ALBERT

Beschreibung: We investigated yellow birch ( Betula alleghaniensis Britt.) growth patterns and disturbance frequency before and after the advent of selection harvesting at the Ford Forestry Center in the Upper Peninsula of Michigan, USA, through the use of tree-ring analysis. Based on the boundary-line release detection procedure, 88% of the trees in our sample (n = 67) displayed evidence of at least one moderate or major release. Prior to active forest management, releases were infrequent, and trees that originated during that period had growth histories consistent with establishment after large-scale disturbances (i.e., large canopy gaps, 〉200 m2). Conversely, tree cohorts that recruited to the canopy more recently displayed a growth pattern suggestive of periodic small gap expansion. Given the declining representation of yellow birch in these forests, the latter strategy, although probably sufficient to prevent extirpation, is unlikely to ensure a sustainable and harvestable population of this and other midtolerants in managed uneven-aged forests. Our results highlight the importance of considering the cumulative influence of infrequent disturbances and chance events on the maintenance of tree species diversity.

Beschreibung: Locating fuel treatments with scarce resources is an important consideration in landscape-level fuel management. This paper developed a mixed integer programming (MIP) model for allocating fuel treatments across a landscape based on spatial information for fire ignition risk, conditional probabilities of fire spread between raster cells, fire intensity levels, and values at risk. The fire ignition risk in each raster cell is defined as the probability of fire burning a cell because of the ignition within that cell. The conditional probability that fire would spread between adjacent cells A and B is defined as the probability of a fire spreading into cell B after burning in cell A. This model locates fuel treatments by using a fire risk distribution map calculated through fire simulation models. Fire risk is assumed to accumulate across a landscape following major wind directions and the MIP model locates fuel treatments to efficiently break this pattern of fire risk accumulation. Fuel treatment resources are scarce and such scarcity is introduced through a budget constraint. A test case is designed based on a portion of the landscape (15 552 ha) within the Southern Sierra fire planning unit to demonstrate the data requirements, solution process, and model results. Fuel treatment schedules, based upon single and dual wind directions, are compared.

Beschreibung: In this paper, we consider a tactical transportation planning problem in forestry to decide the destination of logs. This problem is generally solved by finding the flow between a set of supply points and demand points. It can be formulated as a linear programming problem involving direct flows between supply and demand points. However, better solutions can be found by using additional flow variables representing flow in potential backhaul routes. However, the number of such variables is often very large. In this article, we provide the basis for backhaul flow planning in forestry. This includes defining the underlying operations research models for both the flow problem and the subproblem to find backhaul routes. The size of the problem in terms of the number of variables increases rapidly with the number of supplies and demands and we describe a column generation approach for its solution. We report on some case studies and industrial systems where the approach has been used.

Beschreibung: This paper examines the problem of harvest capacity planning at a tactical level. Annual capacity planning allows planners to determine the number of contractors to hire per period throughout the year and to define the duration of their contracts. In practice, this process usually involves the analysis of historical data regarding the operational use of capacity and aggregated demand forecast, the output of which then serves to plan harvest operations. Although this form of hierarchical planning reduces the complexity of the task, the decomposition into subproblems that must be successively resolved can lead to infeasibility or poor use of harvesting capacity. The specific problem addressed here resides in how one can consider the operational impact of harvesting decisions taken at the tactical level to ensure a plan’s feasibility at the operational level. We present a tactical planning process based on Schneeweiss’ generic hierarchical modeling approach. A computational experiment demonstrates how a tactical planning process is influenced by the input of the operational level anticipation model. The anticipation approach we propose appears to be a valid method to better integrate key operational-level decisions into tactical plans.

Beschreibung: Spatial tree data are required for the development of spatially explicit models and for the estimation of summary statistics such as Ripley’s K function. Such data are rare and expensive to gather. This paper presents an efficient method of synthesizing spatial tree point patterns from nearest neighbour summary statistics (NNSS) sampled in small circular subwindows, which uses a stochastic optimization technique based on simulated annealing and conditional simulation. This nonparametric method was tested by comparing tree point patterns, reconstructed from sample data, with the original woodland patterns of three structurally different tree populations. Analysis and validation show that complex spatial woodland structures, including long-range tree interactions, can be successfully reconstructed from NNSS despite the limited range of the subwindows and statistics. The influence of the NNSS varies depending on the woodland under study. In some cases, the sampling results can be improved by reconstruction. Furthermore, it is clearly shown that it is possible to estimate second-order characteristics such as Ripley’s K function from small circular subwindows through the reconstruction technique. The results offer new opportunities for adding value to woodland surveys by making raw data available for further work such as growth projections, visualization, and modelling.

Beschreibung: The relative importance of fire and flooding on the population dynamics of eastern white-cedar ( Thuja occidentalis L.) and black ash ( Fraxinus nigra Marsh.) was evaluated in eight old-growth riparian stands of southwestern boreal Quebec, Canada. Rising water levels and decreasing fire frequency since the end of the Little Ice Age (ca. 1850) were expected to have favoured an inland migration of the riparian forest fringe, with the flood-tolerant black ash colonizing the lower parts of the shore terraces and eastern white-cedar the upper parts. Black ash was found to be restricted to the riparian zone (

Beschreibung: A modified logistic function was used for modeling specific-gravity profiles obtained from X-ray densitometry analysis in 675 loblolly pine ( Pinus taeda L.) trees in four regeneration trials. Trees were 21 or 22 years old at the time of the study. The function was used for demarcating corewood, transitional, and outerwood zones. Site and silvicultural effects were incorporated into the model. Heteroscedasticity and within-group correlation were accounted for by specifying the variance and serial-correlation structure, respectively. The estimated transition zone was located between rings 5 and 15, and the outerwood demarcation point varied from rings 12 to 15. No effects of treatments on the demarcation points were observed; however, site preparation and fertilization affected the lower asymptotes of the curves in all sites. A geographical trend for the demarcation point was observed, with the northern site requiring more time to reach a plateau in specific gravity compared with the southern sites. The diameter of the juvenile core was increased as a result of the treatments. However, the amount of corewood was not statistically affected, ranging from 55% in the north to 75% in the south, except at one site where fertilization decreased the percentage of corewood.

Beschreibung: A forest plot with a clustered spatial pattern of tree locations was used to investigate the impacts of different kernel functions (fixed vs. adaptive) and different sizes of bandwidth on model fitting, model performance, and spatial characteristics of the geographically weighted regression (GWR) coefficient estimates and model residuals. Our results indicated that (i) the GWR models with smaller bandwidths fit the data better, yielded smaller model residuals across tree sizes, significantly reduced spatial autocorrelation and heterogeneity for model residuals, and generated better spatial patterns for model residuals; however, smaller bandwidth sizes produced a high level of coefficient variability; (ii) the GWR models based on the fixed spatial kernel function produced smoother spatial distributions for the model coefficients than those based on the adaptive kernel function; and (iii) the GWR cross-validation or Akaike’s information criterion (AIC) optimization process may not produce an “optimal” bandwidth for model fitting and performance. It was evident that the selection of spatial kernel function and bandwidth has a strong impact on the descriptive and predictive power of GWR models.

Beschreibung: Using a mail questionnaire targeted at 500 softwood sawmills in the United States and Canada, firm innovativeness was assessed using three methods: (1) current technology, (2) self-evaluation, and (3) a new scale — the propensity to create and adopt scale. The results of these three methods were then compared to assess the performance of each method. Additionally, the relationship between firm innovativeness and financial performance was examined. Based on responses from 85 sawmills (19% adjusted response rate), the results show that both the self-evaluated and the propensity to create and adopt measures differentiate between mills with high and low levels of innovativeness. The composite of the propensity to create and adopt scale shows higher reliability (Chronbach’s α = 0.97) than the self-evaluated scale (Chronbach’s α = 0.68). Significant relationships between sawmill performance and each of the three measures of innovativeness were seen, with the propensity to create and adopt scale generally having the strongest positive relationships. Current technology was significantly related to sales growth, but not gross profit.

Beschreibung: The effects of 20th century spruce budworm ( Choristoneura fumiferana (Clem.)) outbreaks on forest dynamics was examined in the southern and northern parts of the mixedwood forest zone in central Quebec, Canada. In each region, three study areas were placed in unmanaged stands that had not burned for more than 200 years. Disturbance impacts and forest succession were evaluated using aerial photographs and dendrochronology. Spruce budworm outbreaks occurred around 1910, 1950, and 1980 in both regions. The 1910 outbreak seemed to have limited impact in both regions, and the 1950 outbreak caused heavy mortality in conifer stands (mostly of balsam fir, Abies balsamea (L.) Mill.) in the southern region. The 1980 outbreak caused major mortality in the northern region, but had little impact in the southern region. Successive spruce budworm outbreaks led to a massive invasion by hardwood species in the last century in the southern region but not in the northern region. The reason for such contrasting dynamics between regions is unknown, but we hypothesize that differences in disturbance intensities, influenced by climate, played a major role. Results from this study emphasize that generalizations about the effect of spruce budworm outbreaks on forest dynamics cannot be derived from observations made during a single outbreak or at a single location.

Beschreibung: Thinning and thinning followed by prescribed fire are common management practices intended to restore historic conditions in low-elevation ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) forests of the northern Rocky Mountains. While these treatments generally ameliorate the physiology and growth of residual trees, treatment-specific effects on reproductive output are not known. We examined reproductive output of second-growth ponderosa pine in western Montana 9 years after the application of four treatments: thinning, thinning followed by spring prescribed fire, thinning followed by fall prescribed fire, and unthinned control stands. Field and greenhouse observations indicated that reproductive traits vary depending on the specific management treatment. Cone production was significantly higher in trees from all actively managed stands relative to control trees. Trees subjected to prescribed fire produced cones with higher numbers of filled seeds than trees in unburned treatments. Seed mass, percentage germination, and seedling biomass were significantly lower for seeds from trees in spring burn treatments relative to all others and were generally higher in trees from fall burn treatments. We show for the first time that thinning and prescribed-burning treatments can influence reproductive output in ponderosa pine.

Beschreibung: We surveyed and wounded forest-grown sugar maple ( Acer saccharum Marsh.) trees in a long-term, replicated Ca manipulation study at the Hubbard Brook Experimental Forest in New Hampshire, USA. Plots received applications of Ca (to boost Ca availability above depleted ambient levels) or Al (to compete with Ca uptake and further reduce Ca availability). We found significantly greater total foliar and membrane-associated Ca in foliage of trees in plots fertilized with Ca when compared with trees from Al-addition and control plots (P = 0.005). Coinciding with foliar Ca differences, trees exhibited a significant difference in crown vigor and in percent branch dieback among treatments (P 〈 0.05), with a trend towards improved canopy health as Ca levels increased. Annual basal area increment growth for the years following treatment initiation (1998–2004) was significantly greater in trees subjected to Ca addition compared with trees in control and Al treatments. Treatment-related improvements in growth were particularly evident after overstory release following a 1998 ice storm. The amount of wound closure was also greatest for trees in Ca-addition plots relative to Al-addition and control plots (P = 0.041). These findings support evidence that ambient Ca depletion is an important limiting factor regarding sugar maple health and highlight the influence of Ca on wound closure and growth following release from competition.

Beschreibung: Airborne laser scanning (lidar) technology is increasingly being applied in forest ecosystem surveys. This research note proposes a design-based approach for the lidar-assisted estimation of forest standing volume when ground surveys are performed by means of fixed-area plots. The lidar measurement of the height of the upper canopy (digital crown model) is performed for the whole study area, and the resulting pixel heights are adopted as auxiliary information to couple with the standing volume acquired on the ground by means of sample plots. The ratio estimator for the total volume of the forest is derived in a complete design-based framework together with an unbiased estimator of its sampling variance and the corresponding confidence interval. The proposed procedure has been tested in Bosco della Fontana, a lowland forest in Northern Italy, obtaining a 95% confidence interval for the total volume, which is approximately 2/3 smaller than that obtained by solely using information arising from field plots.

Beschreibung: We assessed canopy openness (%) in an old-growth beech–maple forest immediately before and in the 3 years following a severe ice storm. We estimated canopy openness using hemispherical photographs taken at a height of 0.6 m above the soil surface in 101 permanent plots. Mean canopy openness increased from a prestorm value of 7.7% to 16.6% in the summer immediately following the storm. However, the mean canopy openness returned to prestorm levels within 3 years. The changes in canopy openness immediately after the storm were significantly influenced by canopy openness prior to the storm and also by species composition; plots with lower canopy openness prior to the storm and plots that consisted of more shade-tolerant species had greater canopy damage. While canopy gaps are often considered to promote the establishment of shade-intolerant species in the deciduous forests of eastern North America, gaps created by ice storms at our study site may not persist long enough to promote the establishment of these species.

Beschreibung: Plant polyphenolics are receiving increased attention for their influences on belowground processes. Tannins are of particular interest because of their predominance in natural systems, their wide variation in both quality and quantity, and their protein-binding abilities. Current theory holds that simple phenolics increase microbial activity by acting as carbon substrates, while larger tannins decrease microbial activity by binding with organic nitrogen such as proteins. Here, we present results from a simple microcosm experiment that demonstrates that the influence of condensed tannins on soil respiration depends on the availability of additional carbon substrates. We purified tannins from trembling aspen ( Populus tremuloides Michx.) and crossed three levels of tannin additions with three levels of cellulose additions in laboratory microcosms. Soil respiration was measured over 36 days. In the absence of cellulose, high amounts of condensed tannins increased cumulative soil respiration. In the presence of abundant cellulose, condensed tannins decreased cumulative soil respiration. The positive and negative effects of purified tannins on soil respiration are time dependent, such that initial respiration is likely tannin induced, while later respiration is cellulose induced and tannin limited.

Beschreibung: This paper aims to assess the influence of canopy cover on lichen growth in boreal forests along a regional forest gradient. Biomass and area gain, and some acclimation traits, were assessed in the old-forest lichens Lobaria pulmonaria (L.) Hoffm., Pseudocyphellaria crocata (L.) Vain., and Usnea longissima Ach. transplanted 110 days in three successional Norway spruce ( Picea abies (L.) Karst.) forest stands (clearcut, young, and old forest) repeated along a rainfall gradient (continental, suboceanic, and Atlantic zones) in Scandinavia. Lichen growth peaked in Atlantic rainforests with mean dry matter (DM) gain up to 36%–38%. The alectorioid lichen U. longissima showed the widest range of growth responses and no signs of chlorophyll degradation. Its highest DM gain consistently occurred in clearcuts, whereas the DM gain was close to zero in the shadiest young forest. The two foliose lichens L. pulmonaria and P. crocata exhibited maximal growth rates in old forests, but apparently growth was limited by low light even in old forests. Their DM gain was reduced in the most sun-exposed clearcuts due to chlorophyll degradation and was relatively high under closed young canopies, suggesting a better adaptation to shade. The lichen responses show that a high frequency and dominance of young and dense fast-growing forest stands at a landscape level are not compatible with large populations of these old-forest lichens and that a lack of lichens under an industrial forestry regime may not necessarily be determined by low dispersal efficiency only.

Beschreibung: The forwarding of logs at harvest areas once the harvesting is done is planned manually by experienced operators. To improve their efficiency and simplify the planning we have developed and tested a decision support system at a major Swedish forest company. The system is based on a combination of a geographic information system (GIS), global positioning system (GPS), and optimization routines to solve the underlying vehicle routing problem. The routes for the forwarders are found by using a repeated matching algorithm. The solution time is short, and it is possible to find routes dynamically in a real-time environment. The geographic information required is found by using a GPS together with data obtained from the bucking software in the harvesters. To show the routes and location of the forwarder, we make use of a GIS that is connected to the GPS. We report on a study with savings in the distance travelled of 8% and numerical tests on the solution methodology. We also compare the proposed solution method with some well-known routing methods.

Beschreibung: We analyzed spatial patterns of overstory trees in late-successional Abies amabilis (Dougl. ex Loud.) Dougl. ex J. Forbes forests and late-successional Pseudotsuga menziesii (Mirb.) Franco forests to establish reference spatial patterns for restoration thinning treatments, and to determine whether thinning treatments with minimum intertree spacing rules result in spatial patterns characteristic of late-successional forests. On average, 32.7% of overstory trees in Abies plots and 26.3% of overstory trees in Pseudotsuga plots occurred as members of multitree clusters (groups of trees in which trees are spaced within a specified minimum distance of each other) at a distance of 3.0 and 4.0 m, respectively. Multitree clusters occurred throughout the three Abies plots; the distribution of multitree clusters within the two Pseudotsuga plots was variable. Spatial patterns of overstory trees in late-successional forests were significantly different from those created by simulated restoration thinning treatments. Restoration thinning treatments that release both individual trees and multitree clusters promote characteristic late-successional tree spatial patterns at the within-patch scale (

Beschreibung: This paper uses hedonic regression techniques to analyze timber bid transactions in central Georgia. Softwood stumpage prices from pay-as-cut transactions are regressed against timber sale and stand characteristics. We identify observable factors that are statistically associated with the volatility of pine sawtimber stumpage prices in the market. The remaining price volatility, defined as market risk, characterizes undiversifiable price volatility in the market. Isolating market risk in this way has implications for relative price risk across predefined timber markets. Applications of this these techniques suggest that analyzing market price variability with total measures alone, such as standard deviation, may provide false senses of timber price risk.

Beschreibung: In forests historically maintained by frequent fire, reintroducing fire after decades of exclusion often causes widespread overstory mortality. To better understand this phenomenon, we subjected 16 fire-excluded (ca. 40 years since fire) 10 ha longleaf pine ( Pinus palustris Mill.) stands to one of four replicated burning treatments based on volumetric duff moisture content (VDMC): wet (115% VDMC); moist (85% VDMC); dry (55% VDMC); and a no-burn control. During the first 2 years postfire, overstory pines in the dry burns suffered the greatest mortality (mean 20.5%); pine mortality in the wet and moist treatments did not differ from the control treatment. Duff reduction was greatest in the dry burns (mean 46.5%), with minimal reduction in the moist and wet burns (14.5% and 5%, respectively). Nested logistic regression using trees from all treatments revealed that the best predictors of individual pine mortality were duff consumption and crown scorch (P 〈 0.001; R2 = 0.34). Crown scorch was significant only in dry burns, whereas duff consumption was significant across all treatments. Duff consumption was related to moisture content in lower duff (Oa; R2 = 0.78, P 〈 0.001). Restoring fire to long-unburned forests will require development of burn prescriptions that include the effects of duff consumption, an often overlooked fire effect.

Beschreibung: Height observations H1and H2present on the right- and left-hand sides of site index models, respectively. The error terms associated with H1and H2, along with parameter estimate errors, affect the estimate of the site index. Projection error variance (PEV), in a projection from A1to A2, consisted of four components associated with H1, H2, the covariance of H1and H2, and the parameter estimate errors. In this study, behaviors of these components were investigated via simulations on the basis of six equations derived from the Lundqvist–Kerf and the Hossfeld IV functions. Simulation results showed that projection interval, projection direction, and selected site-dependent parameter influenced PEV and its components. PEVs of backward and forward projections with the same projection interval lengths were remarkably different if the underlying model was anamorphic. With increasing projection interval length, the PEV of forward projections monotonically increased to a certain value, whereas the PEV of backward projections decreased to zero after reaching a maximum.

Beschreibung: We describe the development of a statistical model of spatial variation in the area burned by lightning-caused forest fires across the province of Ontario. We partitioned Ontario’s fire region into 35 compartments, each of which is relatively homogeneous with respect to its vegetation, weather, and the level of fire protection it receives. We used linear regression and spatial autoregressive models to relate the average annual area burned in a compartment to its vegetation, weather, and level of protection attributes. We also examined the relationship between burned area and the level of protection in two areas that are relatively homogeneous with respect to vegetation and weather. We found a statistically significant relationship between the average annual fraction of the area of a compartment burned by lightning-caused forest fires and its vegetation, weather, and the level of fire suppression effort it receives.

Beschreibung: Phenolic metabolites are frequently implicated in chemical defense mechanisms against pathogens in woody plants. However, tree breeding programmes for resistance to pathogens and practical tree-protection applications based on these compounds seem to be scarce. To identify gaps in our current knowledge of this subject, we explored some of the recent literature on the involvement of phenolic metabolites in the resistance of northern forest trees (Pinus, Picea, Betula, Populus, and Salix spp.) to pathogens. Although it is evident that the phenolic metabolism of trees is often activated by pathogen attacks, few studies have convincingly established that this induction is due to a specific defense response that is capable of stopping the invading pathogen. The role of constitutive phenolics in the resistance of trees to pathogens has also remained unclear. In future studies, the importance of phenolics in oxidative stress, cell homeostasis and tolerance, and the spatial and temporal localization of phenolics in relation to invading pathogens should be more carefully acknowledged. Possibilities for future studies using advanced methods (e.g., metabolic profiling, confocal laser scanning microscopy, and use of modified tree genotypes) are discussed.

Beschreibung: Due to the scarcity of old-growth forests in much of Europe, there is little quantitative information on disturbance processes that influence forest dynamics. However, this information is crucial for forest management that tries to emulate patterns and processes in natural forests. We quantified the gap disturbance regime in an old-growth forest dominated by European beech ( Fagus sylvatica L.) and silver fir ( Abies alba Miller) in the Dinaric Mountains of Bosnia and Herzegovina. We sampled 87 gaps in four stands using line-intercept sampling. The percentages of forest area in canopy gaps and expanded gaps ranged from 12% to 17.2% and 35.5% to 39.7%, respectively. Although many of the gaps were small (1000 m2 with numerous gapmakers made up a disproportionate amount of the total gap area. More than half the gaps had more than one gapmaker and were often in separate decay classes, indicating gaps had expanded over time during separate disturbance events. Furthermore, 51% of all gapmakers were uprooted or wind-snapped, whereas only 22% died standing. These results suggest that wind disturbance plays an important role in creating intermediate to large canopy openings through both gap formation and gap expansion processes.

Beschreibung: Locating a log landing is an important task in forest operations planning. Several methods have been developed to find an optimal landing location and compute a mean skidding distance, but they simplify harvest unit attributes and do not simultaneously consider multiple design factors influencing optimal landing locations. In this study, we introduce a computerized model developed to determine the optimal landing location for ground-based timber harvesting. Using raster-based GIS data, the model finds skid trails from stump to each of candidate landings and selects the best landing location that minimizes total skidding and spur road costs. The model is applied to several hypothetical harvest units with different terrain and harvest volume attributes to analyze the effects of design factors influencing optimal landing locations. Unit boundary shapes, volume distribution, the presence of obstacles, terrain conditions, and spur road construction are considered as influencing design factors.

Beschreibung: European beech ( Fagus sylvatica L.) is increasingly managed by close-to-nature principles, mimicking the gap dynamics of seminatural forests. The prime aim of this study was to analyse natural regeneration reliability under favourable conditions in newly formed gaps. A total of 12 gaps were created by felling three canopy trees for each gap: six gaps in each of the two winters 1996–1997 and 1997–1998. One-half of the gaps were fenced against deer. We recorded advance regeneration density (1997), regeneration density and height (1997–2002), relative light intensity (1997–2002), and volumetric soil moisture content (1997–2002). We also studied the effect of year of establishment, fenced versus unfenced, and position within gap on regeneration. Three or 4 years after gap formation, most gaps had nearly closed. Response of European beech, European ash ( Fraxinus excelsior L. ), and sycamore maple ( Acer pseudoplatanus L.) regeneration to gap formation was limited, and few seedlings were added to the advance regeneration pool during the study period. Other factors, such as relative light intensity, soil moisture, fencing, year of establishment, and position within gaps, all had rather low effects. Thus, the presence of advance regeneration appeared to be a key factor in explaining regeneration patterns in artificially created gaps.

Beschreibung: Increased forest productivity has been obtained by improving resource availability through water and nutrient amendments. However, more stress-tolerant species that have robust site requirements do not respond consistently to irrigation. An important factor contributing to robust site requirements may be the distribution of biomass belowground, yet available information is limited. We examined the accumulation and distribution of above- and below-ground biomass in sweetgum ( Liquidambar styraciflua L.) and loblolly pine ( Pinus taeda L.) stands receiving irrigation and fertilization. Mean annual aboveground production after 4 years ranged from 2.4 to 5.1 Mg·ha–1·year–1 for sweetgum and from 5.0 to 6.9 Mg·ha–1·year–1 for pine. Sweetgum responded positively to irrigation and fertilization with an additive response to irrigation + fertilization. Pine only responded to fertilization. Sweetgum root mass fraction (RMF) increased with fertilization at 2 years and decreased with fertilization at 4 years. There were no detectable treatment differences in loblolly pine RMF. Development explained from 67% to 98% of variation in shoot versus root allometry for ephemeral and perennial tissues, fertilization explained no more than 5% of the variation in for either species, and irrigation did not explain any. We conclude that shifts in allocation from roots to shoots do not explain nutrient-induced growth stimulations.

Beschreibung: We examined the impacts of a severe, regional drought (2001–2002) on trembling aspen ( Populus tremuloides Michx.) forests in the western Canadian interior. A total of 150 plots were established in 25 study areas as part of a regional-scale study (CIPHA). Aspen health and mortality were assessed annually during 2000–2005, and changes in stem biomass were estimated using tree-ring analysis and plot-based measurements. Net mean increment in living biomass for all plots was 2.2 t·ha–1·year–1 during 2000–2002 but subsequently decreased to near zero. This collapse was driven by a more than two-fold increase in stem mortality and a 30% decrease in regional stem growth during and following the drought. The analysis showed that spatial variation in aspen productivity and biomass across the region was positively related to multiyear mean values of a climate moisture index and mineral soil silt content but was negatively related to levels of insect defoliation and wood-boring insects. In contrast, mortality and dieback was best correlated with minimum annual climate moisture index, which provided a measure of short-term drought severity. The results support previous studies showing that aspen forests are moisture limited in this region, which poses concerns for the future under a changing climate.

Beschreibung: Forest litter and soil may contain 〉10 × 106 individual nematodes·m–2 and, regionally, 〉400 species. Root-feeding nematodes may be pathogenic to young plants; microbial-feeding nematodes may increase turnover of the microbial pool; predacious and omnivorous nematodes represent higher trophic levels. The spatial distribution and abundance of nematode species in forests reflect soil type, soil fertility, climate, canopy and understorey plant species, litter depth, forest age, and management. Nematodes may be important in forest nurseries; they occur throughout the rooting depth of forest trees; hyphal-feeding species may influence mycorrhizae; and insect-vectored Bursaphelenchus species are a quarantine risk. Nematode populations interact with those of other soil animals (e.g., mites, tardigrades, enchytraeids, and protozoa). The diversity and abundance of the nematode assemblage make nematodes a useful indicator of soil condition and soil processes. Information available from forest systems suggests that, as long as physical disturbance is minimized and remaining trees or herb layer moderate the microclimate, logging and other forestry operations have only transitory effects on nematode populations. Extreme disturbance, such as bulldozing and slash-and-burn management, can significantly reduce nematode abundance and diversity. In contrast, management that enhances growth of understorey or herb layer can stimulate nematode populations. Each of these changes can be related to changes in food resource availability and environmental conditions, such as soil temperature and moisture. Although details of soil nematode contributions to nutrient processes in forest soils are sparse, that their populations are maintained through cycles of moderate management practices suggest that their beneficial contributions will also be maintained.

Beschreibung: Two midmontane moist evergreen broad-leaved forests on the Ailao Mountains, Yunnan, and one on Mount Emei, Sichuan, southwestern China were studied to characterize the forest structure and dynamics as manifested in size, age, canopy gaps, regeneration modes, and the survival of seedlings. The most dominant canopy trees were species of Lithocarpus and Castanopsis of Fagaceae along with species of Machilus of Lauraceae and of Schima of Theaceae. The vertical structures of the forests were multilayered. All the canopy species had multimodal-shaped size and age distributions. In each forest of the study sites, the mean size of a canopy gap, caused mainly by the death of canopy trees, was smaller than 65 m2. No surviving seedlings of Fagaceae species were found in understories having bamboo with a coverage greater than 25% in any quadrat of the three forests. The poor seedling bank in the study forests is apparently due to the presence of bamboo in the understory. The tree regeneration may be synchronously related to the bamboo flowering event (interval approximately 55–60 years).

Beschreibung: The mechanisms governing tree mortality in surface fires are poorly understood, owing in large part to the absence of a process-based framework for defining and evaluating these mechanisms. This paper begins the development of such a framework by deriving a first-order process model of tree mortality in surface fires (intensities less than approximately 2500 kW·m–1). A buoyant line-source plume model is used to drive heat transfer models of vascular cambium and vegetative bud necroses, which are linked to tree mortality using an allometrically-based sapwood area budget. Model predictions are illustrated for white spruce ( Picea glauca ), lodgepole pine ( Pinus contorta ), and trembling aspen ( Populus tremuloides ) and are compared with independent mortality data for Engelmann spruce ( Picea engelmannii Parry ex Engelm.) and Pinus contorta Dougl. Results help define first-order mortality mechanisms and suggest second-order mortality mechanisms that should be incorporated into future modeling efforts.

Beschreibung: We compared prefire and postfire organic-layer depths in boreal forest types (14 fires) across Canada, and examined tree recruitment as a function of depth. There was extensive within-stand variation in depth, much of it due to clustering of thinner organic layers around boles. There were no significant differences in postfire organic-layer depth among sites with different prefire forest species composition, but sites in the eastern boreal region had thicker postfire organic layers than those in the western boreal region. Mean organic-layer depth was much greater in intact stands than after fires; overall, fire reduced organic-layer depth by 60%, largely because of increases in the area of thin (

Beschreibung: The purpose of this study is to investigate the suitability of the Birnbaum–Saunders distribution to model diameter at breast height (DBH) distributions of near-natural complex structure silver fir ( Abies alba Mill.) – European beech ( Fagus sylvatica L.) forests. The investigations were carried out in Świętokrzyski National Park, situated in Central Poland. To estimate the parameters of the Birnbaum–Saunders distribution, three methods were used: the maximum likelihood method (MLE) and the mean–mean estimator, the modified moment method (MME), and the graphical method (GME). The empirical DBH distributions in near-natural fir–beech stands, which arose according to the model taking into account the overlapping of fir mortality and beech regeneration, were generally conformed to the Birnbaum–Saunders distribution. In such forests, the Birnbaum–Saunders distribution approximated the empirical DBH distributions more precisely than the Weibull and gamma distributions.

Beschreibung: We compared understory vegetation composition and richness in aspen-dominated boreal mixedwood forest stands in Alberta, Canada, that had been burned by wildfire with those that burned and were subsequently salvage logged. Stands were examined at early and midsuccessional (2 and 34 years after disturbance(s), respectively) developmental stages. In comparison with wildfire stands, understory communities of early successional salvage-logged stands were characterized by greater species richness, weedy species presence, higher shrub abundance, and lower abundances of fire-specialist seed bank species. In constrained ordination, the understory community of early successional wildfire stands was related to greater canopy cover, sapling density, and moss depth, whereas that of salvage-logged stands was related to greater light, volume of downed deadwood, and litter and organic matter. Longer term effects of salvage logging on the understory community were minimal and, instead, reflected the influence of forest canopy redevelopment. In midsuccessional stands, understory composition was related to conifer density, litter cover, soil moisture, organic layer depth, tall shrub density, and bryophyte-covered microsite cover. Postfire salvage logging can have substantial short-term effects on the postfire understory plant community; in the longer term, effects will depend to a large extent on the influence of harvesting and subsequent management on canopy redevelopment.

Beschreibung: We sampled second-growth forests ranging in age from 28 to 98 years and compared them with old-growth forests to quantify rates of terrestrial vegetation recovery following harvesting on the northcentral coast of British Columbia. Species richness approximately doubles, while Simpson’s index of diversity increases from 0.81 to 0.91 from young to old forests. Nonmetric multidimensional scaling ordinations showed differentiation, with some overlap, of old-growth and second-growth forests and a fairly strong correlation of stand age with plot scores, driven by plant species presence and cover. Vegetation succession following logging disturbance is driven primarily by predisturbance species composition; most species found in the young forests are present in old forests and the higher species richness typical of old growth is largely due to the establishment of additional cryptogam and herb species of low cover and constancy. Significantly higher cover of shrub, herb, and bryophyte species differentiates old forests from second-growth forests. Forests 41–100 years old average 63%–73% similarity (depending on site type) to old-growth forests based on species presence–absence and 53%–58% similarity based on species cover. The scarcity of western redcedar ( Thuja plicata Donn ex D. Don) in second-growth stands is of particular concern because of the high ecological, cultural, and economic importance of this tree species.

Beschreibung: Buffer zones are an important component of forest-management strategies and are thought to reduce the impact of nutrients released after harvesting on water quality. Conceptually, steep slopes have shorter water residence times than shallow slopes, have a reduced capacity to moderate water quality, and therefore, require wider buffers. Carbon and N concentrations in riparian zone shallow soil water at 30 cm depth and lake water were measured on shallow and steep slopes at the Esker Lakes Research Area in northeastern Ontario to determine if nutrient concentrations were correlated to catchment terrain attributes. Field measured slope, slope class obtained from a triangular irregular network model, and upslope contributing area and topographic index calculated from a digital elevation model were calculated for each sampling location. Modeled terrain properties, including those currently used during forest-management planning, were not significantly correlated with soil water N and C concentrations, whereas only dissolved organic carbon levels were significantly greater on field measured steep slopes. Forest species composition and soil N levels were positively correlated with soil water N concentrations. These results from the undisturbed boreal ecosystem highlight the potential limitation of using only catchment slope as a tool for prescribing riparian buffers during harvesting when considering terrestrial nutrient export.

Beschreibung: This paper has three main objectives: (i) to investigate whether the four-quadrant approach introduced by     J.S. Maini reveals a useful typology for grouping countries by gross domestic product (GDP) and forest cover per capita, (ii) to determine if the framework can enhance our understanding of the relationship between forest cover and GDP per capita, and (iii) to investigate why countries in the four-quadrant world occupy different quadrants and to determine the principal factors affecting country movement across and within the individual quadrants. The examination reveals that countries can be classified into four broad categories and that GDP and forest cover per capita have a low but consistent level of negative association. After regressing economic, institutional, social capital, and other variables on a country’s occupancy and movement in the four-quadrant world, the results suggest that countries in each quadrant share different characteristics and that factors underlying country movement vary according to the quadrant being observed. Overall, countries with less corruption and higher education are likely to experience increases in both forest cover and GDP per capita, while countries exporting a significant proportion of forest products have a reduced probability of increasing both variables.

Beschreibung: In forest breeding programmes, growth has typically been used as a selection trait of prime importance in Norway spruce ( Picea abies (L.) Karst.), whereas less attention has been given to the wood and fibre characteristics. In the above context, we investigated phenotypic relationships between different fibre properties and growth and wood density traits in 20 cloned Norway spruce based on a clonal trial established in the 1970s in southeastern Finland. We found that fibre width showed, on average (2.9%), the lowest phenotypic variation followed by fibre wall thickness (3.4%), coarseness (5.5%), and fibre length (8.1%). All of the phenotypic correlations between the fibre properties were also positive (p 〈 0.05), ranging from moderate to strong, suggesting that selection for one trait could simultaneously affect the other traits. The phenotypic correlations, on average, were quite weak but positive between growth and fibre properties and slightly negative or weak positive between wood density and different fibre properties (p 〈 0.05). Individually, some of the clones showed negative correlation between growth traits and fibre length. As a result, selection for fibre properties alone could also reduce overall stem volume (or stem mass) and would not directly indicate wood density traits and vice versa.

Beschreibung: In the winter of 1998–1999, two partial harvesting treatments that removed 33% (1/3) and 61% (2/3) of stand basal area were applied to even-aged trembling aspen ( Populus tremuloides Michx.) stands and compared with unharvested control stands. Stands in the 1/3 treatment were low thinned, while stands in the 2/3 removal were crown thinned. Coarse woody debris dynamics were assessed during the following 6 years by means of permanent sampling plots and downed wood inventories. Between 1999 and 2004, tree mortality was, respectively, 18%, 17%, and 32% in control stands and 1/3 and 2/3 harvesting treatments. Although total snag density was similar between controls and partial cutting treatments, total snag basal area was significantly higher in controls in 2004. Between 1999 and 2004, net change in aspen snag density was positive for controls and negative for both partial cutting treatments. Partial cutting also exacerbated mortality of small-diameter white birch ( Betula papyrifera Marsh.). Downed wood volume increased by 35 m3·ha–1 in controls and by 25 m3·ha–1 in the 2/3 harvesting treatment, while it decreased by 7 m3·ha–1 in the 1/3 harvesting treatment. Coarse woody debris goals can be established in silviculture prescriptions; type, timing, and intensity of partial cutting are crucial to the outcome.

Beschreibung: A study was established between 1959 and 1961 to study the long-term responses of balsam fir ( Abies balsamea (L.) Mill.) and red spruce ( Picea rubens Sarg.) to precommercial thinning. Three nominal spacings of 4 ft (1.2 m), 6 ft (1.8 m), and 8 ft (2.4 m) were compared with an unthinned control in a randomized complete block design with five replicates. At the time of thinning, natural regeneration averaged 16 years of age, 8 years after harvest. Although thinning had minimal effect on gross total volume production over a 42 to 44 year observation period, actual spacings between 2.1 and 2.5 m produced an average of 360 m3·ha–1gross merchantable volume (GMV), representing a 21% gain over unthinned stands. The same spacings produced quadratic mean diameters of 21 and 23 cm, respectively, compared with 18 cm in the unthinned stands. These size increases translated to individual stem volume gains of 33% and 62%, significantly reducing the age at which thinned stands would meet a specified minimum requirement for merchantability or habitat. The mean annual increment of GMV ranged from 6 m3·ha–1·year–1in unthinned stands, to more than 7 m3·ha·–1·year–1in the thinned stands, and had not yet culminated an average of 50 years postharvest.

Beschreibung: This work proposes a novel approach to describing intraring wood density characteristics as functions of annual weather events. This approach was tested using three different maritime pine clonal experiments, in which X-ray microdensitometry has revealed conspicuous within-ring patterns affecting most of the trees in 1996. This pattern has been interpreted as the variation of tree response to weather-controlled changes of water balance during the 1996 growing season. The level of tree response was estimated using an original norm of reaction obtained from the microdensity profiles. A 1996 site drought index profile was synchronized with the 1996 microdensity profile by pairing conspicuous points of abrupt change in both profiles (breakpoints). Regression of density breakpoints on drought breakpoints describes the norm of reaction of radial increment to water availability, and the plasticity of radial increment to changes in soil moisture is described by the slope of the regression. The slope showed moderate levels of genetic control that depended on the site and could potentially be used as criteria for the evaluation of tree adaptation to weather.

Beschreibung: Fuel distribution in the southern Appalachian Mountain region was measured in over 1000 study plots that were stratified by topographic position (aspect and slope position) and disturbance history. Few fuel differences occurred among topographic positions for undisturbed plots, indicating that fuel accumulation is no greater on highly productive sites than on less productive sites. Litter was slightly higher on undisturbed upper slopes (4.2 t/ha) than on lower slopes (3.7 t/ha) but woody fuels showed no significant differences. Rhododendron ( Rhododendron spp.) and mountain laurel ( Kalmia latifolia L.) were less common than expected, occurring on 25% and 42% of sampled plots, respectively. Disturbance history and type played a greater role in determining fuel loads than did topographic position. Disturbances had occurred on 30% of sample plots within the past 10 years and were most common on exposed slopes. Litter was significantly lower in burned plots (3.5 t/ha vs. 4.0 t/ha in undisturbed plots). One-hour fuels (1.0 t/ha) were significantly higher on beetle-killed plots than on undisturbed plots (0.7 t/ha) while larger woody fuels tended to be greater in plots subjected to beetle attack, fire, and wind.

Beschreibung: The Earth’s temperature has increased 0.6 °C over the last 100 years, and further climate change is predicted to potentially raise it by 3.5 °C over the next century. More than half of the global annual net primary production of biomass is estimated to occur in the tropics, especially tropical evergreen forest. In temperate forests, increasing temperature may extend the non-frost growing season, and thus increase the CO2 sequestration rate, but some authors have also suggested a negative impact of warming in tropical forests from decreased photosynthetic activity. Using the PL model (Ricker and del Río 2004), we forecast growth of two Mexican tree species after climate warming. The model predicts the high-mountain species Pinus hartwegii Lindl. to decrease its expected relative growth throughout its lifetime by 10.6% as a consequence of a 0.6 °C temperature increase; in contrast, the tropical rainforest species Diospyros digyna Jacq. is predicted to increase its expected relative growth throughout its lifetime by 25.4%. The key factor appears to be the expected relationship between temperature and precipitation, rather than temperature alone. While one cannot expect a universal response across sites, some standing tropical rainforests such as those at Los Tuxtlas in Mexico may constitute a carbon sink in a changing climate.

Beschreibung: Basic density (BD), fibre length (FL), fibre width (FW), and fibre wall thickness (FWT) were investigated in 46 Norway spruce ( Picea abies (L.) Karst.) trees from five different stands in eastern Norway. From each tree, wood samples were collected in different radial and longitudinal positions. Random coefficient mixed models were used to investigate variation within as well as among trees, both within and among stands. The R2 with random effects included, describing the best possible (individual) fit of the observed data to the models, were 0.90 for BD, 0.99 for FL, 0.88 for FW, and 0.91 for FWT. With only fixed effects, the best model explained 56% of the total variation for BD, 94.5% for FL, 61% for FW, and 63% for FWT. A common model for all trees, without tree and site information, predicted FL well but BD, FW, and FWT poorly. Adding site index, breast height diameter, and tree height to the models reduced the residual variance considerably for FW, FWT, and particularly BD, whereas only a minor improvement was gained for FL. The latter type of models might be easier to use for industrial purposes. Although information about ring width gave further improvements, ring width measurements are time consuming and difficult to perform in the forest and in industrial environments.

Beschreibung: A variety of competing hypotheses have been described to explain yield decline in Chinese-fir ( Cunninghamia lanceolata (Lamb.) Hook.) plantations. The difficulty in implementing field experiments suggests ecosystem modeling as a viable option for examining alternative hypotheses. We present a conceptual model of Chinese-fir yield decline and explore its merits using the ecosystem-based FORECAST model. Model results suggest that yield decline is caused primarily by a decline in soil fertility, largely as a consequence of slash burning in conjunction with short rotations. However, as tree leaf area declines, there is a transition (over subsequent rotations) from seed rain based competition to bud bank based competition, increasing the competitive impact of minor vegetation on tree growth. Short rotations increase understory survival between rotations and may cause a gradual shift from tree dominance to shrub/herb dominance over subsequent rotations. These effects are most evident on nutrient-poor sites, but understory competition poses a significant yield decline risk on good sites as well. We conclude that sustainable production in Chinese-fir plantations requires the avoidance of activities that compromise soil fertility and increase understory competition. The risk and severity of yield decline would be reduced by increasing rotation lengths and avoiding plantations on infertile sites.

Beschreibung: Decline index (indicator of crown condition) data from 102 forest plots (approximately 10 000 trees) during 1986–2004 were compiled to derive survival models for south-central Ontario, Canada. The dominant species was sugar maple ( Acer saccharum Marsh.) with approximately 75% occurrence (n = 7640). The predictor variables for sugar maple survivorship included the decline index of 1 or 2 years prior to the beginning of the modelled period and ecological region (Algoma, Georgian Bay, Huron–Ontario, and Upper St. Lawrence). The observed crown condition of sugar maple improved significantly over the study period; in contrast, short-term mortality rate did not improve. The risk of sugar maple mortality could be predicted from decline index data for a single year indicating that the risk of tree death increases with higher decline index values (declining crown condition). Moreover, using 2 years of decline index data indicated that the risk of tree death also increased with the length of consecutive time individual trees have higher decline index values. Trees in the Algoma region, which represent the northern limit of sugar maple distribution in Ontario, were significantly more likely to die than trees in Huron–Ontario region.

Beschreibung: Forest management prescriptions increasingly incorporate snag and downed dead wood (DDW) guidelines. This study utilizes permanent inventory plots to determine dead wood dynamics in 33 balsam fir ( Abies balsamea (L.) Mill.) – spruce ( Picea spp.) (BFSP) and 17 spruce – balsam fir (SPBF) stands in New Brunswick, Canada. Stands were declining, unmanaged, and had a history of recurrent spruce budworm ( Choristoneura fumiferana (Clem.)) outbreaks and aerial insecticide spraying. Fixed-area sampling matched remnants of 1165 dead trees and 864 corresponding pieces of DDW to plot trees that died over the last 15–18 years with known year and cause of death. Declining BFSP stands had the highest accumulation of dead wood (196 m3/ha) compared with SPBF and nondeclining BFSP (122 m3/ha and 77 m3/ha, respectively). Dead wood dynamics were influenced by cause of death, as a function of differences in tree height at death affecting snag decay, fragmentation, and fall. One-half of all dead trees never made a significant contribution to the snag population (25% uprooted and 25% stem breakage), and attrition resulted in only 50% of snags standing with a mean height of 6 m 15–20 years after death. This study will be of direct value to those managing or modeling dead wood dynamics in similar forests.

Beschreibung: The influence of climatic factors on tree-ring width and the formation of double rings was studied in Quercus ilex L. growing in a coppice stand left unmanaged for 22 years. Ten trees were felled and discs were taken every 30 cm from bole and dominant branches. Dendrometer bands were installed on 10 nearby trees and the data recorded were used to confirm the accuracy of our tree-ring identification. They were also used to relate the seasonal radial growth pattern to double-ring formation. Double rings were frequent and occurred consistently along the stem. Two types of double rings could be recognized according to their width: type I, with the extra growth band accounting for approximately 50% of the tree ring; and type II, with a narrow extra growth band. Type I double rings were formed when approximately 1/2 of the growing-season precipitation occurred during the second growth period of the season and after the summer drought. Type II double rings occurred when approximately 1/3 of the precipitation in the growing season occurred after the summer drought. The formation of double rings was triggered by rainfall in summer and the extra growth-band width was related to summer and autumn environmental conditions. Double rings in Q. ilex can potentially be used in dendroclimatological studies, as they are formed in response to climatic conditions within the growing season.

Beschreibung: Using an extensive Douglas-fir data set from southwest Oregon, we examined the (1) performance and suitability of selected prediction strategies, (2) contribution of relative position and stand-density measures in improving tree height (h) prediction values, and (3) effect of different subsampling designs to fill in missing h values in a new stand using a regional nonlinear model. Nonlinear mixed-effects models (NMEM) substantially improved the accuracy and precision of height prediction over the conventional nonlinear fixed-effects model (NFEM) that assumes the observations are independent, particularly when a few trees are subsampled for height. The predictive performance of a correction factor on a NFEM with relative position and stand-density measures was comparable to that of a NMEM when four or more trees were subsampled for height. When two or more heights were randomly subsampled, the NMEM efficiently explained the differences in the height–diameter relationship because of the variations in relative position of trees and stand density without having to incorporate them into the model. When only one height was subsampled, selecting the largest diameter tree in the stand would result in a lower predicted root mean square error (RMSE) than randomly selecting the height, regardless of the model form or fitting strategy used.

Beschreibung: We evaluated agreement in the location and occurrence of 20th century fires recorded in digital fire atlases with those inferred from fire scars that we collected systematically at one site in Idaho and from existing fire-scar reconstructions at four sites in Washington. Fire perimeters were similar for two of three 20th century fires in Idaho (1924 and 1986). Overall spatial agreement was best in 1924 (producer’s accuracy = 94% and 68% and user’s accuracy = 90% and 70% for the 1924 and 1986 fires, respectively). In 1924, fire extent from the atlas was greater than for fire scars, but the reverse was true for 1986. In 1986, fire extent interpreted from the delta normalized burn ratio derived from pre- and post-fire satellite imagery was similar to that inferred from the fire-scar record (producer’s accuracy = 92%, user’s accuracy = 88%). In contrast, agreement between fire-scar and fire-atlas records was poor at the Washington sites. Fire atlases are the most readily available source of information on the extent of late 20th century fires and the only source for the early 20th century. While fire atlases capture broad patterns useful at the regional scale, they should be field validated and used with caution at the local scale.

Beschreibung: Effective fire suspension in Fennoscandian boreal forests has caused a number of species to become threatened. To compensate for the negative ecological impacts of fire elimination, prescribed burning of forests as a restoration method has been introduced recently. We studied the effects of controlled burning on assemblages of wood-decaying polypores (Basidiomycota), including red-listed species, in a large-scale field experiment in Finland. A total of 24 forest sites were included in the factorial study design with two factors: logging and burning. The presence of polypore fruiting bodies was documented 1 year before the treatments, and 1 and 4 years after the treatments. Over 11 000 observations of 104 species of polypores were made. Change in the fungal species composition due to logging and burning was clear after 4 years. At the species level, the responses to logging and fire varied depending on the species. Treatments increased fruiting of pioneer decayers; however, most red-listed species seemed to suffer. Thus, prescribed burning does not offer immediate benefits for most red-listed species. In unlogged forests, the restorative effects of fire are likely to be seen later as the death and decay processes of trees continue and provide more resources for polypores.

Beschreibung: We used General Land Office survey data (1860–1890) and interpreted aerial photography from the 1930s, 1970s, and 1990s to quantify forest disturbance frequency and spatial patterns for four time periods in the Mixed Forest Province of Minnesota. The study region included eight subsections within the Mixed Forest Province of Minnesota’s Ecological Classification System. Presettlement disturbance and spatial pattern estimates varied across the eight subsections indicating a strong relationship to soil and landform characteristics. Land surveyors primarily recorded higher severity disturbances that resulted in significant tree mortality. The 1900–1940 era was characterized by a short-term increase in fire frequency that was relatively uniform across the study region, in contrast to the variability of the presettlement (1860–1890) landscape. In the postsettlement period (1940–1995), timber harvest replaced fire as the dominant disturbance factor. Similar management practices among subsections created similar harvest rates throughout the study region. These management practices imposed a more homogeneous pattern dominated by small (10–25 ha) patches. Management practices now have a greater influence than natural processes in the generation of landscape pattern in the Mixed Forest Province of Minnesota. Information on presettlement forest conditions and subsequent changes can be used by land managers to restore spatial pattern variability in managed forest landscapes.

Beschreibung: The seasonal flight of Hylurgus ligniperda (Fabr.) and Hylastes ater (Paykull) and the effect of log placement in Pinus radiata D. Don plantations on colonization by these species in Chile are described. Seasonal flight activity was monitored with α-pinene and ethanol-baited traps at three sites and colonization synchrony with trap logs. In two experiments with individual logs and log decks, data were recorded on adults found under the logs, attacks on the bark surface, and adults beneath the bark of logs in three storage treatments: individual logs and log decks (1) touching the soil; (2) elevated above the soil; and (3) touching other logs (individual logs) or logs separated with spacers (log decks). Each treatment was replicated in a plantation and a clearcut. At San José (region IX) and Peña Blanca (region VIII), H. ligniperda flew primarily in spring and H. ater flew in spring and fall. At Brasil (region VIII), H. ligniperda and H. ater flew primarily in summer through fall. The flight of H. ligniperda was more synchronized with colonization of logs than was that of H. ater. The mean number of attacks was significantly higher on individual logs and log decks stored on the soil than on those elevated above the soil, and significantly higher in the clearcut than in the plantation. Mean numbers of adults beneath the bark did not vary significantly among individual-log treatments, but were significantly lower in the log decks separated by spacers. Logging during nondispersal periods (June–August in regions VIII–X) and storing log decks above the soil could reduce beetle colonization of newly cut logs. With this information, management of recently cut logs (harvesting, transport, and storage operations) could be scheduled and executed so as to reduce the risk of these species being introduced into log-importing countries.

Beschreibung: Using a long-term silvicultural experiment, we (i) investigated how epiphyte and arthropod communities were affected by height on the stem, bark texture, and stem diameter of red maple ( Acer rubrum L.) trees, (ii) examined how harvest gaps influenced epiphyte and arthropod communities on red maple boles, and (iii) explored whether these effects influenced the relationship between the epiphyte and arthropod communities. Arthropod and epiphyte assemblages dwelling on the bark of red maple trees located in undisturbed forest and harvest gaps varied with height. Bryophytes, Cladonia spp., and cyanolichens were most abundant near the base of the tree, while noncyano, foliose lichens and fruticose lichens were most abundant 4–6 m above the ground. Acari, Araneae, and Collembola were most abundant near the base of the tree, while Diptera were most abundant above 2 m. A previously undocumented assemblage of dipterans (flies), primarily in the suborder Nematocera, was found. Gap harvesting reduced the abundance of bryophytes, Collembola (springtails), Araneae (spiders), and total arthropods on the bark of red maple. Canonical correlation analysis revealed a positive association between bryophytes, Collembola, and Araneae. A strong correlation between Collembola and Araneae suggested a possible trophic interaction that may be affected by gap harvesting through a reduction in bryophyte abundance.

Beschreibung: Leaf physiology and stem growth were assessed in loblolly pine ( Pinus taeda L.) in response to 10 to 11 years of treatment with weed control (W), weed control plus irrigation (WI), weed control plus irrigation and fertigation (WIF), or weed control plus irrigation, fertigation, and pest control (WIFP) to determine whether increased resource availability can push productivity of loblolly pine closer to its biological growth potential expressed in favorable, exotic environments. Maximum basal area and stem biomass were 41 m2·ha–1 and 172 Mg·ha–1, respectively, in response to fertigation. Stemwood biomass production was positively and linearly related to basal area. Belowground woody biomass was highest in the WIF and WIFP treatments and averaged 50 Mg·ha–1, but the W and WI treatments exploited a greater area of soil with low-density coarse roots. Fertigation increased foliar nitrogen concentration and foliage biomass, but treatment had no effect on leaf physiological parameters or growth efficiency. Comparison with growth rates reported for loblolly pine in Hawaii revealed that loblolly pine grown in its native range can produce the high yields observed in exotic environments when stands are below maximum carrying capacity.

Beschreibung: This study contributes to the Pinus pinaster Ait. breeding programme, which is reaching the third generation by adding information on wood quality of 46 open-pollinated families from a progeny trial located in Leiria, Portugal, that originated from seed collected in a clonal seed orchard. A total of 552 seventeen-year-old trees were sampled at 2 m height. Trends were studied from the pith outward in variance components and narrow-sense heritability (h2) of wood density components and ring-width characteristics as well as genetic correlations between cambial ages. Mean ring density (RD), minimum density (MND), maximum density (MXD), earlywood density (EWD), latewood density (LWD), earlywood width, latewood width, ring width, latewood percentage, and heterogeneity index were determined using X-ray densitometry procedures. RD had higher genetic control (h2 = 0.63), and heritability values of earlywood components (h2MND = 0.54, h2EWD = 0.60) exceeded those of latewood components (h2MXD = 0.34, h2LWD = 0.26). Heritabilities increased with ring number from pith for almost all wood density components, and there were high age–age genetic correlations for wood density traits (rg 〉 0.98).

Beschreibung: Persistence of tree species in a habitat depends on their ability to avoid and respond to disturbance-related damage. Responses to stem damage vary among species but typically include bark wound closure and prevention of xylem decay spread. These responses are associated with anatomical, structural, and physiological traits. This study explores how xylem (vessel size and (or) abundance, parenchyma abundance, ray width, and wood density) and phloem (bark thickness, proportion of live inner bark, ray width and (or) dilation, inter-ray distance, and tissue density) traits relate to responses to stem damage in seven species from the Bolivian Amazon. Rates of bark wound closure and radial xylem decay penetration were compared 2 years after experimental damage. A species that closed bark wounds rapidly (100% in Chorisia speciosa A. St.-Hil.) was not efficient at constraining xylem radial decay spread (1.7 mm). The opposite was true for Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr., a species that closed wounds slowly (30%) but efficiently controlled decay spread (0.5 mm). The relationship between anatomical and (or) structural traits and damage response variables revealed that species with favorable traits for rapid wound closure (e.g., widely dilating rays) had traits that favored xylem decay spread (e.g., low wood density). It is plausible that this apparent trade-off is based on physiological and phylogenetic constraints.

Beschreibung: Fungal foliar endophytes (hereinafter endophytes) were isolated on malt agar from black spruce ( Picea mariana (Mill.) BSP) needles from the Quebec boreal forest during 2 successive years to study their distribution along a latitudinal transect. Twenty sites were sampled in 2002, 8 of which were sampled again in 2003, in an area located between 47°N and 54°N in the province of Quebec. The endophytes were named whenever possible, using homologies of ITS rDNA sequences from GenBank. In 2002, 232 morphotypes were classified in 40 operational taxonomic units (OTUs), and in 2003, 93 morphotypes were classified in 16 OTUs, for a total of 44 OTUs. Isolation frequencies of the most common endophyte species were 28% for Lophodermium piceae (Fuchel) Höhn., 17% for Darkera parca Whitney, Reid & Piroz, and 9% for Dwayaangam colodena Sokolski & Bérubé. Of the 44 OTUs identified, 18 were unique to P. mariana and found on one site only, possibly constituting rare endemic species. There was some evidence that needles colonized with rare endophyte species were more abundant in the western and southern regions than in the northern region. This supports our hypothesis that as the diversity of tree species surrounding the P. mariana decrease from the southern region to the northern region, the diversity of endophytes also decreases.

Beschreibung: The evergreen sclerophyllous broadleaf forests composed of alpine oaks in the Hengduan Mountains of southwest China are mainly distributed at altitudes of 1700–4800 m. Gas exchange and related leaf traits of Quercus guyavifolia H. Lév. and Quercus pannosa Hand.-Mazz. were measured at eight sites along an altitudinal gradient to understand their physiological adaptabilities. Both Q. guyavifolia and Q. pannosa showed a significant midday depression in the photosynthesis rate (PN) due to a high vapor pressure deficit and high temperature around noon. The optimum temperature for photosynthesis for the two oak species ranged from 17 to 23 °C depending on altitude. When the temperature exceeded 25 °C, PN decreased sharply. However, PN was not depressed by high irradiance. As altitude increased, the leaf dry mass per unit area of the two oaks increased but the chlorophyll content decreased, while the maximum values of daily mean photosynthesis rate (Pd), maximum photosynthesis rate (Pmax), Pd/Pmax, water-use efficiency, and photosynthetic nitrogen use efficiency occurred at altitudes of 3240–3610 m. Although the photosynthetic capacities of the two oaks were higher in August than in May, altitudinal trends did not change with season. The altitudinal range from 3240 to 3610 m would be optimal for the growth and development of these two alpine oaks in the Hengduan Mountains.

Beschreibung: It is critical to understand how the public prioritizes multiple forestry values when establishing objectives for sustainable forest management. While this is a complex and difficult task, a necessary step is to elicit a broad range of public opinions in forest planning to ensure that decisions serve the needs of various forest stakeholders and society at large. This study seeks to understand how six forest dependent communities in British Columbia prioritize a number of attributes associated with sustainable forest management by using a simple survey-based measurement tool, the Thurstone scale. The results suggest that ecological attributes are a higher priority for survey respondents followed by quality of life, global warming, and economic considerations. This paper explores some of the ramifications of the priorities for sustainable forest management measured in these six communities as well as implications for using the Thurstone scale in processes like Public Advisory Groups.

Beschreibung: Dendrochronology climate reconstruction studies often sample dominant, open-grown trees to reduce competition effects and isolate annual climate influences on radial increment growth. However, there has been no examination of how species respond as stand densities increase or which species in mixed-conifer forests provide a better record of past climate. We sampled 579 trees representing five upper montane mixed-conifer species at the Teakettle Experimental Forest in California’s southern Sierra Nevada to determine species-specific responses to annual climatic fluctuations. Using the Kalman filter, we examined the affect of local stand density on growth response and whether the growth–climate relationship improved with a time lag. The Kalman filter iteratively calculates error for predicted versus actual radial growth and accounts for this variation in the corrector equation. Under current high-density conditions, shade-tolerant white fir ( Abies concolor (Gord. & Glend.) Lindl.) provided the best model for climate reconstruction. Shade-intolerant Jeffrey pine ( Pinus jeffreyi Grev. & Balf.) had a lagged response to annual climatic fluctuations, possibly because its roots may tap water reserves in granitic bedrock fissures. Open-grown trees provided more accurate records of climate. Changes in forest density in this forest may have resulted in changes in species-specific response to annual climatic fluctuations.

Beschreibung: This study provides a comprehensive set of functions for predicting biomass for Common beech ( Fagus sylvatica L.) in Central Europe for all major tree compartments. The equations are based on data of stem, branch, timber, brushwood (wood with diameter below 5 or 7 cm), foliage, root, and total aboveground biomass of 443 trees from 13 studies. We used nonlinear mixed-effects models to assess the contribution of fixed effects (tree dimensions, site descriptors), random effects (grouping according to studies), and residual variance to the total variance and to obtain realistic estimates of uncertainty of biomass on an aggregated level. Candidate models differed in their basic form, the description of the variance, and inclusion of various combinations of additional fixed and random effects and were compared using the Akaike information criterion. Model performance increased most when accounting for between-study differences in the variability of biomass predictions. Further, performance increased with the inclusion of age, site index, and altitude as predictor variables. We show that neglecting variance partitioning and the fact that prediction errors of trees are not independent with respect to their predictor variables will lead to a significant underestimation of prediction variance.

Beschreibung: A midrotation commercial thinning experiment conducted in two independent spruce plantations compared several tree selection strategies as both early (age 19 or 24 years) single and double (again at age 30 or 34 years) entries over a 16 year period. A delayed (age 30 or 32 years) single-entry thinning and unthinned portions of the plantations served as references for the other treatments. With the exception of 50% systematic row thinnings, removal intensities were held relatively constant at about 40% of stand basal area. All of the thinning treatments satisfied the objectives of focusing diameter and volume growth on a reduced number of stems and making merchantable volume available during the rotation. Early single-entry thinning allowed 30–50 m3/ha to be harvested, increasing quadratic mean diameter and mean merchantable volume per stem at the end of the observation period by 10% and 24%, respectively, over unthinned stands. A second thinning removed an additional 48–64 m3/ha and increased diameter and volume gains to 25% and 71%, respectively. Only marginal differences were observed between the tree-selection strategies, and there were no overall gross total or gross merchantable volume gains or losses associated any of the thinning treatments relative to the unthinned plantations.

Beschreibung: A percent stocking change model was developed for lodgepole pine ( Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) in Alberta based on spatially mapped permanent sample plot data. Percent stocking was defined as the percentage of 10 m2 subplots occupied by at least one tree with a minimum height of 1.3 m. The difference equation technique was employed to fit the model. Three model forms were examined and the logistic function was chosen as the final model. Site index was found to be a significant predictor and incorporated into the model. Analyses revealed that the model had correlated, but homoskedastic errors and the correlated errors were modeled by spherical covariance structure using NLINMIX macro in SAS. A percent stocking index, defined as the percent stocking at 50 years total age, was introduced and derived from the developed model. The percent stocking model had both forward and backward projection capabilities. It was demonstrated, both on model fitting and validation data, that the model adequately portrayed the percent stocking dynamics of lodgepole pine stands in Alberta. The model also provided an important basis for creating linkages between reforestation survey results and future yield, which is crucial for sustainable forest management.

Beschreibung: Impacts of elevated tropospheric ozone and soil nitrogen amendment on two native European aspen ( Populus tremula L.) and eight hybrid aspen (P. tremula L. × Populus tremuloides Michx.) clones were studied in a free-air ozone exposure system. Potted saplings were exposed to ambient (ca. 20 ppb) or 1.5× ambient ozone and two levels of soil nitrogen (39 and 78 kg N·ha–1·year–1 in the first year, 60 and 140 kg N·ha–1·year–1 in the second year for low-nitrogen and high-nitrogen treatments, respectively) over two growing seasons. The plants were measured for photosynthesis, chlorophyll fluorescence, and biomass accumulation. Ozone decreased leaf-level net photosynthesis (Asat) in particular early in the growing season and maximum quantum yield of photosystem II (Fv/Fm) at the end of the growing season. Nitrogen amendment increased the growth of all plant parts and mitigated the adverse ozone effects. There were significant differences in ozone responses among the clones, and we were able to cluster the clones into sensitivity groups based on their growth responses. The most ozone-tolerant genotypes were hybrid aspen clones, indicating that populations that have already experienced selection for ozone-tolerant genotypes should be used to cross-breed with ozone-sensitive populations to achieve tolerance of a climate with increasing tropospheric ozone concentrations.

Beschreibung: Real option theory has for some decades been used in forest economics to analyse problems that involve irreversible and stochastic decisions. One of these problems is when to harvest a stand and with what species to regenerate. This problem is analysed here, assuming that the value development of both the present and the next stand is stochastic and that the regeneration species must be chosen. The latter decision is assumed asymmetric, as natural regeneration can be exchanged for a planted regeneration but not vice versa. Consequently, the decision can be postponed for some time if natural regeneration is foreseen, making a species change possible if favoured by the value development or if regeneration of the present species proves unsuccessful. This is an extension of the traditional two-option problem, by involving several stochastic elements that are nested in another option problem: when to harvest a stand. Thus the decision on when to harvest a stand depends on its present value as well as on the value of the future stand, both values evolving stochastically. The problem is formulated as a sequential two-option problem and solved numerically.

Beschreibung: Until recently, natural dynamics of mixedwood stands have been largely ignored, resulting in the transformation of many North American mixedwoods into conifer- or hardwood-dominated stand types. The goal of this study was to examine canopy gap dynamics in balsam fir ( Abies balsamea (L.) Mill.) – yellow birch ( Betula alleghaniensis Britt.) mixedwoods to better understand possible mechanisms for species coexistence. Gap proportion in 12 study stands varied between 9% and 30% of the total stand area, while gap size varied from 20 to 2100 m2. Balsam fir mortality was the primary cause of gap formation. Balsam fir and mountain maple ( Acer spicatum Lamb.) dominated the tree and shrub regeneration layers, respectively. Shrub competition slows the natural filling of gaps by tree species. Our results indicate that yellow birch is most abundant in gaps over 800 m2 and balsam fir in those under 200 m2. Transition models showed that the greater longevity of yellow birch than balsam fir ensured its maintenance as a dominant. Dominant species coexistence thus results from divergent use of available resources through time and space. Forest management should maintain variability in harvest timing and size because the use of one gap size or a single rotation age will lead to an imbalance in species proportion relative to natural stands.

Beschreibung: An extensive survey of spruce growth forms was conducted in frost hollows of the Parc national des Grands-Jardins located in the highlands of the Charlevoix region (Quebec). Frost hollows studied were situated within lichen woodlands, which are a favourable environment for the formation of cold air masses. The mean number of frosts recorded 1 m aboveground during the growing seasons of 1997–2002 indicates that there is no frost-free period in frost hollows. Based on the records of minimum temperature in two frost hollows, the critical threshold for frost-ring formation is a nocturnal temperature below 0 °C for 6.4 h with a climax at around –5.7 °C, i.e., at a cooling rate of 1.78 °C/h under the freezing point for at least 3.2 h. Frequent and severe nocturnal frosts slow down the colonization of black spruce in frost hollows, and are at the origin of the inverted tree line that surrounds frost hollows. Mean stem height of black spruce in frost hollows is 1.97 m ± 2.15. A comparative analysis of aerial photographs between 1950 and 1996 showed a 34% reduction in frost hollow area due to the progressive colonization of black spruce. Massive tree establishment occurred in frost hollows between 1970 and 1980 because of the reduced frequency of freezing temperatures during the growing season.

Beschreibung: We studied responses of Acari, especially oribatid mites, to prescribed low-intensity fire in an east side pine site in the southern Cascade Range in California. We compared oribatid population and assemblage responses to prescribed fire in stands that had been selectively logged to enhance old growth characteristics, in logged stands to minimize old growth characteristics, and in undisturbed forest reference stands. Low-intensity prescribed fire altered habitat characteristics within the organic layer of forest soil. Acarine populations declined following prescribed fire, and oribatid losses accounted for two thirds of that decline. Individual oribatid species responded differently to prescribed fire, with a few populations increasing after fire but most declining. The prescribed fire also altered oribatid assemblages, reducing species richness and species diversity and modifying assemblage dominance relationships. We also identified several oribatid taxa that were potential indicator species of fire effects upon forest soil fauna. Finally, our results suggested that oribatid responses to fire were intensified by stand alteration and especially by removal of old growth structural characteristics. Decline in oribatid abundance, species richness and diversity, and loss of equilibrium dominance relationships was greatest in the low structural diversity plots.

Beschreibung: We evaluated geographic variations in mean fire return intervals and postfire forest succession within a 66 497 km2 land area located in the eastern Quebec boreal forest. Fire return intervals were calculated using a time since last fire map for 1800–2000, and forest dynamics were studied by superimposing 3204 forest inventory plots onto the fire map. Mean fire return interval proved significantly shorter in the western part of the study area, at 270 years, compared with the eastern part, where it was probably more than 500 years. The two main tree species in the study area were balsam fir ( Abies balsamea (L.) P. Mill.) and black spruce ( Picea mariana (Mill.) BSP). Balsam fir abundance increased progressively as a function of time since fire, whereas black spruce abundance increased during the first 90 years after fire and then declined. Balsam fir was significantly more abundant in the southeastern portion of the study area, which we attribute to the combined limitations imposed by temperature along the north–south axis and by fire along the east–west axis. Large forest patches (i.e., ≥200 km2) dominated by early successional tree species, within a matrix of irregular black spruce – balsam fir mixtures, are an important feature of preindustrial forest landscapes in this region.

Beschreibung: We measured soil seed banks in 102 plots within a 110 000 ha Arizona Pinus ponderosa landscape, determined seed-bank responses to fire cues and tree canopy types (open or densely treed patches), compared seed-bank composition among ecosystem types, and assessed the utility of seed banks for ecological restoration. Liquid smoke was associated with increased community-level emergence from seed banks in greenhouse experiments, whereas heating to 100 °C had minimal effect and charred P. ponderosa wood decreased emergence. We detected 103 species in seed-bank samples and 280 species in aboveground vegetation. Erigeron divergens was the commonest seed-bank species; with the exception of Gnaphalium exilifolium , species detected in seed banks also occurred above ground. Although a dry, sandy-textured black-cinder ecosystem exhibited the greatest seed density, seed-bank composition was more ecosystem-specific than was seed density. Native graminoids (e.g., Carex geophila and Muhlenbergia montana ) were common in seed banks, whereas perennial forbs were sparse, particularly under dense tree canopies. Our results suggest that (i) smoke may increase emergence from seed banks in these forests, (ii) seed banks can assist establishment of major graminoids but not forbs during ecological restoration, and (iii) seed-bank composition is partly ecosystem-specific across the landscape.

Beschreibung: The intrinsic variability of volatile organic compound emissions and photosynthetic parameters in response to two drying cycles was investigated on Quercus suber L. saplings originating from three Tunisian populations native to contrasting climates. Emissions mainly included monoterpenes plus traces of sesquiterpenes and methylsalicylate that steadily decreased during the experiment unrelated to treatments. Instead, monoterpene emissions increased by 30% during a first moderate drought and remained enhanced after rewatering, while CO2–H2O gas exchange was reduced by 30%. A second severe drought decreased monoterpene emissions to about two third of controls, strongly inhibited gas exchange and photochemical efficiency, and caused a partial loss of chlorophyll. Rewatering led to only partial recovery of emissions and photosynthetic parameters of stressed plants whose biomasses and leaf nitrogen and chlorophyll contents were lower than in control plants. There was no clear difference among populations in volatile organic compound emissions or photosynthetic parameters. However, the population inhabiting the most arid region had intrinsically smaller leaves with lower specific leaf masses and higher drought-related leaf losses than populations native to less arid climates. These results indicate an adaptive shift from stress-tolerating towards stress-avoiding ecotypes under arid climates involving mainly morphological adaptations without apparent differentiation in volatile organic compound production, probably because of its high phenological plasticity.

Beschreibung: Preindustrial forest composition for 〉180 000 km2 throughout central and northeastern Ontario was recreated from Ontario Crown land survey notes (1816–1955) and compared with existing forest composition derived from current Forest Resource Inventories (1998–2009) in each of Site Regions 3E, 4E, and 5E. A validation analysis was performed using the Forest Resource Inventory data to test the assumption that sampling the land survey tree species composition along township boundaries is adequate in describing the composition of the whole forest. The majority of tree species in each of the three site regions validated successfully. A binary logistic regression model allowed birch genera to be classified at the species level to aid in the interpretation of survey notes. All analyses showed significant reductions in conifers (especially red pine ( Pinus resinosa Ait.), white pine ( Pinus strobus L.), and eastern larch ( Larix laricina (Du Roi) K. Koch)) and significant increases in maple ( Acer spp.), oak ( Quercus spp.), white birch ( Betula papyrifera Marsh.), and poplar ( Populus spp.).

Beschreibung: This study compared the performance of Eucalyptus camaldulensis Dehnh. planted at four spacings (2.0 m × 2.0 m, 2.5 m × 2.5 m, 3.0 m × 3.0 m, and 3.5 m × 3.5 m) at three experimental saline sites in Punjab province of Pakistan over 5 years and assessed the impact of these planting densities on soil salinity amelioration. Tree response was assessed by measuring survival, height, and diameter at breast height (DBH) of trees at 1, 2, and 5 years as well as wood volume after 5 years. Wood volume per hectare, height, and DBH were greater at sites I and II than at site III. The 3.0 m × 3.0 m spacing resulted in taller trees, but DBH and wood volume were greater with the 3.5 m × 3.5 m spacing. The study confirmed that initial tree spacing or density has a significant impact on the subsequent height, DBH, and wood volume of E. camaldulensis plants. The study also showed that broad spacing could be a better option in saline environments. There was a definite reduction in soil salinity (electrical conductivity, ECe) at five soil depths after 5 years of tree growth at all sites and spacing treatments, with reductions varying from 46% to 47% at site I, 31% to 52% at site II, and 25% to 58% at site III. Soil ECewas generally higher in surface (0–15 cm) soils.

Beschreibung: Increases in the emission of greenhouse gases, particularly during the second half of the 20th century, have been associated with climate warming at the global scale. High latitude areas have been reported to be particularly sensitive to such changes, with significant impacts on plant phenology. The objectives of the present study were to (i) estimate changes in the flowering dates of 18 spring-flowering herbaceous plant species typical of the deciduous forests of eastern North America in three areas of eastern Canada (Gatineau–Ottawa, Montréal, and Québec) from 1900 to 2000 and (ii) associate these changes with those of annual and spring local temperatures. My results show a 2–6 days advance in flowering date over 100 years, depending on the region considered (corresponding to a ~2–3 days advance per 1 °C); these values are somewhat lower than those published in other studies, but still support the increasing body of literature on the effects of climate warming on plant phenology. Shifts in flowering phenology were particularly evident for Montréal, a large metropolitan region; this suggests that global climate warming, and its effects on plant phenology, may be exacerbated by local conditions, particularly those associated with large urban areas. Furthermore, species-specific responses to climate warming, as those presented here, might lead to significant changes in community composition and ecosystem functions.

Beschreibung: Episodic stand-replacing wildfire is a significant disturbance in mesic and moist Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco) forests of the Pacific Northwest. We studied 24 forest stands with known fire histories in the western Cascade Range in Oregon to evaluate long-term impacts of stand-replacing wildfire on carbon (C) and nitrogen (N) pools and dynamics within the forest floor (FF, Oe and Oa horizons) and the mineral soil (0–10 cm). Twelve of our stands burned approximately 150 years ago (“young”), and the other 12 burned approximately 550 years ago (“old”). Forest floor mean C and N pools were significantly greater in old stands than young stands (N pools: 1823 ± 132 kg·ha–1vs. 1450 ± 98 kg·ha–1; C pools: 62 980 ± 5403 kg·ha–1vs. 49 032 ± 2965 kg·ha–1, mean ± SE) as a result of significant differences in FF mass. Forest floor C and N concentrations and C/N ratios did not differ by time since fire, yet potential N mineralization rates were significantly higher in FF of old sites. Old and young mineral soils did not differ significantly in pools, concentrations, C/N ratios, or cycling rates. Our results suggest that C and N are sequestered in FF of Pacific Northwest Douglas-fir forests over long (∼400 year) intervals, but that shorter fire return intervals may prevent that accumulation.

Beschreibung: The decay resistance of black locust ( Robinia pseudoacacia L.) heartwood was evaluated on standardized test specimens taken from mature and juvenile wood in 25 trees distributed over five sites in Wallonia, Belgium. The percentage mass losses caused by the wood-destroying fungi Coriolus versicolor CTB 863 A and Coniophora puteana BAM Ebw.15 were evaluated according to Cen/ts 15083-1 (Cen/ts 15083-1. 2005. Durability of wood and wood-based products — determination of the natural durability of solid wood against wood-destroying fungi, test methods — Part 1: Basidiomycetes. European Comm. for Standardization, Brussels). Black locust is classified as a “very durable” wood (class 1), while the juvenile heartwood (located near the pith) is classified in classes 1 to 2. The statistical analysis does not reveal any significant differences among the sites, suggesting that the growth environment of the trees has no effect on their natural durability. However, statistical analysis shows very highly significant differences among trees and between the two radial positions in the tree, with mature wood being more resistant than juvenile wood. Using such a naturally very durable species could reduce reliance on preservative treatment and the use of tropical species generally recommended for outdoor woodwork.

Beschreibung: Caribou Poker Creeks Research Watershed is a boreal forest watershed in Interior Alaska that is susceptible to regional warming and permafrost thaw. We measured seasonal foliar N concentrations of the black spruce understory shrubs, Vaccinium uliginosum L., Vaccinium vitis-idaea L., and Ledum palustre L. on north- and south-facing sides of the watershed. We predicted that the shrubs would respond to small-scale changes in active layer and soil climate, and we expected similar responses according to growth strategy (evergreen or deciduous). Overall, foliar N in shrubs was higher on warmer, drier soils with deep active layers: +7.9% N in V. uliginosum, +11.1% N in V. vitis-idaea, and +9.4% N in L. palustre. Each shrub had species-specific foliar N patterns that could not be categorized by growth strategy and were not well explained by soil climate or active layer. Leaf mineral nutrition is influenced by multiple processes, and foliar N was best explained by the combination of environmental variables operating at the study site. For Caribou Poker Creeks Research Watershed, we can expect increased N status of the black spruce understory along with continued climate warming, but changes cannot be predicted based on growth strategy.

Beschreibung: Tree species can differ in their responses to resource availability during the critical phase of establishment, which could influence forest dynamics. In Mediterranean forests, most of the attention has focused on the effects of shade and summer drought on seedling survival, but little is known about the effect of autumn to spring rains on earlier stages of recruitment. A sowing experiment was set up along natural light and water gradients with three co-occurring oak species ( Quercus suber L. (cork oak), Quercus canariensis Willd. (Algerian oak), and Quercus pyrenaica Willd. (Pyrenean oak)) that show limited natural regeneration in southern Spain. Recruitment stages were monitored for 1 year. Models of seed germination, seedling emergence, and seedling survival as well as of overall recruitment patterns were developed as functions of light, soil moisture, and soil compaction. The influence of intraspecific variation in seed mass and emergence time were also tested. Excess soil water levels during the winter reduced germination and emergence and lengthened time to emergence (in waterlogged open areas), which in turn decreased seedling survival during the dry season. Seedlings from larger seeds were more likely to germinate and emerge. The results suggest that temporal and spatial variability of soil water content, mediated by emergence time and seed size, play a crucial role in the regeneration dynamics of Mediterranean oak forests.

Beschreibung: Perpendicular distance sampling (PDS) has emerged as a compelling alternative to line intersect sampling (LIS) for the inventory of forest fuels and other downed woody materials (DWM), particularly where the aggregate volume of DWM is of primary interest. This article develops a selection protocol and design-unbiased estimators for a new probability proportional-to-volume sampling strategy, termed line intersect distance sampling (LIDS). LIDS combines the distance sampling protocol of PDS with the transect sampling protocol of LIS and provides unbiased estimates of aggregate DWM volume from counts of selected logs or log fragments. Simulations indicate that LIDS along multidirectional (e.g., Y-shaped) transects should perform similarly to PDS in terms of sampling error; however, it remains unclear how LIDS and PDS compare with LIS, especially when interest is attached to multiple DWM population parameters. It is argued that LIDS will be most useful in reducing implementation errors, particularly detection errors, relative to PDS under limited visibility field conditions.

Beschreibung: This 2-year study examined the effect of fertilizers on tree growth and foliar nutrition in a Populus trichocarpa Torr. & A. Gray × Populus deltoides Bartr. ex Marsh. plantation located in southwestern Québec. The treatments included a control that did not receive N or P fertilizer, inorganic NP fertilizers, organic fertilizers applied at 65–70 kg N·ha–1, and organic fertilizers applied at 130–140 kg N·ha–1. Fertilized trees were taller and had larger diameters than control trees. Three methods were used to diagnose limiting nutrients and nutrient imbalances, and compare the nutrient supply from different fertilizer sources. The critical value approach and the compositional nutrient diagnosis methods found below-optimum N and P concentrations, sufficient K and Mg concentrations, and an excessive Ca concentration in foliage. Vector analysis compared the N nutrition in foliage from fertilized trees and the control trees. The compositional nutrient diagnosis r2(nutrient imbalance index) was negatively correlated with annual tree growth in height (r = –0.46, P 〈 0.05) and diameter (r = –0.59, P 〈 0.05), meaning that trees with a greater nutrient imbalance grew less in height and diameter than trees with balanced foliar nutrition. Of these diagnostic methods, compositional nutrient diagnosis holds promise for identifying nutrient limitations and predicting growth responses to fertilization in hybrid poplar plantations.

Beschreibung: Genetic diversity within populations is an important component of adaptive evolution, and recent research has demonstrated that genetic variation within plant populations can have important ecological effects. In this study, we investigate quantitative-genetic variation in several traits within a quaking aspen ( Populus tremuloides Michx.) population. A common garden experiment was planted with replicates of 13 aspen genotypes collected from wet and dry sites within a population in southern Utah, USA. Ten growth, leaf, physiological, and structural traits were measured. There were significant, heritable phenotypic differences among genotypes in every measured trait and differences in 4 of the 10 traits among genotypes originating from wet and dry collection sites. The data were compared with other published studies, showing that aspen heritability (H2) estimates and coefficients of genetic variation (CVG) were comparable or higher than other Populus species and hybrid F1 Populus genotypes, indicating a large amount of quantitative-genetic variation in aspen.

Beschreibung: Down woody debris (DWD) plays a vital role in forest ecosystem structure and function. Although volume is likely the most common metric used to characterize DWD, an evaluation of the formulae used for volume estimation on individual DWD pieces has received little attention. We determined actual volume of 155 diverse DWD pieces (types, species, lengths, and diameters) by detailed field measurements. By comparing the actual and calculated volumes from six commonly used formulae, we assessed their bias, precision, and accuracy. Based on observed DWD forms, we developed a new formula, namely the “conic−paraboloid”, which was included in the assessment. Among the formulae that require length and two end diameter measurements, the conic−paraboloid had the lowest bias, highest precision, and hence greatest accuracy. Newton’s and the centroid formulae had higher accuracy yet require more field measurements. Smalian’s, conical frustum, and average-of-ends formulae had poor performance relative to the others. Accuracy of all formulae decreased with increasing piece length. Thus, partitioning pieces into two, three, and four sections for additional measurement improved accuracy. As decay advances, pieces become progressively more elliptical in cross section. Using the cross-sectional area derived from only the long axis of the ellipse leads to substantial volume overestimates for well-decayed DWD.

Beschreibung: Innovativeness can help companies differentiate themselves, with the ultimate goal of securing survival and improving performance. Modern theories in organizational behavior look at innovation as something that starts with individual creativity but that is also affected by the work environment. Using one broad industry sector, the US forest products industry, this study attempts to integrate into a unifying model the concepts of work climate, innovativeness, and firm performance using structural equation modeling. Results support the proposed theoretical model, with some modifications, finding a positive and significant relationship among all factors. Having innovation as a core part of a company’s strategy and fostering a climate for innovation positively affects the degree of innovativeness and performance of a company. This is especially true for secondary or value-added wood products manufacturers. A climate for innovation is characterized by high levels of autonomy and encouragement, team cohesion, openness to change and risk taking, and sufficient resources available to people. Lack of a validation sample suggests treating the model as tentative until further testing.

Beschreibung: Rising societal demands for forest resources along with existing natural disturbance regimes suggest that sustainable forest management will increasingly depend on better understanding the cumulative effects of natural and anthropogenic disturbances. In North America, for example, there is increasing economic pressure to salvage log burned forests, although the ecological consequences of combining fire and harvesting on the same sites are unclear. We examined the short-term (2 year) responses of boreal forest ground beetles (Coleoptera: Carabidae) to the individual and combined effects of wildfire, harvesting, and herbicide. Ground beetle responses to wildfire and forestry-related disturbances differed strongly and suggested that, although some species may appear to benefit from disturbance combinations (e.g., Sericoda quadripunctata (DeGeer)), these effects are detrimental to others (e.g., Sericoda bembidioides Kirby). Species compositional variability was significantly reduced by disturbance combinations suggesting that multiple disturbances may lead to a simplification of this entire assemblage. In addition, ground beetle responses were correlated with changes in several key habitat parameters such as amount of woody debris, exposed ground, and plant species richness suggesting avenues for future study. Overall, however, our results suggest that efforts to avoid compounding disturbances on any site should be considered when developing current and future forest management guidelines.

Beschreibung: The complex fluxes between standing and harvested carbon stocks, and the linkage between harvested biomass and fossil fuel substitution, call for a holistic, system-wide analysis in a life-cycle perspective to evaluate the impacts of forest management and forest product use on carbon balances. We have analysed the net carbon emission under alternative forest management strategies and product uses, considering the carbon fluxes and stocks associated with tree biomass, soils, and forest products. Simulations were made using three Norway spruce ( Picea abies (L.) Karst.) forest management regimes (traditional, intensive management, and intensive fertilization), three slash management practices (no removal, removal, and removal with stumps), two forest product uses (construction material and biofuel), and two reference fossil fuels (coal and natural gas). The greatest reduction of net carbon emission occurred when the forest was fertilized, slash and stumps were harvested, wood was used as construction material, and the reference fossil fuel was coal. The lowest reduction occurred with a traditional forest management, forest residues retained on site, and harvested biomass was used as biofuel to replace natural gas. Product use had the greatest impact on net carbon emission, whereas forest management regime, reference fossil fuel, and forest residue usage as biofuel were less significant.

Beschreibung: We investigated three primary causes of old-growth forest pedodiversity imposed by top-down trophic interactions, including pit and mound topography from past tree fall events, current canopy gaps from tree falls, and the influence of individual tree species on soil properties and processes. In this paper, we discuss the effects of pits, mounds, gaps, and individual tree species on pedodiversity in a single soil map unit in an old-growth northern hardwood forest. Pits and level areas had significantly greater soil organic matter, cation-exchange capacity, and exchangeable K and Ca contents than mounds. Gap subplots had significantly less cation-exchange capacity, K, Mg, and Ca compared with level areas within the contiguous forest. Base cations (K, Mg, and Ca) were significantly greater under sugar maple ( Acer saccharum Marsh.) compared with eastern hemlock ( Tsuga canadensis (L.) Carr.). Extractable P was significantly greater under yellow birch ( Betula alleghaniensis Britt.) compared with eastern hemlock. We quantified pedodiversity in an old-growth northern hardwood forest stand and single soil map unit using principal components analyses, ArcGIS, and biodiversity indices. Our results suggest that pedodiversity should be considered in soil survey and forest management.

Beschreibung: Partial cutting is increasingly applied in European temperate oak-dominated forests for biofuel harvesting, and to counteract succession in protected stands. Effects on biodiversity of these measures need to be carefully evaluated, and species-rich but neglected taxa such as fungi should be considered. We studied the effects of partial cutting on fungal fruiting bodies on woody debris. In 21 closed canopy forests rich in large oaks in Sweden, on average 25%–30% of the basal area was cut. Fruiting bodies were counted and some were collected in treated and control plots before and after treatment. We found 334 basidiomycete and 47 ascomycete species. Species richness of basidiomycetes declined significantly more in treated plots (on average 26%) than in control plots (on average 13%) between seasons. Species richness of ascomycetes increased by 17% in control plots and decreased by 2% in treated plots. Total species richness was significantly reduced on fine woody debris (1–10 cm in diameter), but not on coarse woody debris (〉10 cm). Overall species composition did not change significantly as a result of partial cutting, but red-listed species tended to decrease more in treated plots. We suggest that approximately 30% of the stands should not be thinned, and dead stems and fallen branches should not be removed, to favor saproxylic fungi and their associated fauna.

Beschreibung: We designed and developed an internet mapping application to collect data on the locations of forest landscape values across a 2.4 million hectare study area in the province of Alberta, Canada. Four communities in the study area were surveyed and 8053 point locations were mapped for 10 different value types. Importance weights of landscape values were determined through a ranking exercise. Nearest-neighbour and second-order spatial point pattern analysis (K functions) suggested that all value types were significantly clustered across the study area. Recreational, wilderness, existence, and biological diversity values exhibited maximum clustering at larger spatial scales in comparison with educational, economic, historic or cultural, and spiritual values. Maximum clustering was positively related to mean road density and negatively related to mean distance to water, which suggests that landscape features influence the spatial pattern of values by acting as focal points or attractors for values. An applied use of the data for values hotspot detection and community protection zoning in forest fire management planning is presented.

Beschreibung: European aspen ( Populus tremula L.) is a keystone species for biodiversity in boreal forests. However, large aspen have largely been removed from managed forests, whereas regeneration and the long-term persistence of mature trees in protected areas are matters of concern. We recorded the numbers of mature (≥20 cm diameter) aspen in old-growth and managed forests in eastern Finland, based on a large-scale inventory (11 400 ha, 36 000 living and dead trees). In addition, saplings and small aspen trees were surveyed on thirty-six 1 ha sample plots. The average volumes of mature living and dead aspen were 4.0 and 1.3 m3/ha in continuous old-growth forests and 0.2 and 0.6 m3/ha in managed forests, respectively. These results indicate that large aspen trees in managed forests are a legacy of the past, when forest landscapes were less intensively managed. We conclude that the long-term persistence of aspen in protected areas can only be secured by means of restoration measures that create gaps large enough for regeneration to occur. More emphasis should be given to sparing aspen during thinning and to retaining mature aspen during regeneration cutting in managed forests.

Beschreibung: Severe droughts have the potential of inducing transient shifts in forest canopy composition by altering species-specific adult tree mortality patterns. However, permanent vegetation change will occur only if tree recruitment patterns are also affected. Here, we analyze how a massive mortality event triggered by the 1998–1999 drought affected adult and sapling mortality and recruitment in a mixed Nothofagus dombeyi (Mirb.) Blume – Austrocedrus chilensis (D. Don) Flor. et Boult. forests of northern Patagonia. Comparing drought-induced and tree-fall gaps, we assessed changes in forest composition, microenvironments, and seedling density and survival of both species. Drought-kill disturbance shifted species composition of both canopy and sapling cohorts in favour of A. chilensis. Drought gaps were characterized by a shadier and more xeric environment, affecting the recruitment pattern of N. dombeyi seedlings. The seedling cohort was composed mostly of A. chilensis, and its survival was always higher than that of N. dombeyi. Additionally, A. chilensis seedlings showed higher plasticity than N. dombeyi seedlings, increasing its root to shoot ratios in drought gaps. The results suggest that extreme drought itself is a strong driving force in forest dynamics, with important imprints on forest landscapes. Future climate-change scenarios, projecting an increased in frequency and severity of droughts, alert us about expected long-term compositional shifts in many forest ecosystems.

Beschreibung: Survival of live retention trees is a key issue for stand-scale applications of natural-disturbance-based silviculture. We explored the survival of 3255 trees in 102 cut areas (mean size 2.3 ha) in Estonia for 6 years, focusing on spatial variation and preadaptation of the trees. Altogether, 35% of the trees died during the study period, contributing 4.4 m3 of downed dead trunks and 1 m3 of standing dead trees per hectare. The annual mortality rates declined over time. The main survival determinants were tree species (higher for hardwood deciduous trees), diameter (species dependent), position relative to forest edge (higher for trees near current or former forest edges), retention density (positive), and exposure (negative). The results suggest that (1) green-tree retention can effectively increase the abundance of large shade-tolerant trees, but it is equally important for producing deadwood; (2) larger individuals, former interior-forest trees near current forest edges, and preadapted trees in open conditions should be preferably retained; (3) there is no obvious necessity to modify tree-retention techniques for tree survival according to geographical region or forest site type.

Beschreibung: The effect of thinning on the relationship of wood quality traits measured on standing trees (dynamic modulus of elasticity (MOE) and outerwood density) and traits measured on logs or short clear specimens was determined using data collected from radiata pine ( Pinus radiata D. Don) trees growing in 22 unthinned and 16 thinned plots of harvest age trees in New South Wales, Australia. Stiffness showed a linear decrease along the stem. Trees growing on thinned sites were, on average, 3% lower in stiffness at each height in the stem. MOE measured on short clear specimens was moderately related to standing tree MOE (R2 = 0.62) and outerwood density (R2 = 0.56) but less well related to MOE of the adjacent log (R2 = 0.30). Standing tree MOE was a better predictor of whole stem MOE for the thinned sites (R2 = 0.60) than for the unthinned sites (R2 = 0.31). Stiffness and density appear to follow different patterns of variation and results for density may not be extrapolated to stiffness. Outerwood density was a very poor predictor of mean whole stem stiffness (R2 = 0.14). Overall, the acoustic tool, TreeTap, was a better predictor of whole stem stiffness than outerwood density, particularly for the thinned sites.

Beschreibung: Following release, mature trees can acclimatively shift aboveground growth allocation from height to diameter at breast height (DBH) growth to adjust their stem form (reducing height/diameter ratio (HDR)) to better withstand increased wind stress. The purpose of this study was to determine if, over a 6-year period, 7- to 12-year-old eastern white pines ( Pinus strobus L.) also responded to different levels of partial release with (i) stem-form adjustments (reduced HDR) through growth-allocation shifts from height to DBH growth and (ii) lower HDR values (shorter heights) than for unreleased trees of the same DBH (“acclimative stem-form development hypothesis”). Over the 6-year postrelease period, juvenile white pine seedlings exhibited unsynchronized height and DBH growth response patterns that depended on their prerelease height growth. Height growth of faster growing white pines was temporarily reduced. Concurrently, DBH growth was enhanced with increasing release intensity. HDR reductions followed a fairly narrow and predictable trajectory, but HDR values of released trees were not lower than those of unreleased trees of the same DBH after 6 years. Juvenile white pine seedlings appear to maintain a balance among aboveground tree parts to enhance future tree stability, which may be an adaptive trait for moderately shade-tolerant species.