ALBERT

Description: Studies on the Mediterranean Undercurrent in the Gulf of Cádiz showed that bacterial abundance and biomass as well as heterotrophic activity were higher in the water of Mediterranean origin in 500–800 m depth than in the adjacent Atlantic water. Upwelling processes off Mauretania and Portugal were accompanied by high bacterial numbers (bacterial plate counts) in the mixed surface layer. Changes in the qualitative composition of the bacterial flora in the waters off West Africa and in the Arabian Gulf were explained by the introduction of dust from desert regions into the sea by aeolian transport. In the Western Baltic migration of fish was detected by the presence of special bacteria, which normally live on or in these animals. Regions with complex hydrographic structures such as the Western and Central Baltic Sea revealed interesting relationships between bacteriological abundance and activity on the one hand and characteristic physical and chemical properties, such as origin, salinity and O2/H2S‐content, on the other. The importance of bacteriological variables for the characterization of different water bodies is discussed.

Description: In this study a scenario is developed of two adjacent Mediterranean Water eddies (meddies) as they were observed merging and drifting through the Iberian Basin. Observations are based on four RAFOS floats (at 850–1050 dbar), two hydrographic surveys (centered roughly at 38°N, 24°W), and trajectories of surface drifters (drogued at 100 m). In April 1991, the meddy A was identified and labeled by surface drifters. During the revisit one month later two meddies were encountered, B1 and B2, in the vicinity of the former meddy A. The coalescence of B1 (subsequently identified as A, one month older) and B2 is inferred from a simple kinematic model describing the observed movement of the RAFOS floats for up to three months after the second CTD survey. The deduced vorticity front, radius ∼15 km, within B1 was of insufficient strength to keep the core waters of B1 isolated and prevent the absorption of B1 by B2. The resulting meddy (B1 + B2) showed a clear near-surface dynamical signal. Its deep root (1800 m) could explain the expulsion from the meddy of the remaining RAFOS float and surface drifter at the time of the meddy's collision with the Josephine Seamount. For the first time, a set of Lagrangian and hydrographic observations give direct evidence that neighboring meddies can merge as predicted by theoretical considerations.

Description: Global mean and eddy fields from a four-year experiment with a 1/6° × 1/5° horizontal resolution implementation of the CME North Atlantic model are presented. The time-averaged wind-driven and thermohaline circulation in the model is compared to the results of a 1/3° × 2/5° model run in very similar configuration. In general, the higher resolution results are found to confirm that the resolution of previous CME experiments is sufficient to describe many features of the large-scale circulation and water mass distribution quite well. While the increased resolution does not lead to large changes in the mean flow patterns, the variability in the model is enhanced significantly. On the other hand, however, not all aspects of the circulation have improved with resolution. The Azores Current Frontal Zone with its variability in the eastern basin is still represented very poorly. Particular attention is also directed toward the unrealistic stationary anticyclones north of Cape Hatteras and in the Gulf of Mexico.

Description: To avoid an explicit simulation of the overflows across the Greenland-Scotland ridge, many models of the large-scale ocean circulation seek to include the net effect of the inflowing dense water masses by restoring temperature and salinity near the ridge to observed conditions. In this paper the authors examine the effect of different datasets for the northern restoring condition in two versions, eddy resolving and non-eddy resolving, of the model of the North and equatorial Atlantic that has been developed in recent years as a Community Modeling Effort for WOCE. It is shown that the use of smoothed climatological fields of temperature and salinity south of the Denmark Strait leads to strong deficiencies in the simulation of the deep flow field in the basin. A switch to actual hydrographic data from the Denmark Strait ignites a rapid dynamic response throughout the North Atlantic, affecting the transport and vertical structure of the deep western boundary current and, by virtue of the JEBAR efffect, the transport of the horizontal gyres. Meridional overturning and northward heat transport too weak in the cases with climatological boundary conditions, increase to more realistic levels in the subtropical North Atlantic. The initial response to switches in the high-latitude thermohaline forcing is mediated by fast waves along the westurn boundary, leading to changes in the deep western boundary current in low latitudes after about two years in the non-eddy-resolving cast. The initial timescale depends on the horizontal grid spacing of the model; in the high-resolution case, the first signal reaches the equator in a few months. The adjustment to a new, dynamic quasi equilibrium involves Kelvin waves along the equator and Rossby wave in the interior and is attained in less than two decades throughout the North Atlantic. It is suggested that these fast dynamic adjustment processes could play an important role in possible fluctuations of the thermohaline circulation, or transitions between different equilibrium states of the coupled ocean–atmosphere system, and may have determined the timescale of the observed climatic transitions before and during the last deglaciation.

Description: We have investigated the seasonal cycle and the interannual variability of the tropical Indian Ocean circulation and the Indian summer monsoon simulated by a coupled ocean-atmosphere general circulation model in a 26- year integration. Although the model exhibits significant climate drift, overall, the coupled GCM simulates realistically the seasonal changes in the tropical Indian Ocean and the onset and evolution of the Indian summer monsoon. The amplitudes of the seasonal changes, however, are underestimated. The coupled GCM also simulates considerable interannual variability in the tropical Indian Ocean circulation, which is partly related to the El Niño/Southern Oscillation phenomenon and the associated changes in the Walker circulation. Changes in the surface wind stress appear to be crucial in forcing interannual variations in the Indian Ocean SST. As in the Pacific Ocean, the net surface beat flux acts as a negative feedback on the SST anomalies. The interannual variability in monsoon rainfall, simulated by the coupled GCM, is only about half as strong as observed. The reason for this is that the simulated interannual variability in the Indian monsoon appears to be related to internal processes within the atmosphere only. In contrast, an investigation based on observations shows a clear lead-lag relationship between interannual variations in the monsoon rainfall and tropical Pacific SST anomalies. Furthermore, the atmospheric GCM also fails to reproduce this lead-lag relationship between monsoon rainfall and tropical Pacific SST when run in a stand-alone integration with observed SSTs prescribed during the period 1970–1988. These results indicate that important physical processes relating tropical Pacific SST to Indian monsoon rainfall are not adequately modeled in our atmospheric GCM. Monsoon rainfall predictions appear therefore premature.

Description: The cause of decadal climate variability over the North Pacific Ocean and North America is investigated by the analysis of data from a multidecadal integration with a state-of-the-art coupled ocean-atmosphere model and observations. About one-third of the low-frequency climate variability in the region of interest can be attributed to a cycle involving unstable air-sea interactions between the subtropical gyre circulation in the North Pacific and the Aleutian low-pressure system. The existence of this cycle provides a basis for long-range climate forecasting over the western United States at decadal time scales.

Description: The annual cycle of meridional heat transport in the North and equatorial Atlantic Ocean is studied by means of the high-resolution numerical model that had been developed in recent years as a Community Modeling Effort for the World Ocean Circulation Experiment. Similar to previous model studies, there is a winter maximum in northward heat transport in the equatorial Atlantic and a summer maximum in midlatitudes. The seasonal variation in heat transport in the equatorial Atlantic, with a maximum near 8°N, is associated with the out-of-phase changes in heat content to the north and south of that latitude in connection with the seasonal reversal of the North Equatorial Countercurrent. The amplitude of the heat transport variation at 8°N depends on model resolution: forcing with the monthly mean wind stresses of Hellerman–Rosenstein (HR) gives an annual range of 2.1 PW in the case of a 1/3° meridional grid, and 1.7 PW in the case of a 1° grid, compared to 1.4 PW in a previous 2° model. Forcing with the wind stresses of Isemer–Hasse (IH) gives 2.5 PW in the 1/3° and 2.2 PW in the 1° model case. The annual range of heat transport in the subtropical North Atlantic is much less dependent on resolution but sensitive to the wind stress: it increases from 0.5 PW in the case of HR forcing to almost 0.8 PW with IH forcing. The annual cycle of heat transport can be understood in terms of wind-driven variations in the meridional overturning; variations in horizontal gyre transport have only little effect both in the equatorial and in the subtropical Atlantic. In all model solutions the seasonal variations in the near-surface meridional Ekman transport are associated with deep seasonal overturning cells. The weak shear of the deep response suggests that the large variations in heat transport on seasonal and shorter time scales should be of little consequence for observational estimates of mean oceanic heat transports relying on one-time hydrographic surveys.

Description: Cicosal sea surface height (SSH) data in the tropical and midlatitude North Atlantic are analyzed with and without water vapor (WV) correction to study the WV influence on along-track SSH anomaly profits, mesoscale SSH variability, wavenumber spectra, and objectively mapped fields of SSH anomaly. Three different WV datasets were used, one from the Fleet Numerical Oceanographic Center (FNOC) model and two from the Special Sensor Microwave/Imager (SSM/I) based on different WV retrieval algorithms. These WV dataset show significant differences, in particular in the tropics. However, the method for deriving SSH anomalies from altimeter height data Alters out much of the WV corrections. The residual WV effect on SSH anomaly is shown to be most significant in the seasonally migrating intertropical convergence zone of the tropical Atlantic: there the SSM/I corrections reduce the along-track mesoscale SSH variability by typically 1–1.5 cm. On seasonal timescales the maximum WV effect in this region is characterized by a 2–3-cm rms difference between SSH anomaly with and without SSM/I WV corrections, whereas FNOC corrections have almost no effect. Inferred seasonal velocity variations in the North Equatorial Countercurrent core (4° – 6°N) in the region of maximum WY influence (30° – 40°W) are reduced by about 20% and 30%, depending on whether SSM/I corrections by Emery or Wentz are used

Description: Early Tertiary lithospheric breakup between Eurasia and Greenland was accompanied by a transient (∼3 m.y.) igneous event emplacing both the onshore flood basalts of the North Atlantic Volcanic Province (NAVP) and huge extrusive complexes along the continent‐ocean transition on the rifted continental margins. Seismic data show that volcanic margins extend 〉2600 km along the early Eocene plate boundary, in places underlain by high‐velocity (7.2–7.7 km/s) lower crustal bodies. Quantitative calculations of NAVP dimensions, considered minimum estimates, reveal an areal extent of 1.3×106 km2 and a volume of flood basalts of 1.8×106 km3, yielding a mean eruption rate of 0.6 km3/yr or 2.4 km3/yr if two‐thirds of the basalts were emplaced within 0.5 m.y. The total crustal volume is 6.6×106 km3, resulting in a mean crustal accretion rate of 2.2 km3/yr. Thus NAVP ranks among the world's larger igneous provinces if the volcanic margins are considered. The velocity structure of the expanded crust seaward of the continent‐ocean boundary differs from standard oceanic and continental crustal models. Based on seismic velocities this “volcanic margin” crust can be divided into three units of which the upper unit corresponds to basaltic extrusives. The regionally consistent velocity structure and geometry of the crustal units suggest that the expanded crust, including the high‐velocity lower crust which extends some distance landward of the continent‐ocean boundary, was emplaced during and subsequent to breakup. The volcanic margin crust was formed by excess melting within a wide zone of asthenospheric upwelling, probably reflecting the interaction of a mantle plume and a lithosphere already extending.

Description: Observations of upper-ocean western boundary current (WBC) transports reveal asymmetries between the Northern and the Southern Hemispheres of the Atlantic Ocean. To find out what mechanism might cause these asymmetries the linearized steady-state vorticity equation is applied to the interior of a layer of constant thickness representing the upper Atlantic Ocean. WBC transports are then required to balance the interior volume flux deficit. The ocean is forced by climatological wind stress at the surface; thermohaline forcing is introduced by vertical motion at the lower boundary. A series of model runs using selected combinations of different basin geometries, wind stress fields, and thermohaline forcing patterns yields the following results: asymmetries of WBC transports cannot be explained by the topography shape of coastlines. The wind stress causes 12 Sv (Sv ≡ 1 × 106 m3 s −1) cross-equatorial transport to the north but it cannot account for the other WBC asymmetries. These can be explained by superimposing a thermohaline flow component to the wind-driven circulation. The best agreement with observations could be obtained from a model run driven by a sinking rate of 20 Sv in the northern North Atlantic and 4 Sv in the Weddell Sea compensated by 15 Sv return flow from other oceans via the Agulhas Current or Drake Passage and uniform upwelling of 9 Sv in the Atlantic. In tropical and subtropical latitudes this run reproduces all observed asymmetries, but in subpolar latitudes the model fails. Further conclusions can be drawn from the model results. (i) Up to 20 Sv northward transport of Antarctic Intermediate Water is needed at about 10°S to explain the difference of modeled transports and observations. For the same reasons an Antilles Current of up to 16 Sv is required. (ii) The major part of the northward heat transport in the North Atlantic has to occur via the tropical countercurrents and the North Equatorial Current. Only less than 7 Sv take the shortest way to the Caribbean via the Guyana Current. (iii) Fifty-six percent of the Florida Straits transport is wind driven.

Description: The effects of a floating fish farm in Kiel Fjord, Western Baltic, have been studied in the summer 1991 by underwater video, sediment and benthos samples. Significant alterations of the benthos and sediment geochemistry as compared to control stations were documented. The sediment under the farm is anoxic, organically enriched (1.5 to 3.5 fold), covered by sulfur bacteria, and almost free of benthic macro fauna. Rates of decay of organic carbon and oxygen uptake (derived from porewater profiles) are high and account for 100–150 mmol m‐2 d‐1 in summer.

Description: Nicht leicht haben es die Vertreter einer wenig beschriebenen Terordnung, nach denen wir diesmal fragen. Birgt doch ihr Parasitendasein mit mehrfachem Wirtswechsel in den Weiten der Ozeane nicht wenige Gefahren.

Description: Accurate measurement of fluctuations in temperature and humidity are needed for determination of the surface evaporation rate and the air-sea sensible heat flux using either the eddy correlation or inertial dissipation method for flux calculations. These measurements are difficult to make over the ocean, and are subject to large errors when sensors are exposed to marine air containing spray droplets. All currently available commercial measurement devices for atmospheric humidity require frequent maintenance. Included in the objectives of the Humidity Exchange over the Sea program were testing and comparison of sensors used for measuring both the fluctuating and mean humidity in the marine atmosphere at high wind speeds and development of techniques for the protection of these sensors against contamination by oceanic aerosols. These sensors and droplet removal techniques are described and comparisons between measurements from several different systems are discussed in this paper. To accomplish these goals, participating groups devised and tested three methods of removing sea spray from the sample airstream. The best performance was given by a rotating semen device, the “spray Ringer.” Several high-frequency temperature and humidity instruments, based on different physical principles, were used in the collaborative field experiment. Temperature and humidity fluctuations were measured with sufficient accuracy inside the spray removal devices using Lyman-α hygrometers and a fast thermocouple psychrometer. Comparison of several types of psychrometers (using electric thermometers) and a Rotronic MP-100 humidity sensor for measuring the mean humidity illustrated the hysteresis of the Rotronic MP-100 device after periods of high relative humidity. Confidence in the readings of the electronic psychrometer was established by in situ calibration with repeated and careful readings of ordinary hand-held Assman psychrometers (based on mercury thermometers). Electronic psychrometer employing platinum resistance thermometers perform very well.

Description: Changes in permeability and porosity during shortening deformation of Carrara marble and hot-pressed calcite aggregates were measured under high pressure at room temperature using argon as pore fluid. At effective pressures of 30 and 50 MPa, the permeability of Carrara marble increased by up to 2 orders of magnitude with less than 2% strain during which the connected porosity increased by only 0.005. The permeability increased more slowly with further strain up to 18%, during which the connected porosity increased by a further 0.05 to 0.06. At effective pressures of 100 MPa to 200 MPa, these effects were much less marked. In hot-pressed calcite aggregates, deformed at an effective pressure of 50 MPa, the permeability increased by about 2 orders of magnitude after about 12% strain and an increase in connected porosity of about 0.03. Microstructural studies indicate that, in the coarse-grained Carrara marble specimens, both transgranular and grain boundary cracks are present after room temperature deformation. For a given strain, the average length and the linear density of transgranular cracks decrease with increasing effective pressure. In fine-grained, hot-pressed calcite aggregates, dilatancy is mainly due to opening of grain boundary cracks. The very marked increase in permeability with small strain at low effective pressure can be correlated with the proliferation of connected microcracks of relatively large apertures, deduced on the basis of theoretical models.

Description: New light is shed on Worthington's concept of the North Atlantic circulation, postulating the existence of two anticyclonic gyres. This concept, which seems to have been laid to rest in the last decade, has now been reinforced by the results of a simple linear Sverdrup circulation model yielding a band of westward transport all across the North Atlantic at about the Azores latitude. This narrow band is called the Azores Countercurrent (AzCC) and matches the position of westward flow required by Worthington's “northern gyre.” An anomaly in the meridional change of the wind-stress curl in the eastern North Atlantic has been identified as the driving mechanism. A comparison with observations shows that the AzCC is verified in many analyses of historical datasets and synoptic surveys. A lack of the AzCC in other analyses is probably due to missing meridional sections, strong smoothing, and the superimposed Ekman flow close to the sea surface directed to the southeast. The AzCC has not been verified in low-resolution general circulation models applying simplified wind-stress fields and large friction coefficients, but there is evidence for its existence in recent high-resolution models driven by realistic wind stresses. Based on these findings, a new pattern for the wind-driven upper ocean circulation of the midlatitude North Atlantic is presented.

Description: A sigma-coordinate, primitive equation ocean circulation model is used to explore the problem of the remnant generation of trapped waves about a tall, circular, isolated seamount by an incident oscillatory barotropic current. The numerical solutions are used to extend prior studies into the fully nonlinear regime, and in particular to quantify and interpret the occurrence of residual circulation. Specific attention is also devoted to the dependence of the resonance and rectification mechanisms on stratification, forcing frequency, and choice of subgrid-scale viscous closure. Resonantly generated trapped waves of significant amplitude are found to occur broadly in parameter space; a precise match between the frequency of the imposed incident current and the frequency of the trapped free wave is not necessary to produce substantial excitation of the trapped wave. The maximum amplification factors produced in these numerical solutions, O(100) times the strength of the incident current, are consistent with previous studies. In the presence of nonlinear advection, strong residual currents are produced. The time-mean circulation about the seamount is dominated by a strong bottom-intensified, anticyclonic circulation closely trapped to the seamount. Maximum local time-mean current amplitudes are found to be as large as 37% of the magnitude of the propagating waves. In addition to the strong anticyclonic residual flow, there is a weaker secondary circulation in the vertical-radial plane characterized by downwelling over the top of the seamount at all depths. Maximum vertical downwelling rates of several tens of meters per day occur at the summit of the seamount. The vertical mass flux implied by this systematic downwelling is balanced by a slow radial flux of mass directed outward along the flanks of the seamount. Time-mean budgets for the radial and azimuthal components of momentum show that horizontal eddy fluxes of momentum are responsible for transporting net radial and azimuthal momentum from the far field to the upper flanks of the seamount. There, Coriolis and pressure gradient forces provide the dominant balances in the radial direction. However, the Coriolis force and viscous effects provide the primary balance for the azimuthal component.

Description: Current measures of microbe‐mediated biogeochemical processes in sediments were examined for their potential use as indicators of heavy metal ecotoxicity in both river sediments and bacterial cultures. Assays were carried out with HgCl2, CuSO4, and 3CdSO4 · 8H2O added to sediment samples and bacterial cell suspensions at concentrations ranging from 0.1 to 10 mM and 0.1 μM to 1 mM, respectively. Chemoautotrophic CO2 fixation by Elbe River sediment microbiota was most sensitive to Hg2+ and Cd2+, but not to Cu2+. Among the estimates of heterotrophic productivity, incorporation of leucine into cellular protein showed clearer dose responses than incorporation of thymidine into bacterial DNA. Thymidine incorporation was highly resistant to and even stimulated by metal ions, particularly in starved and anaerobic cultures of a test strain of Vibrio anguillarum. Similar metal ion induced “overshoot” responses beyond the levels of untreated controls were noted for mineralization of 14C‐glucose by V. anguillarum and, in the case of Cd2+, also in sediment. As a less complex measure of microbial respiratory activity, succinate dehydrogenase (SDH) showed normal dose responses without stimulatory effects, as long as bacterial cell homogenates were assayed. Despite this result, it is concluded that levels of SDH in natural sediment microbiota are inevitably affected by metal‐induced processes of selection and enzyme synthesis, and would thus fail to provide an appropriate measure of metal ecotoxicity. The final conclusion is that current parameters of microbial production and activity often reveal dose responses that do not fulfill basic requirements of ecotoxicity testing in metal‐polluted sediments.

Description: The total transport of Antarctic Bottom Water across the Rio Grande Rise, including the western boundary, the Vema Channel, and the Hunter Channel is estimated from hydrographic measurements across these pathways. The contribution of the Vema Channel is greatest at 3.9 × 106 m3 s−1, which is very close to earlier estimates. The western boundary current contribution is 2.0 × 106 m3 s−1 and that of the Hunter Channel 0.7 × 106 m3 s−1. The lower values outside the Vema Channel are offset by the important source of mass they form to the lower density classes of bottom water. About 40% of the flow is concentrated in the highest density class representing the source of Weddell Sea Deep Water to the Brazil Basin. The flow structure is characterized by horizontal and vertical recirculation.

Description: On a transect between 20° and 70°S in the eastern Atlantic Ocean and Weddell Sea, water samples from 19 hydrographic stations and bottom water from 55 surface sediment samples taken with a multiple corer were investigated for the stable carbon isotopic composition of the total dissolved inorganic carbon (δ13CΣCO2). These measurements were compared to δ13C values determined on live specimens of the benthic foraminifer Fontbotia wuellerstorfi and closely related genera from the same stations. In addition, at 16 stations the stable carbon isotope composition of sedimentary organic carbon was measured. General deepwater and bottom-water mass circulation patterns as inferred from the δ13CΣCO2 are in close agreement with those known from other nonconservative tracers. Very low δ13C values of upper Circumpolar Deep Water (〈0.3‰ Pee Dee belemnite (PDB)) in the Polar Front region and the eastern limb of the Weddell gyre coincide with nutrient maxima. However, a significant decoupling of the dissolved phosphate signal from the δ13CΣCO2 signal is indicated in the abyssal Weddell Sea. We attribute this to temperature-dependent fractionation processes during gas exchange of surface waters with the atmosphere at sites of bottom-water formation. Multiple corer water from the sediment/water interface is slightly δ13C depleted relative to deepwater and bottom-water δ13ΣCO2. The surface sediment organic carbon δ13C is 3 to 4‰ lower south of the Polar Front than north of it, and the δ13Corg in freshly accumulated phytodetritus is 3 to 4‰ lower than surface sediment organic carbon δ13C. Comparison of live F. wuellerstorfi δ13C and related genera with bottom-water δ13CΣCO2 exhibits at most stations between the Subtropical Front (≈41°S) and the southern boundary of the Antarctic Circumpolar Current (≈55°S) a significant lowering of foraminiferal δ13C values. Compilation of a mean last glacial/interglacial δ13C amplitude (Δδ13C) from six published southern ocean cores results in a shift of −0.99± 0.13‰ PDB; this shift is greater than that in all other regions. However, all of these cores are from positions close to Recent oceanic fronts. Thus, for these peripheral areas of the southern ocean, we suggest about half of the glacial/interglacial shift can be explained by varying frontal zone positions and widths accompanied by a change in mode and height of export production.

Description: What limits phytoplankton growth in nature? The answer is elusive because of methodological problems associated with bottle incubations and nutrient addition experiments. We are investigating the possibility that antibodies to proteins repressed by a specific nutrient can be used as probes to indicate which nutrient limits photosynthetic carbon fixation in the ocean. The diatom Phaeodactylum tricornutum Bohlin and the chlorophyte Dunaliella tertiolecta Butcher were grown in batch cultures in artificial seawater and f/2 nutrient lacking either phosphorus, iron, or nitrogen. Chlorosis was induced by nutrient limitation in both species with the exception of phosphorus‐limited D. tertiolecta. The synthesis and appearance of specific proteins were followed by labeling with 14C‐bicarbonate. Nutrient limitation in general leads to a decrease in the quantum efficiency of photosystem II, suggesting that deficiency of any nutrient affects the photosynthetic apparatus to some degree: however, the effect of nitrogen and iron limitation on quantum efficiency is more severe than that of phosphorus. A crude fractionation of the soluble and membrane proteins demonstrated that the large proteins induced under limitation by phosphorus and iron were associated with the membranes. However, small iron‐repressible proteins were located in the soluble fraction. Isolation with anion‐exchange chromatography and N‐terminal sequencing of iron‐repressible, 23‐kDa Proteins from D. tertiolecta, P. tricornutum, and Chaetoceros gracilis revealed that these small soluble proteins have strong homology with the N‐terminal sequence of flavodoxins from Azotobacter and Clostridium. The identity of the flavodoxin from D. tertiolecta was confirmed by immunodetection using antiflavodoxin raised against Chlorella. Flavodoxin was detected only under iron deprivation and was absent from nitrogen‐and phosphorus‐limited algae. Flavodoxin is a prime candidate for a molecular probe of iron limitation in the ocean. The requirements to confirm its utility in nature are discussed.

Description: Etwa ein Drittel des von Menschen freigesetzten Treibhausgases Kohlendioxid (CO2) reichert sich in der Atmosphäre an und verstärkt dort den Treibhauseffekt. Zwei Drittel dieses Eintrags werden der Atmosphäre wieder entzogen und an anderer Stelle deponiert. Doch nur für etwa die Hälfte des wieder gebundenen Kohlendioxids kennen wir bisher die Senken. Wo bleibt der Rest? Um Aussagen über den Verbleib weiterer CO2-Emissionen und damit über die zukünftige Entwicklung des Treibhauseffektes machen zu können, bedarf es der Lösung des Kohlenstoffrätsels.

Description: The effects of nitrate, phosphate, and iron starvation and resupply on photosynthetic pigments, selected photosynthetic proteins, and photosystem II (PSII) photochemistry were examined in the diatom Phaeodactylum tricornutum Bohlin (CCMP 1327). Although cell chlorophyll a (chl a) content decreased in nutrient‐starved cells, the ratios of light‐harvesting accessory pigments (chl c and fucoxanthin) to chl a were unaffected by nutrient starvation. The chl a‐specific light absorpition coefficient (a*) and the functional absorption cross‐section of PSII (σ) increased during nutrient starvation, consistent with reduction of intracellular self‐shading (i.e. a reduction of the “package effect”) as cells became chlorotic. The light‐harvesting complex proteins remained a constant proportion of total cell protein during nutrient starvation, indicating that chlorosis mirrored a general reduction in cell protein content. The ratio of the xanthophylls cycle pigments diatoxanthin and diadinoxanthin to chl a increased during nutrient starvation. These pigments are thought to play a photo‐protective role by increasing dissipation of excitation energy in the pigment bed upstream from the reaction centers. Despite the increase in diatoxanthin and diadinoxanthin, the efficiency of PSII photochemistry, as measured by the ration of variable to maximum fluorescence (Fv/Fm) of dark‐adapted cells, declined markedly under nitrate and iron starvation and moderately under phosphate starvation. Parallel to changes in Fv/Fm were decreases in abundance of the reaction center protein D1 consistent with damage of PSII reaction centers in nutrient‐starved cells. The relative abundance of the carboxylating enzyme, ribulose bisphosphate carboxylase/oxygenase (RUBISCO), decreased in response to nitrate and iron starvation but not phosphate starvation. Most marked was the decline in the abundance of the small subunit of RUBISCO in nitrate‐starved cells. The changes in pigment content and fluorescence characteristics were typically reversed within 24 h of resupply of the limiting nutrient.

Description: According to Sverdrup's (1953) model of the spring bloom, phytoplankton biomass decreases in winter when the mixed layer depth exceeds the critical depth. We have used a one-dimensional mathematical model integrated by the Lagrangian Ensemble method to simulate a population of diatoms during the winter between two growing seasons off the Azores. The model allows us to diagnose the demographic changes in the simulated diatom population from a variety of perspectives. The total population falls to a minimum of 70 million diatoms m-2 at the end of February. The vertical distribution of the population dynamics is first analysed in terms of daily Eulerian averages over 1 m depth intervals. Growth starts in February when the diurnal thermocline becomes shallower than 50 m, but while the mixed layer is still 200 m deep. The natural mortality has a minimum in winter because it is reduced (in the model) with temperature and population density. Eulerian analysis suggests that in winter, diatoms have a life expectancy of more than 3 months, so a significant number will survive the months of December, January and February when there is very little growth. Losses to grazing are negligible in winter. Lagrangian analysis shows how an individual diatom responds to its changing ambient environment caused by variation in depth (due to turbulent mixing) and the diurnal and seasonal changes in the photosynthetically active radiance. The different trajectories followed by the thousands of plankton particles simulated by the model produce diversity in growth rate ranging over several orders of magnitude, so care has to be taken in statistical analysis. The paper ends with a re-assessment of the value of the critical depth and compensation depth as predictors for onset of the spring bloom. The compensation depth was computed by Eulerian averaging over 1 m depth inter-vals each day. For 1 month after the vernal equinox the compensation depth follows the ascent of the mixed layer as it rises from a depth of 100 m to 40 m. Lagrangian analysis reveals that this is due to the photo-adaptation better matching the ambient irradiance experienced by diatoms in the mixed layer compared with those at the same depth in the seasonal thermo-cline. By mid-April the spring bloom has already ad-vanced so far that self shading influences the compensation depth, which then rises into the mixed layer. We conclude that Sverdrup's criterion is not useful for predicting changes in the diatom population simulated by our model.

Description: The space-time structure and predictability of the El Niño/Southern Oscillation (ENSO) phenomenon was investigated. Two comprehensive datasets were analyzed by means of an advanced statistical method, one based on observational data and the other on data derived from an extended-range integration performed with a coupled ocean-atmosphere general circulation model. It is shown that a considerable portion of the ENSO-related low-frequency climate variability in both datasets is associated with a cycle involving slow propagation in the equatorial oceanic beat content and the surface wind field. The existence of this cycle implies the ability of climate predictions in the tropics up to lead times of about one year. This is shown by conducting an ensemble of predictions with our coupled general circulation model. For the first time a coupled model of this type was successfully applied to ENSO predictions.

Description: A hybrid coupled model (HCM) of the tropical ocean–atmosphere system is described. The ocean component is a fully nonlinear ocean general circulation model (OGCM). The atmospheric element is a statistical model that specifies wind stress from ocean-model sea surface temperatures (SST). The coupled model demonstrates a chaotic behavior during extended integration that is related to slow changes in the background mean state of the ocean. The HCM also reproduces many of the observed variations in the tropical Pacific ocean-atmosphere system. The physical processes operative in the model together describe a natural mode of climate variability in the tropical Pacific ocean–atmosphere system. The mode is composed of (i) westward-propagating Rossby waves and (ii) an equatorially confined air–sea element that propagates eastward. Additional results showed that the seasonal dependence of the anomalous ocean–atmosphere coupling was vital to the model's ability to both replicate and forecast key features of the tropical Pacific climate system. A series of hindcast and forecast experiments was conducted with the model. It showed real skill in forecasting fall/winter tropical Pacific SST at a lead time of up to 18 months. This skill was largely confined to the central equatorial Pacific, just the region that is most prominent in teleconnections with the Northern Hemisphere during winter. This result suggests the model forecasts of winter SST at leads times of at least 6 months are good enough to be used with atmospheric models (statistical or OGCM) to attempt long-range winter forecasts for the North American continent. This suggestion is confirmed in Part II of this paper.

Description: A 26-year integration has been performed with a coupled ocean-atmosphere general circulation model (CGCM). The oceanic part resolves all three oceans in the latitude band 70°N–70°S but is dynamically active only between 30°N and 30°S. The atmosphere is represented by a global low-order spectral model. The coupled model was forced by seasonally varying insolation. Although the simulated time-averaged mean conditions in both atmosphere and ocean show significant deviations from the observed climatology, the CGCM realistically simulates the interannual variability in the tropical Pacific. In particular, the CGCM simulates an irregular ENSO with a preferred time scale of about 3 years. The mechanism for the simulated interannual variability in the tropical Pacific is related to both the “delayed action oscillator” and the “slow SST mode.” It therefore appears likely that either both modes can coexist or they degenerate to one mode within certain locations of the parameter space. This hypothesis is also supported by calculations performed with simplified coupled models, in which the atmospheric GCM was replaced by linear steady-state atmosphere models. Further, evidence is found for an eastward migration of zonal wind anomalies over the western Pacific prior to the extremes of the simulated ENSO, indicating a link to circulation systems over Asia. Because an earlier version of the CGCM did not simulate interannual variability in the tropical Pacific, additional experiments with a simplified coupled model have been conducted to study the sensitivity of coupled systems to varying mean oceanic background conditions. It is shown that even modest changes in the background conditions can push the coupled system from one flow regime into another.

Description: Long-range global climate forecasts were made by use of a model for predicting a tropical Pacific sea-surface temperature (SST) in tandem with an atmospheric general circulation model. The SST is predicted first at long lead times into the future. These ocean forecasts are then used to force the atmospheric model and so produce climate forecasts at lead times of the SST forecasts. Prediction of seven large climatic events of the 1970s to 1990s by this technique are in good agreement with observations over many regions of the globe.

Description: The demersal fish and cephalopod communities of the continental shelf and upper slope from 17 to 395m deep were studied during five annual cruises between Cape Agulhas and Port Alfred, South Africa. The cruises showed a consistent pattern of an inshore community (〈100m), a shelf community (c. 90–190m) and a shelf‐edge/upper slope fauna (〉200m). These groups were identified by dendrograms and multidimensional scaling cluster analysis, which supported on‐board observations of catch variation with depth. Although the boundaries are not clearly defined, examination of physical features at the clustered stations suggests that depth, temperature and, to a lesser extent, oxygen concentration are important in the grouping. Occasional, apparently anomalous associations of inshore stations suggested that water temperature and oxygen may over‐ride the normal depth distributions of the species groups. This intimates that patterns offish and cephalopod distribution may be dynamic and in part related to the physical parameters of the water body.

Description: 1. Survival, growth and downstream dispersal of trout (especially 0 group) and the relationships of these variables to initial stocking density were studied in north Pennine streams. 2. Two methods were used. First, electrofishing censuses were made in a marked reach of each of four streams over a period of about 20 years. Second, downstream moving trout were trapped in two streams over a 10-year period. Each stream upstream of the trap was experimentally stocked with `swim-up' trout fry, using a different population density each year. 3. Before 1970 the four census reaches showed very large year-upon-year variations in August trout parr densities, with local failures of recruitment in some years. Population densities after completion of Cow Green Reservoir (1970) were generally higher but still showed wide fluctuations. 4. Survival (including the effects of losses by dispersal) from swim-up to early August, for starting population densities of 0-10 fry m-2, was about 10% regardless of initial density. Estimates of survival from August to early October were 30-50% for the census reaches and 55-65% for the areas upstream of the traps. However, for August 0 group densities of 0-0.9 m-2, estimated instantaneous loss rate from August of the first year of life up to age 40-65 months showed a positive curvilinear relationship to population density in the first year of life. Loss rate was, therefore, density-dependent during this period. 5. Estimated instantaneous growth rate day-1 of 0 group fish from swim-up to August and from swim-up to October was inversely related to the natural logarithm of August population density and this was most apparent for August densities of 〈0.15 fish m-2. 6. Although survival from swim-up to August was proportional (about 10%, at starting densities of 10 m-2 or less), the percentages of the total loss attributable to mortality and to downstream dispersal varied considerably with starting density. At starting densities around 4-5 fish m-2 dispersal was negligible. As initial density rose above 4-5 fish m-2 and towards 10 fish m-2 the percentage of loss attributable to dispersal rose towards 30%. As initial densities decreased from 4 to 1.4 fish m-2, the percentage rose to around 20%. Below a starting density of 1.4 fish m-2 the percentage decreased.

Description: Ocean deep velocity profiles were obtained by lowering a self-contained 153.6-kHz acoustic Doppler current profiler (ADCP) attached to a CTD-rosette sampler. The data were sampled during two Meteor cruises in the western tropical Atlantic. The ADCP depth was determined by integration of the vertical velocity measurements, and the maximum depth of the cast was in good agreement with the CTD depth. Vertical shears were calculated for individual ADCP velocity profiles of 140-300-m range to eliminate the unknown horizontal motion of the instrument package. Subsequent raw shear profiles were then averaged with respect to depth to obtain a mean shear profile and its statistics. Typically, the shear standard deviations were about 10(-3) s-1 when using up and down traces simultaneously. The shear profiles were then vertically integrated to get relative velocity profiles. Different methods were tested to transform the relative velocities into absolute velocity profiles, and the results were compared with Pegasus dropsonde measurements. The best results were obtained by integrating the raw velocities and relative velocities over the duration of the cast and correcting for the ship drift determined from the Global Positioning System. Below 1000-m depth a reduction of the measurement range was observed, which results either from a lack of scatterers or instrumental problems at higher pressures.

Description: Inertial separation of a western boundary current from an idealized continent is studied in a homogeneous ocean circulation model. A number of processes are identified that either encourage or prevent separation at a coastal promontory in this model. For a single-gyre wind forcing a free-slip boundary condition forces the stream to follow the coastline, whereas the no-slip condition allows separation at a sharp corner. A prescribed countergyre to the north of the stream is not necessary to achieve separation if the no-slip condition is used. "Premature" separation occurs for wind fields that do not extend beyond the latitude of the cape. For a more realistic wind field and coastline two distinct states of the stream are found. At small Reynolds numbers the current fails to separate and develops a stationary anticyclonic meander north of the cape. Stronger currents separate and drive a recirculation in the lee of the continent.

Description: Seismically derived depth estimates to the top of the oceanic crust beneath the Hawaiian Islands indicate that the curvature of the deflected lithosphère is much larger than commonly believed. The conservative and model-independent curvature estimates exceed 10−7 m−1 and are comparable in magnitude to curvatures at trenches and outer rise systems. The depth estimates are used to constrain both two-dimensional (2-D) and three-dimensional (3-D) flexural models. The curvature constraints require a 2-D variable elastic thickness that decreases from 35 km in areas away from the volcanic load to 25 km directly beneath the load. In an attempt to understand the nature of the yielding beneath the Hawaiian Islands we introduce two new 3-D models. The first model combines a realistic yield strength based rheology with a new technique for 3-D flexure calculations in which the elastic plate thickness is curvature-dependent. The new variable rigidity model predicts an undeformed (mechanical) plate thickness of 44 km, decreasing to 33 km beneath the big island of Hawaii. The best-fitting mechanical thickness corresponds approximately to the depth to the 600 °C isotherm in 90-m.y.-old lithosphere. The second model uses a broken plate, but here the crack is oriented along the weak Molokai fracture zone rather than along the island chain trend. This unconventional flexure model can explain the observed asymmetry in the depth data across the fracture zone without requiring the excessively large elastic thickness of more conventional broken plate models. Both the proposed models imply that modeling with constant thickness plates may underestimate the true mechanical plate thickness by being unduly influenced by the weak zone beneath the seamounts.

Description: Studies of laboratory cultures of Chordaria linearis (Hooker et Harvey) Cotton from southernmost South America revealed that this species has an obligate sexual life history in which a macroscopic sporophyte alternates with a monoecious microscopic gametophyte. Sexual reproduction is isogamous and under photoperiodic control. Gametes are produced only in short days, whereas in long days, asexual zoospores are formed that recycle the gametophyte generation. Unfused gametes develop into gametophytes, and sporophytes originate only from zygotes. Unlike other sexual members of the Chordariales, gametes of C. linearis have a reduced stigma and do not show phototaxis. They are released at the beginning of the night, not in the morning. In nature, C. linearis seems to be regularly infected by a dictyosiphonalean epiphyte resembling the rare arctic species Trachynema groenlandicum (Lund) Pedersen. The epiphyte is responsible for previous contradictory results obtained in laboratory cultures of C. linearis. This is the first record of Trachynema in the southern hemisphere.

Description: The constancy of postmoult/premoult ratios of measures of linear size during ontogeny in insect and other arthropods is widely known as Dyar's rule. We tested this rule in nine species of the waterstrider genera Gerris and Aquarius (Heteroptera: Gerridae), using two size variables: head width and a multivariate measure derived from the pattern of multivariate allometry common to the species considered. Allometric patterns were similar in two independent datasets of laboratory-reared and field-caught specimens. Although our data strictly followed Dyar's rule injust a few instances, all growth ratios varied within a limited range only. Growth ratios for head width differed more between moults than those for multivariate size. The relationship between growth ratios for the two size measures conformed to the predictions based on allometry. We discuss hypotheses of the possible adaptive significance of growth ratios, such as their relation to mobility and systematic differences between hemimetabolous and holometabolous insects, and emphasize the importance of allometry. Since Dyar's rule is consistent with available evidence of physiological mechanisms underlying growth and moulting control of insects and crustaceans, it can be used as a general frame of reference to test alternative growth models.

Description: The ultimate cause of the onset of glaciations remains elusive, but in the case of northem hemisphere glaciation it is probable that several factors acted in combination. General global cooling resulted from reduction of atmospheric C02 by weathering of silicate rocks exposed by erosion of late Cenozoic uplifts. Uplifts in south Asia, southwestern North America and Scandinavia occurred at distances appropriate for the generation of quasi-permanent Rossby waves in the atmosphere. The resulting winds, given suitable moisture sources, were favourable for causing large-scale precipitation at mid-latitudes on the northern continents. Moisture sources were provided by the closure of the Central American isthmus. Gulf Stream flow increased, carrying warm subtropical waters to high latitudes. The Denmark Strait deepened permitting greater outflow of deep water from the Norwegian-Greenland Sea. The relative importance of each of these factors should be investigated by additional atmospheric and ocean climate model sensitivity studies.

Description: King Penguins are the second largest of all diving birds and share with their congener, Emperor Penguins, breeding habits strikingly different from other penguins. Our purpose was to determine the feeding behavior, energetics of foraging and the prey species, and compare these to other sympatric species of subantarctic divers. We determined: (1) general features of foraging behavior using time—depth recorders, velocity meters, and radio transmitters, (2) energetics by doubly labeled water, (3) food habits and energy content from stomach lavage samples, and (4) resting and swimming metabolic rate by oxygen consumption measurements. The average foraging cycle was ≈6 d, during which the mass gain of 30 birds was ≈2 kg. When at sea, the birds exhibit a marked pattern of shallow dives during the night, whereas deep dives of 〉100 m only occurred during the day. Maximum depth measured from 34 birds and 18 537 dives was 304 m, and maximum dive duration from 23 birds and 11 874 dives was 7.7 min. The frequency distribution of dive depth was bimodal, with few dives between 40 and 100 m. Overall, swim velocities when a bird was at sea averaged 2.1 m/s (N = 5), while descent and ascent rates of change in depth averaged 0.6 m/s for dives 〈60 m (N = 74) and 1.4 m/s for dives 〉150 m (N = 90). Night feeding dives occurred at a rate of ≈20 dives/h, and deep dives occurred at a rate of ≈5 dives/h. The energy consumption rate while resting ashore was 3.3 W/kg (N = 3) or 1.6 times the predicted standard metabolic rate (SMR). The average energy consumption rate while away from the colony was 10 W/kg (N = 8) or 4.6 x SMR, compared to 4.3 x SMR estimated from a time—energy budget. The latter value is based on an average metabolic rate of 4.2 W/kg for three birds while resting in 5°C water and 9.6 W/kg while swimming at 2 m/s, which was extrapolated from the average of three birds swimming at 1 m/s. The average energy intake based on 9 stomach content samples was nearly 24.6 kJ/g dry mass. The main prey by number are myctophid fish of the species Krefftichthys anderssoni and Electrona carlsbergi. It was concluded that: (1) feeding begins ≈28 km from the colony, (2) prey is pursued night and day through its vertical movements, (3) vertical distribution of the prey is reflected closely by diving habits of the birds, (4) deep—diving, for unknown reasons, is an important component of foraging success, (5) diving capacities of King Penguins are remarkable compared to other birds and many pinnipeds, and (6) calculated foraging energetics can be closely estimated from time—energy budgets.

Description: Routine oxygen consumption of turbot, Scophthalmus maximus, was determined in relation to temperature, salinity, body wet weight, and time of day. The highest routine oxygen consumption rates measured roughly followed a arabolic curve over the temperature range tested (8 to 24°C). The lowest rates showed a more linear refationship over the same temperature range. It is argued that lowest rates correspond to the standard oxygen consumption. Between 16 and 19°C, routine oxygen consumption reached a maximum. It is suggested that these temperatures correspond to the preferred ternerature of the species and are within the range of optimum temperature for growth of specimens weigkng about 100 g. Salinity effect on oxygen consumption rates was studied in five groups acclimated over 4 weeks to 8, 15, 22, 29, and 35%. salinity. Routine oxygen consumption rates were lowest at 8% salinity with no significant differences in higher acclimation salinities. Routine respiration of turbots showed conspicuous daily fluctuations. During spring, summer, and autumn, oxygen Consumption was higher during morning hours and at night. In winter, higher rates were measured only once (during morning and early afternoon). The relationship between routine oxygen consumption and body weight of turbots followed an exponential function with a slope of 0.7, which was lower compared to the slope of 0.8 usually given for roundfish-species.

Description: North Atlantic air-sea heat and freshwater flux data from several sources are used to estimate the conversion rate of water from one density to another throughout the range of sea surface density. This cross-isopycnal mass flux varies greatly over the ocean, with a maximum of 32.2 × 106 m3 s−1 at σ = 26.1 kg m−3 (toward greater densities) and a minimum of −7.6 × 106 m3 s−1 (toward lesser densities) at σ = 23.0 kg m−3. The air-sea fluxes force water to accumulate in three density bands: one at the lowest sea surface densities generated by heating; one centered near the density of subtropical mode water; and one spanning subpolar mode water densities. The transfer of water to the highest and lowest densities is balanced by mixing, which returns water to the middle density range, and also by boundary sources or sinks. Integrating the cross-isopycnal flux over all densities gives an annual average sinking of about 9 × 1O6 m3 s−1, which presumably escapes across the equator and must be balanced by a similar inflow. Comparison with estimates from tracer studies suggests that the renewal of tracer characteristics at a given density may occur without the existence of an annual average mass source at that density, because along- and cross-isopycnal mixing can renew a tracer without supplying mass.

Description: Haplogloia andersonii (Farlow) Levring is an anti-tropical species that occurs on cold and warm-temperate Pacific coasts of both Americas. In its habit it resembles the subantarctic species Chordaria linearis (Hooker et Harvey) Cotton. Culture studies show that the species differ in morphology and ecophysiology of their microscopic gametophytes and in gamete behavior. Details of sporophyte anatomy are presented that also allow the distinction of field plants. In South America, H. andersonii occurs only on the Pacific coast, from central Perú (14°S) to southern Chile (50°S). Chordaria linearis occurs on the Pacific coast from Chiloé Island (43°S) to Cape Horn (56°S). In the shared area the species may co-occur. On the Atlantic coast, C. linearis was newly collected at a locality in northern Patagonia (41°S). In addition, C. linearis occurs in Antarctica. Haplogloia moniliformis Richer, recently described from Macquarie Island, is probably synonymous with Chordaria linearis.

Description: We examined the temperature tolerance of microscopic phases from geographically disjunct isolates of eight species or closely related, putatively conspecific taxa of temperate brown algae with disjunct distributions. Maximum within-taxon differences were small and ranged from 1.6° to 4.3° C. Desmarestia aculeata and Sphaerotrichia divaricata, both with northern hemisphere amphioceanic distributions, showed little or no significant intraspecific variation between the mean upper survival limits (USL) of Atlantic and Pacific strains (δUSL ≤ 1.4°C), which would agree with a relatively recent separation of the respective populations. Among the plants with bipolar distributions, there was likewise very little difference (δUSL 0–1.1°C) between northern and southern hemisphere strains in Striaria attenuata and in the species pair Desmarestia viridis/D. willii. In Desmarestia ligulata, and in the species pairs Desmarestia firma/D. munda, Dictyosiphon foeniculaceus/D. hirsutus, and Scytothamnus australis/Scytothamnus sp., significant differences occurred, which indicate longer divergence times. δUSL in these cases ranged from 1.7° to 2.7°C, without overlap between strains from the northern and southern hemispheres. All species that passed the equator during cooler epochs had a USL of 26–27°C, at least in some geographical isolates. The NE Asian kelp Undaria pinnatifida, which passed the equator in recent times, had a USL of 29.6°C. We hypothesize that the mechanism of spreading in the amphipolar species studied was migration of vegetative microthalli. The more unlikely alternative hypothesis of continuous populations through the tropics during a cooler epoch would imply a drop in seawater temperatures to approximately 20° C in summer and 15° C in winter, which is not supported by paleoclimatic evidence.

Description: The survival of cod Gadus morhua, plaice Pleuronectes platessa, and dab Limanda limanda was determined in relation to ambient oxygen saturation at 8° C and 35% m salinity. Mortalit rates were observed in fish exposed to constant oxygen levels for 24h. First mortality occurred around 60 % oxygen saturation in cod and around 30% oxygen saturation in dab and plaice. Below these thresholds mortality increased linearly with decreasing oxygen levels. If cod were infested with 1 or 2 individuals of Lernaeocera branchialis (Copepoda), their tolerance was significantly lower; under such circumstances the incipient lethal oxygen saturation was 66 %.

Description: The responses of sea ice microalgae to variation in ambient irradiance (0 to 150 μE · m−2· s−1), temperature (–6° to + 6° C), and salinity (0 to 100 ppt) were tested to determine whether these variables act independently or in concert to influence rates of microalgal photosynthesis. The photosynthetic efficiency and maximum photosynthetic rate for sea ice microalgae increased as a function of incubation temperature between -6° and + 6° C. Furthermore, photosynthetic efficiency, maximum photosynthetic rate, and quantum yield were greatest at salinities between SO and 50 ppt. In contrast, the mean specific absorption coefficients were lowest near seawater salinities, and the saturating irradiance, Is, appeared to be inversely proportional to salinity. Results also suggest that the effects of salinity on the growth of sea ice microalgae are independent of those elicited by temperature or light, and that the functional relationship between salinity and light or temperature is multiplicative. This information is essential to the proper formulation of algorithms used to describe algal growth in environments where light, temperature, and salinity are changing simultaneously, such as within sea ice or within the water column at the marginal ice edge zone.

Description: Antarctic Bottom Water flows into the western North Atlantic across the equator, shifting from the western side to the eastern side of the trough between the American continents and the Mid-Atlantic Ridge as it continues north. This is puzzling because such large-scale motion is thought to be controlled by dynamics that disallows an eastern boundary current. Previous explanations for the transposition involve a (necessarily small-scale) density current that changes sides because of the change in sign of rotation across the equator, or a topographic effect that changes the sign of the effective mean vorticity gradient and thus requires an eastern boundary current. Here an alternative explanation for the overall structure of bottom flow is given. A source of mass to a thin bottom layer is assumed to upwell uniformly across its interface into a less dense layer at rest. A simple formula for the magnitude of the upwelling and thickness of the layer is derived that depends on the source strength to the bottom layer. For a strong enough source, the bottom layer thickness is zero along a grounding curve that separates the bottom water from the western boundary and confines it to the east. A band of recirculating interior flow occurs, supplied by an isolated northern and western boundary current. Similar structures appear to exist in the Antarctic Bottom Water of the western North Atlantic.

Description: Results of a three-dimensional primitive equation model are presented simulating turbulent mesoscale motions in the seasonal thermocline on an f plane. The model is based on a hybrid vertical coordinate scheme and conserves isopycnic potential vorticity. Mesoscale turbulence is modeled in terms of an unstable potential vorticity front. The model integration starts from a purely zonal, 60-km-wide geostrophically balanced jet, on which is superimposed a small initial perturbation. The most unstable mode exhibits a wavelength of 85 km and is driven by a mixed type of instability. Characteristic dynamical ingredients of the wave are enhanced cyclonic and anticyclonic relative vorticity in the troughs and the ridges, respectively, due to the curvature of the flow. Vertical motion of up to 10 m d−1 occurring downstream of the ridges (downwelling) and downstream of the troughs (upwelling) is driven by geostrophic advection of relative vorticity. The contrast of static stability across the front is changing during amplification of the instability: in troughs the stability is decreasing whereas in ridges it is increasing. The density field exhibits local anomalies of the isopycnals' depths (bumps) due to the ageostrophic cross-jet advection of potential vorticity streamers wound up in cyclones and anticyclones. Locally, the potential vorticity gradients are enhanced, creating a multiple front structure. The model results support observations and findings of earlier atmospheric and oceanic models. It is emphasized that mesoscale turbulent structures may have a profound influence on primary productivity, mixed-layer, and internal wave dynamics.

Description: In this paper, the historical hydrographic database for the south Indian Ocean is used to investigate (i) the hydrographic boundary between the subtropical gyre and the Antarctic Circumpolar Current (ACC), the subtropical front (STF), and especially (ii) the southern current band of the gyre. A current band of increased zonal speeds in the upper 1000 m is found just north of the STF in the west near South Africa and at the surface STF in the open Indian Ocean until the waters off the coast of Australia are reached. As neither any other investigation of this current nor a name for it are known, the flow has been called the South Indian Ocean Current (SIOC). This name is anologous to the same current band in the South Atlantic Ocean, the South Atlantic Current. The STF is located in the entire south Indian Ocean near 40-degrees-S. The associated current band of increased zonal speeds is the SIOC, which is found at or north of the STF. East of 100-degrees-E the SIOC separates from the STF and continues to the northeast. The zonal flow south of the STF is normally weak and serves to separate the South Indian Ocean and Circumpolar currents. Near Africa the SIOC has a typical volume transport of 60 Sv (1 Sv = 10(6) m3 s-1) in the upper 1000 m relative to deep potential density surfaces of sigma(4) = 45.87 kg m-3 (2800-3500 m) or sigma(2) = 36.94 kg m-3 (1500-2500 m). Near western Australia the SIOC is reduced to about 10 Sv as it turns to the northeast.

Description: Eight species of marine phytoplankton showed significant variation in the relative amount of some fatty acids (FAs) in response to variation in temperature. Large changes in relative amounts of certain FAs occurred as a result of a 15° C change in growth temperature. For example, 14:0 increased from ≃4% of total FAs at 10° C to 〉 20% at 25° C for Chaetoceros simplex and Isochrysis aff. galbana but decreased for Phaeodactylum tricornutum. The percentage of the polyunsaturated fatty acid (PUFA) 16:ω1 was consistently greater at 10° C than at 25° C, and the converse was usually true for 16: 4ω3. Calculated over all eight species, there was a modest but significant inverse relationship between the percentage of PUFAs and temperature. Only for Thalassiosira pseudonana was the percentage of either of the PUFAs and nutritionally essential fatty acids (EFAs) also an inverse function of temperature. For T. pseudonana, the percentage of the EFA 22:6ω3 decreased linearly with increasing temperature over the range from 10 to 25° C. For three species, the ratio of unsaturated/saturated FAs was correlated with growth rate when growth rate was controlled by variation in irradiance and temperature. Only for Thalassiosira pseudonana was the ratio of unsaturated/saturated FAs also an inverse function of temperature alone.

Description: The Hindak strain of a Cryptomonas species (Cryptophyceae) produces extracellular polysaccharides. Because there is no information on the structure of these compounds in the Cryptophyceae we conducted structural studies. Gas–liquid chromatographic analyses showed that the polysaccharide is composed of fucose, rhamnose, xylose, mannose, glucose, galactose, galacturonic acid, glucuronic acid, and traces of 3-O-methyl galactose. The polysaccharide was separated into two subtractions by ion-exchange chromatography. Fraction A consisted mainly of 1,3-linked galactose units and 1,4-linked galacturonic acid. Unlike fraction B, fraction A did not have xylose, 3-O-methyl galactose, or glucuronic acid. Also, its degree of branching was low compared to that of fraction B. Only traces of sulfate were present infraction A, but fraction B was 10–15% sulfated. Protein was approximately 1% in both fractions. These polysaccharides appear to be a novel type of polymer in algae.

Description: Characteristic of the mesoscale variability in the Atlantic Ocean are investigated by analyzing the Geosat altimeter signal between 60°S and 60°N. The rms sea-surface variability for various frequency bands is studied, including the high-frequency eddy-containing band with periods 〈150 days. Wavenumber spectra and spatial eddy characteristics are analyzed over 10° by 10° boxes covering both hemispheres of the Atlantic Ocean. A comparison, with solutions of a high-resolution numerical experiment, developed as the Community Modeling Effort of the World Ocean Circulation Experiment, aids interpretation of the Geosat results in the tropical and subtropical Atlantic and provides a test of the model fluctuating eddy field. Results from Geosat altimetry show a wavenumber dependence close to k1−5 (k1 being the alongtrack wave-number) over almost the entire Atlantic Ocean except for areas in the tropical and subtropical Atlantic where the rms variability in the eddy-containing band is less than 5 cm, that is, not significantly different from the altimeter noise level. Characteristic eddy length scales inferred from Geosat data are linearly related with the deformation radius of the first baroclinic mode over the whole Atlantic Ocean, except for the equatorial regime (10°S to 10°N). The data-model comparison indicates that the high-resolution model with horizontal grid size of ⅓° and ° in latitude and longitude is quite capable of simulating observed eddy characteristics in the tropics and subtropics. In mid- and high latitudes, however, the model fails to simulate the pronounced poleward decrease in eddy scales. This leads to systematic discrepancies between the model and Geosat observation, with model scales being up to 50% larger than deduced from altimetry.

Description: The time history of upper-ocean temperatures in the tropical Pacific has been used as a predictor in a statistical prediction scheme to forecast SST anomalies in this region. The temperature variations were taken from the output of an oceanic general circulation model that was forced by observed winds for the period 1961 to 1985. Since such model data are presently used as initial conditions in prediction experiments with coupled ocean–atmosphere models, it is of particular interest to investigate up to what lead time tropical Pacific SST is predictable without the coupling of an atmosphere model to the ocean model. We compared our results with those obtained by the persistence forecast and with those obtained by using the wind stresses themselves as predictors in a statistical forecast model. It is shown that using the upper ocean temperatures from the ocean model forced by observed winds gives significantly better skills at lead times of 6 to 12 months compared to persistence and to the pure wind-stress model. Off-equatorial heat content anomalies at 5°N are shown to contribute significantly to the predictability at these lead times, while those at 12°N do not.

Description: Preexisting developmental plasticity in feeding larvae may contribute to the evolutionary transition from development with a feeding larva to nonfeeding larval development. Differences in timing of development of larval and juvenile structures (heterochronic shifts) and differences in the size of the larval body (shifts in allocation) were produced in sea urchin larvae exposed to different amounts of food in the laboratory and in the field. The changes in larval form in response to food appear to be adaptive, with increased allocation of growth to the larval apparatus for catching food when food is scarce and earlier allocation to juvenile structures when food is abundant. This phenotypic plasticity among full siblings is similar in direction to the heterochronic evolutionary changes in species that have greater nutrient reserves within the ova and do not depend on particulate planktonic food. This similarity suggests that developmental plasticity that is adaptive for feeding larvae also contributes to correlated and adaptive evolutionary changes in the transition to nonfeeding larval development. If endogenous food supplies have the same effect on morphogenesis as exogenous food supplies, then changes in genes that act during oogenesis to affect nutrient stores may be sufficient to produce correlated adaptive changes in larval development.

Description: The seasonal cycles found in moored current measurements in the equatorial Somali Current region and along the equator between 50° and 60°E are compared with the multilayer Geophysical Fluid Dynamics Laboratory model for the tropical Indian Ocean. The remote forcing of Somali Current transport variations by incident long equatorial waves from the equatorial interior subthermocline region is investigated by analyzing the model velocities of annual and semiannual period. Amplitudes and phases of linear equatorial Rossby and Kelvin waves were least-squares fitted to the model velocities between 5°S and 5°N, 55° and 86°E from 100-m to 1000-m depth. Two cases of wave fits are distinguished: the “free” Kelvin wave case, where the Kelvin waves were fitted independently, and the “reflected” Kelvin wave case, where they were coupled to the Rossby waves by the western boundary condition for a straight slanted (45° to the north) coastline. The wave field velocities explained 70% of the spatial variance in the equatorial model subregion and also compared reasonably well with observed current variations along the equator. At the western boundary, the short-wave alongshore transport due to reflected incident long waves was determined and found to be antisymmetric about the equator. The maximum transport variation for the semiannual period due to the short waves was about 5 × 106 m3 s−1 between 150- and 800-m depth at 3° north and south of the equator. Observational evidence for the western boundary transport variations and the sensitivity to changes in the incident wave field are discussed.

Description: Seasonal records of tropical sea-surface temperature (SST) over the past 10(5) years can be recovered from high-precision measurements of coral strontium/calcium ratios with the use of thermal ionization mass spectrometry. The temperature dependence of these ratios was calibrated with corals collected at SST recording stations and by (18)O/(16)O thermometry. The results suggest that mean monthly SST may be determined with an apparent accuracy of better than 0.5 degrees C. Measurements on a fossil coral indicate that 10,200 years ago mean annual SSTs near Vanuatu in the southwestern Pacific Ocean were about 5 degrees C colder than today and that seasonal variations in SST were larger. These data suggest that tropical climate zones were compressed toward the equator during deglaciation.

Description: Sampling inadequacies and an inability to distinguish age classes have limited our knowledge of octopus biology in nature. Using an artificial shelter sampling technique (Voight, 1988a), and defining mature males by the presence of enlarged suckers (Voight, In press), an intertidal population of Octopus digueti was monitored for one year. In total, 803 octopuses were narcotized; the mass, sex, arm injuries and reproductive condition of each octopus were recorded. Captures were more frequent in lower intertidal areas offering higher shelter availability and a more moderate environment. Capture rates, assumed to indicate octopus movement, correlated with sea temperature except during full moon periods when they were reduced. Over 26%, of the octopuses handled had damaged arms or arm tips, with dorsal arm pairs more often injured. The overall sex ratio was significantly male biased, probably due to maturity‐linked mobility differences between the sexes. Reproduction occurred throughout the year; reproductively competent adults, brooding females and juveniles were present every month. However, annual temperature oscillations synchronize spring hatching of eggs spawned from winter to early spring, creating a clear spring cohort. Growth and age at maturity of males in the spring and autumn cohorts were estimated. Variance was too high for these parameters to be estimated in the winter cohort. Growth rates of males over 12 weeks of age did not differ from those reported in laboratory rearing studies. Estimated average age at maturity ranged from 20 to 32 weeks, depending on temperature.

Description: Among King Penguins Aptenodytes patagonica at Possession Island, one of the Crozet Islands, the length of the moult period, pre‐laying period, incubating and brooding shifts were highly variable according to the year and to the stage of the breeding season. The moulting period was shorter in late breeders than in early breeders. Only half of the birds which successfully reared a chick bred the following cycle, but late in the season. Almost all these late breeders were unsuccessful. The reasons for the high variability in the breeding pattern observed in this species between years, as well as between colonies and between individuals are discussed. Breeding success was on average 30.6% and survival during the first year at sea could reach 50%. The survival of adult birds has increased during the past 10 years from 90.7% to 95.2% per annum. Despite an almost biennial breeding frequency and a very high rate of chick loss during the winter fast, the King Penguin population of Possession Island has doubled between 1966 and 1985 due to a high survival rate of adult and immature birds. The increase during the last decade in adult survival and in adult and chick condition suggests that the population increase could be the result of an improvement in food availability.

Description: A primitive equation model of an idealized ocean basin, driven by simple, study wind and buoyancy forcing at the surface, is used to study the dynamics of mesoscale eddies. Model statistics of a six-year integration using a fine grid (1/6° × 0.2°), with reduced coefficients of horizontal friction, are compared to those using a coarser grid (1/3° × 0.4°), but otherwise identical configuration. Eddy generation in both model cases is primarily due to the release of mean potential energy by baroclinic instability. Horizontal Reynolds stresses become significant near the midlatitude jet of the fine-grid case, with a tendency for preferred energy transfers from the eddies to the mean flow. Using the finer resolution, eddy kinetic energy nearly doubles at the surface of the subtropical gyre, and increases by factors of 3–4 over the jet region and in higher latitudes. The spatial characteristics of the mesoscale fluctuations are examined by calculating zonal wavenumber spectra and velocity autocorrelation functions. With the higher resolution, the dominant eddy scale remains approximately the same in the subtropical gyre but decreases by a factor of 2 in the subpolar areas. The wavenumber spectra indicate a strong influence of the model friction in the coarse-grid case, especially in higher latitudes. Using the coarse grid, there is almost no separation between the energetic eddy scale and the scale where friction begins to dominate, leading to steep spectra beyond the cutoff wavenumber. Using the finer resolution an inertial subrange with a k−3 power law begins to emerge in all model regions outside the equatorial belt. Despite the large increase of eddy intensity in the fine-grid model, effects on the mean northward transport of heat are negligible. Strong eddy fluxes of heat across the midlatitude jet are almost exactly compensated by changes of the heat transport due to the mean flow.

Description: The Ivrea‐Verbano Zone in northern Italy represents a section through the lower continental crust which has been tilted and emplaced into its present position during the Alpine orogeny. Recent and on‐going structurally‐oriented geological mapping in this region is providing new information about the geometry of the complex. The central part of the zone is dominated by a large basic complex (the ‘mafic formation') which is intrusive into the surrounding gneisses. The foliation within the envelope of gneisses is deflected around the intrusive complex as if by ballooning, but in the region south‐west of Monte Capio both units are folded together into a tight to isoclinal steeply plunging fold with an amplitude of c. 10 km. This fold locally inverts the stratigraphy of the layered basic group of the complex, and is thought to be the result of gravitational collapse following intrusion and inflation of a large magma body into the lower crust. Several high‐temperature shear zones have now been traced within the country rock for distances up to 20 km. The geometry of these, and their relationship to the basic complex suggests that at least some of the extensional collapse of the mafic body is related to uplift caused by intrusion of this body. Close parallels can be drawn between the observed structure in the Ivrea‐Verbano Zone (after removing the effects of late, low‐temperature faulting and folding related to emplacement of the rocks into their present position), and those inferred from deep seismic reflection profiling in areas of current extension such as parts of the US Basin and Range province.

Description: The development of egg/follicular cell complexes is described in maturing females of the octopus Eledone cirrhosa. Follicle cells proliferate to enclose the oocyte in a single epithelial layer which becomes deeply infolded. Active cell division of the follicle cells and recruitment of cells from an outer (thecal) layer generate this expansion of follicle cell epithelium. The onset of the main phase of vitellogenesis, secretion of protein yolk, occurs when eggs reach about 2mm in length and is marked by the columnar appearance of the follicle cells and an increased number of larger and more complex nuclei. A significant proportion of the egg population fails to develop beyond 2-3mm in length and these eggs subsequently degenerate.

Description: Shipboard bathymetry and gravity data from 30 crossings of 6 great Pacific fracture zones (FZs), the Mendocino, Murray, Molokai, Clarion, Clipperton, and Udintsev, are compared with the predictions of a model in which FZs are locked beyond the ridge-transform intersection, such that no vertical motion occurs on the fault in response to differential thermal subsidence. At least some sections of all of these FZs, except the Molokai, are consistent with this model and sustain shear stresses as high as 20 MPa. However, none of the FZs is locked along its entire length, as inferred from observed shear stresses dropping below 75% of the value necessary to maintain a locked fault. There is some suggestion that the unlocking may be related to excess volcanism.

Description: A high-resolution model of the wind-driven and thermohaline circulation in the North and equatorial Atlantic Ocean is used to study the structure and variability of the boundary current system at 26°N, including the Florida Current, the Antilles Current, and the Deep Western Boundary Current (DWBC). The model was developed by Bryan and Holland as a Community Modeling Effort of the World Ocean Circulation Experiment. Subsequent experiments have been performed at IfM Kiel, with different friction coefficients, and different climatologies of monthly mean wind stress: Hellerman–Rosenstein (HR) and Isemer–Hasse (IH). The southward volume transports in the upper 1000 m of the interior Atlantic, at 26°N, are 25.0 Sv (Sv ≡ 106m3s−1) for HR, and 34.9 Sv for IH forcing, in good agreement with the transport from the integrated Sverdrup balance at this latitude (23.9 Sv for HR, 35.6 Sv for IH). The return flow of this wind-driven transport, plus the southward transport of the DWBC (6–8 Sv), is partitioned between the Florida Current and Antilles Current. With HR forcing, the transport through the Straits of Florida is 23.2 Sv; this increases to 29.1 Sv when the wind stresses of IH are used. The annual variation of the simulated Florida Current is very similar to previous, coarse-resolution models when using the same wind-stress climatology (HR); the annual range (3.4 Sv) obtained with HR forcing is strongly enhanced (6.3 Sv) with IH forcing. The meridional heat transport at 26°N, zonally integrated across the basin, is in phase with the Florida Current; its annual range increases from 0.44 PW (HR) to 0.80 PW (IH). The annual signal east of the Bahamas is masked by strong transport fluctuations on a time scale of O(100 days), caused by an instability of the Antilles Current. By averaging over several model years, an annual cycle is extracted, which is in phase with the wind stress curl over the western part of the basin.

Description: Two extended integrations of general circulation models (GCMs) are examined to determine the physical processes operating during an ENSO cycle. The first integration is from the Hamburg version of the ECMWF T21 atmospheric model forced with observed global sea surface temperatures (SST) over the period 1970–85. The second integration is from a Max Planck Institut model of the tropical Pacific forced by observed wind stress for the same period. Both integrations produce key observed features of the tropical ocean-atmosphere system during the 1970–85 period. The atmospheric model results show an eastward propagation of information from the western to eastern Pacific along the equator, although this signal is somewhat weaker than observed. The Laplacian of SST largely drives the surface wind field convergence and hence determines the position of large scale precipitation-condensation heating. This statement is valid only in the near-equatorial zone. Air-sea heat exchange is important in the planetary boundary layer in forcing the wind field convergence but not so important to the main troposphere, which is heated largely by condensation heating. The monopole response seen in the atmosphere above about 500 mb is due to a combination of factors, the most important being adiabatic heating associated with subsidence and tropic-wide variations in precipitation. The models show the role of air-sea heat exchange in the ocean heat balance in the wave guide is one of dissipation/damping. Total air-sea heat exchange is well represented by a simple Newtonian cooling parameterization in the near-equatorial region. In the wave guide, advection dominates the oceanic heat balance with meridional advection being numerically the most important in all regions except right on the equator. The meridional term is largely explained by local Ekman dynamics that generally overwhelm other processes in the regions of significant wind stress. The principal element in this advection term is the anomalous meridional current acting on the climatological mean meridional SST gradient. The eastward motion of the anomalies seen in both models is driven primarily by the ocean. The wind stress associated with the SST anomalies forces an equatorial convergence of heat and mass in the ocean. Outside the region of significant wind forcing, the mass source leads to a convergent geostrophic flow, which drives the meridional heat flux and causes warming to the east of the main wind anomaly. West of the main anomaly the wind and geostrophic divergence cause advective cooling. The result is that the main SST anomaly appears to move eastward. Since the direct SST forcing drives the anomalous wind, surface wind convergence, and associated precipitation, these fields are seen also to move eastward.

Description: From the aerial parts of Lagotis stolonifera (Scrophulariaccae), a new phenylpropanoid glycoside, lagotoside (8), and the three known glycosides ehrenoside (5), verbascoside (= acteoside; 6), and plantamajoside (7) were isolated, together with the four known iridoid glucosides aucubin (1), catalpol(2), globularin (4), and lythantosalin (3). The structure of the new compound 8 was elucidated on the basis of chemical and spectral data as 2-(3-hy-droxy-4-methoxyphenyl)ethyl O-[α-L-arabinopyranosyl-(1 [RIGHTWARDS ARROW] 2)]-O-[α-L-rhamnopyranosyl-(1 [RIGHTWARDS ARROW] 3)]-4-O-feruloyl-β-D-glucopyranoside.

Description: The monthly mean wind stress climatology of Hellerman and Rosenstein (HR) is compared with the climatology of Isemer and Hasse (IH), which represents a version of the Bunker atlas (BU) for the North Atlantic based on revised parameterizations. The drag coefficients adopted by IH are 21% smaller than the values of BU and HR, and the calculation of wind speed from marine estimates of Beaufort force (Bft) is based on a revised Beaufort equivalent scale similar to the scientific scale recommended by WMO. The latter choice significantly increases wind speed below Bft 8, and effectively counteracts the reduction of the drag coefficients. Comparing the IH stresses with HR reveals substantially enhanced magnitudes in the trade wind region throughout the year. At 15°N the mean easterly stress increases from about 0.9 (HR) to about 1.2 dyn cm−1 (IH). Annual mean differences are smaller in the region of the westerlies. In winter, the effect due to the reduced drag coefficient dominates and leads to smaller stress values in IH; during summer season the revision of the Beaufort equivalents is more effective and leads to increased stresses. Implications of the different wind stress climatologies for forcing the large-scale ocean circulation are discussed by means of the Sverdrup transport streamfunction (ψs): Throughout the subtropical gyre a significant intensification of ψs takes place with IH. At 27°N, differences of more than 10 Sv (1 Sv ≡ 106 m3 s−1) are found near the western boundary. Differences in the seasonality of ψs are more pronounced in near-equatorial regions where IH increase the amplitude of the annual cycle by about 50%. An eddy-resolving model of the North Atlantic circulation is used to examine the effect of the different wind stresses on the seasonal cycle of the Florida Current. The transport predicted by the numerical model is in much better agreement with observations when the circulation is forced by IH than by HR, regarding both the annual mean (29.1 Sv vs 23.2 Sv) and the seasonal range (6.3 Sv vs 3.4 Sv).

Description: A low-cost underwater sound recorder has been developed and tested. It is designed to receive signals from sound sources that serve as navigation aids for RAFOS floats. This moored version of the RAFOS float (MAFOS) can monitor sound sources over many months and several hundred kilometers. It thus improves RAFOS navigation accuracy by enabling corrections for potential long-term clock drifts of the sound sources. MAFOS can also provide information on the local variation in the speed of sound due to natural hydrographic variability. In a first test, this usefulness has been proven and a warm, salty inhomogenity that traveled through a sound-source mooring array in the Iberian Basin has been observed.

Description: A mechanism exists whereby global greenhouse warning could, by intensifying the alongshore wind stress on the ocean surface, lead to acceleration of coastal upwelling. Evidence from several different regions suggests that the major coastal upwelling systems of the world have been growing in upwelling intensity as greenhouse gases have accumulated in the earth's atmosphere. Thus the cool foggy summer conditions that typify the coastlands of northern California and other similar upwelling regions might, under global warming, become even more pronounced. Effects of enhanced upwelling on the marine ecosystem are uncertain but potentially dramatic.

Description: In this paper we use the historical hydrographic data base for the South Atlantic Ocean to investigate (i) the hydrographic boundary between the subtropical gyre and the Antarctic Circumpolar Current (ACC), the Sub-tropical Front (STF), and (ii) the southern current band of the gyre, which is called the South Atlantic Current (SAC). The STF begins in the west in the Brazil-Falkland (Malvinas) confluence zone, but at locations at and west of 45°W this front is often coincident with the Brazil Current front. East of 45°W the STF appears to be a distinct feature to at least the region south of Africa, whereupon it continues into the Indian Ocean. The associated current band of increased zonal speed is the SAC, which, except for one instance, is found at or north of the surface STF until Indian Ocean water from the Agulhas retroflection is reached. A reversal of baroclinicity in the STF is observed south of a highly saline Agulhas ring, causing the SAC to separate from the STF and turn north into the Benguela Current. Zonal flow south of the STF is generally weak and serves to separate the South Atlantic and circumpolar currents. In the Argentine Basin, the SAC has a typical volume transport of 30 Sv (1 Sv = 106m3s−1) in the upper 1000 m relative to a deep potential density surface (σ4 = 45.87 kg m−3), and can be as high as 37 Sv. It is thus comparable to, or stronger than, the Brazil Current. In the Cape Basin, the transport of the SAC is reduced to about 15 SY before it turns north to feed the Benguela Current. In late 1983 this flow was joined by about 8 Sv of water from the Agulhas Current.

Description: The ECMWF-T21 atmospheric GCM is forced by observed near-global SST from January 1970 to December 1985. Its response in low level winds and surface wind stress over the Pacific Ocean is compared with various observations. The time dependent SST clearly induces a Southern Oscillation (SO) in the model run which is apparent in the time series of all variables considered. The phase of the GCM SO is as observed, but its low frequency variance is too weak and is mainly confined to the western Pacific. Because of the GCM's use as the atmospheric component in a coupled ocean-atmosphere model, the response of an equatorial oceanic primitive equation model to both the modeled and observed wind stress is examined. The ocean model responds to the full observed wind stress forcing in a manner almost identical to that when it is forced by the first two low frequency EOFs of the observations only. These first two EOFs describe a regular eastward propagation of the SO signal from the western Pacific to the central Pacific within about a year. The ocean model's response to the modeled wind stress is too weak and similar to the response when the observed forcing is truncated to the first EOF only. In other words, the observed SO appears as a sequence of propagating patterns but the simulated SO as a standing oscillation. The nature of the deviation of the simulated wind stress from observations is analyzed by means of Model Output Statistics (MOS). It is shown that a MOS-corrected simulated wind stress, if used to force an ocean GCM, leads to a significant enhancement of low frequency SST variance, which is most pronounced in the western Pacific.

Description: To study the resolving power of teleseismic P waveforms for receiver structure, we model synthetic waveforms using a time domain waveform inversion scheme beginning with a range of initial models to estimate the range of acceptable velocity structures. To speed up the waveform inversions, we implement Randall's (1989) efficient algorithms for calculating differential seismograms and include a smoothness constraint on all the resulting velocity models utilizing the “jumping” inversion technique of Shaw and Orcutt (1985). We present the results of more than 235 waveform inversions for one‐dimensional velocity structures that indicate that the primary sensitivity of a receiver function is to high wavenumber velocity changes, and a depth‐velocity product, not simply velocity. The range of slownesses in a typical receiver function study does not appear to be broad enough to remove the depth‐velocity ambiguity; the inclusion of a priori information is necessary. We also present inversion results for station RSCP, located in the Cumberland Plateau, Tennessee. Our results are similar to those from a previous study by Owens et al. (1984) and demonstrate the uncertainties in the resulting velocity estimate more clearly.