ALBERT

Description: Males of Eledone cirrhosa grow to a size little over 600 g and normally have well-developed, and presumably active, reproductive organs from about 200 g upwards. Total weight of the genital bag is well correlated with total body weight (r= 0·906). Growth of the testis precedes that of the spermatophoric sac, and the size of neither of these reproductive components is predictable from body weight. The sizes of these organs and the estimated number and length of stored spermatophores are given for 100 g intervals of total body weight. No evidence was obtained for a seasonal trend in male maturity.

Description: We present chemical data on magmatically heterogeneous pyroclastic deposits of late Quaternary age erupted from zoned magma systems underlying Tenerife (Canary Islands), Sao Miguel and Faial (Azores), and Vesuvius. The most fractionated magmas present at each centre are respectively Na-rich phonolite, trachyte, and K-rich phonolite. Within any one deposit, chemical variation is either accompanied by changes in the phenocryst assemblage (petrographic zonation) or is largely manifested in trace element abundances, unaccompanied by any petrographic change (occult zonation). Zoning is analogous to that in calc-alkaline systems where the most fractionated products are high-silica rhyolites. When a range of magma types are considered, a correlation emerges between roofward depletion of trace elements (especially REE) in the zoned system and compatability of those same trace elements in the accessory phenocryst phases present. Thus, allanite- or chevkinite-bearing rhyolitic systems are light-REE depleted roofwards, the sphene-bearing Tenerife system is middle-REE depleted roofwards, the melanite-bearing Vesuvius system is heavy-REE depleted roofwards, while the Azores systems, which lack these phases, display roofward REE enrichment. Therefore, the behaviour of trace elements may in each case be explained by fractionation of observed phenocryst assemblages. The resemblance between features of zoned magma systems and published work on the dynamic consequences of cooling saturated aqueous solutions prompts us to suggest that sidewall crystallization and consequent boundary-layer uprise to form a capping layer at top of the system may be a plausible mechanism for the generation of both petrographic and occult zonation. Reverse zoning occurs among the first-erupted tephra of some deposits, demonstrating that the most highly differentiated magma available is not always the first to be tapped during an eruption from a zoned system.

Description: Female Eledone cirrhosa held in aquarium conditions for periods of time of five daysand over show relatively enlarged ovary sizes. Values for ovary index considerably exceed thoseof freshly caught animals and the incidence of the final stage of maturity, in which eggs pack the oviducts, is greater. A comparison of maturity indices for fresh and aquarium males was inconclusive. The range of factors associated with aquarium conditions is briefly reviewed and it is concluded that studies of cephalopod reproductive maturation must distinguish fresh and aquarium animals. Introduction External factors effective in inducing sexual maturation in cephalopods have been suggested several times. The influence of the absence of light has been implicated since the experiments of Wells & Wells (1959) showed that blinded Octopus vulgar is matured precociously. An effect of short day length in stimulating the optic glands of Sepia has been found by Defretin & Richard (1967) and Richard (1967) but this is not clearly the case for Octopus (Buckley, 1977). Octopuses kept in aquarium conditions for lengthy periods are said to have larger relative gonad sizes than those fresh from the sea (Wells & Wells, 1975). One of the factors associated with aquarium conditions is often a degree of starvation, and this circumstance alone is held to be a factor in inducing precocious sexual maturation in Eledone (Mangold & Boucher-Rodoni, 1973). In the course of recent studies on the growth and reproduction of Eledone cirrhosa from the North Sea (Boyle & Knobloch, 1982,1983,1984) animals which had remained in aquarium conditions for 5 days or over were separated from the analysis.

Description: A new species of a monogenean Isancistrum subulatae (Gyrodactylidae) has been discovered on the arms and tentacles of the cephalopod mollusc Alloteuthis subulata at Plymouth and I. loliginis, on the gills of the same host, has been re-discovered for the first time since its original description in 1912. I. subulatae, like other gyrodactylids, is viviparous, and has been shown by experiments to transfer to new hosts by contagion. In nature such transfers probably take place during copulation of the hosts and since the parasite may occur in numbers of several thousands per host, it may thereby constitute a venereal disease.

Description: This is the first detailed analysis of cephalopod beaks from the stomach of a northern bottlenosed whale, Hyperoodon ampullatus (Forster, 1770). The digestive action of many predators barely affects the chitinous beaks of cephalopods and some cetaceans accumulate the beaks in considerable numbers in their stomachs. The present beaks are clean and unbroken. Identification of cephalopod beaks from stomachs of predators such as sperm whales (see Clarke, 1977), seals (Clarke & Trillmich, 1980) and albatrosses (Clarke, Croxall & Prince, 1980) throws considerable light on the biology and relative ecological importance of the species of cephalopods concerned as well as providing useful information on the diet of the predators.

Description: Cephalopod statoliths are paired calcareous stones which lie in cavities, the statocysts, within the skull. They have a form which, though variable, shows promise as a source of criteria for taxonomic and evolutionary studies. As a preliminary to more detailed studies, Clarke (1978) published a description of the form of a generalized teuthoid statolith, coined nomenclature for the various parts and gave a very brief survey of variation of statoliths within the living Cephalopoda. This nomenclature was used in a detailed description of fossilized teuthoid statoliths by Clarke & Fitch (1979). Here, descriptions of the statoliths of the living species Berryteuthis magister (Berry, 1913), Gonatopsis borealis Sasaki, 1923, Gonatopsis (Boreoteuthis) makko Okutani & Nemoto, 1964 and Gonatus fabricii (Lichtenstein, 1818) are given and the fossil Berryteuthis species described in outline by Clarke & Fitch (1979) is compared with B. magister. A statistical analysis of measurements of the statoliths of these five species has been made and the results are presented. This forms the first part of a general description of teuthoid statoliths and similar studies on the Ommastrephidae and the Loliginidae are in preparation.

Description: A new technique for the harmonic analysis of current observations is described. lt consists in applying a linear band pass filter which separates the various species and removes the contribution of non-tidal effects at intertidal frequencies. The tidal constituents are then evaluated through the method of least squares. In spite of the narrowness of the fitter, only three days of data are lost through the filtering procedure and the only requirement on the data is that the time interval between samples be an integer fraction of one day. This technique is illustrated through the analysis of a few French current observations from the English Channel within the framework of INOUT. The characteristics of the main tidal constituents are given.

Description: 14C concentrations, as weil as 13C, hydrographic and nutrient data are reported for 5 hydrographic stations that form a transatlantic section near 40° N ("Meteor" cruise no. 23, 1971). Precision (for 14C ± 0.3% or better) and comparability with literature data are specified. A planned intercomparison with the US GEOSECS program within the Newfoundland Basin deep water failed because of variability of water characteristics. The observed 14C values decrease from about Δ 14C = + 80‰ at the surface to -70‰ at 2000 m depth. Deeper down, the values west of the Midatlantic Ridge remain similar, whereas those east of the ridge decrease further, to about -110‰. It is shown that bomb-14C is prominent down to about 1500 m depth. Beyond this depth the bomb 14C component is small and is negligible in the eastern basin below 2800 m. On the basis of the 14C-tritium correlation, the distribution of natural 14C below about 1500 m depth is derived from the observations. In the deep and bottom water east of the ridge the 14C-salinity relationship seemingly is non-linear. Contrary to expectation, the 14C concentration in the bottom water is not lower than found on an US GEOSECS station near 10° N. Apparently, lateral concentration differences in the Northeast Atlantic bottom water as well as nonlinearity of the 14C-salinity relationship at 40°N do not exceed 10‰ in Δ 14C.

Description: The grazing of different copepodite stages of Calanus finmarchicus on natural phytoplankton populations was measured at different times of day and night during the Fladen Ground Experiment (FLEX), from the end of May to the beginning of June 1976, at the central position of the "FLEX box" (58° 55' 00" N, 0° 32' 05" E), on board of the R.V. "Meteor". A big difference between nocturnal and diurnal food intake was found, particularly for the stages V & IV, which at night ingest 4 to 10 times more than during daytime. The maximal daily food intake (24 hours), taking into account the different feeding rates at the different times of the day and expressed as percent of own carbon weight, was found to be 31 %, 83 %, 115 % and 148 % for the copepodite stages V, IV, III, II and I together, respectively, with an initial feeding threshold at about 50-60 µg C · l-1. The total grazing for the upper 40 m depth layer was calculated for each sampling time; a very pronounced grazing rhythm was found, due to the higher ingestion rates at night combined with the vertical migrations. Maximal night ingestions were 100-200 mg C/m2/hour at the end of May to 300-400 mg C/m2/hour at the beginning of June, compared with 1-20 mg C/m2/hour at midday. Making a balance of the positive influence of particulate primary production and the negative influence of grazing on the stock of particulate carbon, we could show that the main cause of the decline of the phytoplankton bloom in this period was Calanus grazing. Other zooplankton activities, namely respiration, production and faecal pellets excretion as measured by four other authors are in balance with our results of ingestion.

Description: "Auftrieb 75" was a joint cruise of R.V. "Meteor" and R.R.S. "Discovery" performed in the area 21-26° N off the northwest African coast during January-March, 1975. Current meter observations from six moorings and CTD/STD observations from three repeatedly covered sections of the cruise are analysed in order to describe the distribution of the water masses and its variability with the changes in the current system. The Canary Current is seen to pass through the northwestern corner of the expedition area, carrying with it a geostrophic eddy with a period in excess of 30 days. The eddy is probably another example of the type generally found in the open ocean circulation. Further south, offshore currents display large variations in strength and direction with a typical time scale of 10-14 days which are believed to reflect largescale mixing processes along the boundary of the Canary Current and the poleward flow of the coastal region, involving North Atlantic Central Water (NACW) and South Atlantic Central Water (SACW). The coastal upwelling undercurrent has been observed along the shelf edge but undergoes large variations in response to the wind and to variations in the horizontal divergence of the offshore currents. On the shelf the circulation responds to the local longshore wind to some extent but is also strongly affected by variations of the currents at the shelf edge. The distribution of water masses at intermediate depth ( 100-800 m) is analysed quantitatively based on the assumption that mixing between NACW and SACW is effective along surfaces of constant density only. lt is shown that most intense mixing occurs at about 22-23° N and is still very important further south. At 25-26° N SACW has been absorbed into the surrounding NACW water mass to a high degree, even in the undercurrent. Isolated bodies of water are found in the southern part of the expedition area, some of which are carried north in the undercurrent. The large variability in the water mass distribution at the source level of the upwelling probably has important consequences for the variations in the efficiency of biological production of the upwelling region.

Description: The main objectives of the Fladen Ground Experiment (FLEX) 1976 were an investigation of the dynamics of the mixed layer and the development of the spring plankton bloom. A quantitative consideration of individual chemical parameters (nutrients, particulate nitrogen and phosphorus) showed it to be advantageous to divide the water column into several layers which are separated from one another by measured temperature gradients. Measurements of particulate matter (phosphorus and nitrogen) revealed two plankton blooms at the central station. The first one (21.4.-14.5.76) was much more extensive than the second one, since the limiting thermocline was at a depth of about 60 m. Silicate was consumed most rapidly during this bloom and almost completely exhausted. The second bloom (19.-30.5.76) produced concentrations of particulate matter which were almost as high as the first one but were limited in only a few meters' water depth by a secondary thermocline, which prevented an influx of nutrients. Nitrate was completely exhausted during the second bloom. A high degree of nutrient depletion in the mixed layer from 24.4.-29.4. and from 22.5.-27.5.76 make these periods appear well-suited for determination of uptake rates. These periods are characterized. by a parallel course of concentration curves in the upper water layers for all nutrients except ammonium, and by the absence of !arge short-term fluctuations. Thus, disturbances due to hydrodynamic effects were minimal. The effects of tidal changes were minimized by using daily averages of the various concentrations. However, from 9.5.-13.5.7 6 in the upper layer a relatively strong and rapid increase was observed in the concentrations of all nutrients, which can probably only be explained by hydrodynamic influences. The nitrogen and phosphorus budgets can only be approximately balanced without considering dissolved organic matter. The strong decrease in dissolved inorganic nutrients during the first plankton bloom is almost comple,tely compensated by the increase in particulate matter. Not considering hydrodynamic influences, deficits after the plankton bloom could be explained by the fact that dissolved organic substances, as weil as sedimentaty matter and zooplankton, either were not considered at all, or at least not quantitatively. The time periods from 2.-7.4. and from 21.4.-14.5.76 show no strong short-term fluctuations in the nitrogen and phosphorus budgets and appear most suitable for calculations of mass fluxes, uptake and production rates, since biological-chemical processes seem to be dominant over hydrodynamic ones during these periods.

Description: The monthly mean meteorologically induced circulation of the North Sea for the period 15 March to 15 April 1976 (JONSDAP '76 INOUT period) is computed using two numerical models. One has a fine grid resolution, is two-dimensional, comprehensively non-linear and covers the North Sea. The other has a coarser grid resolution, is three-dimensional, and linear except for the inclusion of quadratic bottom friction. The depth variation of the two horizontal components of current in this model is accomplished using an expansion of time and horizontally dependent coefficients and depth dependent functions. The three-dimensional modelling technique presented here is sufficiently flexible to allow an arbitraty space and time variation of vertical eddy viscosity, although still retaining a continuous representation of current in the vertical, and allowing for variable bottom topography. Preliminary results presented here for the meteorologically induced circulation of the North Sea during JONSDAP '76 agree weil with present knowledge of the circulation of the North Sea. Comparing surface and bottom currents computed with the 3 D shelf model, it is evident that the windinduced flow varies considerably in both magnitude and direction between sea surface and sea bottom. The distribution of depth mean current computed with the 3 D shelf model over the North Sea is nearly identical to that computed with the 2 D North Sea model. This is particularly encouraging since the 3 D shelf model does not contain the advective terms, which are present in the 2 D model, and has a coarser resolution.

Description: The ecology of the lower marine fungi, namely the thraustochytrids, in the Fladen Ground area (FLEX box) and other parts of the North Sea was studied during 5 cruises in 1975 and 1976. The number of fungi/liter and the number of species showed seasonal fluctuations in the surface water samples from all the stations. A high number was found in September 1976 and a low number in March 1976. These numbers, however, revealed no seasonal fluctuations in the underlying sediments. In both the surface waters and the sediments, a consistently low number of fungi was recorded for certain stations and a high number of fungi for others, the reason for this being unknown. The sediments revealed a very high number of fungi/liter. Observations on the distribution of various species are presented. Certain species occurred more frequently at some stations than at others; certain species occurred more in the water column, e. g. Ulkenia minuta, and still others in the sediments, e. g. Thraustochytrium multirudimentale.

Description: In the framework of the Fladen Ground Experiment 1976, zooplankton samples were taken during the phytoplankton spring bloom in April-May and after the bloom in June. The plankton was collected with two samplers, a towed multiple high-speed sampler and a newly developed swing net for local sampling. The results of both methods are compared. The mean number and dry weight per m3 are given for the common species, as weil as the density of fecal pellets. The calanoid copepod Calanus finmarchicus formed 63 %, resp. 90 % of the biomass. The population composition, distribution, dry weight/length relationship and productivity of this species is studied in more detail. The high abundance of fecal pellets, eggs and young stages during the spring bloom indicate a high zooplankton activity in this period. The low density of these indicators and the apparent descent of the older Calanus stages to deep water layers in June signify the dependency of the zooplankton activity on the phytoplankton density. The estimated daily production of Calanus during the phytoplankton bloom was 20 mgC/m2. The significance of the seasonal vertical migration and the role of zooplankton in the northern North Sea ecosystem is discussed. lt is concluded that the Zooplankton could have checked the low algal density in May-June after the spring bloom, but it is far from certain that it stopped the phytoplankton outburst at the end of April and early May.

Description: During the culmination of the phytoplankton spring bloom in the Fladen Ground area in April-Mai 1976, gross primary production was between 1500 and 2000 mg particulate C m-2 day-1, at a crop density (mainly diatoms of the genus Chaetoceros) of about 1500-3500 mg C m-2. Estimates of the C : chlorophyll a ratio in living cells were much lower than those reported in the literature, possibly because part of what is measured as "chlorophyll a" by the common fluorometric method is associated with particles that are not reported as cells. Most of the dark 14C fixation during the bloom's climax was due to abiotic processes. Excretion of 14C-labeled carbohydrates did not account for a significant fraction of the total photosynthetic rate. The low crop after the bloom period, in June, corresponded with nutrient depletion of the euphotic zone. The low photosynthetic efficiency in June may have been a gross underestimate. The presence of relatively high concentrations of chlorophyll derivatives signifies that the algal crop was consumed by heterotrophs, but at a lower rate in April/May than during the June cruise when particularly high molar ratios of phaeophorbide a and phaeophytin a relative to chlorophyll a were found. The high respiratory rate relative to autotrophic production in June manifested itself also in high dark 14C fixation values. The high concentration of phaeophorbide a in the upper 40 m and its scarcity below this depth during the spring bloom climax in April/May implies that copepod grazing at that time took place principally in the euphotic zone. The remarkably high concentration of chlorophyllide a in the surface layer during the bloom period indicates that the part of the crop that was destroyed by the grazers while eating was occasionally as high as the part that was actually ingested.

Description: During the Fladen Ground Experiment (FLEX, March 26-June 6, 1976) quantitative investigations of herbivorous zooplankton were performed. For that purpose 1785 water samples (5 or 10 l) were taken with a rosette sampler at the central station (pos. 58° 55' N, 0° 32' E) from nine to ten standard depths and filtered through 30 µm gauze filters. The development of the most important Zooplankton groups at the central Station is presented in depth-integrated time series (0-100 m) of individuals per m2. The number of individuals of Calanus finmarchicus was converted to biomass (dry weight). At the end of the sampling at the central station the standing stock of Calanus finmarchicus showed about 18 g dry weight per m2. Numerical values for the time lags between different plankton groups are calculated with the aid of the cross-correlation function. The time lag between the diatom ad copepod egg peaks is only up to 2.75 days; that between diatoms and the larvae of bottom invertegrates is 5 days. Between the copepod eggs and the maximum number of nauplii (N I-N VI) there is a time lag of 10.75 days. Another 10.25 days lie between the maximum numbers of nauplii and copepodites (C I-C VI). Considering the depth-integrated time series of the abundance of Calanus finmarchicus copepodites, a succession of the Stages seems to exist. Nevertheless, numerical values for the time lags between the different developmental stages cannot be calculated unambiguously by the cross-correlation technique. This is probably caused by extreme simultaneous fluctuations, especially of the numbers of all stages of Calanus finmarchicus in the last phase of the sampling at the central station.

Description: The third phase of the cooperative data collection program JONSDAP begun in 1970 by the countries bordering on the North Sea was planned to include for the first time biological and chemical measurements in 1976, as well as physical ones. Intensive measurement programs were performed, one concentrating on the currents flowing into and out of the open boarders of the North Sea (INOUT) for 40 days in March/ April and one in which the dependence of the spring plankton bloom on the vertical temperature structure was studied in the Fladen Ground area for 100 days from March to June (FLEX). Around 90 % of the planned program with moored systems and 75 % of the measurements from moving ships were able to be carried out successfully. The large quantities of data required the establishment of two data centers. The current data were collected in Liege, all data from the FLEX box in Hamburg. A cooperative JONSDAP format was agreed on for the exchange of data. Graphic presentations of the data have been compiled in the two-volume Draft FLEX/IN OUT Atlas and distributed to all JONSDAP '76 participants, one of the main purposes of which being the facilitation of interdisciplinary evaluations, now that the intradisciplinary data processing has almost been completed - as presented in this volume.

Description: Measurements of nutrients and the elements phosphorus and nitrogen from the central station in FLEX '76 were used to calculate mass fluxes and budgets for these elements in defined time and depth intervals. The intervals were defined using hydrodynamic and biological criteria. Within these periods rates of increase and decrease were calculated for nitrate, ammonium, phosphate, particulate nitrogen and phosphorus. These rates were also calculated for the layer above and below the main thermocline. The element budget was balanced within some of the defined intervals. This was possible by only considering biological, chemical, and vertical physical interactions at the central station. At the beginning of the phytoplankton bloom we found fluxes with a maximum of 28 ngat · l-1 · d-1 for phosphorus and 590 ngat · l-1 · d-1 for nitrogen. Interaction diagrams are shown for mass fluxes for three periods during the plankton bloom.

Description: Since both German research vessels „Meteor" played a major role in G. Dietrich's work, it seemed appropriate to present his bibliography in this series. An assessment of his scientific achievements was given at an earlier date (H. U. ROLL 1973: In memoriam GüNTER DIETRICH 1911-1972. - ,,Meteor" Forsch.­ Ergebn. A, No. 12: V-X).

Description: Vertical velocity spectra can be determined by three different indirect methods: Following isopycnal surface displacements, following isothermal surface displacements, or calculating water particle displacements from temperature time series at fixed pressure levels, with the mean vertical temperature gradient used for the conversion factor. This third method is the easiest to perform. Data from GATE 1974, obtained by CTD profiling onboard R. V. "Meteor" and by temperature measurements on moorings, are used to check whether the three methods are equivalent. lt is shown that autospectra do not differ significantly within the 95 % confidence intervals, and either method can be applied. This does not infer an equivalence with respect to the actual data points since individual events were found in the low-passed time series with deviations occurring between displacement curves obtained by the different methods. These events were apparently due to the advection of another water mass.

Description: The research programmes of the German groups in JASIN 1978 are summarized, comprising projects that concern the planetary boundary layer, the radiation budget, the oceanic mixed layer and thermocline and the trace element exchange. Lists of the participating scientists and technicians are included. Brief descriptions of the field activities on board the three ships and the aircraft are given, including maps and stations lists.

Description: A first survey of the entire temperature data base from the central station is presented. The development and the changes of the sea surface temperature, the thermocline, and the heat content due to meteorological forces, are described.

Description: A description is given of part of the data on residual currents which were obtained during the INOUT phase of JONSDAP '76. This current measuring programme was the result of a concerted effort of North Sea bordering countries and was aimed at a better understanding of residual water movements in the North Sea. One of the first results of INOUT is the possibility to compare estimates of simultaneously occurrmg inflow/outflow through several North Sea boundary sections. It appears that major inflow/outfow occurs in the neighbourhood of the Norwegian Trench. Estimates for other boundaries are significantly lower. Matching the objectives of INOUT several charts are presented showing measured residual water movements under influence of some "persistent" winds that is to say, during situations with spatial homogeneous windfields not changing too rapidly in time. Under such conditions it is possible to identify residual current systems. However, as can be expected, in cases of spatially inhomogeneous winds and/or winds changing rapidly in time it is harder to interpret the current measurements. In those cases it is necessary to use mathematical North Sea models with the proper wind field input. The Royal Netherlands Meteorological Institute (KNMI) took care that these windfields were stored on magnetic tape during INOUT for this purpose.

Description: Recordings were made of the voltage differences between the ends of two submarine cables crossing the Pentland Firth; the recordings covered the period from February to October 1976 and thus included the period of the JONSDAP '76 oceanographic experiment in the North Sea. The noise level on these recordings was found to be greater than for similar recordings on cables crossing the Dover Strait and the North Channel of the Irish Sea; reasons for this increased noise level are discussed. Tidal analyses of the recordings were made for six discrete sets of recordings, each of 29 days duration; these analyses produced consistent results for the five major semi-diurnal constituents, namely N2, V2, M2, S2 and K2. However, an examination of the propagation of the M2 tidal constituent in this region showed that the phase of M2, as calculated from the cable recordings, did not obviously aeeord with the ove1·all pattern of propagation of that constituent. Tentatively it was concluded that cable measurements represent the integrated effect of flow over an area surrounding the Pentland Firth where this flow varies widely in both phase and direction. A calibration coefficient for the cable, relating voltage to flow, was calculated by equating the annual variation in the amplitude of the M2 constituent (measured by the cable) with the annual variation in the conduetivity of sea water in the region. The residual component of cable voltage was compared with residual components of (a) the sea surface elevation at Wick and (b) currents derived from current meter measurements at a nearby location in the North Sea; also with wind recordings at Wick. These comparisons indicated that there was neither a significant wind-driven flow through the Pentland Firth, nor a seasonally varying residual flow. It is suggested that a significant component of both the residual flow (as measured by the cables and by the current meters) and the residual elevations at Wick is associated with the propagation of "external surges" into this region.

Description: A simple model is used to simulate the dynamics of phytoplankton and phosphate from April 19 to May 16 during the Fladen Ground Experiment. The model describes the development of vertical profiles in time. lt allows changes of phytoplankton standing stock by turbulent diffusion, sinking of cells, primary production, respiration, mortality and grazing, and changes of phosphate concentration by turbulent diffusion, nutrient uptake and remineralization. The model is used as a means of checking the consistency of the biological assumptions made with the observed phosphate concentration, phytoplankton cell counts and primary production. Local turbulent diffusion coefficients, underwater light climate and Zooplankton standing stock in terms of total copepods are given quantities, the former from a physical upper layer model, the latter two from data. Simulations show that if phytoplankton standing stock and primary production are reproduced fairly well, then phosphate is still too high and vice versa. lt appears that the conversion factor expressing the amount of phosphate necessary to produce l g carbon is crucial. Relating this ratio to the observed decrease of phosphate and the primary production the simulation reproduces the exponential growth phase weil but fails for the rest. This indicates the need of a variable conversion factor in the model, depending on the status of the bloom. lt can be shown that turbulent diffusion is indispensable for the development of the spring bloom. Further it is shown that as a consequence of the assumptions made about two thirds of primary production are taken up by zooplankton. An estimate of potential grazing of copepods shows that copepods starve or have other food sources than living phytoplankton.

Description: The figures contain section data of S, t, total P, N03, chlorophyll a, total particulate P and total phytoplankton. The data indicate that primary production had started only along Norway, probably due to the high stability caused by the presence of water of Baltic origin. Three runs were made; differences between them are discussed in relation to results of current measurements.

Description: From 8 April to 4 May 1976 chemical changes in the marine environment and primary production were studied in the area of the northern North Sea (Fladen Ground). During most of the observation time, when the ship was keeping close to the central position, the influence of patchiness and advection by currents was relatively small. Under such favourable conditions it was possible to follow the temporal changes and processes in the water during the first phase of the spring phytoplankton bloom. After 19 April a distinct thermocline was established. At the same time the nutrient and C02 concentrations in the surface water began to decrease, due to the uptake by the growing phytoplankton. Before the start of the phytoplankton bloom the ratios (by atoms) were N : Si : P = 14 : 6.5 : 1.0. The ratios of nutrient and C02 uptake into the phytoplankton were ΔC : ΔN : ΔSi : ΔP = 120 : 19 : 9 : 1.0. During periods with high irradiance a remarkable diurnal rhythm of temperature and 02 concentration in the surface water could be observed. From the pH, temperature, salinity und alkalinity data the mean net primary production during the first 12 days of the phytoplankton bloom ( 19 April to 1 May) was calculated. lt was about 2.0 ± 0.4 gC · m-2 • d-1. The same result was obtained from the phosphate data.

Description: From 27 Janua1y to 23 June 1979 R. V. "Meteor" surveyed the central equatorial Atlantic on a section along 22° W from 3° N to 2° S. During the observation period, a hydrographic section down to 600 m was repeated ten times with a continuous "Howaldt-Bathysonde" CTD and a rosette sampler. The station distance was 10 to 15 nm. The water samples were used to recalibrate salinity and to determine oxygen, nutrients and chlorophyll a. An undulating CTD system ("Delphin") was towed on 11 sections. A profiling distance of one to two nautical miles and a profile depth of 90 m was obtained. Five current meter arrays were moored along 22° W between 3° N and 1° S from January to March 1979. In May and June two moorings were installed at 2° N and at the equator. On the buoys measurements of wind speed and direction were obtained. At 43 stations a wire-guided Aanderaa profiling current meter was successfully lowered. Drifting buoy experiments were repeated three times with clusters of 5 to 10 buoys. A fourth experiment took place in 1978 in the Gulf of Guinea. On the way from and to port XBT sections were carried out. The data sets obtained by these instruments are presented in this data report.

Description: The benthic foraminiferal populations along three traverses across the Northwest African continental margin were analyzed on the base of ea. 60 surface sediment samples. Depth ranges of 213 species were established and the main trends of vertical distribution are compared with those known from adjacent regions. Main faunal breaks occur at 100/200 m and 1OOO/ 1500 m depth of water. Some species show latitudinal distribution boundaries and the same applies to population density (standing stock), reflecting the regional distribution of nutrients supply by river discharge and upwelling processes. - High proportions of Bolivina tests at the lower slope indicate extended downslope transport.

Description: Surface sediments from the continental slope and rise off North-West Africa between the Canary Islands and the Cape Verde Islands are mainly composed of silt-sized material (2-63 µm). A number of sampling profiles were run normal to the coast and the composition of the silt fraction was determined quantitatively by scanning electron microscope analysis. The carbonate portion of the sediment was found to be nearly exclusively of biogenic origin. The most important contributors are planktonic foraminifers and coccoliths with minor contributions derived from pteropods. Plankton-produced biogenic opal such as diatoms and radiolarians play a very minor role. The high production rates of opal-silica plankton which exists in the surface waters of the NW-African upwelling system does not give rise to corresponding increases of opal accumulation in the bottom sediment. Benthic producers consist mainly of foraminifers and molluscs but the entire input from benthic producers is extremely small. An exception to this occurs in the prodelta sediments of the Senegal river. Downslope particle transport is indicated by the occurrence of shallow-water coralline algae, ascidian sclerites and cliona boring chips and can be traced as far down as the continental rise. The non-carbonate silt fraction mostly consists of quartz which is derived as eolian dust from the Sahara desert by the Harmattan and the NE-Trade-wind system. The percentage of carbonate in the surface sediments directly indicates the relative proportions of autochthonous biogenic components and terrigenous allochthonous quartz particles.

Description: Water depth zonation of fifty nine benthonic foraminiferal species in marine sediment surfaces has been described. Some species are combined to groups which mark particular depth zones: an upper and lower shelf-fauna, an upper and lower slope fauna, and a shelf-slope fauna. Dependence on latitude could be ascertained for Textularia panamensis) and submergence effects for Hyalinea balthica.

Description: During cruise VA 18 of R.V. "Valdivia" an area of 15 X 18 km, lying on a horst within the Clarion fracture zone south-southeast of Hawaii at 14° N and 153° W was studied in detail by reflection seismic measurements and geological sampling. The horst lies between two fracture-zone-parallel grabens. Its top displays a typical abyssal hill topography which is a result of block faulting within the oceanic crust, with faults parallel to the fracture zone and in north-southerly directions. It is assumed that the horst represents an already faulted block of oceanic crust which was included into the fracture zone when it tectonically developed into a system of horsts and grabens upon its transform stage. The surveyed horst differs from its neighbouring grabens by reduced sediment-accumulation rates and hiatuses. Its basement relief has led to a further differentiation of the internal sedimentary pattern. The northward-tilted horse block shows on its southern, more elevated part, frequently signs of erosion or reduced sedimentation, whereas the low lying part is characterized by an increased sediment accumulation. In the area of low accumulation rates and erosion volcanic ash layers with manganese-oxide crusts occur at or near the surface of the sediment layer. Also manganese nodules occur in this area. In the area of high accumulation rates the volcanic ash layers were observed with increased thicknesses in deeper sections of the cores. Crusts and nodules were not found here. This is also the case for the graben south of the horst, where high accumulation rates prevented the formation of manganese crusts and nodules. The reduced accumulation in the southern part of the horst has favored the dissolution of radiolarians as well as the authigenic formation of zeolites and manganese micronodules. The correlation between the reflectors seen in the 3.5 kHz subbottom profiler records and the lithostratigraphic boundaries found via geological sampling was generally good. It allowed the determination of the areal distribution of sedimentary sequences and magmatic rock outcrops.

Description: Along the Sierra Leone continental shelf edge, bedded rocks, presumably mostly calcareous sandstone, calcarenites and siltstones form exposures up to 10 km long in water depths ranging from 80 to 110 m. These outcrops are found in a 6 km broad zone along the shelf edge; landward they get covered by recent shelf sands. Their apparent strike direction parallds the shelf edge indicating nearly horizontal bedding which locally is interrupted by faulting. On the middle shelf, a terrace is well developed at a water depth of 56 m. Ripples, megaripples, pockmarks 1-5 m in diameter, and canyon-heads form other significant features of the shelf.

Description: The sediments of a 10.67 m long piston core recovered from the Sierra Leone Rise ("Meteor" Core 13519) have been analysed for their paleomagnetic properties. Using detailed demagnetization techniques, the geomagnetic Brunhes/Matuyama boundary could be identified between 9.81 and 9.89 m sub-bottom. This is in excellent agreement with the oxygen isotope stratigraphy by SARNTHEIN et al., this volume. Down to this level, therefore, the average apparent accumulation rate amounts to 13.5 m/m.y. assuming an age of 0.73 m.y. for the reversal boundary (BERGGREN et al. 1983). A second transition in magnetization polarity was recognized between 10.51 and 10.54 m sub-bottom. According to its magnetic signature it was tentatively correlated to the beginning of the Jaramillo event (0.98 m.y.). This would imply that in the lowermost part of the core the apparent accumulation rate decreases to less than 20% of that found in the upper sequences. However, as obviously only a small portion of the Jaramillo event is recorded in the sediment column, a more plausible solution is to keep the overall sedimentation rate about constant by introducing a hiatus of some 2·105 years shortly after the lower Jaramillo/Matuyama reversal boundary. Alternative interpretations are also discussed.

Description: From a 10.7 m long gravity core from the Sierra Leone Rise (5° 39,5' N, 19° 51'W) a detailed oxygen and carbon isotope record of both planktonic and benthonic foraminifera species was obtained extending from the Recent to the Jaramillo event. The analysis yielded six major results. 1. Benthos oxygen isotopes varied by 1.8-2.2‰ from interglacial to glacial times and may indicate a synglacial cooling of North Atlantic Deep Water at 2800 m depth by 1-3° C. 2. Variable anomalies between the benthos and plankton δ18 O record indicate a cooling of sea-surface temperatures (SST) by up to 6° C during some glacial stages. 3. Southerly trade winds and equatorial upwelling may exert the primary control of SST variations, in particular of extreme values of cold and warm stages and of the abrupt character of climate transitions and their leads and lags, and finally, of variable sedimentation rates. 4. The benthos δ13C record correlates well with the flux and preservation of organic matter. 5. A new time scale, CARPOR, was established from the assumption that terrigenous sediment supply was ± constant but CaC03 varied considerably. When applied to the δ18O record, three major and numerous short-term variations of sedimentation rates (0.8 to 4.0 cm/kyr) can be distinguished. 6. The climatic record was modified by bioturbation much more strongly during cold than during warm stages.

Description: Planktonic foraminiferal tests of the spinose species Orbulina universa, of the non-spinose Globorotalia tumida-menardii complex, and of a mixed species assemblage (grain size fraction 200-400 μm) were isolated from Sierra Leone Rise core 13 519-2 and analyzed for free, total, and bound (by difference) amino acids to study the isoleucine epimerization mechanism in fossil foraminiferal tests and to define empirical calibration curves for dating deep-sea sediments over the past 900,000 years. Total isoleucine epimerization curves typically separate into three "linear" segments of decreasing apparent rates with increasing time and exhibit a pronounced "species effect". The degree of epimerization attained at time is considerably lower in 0. universa than in G. tumida-menardii while the mixed species results scatter between the limits delineated by the two monospecific curves. Total allo/iso ratios are closely related to the proportion of free to total isoleucine accumulating in the tests indicating that the rate of hydrolysis of matrix proteins and peptides controls the overall epimerization reaction. The results are consistent with experimental evidence (KRIAUSAKUL & M1TTERER 1978, 1980 a, b) whereupon isoleucine epimerizes at a rapid rate in terminal positions but at slow rates in interior positions as well as in the free state. Notwithstanding free isoleucine exhibits the highest degree of epimerization due to preferential hydrolysis of extensively epimerized terminal isoleucine. Species-specific hydrolysis and epimerization rates are maintained until about 50% of bound isoleucine have been hydrolyzed to the free state corresponding to a total allo/iso ratio of about 0.5. Remaining peptide units appear to be more resistent against hydrolysis and separate species then show the same apparent epimerization rate dominantly controlled by the slow conversion rate in the free state until equilibrium is achieved in Miocene samples under deep-ocean temperature conditions (KING & HARE 1972 a). The degree of epimerization attained at comparable time in separate species will, however, remain different due to different initial rates of hydrolysis. Selective leaching of free isoleucine from the tests results in a lowering of total allo/iso ratios and apparent rate constants when significant amounts have been generated by diagenetic hydrolysis. Comparison with results based on Pacific core V 28-238 (KING & NEVILLE 1977) indicates an inverse relationship between the intensity of surficial carbonate dissolution and post-depositional leaching. Tests deposited under less corrosive conditions (Atlantic core 13519-2) will more significantly deviate from a closed system during diagenesis than those deposited in more corrosive bottom waters (Pacific core V 28-238). Carbonate dissolution removes proteinaceous lamellae from more external shell structures, which otherwise - if preserved during deposition - will preferentially be subject to leaching of free amino acids during diagenesis. This effect may qualify the applicability of empirical epimerization calibration curves to different depositional environments although the general agreement of the epimerization curves obtained for both sites is promising for future research. Identical apparent epimerization rates in G. tumidamenardii tests younger than 120,000 years (when leaching is stiH insignificant) at both Atlantic and Pacific sites are not consistent with the presently different bottom water temperatures in these regions. It is concluded that glacial North Atlantic Deep Water was cooler than at the present at the Sierra Leone Rise.