ALBERT

Description: Twelve species of terricolous microlichens from the Angmagssalik District, Southeast Greenland, are reported: Caloplaca friesii, C. livida, Lecanora boligera, Lecidea oligotropha and Leciophysma arctophila, which are new to the lichen flora of Greenland, Rinodina conradi, which is new to the eastcoast, and Baeomyces roseus, B. rufus, Buellia geophila, B. punctata, Caloplaca tornoensis and Mycoblastus tornoensis, new to Southeast Greenland. In a discussion of the greenlandic distribution, unpublished records from the herbarium of Copenhagen (C) are incorporated. Notes on the habitats are given and the pertinent phytosociological units indicated. Some morphological and anatomical characters are commented upon briefly.

Description: The 16 recognized species of Sorocea are listed with their synonyms and distribution. Two new taxa are described: S. steinbachii C.C. Berg and S. hirtella Mildbread ssp. oligotricha Akkermans & Berg. Three new combinations are made: S. muriculata Miquel ssp. uaupensis (Baillon) C.C. Berg, S. trophoides W. Burger ssp. rhodorachis (Cuatrecasas) C.C. Berg, and S. sprucei (Baillon) Macbride ssp. saxicola (Hassler) C.C. Berg. A key to the species is presented.

Description: The species Polypodium banaense C.Chr. is transferred to Crypsinus. The recognition of a genus Phymatopteris Pic. Ser. (= Phymatopsis J.Sm.) separate from Crypsinus is discussed.

Description: The six species of Curtia, including a hitherto undescribed species published here, as well as the monotypic genus Hockinia can be distinguished from each other by the seed coat structure. The anticlinal walls and the cuticle provide the most useful information. Curtia tenuifolia appears to be a complex species, but subsp. tenella can be readily separated from this complex by the seed coat structure. Heterostyly has been found in C. tenuifolia subsp. tenuifolia, C. obtusifolia, and Hockinia montana, but differences in seed coat structure can not be correlated with long-, short-, and equal-styled flowers. The differences in seed coat structure, the length of the seeds, and the number of cells per seed plead for maintaining Hockinia (closely related to Curtia) as a distinct genus. One new species of Curtia is described and a new combination is made.

Description: SETTEN, A. K. van & KOEK-NOORMAN, J.: Studies in Annonaceae. VI. A leafanatomical survey of genera of Annonaceae in the Neotropics. — Bot. Jahrb. Syst. 108: 17—50. 1986. — ISSN 0006-8152. Within the scope of the multidisciplinary research project on systematics of Annonaceae, a survey of the leafanatomical features and their distribution in the neotropical Annonaceae is presented. The studied specimens form a rather homogeneous group, as may appear from the family description given here. A detailed study of the leafanatomical features reveals, that differences are mainly found in the indument, the position and contents of the idioblasts, the structure of the primary vein, the type of crystals in the epidermal cells, and the type of sclereids. Based on character states, phenetic similarities and differences are discussed and compared with the classifications of FRIES (1959) and WALKER (1971).

Description: One new species of Dorstenia from Brazil is described: D. carautae C.C. Berg, and four new combinations are made: D. cayapia Vellozo subsp. asaroides (Hooker) C.C. Berg, D. cayapia Vellozo subsp. paraguariensis (Hassler) C.C. Berg, D. cayapia Vellozo subsp. vitifolia (Gardner) C.C. Berg, and D. ramosa (Desvaux) Carauta, Valente & Sucre subsp. dolichocaula (Pilger) C.C. Berg. A list of and a key to the 22 Dorstenia species distinguished in south-eastern tropical America are presented, together with synonymy and distributional data.

Description: Recently a multidisciplinary investigation program on the systematics of Annonaceae was started at Utrecht with special emphasis on the Neotropics. This project will be carried out largely within the framework of the UNESCO-project Flora Neotronica. The first goal is to provide a modern classification of the family as a whole, the second is the publication of a series of monographs for Flora Neotropica. The project has been planned and started in close consultation with leading botanists on the Neotropical flora. The Annonaceae are a family of pantropical distribution with between 2000 and 2500 species in ca. 130 genera as presently understood. In the Neotropics the family is represented by ca. 750 species and 35 genera. It is a family of trees, shrubs, and lianas. Its place is within the order of the Magnoliales and its supposedly closest relative is the family of the Myristicaceae. The Annonaceae, although generally considered primitive in many features, nevertheless offer a number of specialized features as well This makes it a promising object using various kinds of comparative morphological, karyological, and anatomical data. Besides, many species are of medicinal or commercial value, such as various species of Annona and Rollinia, the fruits of which are commonly eaten in most countries of Central America and South America; the Soursop (Annona muricata) is widely cultivated throughout the tropics.

Description: A new species of Asterophorum, A. mennegae, is described from the Sipaliwini Savanne (Suriname). The position of Asterophorum within the family is briefly reviewed, and a key to the 2 species is given.

Description: The wood and leaf anatomy of representatives of the 9 genera of the Opiliaceae are described in detail. It is possible to separate the genera on the base of both wood- and leaf anatomical characters. Herein the presence of cystoliths of varying shape and size is important. Some comments on the taxonomy and possible phylogeny of the familiy are given.

Description: In his introductory statements to 'The Symposium on the Phylogeny and Classification of the Filicopsida' which was held in London, 1972, HOLTTUM, when dealing with 'dubious groups of relationships which would particularly repay investigation', mentioned the Polypodiaceae first (HOLTTUM, 1973: 6). Talking about Polypodiaceae the present authors deal with the Polypodiaceae sensu stricto only, thus excluding the Cheiropleuriaceae, Dipteridaceae, Grammitidaceae, and also the Loxogrammaceae, taxa which were formerly (or are still) included in the Polypodiaceae sensu lato. As delineated in this way, this almost exclusively pantropical family consists of about 600 species and an indefinite number of genera.

Description: The new species Coussapoa manuënsis C.C. Berg is described.

Description: IUCN, says the paper Categories, Objectives and Criteria for Protected Areas, ”is dedicated to the wise use of the Earth’s natural resources and to the maintenance of the Planet’s natural diversity.” What to think of the sequence? Use first, maintain second? And this comes from the International Union for the Conservation of Nature and Natural Resources? ”The World National Parks Congress, taking place in Bali, Indonesia, October 11-22, 1982, will provide case studies from around the world to illustrate how the various categories of protected areas are meeting the needs of countries of all economic, social, cultural, and political backgrounds,” writes J.A. McNeely, the secretary of the Commission on National Parks and Protected Areas, in a special issue of the Swedish journal Ambio (11: 237. 1982). ”No longer just playgrounds for vacationers and means for conserving natural heritage, protected areas have become an inseparable part of the modern human ecosystem.”

Description: Mrs. Delia D. Adefuin, Museum Research Assistant, Manila, is pursuing her M.S. in Botany degree. She is currently the Secretary of the Fern Society of the Philippines. She is working on the Fern Flora of Metro Manila and is preparing the manuscript of a pictorial encyclopedia which will include descriptions of species and horticultural recommendations. Miss Barbro Axelius (S) collected and studied Xanthophytum and Lerchea (Rubiaceae) in Sarawak, Kalimantan and Sumatra, August 1982- February 1983.

Description: Tropical Botany in Aberdeen University. This was started by Professor J.W.H. Trail, who held the chair from 1877 to 1919, and travelled in the Amazon Valley (1873-75) mainly collecting cryptogams and studying palms. He was succeeded by Prof. W.G. Craib (1920-33) who was never in the tropics but devoted his work to the Flora of Siam, based on the collections of A.F.G. Kerr, and assisted by Miss E.C. Barnett. After a considerable lag, tropical botany was revived by the energetic efforts of Dr. P. Ashton as lecturer in systematics and ecology of the eastern tropics, establishing ties with Malayan colleges in teaching and research. This is at present perpetuated by two lecturers, Dr. K. Jong and Dr. M.D. Swaine, the latter’s experience lying largely in the tropics of West Africa. In addition Dr. N.M. Pritchard, Dr. J.B. Kenworthy and Dr. G. Hadley have been on secondment to the University of Malaya, while Dr. I. Alexander made research visits to India, Ghana and Peru. Over the years the Department has provided undergraduate and research training to innumerable students from many different tropical countries, some of which attained responsible posts, e.g. Prof. E. Soepadmo. Important courses in tropical biology are given, not available elsewhere in the U.K. (started 1973). The benefits for Aberdeen students is important: amongst others they led to expeditions to various parts of the tropics, recently to Sabah and to the Ivory Coast. Royal Society Tropical Rain Forest Collaborative Research Programme. Arising out of a feasibility study by Dr. T.C. Whitmore and P.F. Cockburn, the theme ’Recovery of tropical rain forest after disturbance’ was adopted as the initial basis of the programme. Possible territories for the research include Sabah and the Philippines. Detailed plans for a 5-year project are being prepared in consultation with colleagues in Southeast Asia.

Description: Mr. N. GAPAS (PNH) has made a survey of the phytoplankton of Aklan Bay, the Philippines. Dr. R.J. KING with Mr. C.F. PUTTOCK (Univ. NSW) has finished a complete revision of the Bostrychioideae (Rhodomelaceae) on a worldwide basis which has been accepted for publication in the Austr. Syst. Bot.

Description: Evolution is a complex process and the species produced by evolutionary processes are therefore of necessity not always clear-cut. Problems associated with definition of species boundaries have always plagued taxonomists and, no doubt, always will. However, I would maintain that in most situations there really is a ‘best’ solution to what kind of taxonomic recognition to accord a specific evolutionary pattern. The optimum solution to these kinds of problems depends in part on taxonomic philosophy. Are species real objective evolutionary entities? (Our job being to find out what they are.) Or are they essentially artificial constructs whose delimitation more or less depends on taxonomic convenience and preference? In the latter case, there is no ‘solution’ to the taxonomists’ dilemma. Certainly, many people working in the temperate zone, where rampant autogamy means that every clone of Taraxacum is essentially a different microspecies, tend to feel that this is the case. However, many of us working in the Neotropics feel that species are mostly far better defined and are, for the most part, quite objective entities. Indeed, as I see it, if species are not in some sense real units whose nature and limits we are trying to discover via scientific thought processes, taxonomy would hardly be worth doing, and we should turn our efforts to some other field of endeavor, say, population genetics, where the scientific method does provide insight into the real world.

Description: The Forest Research Institute of Malaysia (FRIM) has started a project on the conservation of fruit trees. Collecting expeditions have been made in 1988 to the Kraun Game Reserve (c. 102° 30’ E, 3° 40’ N), Maxwell’s Hill, Endau-Rompin. Collections were not only made of fruit trees, but also of any other flowering or fruiting tree. On 16 January, 1988, Mr. A.Z. IBRAHIM and Dr. A.M. LATIFF (UKMB) collected c. 15 numbers on Gunung Pulai, Johor.

Description: It is with the utmost regret that we announce the sudden and quite unexpected death of Dr. Marius Jacobs, editor of this Bulletin. See the obituary on page 3869. He was co-editor of the Flora Malesiana Bulletin for nr. 17 (1962) to nr. 22 (1968) and took full responsibility onwards of nr. 27 (1974). He showed great ability in enlarging its scope and we have many letters in our archives expressing appreciation and admiration for the lively and informative style in which he edited the Bulletin. I had to take over the editorial work for this number at short notice, but I was greatly helped by a number of Rijksherbarium colleagues, which help is gratefully acknowledged. In this way the delay has been kept to a minimum. It is, however, possible that some news items etc. have not been printed and that information submitted to Dr. Jacobs has not been entered due to this sudden change of editorship. I offer my apologies if this has happened and hope that (if still relevant) the news will again be forwarded to the new editor, Dr. J.F. Veldkamp, Rijksherbarium, who will take over starting next number.

Description: The symbol of the Flora Malesiana is the Rafflesia, a picture of which adorns the front cover of this Bulletin. It is therefore with mixed feelings that we have heard the news that a new species was discovered in Sabah, but its type locality was nearly immediately destroyed by logging activities (Salleh & Latiff, 1989, and Chapter IX). Fortunately another one seems to have been discovered. This is but one example of many of the irreplaceable losses Nature suffers through the thoughtless activities of Man. The instigators of this instance are known, but will remain free to continue to make a buck. It is a strange world where fines are meted out to smugglers of protected species. One of these defended himself by saying that he had played such a positive role in nature conservation. Of course the judge disagreed, but considering the on-going loss of natural habitats by exploitation and pollution, the defendant did make a point to ponder. What is better: a species irrevocably gone, or one in a zoo or a botanical garden? In the history of the earth a score of periods of extinctions have taken place, the cause of which is a matter of speculation: comet impacts, the passing through galactic dust clouds, a mysterious sister-star, extensive ice ages that drained the seas? Most spectacular were the ones of the middle Perm (c. 250 My) and on the Cretaceous-Tertiary border (c. 65 My), where 90% and 75% of all species respectively disappeared.

Description: The Malayan Nature Society (MNS), a non-governmental organisation with about 3,000 members, has since its formation in 1940 always had a strong emphasis on conservation. The Society’s objective in mounting this expedition, a mammoth undertaking for any Society as it included raising funds to a tune of almost M $ 400,000 (about £ 80, 000), were several: 1. To gain permanent legal protection for the only area in Malaysia where there still is a viable population (estimated at between 20 and 25 animals) of the rare Sumatran rhinoceros. 2. To gain legal protection for one of the few remaining extensive areas of lowland forest left in the Southern part of Peninsular Malaysia. 3. To explore the area and to document the flora and fauna of this Southern forest type, which until then was relatively poorly known scientifically. 4. To increase the awareness among school children and the general public of the beauty and value of Malaysia’s natural heritage.

Description: The Botanical Survey of India continued to make collections during 1982, with the following results: Andaman & Nicobar Is.: Shola Bag, Mt Harriet, Jirkathang, Poona Nallah, Saddle peak, Diglipur, Rutland I. & Little I., 2875 specimens. Arunachal Pradesh: Various areas of Kameng Distr., Subansiri Distr., 9750 specimens. Assam: Garampani, 60 specimens. West Bengal: Darjeeling, Jalpaiguri, Hollong, Jaldapara, Chilapata, Salkumar, Daidaighat, Barasat, 2665 specimens. Bihar: Madhuban, Nimiaghat, Paresnath Hills, 315 specimens. Dehra Dun: Chakrata, Missoori, 325 specimens. Gujrat: Catchment and submergence areas of Sipu Reservoir Project, 1505 specimens. Kerala: Trichur, Idduki, Silent Valley, Valra Reserve Forest, Cannanore, Trivandrum, 3770 specimens. Madhya Pradesh: Kanha National Park, Chhodarpur Distr., 1190 specimens. Maharashtra: Areas of Jalgaon Distr. and Buldhana Distr., 4390 specimens. Manipur: Cherrapunjee, Mawphlong, Sorharim, areas of West Khasi Hills Districts, 2000 specimens. Rajasthan: Bharatpur, Desert National Park, 1605 specimens. Sikkim: Rangpo, Singtham, Bumbing, Manuring, Duga, Pandam, Takchi, Meli, areas of Gangtok, Chungtham, Lachi, Thanga, Panthang, 2590 specimens. Uttar Pradesh: Gori & Kali Valley, Chittoragarh Distr., 500 specimens.

Description: The ultrastructure of the spore walls of Beenakia dacostae (Beenakiaceae, Gomphales) has been studied. Spore walls are mainly composed of a distinct episporium and a thick, dark, ornamented ectosporium. The general structure is identical with that of other members of the Gomphales, such as Gomphus and Ramaria

Description: An illustrated revision of the 65 species of the genus Neonauclea (Rubiaceae-Naucleeae) with a key to all species. The 61 Malesian species are treated in full, 28 new species are described and 5 new combinations are made.

Description: In Malesia the genus Tephrosia is represented by 20 species, native or introduced and naturalized, including 6 subspecies and 5 varieties; 4 species are restricted to Malesia. Two of these species are newly described: T. barbatala and T. elliptica; the former includes one new variety (var. glabra). Two new subspecies and one new variety are distinguished: T. filipes subsp. longifolia, T. purpurea subsp. barbigera, and T. maculata var. elongata. Two taxa are given a new status: T. maculata var. appressepilosa and T. pumila subsp. aldabrensis. Four species, T. dichotoma, T. repentina, T. coarctata, and Kiesera sumatrana are for the first time sunk into synonymy. A key to the taxa, synonymy, and full descriptions with plates and distribution maps are given.

Description: A pollen morphological survey of the genus Guioa is presented. Based on a study of the harmomegathy in relation to morphological series, evolutionary trends are postulated: 1. wide ectoapertures → narrow ectoapertures 2. grain parasyncolporate → grain colporate 3. grain parasyncolporate → grain syncolporate 4. ornamentation rugulate → ornamentation psilate-imperforate. It is suggested that the most ‘primitive’ species of Guioa occur in the southeastern part of the distribution area of the genus. Guioa can be subdivided into four poorly delimited pollen morphological groups. Its affinities to other genera in the Cupanieae are briefly discussed. Pollen of the most ‘primitive’ group is similar to that of several ‘primitive’ genera (e.g. Arytera, Matayba, Molinaea, and Tina) in the tribe.

Description: During identification and study of the materials of the genus Blepharis Juss. in the Central National Herbarium (CAL) of the Botanical Survey of India the authors came across some specimens (Lawson 314) from Tavancore, which was identified as B. molluginifolia aff But critical study revealed that the typical molluginifolia is a different species from which the new taxon can be distinguished by several characters. The species is named after M.A. Lawson who collected the specimen for the first time.

Description: Panicum smithii is newly described from North Vietnam; P. amoenum Balansa is lectotypified.

Description: One new orchid, Trias bonaccordensis Sathish, is described from the Bonaccord forests of Trivandrum, Kerala State, India. Affinities with the related T. stocksii Benth. ex Hook, f., T. disciflora (Rolfe) Rolfe and T. nasuta (Reichb. f.) Stapf are discussed. One new combination, viz. T. crassifolia (Thw. ex Trimen) Sathish, is proposed for the Sri Lankan Bulbophyllum crassifolium. Operculum features of the genus are illustrated and a distribution map is supplied.

Description: One new species of Aristolochia, A. singalangensis, from Sumatra is described here. This is the only Malesian species to have the fruit dehiscing from the apex towards the base. Remarks are given for some Asiatic and Malesian species, all belonging to Aristolochia except one to Thottea, on their taxonomy, nomenclature, typification, characteristics for identification, relationship, distribution, etc. The phenomena and significance of aristolochiaceous plants-butterflies relationship have been discussed. Germinated pollen grains have been found in dehisced anthers of open flowers in both Aristolochia and Thottea. Pollination of the Aristolochiaceae has been concisely reviewed.

Description: There are only two genera of the Aristolochiaceae, Aristolochia and Thottea, so far known to occur in Malesia. In the course of a revision of this family for the Flora Malesiana, some new species of both genera have been found. Eight new ones of Thottea were published in a precursor on that genus (Blumea 27, 1981, 301-332, f. 1-72). There are four new species of Aristolochia from Borneo and one more new Thottea from Sumatra to be described here.

Description: Carpels develop centripetally as oblique slightly ascidiform structures provided with a seat-like support. The terminal mouth of the ascidiform primordium becomes the lateral cleft of the carpel. Solitary terminal carpels develop as lateral structures. The sections Tasmannia and Drimys of the genus Drimys differ by the varying degree of ascidiform development. Austrobaileya is like Drimys. The structure of the carpel margins with submarginal placentation may have evolved from a slit like that in Winteraceae. Allometric development of an oblique ascidiform structure may have formed a large basal ovary, and may thus have moved the stigmatic part apically. A possible cupular origin of carpels is discussed.

Description: Being engaged during several years with a revision of the grasses preserved in the Rijks-Herbarium at the University of Leyden, my attention was called to the group of the Stipeae, and especially to the very difficult genus of Aristida. After an exhaustive study of the literature, I thought it desirable to have a monograph of this genus, containing extensive keys for the determination of all the species hitherto known, and I resolved to prepare such a work. It has been my good fortune that I had at my disposal not only the valuable collections of the Rijks-Herbarium, but that by the courtesy of the directors of the great herbaria in Europe and in America, I could study many thousands of specimens, among them authentic specimens and types. So several years elapsed before the revision was finished. Before I am going to publish my work, it seemed desirable to prepare a preliminary paper on the subject, dealing with the literature studied and the results of the critical examination of the types, moreover the new species found in herbaria are included in this paper. To find easily the original description and the type specimen, I give in alphabetical order all the species and varieties hitherto described, no matter if they are accepted in my monograph as valid or not.

Description: Ascension Island (08° S 14°25’ W) occupies a very peculiar place in our current research project on the biological properties of ground waters of the the Mid-Atlantic islands: -- The island lies closer to the equator than any of the other Mid-Atlantic islands. -- The island lies almost on the Mid-Atlantic Ridge, and far remote from any other insular or continental region. -- The island’s subaerial part consists of very young volcanic outcrops (dated radiometrically at 1-2 My). Through these properties, the comparison of Ascension with the intensively studied Canary Islands (see the series of publications of our team under the collective title “Stygofauna of the Canary Islands”, parts 1 to 19) is of great importance for a better understanding of dispersal and/or vicariant processes in the evolution of insular groundwater organisms (or stygobionts as these are called scientifically).

Description: A monograph of the genus Guioa Cav. (Sapindaceae) is presented. 64 species are recognized; one species, Guioa subfalcata, is regarded as a dubious species. A general key and several regional keys provide access to the species. One species, Guioa glauca, is subdivided into two varieties. The infrageneric classification contains three levels: subgenera, sections, and subsections. The classification is based on a phylogenetic analysis of the genus. The leaf anatomy was studied besides the macromorphology. This resulted in 67 characters for a cladistic analysis. The outcome of the initial analysis was rejected for several reasons (chapter 11.6). An alteration of the method resulted in an acceptable cladogram: the genus was subdivided, based on several characters, into 5 groups, 4 of which were subordinate to each other. Each subgroup was analysed separately and contained a representative species for a higher group and an outgroup which was obtained from a lower group. The cladograms of the 5 subgroups were pieced together to form the accepted cladogram. The computer programs HENNIG86 and PAUP provided the best method for a cladistic analysis (Wagner algorithm combined with some kind of ‘branch and bound’-option). A historical biogeographic analysis was performed with two aims: a) to find the historical relationships between the different distribution areas and b) to test the generic cladogram by comparing it with cladograms of independent groups of plants and animals. Like with the genus, the areas were subdivided into several groups, which were analysed separately under assumption 0 via a parsimony method instead of a consensus method. The complete generalized areagram was later on pieced together. The partial generalized areagrams did not falsify the accepted cladogram of Guioa. In two areas, the Pacific and West Malesia, Guioa mainly showed a pattern of dispersal and not of vicariance. The computer program CAFCA (group- and component-compatibility) presented the best method for a historical biogeograpic analysis. A comparison between the presumed results of the classical, evolutionary school and the phylogenetic school showed that the results of the phylogenetic school had a higher information content in the sense of natural relations and that only they could be used for a historical biogeographic analysis. The ecological part contains data on the habitat of some Guioa species, a compilation of mainly literature data on floral biology of Sapindaceae in general and on fruit biology, data on seed germination and seedling morphology. In order to unravel the species complex ( Guioa rigidiuscula group) on New Guinea only an artificial technique could be used to separate the 14 species. These appear to be natural due to correlating characters and coherent, non-disjunct distributions. It is discussed that species complexes are mainly found in especially geologically dynamic areas and that the mode of speciation is then best explained by the model of punctuated equilibria.

Description: The present study deals with the systematics and taxonomy of the genus Lamium (Lamiaceae). The taxonomic revision is mainly based on the study of herbarium collections, and to a smaller degree on field observations and abstracts from literature. The research was done at the Rijksherbarium, Leyden, while many other Herbaria were shortly visited. As the genus Lamium was never before monographed, a short survey is given of the taxonomic history of Lamium, which goes back as far as Linnaeus (1753). In this short survey of the taxonomic history of the genus Lamium the general history of taxonomy and its changing emphasis on different geographical regions can be recognized. It is difficult to give a definition of Lamium in which it stands out from related genera by means of one or preferably two characters. As with so many genera, Lamium can only be defined polythetically: no single character suffices. An important character is the chromosome number: the base number x = 9 is rather uncommon in other genera of the Stachyioideae to which Lamium belongs. As morphological outline of the genus Lamium can be given: annual or perennial herbs, usually not higher than about ½ m, with cordate or reniform, shallow incised leaves; flowers in verticillasters in the axils of the uppermost (= floral) leaves; calyx campanulate with an oblique mouth and with subequal teeth; corolla 2-lipped, the tube straight or sigmoid, the lower lip usually with two rounded lateral lobes, each with one or two teeth; nutlets obovoid with a truncate apex and in cross-section trigonous with one convex and two flat sides. The genus is divided into 3 subgenera; one subgenus ( Lamium) 3 sections, of which one section (Amplexicaule) is divided into is new. A number of 162 names of Lamium species and a more or less equal number of names of infraspecific taxa were known. In this revision 16 Lamium species, 11 subspecies, and 13 varieties are accepted, and also one hybrid. 9 New combinations are made. Type specimens are recorded for all taxa and an identification key to all taxa is given. In all c. 10,000 collections were examined. The most important quantitative characters were measured in 2700 collections, representing specimens from the entire distribution area of individual taxa. To show the variability of these quantitative characters diagrams are added, as diagrams give more information than minimum, average and maximum values. The percentage of ripe fruits appears to vary strongly in herbarium specimens of Lamium: all species with a percentage of ripe fruits higher than 55% are annuals, and all species with a percentage of ripe fruits lower than 20% are perennials. Random studies in other genera led to the same result: the ripening of fruits of annual species takes place in a shorter period than the ripening of fruits of perennial species. Distribution maps of all Lamium taxa are given. The distribution of the genus Lamium may be characterized as temperate Eurasiatic (holarctic), but the centre of diversity is obviously found in the Irano-Turanian and the Mediterranean regions. The habitats of the taxa of the genus Lamium roughly can be listed under two categories: forests on the one hand and rocky mountain slopes/cultivated fields on the other. The differences in habitats and also in distribution areas were the starting point for an intuitive phylogenetic reconstruction of the genus Lamium, in which Lamium galeobdolon is included, in spite of a number of deviating morphological and chemical characters.

Description: The buthid scorpions of the genus Centruroides are widely distributed in the Antillean area. They are also the most common scorpions in the majority of these islands. Nevertheless they remained almost forgotten until recently (STAHNKE 1970; ARMAS 1976, 1977, 1980, 1981, 1982; FRANCKE & SISSOM 1980). For the purpose of this paper the Lesser Antilles are defined as the islands situated between the Virgin Islands and Trinidad & Tobago, according to BOND’S 1978 zoogeographical point of view.

Description: In der obersenonen Mastrichter Tuffkreide finden sich kleine Zähne, die durch ihre glatten Kauflächen und die Furchen an den Seiten des oberen Teiles an Kauplatten von Myliobatis erinneren, einen Rochentypus, der ein an durophage Lebensweise angepasstes Gebiss hat. Niemals findet man aber die für diese Familie so typische langgestreckte Form der Zahnplatten; die Zahnoberfläche hat immer rhombische Form. Dames hat eine ausführliche Beschreibung von diesen Zähnen gegeben, die er für Reste eines Cestracion-artigen Namen Rhombodus Binkhorsti Haies hielt, dem er den gab. Ich möchte hier nur noch einige kurze Bemerkungen hinzufügen. Die Abbildungen (fig. 1) zeigen den typischen rhombenförmigen Umriss der Kaufläche (d). Die durch eine in der Richtung der kurzen Diagonale verlaufende, tiefe Rinne in zwei Hälften geteilte Wurzel hat ebenfalls die Gestalt eines Rhombus (fig. 1, b, e). An der Grenze von Krone und Wurzel findet sich an der einen Seite eine Rinne, an der anderen Seite eine vorspringende Leiste (fig. 1 c). Zusammen mit den verticalen Furchen, mit denen die Seiten versehen sind, hat diese Leiste zur Verbindung der Zähne untereinander zu einem Mahlpflaster gedient. Neben dieser regelmässigen Form, die besonders den grösseren Zähnen eigen ist, fanden sich aber Exemplare, die eine Abweichung zeigen, indem nämlich entweder zwei Seiten eines spitzen Winkels des Rhomboïds länger sind wie die beiden anderen, oder das Rhomboïd unsymmetrisch zusammengepresst ist. Es scheint mir, dass dies nicht eine zufällige Variation ist, sondern dass wir gerade durch diese Eigentümlichkeit etwas mehr über die ganze Zusammenstellung des Gebisses erfahren können. Wie ich unten noch näher auseinandersetzen werden, muss man nämlich Rhombodus zu den durophagen Stachelrochen stellen. Bei diesen findet man sehr oft gerade die grössten Zähne in der Mitte des Kiefers. Wenn man nun die Zahl der Zahnreihen, wie es gewöhnlich bei den grosszähnigen Rochen der Fall ist Rhombodus-Unterkiefers zu 7 bis 9 annimmt, so könnte man das Gebiss eines auf eine Weise rekonstruieren, wie es fig. 3 A zeigt, (wobei die verschiedenen obengenannten Formen vorkommen). Es wäre wohl ein grosser Zufall wenn man noch einige Zähne im ursprünglichen Verband finden würde. Wenn einmal die knorpeligen Kiefer aufgelöst sind, bieten die Seitenfurchen nicht genug Festigkeit und fallen die einzelnen Zähne auseinander.

Description: Da die Originale der von Göppert aus dem Tertiär von Java beschriebenen Arten Piperites Hasskarlianus und Junghuhnites javanicus nicht mehr vorhanden sind, die vorliegenden Beschreibungen für eine Bestimmung aber nicht ausreichen, so sind sie aus der fossilen Flora Javas zu streichen. Das gilt auch von Miquelites elegans, dessen schlechte Erhaltung eine sichere Bestimmung unmöglich macht. Bredaea moroides dagegen ist ebenso wie Naucleoxylon spectabile Crié sowie ein bisher unbeschriebenes Kieselholz von Java eine Dipterocarpacee. Die Stücke werden beschrieben als Dipterocarpoxylon moroides, D. spectabile und D. Göpperti n. sp. Die Frage, ob es möglich ist, diese wie andere fossile Dipterocarpoxyla bestimmten rezenten Dipterocarpaceengattungen zuzuweisen, soll später erörtert werden. Frankfurt a/M. Geologisch-Paläontologisches Institut der Universität.

Description: During the Late Devonian, deposition in the Cantabrian Mountains was largely controlled by movements along faults. By way of intermitting subsidence of the area south of the Sabero-Gordón line and the connected progradation of the coast during the Frasnian and early Famennian, three regressive sequences were deposited. On account of these sequences the Nocedo Formation is divided into three units. Before the late Famennian transgression, after which the upper part of the Ermita Formation was deposited, the area was peneplained. Six facies maps show the changes in palaeogeography during the Late Devonian.

Description: The area described comprises the SE plunging extension of the anticlinorium of Mondoñedo-Lugo-Sarria and several structural units to the NE of it. The stratigraphy compares with that of neighbouring areas. Certain new observations have been made regarding the west flank of the East Galician-West Asturian miogeosyncline at the end of the Precambrium and during the Early Palaeozoic. Stromatolites or algae played an important part in the formation of carbonate deposits during late Precambrian (Cándana limestone) and Lower Cambrian time (Vegadeo limestone). An imprint, possibly attributable to a specimen belonging to the Precambrian Ediacara fauna, was found in the Cándana Schist Formation. A shallow or relatively high zone existed during that time in the region around Incio. The same general area formed a high again at the end of the Ordovician, causing erosion and a marked disconformity of the Silurian. The Hercynian structures can be explained by assuming roughly WSW-ENE compression and shortening. In the NW of the area this is expressed in the first place by the recumbent folds of the Lugo-Sarria anticlinorium, and in the SE by the narrowness of the steep folds and their slightly changed direction in the general area of Seoane and El Cebrero. Normal crossfaults are related to the folding and the compression. They show locally a deviation in accordance with the direction of shear stress. Horizontal displacement along such faults in an EW direction was probably of little importance. Finally a few observations are given on the occurrence of Pb-Zn ores at the mine of Rubiales.

Description: Species of the heteropod families Carinariidae and Pterotracheidae collected in the Mid North Atlantic Ocean in 1980 have been studied. The distribution of the species is given, as well as the morphological variation. It is concluded that two subspecies of Carinaria lamarcki actually have to be considered distinctly recognisable and sympatric species, to be called C. lamarcki and C. challengeri. The vertical distribution of the heteropods studied proves not to be restricted to the photic zone and diurnal vertical migration occurs among the larger species. The horizontal distribution of the two Carinaria species and of at least two populations of the Pterotrachea species coincides with the southern branch of the North Atlantic Current, while others are restricted to the subtropical waters. Consequently, the present heteropods have to be considered good indicators of water masses and currents.