from the central boreal plains of Saskatchewan and Alberta, and populations from Minnesota exhibit moderately low heat-sum requirements for budbreak. Analysis of corresponding climate normal data shows that late budbreak is strongly associated with the driest winter and spring environments, which suggests selection pressures for late budbreak due to both frost and drought risks in early spring. We therefore caution against long-distance seed transfer of Minnesota provenances to the boreal plains of Alberta and Saskatchewan. Although such transfers have been shown to increase tree growth in short-term field tests, this planting material may be susceptible to exceptional spring droughts.

Description: This paper attempts to address the question of how the value of the forest, the land, and the standing timber should be determined under the generalized Faustmann formula when the beginning and ending value of the land may be different. First, the formulas to determine the value of the forest and the land under such a situation were derived. These formulas were then used to separate the value of the trees from that of the forest. A comparison of the correctly determined valuations of the land against those obtained through a frequently used approximation method showed that at interest rates commonly used, the approximation method overestimates the land value and underestimates the value of the standing timber. Sensitivity analyses showed that higher future land value tends to affect the value of the land less at the beginning of the rotation and more at the end. Its impact on the value of the standing timber may or may not be affected, depending on whether the optimal harvest age is affected or not. Higher final harvest value and higher interest rate affect both the value of the land and the value of the standing timber throughout the entire rotation. Lastly, higher regeneration at the beginning of the rotation simply re-allocates the forest value between that of the land and that of the standing timber, reducing the former while increasing the latter.

Description: Large eddy simulation (LES) based computational fluid dynamics (CFD) simulators have obtained increasing attention in the wildland fire research community, as these tools allow the inclusion of important driving physics. However, due to the complexity of the models, individual aspects must be isolated and tested rigorously to ensure meaningful results. As wind is a driving force that can significantly dictate the behavior of a wildfire, the simulation of wind is studied in the context of a particular LES CFD model, the Wildland–urban interface Fire Dynamics Simulator (WFDS). As WFDS has yet to be tested extensively with regard to wind flow within and above forest canopies, a study of its ability to do so is carried out. First, three simulations are conducted using periodic boundary conditions. Two of these assume a spatially heterogeneous forest and one models wind downstream of a canopy edge. Second, two simulations are conducted with specified “inflow” conditions using two inflow profiles: one static and one dynamic (driven by a precursor simulation). Using periodic boundary conditions, the model is found to generate profiles of mean velocity and turbulent statistics that are representative of experimental measurements. The dynamic inflow scenario is found to perform better than the static case.

Description: Liming and wood-ash addition have long been used to attenuate the effects of acidic deposition on forest soils with the goal of promoting tree growth. We performed quantitative meta-analyses of treatment studies from managed forest ecosystems to assess general tendencies of effects of treatment on seven selected measures of performance thought to reasonably reflect the effects of Ca-addition treatment. We retrieved over 350 independent trials from 110 peer-reviewed liming and wood-ash addition studies that were integrated to determine soil pH, base saturation (BS), tree foliar Ca concentration, tree growth, ectomychorrhizae root colonization, soil C-to-N ratio, and microbial indices. The results were quantified through three separate meta-analysis effect size metrics: unweighted relative values and two weighted metrics, Hedges’ d and ln R. A surprising number of treatment trials (22%–85%) reported no significant effect, and soil pH and foliar Ca appeared more responsive to liming than to wood-ash addition, whereas BS and tree growth appeared more responsive to wood-ash addition. For six of the seven parameters, estimated mean effect sizes were similar in magnitude and positive in direction for all three meta-analysis metrics. Regression tree optimal models explained 38% of the variation in pH, 47% of the variation in BS, 51% of the variation in foliar Ca concentration, and 26% of the variation in tree growth. The largest predictors of effect size, within our selected group, were as follows: soil type for pH; soil type, trial duration in years, and species (hardwood or softwood) for BS; treatment dose and type for foliar Ca concentration; and trial duration, initial soil pH, and tree species for tree growth. This analysis shows that Ca additions are not universally beneficial and provides insight into when Ca additions to forest soils are likely to be most effective.

Description: There is increasing recognition that forestry provides a low cost and robust means of climate change abatement through carbon sequestration and substitution. However, current understanding of forest ecosystem carbon exchange and forest–atmosphere interactions are often inadequately characterized by existing empirical growth models with resulting poor representation for regional extrapolations. In this paper, we describe the parameterisation and independent validation, against both eddy covariance and forest growth experimental data, of a process-oriented model 3PGN to provide assessments of carbon sequestration of Sitka spruce (Picea sitchensis (Bong.) Carrière) plantations across Scotland. In comparison with eddy covariance measurements, the model predicted all of the major annual carbon fluxes, i.e., gross primary production (PG), net ecosystem production (PE), and ecosystem respiration (RE), with biases lower than 10%. At a monthly time step, only PG and PE were accurately estimated, whereas RE was not. At longer time scales (i.e., several decades), the model reliably represented the major patterns of the carbon balance. Soil type was identified as the important factor influencing site productivity; fertilization practices did not alter long-term site nutritional status. The analyses also highlighted the potential impact of carbon loss from carbon-rich soils, which can result in differences between optimal rotation length for carbon sequestration and for timber production.

Description: Understanding of the natural factors that lead to complex changes in forest ecosystems is limited. Worldwide, there are only a few forests as pristine and isolated as the Sierra de La Laguna in the southernmost range of the arid Baja California, Mexico. Its outstanding trait as a model system is that anthropogenic stressors are notably absent, which facilitates the study of natural ecological processes of the forest because separating human-induced ecological changes from natural ones is not a simple matter. In this study, we sampled sites and defined vegetation units on the basis of dominance of the canopy by the main tree species. We identified three forest types: the pine and encino forests that occupy the higher areas and the roble forest at lower elevations. For each living tree in the sampling plots, we measured height, canopy coverage per tree, diameter at breast height, as well as the amount of deadwood, leaf litter, and abundance of young trees. A succesional competition occurs between Pinus and Quercus sensu lato; we conclude that the encino forest represents a climax condition, the pine type represents an early succesional stage, and the roble forest type is a simple climax community.

Description: Duff fires (smouldering in fermentation and humus forest floor horizons) and their consequences have been documented in fire-excluded ecosystems but with little attention to their underlying drivers. Duff characteristics influence the ignition and spread of smouldering fires, and their spatial patterns on the forest floor may be an important link to the heterogeneity of consumption observed following fires. We evaluated fuel bed characteristics (depths, bulk densities, and moisture) of duff in a long-unburned longleaf pine (Pinus palustris Mill.) forest and corresponding spatial variation across 100to 103m scales. Fermentation and humus horizon depths both varied (∼100% coefficient of variation) but with moderate to strong spatial autocorrelation at fine scales. Fermentation bulk density varied less than humus bulk density, which varied considerably at fine scales. Fermentation horizons held more moisture (average 49%–172%) and were much more variable than humus following rainfall, which remained stable and relatively dry (average 28%–62%). Humus moisture was moderately autocorrelated at fine scales, but fermentation moisture was highly variable, showing no evidence of spatial autocorrelation under dry, intermediate, or wet conditions. Observations from this study highlight the underlying spatial variability in duff, informing future sampling and fire management efforts in these long-unburned coniferous forests.

Description: Forest structure, as measured by the physical arrangement of trees and their crowns, is a fundamental attribute of forest ecosystems that changes as forests progress through suc;cessional stages. We examined whether LiDAR data could be used to directly assess the successional stage of forests by determining the degree to which the LiDAR data would show the same relative ranking of structural development among sites as would traditional field measurements. We sampled 94 primary and secondary sites (19–93, 223–350, and 600 years old) from three conifer forest zones in western Washington state, USA, in the field and with small-footprint, discrete return LiDAR. Seven sets of LiDAR metrics were tested to measure canopy structure. Ordinations using the of LiDAR 95th percentile height, rumple, and canopy density metrics had the strongest correlations with ordinations using two sets of field metrics (Procrustes R = 0.72 and 0.78) and a combined set of LiDAR and field metrics (Procrustes R = 0.95). These results suggest that LiDAR can accurately characterize forest successional stage where field measurements are not available. This has important implications for enabling basic and applied studies of forest structure at stand to landscape scales.

Description: Sustainable forest management requires timely, detailed forest inventory data across large areas, which is difficult to obtain via traditional forest inventory techniques. This study evaluated k-nearest neighbor imputation models incorporating LiDAR data to predict tree-level inventory data (individual tree height, diameter at breast height, and species) across a 12 100 ha study area in northeastern Oregon, USA. The primary objective was to provide spatially explicit data to parameterize the Forest Vegetation Simulator, a tree-level forest growth model. The final imputation model utilized LiDAR-derived height measurements and topographic variables to spatially predict tree-level forest inventory data. When compared with an independent data set, the accuracy of forest inventory metrics was high; the root mean square difference of imputed basal area and stem volume estimates were 5 m2·ha–1 and 16 m3·ha–1, respectively. However, the error of imputed forest inventory metrics incorporating small trees (e.g., quadratic mean diameter, tree density) was considerably higher. Forest Vegetation Simulator growth projections based upon imputed forest inventory data follow trends similar to growth projections based upon independent inventory data. This study represents a significant improvement in our capabilities to predict detailed, tree-level forest inventory data across large areas, which could ultimately lead to more informed forest management practices and policies.

Description: Characteristics of annual rings are reliable indicators of growth and wood quality in trees. The main objective of our study was to model the variation in annual ring attributes due to intensive silviculture and inherent regional differences in climate and site across a wide geographic range of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Ring specific gravity and ring width of Douglas-fir were examined at five long-term Levels-Of-Growing-Stock (LOGS) installations, three in the US and two in Canada, covering a latitudinal gradient between 43°N and 50°N. At each location, increment cores were collected from replicated plots with three levels of stocking: control (unthinned), lightly thinned (70% basal area retention), and heavily thinned (30% basal area retention). X-ray densitometry analysis provided ring specific gravity and width profiles for 5676 rings from 134 trees. The reduction of stand density through repeated entries resulted in decreased ring specific gravity and increased ring width. A four-parameter mixed-effects logistic model was used to predict ring specific gravity using cambial age, stand density (as number of stems per hectare), and two climatic variables: average temperature from March to May and total precipitation from April to August. A three-parameter mixed-effects logistic model was used to predict ring width using cambial age, stand density (as stand density index), and total climatic moisture deficit of June and July. Both models indicated significant site differences that were included in the models through indicator variables. Ring specific gravity increased slightly with increasing average temperature from March to May and decreasing total precipitation from April to August. Predictions of ring specific gravity of Douglas-fir appear to be more sensitive to changes in temperature compared with changes in precipitation.

Description: Radial patterns of modulus of elasticity (MOE) were examined for white spruce (Picea glauca (Moench) Voss) and trembling aspen (Populus tremuoides Michx.) from 19 mature, uneven-aged stands in the boreal mixedwood region of northern Alberta, Canada. The main objectives were to (1) evaluate the relationship between pith-to-bark changes in MOE and cambial age or distance from pith; (2) develop species-specific models to predict pith-to-bark changes in MOE; and (3) to test the influences of radial growth, relative vertical height, and tree slenderness (tree height/DBH) on MOE. For both species, cambial age was selected as the best explanatory variable with which to build pith-to-bark models of MOE. For white spruce and trembling aspen, the final nonlinear mixed-effect models indicated that an augmented rate of increase in MOE occurred with increasing vertical position within the tree. For white spruce trees, radial growth and slenderness were found to positively influence maximum estimated MOE. For trembling aspen, there was no apparent effect of vertical position or radial growth on maximum MOE. The results shed light on potential drivers of radial patterns of MOE and will be useful in guiding silvicultural prescriptions.

Description: A method for the production of multiple clonal plantlets of Pinuscaribaea var. hondurensis Morlet, P. oocarpa Schiede, and P. tecunumanii Equiluz and Perry (P. patula Schiede and Deppe ssp. tecunumanii (Equiluz and Perry) Styles) from juvenile sources is described. The procedure is based on the spontaneous production of axillary shoots following shoot elongation. Much interclonal variation exists with respect to expiant multiplication. The addition of activated charcoal (0.1%) stimulates shoot elongation, especially in recalcitrant clones of P. caribaea. Axillary bud initiation may be stimulated by 6-benzylaminopurine, but levels above 1 μM can result in failure of induced buds to form shoots. Rooting occurs at about 50% in nonsterile peat–perlite medium without auxin pretreatment. Rooted plantlets can be readily hardened and will grow normally in the glasshouse.

Description: In 2009, the Swiss National Forest Inventory (NFI) turned from a periodic into an annual measurement design in which only one-ninth of the overall sample of permanent plots is measured every year. The reduction in sample size due to the implementation of the annual design results in an unacceptably large increase in variance when using the standard simple random sampling estimator. Thus, a flexible estimation procedure using two- and three-phase regression estimators is presented with a special focus on utilizing updating techniques to account for disturbances and growth and is applied to the second and third Swiss NFIs. The first phase consists of a dense sample of systematically distributed plots on a 500 m × 500 m grid for which auxiliary variables are obtained through the interpretation of aerial photographs. The second phase is an eightfold looser subgrid with terrestrial plot data collected from the past inventory, and the third and final phase consists of the three most recent annual subgrids with the current state of the target variable (stem volume). The proposed three-phase estimators reduce the increase in variance from 294% to 145% compared with the estimator based on the full periodic sample while remaining unbiased.

Description: Over the last decades, continuous signs of sugar maple ( Acer saccharum Marsh.) dieback in stands of northeastern North America have promoted the experimentation of corrective measures to restore sugar maple vitality. To verify the hypothesis that K–Mg antagonism may have limited the full response of sugar maple to dolomitic lime application in a previous experiment (CaMg(CO3)2, 12% Mg), two Ca fertilizers (CaCO3 and CaSO4·2H2O), having negligible Mg content, were applied at rates of 1, 2, and 4 t Ca·ha–1 on sugar maple trees adjacent to the limed area. After 3 years, most of the foliar nutrient concentrations of treated trees were improved, particularly Ca, for both Ca fertilizers, in line with published ranges for healthy sugar maple trees, except for Mg. Moreover, no persistent nutrient antagonism was observed. The crown dieback rate of treated sugar maple was ≤5.8% after 3 years, while it reached 12% for the controls. Also, relative basal area growth showed that both Ca sources can improve growth rate. Growth response following Ca treatments was, however, lower than for the former lime experiment after the same period of time. In this context, our results suggest that Mg nutrition could be more important for sugar maple in this ecosystem than initially thought.

Description: Soil compaction often limits conifer regeneration on sites degraded by landings and roads, but inadequate understanding of the relationship between compaction and tree growth could lead to inappropriate soil conservation and rehabilitation efforts. We tested liquid and plastic limits, oxidizable organic matter, total carbon, particle size distribution, and iron and aluminum oxides on soil samples collected from five forest experiments in interior British Columbia. These data were used to estimate soil maximum bulk density (MBD) and relative bulk density (RBD); our objective was to relate RBD to tree growth. Height of interior Douglas-fir ( Pseudotsuga menziesii var. glauca (Bessin) Franco) was limited when RBD was 〉0.72. For lodgepole pine ( Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) and hybrid white spruce ( Picea glauca (Moench) Voss × Picea engelmannii Parry ex Engelm.), RBDs of 0.60–0.68 corresponded to maximum height, whereas RBDs of 0.78–0.87 appeared to limit height growth. The presence of surface organic material mitigated compaction and was often associated with lower RBD. Our results illustrate the usefulness of RBD to assess compaction and suggest that soil rehabilitation should be considered on disturbed sites where soil RBD is 〉0.80.

Description: Piceaglauca var. albertiana (S. Brown) Sarg. shoot phenology and water relation parameters were monitored monthly for 1 year. Seedlings were kept outdoors, well watered, and exposed to seasonal changes in temperature and day length. Changes in shoot water relation parameters corresponded with changes in phenology. During spring, shoot elongation, osmotic potential at saturation, and turgor loss point were least negative, −1.30 and −1.56 MPa, respectively, whereas bulk modulus of elasticity at full turgor was at its highest, 22 MPa. Both osmotic potential at saturation and turgor loss point were most negative, −2.01 and −2.73 MPa, respectively, during late winter just before bud break. Shoot dry weight fraction was at its lowest, 0.33 g dry wt./g shoot weight and maximum symplastic water per unit weight of shoot tissue was at its highest, 3.99 g H2O/g dry wt., during spring shoot elongation. Number of osmoles of solute per kilogram shoot dry weight was highest during spring and late summer shoot elongation phases, 1.03 and 0.91 osmol/kg dry wt., respectively. These data suggest that P. glauca seedlings do not adjust well to dry site conditions.

Description: A novel “varying-centroid” method is presented for predicting whole-tree, aboveground stem volume (i.e., bole plus branch volume) to bole volume ratios from changes in the centroid of tree bole volume associated with branching of the bole. The method was derived from a simple fractal-like tree model based on a conceptualization of tree branching architecture by Leonardo da Vinci. The method recognizes that the centroid of bole volume (the point at which one half of bole volume is above and one half is below) is always lower than the centroid of whole-tree volume and that shifts in the centroid of bole volume should be predictably related to the size of a tree’s crown. The method assumes that branch-displaced bole volume profiles can be compared with reference bole profiles that are not significantly influenced by branching, at the centroid of bole volume, and that the magnitude of bole centroid displacement predicts the branch volume necessary to cause it. When the method was applied to hardwood trees representing diverse species, sizes, and stand conditions across Michigan, the centroid of bole volume was found to vary predictably with measurable tree crown attributes and bole plus branch wood to bole wood volume ratios were generally predicted within 10% of the true value using the new method.

Description: Because of the importance of seed surface area, volume, and fill to hydraulic and thermal exchanges with the soil, mechanistic simulation of seed physiological processes associated with tree migration dynamics and the spread of invasive species require accurate equations to model seed shape. Seed dimensions have previously been described with measurements of the three principal axes, assuming an implied single ellipsoid. However, conifer seeds often exhibit anisotropy that results from bilaterally symmetric pairing on cone scales. We developed a method for measuring and modeling conifer seed shape as a sum of 2jpartial ellipsoids fused at their equators, where j = 0, …, 3. We demonstrate the use of the methods in the study of shape characteristics of ponderosa pine (Pinus ponderosa P.& C. Lawson) seeds from four families in Montana and among commercial lots of ponderosa pine, lodgepole pine (Pinus contorta Dougl. ex Loud.), and Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco). The shapes of 92%, 73%, and 47% of seeds in commercial lots studied had eight unique ellipsoids when classified with 1%, 5%, and 10% difference classification rules, respectively. Ponderosa pine seeds with longer minor axes were less well filled with storage reserves. Three-dimensional surface areas of lodgepole and ponderosa pine were approximately 2 and 3.4 times larger, respectively, than previously reported one-sided surface areas.

Description: Small-scale disturbance is a significant process in all major forest biomes. Some silvicultural practices, particularly group selection harvesting, intend to emulate natural small-scale disturbance by harvesting small clearcuts in the continuous forest. We conducted a meta-analysis on the effects of small-scale harvesting on North American breeding forest birds. We extracted species richness and relative abundance of several functional bird groups and guilds from published studies and compared them between gap-dominated and unlogged forest as a function of forest type and the size and age of the gap. The abundance of many bird groups was higher in the gap-dominated than in the continuous forest. Species preferring interior parts of the forest had the most negative association with the presence of gaps but this relationship was not statistically significant. Abundances of many bird groups increased with increasing gap size, while its effect on abundance of some bird groups disappeared quickly. Our review suggests that silvicultural practices that bring about small gaps do not negatively affect the abundances of most forest birds and often even enhance it. However, more studies are needed to examine optimal size and abundance of gaps in a forest and whether emulated small-scale disturbance effectively mimics natural processes.

Description: Crown dimensions and sapwood area near crown base were measured on 189 Douglas-fir trees in southwestern Oregon. Sapwood areas were interpolated or extrapolated to crown base with a sapwood taper function. Various transformations of crown length and crown radius (as well as crown base stem diameter as a surrogate for crown diameter) were assessed for their ability to predict crown base sapwood area. Regression analyses indicated that no single untransformed variable was a good predictor of sapwood area at crown base, but that combinations representing conic surface area performed quite well. Given the consistently strong relationship between total leaf area and sapwood area at crown base, conic surface area should accurately reflect total leaf area and relative photosynthetic potential of the tree. Gross crown dimensions and crown base sapwood area prove complementary in a forest modeling context, owing to the structural appeal of the former and the physiological appeal of the latter.

Description: A system of equations for compatible prediction of total and merchantable volumes that allows for different definitions of tree volume was developed in this study. The use of the developed system will allow the conversion and subsequent comparison of results from forest inventories using different definitions of tree volume (e.g., including or not the top material of the tree and (or) the stump, inside or outside bark). The compatibility between taper, total volume, and volume ratio equations is ensured by properly integrating the taper equation. The diameter under the bark at any height is modeled with the Demaerschalk taper equation, and the corresponding diameter over the bark is obtained by assuming that bark thickness is also modeled with Demaerschalk’s function. The set of equations that has contemporaneous cross-equation error correlation (known as nonlinear seemingly unrelated regression equations) was fit using nonlinear joint generalized least squares regression. The predictive ability was evaluated using an independent data set. The system is consistent and performs well when applied to maritime pine ( Pinus pinaster Ait.) trees in Portugal, showing better performance than do other total volume equations for maritime pine used in the latest Portuguese national forest inventories.

Description: Loblolly pine (Pinus taeda L.) is a major plantation species grown in the southern United States, producing wood having a multitude of uses including pulp and lumber production. Specific gravity (SG) is an important property used to measure the quality of wood produced, and it varies regionally and within the tree with height and radius. SG at different height levels was measured from 407 trees representing 135 plantations across the natural range of loblolly pine. A three-segment quadratic model and a semiparametric model were proposed to explain the vertical and regional variations in SG. Both models were in agreement that a stem can be divided into three segments based on the vertical variation in SG. Based on the fitted models, the mean trend in SG of trees from the southern Atlantic Coastal Plain and Gulf Coastal Plain was observed to be higher than in other physiographical regions (Upper Coastal Plain, Hilly Coastal Plain, northern Atlantic Coastal Plain, and Piedmont). Maps showing the regional variation in disk SG at a specified height were also developed. Maps indicated that the stands in the southern Atlantic Coastal Plain and Gulf Coastal Plain have the highest SG at a given height level.

Description: Subsistence harvesting and wild food production by Athabascan peoples is part of an integrated social–ecological system of interior Alaska. We describe effects of recent trends and future climate change projections on the boreal ecosystem of interior Alaska and relate changes in ecosystem services to Athabascan subsistence. We focus primarily on moose, a keystone terrestrial subsistence resource of villages in that region. Although recent climate change has affected the boreal forest, moose, and Athabascan moose harvesting, a high dependence by village households on moose persists. An historical account of 20th century socioeconomic changes demonstrates that the vulnerability of Athabascan subsistence systems to climatic change has in some respects increased while at the same time has improved aspects of village resilience. In the face of future climate and socioeconomic changes, communities have limited but potentially effective mitigation and adaptation opportunities. The extent to which residents can realize those opportunities depends on the responsiveness of formal and informal institutions to local needs. For example, increases in Alaska’s urban population coupled with climate-induced habitat shifts may increase hunting conflicts in low-moose years. This problem could be mitigated through adaptive co-management strategies that project future moose densities and redirect urban hunters to areas of lower conflict.

Description: Forest compartments are usually delineated according to artificial or natural boundaries and usually include portions of different strata. While volume estimation of each stratum can be performed from field plots located within each stratum, volume estimation in portions of the stratum may be problematic owing to the small number (or even the absence) of plots falling in those portions. If upper canopy heights from airborne laser scanning are available at the pixel level for the whole survey area, these data are used as auxiliary information. A ratio model presuming a proportional relationship between transformed heights (e.g., power of heights) and volumes at the pixel level is adopted to guide estimation. From this model, the volume within any portion of the survey area is estimated as the proportionality factor estimate multiplied by the total of transformed heights in that portion. This estimator is considered from the model-based, design-based, and hybrid perspectives. Variances and their estimators are derived under the three approaches together with the corresponding confidence intervals. The volume estimator and the variance estimators are checked from the design-based point of view by a simulation study performed on a real forest in northwestern Italy. An application to a public forest estate in the same zone is performed.

Description: Forest management planners usually treat potential fire loss estimates as exogenous parameters in their timber production planning processes. When they do so, they do not account for the fact that forest access road construction, timber harvesting, and silvicultural activities can alter a landscape’s vegetation or fuel composition, and they ignore the possibility that such activities may influence future fire losses. We develop an integrated fire and forest management planning methodology that accounts for and exploits such interactions. Our methodology is based on fire occurrence, suppression, and spread models, a fire protection value model that identifies crucial stands, the harvesting of which can have a significant influence on the spread of fires across the landscape, and a spatially explicit timber harvest scheduling model. We illustrate its use by applying it to a forest management unit in the boreal forest region of the province of Alberta in western Canada. We found that for our study area, integrated fire – forest management planning based on our methodology could result in an 8.1% increase in net present value when compared with traditional planning in which fire loss is treated as an exogenous factor.

Description: White poplar ( Populus alba L.) is a native species in Europe, but its growth potential is largely unknown. The general objectives of our study were to determine the impact of contrasted environments across Europe and the influence of parental characteristics on the growth potential of an intraspecific F1 white poplar family originating from a cross between parents native from the south and the north of Italy. The growth of the family was monitored at three sites located in the north of Italy, in central France, and in the southern United Kingdom. The family showed a highly superior productivity in Italy. A pronounced plasticity among sites was found for the male parent only. Indeed, for this parent, the highest growth was observed in northern Italy, its area of origin. A positive heterosis was observed mainly in France and in the United Kingdom. Broad-sense heritability values were moderate in most cases. However, the growth of the family was in some cases superior to the one of several other interspecific hybrid families growing under the same conditions, underlying the poorly known growth potential of such intraspecific hybrids for biomass production under European conditions.

Description: The natural phenotypic variation in Chinese white poplar (Populus tomentosa Carr.), which is distributed across a wide geographical area of northern China (30°N–40°N, 105°E–125°E), is a potential source of beneficial variation for poplar breeding. Thirteen traits related to growth, leaf, and wood properties were quantified in 460 P. tomentosa individuals grown in a common garden plot. There was considerable range-wide phenotypic variation in all traits across individuals according to the patterns of ANOVA among hierarchical groups (populations and regions, respectively). A clear sexual dimorphism for seven traits was examined. In total, 32 trait–trait phenotypic correlations (P ≤ 0.05), 10 trait–geographical factor correlations (P ≤ 0.05), and a highly interrelated structure network were identified, which was further supported by principal component analysis (PCA). These associations can be used in multiple-trait selective breeding programs for advantageous phenotypic traits. A hierarchical cluster analysis was used to classify four groups (southeastern, central, northeastern, and southwestern populations) among the natural populations using these 13 phenotypic traits. This study provides important perspectives into the use of direct breeding to potentially improve economic traits and provides a starting point for genome-wide association studies in P. tomentosa in the near future.

Description: Declines of whitebark pine ( Pinus albicaulis Engelm.) have occurred across much of the species’ range over the last 40 years due to mountain pine beetle outbreaks and white pine blister rust infection. Management efforts to stem these declines are increasing, yet the long-term success of whitebark pine depends on the species itself adapting to the modern environment. Natural regeneration will be a critical part of this process. We examined patterns in natural whitebark pine regeneration as related to the biophysical environment in sixty 0.1 ha plots in Montana, Idaho, and Oregon. Whitebark pine regeneration was present in 97% of our plots and varied widely in density from 0 to 17 000 seedlings/ha and 0 to 2680 saplings/ha. Using nonparametric correlation analysis and ordination techniques, we found whitebark pine regeneration abundance was unrelated to stand age but significantly related to several biophysical site characteristics, including positive relationships with elevation and canopy tree mortality caused by mountain pine beetle and negative relationships with moisture availability, temperature, and subalpine fir importance. Our findings indicate that whitebark pine is regenerating in many areas and that the widespread mortality from recent mountain pine beetle outbreaks may provide suitable settings for whitebark pine regeneration given sufficient seed sources.

Description: We evaluated spruce budworm ( Choristoneura fumiferana (Clem.)) outbreak effects in nine study areas (60–86 ha each) located in the boreal forest of eastern Quebec (Canada). In each area, spruce budworm outbreak effects were measured from vegetation plots, dominant canopy and understory tree age structures, retrospective analysis of aerial photographs, defoliation records, and host tree growth reductions (dendrochronology). Large-scale synchronous outbreaks were detected across the region around the years 1880, 1915, 1950, and 1980. Overall, contrarily to what was expected for a region where host species (balsam fir ( Abies balsamea (L.) Mill.), Picea spp.) content is relatively high, these spruce budworm outbreaks seemed to have a relatively minor influence on stand dynamics, with the exception of the most recent outbreak (1980). This outbreak resulted in major stand mortality in the southern part of the region and favored the establishment of extensive tracts of young even-aged stands with few residual mature trees. This very abrupt increase in outbreak severity compared with earlier outbreaks, perhaps due to climatic or random factors, suggests that historical trends in successive outbreak severity should be extrapolated very cautiously and that the study of several outbreak cycles is needed to establish a range of natural variability that can be used to develop an ecosystem forest management strategy.

Description: Effects of short-term flooding on stomatal conductance, net photosynthesis, and water status of sweet gum (Liquidambarstyraciflua L.) seedlings were studied under controlled environment conditions. Flooding for 9 days induced partial stomatal closure, resulting in significant declines in transpiration and net photosynthesis. The response to flooding was rapid with an average daily stomatal conductance declining from a preflood level of 0.43 cm•s−1 to 0.26 cm•s−1 by 24 h after flooding began (40% reduction). The average preflooding daily net photosynthesis was reduced from 13.7 to 10.2 mg CO2•dm−2•h−1 (25% reduction) during the same period and the average daily stomatal conductance and net photosynthesis for the 9th day of flooding were reduced by 70 and 77%, respectively, compared with preflood levels. The leaf xylem pressure potential measurements, however, indicated that water deficits did not develop as a result of flooding. Partial stomatal reopening 3 days after termination of flooding was noted with an average daily stomatal conductance approaching 63% of the preflood levels and an average daily net photosynthesis reaching 46% of its preflood levels. Maintenance of positive net photosynthesis throughout flooding, and partial stomatal and photosynthetic recovery following drainage may account for the tolerance of sweet gum seedlings to short-term flooding.

Description: Forest managers are seeking strategies to create stands that can adapt to new climatic conditions and simultaneously help mitigate increases in atmospheric CO2. Adaptation strategies often focus on enhancing resilience by maximizing forest complexity in terms of species composition and size structure, while mitigation involves sustaining carbon storage and sequestration. Altered stand age is a fundamental consequence of forest management, and stand age is a powerful predictor of ecosystem structure and function in even-aged stands. However, the relationship between stand age and either complexity or carbon storage and sequestration, especially trade-offs between the two, are not well characterized. We quantified these relationships in clearcut-origin, unmanaged pine and aspen chronosequences ranging from 130 years in northern Minnesota. Complexity generally increased with age, although compositional complexity changed more over time in aspen forests and structural complexity changed more over time in pine stands. Although individual carbon pools displayed various relationships with stand age, total carbon storage increased with age, whereas carbon sequestration, inferred from changes in storage, decreased sharply with age. These results illustrate the carbon and complexity consequences of varying forest harvest rotation length to favor younger or older forests and provide insight into trade-offs between these potentially conflicting management objectives.

Description: Disturbances such as fire and harvesting shape forest dynamics and must be accounted for when modelling forest properties. However, acquiring timely disturbance information for all of Canada’s large forest area has always been challenging. Therefore, we developed an approach to detect annual forest change resulting from fire, harvesting, or flooding using Moderate Resolution Imaging Spectroradiometer (MODIS) imagery at 250 m spatial resolution across Canada and to estimate the within-pixel fractional change (FC). When this approach was applied to the period from 2000 to 2011, the accuracy of detection of burnt, harvested, or flooded areas against our validation dataset was 82%, 80%, and 85%, respectively. With FC, 77% of the area burnt and 82% of the area harvested within the validation dataset were correctly identified. The methodology was optimized to reduce the commission error but tended to omit smaller disturbances as a result. For example, the omitted area for harvest blocks greater than 80 ha was less than 14% but increased to between 38% and 50% for harvest blocks of 20 to 30 ha. Detection of burnt and harvested areas in some regions was hindered by persistent haze or cloud cover or by insect outbreaks. All resulting data layers are available as supplementary material.

Description: Pre- and post-harvest regeneration levels were compared for Abiesbalsamea (L.) Mill. –Betulapapyrifera Marsh. –Picea spp. forests in an area of the southern clay belt of northwestern Quebec. Results revealed abundant advance softwood regeneration (mean = 65 000 stems/ha), almost entirely of Abies prior to harvest. The survey following mechanical and manual whole-tree harvesting suggested a 92% reduction of softwood regeneration and a shift from softwood to a mixed or hardwood-shrub dominated regeneration. Ninety percent of softwood seedlings collected after harvest were pre-established. Destruction of advance regeneration was generally greater on fine-textured soils. Hierarchic cluster analysis of ecological types based on softwood, hardwood, and shrub tree regeneration data as variables, revealed nine groups that could serve as a basis for operational silvicultural decision making. In general, Salix spp. and Alnusrugosa (Du Roi) Spreng. are the major competitors on poorly drained sites; Betula sp., Acerspicatum Lam., and Prunuspensylvanica L.f. dominate on thin organic deposits and coarse deposits, whereas Populustremuloides Michx. and Acerspicatum dominate on fine-textured deposits. These findings suggest that a good understanding of physical site factors can provide useful information for harvesting and silvicultural planning.

Description: Proportional basal area (Pro-B) was developed as an accurate, easy-to-use method for making uneven-aged silviculture a practical management option. Following less than 3 h of training, forest staff from a range of professional backgrounds used Pro-B in an operational-scale field study to apply single-tree selection and group selection systems in longleaf pine (Pinus palustris Mill.) stands. Field crews achieved precision levels often within 3%–5% of the 11.5 m2·ha−1 target residual basal area. By aggregating many diameter classes into only three diameter-class groups, Pro-B improves efficiency by requiring tree markers to remember only three fractions, while making a single pass through the stand. Trees of large size, specific species and with good form, broad crowns and cavities can be retained, while adjusting spacing to release residuals. Systematic quantification of marking trees for removal enables different individuals to obtain similar results. Early observations revealed encouraging levels of pine regeneration and stand development, along with continuing good volume growth rates of 3% per year. Although less certain until one or more cutting cycles are completed, these early tests indicate that a stable mature forest structure should develop, which is characterized by the presence of large trees and natural regeneration.

Description: Tree mortality because of beetle outbreaks has become substantial and widespread in conifer forests in western North America. A number of environmental and physiological factors influence patterns of mortality. Tree diversity may reduce the severity and extent of insect damage to host trees by providing associational resistance, but the existence and importance of associational resistance varies by forest type and by tree and insect species. We assessed whether plot-level tree diversity contributed to survival of Engelmann spruce (Picea engelmannii Parry ex Engelm.) following a spruce beetle (Dendroctonus rufipennis Kirby) epidemic. Our study plots comprised 2 to 5 tree species including Engelmann spruce, subalpine fir (Abies lasiocarpa (Hook) Nutt.), Douglas-fir (Pseudostuga menziesii (Mirb.) Franco), quaking aspen (Populus tremuloides Michx.), and white fir (Abies concolor (Gordon & Glend.) Hildebr.). We used a model-selection analysis to compare the importance of tree diversity with other known factors that influence spruce survival. We found lower rates of spruce survival in stands where spruce was the dominant tree species (by percent of stand density index) and higher survival in stands where nonspruce conifers (Douglas-fir, subalpine, and white fir) were dominant. We also found that tree diversity (Shannon index) did not show a positive correlation to spruce survival and that there was no additional benefit derived from the presence of aspen, which has higher phylogenetic distance from Engelmann spruce than the other trees in this study. The relationship between diversity and survival is complicated by factors that naturally co-vary with diversity, such as elevation, aspect, and stand density of spruce. Our results best support an explanation that if associational resistance does increase spruce survival during a beetle epidemic, it is due to host or resource dilution, which may be an indirect effect of higher stand diversity.

Description: Tree species distributions and diversity could be explained by rank changes in performance over multiple spatiotemporal resource gradients, i.e., resource partitioning. For 14 species planted in 45 harvest gap and closed canopy locations in a mesic northern hardwood forest community, Michigan, USA, we asked the following questions: (i) are species growth responses to light, nitrogen (N), or N form (ammonium vs. nitrate) related to their ecological distributions and phylogenies? and (ii) is there evidence of growth-based resource partitioning over measured resource gradients? Growth responses to the N form were consistent with both differences in uptake energy requirements between N forms and their availability through succession and across fertility gradients; height growth was negatively related to the species shade-tolerance score, especially in high light, i.e., shade-intolerant species responded to soil nitrate-N and shade-tolerant species responded to ammonium-N; fertile soil associated species responded to nitrate-N and infertile soil associated species to ammonium-N; and gymnosperms responded to ammonium-N and angiosperm responses varied. Modeled growth responses to resources showed only modest evidence for rank changes over resource gradients, with N contributing less to rank changes than light. Thus, growth responses to resources were accurately predicted by species ecology and (or) phylogeny; however, there was only modest support for the notion that growth-based resource partitioning underlies community-scale diversity in a northern hardwood forest.

Description: An integrated forest management optimization model was developed to calculate potential spruce budworm ( Choristoneura fumiferana Clemens) effects on forest and wood product carbon (C) from 2007 to 2057 and to evaluate potential C sequestration benefits of alternative management strategies (salvage, biological insecticide application). The model was tested using simulated spruce budworm outbreaks on a 210 000 ha intensively managed forest in northwestern New Brunswick, Canada. Under a severe spruce budworm outbreak scenario from 2007 to 2020, harvest volume and forest and wood product C storage in 2027 were projected to be reduced by 1.34 Mm3, 1.48 Mt, and 0.26 Mt, respectively, compared with the levels under no defoliation. Under the same severe outbreak scenario, implementation of salvage and harvest replanning plus a biological insecticide applied aerially to 40% of susceptible forest area, reduced harvest, forest C, and wood product C impacts by 73%, 41%, and 56%, respectively. Extrapolation of these results to all of New Brunswick suggests that a future severe spruce budworm outbreak could effectively increase total provincial annual C emissions (all sources) by up to 40%, on average, over the next 20 years. This modeling approach can be used to identify to what extent insecticide application, as a forest-C-offset project, could result in additional C storage than without forest and pest management.

Description: With a growing interest in the diversification (e.g., bioenergy, biochemicals) of the forest industry beyond the traditional product streams, concerns that higher harvest utilization levels may compromise site productivity have been heightened. This study reports on 15-year tree growth responses to varying levels of biomass removals conducted on four soil types: loamy tills, outwash sands, wet mineral, and peatlands. Experimental harvest treatments included stem-only, full-tree, full-tree chipping (a full-tree harvest with the roadside material chipped and returned to the site), and full-tree + bladed (a full-tree harvest followed by forest floor removal). Results indicated no significant effect on height growth on the loamy tills, a significant decline for the blading treatment on the sandy soils, and an increase when the blading treatment was applied to the peatland sites. At the stand level, better planted seedling survival and higher recruitment of naturals on the more extreme removal treatment (forest floor removal on sandy sites) tended to nullify any negative impacts identified in the individual-tree growth measurements. The more than doubling of the slash loading on the stem-only treatment plots compared with the full-tree plots did not result in differences in tree productivity levels between these two operational treatments. The stands, however, were just approaching crown closure by year 15, suggesting that ongoing monitoring will be required to confirm that the growth trajectories for the various harvest treatment – soil type combinations can be maintained.

Description: Previous reconstructions of the late Quaternary biogeographical history of lodgepole pine (Pinuscontorta Dougl.) have been based upon inferences from the modern geographical distribution of morphological and genetic variation. These studies have led to the widely accepted conclusion that relict populations of the Rocky Mountain subspecies of lodgepole pine (ssp. latifolia Engelm.) persisted in glacial refugia located in northwestern Canada. New fossil pollen evidence of the late Pleistocene and Holocene distribution of lodgepole pine in the western interior of Canada contradicts this view. Pinuscontorta ssp. latifolia migrated northward into Canada from refugia located south of the continental glacial limits and did not reach its northern range limits in the southern Yukon until the late Holocene.

Description: Managed coniferous forest dominates much of the black-backed woodpecker’s ( Picoides arcticus Swainson) breeding range. Despite this, little is known about the fine-scale foraging behaviour of this focal species in unburned managed forest stands in the absence of insect outbreaks. To investigate the foraging substrates used in such a habitat, we employed radio-telemetry to track a total of 27 black-backed woodpeckers. During two successive summers (2005–2006), 279 foraging observations were recorded, most of which were on dying trees, snags, and downed woody debris. Individuals frequently foraged by excavation, suggesting that in the absence of insect outbreaks the black-backed woodpecker forages mainly by drilling. The majority of foraging events occurred on recently dead snags with a mean dbh (±SE) of 18.3 ± 0.4 cm. Our results suggest that in unburned boreal forest stands, substrate diameter and decay class are important predictors of suitable foraging substrates for black-backed woodpeckers. We suggest that conservation efforts aimed at maintaining this dead-wood dependent cavity nesting species within the landscape, should endeavour to maintain 100 ha patches of old-growth coniferous forest. This would ensure the continuous production of a sufficient quantity of recently dead or dying trees to meet the foraging needs of this species.

Description: A procedure is presented for estimating the coefficients of allometric models for predicting tree component biomass. The resulting equations force the sum of the component estimates to be equal to the estimate of total biomass. An illustration of the procedure is given using published biomass data and the relationship of this procedure to previously published procedures is discussed.

Description: The identification of low-vigor trees with potential for sawlog production is a key objective of tree marking guidelines used for partial cuts in northern hardwoods. The aim of this study was to measure the impact of various vigor-related defects on the monetary value of hardwoods. To achieve this, we sampled 64 sugar maple (Acer saccharum Marshall) and 32 yellow birch (Betula alleghaniensis Britton) trees from two locations in southern Quebec, Canada. We identified over 420 defects, which were grouped into 8 categories. The trees were then harvested and processed into lumber, and the value per unit volume of each stem was calculated from the value of the product assortment (lumber, chips, and sawdust). We found that visible evidence of fungal infections (sporocarps and (or) stroma) and cracks had the largest negative influence on value in both species. A model that included these defects was almost as good at predicting value as one that included a specifically designed quality classification. A more accurate assessment of value could be achieved using wood decay assessment tools and (or) by considering site-specific variables. Results from this study showed that visual identification of fungal infections and cracks could be used to enhance tree marking guidelines for hardwoods. This would meet both the silvicultural objective of selection cuts, by removing low-vigor trees, and the wood supply objective, by improving stem quality assessment prior to harvest.

Description: Height-growth patterns for jack pine (Pinusbanksiana Lamb.) were studied using stem analyses from dominant and codominant trees on 141 plots in north central Ontario. All plots were in natural, well-stocked, even-aged stands 50 years of age or older. Data from 32 of the 141 plots were randomly selected to confirm results, the remaining 109 plots were used for computing the curves. Height-growth curves were developed using a five-parameter Chapman–Richards nonlinear regression that expressed height as a function of age and site index. A site-index prediction equation was also computed using a similar model that expressed site index as a function of age and tree height. Estimated site index using height-growth curves based on the 109 computation plots agreed closely with site index observed from stem analyses on the 32 confirmation plots. Major results were as follows: (i) height-growth curves based on breast-height age were more accurate than curves based on total age; (ii) polymorphic height-growth patterns were related to site index, becoming more curvilinear as site index increased; (iii) average height-growth patterns were similar for jack pine growing on four glacial landforms: shallow and deep moraines, outwashed glacial sands, and lacustrine clays and silts; and (iv) height-growth patterns for ages less than 50 years were very similar to patterns of the commonly used Plonski curves for jack pine site classes in Ontario, but after 50 years, height growth was somewhat better for all sites than predicted by the Plonski curves.

Description: The most common method for determining tree profile models is by fitting an analytical function to a set of sample trees. In situations where the forester has relatively little a priori knowledge about the nature of profiles, choosing the analytical form of the parametric model is a critical problem without a satisfactory solution. An alternative is to use a nonparametric approach in which the model of the profile is explicitly specified by a tabulation of diameters at discrete heights. In this paper, it is shown how the model can be determined from the sample trees by means of a classical nonparametric probability density function estimation technique. Field tests and a measure of goodness of fit are used to express how well the models match the actual stem profiles. Using the Moroccan cedar (Cedrusatlantica Manetti) tree as an example, it is shown how the new nonparametric model can be compared with any classical parametric model. The results achieved demonstrate the advantages in using a nonparametric representation of stem profiles, which, moreover, is well suited to computer calculation constraints.

Description: The use of principal components analysis to study tree stem profiles was critically analyzed during 1085 destructively sampled Douglas-fir trees and 1260 simulated trees with known geometric shapes. Interpretation about the meaning of each principal component is provided and contrasted with others in the forestry literature. Nearly identical results with both the destructively sampled and simulated trees, along with certain theoretical consideratons, indicate that the principal components are related to tree form as opposed to tree profile or taper. Therefore, principal components analysis is a useful analytical tool for stratifying trees into different form groups.

Description: New sensor-based approaches for assessing the quantity, quality, and value of timber are being developed with the goals of improving the accuracy and economics of forest measurements. One new approach is based on terrestrial laser scanning (TLS). Thirty-three plots in six radiata pine (Pinus radiata D. Don) stands were scanned using TLS. Tree locations were automatically detected. Stem profiles were measured using three methods: (i) TLS scans, (ii) Atlas Cruiser inventory procedures, and (iii) manual measurement after harvesting. Stems were optimally bucked based on log specifications and prices for Australian markets. Tree values and log product yields were estimated for the TLS data and compared with estimates based on Cruiser and actual manual measurements of stem profiles. TLS volume and value recovery were within 8% and 7%, respectively, of actual harvester recovery for five of the six stands in which it was used. Cruiser volume and value estimates were both within 4% of actual harvester recovery. Plot preparation procedures, tree characteristics, and taper equations used to model diameters on hidden stem sections affected the accuracy of automated stem detection and profile measurements for the TLS system. Improvements in data capture and analytical procedures should improve the accuracy of TLS-based volume and value estimates.

Description: While the behavior and objectives of non-industrial private forest (NIPF) owners have been studied extensively, studies that systematically test the underlying measurement model are lacking in forest economic literature. Our paper reports the results obtained from a recent large-scale survey conducted in Finland in 2011 (n = 557). Results indicate a novel way to systematically analyze the objectives of forest ownership by testing the validity of the developed measurement scale using the structural equations modeling technique. From an exploratory factor analysis of 22 items measuring forest owner objectives, a four-dimensional structure is identified in the background objectives of NIPF owners. These dimensions are labeled as recreation and leisure time, sense of economic security, nature conservation and aesthetics, and timber sales income objective. Having undergone a confirmatory testing process, results from the four-dimensional model support the validity of the developed 16-item measurement model. Based on these findings, we argue that the logical NIPF owner objective structure in Finland consists of experiential forest value, as perceived in current and future time contexts, as well as of current and future economic objectives. As the theoretical structure divides forest owner objectives into the evaluation of the present objectives, supplemented with a psychological evaluation of the future objectives, a novel classification of NIPF owner objectives is suggested.

Description: By using more sophisticated sampling designs in forest field inventories, it is possible to select more representative field samples. When full cover auxiliary information is available at the planning stage of a forest inventory, an efficient strategy for sampling is formed by making sure that the sample is well spread in the space spanned by the auxiliary variables. We show that by using such a sampling design, we can improve not only design-based estimation, but also estimation based on nearest neighbour techniques. A new technique to select well-spread probability samples, in multidimensional spaces, from larger populations is introduced. As an application, we illustrate how this strategy can be applied to a forest field inventory. We use an artificial dataset corresponding to a full cover forest remote sensing inventory of a 30 000 ha area of Kuortane, western Finland. The target variable (growing stock volume) has been generated for the entire area by a copula technique. The artificial population has been validated by utilizing the Finnish National Forest Inventory.

Description: Seventy-five eastern Cottonwood (Populusdeltoides Bartr.) clones, selected from tests representing the top one-third of clones tested throughout the Lower Mississippi River Valley, were analyzed for alpha cellulose content, specific gravity, and volume after three growing seasons. All traits were found to differ significantly among clones. Mean clonal alpha cellulose content ranged from 48.2 to 55.8% of oven-dry, extractive-free wood with an average of 51.1%. Specific gravity averaged 0.33, with clones ranging from 0.27 to 0.39. Specific gravity was highly inherited, while alpha cellulose content was found to be moderately heritable. Negative genetic correlations between volume and both wood properties indicate that using a selection index to simultaneously improve all three traits is currently not possible. Two other selection methods were discussed. These selections showed substantial gain differentials in volume, but either a small loss or improvement in specific gravity and alpha cellulose content. This indicates that at age 3 years only volume should be considered in a selection program.

Description: Declining biodiversity is a critical component of global change owing to its influence on ecosystem functioning. Decomposition rate frequently increases with fungal species number, but the responses of extracellular enzymes to fungal species number have not been tested. To test the effect of biodiversity on decomposition and enzyme activities, quaking aspen ( Populus tremuloides Michx.) litter was inoculated with mixtures of one, two, four, or eight fungi from a pool of 16 fungi that had been isolated from a boreal forest in Alaska. Total CO2 release and the activities of β-glucosidase, which targets cellulose, and polyphenol oxidase, which targets lignin and other recalcitrant phenolic compounds, were observed across the range of species numbers in the mixtures. Total CO2 release and β-glucosidase activity increased with number of species but were only weakly correlated with each other; polyphenol oxidase activity had no correlation with number of species or CO2 release. The results indicate that, over 4 months, decomposition of labile carbon is positively correlated with number of species.

Description: In recent years, the annual imports of wooden bedroom furniture by the United States have been over five billion dollars, with more than two billion dollars of that coming from China. This trend led to an antidumping action against China in October 2003. Since January 2005, antidumping duties of 0.83% to 198.08% have been imposed on individual Chinese firms. To assess the impact of this antidumping action, intervention analysis was employed to examine the import values of four furniture commodities and the prices of two of them over 1997–2008. China and six other major competing countries were included in the analysis. With regard to import values from China, significant trade investigation effects were identified: the petition announcement generated a positive impact in March 2004; the preliminary less-than-fair-value (LTFV) determination had a negative impact from July to December 2004. However, the final implementation did not show any expected trade duty effect. The aggregate impact of the antidumping action on import values from China over 2003–2008 was approximately equivalent to a 1-month import reduction. The impact on the unit prices for China was insignificant. For the six competing countries, intervention analyses revealed that the antidumping action generated a positive trade diversion effect, with the magnitude smaller than the trade depression effect on China.

Description: Longleaf pine (Pinus palustris Mill.) restoration in the southeastern United States offers opportunities for carbon (C) sequestration. Ecosystem C stocks are not well understood in longleaf pine forests, which are typically of low density and maintained by prescribed fire. The objectives of this research were to develop allometric equations for above- and below-ground biomass and quantify ecosystem C stocks in five longleaf pine forests ranging in age from 5 to 87 years and in basal area from 0.4 to 22.6 m2·ha−1. Live aboveground C (woody plant + ground cover) and live root C (longleaf pine below stump + plot level coarse roots + plot level fine roots) ranged from 1.4 and 2.9 Mg C·ha−1, respectively, in the 5-year-old stand to 78.4 and 19.2 Mg C·ha−1, respectively, in the 87-year-old stand. Total ecosystem C (live plant + dead organic matter + mineral soil) values were 71.6, 110.1, 124.6, 141.4, and 185.4 Mg C·ha−1 in the 5-, 12-, 21-, 64-, and 87-year-old stands, respectively, and dominated by tree C and soil C. In the 5-year-old stand, ground cover C and residual taproot C were significant C stocks. This unique, in-depth assessment of above- and below-ground C across a series of longleaf pine stands will improve estimates of C in longleaf pine ecosystems and contribute to development of general biomass models that account for variation in climate, site, and management history in an important but understudied ecosystem.

Description: Microplot and conventional plot trials were used to determine the nutritional status and required nutrient additions to bring young regenerations of western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western red cedar (Thujaplicata Donn ex D. Don) out of "check." The trees were growing on deep mor-humus Podzols invaded by dense salal (Gaultheriashallon). Salal removal by grubbing and application of Garlon was also tested. Foliar vector analysis, used for hemlock, identified a response to N and P that was confirmed by subsequent 3-year height growth response. Salal removal resulted in increased N uptake only in cedar. Cedar also responded to N and P additions, but vector analysis was not feasible owing to indeterminate growth. Foliar analysis values are compared with published data. It is suggested that salal competition or allelopathy may be the primary cause of inadequate N and P nutrition.

Description: The comparative analysis of a large set of long-term fertilization and thinning studies in the major forest types of interior Alaska is summarized. Results indicate that nutrient limitations may only occur during the early spring growth period, after which moisture availability is the primary control of tree growth on warm sites. The temperature dynamics of both air and soil set seasonal bounds on the nutrient and moisture dynamics for all forest types. Air and soil temperature limitations are the primary control of intraseasonal growth in the colder topographic locations in interior Alaska. These locations are usually dominated by black spruce (Picea mariana (Mill.) Britton, Sterns, Poggenb.) vegetation types. The seasonal progression of factors controlling growth is strongly tied to the state factor structure of the landscape.

Description: Progenies belonging to 49 open-pollinated families of paper birch (Betulapapyrifera Marsh.) representing 13 provenances were evaluated sequentially for rate of root elongation in solutions containing only calcium nitrate and solutions containing calcium nitrate + 120 ppm aluminum. Provenances, but not families within provenances, differed significantly in both rate of root elongation and apparent response to Al. Provenance mean tolerance indices (root growth in Al solutions/root growth in non-Al solutions) varied from 0.10 to 0.37. Exposure to 120 ppm Al caused decreased concentrations of Ca and Cu in the foliage, decreased or nearly unchanged concentrations of P and Mg, and increased concentrations of Al, B, and Mn. Compared with the more tolerant provenances, intolerant provenances exhibited generally greater increases in foliar Al concentration after exposure to Al. Differential tolerance to Al will be useful in improving the performance of planted trees on Al-toxic sites.

Description: Short-rotation intensive culture (SRIC) of willow (Salix spp.) or hybrid poplar (Populus spp.) is currently at a precommercial stage with a potential to be applied economically across important areas to produce lignocellulosic biomass and environmental services in Canada. A two-round Delphi survey was conducted among 50 experts to assess the future deployment of SRIC in this country. The total area in 10 years (2011 base year) was forecasted as 1330, 4100, and 11 400 ha under pessimistic, realistic, and optimistic scenarios, respectively. The deployment of SRIC in the next decade depends mainly on the development of the demand for SRIC biomass and services, that of production technologies, and the establishment of policies and programs promoting its application. In the short term, research and development (R&D) and policy initiatives should be funded or implemented by various stakeholders to facilitate the deployment of the system. On average, respondents deemed that the potential for long-term (20 years) deployment of SRIC in Canada was good. Some of the conclusions and the methodological approach of this study could apply to short-rotation woody crop systems and to agroforestry systems in Canada and elsewhere.

Description: Restoration by imitating natural disturbances is widely practised in boreal forests to increase the availability of habitats for specialized species. We studied the abundance and species richness of saproxylic beetles on different types of created dead wood during 2 years after restoration. The study was conducted on areas of a large-scale experiment in which Norway spruce ( Picea abies (L.) Karst.) forests were restored by controlled burning and partial harvesting with down wood retention in southern Finland. More beetle species were attracted to spruces than to birches and more species were attracted to burnt trees than to unburnt trees killed by girdling. Birch-living species consistently benefited from fire, but on spruce, the abundance of cambium consumers and their associates was negatively affected by fire. Trees at harvested sites attracted more beetles in the first year, but the volume of down wood retention had only minor effects. Beetle assemblages were strongly altered by burning and harvesting. We conclude that burning and harvesting are efficient tools to promote species richness within a short time period, but there is a risk that the dead wood resource may be rapidly exhausted. Moreover, many saproxylic species of spruce forests may not be adapted to open habitats formed by stand-replacing disturbances.

Description: This study evaluated the potential for the selective genetic improvement of the structural quality traits important in sawn Norway spruce (Picea abies (L.) Karst.) timber based on early and nondestructively assessed field traits. From a 34-year-old Norway spruce trial situated in southern Sweden, 401 butt logs were sampled and sawn to produce two 50 × 100 mm boards that were dried to an 18% moisture content. Structural quality traits were assessed, and genetic parameters were estimated, including additive genetic variance, heritability, and their genetic correlations with field traits. Board twisting, density, and modulus of elasticity (MOE, stiffness) were found to have appreciable heritabilities (0.23–0.44). Board twist was found to have a strong genetic correlation with grain angle measured under bark in the field (0.93), and both board MOE and density exhibited strong genetic correlations with field-assessed pilodyn penetration (–0.75 and –0.91, respectively). Although these observations were made on a thinning material comprising mainly juvenile wood, they nonetheless suggest grain angle and pilodyn penetration to be promising candidates as selection criteria for Norway spruce breeding. Heritabilities of other sawn timber traits were lower and the genetic correlations between these traits and field traits were also lower, variable, and had large estimation errors.

Description: The current mountain pine beetle infestation in British Columbia’s lodgepole pine forests has raised concerns about potential impacts on water resources. Changes in forest structure resulting from defoliation, windthrow, and salvage harvesting may increase snow accumulation and ablation (i.e., spring runoff and flooding risk) below the forest canopy because of reduced snow interception and higher levels of radiation reaching the surface. Quantifying these effects requires a better understanding of the link between forest structure and snow processes. Light detection and ranging (lidar) is an innovative technology capable of estimating forest structure metrics in a detailed, three-dimensional approach not easily obtained from manual measurements. While a number of previous studies have shown that increased snow accumulation and ablation occur as forest cover decreases, the potential improvement of these relationships based on lidar metrics has not been quantified. We investigated the correlation between lidar-derived and ground-based traditional canopy metrics with snow accumulation and ablation indicators, demonstrating that a lidar-derived forest cover parameter was the strongest predictor of peak snow accumulation (r2 = 0.70, p 〈 0.001) and maximum snow ablation rate (r2 = 0.59, p 〈 0.01). Improving our ability to quantify changes in forest structure in extensive areas will assist in developing more robust models of watershed processes.

Description: Black spruce seedlings (Piceamariana Mill.) were exposed to either elevated (1000 ppm) or ambient (340 ppm) atmospheric CO2 levels at different stages of seedling development over a winter greenhouse production cycle. Seedlings germinated in early February and were placed in CO2 chambers for either 3 or 6 weeks during March, April, May, or August. Total seedling biomass increased under high CO2 conditions for the March, April, and May stages of development, but showed no significant response in August. The greater part of the CO2 response occurred during the second 3 weeks of exposure in March and April but during the first 3 weeks of exposure in May. In September, those seedlings exposed to CO2 in April and May had 30 and 14%, respectively, greater biomass than control seedlings, but seedlings from the other stages of development no longer had significant differences remaining from the CO2 treatment. This suggests that it could be very efficient to give a short well-timed CO2 pulse at the beginning of the production cycle in hopes of producing a size difference that is maintained throughout the remainder of the greenhouse production cycle under ambient levels of CO2. Short-term exposure to elevated CO2 also increased the ratio of shoot dry weight to total height for the March, April, and May stages of development. The ratio of total nonstructural carbohydrates to free amino acids was negatively correlated (r2 = 0.98) with the allocation of new growth between shoots and roots as measured by the allocation coefficient, k (milligrams shoot growth per milligrams root growth). As seedlings developed along their seasonal growth cycle, ratios of total nonstructural carbohydrates to free amino acids increased and the values for k decreased. The effect of CO2 enrichment on these two factors is discussed. Monitoring total nonstructural carbohydrate and free amino acid concentrations in foliage could have potential as a method to predict the percentage of carbon allocated to root systems of entire forest stands as well as of individual tree seedlings.

Description: Seasonal patterns of change in lipids, sugars, starch, labile (ethanol soluble) constituents, holocellulose, and lignin were studied in six forest-grown white oak (Quercusalba L.) trees. Contents of metabolically active constituents in leaves, twigs, branches, boles (upper and lower), and roots (support and small lateral) were used to construct whole-tree budgets of energy allocation. [14C]Sucrose was also concurrently supplied to the study trees to follow the fate and efficiency of utilization of food reserves. Results showed that white oak rapidly mobilized and replaced food reserves during the critical period of canopy generation in the spring. Starch was more important as a reserve food than lipids or sugar. Large fluctuations in starch in roots in spring and fall suggested a bimodal belowground growth pattern. Labile constituents showed the most pronounced seasonal changes and dominated the calculated whole-tree energy flux patterns. Rapid decline in labile compounds in early spring and a parallel increase in holocellulose suggested a possible pattern of mobilization and resupply of stored reserves associated with in cell wells. This possibility was supported by a concurrent shift of labile 14C to nonlabile 14C in tissues. Canopy generation was calculated to have cost ≤17.7 kg of glucose (1.6 g glucose/g of canopy) of which 13 kg appeared to have come from within the canopy.

Description: Predation by curculionid larvae, tannic acid content, and germination were measured in acorns from individual trees of Quercusalba and Quercusrubra from two sites at the Meeman Biological Field Station, Shelby County, Tennessee, U.S.A. A crossed and nested analysis of variance design was used, and no significant differences were found among trees or between sites in percentages of acorns that were attacked nor in tannic acid contents. Quercusrubra acorns had significantly less predation and higher amounts of tannic acid than Q. alba acorns. Acorns of both species that were not attacked had significantly higher germination success (82% for Q. alba and 98% for Q. rubra) than acorns that were attacked. There was no significant relationship between amount of predation and tannic acid content for acorns from trees of either species. Tannic acid content appears to have an impact by reducing seed predation and may interact with seed crop size to increase annual germination success of trees.

Description: Forest bryophytes are sensitive to the disturbances and environmental changes associated with forest management. We asked whether the substrates on which bryophytes grow mediate responses to exposure following canopy removal. We measured bryophyte cover and richness in 0.1 m2 quadrats on the forest floor, decayed logs, and tree bases along a gradient of dispersed overstory retention (100%, 40%, and 15% of initial basal area) 7 to 8 years after harvest of mature Pseudotsuga forests. Cover, local richness, and, to a lesser degree, species evenness declined steeply across the retention gradient on decayed logs and tree bases but not on the forest floor. Liverworts were more sensitive than mosses, particularly on decayed logs and on the southwestern aspects of trees (〉97% declines in cover under 15% retention). Richness and evenness at the treatment scale also declined sharply on decayed logs and on the southwestern aspects of trees but changed little or increased under 40% retention on the forest floor. Our results indicate that even moderate levels of dispersed retention cannot sustain the abundance and overall diversity of wood-associated bryophytes in these forests. During regeneration harvests, conservation of these species may require retention of intact forest aggregates in which substrate quality and microclimatic stability can be maintained.

Description: Forest soil respiration insitu was used as a comparative measure of the metabolic activity of substrate in eastern Ontario jack pine (Pinusbanksiana Lamb.) ecosystems that had been exposed to various burning treatments, including wildfire. The five burning treatments consisted of a 1920 wildfire, experimental understorey burning (nonlethal to the overstorey) of this age-class in 1962 and 1963, a 1964 wildfire, and experimental burning of this age-class in 1977. Seasonal respiration trends were similar on all treatments. Carbon dioxide evolution increased in the spring (4000 mg•m−2•d−1) in response to ambient warming (5000 mg•m−2•d−1 in August) and decreased in late fall as seasonal temperatures declined (4000 mg•m−2•d−1 in November). Precipitation and autumnal litter fall apparently acted as secondary modifiers of this general trend by affecting substrate moisture content and nutrient quality, respectively. Highest metabolic activities were measured on the 1963 understorey burning treatment followed in decreasing order by the 1920 wildfire, the 1964 wildfire, the 1962 experimental understorey burn, and the 1977 burn of the 1964 age-class. Multiple comparisons of overall seasonal respiration means revealed lower rates (P 

Description: Productivities of 5 black cottonwood clones from each of 10 source populations (50 clones in total) were compared at one plantation site in western Washington. The source populations are located west of the Cascade Mountains, between central Oregon and southern British Columbia. Most source stands represented populations in major river valleys. Individual clones were selected for superior form and growth rate. Spacing was 1.2 × 1.2 m (6944 stems•ha−1) and trees were harvested after 4 years. Considerable variation in mortality, height, and productivity were found among clones within individual stands and among stands. Southwestern clones were generally more productive. Heights of individual clones averaged between 8.5 and 11.8 m at 4 years. Average dry weight production of black cottonwood clones was 12.5 Mg•ha−1•year−1 and varied between 5.2 and 23.1 Mg•ha−1•year−1 for individual clones. Three new Populustrichocarpa × P. deltoides hybrids planted in the trial were generally more productive than black cottonwood or the 'Robusta' hybrid, with average heights varying from 11.5 to 12.2 m and dry weight production ranging from 15.6 to 27.8 Mg•ha−1•year−1 for an average of 23.6 Mg•ha−1•year−1. The high yields in this experiment are attributed to favorable climate (1940 degree-days•year−1)3, cultural treatments (irrigation, N fertilization) and genetic constitution of certain clones. The data promise substantial gains in short-rotation productivity from combining clonal selection with interspecific hybridization.

Description: Methods for constructing a sampling design for large area forest inventories are presented. The methods, data sets used, and the procedures are demonstrated in a real setting: constructing a sampling design for the first national forest inventory for Tanzania. The approach of the paper constructs a spatial model of forests, landscape, and land use. Sampling errors of the key parameters as well as the field measurement costs of the inventory were estimated using sampling simulation on data. Forests and land use often vary within a country or an area of interest, implying that stratified sampling is an efficient inventory design. Double sampling for stratification was taken for the statistical framework. The work was motivated by the approach used by The Food and Agriculture Organization of the United Nations (FAO) in supporting nations to establish forest inventories. The approach taken deviates significantly from the traditional FAO approaches, making it possible to calculate forest resource estimates at the subnational level without increasing the costs.

Description: The acetylene reduction assay was used to estimate amounts of nitrogen fixed by Lupinusarcticus Wats. (1.97 kg•ha−1•year−1) and Shepherdiacanadensis (L.) Nutt. (0.78 kg•ha−1•year−1) in a regenerating lodgepole pine (Pinuscontorta Dougl.) stand in southern British Columbia. These amounts appear to be significant, as lodgepole pine ecosystems are generally highly deficient in nitrogen. Thinning of excessively dense pine stands may provide an added growth benefit by increasing nitrogen fixation activity of understory vegetation.

Description: Automated individual tree detection and delineation from high spatial resolution imagery provides good opportunities for forest inventory at a large scale. However, the accuracy of delineated crown size compared with ground measurements may not be sufficient. Thus, ordinary least squares (OLS) regression is no longer an appropriate approach to estimating and predicting variables from the delineated tree crown because both response variable and regressor are subject to measurement errors. In this study, we describe the functional and structural relationships between field-measured tree variables (i.e., tree diameter and crown width) and delineated tree crown width from remotely sensed imagery. We investigated the performance of OLS and three error-in-variable regression techniques including maximum likelihood estimator (MLE), major axis (MA) regression, and reduced major axis (RMA) regression using field-measured data and simulated data under different conditions. Our results indicated that MLE was desirable for estimating unbiased model coefficients. However, the adjustment assumption of the MLE model should be checked for predicting tree variables from remotely sensed imagery. When the assumption holds, the MLE model performed better for predicting the response variables than did the OLS model. Otherwise, the MLE model produced biased predictions for the response variables.

Description: It is widely accepted that N limits primary production in temperate forests, although colimitation by N and P has also been suggested, and on some soils, Ca and base cations are in short supply. I conducted a meta-analysis to assess the strength of existing experimental evidence for limitation of primary production by N, P, and Ca in hardwood forests of the northeastern United States and southeastern Canada using data from 35 fertilization experiments in deciduous forests on glaciated soils across the region. There is strong evidence for N limitation (formal meta-analysis weighted mean response ratio = 1.51, p 〈 0.01; simple mean = 1.42, p 〈 0.001). Forest productivity also tended to increase with additions of P (simple mean = 1.15, p = 0.05) and Ca (simple mean = 1.36, p 〈 0.001). Across all treatments, 85% of response ratios were positive. Multiple-element additions had larger effects than single elements, but factorial experiments showed little evidence of synergistic effects between nutrient additions. Production responses correlated positively with the rate of N fertilization, but this effect was reduced at high rates of ambient N deposition.

Description: The resilience and vulnerability of permafrost to climate change depends on complex interactions among topography, water, soil, vegetation, and snow, which allow permafrost to persist at mean annual air temperatures (MAATs) as high as +2 °C and degrade at MAATs as low as –20 °C. To assess these interactions, we compiled existing data and tested effects of varying conditions on mean annual surface temperatures (MASTs) and 2 m deep temperatures (MADTs) through modeling. Surface water had the largest effect, with water sediment temperatures being ~10 °C above MAAT. A 50% reduction in snow depth reduces MADT by 2 °C. Elevation changes between 200 and 800 m increases MAAT by up to 2.3 °C and snow depths by ~40%. Aspect caused only a ~1 °C difference in MAST. Covarying vegetation structure, organic matter thickness, soil moisture, and snow depth of terrestrial ecosystems, ranging from barren silt to white spruce ( Picea glauca (Moench) Voss) forest to tussock shrub, affect MASTs by ~6 °C and MADTs by ~7 °C. Groundwater at 2–7 °C greatly affects lateral and internal permafrost thawing. Analyses show that vegetation succession provides strong negative feedbacks that make permafrost resilient to even large increases in air temperatures. Surface water, which is affected by topography and ground ice, provides even stronger negative feedbacks that make permafrost vulnerable to thawing even under cold temperatures.

Description: Three indices of nitrogen (N) availability were compared in the field over a 1-year period in an old-growth and a young-growth mixed-conifer forest. The indices utilized were ion exchange resin (IER) bags, buried bags, and a core-IER method employing intact soil cores enclosed in tubes capped at both ends by IER bags. The results from all three methods indicated that in the surface mineral soil, N availability was higher in the young-growth stand than in the old-growth stand. However, seasonal patterns of N availability were generally not well correlated among the methods (correlation coefficients ranged from 0.32 to 0.62). For a given amount of net N mineralized in buried bags, more N accumulated on IER bags placed in the young-growth stand than in those placed in the old-growth stand. This was the result of greater net nitrification in the young-growth stand coupled with the greater mobility of [Formula: see text] relative to [Formula: see text] in soil. Ten-month estimates of net N mineralization measured by the core-IER and buried-bag methods were similar in the young-growth stand (about 42 mg•kg soil−1), but the core-IER estimate was almost twice that of the buried-bag estimate in the old-growth stand (31.7 and 16.8 mg•kg soil−1, respectively). The different sensitivities of the core-IER and buried-bag methods to changes in soil moisture and leaching probably account for much of the difference in their N availability estimates. Results from the core-IER method did reflect the effects of leaching; however, soil water content within the core did not follow changes in soil water content effectively. Because of the greater labor involved in using the core-IER method, its use may be most efficacious in high-precipitation environments, or when in-field soil incubations must be conducted for extended periods of time.

Description: In British Columbia, Canada, a recent epidemic of mountain pine beetle (Dendroctonus ponderosae Hopkins, 1902) caused widespread forest mortality. This epidemic was due in part to the changing climate, and damage from pests and diseases is expected to increase in the future. Therefore, we used a historical retrospective approach as a proxy to evaluate management options on reducing the forest health damage that may occur under a future changing climate. We assessed two landscape-scale strategies, intended to increase tree species diversity, for the response in ecosystem resilience and compared the results with the business-as-usual strategy. The assessment was based on simulation modelling of the Merritt Timber Supply Area for 1980–2060. We applied a strategy to increase the harvest of the most dominant tree species, plant more diverse species, and increase natural regeneration. This strategy resulted in greater ecological resilience (higher diversity and growing stocks), higher harvest rates, and higher, more consistent net revenue over time than the business-as-usual strategy or the strategy that only employed a diversity of planting. A sensitivity analysis indicated a high level of robustness in the results. Our study showed that it may not be necessary to compromise economic viability to reduce forest health risks and consequently improve socio-ecological resilience.

Description: This paper assesses the resilience of Alaska’s boreal forest system to rapid climatic change. Recent warming is associated with reduced growth of dominant tree species, plant disease and insect outbreaks, warming and thawing of permafrost, drying of lakes, increased wildfire extent, increased postfire recruitment of deciduous trees, and reduced safety of hunters traveling on river ice. These changes have modified key structural features, feedbacks, and interactions in the boreal forest, including reduced effects of upland permafrost on regional hydrology, expansion of boreal forest into tundra, and amplification of climate warming because of reduced albedo (shorter winter season) and carbon release from wildfires. Other temperature-sensitive processes for which no trends have been detected include composition of plant and microbial communities, long-term landscape-scale change in carbon stocks, stream discharge, mammalian population dynamics, and river access and subsistence opportunities for rural indigenous communities. Projections of continued warming suggest that Alaska’s boreal forest will undergo significant functional and structural changes within the next few decades that are unprecedented in the last 6000 years. The impact of these social–ecological changes will depend in part on the extent of landscape reorganization between uplands and lowlands and on policies regulating subsistence opportunities for rural communities.

Description: Multiple-use forestry requires comprehensive planning to maximize the utilization and sustainability of many forest resources whose growth and productivity are interconnected. Forest fungi represent an economically important nonwood forest resource that provides food, medicine, and recreation worldwide. A vast majority of edible and marketed forest mushrooms belong to fungi that grow symbiotically with forest trees. To respond to the need for planning tools for multiple-use forestry, we developed empirical models for predicting the production of wild mushrooms in pine forests in the South-Central Pyrenees using forest stand and site characteristics as predictors. Mushroom production and species richness data from 45 plots were used. A mixed modelling technique was used to account for between-plot and between-year variation in the mushroom production data. The most significant stand structure variable for predicting mushroom yield was stand basal area. The stand basal area associated with maximum mushroom productivity (15–20 m2·ha–1) coincides with the peak of annual basal area increment in these pine forests. Other important predictors were slope, elevation, aspect, and autumn rainfall. The models are aimed at supporting forest management decisions and forecasting mushroom yields in forest planning.

Description: Exotic Pinus taeda L. plantations may be more productive than native ones. Several hypotheses may explain this difference; however, process models with a light-interception-driving variable cannot test these hypotheses without foliage display first being quantified in native and exotic trees. We quantified leaf area duration in North Carolina, USA (natural), and Gobernador Virasoro, Argentina (exotic), with no additional nutrients and optimum fertilizer treatments. More (60%–100%) foliage was displayed but for a shorter (∼86 fewer days) time per fascicle in the exotics than in the naturals. Study inference was limited, with only one native and one exotic site. However, while the sites were markedly different in soils, climate, resource availability, and genetics, and we observed significant differences in fascicle display and longevity, most fascicles at both sites survived two growing seasons: the one in which they were produced and the subsequent one. This robust finding indicates it would be reasonable to use two growing seasons for fascicle longevity in process modeling to test hypotheses explaining growth differences in native and exotic loblolly. Fertilization had no effect on any exotic tree parameter, but it increased natural tree fascicle number (24%) and length (30%).

Description: This study introduces five facets that can improve inference in small area estimation (SAE) problems: (1) model groups, (2) test of area effects, (3) conditional EBLUPs, (4) model selection, and (5) model averaging. Two contrasting case studies with data from the Swiss and Norwegian national forest inventories demonstrate the five facets. The target variable of interest was mean stem volume per hectare on forested land in 108 Swiss forest districts (FD) and in 14 Norwegian municipalities (KOM) in the County of Vestfold. Auxiliary variables from airborne laser scanning (Switzerland) and photogrammetric point clouds (Vestfold) with full coverage and a resolution of 25 m × 25 m (Switzerland) and 16 m × 16 m (Vestfold) were available. Only the data metric mean canopy height was statistically significant. Ten linear fixed-effects models and three mixed linear models were assessed. Area effects were statistically significant in the Swiss case but not in Vestfold case. A model selection based on AIC favored separate linear regression models for each FD and a single common regression model in Vestfold. Model averaging increased, on average, an estimated variance by 15%. Reported estimates of uncertainty were consistently larger than corresponding unconditional EBLUPs.

Description: Northern Great Lakes forests represent an ecotone in the boreal–temperate transition zone and are expected to change dramatically with climate change. Managers are increasingly seeking adaptation strategies to manage these forests. We explored the efficacy of two alternative management scenarios compared with business-as-usual (BAU) management: expanding forest reserves meant to preserve forest identity and increase resistance, and modified silviculture meant to preserve forest function and increase adaptive capacity. Our study landscapes encompassed northeastern Minnesota and northern Lower Michigan, which are predicted to experience significant changes in a future climate and represent a gradient of latitude, forest type, and management. We used the LANDIS-II forest simulation model to simulate forest change under current climate, low emissions climate, and high emissions climate futures. Our results suggest that under a low emissions climate scenario, expanded reserves and modified silviculture strategies can be effective at increasing resistance by preserving forest composition, including legacy species (e.g., balsam fir (Abies balsamea (L.) Mill.)), and increasing adaptive capacity by maintaining or increasing aboveground biomass compared with BAU management. Under a high emissions climate scenario, the expanded reserve strategy was not effective at preserving legacy species; however, the modified silviculture strategy was effective at increasing aboveground biomass compared with BAU management. These results highlight alternative management options and limitations in the face of climate change.

Description: The induction of the enzyme nitrate reductase in needles may be a prerequisite for the assimilation of foliar-absorbed nitrogen oxide pollutants by red spruce (Picearubens Sarg.) trees. To test for induction of nitrate reductase, 1-year-old red spruce seedlings were exposed to NO2, HNO3 vapor, or acid mist containing nitrate, and the activity of nitrate reductase in needles was measured. One day after exposure to NO2 (75 nL•L−1) began, nitrate reductase activity was three times greater than that of unexposed control plants. One day after exposure ended, the nitrate reductase activity returned to the control level. Older red spruce seedlings that had been excavated from a spruce–fir stand exhibited a similar pattern of response, but the level of nitrate reductase activity was much lower than that of the 1-year-old seedlings. Nitric acid vapor (75 nL•L−1) also induced nitrate reductase in red spruce needles, and the pattern of response was similar to that with NO2, except that the nitrate reductase activity did not return to control levels until 2 days after exposure ended. Exposure of seedlings to acid mist containing nitrate (pH 3.5 and 5.0) did not result in a change in nitrate reductase activity. These results indicate that red spruce is capable of assimilating NO2 and HNO3 vapor and that hypotheses of forest decline based on foliar assimilation of pollutant nitrogen oxides are tenable.

Description: From 1983 to 1987 a survey was made to determine the hosts and geographic distribution of the pinewood nematode Bursaphelenchusxylophilus (Steiner and Buhrer), the cause of pine wilt disease, in China. Samples (669) were obtained from dead and dying conifers in 18 provinces and in the federally administrated cities of Beijing and Shanghai. The nematode was only found in samples from Pinusthunbergii and P. massoniana in and around the cities of Nanjing and Zhenjiang (both in Jiansu Province). To determine the susceptibility of Chinese and exotic pines to the pinewood nematode, 15 pine species were inoculated with a Chinese isolate. The 2- to 5-year-old trees, growing at Qingdao (Shandong Province) and Hongjiang (Hunan Province), were inoculated in mid-August to mid-October 1985. The results showed that one pine (P. bungeana) was extremely susceptible to the pinewood nematode, seven were highly susceptible (P. yunnanensis, P. densiflora, P. armandii, P. kwangtungensis, P. elliottii, P. koraiensis, and P. thunbergii), four were moderately susceptible (P. taiwanensis, P. griffithii, P. fenzeliana, and P. sylvestris var. mongolica), and three were resistant (P. taeda, P. massoniana, and P. banksiana). The results are discussed in relation to the needs for quarantine and management of pine wilt disease in the People's Republic of China.

Description: We studied the effect of water availability on basal area growth and wood properties of 11-year-old loblolly pine ( Pinus taeda L.) trees from contrasting Florida (FL) (a mix of half-sib families) and South Carolina coastal plain (SC) (a single, half-sib family) genetic material. Increasing soil water availability via irrigation increased average whole-core specific gravity (SG) and latewood percentage (LW%) by 0.036 and 6.93%, respectively. Irrigation did not affect latewood SG or wood stiffness, but irrigated FL and SC trees had more latewood due to a 29 day longer growing season. Irrigation did not affect the length of corewood production, but irrigated trees had earlier transition ages, producing outerwood ~3 years before rainfed trees. The increase in whole-core SG and LW% was moderate because irrigation promoted earlywood growth in corewood formed before canopy closure, but after year 7, rain-fed and irrigated trees had similar earlywood growth but irrigated trees had more latewood growth, increasing ring SG and LW%. The SC half-sib family had higher SG and greater LW% than trees from FL independent of irrigation due to greater yearly latewood growth. Thus, absence of soil water stress extended seasonal diameter cessation date but did not change latewood SG or wood stiffness.

Description: Wildfire risk assessment research has made considerable progress towards estimating the probability of wildfires but comparatively little progress towards estimating the expected consequences of potential fires. One challenge with estimating wildfire consequences has been to identify a common metric that can be applied to consequences measured in different units. In this paper, we use the preferences of representatives of local fire management agencies as the common consequences metric and apply it to a case study in the southern Gulf Islands, British Columbia, Canada. The method uses an expert survey and a maximum-difference conjoint analysis to establish the relative importance of specific fire consequences. A fire with a major potential for loss of life was considered to be about three times worse than major damage to houses and 4.5 times worse than loss of a rare species. Risk ratings were very sensitive to changes in fire consequences ratings. As the complexity of values at risk and number of stakeholders increase, the most efficient allocation of wildfire prevention, protection, and suppression resources becomes increasingly challenging to determine. Thus, as the complexity of stakeholder representation and values at risk increases, we need to pay increasing attention to quantitative methods for measuring wildfire consequences.

Description: In their recent review of arrested succession, Royo and Carson ( A.A. Royo and W.P. Carson. 2006. Can. J. For. Res. 36: 1345–1362 ) demonstrate that “recalcitrant understory layers” are widespread and pervasive modifiers of ecosystems and disruptors of forest regeneration. They rightly point out that many plant species associated with arrested succession are characterized by rapid vegetative spread. Extending their review, we point out that most of such species are clonal or thicket-forming and suggest that an additional reason why these plants so effectively suppress succession for extended periods is their long life-spans.

Description: Although considerable research has focused on the influences of logging debris treatments on soil and forest regeneration responses, few studies have identified whether debris effects are mediated by associated changes in competing vegetation abundance. At sites near Matlock, Washington, and Molalla, Oregon, studies were initiated after timber harvest to quantify the effects of three logging debris treatments (dispersed, piled, or removed) on the development of competing vegetation and planted Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco var. menziesii ). Each debris treatment was replicated with initial and annual vegetation control treatments, resulting in high and low vegetation abundances, respectively. This experimental design enabled debris effects on regeneration to be separated into effects mediated by vegetation abundance and those independent of vegetation abundance. Two to three years after treatment, covers of Scotch broom ( Cytisus scoparius (L.) Link) at Matlock and trailing blackberry ( Rubus ursinus Cham. & Schltdl.) at Molalla were over 20% greater where debris was piled than where it was dispersed. Debris effects on vegetation abundance were associated with 30% reductions in the survival of Douglas-fir at Matlock (r2 = 0.62) and the stem diameter at Molalla (r2 = 0.39). Douglas-fir survival and growth did not differ among debris treatments when effects were evaluated independent of vegetation abundance (i.e., with annual vegetation control), suggesting negligible short-term effects of debris manipulation on soil productivity.

Description: Natural regeneration-based silviculture has been increasingly regarded as a reliable option in sustainable forest management. However, successful natural regeneration is not always easy to achieve. Recently, new concerns have arisen because of changing future climate. To date, regeneration models have proved helpful in decision-making concerning natural regeneration. The implementation of such models into optimization routines is a promising approach in providing forest managers with accurate tools for forest planning. In the present study, we present a stochastic multistage regeneration model for Pinus pinea L. managed woodlands in Central Spain, where regeneration has been historically unsuccessful. The model is able to quantify recruitment under different silviculture alternatives and varying climatic scenarios, with further application to optimize management scheduling. The regeneration process in the species showed high between-year variation, with all subprocesses (seed production, dispersal, germination, predation, and seedling survival) having the potential to become bottlenecks. However, model simulations demonstrate that current intensive management is responsible for regeneration failure in the long term. Specifically, stand densities at rotation age are too low to guarantee adequate dispersal, the optimal density of seed-producing trees being around 150 stems·ha−1. In addition, rotation length needs to be extended up to 120 years to benefit from the higher seed production of older trees. Stochastic optimization confirms these results. Regeneration does not appear to worsen under climate change conditions; the species exhibiting resilience worthy of broader consideration in Mediterranean silviculture.

Description: Disturbance–succession models describe the relationship between the disturbance regime and the dominant tree species of a forest type. Such models are useful tools in ecosystem management and restoration, provided they are accurate. We tested a disturbance–succession model for the oak–pine ( Quercus spp. – Pinus spp.) forests of the Appalachian Mountains region using dendrochronological techniques. In this model, fire promotes pines, while fire suppression, bark beetle outbreaks, and ice storms encourage oaks. We analyzed nine Appalachian oak–pine stands for species establishment dates and the occurrence of fires and canopy disturbances. We found no evidence that fire preferentially promoted the establishment of pine more than oak, nor did we find any evidence that canopy disturbances or periods of no disturbance facilitated the establishment of oak more than pine. Rather, we found that both species groups originated primarily after combined canopy and fire disturbances, and reduction of fire frequency and scope coincided with the cessation of successful oak and pine regeneration. Currently, heath shrubs are slowly dominating these stands, so we present a revised disturbance–succession model for land managers struggling to manage or restore oak–pine forests containing a dense ericaceous understory.

Description: Based on individual tree damage data dating back to the gale “Lothar” (winter 1999) in Baden-Württemberg, Germany, a statistical model was developed to estimate the risk of storm damage for individual trees. The data were compiled from the National German Forest Inventory. The model attempts to separate the effects of tree-specific variables, topography, site conditions and flow field related effects on damage probability. The crucial problem of missing information on the actual flow field parameters was solved by applying a generalized additive model that enables the simultaneous fit of a spatial trend function. The geographical location of risk hotspots as predicted by the model correspond well to the actual distribution pattern of storm damage as assessed by the forest service. Tree height proved to be one of the most important factors affecting the level of damage, while height to diameter at breast height ratio influences damage probability to a much lesser extent. The Norway spruce ( Picea abies (L.) Karst.) group has the highest potential to be damaged followed by the silver fir ( Abies alba Miller) – Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco) group and the Scots pine ( Pinus sylvestris L.) – larches ( Larix spp.) group. Predicted probabilities for deciduous trees are generally lower than those of conifers. West- to south-exposed locations bear a considerably higher damage risk and waterlogged soils show an increased predicted probability compared with slightly or not waterlogged soils.

Description: Fire suppression has facilitated the spread of red maple ( Acer rubrum L.), a fire-sensitive, yet highly adaptable species, in historically oak-dominated forests of the eastern United States. Here, we address whether a shift from upland oaks to red maple could influence forest hydrology and nutrient availability because of species-specific effects on precipitation distribution and inorganic nitrogen (N) cycling. In eastern Kentucky, we measured seasonal variations in red maple, chestnut oak ( Quercus montana Willd.), and scarlet oak ( Quercus coccinea Münchh.) throughfall and stemflow quantity and quality following discrete precipitation events, and we assessed net N mineralization rates in underlying soils over a 2-year period (2006–2008). Throughfall was 3%–9% lower underneath red maple than both oaks, but red maple generated 2–3× more stemflow. Consequently, NH4+ throughfall deposition was less under red maple than chestnut oak, whereas stemflow-derived nutrient inputs were substantially larger for red maple than both oaks. Soils underlying red maple had 5–13× greater winter net nitrification rates than soils under both oaks and 20%–30% greater rates of seasonal net ammonification than soils under chestnut oak. These findings suggest a spatial redistribution of water and nutrients via precipitation as red maple dominance increases and point to stemflow as an important mechanism that may foster red maple competitive success, further bolstering the mesophication process in the United States.

Description: The effects of precommercial thinning on the understory vegetative cover of 16- to 18-year-old spruce–hemlock ( Picea sitchensis (Bong.) Carrière – Tsuga heterophylla (Raf.) Sarg.) stands were studied in seven replicate areas over seven growing seasons postthinning. Vegetative cover was analyzed at the class level, but species-specific effects were examined in relation to their value as food for Sitka black-tailed deer ( Odocoileus hemionus sitkensis Cowan). When compared with unthinned controls, thinned stands (3.6–6.3 m spacing) had significantly greater understory cover. However, all thinned spacings led to similar understory cover. Conifer cover recovered to about two-thirds of its prethinning level within seven growing seasons posttreatment. Understory nonconiferous cover increased during the first 2–4 years postthinning but began to decline with increasing conifer cover during the next 3 years, nearly reaching pretreatment levels by year 7. In unthinned understories, vegetative cover had declined and was significantly lower than that beneath thinned stands. Summer food resource values for deer were increased by thinning. Winter food resource values were increased by thinning for snow-free conditions but were unaffected for conditions when herb-layer forbs were buried by snow.

Description: The uniform angle index (UAI) is used to characterize the spatial distribution of a forest community or of individual tree species within that community. This study presents an empirical assessment of the performance of a new statistical test used in conjunction with the UAI. In this study, the UAI is applied to observed and simulated communities. The effect of plot size and density are examined and the results are compared with the criteria of the aggregation index R of Clark and Evan (CE-R) and Ripley’s L function (RL). The results suggest that the UAI performs well in determining a particular class of spatial pattern and that, in contrast to the CE-R index, it is unaffected by population density because the standard deviation can be estimated when the density is known using a new empirical relationship presented in this paper. The results of this study represent an improvement in the development of the UAI, verifying its performance and confirming its overall utility. Our comparative analysis positions the UAI among its historical competitors, the CE-R and RL criteria. The UAI index, which does not require expensive mapping of tree positions, can produce results that are equivalent to those obtained by RL and more robust and reliable than those obtained by CE-R.

Description: Ecological land classification (ELC) is central to forestry and environmental management. Few methods exist for the statistical confirmation of the distinctness and continued integrity of the ecological character of ELC regions. Consequently, forest managers lack the tools to measure the impact of ecosystem stressors such as harvest practices and climate change. We develop a framework for tracking the distinctness and modification of vegetative communities of ELC natural regions. We base the framework on principles of numerical taxonomy using the Kolmogorov–Smirnov measure of distributional difference. We demonstrate the utility of the framework using data from a 1986 Forest Development Survey of tree species abundances on 13 508 sample plots from natural regions of the New Brunswick, Canada, ELC. Using the framework, we found vegetative communities of the ELC statistically distinct in the 1986 sampling. We also investigated the impacts of plantations on forest composition using vegetation profiles from a 1999 Forest Development Survey of planted stands. Simulated planting to 20% levels suggests that past and projected planting practices will modify the vegetative character of several natural regions on a scale comparable with interregion variation. The results demonstrate the potential of the framework to track changes to a variety of biotic communities impacted by natural and anthropogenic disturbance.