ALBERT

Description: Twelve species of terricolous microlichens from the Angmagssalik District, Southeast Greenland, are reported: Caloplaca friesii, C. livida, Lecanora boligera, Lecidea oligotropha and Leciophysma arctophila, which are new to the lichen flora of Greenland, Rinodina conradi, which is new to the eastcoast, and Baeomyces roseus, B. rufus, Buellia geophila, B. punctata, Caloplaca tornoensis and Mycoblastus tornoensis, new to Southeast Greenland. In a discussion of the greenlandic distribution, unpublished records from the herbarium of Copenhagen (C) are incorporated. Notes on the habitats are given and the pertinent phytosociological units indicated. Some morphological and anatomical characters are commented upon briefly.

Description: The 16 recognized species of Sorocea are listed with their synonyms and distribution. Two new taxa are described: S. steinbachii C.C. Berg and S. hirtella Mildbread ssp. oligotricha Akkermans & Berg. Three new combinations are made: S. muriculata Miquel ssp. uaupensis (Baillon) C.C. Berg, S. trophoides W. Burger ssp. rhodorachis (Cuatrecasas) C.C. Berg, and S. sprucei (Baillon) Macbride ssp. saxicola (Hassler) C.C. Berg. A key to the species is presented.

Description: Neohattoria Kamim. is a monotypic genus of the Jubulaceae (= Frullaniaceae) with a single species, N. herzogii (Hatt.) Kamim., known from central to northern Japan and the southern part of the Kurile Islands. The present genus was segregated from Frullania by Kamimura (1961; sub. nom. Hattoria Kamim. nom. illeg., non Schust., 1961) on the basis of the branching type, the shape of the first leaf and underleaf on branch, the total lack of secondary pigmentation, the uniform cell structure of the stem in cross section, and the strongly toothed leaf lobes. The generic concept of Neohattoria was greatly expanded by Schuster (1970), who included eight species and classified them into two subgenera, subgen. Neohattoria (with a single species) and subgen. Microfrullania Schust. (with seven species); however, Hattori et al. (1972) transferred all species of subgen. Microfrullania to a newly segregated genus Schusterella Hatt. et al., thus retaining the monotypic status of Neohattoria. As already described and illustrated by Hattori (1955), Kamimura (1961), Mizutani (1961), Ladyzhenskaja (1963), Schuster (1970), and Hattori et al. (1972), Neohattoria herzogii is closely related to species of both Jubula and Frullania. Regarding the taxonomic desposition of Neohattoria, Mizutani (1961) and Mizutani & Hattori (1969) placed it with Jubula in a subfamily Jubuloideae of Lejeuneaceae and Hattori et al. placed it in Jubulaceae (s. lat.). But, Kamimura (1961), Schuster (1970, 1979), and Guercke (1978) placed it more close to Frullania, e.g. in a subfamily Frullanioideae of Jubulaceae (s. lat.); more recently, Asakawa et al. (1979b), admitting three distinct families, Jubulaceae, Frullaniaceae, and Lejeuneaceae, placed Neohattoria and Jubula in the Jubulaceae (s. str.) but Frullania and Schusterella in the Frullaniaceae.

Description: The species Polypodium banaense C.Chr. is transferred to Crypsinus. The recognition of a genus Phymatopteris Pic. Ser. (= Phymatopsis J.Sm.) separate from Crypsinus is discussed.

Description: The six species of Curtia, including a hitherto undescribed species published here, as well as the monotypic genus Hockinia can be distinguished from each other by the seed coat structure. The anticlinal walls and the cuticle provide the most useful information. Curtia tenuifolia appears to be a complex species, but subsp. tenella can be readily separated from this complex by the seed coat structure. Heterostyly has been found in C. tenuifolia subsp. tenuifolia, C. obtusifolia, and Hockinia montana, but differences in seed coat structure can not be correlated with long-, short-, and equal-styled flowers. The differences in seed coat structure, the length of the seeds, and the number of cells per seed plead for maintaining Hockinia (closely related to Curtia) as a distinct genus. One new species of Curtia is described and a new combination is made.

Description: SETTEN, A. K. van & KOEK-NOORMAN, J.: Studies in Annonaceae. VI. A leafanatomical survey of genera of Annonaceae in the Neotropics. — Bot. Jahrb. Syst. 108: 17—50. 1986. — ISSN 0006-8152. Within the scope of the multidisciplinary research project on systematics of Annonaceae, a survey of the leafanatomical features and their distribution in the neotropical Annonaceae is presented. The studied specimens form a rather homogeneous group, as may appear from the family description given here. A detailed study of the leafanatomical features reveals, that differences are mainly found in the indument, the position and contents of the idioblasts, the structure of the primary vein, the type of crystals in the epidermal cells, and the type of sclereids. Based on character states, phenetic similarities and differences are discussed and compared with the classifications of FRIES (1959) and WALKER (1971).

Description: One new species of Dorstenia from Brazil is described: D. carautae C.C. Berg, and four new combinations are made: D. cayapia Vellozo subsp. asaroides (Hooker) C.C. Berg, D. cayapia Vellozo subsp. paraguariensis (Hassler) C.C. Berg, D. cayapia Vellozo subsp. vitifolia (Gardner) C.C. Berg, and D. ramosa (Desvaux) Carauta, Valente & Sucre subsp. dolichocaula (Pilger) C.C. Berg. A list of and a key to the 22 Dorstenia species distinguished in south-eastern tropical America are presented, together with synonymy and distributional data.

Description: Recently a multidisciplinary investigation program on the systematics of Annonaceae was started at Utrecht with special emphasis on the Neotropics. This project will be carried out largely within the framework of the UNESCO-project Flora Neotronica. The first goal is to provide a modern classification of the family as a whole, the second is the publication of a series of monographs for Flora Neotropica. The project has been planned and started in close consultation with leading botanists on the Neotropical flora. The Annonaceae are a family of pantropical distribution with between 2000 and 2500 species in ca. 130 genera as presently understood. In the Neotropics the family is represented by ca. 750 species and 35 genera. It is a family of trees, shrubs, and lianas. Its place is within the order of the Magnoliales and its supposedly closest relative is the family of the Myristicaceae. The Annonaceae, although generally considered primitive in many features, nevertheless offer a number of specialized features as well This makes it a promising object using various kinds of comparative morphological, karyological, and anatomical data. Besides, many species are of medicinal or commercial value, such as various species of Annona and Rollinia, the fruits of which are commonly eaten in most countries of Central America and South America; the Soursop (Annona muricata) is widely cultivated throughout the tropics.

Description: Campylopus galapagensis J.-P. Frahm & Sipman spec. nov. is described. It is closely related to C. pilifer Brid., from which it differs mainly by the presence of substereids in the ventral layer of the costa. It is endemic on the Galapagos Islands, where it occurs frequently from sea level to the highest summits at 1500 m.

Description: Tortula grandiretis Broth., differing from T. muralis Hedw. mainly in the larger, quite smooth lamina cells, is reported from three localities in the SW-Netherlands, where it occurred on open, sandy or clayey, brackish soil on recently enclosed mud flats or salt-marshes. It is also reported from one locality in Turkey. It was formerly known only from Turkestan (U.S.S.R.).

Description: A new species of Asterophorum, A. mennegae, is described from the Sipaliwini Savanne (Suriname). The position of Asterophorum within the family is briefly reviewed, and a key to the 2 species is given.

Description: The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.

Description: A phytosociological survey based on methods of the Zürich-Montpellier School was carried out in the páramo vegetation of the Cordillera Oriental, Colombia. The study area covers about 10,000 and comprises the páramo between the Nevado de Sumapaz (3°55'N, 4250 m), the Sierra Nevada del Cocuy (6°25'N, 5493 m) and the Páramo del Almorzadero (7°N, 4375 m). The páramo vegetation was studied along various altitudinal transects from the upper forest line (3000-3500 m) up to the lower limit of the snowcap (4800 m). A general description of the study area includes data on geology, geomorphology, soils, climate, flora, phytogeography, morphological characters of the vegetation, fauna and landuse. The evolution and Quaternary history of páramo vegetation and climate is reviewed, incorporating the first data from the Lateglacial and Holocene of the Páramo de Sumapaz. The general altitudinal zonation of the páramo vegetation was studied and is presented for both the dry and the humid side of the Cordillera. The zonal and azonal plant communities are described including their physiognomy, composition and syntaxonomy, habitat and distribution. Eighty five syntaxa from the rank of variant to that of the class are newly described, 17 of which are provisional. The vegetation is not ranked syntaxonomically yet, but described on the basis of preliminary tables. A number of azonal communities, part of them of lesser extent, are described in a similar way. The páramo vegetation is primarily determined by the tropical diurnal high mountain climate. The diversity of the páramo vegetation is related to temperature (altitudinal gradient) and to humidity (dry and wet climate). The presence of zonal bunchgrass páramo, bamboo-bunchgrass páramo or bamboo páramo mainly depends on the complex interrelation between these factors. Finally a synthesis is provided on ecology, morphology and phytogeography of the páramo vegetation of the study area.

Description: The wood and leaf anatomy of representatives of the 9 genera of the Opiliaceae are described in detail. It is possible to separate the genera on the base of both wood- and leaf anatomical characters. Herein the presence of cystoliths of varying shape and size is important. Some comments on the taxonomy and possible phylogeny of the familiy are given.

Description: Isoëtes Palmeri with a distribution in the High Andes from the Páramo of Venezuela to the Páramo of Ecuador is described as a new taxon, and dedicated to the then American specialist of the genus, Thomas Chalkley Palmer (1860-1934). The new species belongs to the tropical-Andeanaustral-antarctic section Laeves, described as new here as well. The publication of the new species had to be anticipated to the projected monographic treatment of the South-American representatives of the genus Isoëtes, as A.M. Cleef, Utrecht intends to base a new association, the Isoëtetum Palmeri on this new taxon, observed and collected by him at many instances within the Colombian Páramo between 1971 and 1980 in the context of the preparation of his doctoral thesis now under way.

Description: This year Prof. Dr. F.P. Jonker, Frits as he is known among his friends, will retire from the formal academic life at the State University of Utrecht: a long and busy life of 49 years, devoted to teaching, administration, and scientific research. Looking back on all these years, one realises the important contributions that Jonker has made to botanical science in general and to palaeobotany in particular, both in The Netherlands and abroad, as well as the impact he has exerted on his surroundings, culminating in the vigorous activities of the Laboratory of Palaeobotany and Palynology at Utrecht. To describe Jonker’s life history is indeed to describe the history of his laboratory. To understand the significance of Jonker and the character of the “lab”, we have to trace his life from its very beginnings at the town of Almelo in the eastern Netherlands, where he was born in 1912. His father and mother were teachers and both liked (wild) flowers. Thus both an intellectual and botanical background were already part of his life at a very young age. Soon Jonker joined a group of boy-scouts, where he combined his love for the outdoors with his interest in nature. In high school the biology teacher was Dr. J. Van Beusekom, an Utrecht botanist, who was at the same time scout-master of the scout group. In these formative years, “de Beus” was a decisive factor in influencing Jonker’s career. It was largely because of Van Beusekom that Jonker went to Utrecht University as a student.

Description: In his introductory statements to 'The Symposium on the Phylogeny and Classification of the Filicopsida' which was held in London, 1972, HOLTTUM, when dealing with 'dubious groups of relationships which would particularly repay investigation', mentioned the Polypodiaceae first (HOLTTUM, 1973: 6). Talking about Polypodiaceae the present authors deal with the Polypodiaceae sensu stricto only, thus excluding the Cheiropleuriaceae, Dipteridaceae, Grammitidaceae, and also the Loxogrammaceae, taxa which were formerly (or are still) included in the Polypodiaceae sensu lato. As delineated in this way, this almost exclusively pantropical family consists of about 600 species and an indefinite number of genera.

Description: El 25 de mayo de 1976 falleció inesperadamente, a la edad de 53 años, Peter Arnold Florschütz, eminente briólogo y profesor de Botánica Sistemática en Utrecht (Holanda). Era bien conocido por sus estudios de los musgos de Surinam. Fue coauter del "Index Muscorum”, miembro de la comisión de la Flora Neotrópica y tesorero del IAPT. Durante sus últimos 10 años estudiaba, junto con la señora Florschütz, los musgos de los Andes colombianos. En 1972 visitó muchas zonas de páramos y selvas andinas, especialmente en los alrededores de Bogotá (Cundinamarca), la Sierra Nevada del Cocuy (Boyacá, Arauca) y el Nevado del Ruiz (Caldas). En 1975 tuvo la oportunidad de visitar nuevamente algunos páramos cercanos a la capital colombiana. Sus colecciones de 1972 y 1975 (con cerca de 1.000 números) se conservan en Bogotá (COL) con duplicados en Utrecht (U).

Description: The new species Coussapoa manuënsis C.C. Berg is described.

Description: Until recently relatively little attention has been paid to the study of chromosomes in liverworts. The first substantial contributions were made by Heitz (1927, 1928) and Lorbeer (1934). In the second half of this century chromosome studies on liverworts were mainly carried out in Europe (e.g. Fritsch 1972; Newton 1977, 1979) and Japan (e.g. Tatuno 1959; Segawa 1965a, b, c; Inoue 1968). Inoue (in Koponen 1979) reports that until now 28% of all bryophyte species in Japan have been investigated as to their chromosome complement. A comprehensive, but rather outdated, survey of chromosome numbers in Hepaticae and Anthocerotae was given by Berrie (1960). Work on a new, updated survey is now underway (Fritsch, in prep.). In the present article results are presented of a cytotaxonomic investigation of European species of the genera Aneura and Riccardia (Aneuraceae). Most specimens were gathered in the Netherlands, but some chromosome counts based on French and German plants are also included.

Description: IUCN, says the paper Categories, Objectives and Criteria for Protected Areas, ”is dedicated to the wise use of the Earth’s natural resources and to the maintenance of the Planet’s natural diversity.” What to think of the sequence? Use first, maintain second? And this comes from the International Union for the Conservation of Nature and Natural Resources? ”The World National Parks Congress, taking place in Bali, Indonesia, October 11-22, 1982, will provide case studies from around the world to illustrate how the various categories of protected areas are meeting the needs of countries of all economic, social, cultural, and political backgrounds,” writes J.A. McNeely, the secretary of the Commission on National Parks and Protected Areas, in a special issue of the Swedish journal Ambio (11: 237. 1982). ”No longer just playgrounds for vacationers and means for conserving natural heritage, protected areas have become an inseparable part of the modern human ecosystem.”

Description: Mrs. Delia D. Adefuin, Museum Research Assistant, Manila, is pursuing her M.S. in Botany degree. She is currently the Secretary of the Fern Society of the Philippines. She is working on the Fern Flora of Metro Manila and is preparing the manuscript of a pictorial encyclopedia which will include descriptions of species and horticultural recommendations. Miss Barbro Axelius (S) collected and studied Xanthophytum and Lerchea (Rubiaceae) in Sarawak, Kalimantan and Sumatra, August 1982- February 1983.

Description: Tropical Botany in Aberdeen University. This was started by Professor J.W.H. Trail, who held the chair from 1877 to 1919, and travelled in the Amazon Valley (1873-75) mainly collecting cryptogams and studying palms. He was succeeded by Prof. W.G. Craib (1920-33) who was never in the tropics but devoted his work to the Flora of Siam, based on the collections of A.F.G. Kerr, and assisted by Miss E.C. Barnett. After a considerable lag, tropical botany was revived by the energetic efforts of Dr. P. Ashton as lecturer in systematics and ecology of the eastern tropics, establishing ties with Malayan colleges in teaching and research. This is at present perpetuated by two lecturers, Dr. K. Jong and Dr. M.D. Swaine, the latter’s experience lying largely in the tropics of West Africa. In addition Dr. N.M. Pritchard, Dr. J.B. Kenworthy and Dr. G. Hadley have been on secondment to the University of Malaya, while Dr. I. Alexander made research visits to India, Ghana and Peru. Over the years the Department has provided undergraduate and research training to innumerable students from many different tropical countries, some of which attained responsible posts, e.g. Prof. E. Soepadmo. Important courses in tropical biology are given, not available elsewhere in the U.K. (started 1973). The benefits for Aberdeen students is important: amongst others they led to expeditions to various parts of the tropics, recently to Sabah and to the Ivory Coast. Royal Society Tropical Rain Forest Collaborative Research Programme. Arising out of a feasibility study by Dr. T.C. Whitmore and P.F. Cockburn, the theme ’Recovery of tropical rain forest after disturbance’ was adopted as the initial basis of the programme. Possible territories for the research include Sabah and the Philippines. Detailed plans for a 5-year project are being prepared in consultation with colleagues in Southeast Asia.

Description: It is with the utmost regret that we announce the sudden and quite unexpected death of Dr. Marius Jacobs, editor of this Bulletin. See the obituary on page 3869. He was co-editor of the Flora Malesiana Bulletin for nr. 17 (1962) to nr. 22 (1968) and took full responsibility onwards of nr. 27 (1974). He showed great ability in enlarging its scope and we have many letters in our archives expressing appreciation and admiration for the lively and informative style in which he edited the Bulletin. I had to take over the editorial work for this number at short notice, but I was greatly helped by a number of Rijksherbarium colleagues, which help is gratefully acknowledged. In this way the delay has been kept to a minimum. It is, however, possible that some news items etc. have not been printed and that information submitted to Dr. Jacobs has not been entered due to this sudden change of editorship. I offer my apologies if this has happened and hope that (if still relevant) the news will again be forwarded to the new editor, Dr. J.F. Veldkamp, Rijksherbarium, who will take over starting next number.

Description: The Botanical Survey of India continued to make collections during 1982, with the following results: Andaman & Nicobar Is.: Shola Bag, Mt Harriet, Jirkathang, Poona Nallah, Saddle peak, Diglipur, Rutland I. & Little I., 2875 specimens. Arunachal Pradesh: Various areas of Kameng Distr., Subansiri Distr., 9750 specimens. Assam: Garampani, 60 specimens. West Bengal: Darjeeling, Jalpaiguri, Hollong, Jaldapara, Chilapata, Salkumar, Daidaighat, Barasat, 2665 specimens. Bihar: Madhuban, Nimiaghat, Paresnath Hills, 315 specimens. Dehra Dun: Chakrata, Missoori, 325 specimens. Gujrat: Catchment and submergence areas of Sipu Reservoir Project, 1505 specimens. Kerala: Trichur, Idduki, Silent Valley, Valra Reserve Forest, Cannanore, Trivandrum, 3770 specimens. Madhya Pradesh: Kanha National Park, Chhodarpur Distr., 1190 specimens. Maharashtra: Areas of Jalgaon Distr. and Buldhana Distr., 4390 specimens. Manipur: Cherrapunjee, Mawphlong, Sorharim, areas of West Khasi Hills Districts, 2000 specimens. Rajasthan: Bharatpur, Desert National Park, 1605 specimens. Sikkim: Rangpo, Singtham, Bumbing, Manuring, Duga, Pandam, Takchi, Meli, areas of Gangtok, Chungtham, Lachi, Thanga, Panthang, 2590 specimens. Uttar Pradesh: Gori & Kali Valley, Chittoragarh Distr., 500 specimens.

Description: Some of the species and names of hydnoid fungi treated in Furukawa’s work are discussed. Hydnum albidum is recorded in Europe for the first time. Further finds of some interesting species are reported. Auriscalpium barbatum (Western Australia) and Steccherinum peruvianum (Peru) are described as new species. A key to the species of Auriscalpium is given.

Description: The ultrastructure of the spore walls of Beenakia dacostae (Beenakiaceae, Gomphales) has been studied. Spore walls are mainly composed of a distinct episporium and a thick, dark, ornamented ectosporium. The general structure is identical with that of other members of the Gomphales, such as Gomphus and Ramaria

Description: Subramanian’s concept of true and false chains of phialoconidia is rejected and replaced by a distinction between connected and disconnected chains. In connected conidial chains the primary conidial wall is strongly thickened at both ends and a connective is formed. This criterion allows the distinction between trichocomaceous or eurotiaceous (connected) and sphaeriaceous (disconnected) catenulate phialoconidia. The ultrastructure of conidiogenesis is described. On the basis of this criterion, the species of the Acremonium diversisporum series as well as the anamorph of Sagenoma viride Stolk & Orr with connected chains are transferred from Acremonium to the new genus Sagenomella to which four new species are added.

Description: During an ecological study of fungi of the tidal mudflats in Kuwait, a Sporothrix species has been recorded twice, in 1977 and 1980. It differs from other species of the genus (de Hoog, 1974, 1978) in several characters and is here described as a new species. A comparison with similar species of the genus is added.

Description: In the course of a study of coprophilous fungi collected in the Zoological Garden in Delhi, an ascomycete belonging to the genus Achaetomium was isolated. It differs from previously described species by larger ascospores and almost colourless ascomata with a wide apical opening.

Description: Among the collections of Knema acquired by the Rijksherbarium since the publication of my new account of the genus Knema, in Blumea 25, 1979: 321 — 478, a few specimens caused problems with the identification, and at closer examination these yielded facts of interest which are published here. Some specimens represented stages not yet known, for instance fruits, or male flowers, while other specimens meant a significant range extension of the species. Two new species and one new subspecies are described. For easy reference, the sequence and numbers of the species presently treated correspond with the numbers as used in the account of 1979. The new species bear the number of the species after which they appear in the general key of 1979, with the addition ‘-bis’.

Description: The early development (ontogeny) of the carpels of 20 species belonging to 8 apocarpous families was investigated with the scanning electron microscope. The results indicate that on the floral apex a circular or a convex meristem develops into an obliquely ascidiate primordium by unequal growth of its periphery. By further unequal growth it develops into a young carpel. The terminal mouth of a cup becomes the lateral cleft of a carpel. The different forms of the young carpels in different species are defined by the varying degree of development of the adaxial region of the initial meristem and/or its margin on the side of the floral apex. This hypothesis is theoretically evaluated.

Description: In Malesia the genus Tephrosia is represented by 20 species, native or introduced and naturalized, including 6 subspecies and 5 varieties; 4 species are restricted to Malesia. Two of these species are newly described: T. barbatala and T. elliptica; the former includes one new variety (var. glabra). Two new subspecies and one new variety are distinguished: T. filipes subsp. longifolia, T. purpurea subsp. barbigera, and T. maculata var. elongata. Two taxa are given a new status: T. maculata var. appressepilosa and T. pumila subsp. aldabrensis. Four species, T. dichotoma, T. repentina, T. coarctata, and Kiesera sumatrana are for the first time sunk into synonymy. A key to the taxa, synonymy, and full descriptions with plates and distribution maps are given.

Description: During identification and study of the materials of the genus Blepharis Juss. in the Central National Herbarium (CAL) of the Botanical Survey of India the authors came across some specimens (Lawson 314) from Tavancore, which was identified as B. molluginifolia aff But critical study revealed that the typical molluginifolia is a different species from which the new taxon can be distinguished by several characters. The species is named after M.A. Lawson who collected the specimen for the first time.

Description: Identities are given for all 35 combinations published in Anplectrum A. Gray ( Melastomataceae) with additional notes on some of the taxa involved. Two new combinations are proposed in Creochiton Bl. and one in Dissochaeta Bl.

Description: A new monotypic genus Tamaricaria Qaiser & Ali of Tamaricaceae is described with a new combination i.e. Tamaricaria elegans (Royle) Qaiser & Ali.

Description: 1. The interrelations between the genera together constituting the Dimocarpus group in the tribe Nephelieae are represented in a scheme. In this scheme are added the main characters that are thought to be of phylogenetic importance. 2. A neotype.is proposed for Cubilia cubili (Blanco) Adelb., the single species of its genus. To its distribution can be added the eastern half of Borneo, incl. also the Island of P. Laut. Mention is made of a geographic clinal variation in a few macromorphological characters. 3. Lilchi is considered to comprise only one species, L. chinensis Sonn., which is subdivided into three subspecies: subsp. chinensis, the commonly grown form, cultivated for thousands of years already, apparently adapted (by nature or partly by selection by man?) to a monsoon climate, if actually wild probably originating from northern Indo China; subsp. philippinensis (Radlk.) Leenh., a wild form closely related to subsp. chinensis, known from the Philippines and New Guinea; and subsp. javensis Leenh., strikingly different from both other forms, known only as a cultivated fruit tree from southern Indo China and Java, apparently adapted to an everwet tropical climate. For subsp. philippinensis a lectotype is proposed. 4. Pometia, though macromorphologically distinctly derived and, moreover, palynologically apparently very exclusive in the alliance under discussion, seems clearly connected with Dimocarpus, the central genus in the group.

Description: The stomata as occurring on the fronds of the sporophytes of a large number of Polypodiaceae s.s. (Filicales) are investigated. A number of different stomatal types is recognised, (newly) described, and their ontogeny investigated. The different types of stomata are discussed in relation to their possible significance for tracing phylogenetic relationships in the Polypodiaceae following a cladistic analysis.

Description: Description of a new species, Diploglottis bracteata Leenh., from Queensland, Australia. Reduction of Euphorianthus (E. Malesia) to Diploglottis (NE. Australia). Discussion of the occurrence of actinomorphic and zygomorphic flowers in the Sapindaceae in nearly all tribes and even within ten genera. Discussion of the systematic position of Diploglottis bracteata: this species seems distinctly allied to and more derived than the New Caledonian genus Storthocalyx, and thus may belong to an old element of the Queensland flora allied with that of New Caledonia. On the other hand, D. bracteata is within Diploglottis closest to the East Malesian species, whereas the further Australian species are distinctly more derived. They may belong to a younger element in the Queensland flora of Malesian derivation.

Description: Herbae erectae caespitosae 1.5—2.0 m altae; rhizoma suberectum 1—2 cm diametro. Folia multa apice rhizomatis spiratim aggregata; petioli usque ad 1 m longi basi vaginantes, sparsim pubescentes; laminae ellipticae oblongo-lanceolatae glabrae 18—35 cm longae, 8—15 cm latae, basi acutae vel cuneatae, apice breviter acuminatae, infra subvirides, nervis lateralibus multis parallelis, nervulis multis scalariformibus inter nervos subtiliter transverse reticulatis; costa basi incrassata gradatim angustata et a nervis lateralibus haud distincta. Pedunculus principalis terminalis usque ad 1.5 m altus ad apicem folio uno inflorescentia unaque, raro folium deest; folium pedunculare foliis primariis simile petiolo ad medium usque vaginante parce piloso, apicem versus pubescente. Inflorescentia laxe paniculata e spicis spiculisque constans, in axillam vaginae folii pedunculati, pedunculo ad 15 cm longo, bractea basi vaginante lineari-lanceolata puberula ad 10 cm longa; panicula puberula, 2—10-ramosa usque ad 15 cm longa et 10 cm lata; spicae 2—10, longitudine variantes, 3—8 cm longae usque ad 4 cm distantes e axillis bractearum principaliarum spiratim dispositae in rachibus principalibus; bracteae principaliae ovati-lanceolatae ± 2 cm longae, ± 1 cm latae, extus puberulae margine hirtae, cito ad medium usque, tandem basem versus in fibras multas secedentes; spiculae multae e axillis bractearum primariarum subspiraliter in spicis dispositae, usque ad 2 cm longae; bracteae primariac ovatae obtusae mucronatae ± 1.5 cm longae, ± 1 cm latae, extus puberulae margine hirtae bracteis principalibus similes. Flores albi in cymis parorum 1—3 vel plurorum; omnis par e axilla prophylli, flos unus paris omnis primo aperiens, ceterus invicem; prophyllum externum magnum dorsaliter 2-carinatum ovato-lanceolatum margine incurvum ± 14 mm longum et ± 5 mm latum, alis carinarum fimbriatis; prophylla interiora dorsaliter 3-carinata parva; pedicelli omnis paris basi connati ± 1 mm longi. Calycis lobi usque ad basim discreti, lineari-lanceolati extra hirti,± 1.2 cm longi, ± 2 mm lati. Corolla basi tubularis, tubo ± 7 mm longo lobis oblongis ovatis erectis obtusis ± 8 mm longis. Staminodia 4 in tubum connata tubum corollae adnata; lobi staminodiorum extemorum subaequales; lobus magnus obovatus obtusus marginibus recurvatis, ± 1 cm longus, ± 8 mm latus; lobus parvus subspathulatus recurvatus, ± 7 mm longus, ± 3 mm latus; staminodium interioris carnosum cucullatum, lobo laterali apicem styli includenti, ± 6 mm longum et ± 3 mm latum; staminodium interiorissimum cucullatum, staminodium extimum magnum omnino adnatum apicem stigmatis includens, infra stigma callo carnoso puberulo instructum, lobo ad laterem stigmatis auriculato. Stamen fertile 1-cellulare appendice subulata ± 1 mm longa uno latere instructum; filamenti segmentum librum ± 3 mm longum; anthera oblonga apiculata lutea ± 2 mm longa. Ovarium ±2 mm longum dense aureo villosum; stylus tubum staminalem adnatus ± 1 cm longus segmento libro cucullato ± 4 mm longo; stigma truncata irregulariter lobata. Fructus 3-lobatus sparsim breviter hirtus rubiginosus ± 9 mm longus, ± 7 mm latus; semina 3 oblonga subtrigona lurida ± 7 mm longa et ± 4 mm diametro arillis 2 lobatis albis lobis lineari-acuminatis ± 3 mm longis. Erect caespitose herbs, 1.5—2.0 m tall; rhizomes suberect, 1—2 cm thick. Leaves many, spirally crowded at apex of rhizome; petiole sheathing at base, up to 1 m long, sparsely pubescent; leaf-blade elliptic to oblong-lanceolate, acute to cuneate at base, shortly acuminate at apex, 18—35 cm long, 8—15 cm wide, glabrous, whitish green beneath; lateral nerves many, parallel, with many fine scalariform transversely reticulate nervules in between; midrib thickened towards base, gradually narrowed and indistinct from lateral nerves towards apex. Main peduncle terminal, up to 1.5 m tall, bearing one leaf and inflorescence at apex, rarely without leaf; peduncular leaf similar to primary leaves; petiole up to 15 cm long, sheathing at basal half, scattered hairy, pubescent towards apex. Inflorescence Inflorescence a lax panicle of spikes and spikelets, arising from axil of peduncular leafsheath with up to 15 cm long stalk, subtended by up to 10 cm long, linear-lanceolate, sheathing, puberulous bract at base; panicle up to 15 cm long, up to 10 cm wide, 2—10- branched, puberulous; spikes 2—10, of various length, 3—8 cm long, arranged at distances of up to 4 cm apart in axils of spirally arranged bracts on main rachis; bracts ovate-lanceolate, ± 2 cm long, ± 1 cm wide, puberulous outside, hairy at margins, soon splitting up in upper half into many fibres, later up to base; spikelets many, subspirally arranged on spikes in axils of primary bracts, up to 2 cm long; primary bracts ovate, obtuse, mucronate, ± 1.5 cm long, ± 1 cm wide, puberulous outside and along margins, similar to main bracts. Flowers white, in cymes of 1—3 or more pairs; each cymule (pair of flowers) in axil of a prophyll, one flower in each pair opening first, the other next; outer prophyll large, 2-keeled at back, ovate-lanceolate, ±14 mm long, ± 5 mm wide, incurved at margins, minutely fringed on wings of keels; inner prophylls 3-keeled at back, smaller; pedicels of each pair of flowers united at base, ± 1 mm long. Calyx-lobes free up to base, linear-lanceolate, ± 1.2 cm long, ± 2 mm wide, hairy outside. Corolla-tube ± 7 mm long; lobes oblong, ovate, obtuse, ± 8 mm long, erect. Staminodes 4. united into a tube and adnate to the corolla-tube; lobes of outer staminodes subequal; larger lobe obovate, obtuse, with recurved margins, ± 1 cm long, ± 8 mm wide; smaller lobe subspathulate, recurved, ± 7 mm long, ± 3 mm wide; fleshy inner staminode hood-shaped with a lateral lobe enclosing style apex, ± 6 mm long, ± 3 mm wide; innermost staminode hooded, entirely adnate to outer large staminode, enclosing lip of stigma, with a fleshy puberulous callus below stigma and an auricular lobe at side of stigma. Fertile stamen 1-celled, with a thin, subulate, ± 1 mm long appendage on one side; free portion of filament ± 3 mm long; anther ± 2 mm long, oblong, apiculate, yellow. Ovary ± 2 mm long, densely golden hairy outside; style adnate to staminodial tube, ± 1 cm long; free portion curved, hooded, ± 4 mm long; stigma irregularly lobed, truncate. Fruit 3-lobed, ± 9 mm long, ± 7 mm wide, sparsely short hairy, reddish-brown; seeds 3, oblong, subtrigonous, ± 7 mm long, ± 4 mm thick, dull brown; arils 2, lobed, white; lobes linear-acuminate, ± 3 mm long.

Description: The Star Mountains of New Guinea are situated at the geographic center of the Island of New Guinea extending on both sides of the Indonesian-Papua New Guinea border. Access to these mountains from either side of the border which divides the island is relatively difficult and as a result few collections have come from the area. A Dutch expedition traveled to the western Star Mountains in 1959, but ran into various difficulties and as a result did little collecting above 1500 m. In 1975 an expedition sponsored jointly by the Division of Botany, Lae, and the Rijksherbarium, Leiden, returned to the Star Mountains collecting extensively throughout the eastern half of the range. The results of this expedition include the first extensive collections of material from the higher altitudes within the Star Mountains. Material collected for the Division of Botany, Lae, by J. R. Croft and G. S. Hope while on the 1975 expedition is surprisingly rich in species of Rhododendron. I was asked by Mr. Croft to examine the Lae material prior to its distribution. The collections contain representatives of several poorly known species of the genus, at least one new plant record for Papua New Guinea, Rhododendron rubrobracteatum Sleumer, and the new taxon described below.

Description: One new species of Aristolochia, A. singalangensis, from Sumatra is described here. This is the only Malesian species to have the fruit dehiscing from the apex towards the base. Remarks are given for some Asiatic and Malesian species, all belonging to Aristolochia except one to Thottea, on their taxonomy, nomenclature, typification, characteristics for identification, relationship, distribution, etc. The phenomena and significance of aristolochiaceous plants-butterflies relationship have been discussed. Germinated pollen grains have been found in dehisced anthers of open flowers in both Aristolochia and Thottea. Pollination of the Aristolochiaceae has been concisely reviewed.

Description: A world-revision of Arthraxon Beauv. ( Gramineae) is presented. Three wide-spread species, A. hispidus (Thunb.) Makino, A. lanceolatus (Roxb.) Hochst., and A. lancifolius (Trin.) Hochst. are very variable and have caused the description of a great number of taxa, most of which are here reduced to synonomy. There are now 7 species and 9 varieties; for 6 of the latter new combinations are proposed. No new taxa are described.

Description: In this fourth report on the taxonomy of Ulothrix Kützing a new classification of the marine and brackish-water species in western Europe is proposed. Comparative studies on field collections, uni-algal cultures, herbarium collections and sections prepared for electron microscopy lead to the recognition of three marine species, viz. Ulothrix speciosa (Carmichael ex Harvey in Hooker) Kützing, U. flacca (Dillwyn) Thuret in Le Jolis, and U. palusalsa Lokhorst (nov. nom.), and two brackish-water species, viz. U. implexa (Kützing) Kützing and U. subflaccida Wille. The vegetative anatomy, the life history, the fine structure of the vegetative thallus and the distributional pattern in nature are amply discussed. Salient, reliable characters proved to be, e.g., the nature and construction of the cell wall, the texture of the cell wall’s surface, the fine structure of the pyrenoid, the developmental stages of germinating zoospores, the coalescense of filaments, the shape of the gametangial filament, and the limited variation of the number of zoospores and gametes. A brief discussion is given of the ecological status of the individual species. In addition there is a brief comment on the taxonomic affinity of Ulothrix with the morphologically related genus Urospora Fries and on the phyletic relationship of Ulothrix with the progenitors of the higher land plants. The reproductive behaviour of the species under different photo periods in culture appeared to be correlated with the seasonal periodicity expressed by the algae in nature.

Description: There has been a notable divergence of opinion in the recent literature concerning the number of species of dammar (Agathis) that might occur in the general area of the Moluccas. Even early literature presents a tangled nomenclatural trail. Perhaps the difficulty in obtaining good representative collections from these huge rainforest emergents may explain the general lack of careful diagnostic descriptions that bedevils most contributions. Among the hundreds of specimens I have been able to study, however, I have found enough data to support a clear conclusion. The important dammar tree was among those described in the early work by Rumphius (1741) that dealt with Ambon. Meijer Drees (1940) reports that natives in the Moluccas recognize two types of dammar, the ‘white dammar’ damar putih) with abundant resin production and the ‘brown dammar’ damar merah) with poor resin production (the ‘white’ or clear resin does turn brown upon aging about a year). Presumably, Rumphius, who spoke of abundant resin, had in mind the ‘white dammar’ when he referred to this tree as Dammara alba.

Description: There are only two genera of the Aristolochiaceae, Aristolochia and Thottea, so far known to occur in Malesia. In the course of a revision of this family for the Flora Malesiana, some new species of both genera have been found. Eight new ones of Thottea were published in a precursor on that genus (Blumea 27, 1981, 301-332, f. 1-72). There are four new species of Aristolochia from Borneo and one more new Thottea from Sumatra to be described here.

Description: A world-wide revision of the tribe Naucleeae with a general discussion of the systematic position and affinities of the tribe and the genera. The generic concepts have been modified and 21 genera are recognized (Ochreinauclea, Ludekia, Diyaminauclea, Khasiaclunea, Adinauclea, Sinoadina, Pertusadina, and Haldina being new), which are placed in three subtribes, Anthocephalinae, Naucleinae, and Adininae, sublrib. now There are keys to the subtribes, genera, and species, followed by descriptions of the Asiatic and Malesian genera. The Asiatic species are described and accompanied by complete synonymy, but the Malesian species are treated in an abbreviated form. Three new species are described: Myrmeconauclea stipulacea, Ludekia borneensis, and Pertusadina malaccensis.

Description: The species related to Medinilla myrtiformis (Naud.) Triana are described and a key is given. A variety of M. monantha Merr. is recognized as a distinct species, while the typical form is considered a synonym of M. myrtiformis. M. neglecta Nayar is reduced to M. rubrifructus Ohwi.

Description: Carpels develop centripetally as oblique slightly ascidiform structures provided with a seat-like support. The terminal mouth of the ascidiform primordium becomes the lateral cleft of the carpel. Solitary terminal carpels develop as lateral structures. The sections Tasmannia and Drimys of the genus Drimys differ by the varying degree of ascidiform development. Austrobaileya is like Drimys. The structure of the carpel margins with submarginal placentation may have evolved from a slit like that in Winteraceae. Allometric development of an oblique ascidiform structure may have formed a large basal ovary, and may thus have moved the stigmatic part apically. A possible cupular origin of carpels is discussed.

Description: The type species of tie genus Polyscias (P. pinnata J. R. & G. Forst.) is closely related to a small number of Pacific and Indo-Malayan species, several of which have long been in cultivation. This group of species have a distinctive facies but can be defined most readily by the elongated sheathing leaf-base. The genus has usually been extended beyond this group to include other pinnate-leaved members of sub-family Schefflereae in which the pedicel is articulated below the flower. There has been uncertainty whether to restrict the genus to species in which the style arms are free or also to include species with connate styles. In his treatment of the New Guinea species, Harms (Bot. Jahrb. 56,1920: 374—414) does include some species with connate styles within Polyscias though also retaining the genus Kissodendron, a genus distinguished from Polyscias mainly by the united style arms. Bernardi (Candollea 26, 1971: 13—89) resolved this difficulty by uniting Kissodendron, and also Palmervandenbroekia, with Polyscias and this treatment is followed here. As thus defined, the genus comprises rather diverse elements, and these have been given sectional status.

Description: A species with the habit of Aristida divaricata H. et B., but welldistinguished by the wanting column and the curious tuberculate lemmas. Closely allied to Aristida gentilis HENR., which differs however in the other position of the glumes and in the smooth lemma. The species resembles in some characters the Aristida Parishii HITCHC., the latter has however a totally different shape of the panicle and the lemma is not tuberculate-hispid, but scabrous only on the upper half. Among duplicates from the U. S. Nat. Herbarium, kindly received from Mrs. A. CHASE, I found a second plant belonging to the species, a plant also collected in Arizona, north slope of Santa Rita Mountains, leg. D. GRIFFITHS no. 7269.

Description: Owing to their limited possibilities for either active or passive dispersal, their association with the soil habitat, their vulnerability towards a dry atmosphere, and, in fact, on account of their general ecology and ethology, Diplopoda among arthropods are surely one of the most important classes in relation to the study of historical biogeography. For the class as a whole the sea appears to be an unsuperable barrier as is proved by the almost complete absence of endemic taxa on oceanic islands. In many cases lowland plains also act as severe obstacles against the dispersal of millipedes. The presence or absence of diplopods on islands or continents, therefore, may give a strong argument in favour or against any supposed former land connection. The long geographical isolation of the Australian continent and the absence of endemic higher taxa seems to imply that most, if not all, of its diplopod fauna dates from the time this continent was solidly attached to other southern continents, i.e. the Mesozoic. Subsequent penetration of fauna elements from the north or northwest seems utterly unlikely, although perhaps not entirely impossible.

Description: 1. Samples of benthic invertebrate fauna were collected in – a ditch, situated in a protected area in the peaty area near the Maarsseveense plassen (prov. Utrecht), – four ditches in the neighbourhood of Castricum (N-Holland), not far from the dune district, – two ditches with water of a high chloride-content on Texel (N- Holland). 2. The faunistic results, with the additional information of some physical and chemical data on the different sampling points, made it possible to draw some conclusions concerning – the importance of the factor salinity for a faunistically oriented typology of ditches, – the importance of the group Hydracarina for a classification of ditch-biocenoses, – the influence of artificial current, caused by the inlet of water, on the invertebrate fauna of ditches and on the faunistical evaluation of the pollution-level in ditches.

Description: The buthid scorpions of the genus Centruroides are widely distributed in the Antillean area. They are also the most common scorpions in the majority of these islands. Nevertheless they remained almost forgotten until recently (STAHNKE 1970; ARMAS 1976, 1977, 1980, 1981, 1982; FRANCKE & SISSOM 1980). For the purpose of this paper the Lesser Antilles are defined as the islands situated between the Virgin Islands and Trinidad & Tobago, according to BOND’S 1978 zoogeographical point of view.

Description: Recent investigations of the distribution of trace elements in metamorphic index minerals of metapelites have revealed, that the plurifacial character of the Hercynian metamorphism in this area is confirmed by the distribution of Yttrium in Hercynian garnets of the metamorphic series.

Description: The formation of thick piles of flysch-like sediments needs the existence of narrowed seas, active denouement of neighbouring continents, and generalized marginal subsidence. These conditions are present during the initial and final stages of Wilson’s perceptive cycle. In this context, the Late Precambrian flysch of the Iberian Massif must be related to the initial rifting, whilst the Culm of southwestern Iberia was accumulated during an episode of Upper Palaeozoic subduction that remained active after the impingement of Iberia against North America. Culm sediments shed from the uplifted collision zone and fed into a remnant ocean that remained at the nonsutured southern border of Iberia. This model of synorogenic flysch formation has been described elsewhere for similar plate arrangements. On other grounds this model provides a framework that explains the different structural and magmatic trends of the Ossa-Morena Zone (near the active margin) in the context of the rest of the Massif (basement reactivation). In addition to this, it seems to support a partly primary origin for the Iberian arc versus a secondary origin.

Description: During the Late Devonian, deposition in the Cantabrian Mountains was largely controlled by movements along faults. By way of intermitting subsidence of the area south of the Sabero-Gordón line and the connected progradation of the coast during the Frasnian and early Famennian, three regressive sequences were deposited. On account of these sequences the Nocedo Formation is divided into three units. Before the late Famennian transgression, after which the upper part of the Ermita Formation was deposited, the area was peneplained. Six facies maps show the changes in palaeogeography during the Late Devonian.

Description: The area described comprises the SE plunging extension of the anticlinorium of Mondoñedo-Lugo-Sarria and several structural units to the NE of it. The stratigraphy compares with that of neighbouring areas. Certain new observations have been made regarding the west flank of the East Galician-West Asturian miogeosyncline at the end of the Precambrium and during the Early Palaeozoic. Stromatolites or algae played an important part in the formation of carbonate deposits during late Precambrian (Cándana limestone) and Lower Cambrian time (Vegadeo limestone). An imprint, possibly attributable to a specimen belonging to the Precambrian Ediacara fauna, was found in the Cándana Schist Formation. A shallow or relatively high zone existed during that time in the region around Incio. The same general area formed a high again at the end of the Ordovician, causing erosion and a marked disconformity of the Silurian. The Hercynian structures can be explained by assuming roughly WSW-ENE compression and shortening. In the NW of the area this is expressed in the first place by the recumbent folds of the Lugo-Sarria anticlinorium, and in the SE by the narrowness of the steep folds and their slightly changed direction in the general area of Seoane and El Cebrero. Normal crossfaults are related to the folding and the compression. They show locally a deviation in accordance with the direction of shear stress. Horizontal displacement along such faults in an EW direction was probably of little importance. Finally a few observations are given on the occurrence of Pb-Zn ores at the mine of Rubiales.

Description: Rosacea flaccida, a new prayine siphonophore, is described from specimens collected by SCUBA divers in the upper 30m of the subtropical and temperate North Atlantic Ocean. The new species has stoutly cylindrical, flaccid nectophores and delicate flattened bracts. The nectophores are morphologically similar to those of R. plicata sensu Bigelow, 1911 and R. cymbiformis (delle Chiaje, 1822) having a deep hydroecial groove and meandering lateral radial canals in the nectosac. In one of the nectophores there is a slight dorsal prolongation of the somatocyst at its apical end into the mesoglea. The eudoxid bracts are distinctive, being flattened dorsoventrally and divided, on the proximal side of the stem, into two lobes which are twisted at an angle of approximately 90° to the lobe on the distal side. Right and left longitudinal bracteal canals are well developed. The origin of the dorsal bracteal canal from the right longitudinal canal differs from that in the other Rosacea species but resembles the configuration found in the bracts of species of the genus Praya.

Description: Species of the heteropod families Carinariidae and Pterotracheidae collected in the Mid North Atlantic Ocean in 1980 have been studied. The distribution of the species is given, as well as the morphological variation. It is concluded that two subspecies of Carinaria lamarcki actually have to be considered distinctly recognisable and sympatric species, to be called C. lamarcki and C. challengeri. The vertical distribution of the heteropods studied proves not to be restricted to the photic zone and diurnal vertical migration occurs among the larger species. The horizontal distribution of the two Carinaria species and of at least two populations of the Pterotrachea species coincides with the southern branch of the North Atlantic Current, while others are restricted to the subtropical waters. Consequently, the present heteropods have to be considered good indicators of water masses and currents.

Description: Moult cycle and morphogenesis in larval instars (zoea I, zoea II, megalopa) of the spider crab Hyas araneus were studied in the laboratory. Changes in the epidermis and cuticle were documented photographically at daily intervals to characterize the stages of the moult cycle. Stage A (early postmoult) is a very short period during which the larva takes up water. During late postmoult (B) and intermoult (C) the endocuticle is secreted, and there is conspicuous epidermal tissue condensation and growth. The onset of early premoult (D sub(0)) is characterized by epidermal apolysis, occurring first at the bases of the setae in the telson of zoeal instars or in the rostrum of the megalopa.

Description: Larvae of the crabs Menippe mercenaria Say (Menippidae), Panopeus herbstii Milne-Edwards, Neopanope sayi Smith (Xanthidae), Sesarma cinereum Bosc (Grapsidae), and Libinia emerginata Leach (Majidae) were reared in the laboratory. Starvation periods different in length and timing within the first zoeal stage were studied as to their effects on later development and survival rate. After 1-3 days of initial feeding, most larvae had accumulated enough reserves to reach the second stage, independently of further food availability. The development of the survivors was delayed in the following stages, and their later mortality rate was higher than the fed controls. Starvation periods commencing directly after hatching of the larvae exert far stronger negative effects than those beginning later. All observations suggest a particularly sensitive phase in the beginning of larval life in brachyurans. When initial starvation periods exceed the point-of-no-return (PNR), the larvae will die later, even if feeding begins long before the energy reserves are depleted. Temporary lack of suitable prey may be an ecological factor controlling the survival of crab larvae as effectively as physical factors.

Description: Resistance to starvation in early larval stages of six species of brachyuran crabs representing four families was observed at various constant temperatures. In the optimal temperature range of 25-30°C for these warm temperate crab larvae, survival time of starved zoeae was longer than the development duration time in fed zoeae, while at lower temperatures the relationship of these two duration periods became inversed. This response pattern is found in larvae of the mud crab Rhithropanopeus harrisii and is considered to be typical for warm temperature brachyuran larvae. It indicates that reserved utilization is strongly controlled by temperature, but not to the same degree as development.

Description: Larval and early post-larval growth has been investigated in H. araneus L. (Majidae) reared in the laboratory. Growth was measured as dry weight (DW), ash-free dry weight (AFDW), carbon (C), nitrogen (N), hydrogen (H), gross biochemical constituents (protein, lipid, carbohydrate, chitin, ash) and energy (calculated separately from carbon and biochemical composition). During larval development, i.e. from freshly hatched zoea-I to late megalopa, all these criteria of biomass increase by factors ranging between 5 and 14; carbohydrate shows the lowest, chitin the highest increment. There are indications of loss in organic body weight during the latest period preceding metamorphosis to the crab stage. When no food is offered during this time, megalopae lose significantly more biomass than control larvae. This suggests that food is still required, but feeding activity is reduced to a level below maintenance ingestion rate. Following metamorphosis, the juvenile crab accumulates biomass at a far higher absolute rate (expressed as µg/d) than all larval stages.

Description: Epifaunal community development on plexiglas panels was observed over one year. The course ofcolonization is described, and some data are presented on autecology, reproduction, and growth rate ofparticular species. About three months after initial settlement, conditions of coexistence in a mixed barnacle-ascidian community began to change increasingly due to heavy competition for space. The colonial species Botryllus schlosseri proved to be potentially dominant. Shortly before it attained monopolization by replacing barnacles (mainly Elminius modestus), a major physical disturbance eliminated the fast growing ascidian. The roles of physical factors, of biological interactions, and of historical events in community development are discussed in context with succession theory and other concepts evolved more recently. It is concluded that succession-like processes can occur in subtidal fouling communities, but there the existence of a globally stable climax is unlikely. Generally the concept of multistable points seems to be better applicable to marine ecosystems than that of succession in the classical sense.

Description: The development of hard bottom communities has been studied on test panels in Helgoland Harbour (German Bight) since 1977. Settlement and growth of epibenthic species was examined monthly. Natural variation in different seasons, years, and at three stations (the latter, only in 1981 and 1982) was investigated. At Station A (Binnenhafen), barnacles (Balanus crenatus ) and polychaetes (Polydora ciliata ) were always among the first settlers in spring. They were followed by other barnacles (Elminius modestus, Balanus improvisus ) and by colonical ascidians (Botryllus schlosseri ). The latter species often dominated from August to October, and tended to overgrow the barnacle populations.

Description: The larval development f the spider crab Hyas araneus L. was studied in the laboratory at different constant temperatures (2,6,12, and 18 °C). Linear relationships between log temperature and log stage duration were described by means of regression equations. They were used in a simple simulation model predicting larval moulting and metamorphosis at different temperatures. The most important predictions were : (1) Settlement of H. araneus takes place mainly from late May to mid-June; (2) assuming wide annual fluctuations in the average spring temperature ( plus or minus 2 °C), a total settlement period ranging from late April to early July can be expected; (3) delay of larval development due to a decrease in temperature is stronger than the acceleration caused by an equally great increase; and (4) both delay and acceleration effects become weaker during the hatching season and from stage to stage.