ALBERT

Description: We tested the efficacy of a range of herbicides to see if they might be potential substitutes for the use of propyzamide or glyphosate to control grass weed species that, if left unchecked, often outcompete recently planted trees in UK forests. Applications of 0.45kg a.i. ha−1 cycloxydim in late April–early May gave good control of wood small reed (Calamagrostis epigejos (L.) Roth), Yorkshire fog (Holcus lanatus L.), and wavy hair grass (Deschampsia flexuosa (L.) Trin.)) for at least 12 weeks after application. When coupled with a sequential application of 0.15kg a.i. ha−1 propaquizafop made 5 weeks later, purple moor grass (Molinia caerulea (L.) Moench) was controlled for at least 21 weeks. Sequential applications of 0.45 kg a.i. ha−1 cycloxydim in October, then 0.45 kg a.i. ha−1 cycloxydim in late April, was the most effective treatment for minimizing competition from purple moor grass, wood small reed or wavy hair grass and equally effective as propyzamide on Yorkshire fog. The herbicides carbetamide, clethodim, clodinafop-propargyl and fenoxaprop-p-ethyl were not as effective as cycloxydim. We conclude that cycloxydim is effective on a broader range of grass weeds than previously thought. Cycloxydim and propaquizafop are both highly selective and unlike the broad-spectrum herbicide glyphosate will not harm young trees, even if they are oversprayed when in active growth. If applied at the correct time of year, cycloxydim is likely to be as effective as glyphosate on most grasses, and for wood small reed, purple moor grass and wavy hair grass considerably more effective than propyzamide. A sequential application of cycloxydim or possibly propaquizafop in the autumn, followed by a second spray in late April of the following spring, is likely to be the most effective means of maximizing tree growth and survival in grass dominated new planting or restock sites in UK forests.

Description: For the first time, an economic analysis of a fully simulated forest transformation process from even-aged to uneven-aged stand structure using natural regeneration was performed using an individual tree growth simulator. A comparison to a ‘business as usual’ management approach (i.e. age-class scenario) was also made. Norway spruce (Picea abies (L.) Karst.)-dominated forests of different ages were selected for the simulation: forest at thinning stage (52 years; hereafter, ‘younger stand’) and a mature stand (95 years) to explore different starting points for the transformation. The harvested and remaining stand volume showed only little fluctuation over the course of the simulation period in the case of the transformation (uneven-aged) scenario. The age-class scenario, on the other hand, showed peaks and drops in the timber volume. The land expectation values at the end of transformation were higher (i = 1 or 2 per cent) or equal (i = 3 per cent) to those of the age-class scenario for the younger stand. The transformation of younger forests appears to be economically interesting (under interest rates above 2 per cent) as the early revenues of more intensive thinnings achieve an equilibrium growing stock well below the potential maximum growing stock of an even-aged stand. This suggests transformation management to be economically efficient under the given silvicultural and economic conditions. The transformation of mature stands was found to be less economically viable due to the lower holding values for the transformation (uneven-aged) scenario compared with the age-class scenario. However, the holding values of the younger stand were higher for all interest rates compared with the age-class scenario. This study concludes that transformation (uneven-aged) scenario yielded more economically viable silvicultural approach (in the case of younger forests) and a steadier trend in harvested and remaining timber volume showing no major fluctuations. Transformation approach also forms a better foundation where other ecosystem services can be built.

Description: Bush encroachment affects ~45 million ha of Namibia and, without appropriate restoration measures, it negatively affects rangeland productivity and biodiversity. Thinning is a common method to counteract bush encroachment. The thinning strategy applied in north-central Namibia was assessed to examine how effective it has been in reducing bush encroachment. Trees/shrubs were selectively thinned manually, targeting all height classes, except individuals with stem diameters ≥18 cm. We investigated the effects on the vegetation and soil properties using surveys on three freehold farms (in 2016 and 2017) in bush-encroached and previously thinned habitats. Our results revealed significant differences in the mean total nitrogen (TN) content between the treatments; thinned areas had higher TN content which would be beneficial for fast-growing grasses. In the thinned plots, the occurrence probability of red umbrella thorn (Vachellia reficiens Warwa) was significantly reduced, indicating that it was the most harvested species; and umbrella thorn (Vachellia tortilis (Burch.) Brenan spp. heteracantha) was increased, indicating that it favoured reduced densities of dominant species. Natural regeneration was rapid; the tree/shrub abundance in the 0–1-m height class in the thinned area surpassed those in the non-thinned by 34 per cent, ~7.2 years since thinning. Thinning significantly reduced tree/shrub abundances of the 1–3- and 〉3-m height classes, which was still evident 7.2 years since thinning. Based upon the generalized linear mixed-effects model, tree/shrub counts between treatments may equalize in ~14 and 15 years for the 1–3- and 〉3-m height classes, respectively. Thinning was effective in reducing tree/shrub abundances and can be used to restore wildlife habitat on the Namibian farmland: however, post-thinning management is required to maintain an open savannah vegetation structure as the 0–1-m height class cohort will eventually grow into mature trees/shrubs.

Description: Norway spruce (Picea abies Karst. (L.)) in the boreal zone can be managed as even-aged or uneven-aged stands, or be grown with no management at all. Here, we investigated how these management options affect carbon dynamics, particularly the carbon stocks in the forest ecosystem (trees and soil), and albedo, and their combined effect on radiative forcing compared to a reference case, clear-cut site before planting seedlings. This allowed us to assess the potential of different management regimes to mitigate global warming. We ran long-term simulations under the current climate on a sub-mesic site in central Finland (62oN) using an eco-physiological forest-ecosystem model. Compared to even-aged management, no management (old-growth forest) increased ecosystem carbon stocks by 47 per cent and decreased albedo by 15 per cent, whereas uneven-aged management reduced ecosystem carbon stocks by 16 per cent and increased albedo by 10 per cent. Only the no management option resulted in a significant net cooling effect whereas for even-aged and uneven-aged management, the opposing effects of changes in albedo and carbon stocks largely cancelled each other with little remaining net effect. On the other hand, the latter one even made a small net warming contribution. Overall, maintaining higher ecosystem carbon stocks implied the larger cooling benefits. This was evident even though lower albedo enhanced radiation absorption, and thus warming. Increasing use of the no management option by forest owners may require proper incentives such as compensation for lost harvest incomes.

Description: Climate change is projected to have a major influence on forest tree populations and composition. Translocation of species outside their historic range has been suggested to maintain healthy forests and tree species. The introduction of exotic species into botanical gardens and arboretums worldwide demonstrates the ability of many trees to grow outside their natural habitat and may play an important part in avoiding climate driven extinction if grown in a matching climate. However, it remains to be determined which climatic factors are the most important predictors of climatic match. In this study we use information from the arboretum in Hørsholm, Denmark, to analyse differences in performance of translocated Oak (Quercus) and show how data from tree collections can be used to predict success of assisted migration. Our data included archive lists of georeferenced Northern hemisphere introductions of Quercus, and assessments of their survival and growth rates in nursery and the Hørsholm arboretum. Using logistic and linear regression we modelled the importance of different bioclimatic predictor variables for survival and growth rate. Several correlations were identified across the Quercus genus. Survival of Quercus species depended primarily on the temperatures at the origin, whereas growth on the other hand was more dependent on a match in precipitation. The negative correlations indicated that introductions were less successful from sites with higher temperatures and wetter conditions. The study demonstrates an approach to use historical data collected from arboreta and botanical gardens in climate change research. This new approach can provide useful information in relation to assisted migration for an array of poorly investigated species where this may be the only source of information.

Description: In this study we investigated the behaviour of aggregate prediction errors in a forest inventory augmented with multispectral Airborne Laser Scanning and airborne imagery. We compared an Area-Based Approach (ABA), Edge-tree corrected ABA (EABA) and Individual Tree Detection (ITD). The study used 109 large 30 × 30 m sample plots, which were divided into four 15 × 15 m subplots. Four different levels of aggregation were examined: all four subplots (quartet), two diagonal subplots (diagonal), two edge-adjacent subplots (adjacent) and subplots without aggregation. We noted that the errors at aggregated levels depend on the selected predictor variables, and therefore, this effect was studied by repeating the variable selection 200 times. At the subplot level, EABA provided the lowest mean of root mean square error ($overline{mathrm{RMSE}}$) values of 200 repetitions for total stem volume (EABA 21.1 percent, ABA 23.5 percent, ITD 26.2 percent). EABA also fared the best for diagonal and adjacent aggregation ($overline{mathrm{RMSE}}$: 17.6 percent, 17.4 percent), followed by ABA ($overline{mathrm{RMSE}}$: 19.3 percent, 18.2 percent) and ITD ($overline{mathrm{RMSE}}$: 21.8, 21.9 percent). Adjacent subplot errors of ABA were less correlated than errors of diagonal subplots, which resulted also in clearly lower RMSEs for adjacent subplots. This appears to result from edge tree effects, where omission and commission errors cancel for trees leaning from one subplot into the other. The best aggregate performance was achieved at the quartet level, as expected from fundamental properties of variance. ABA and EABA had similar RMSEs at the quartet level ($overline{mathrm{RMSE}}$ 15.5 and 15.3 percent), with poorer ITD performance ($overline{mathrm{RMSE}}$ 19.4 percent).

Description: Water stress is the most important factor limiting early survival and growth of seedlings in arid and semi-arid zones. Many woody species develop adaptive mechanisms in response to water stress. This study assesses survival rate, growth and dry matter distribution in response to water stress in four dryland species (Senegalia dudgeonii, Senegalia gourmaensis, Vachellia nilotica and Vachellia tortilis). A total of 240 seedlings (60 per species) were grown using a completely randomized block design with three replicates in each of three water treatments (control = 100 percent of field capacity; moderate water stress = 50 percent of field capacity; severe water stress = 25 percent of field capacity) with a 7 day watering frequency. Data were collected on seedling survival, height, collar diameter, leaf production and dry matter distribution. Survival rate of the four species was not affected by water stress. However, the morphological responses to water stress were significantly different between species. Water stress resulted in significant reductions in height and collar diameter, higher below-ground biomass and significant increase in shedding of leaves. Under water stress, S. dudgeonii, S. gourmaensis and V. tortilis invested more in root growth. The allocation pattern from above-ground to below-ground parts was found to be the main adaption to drought. In contrast, V. nilotica gave preference to above-ground development, which could be an indication that the seedlings are adapted to grow under severe water stress. The results confirm the interspecific genetic differences in growth and dry matter distribution among species. These variations in water stress response may be used as criteria for species selection for degraded land reforestation.

Description: Armillaria (Armillaria solidipes) and laminated root diseases (Inonotus sulphurascens) are two wide-ranging fungal pathogens that occur in the southern half of British Columbia (BC), Canada, and can infect economically and biologically important tree species such as interior Douglas-fir (Pseudotsuga menziesii var. glauca). In northern, temperate locations, Douglas-fir forests serve as winter habitat for ungulates. When these fungal infections are in Douglas-fir forests, core components of winter ranges are altered, including canopy cover, snowpack and understory vegetation. In this study, we investigated how Rocky Mountain mule deer (Odocoileus hemionus hemionus) of central BC used winter range habitats that included root disease (A. solidipes and I. sulphurascens) centres. We used remote camera-traps to collect data from September 2017 to April 2019, and we assessed those habitats in which the cameras were located during the summer and winter of 2018. We used logistic regression and an information theoretic approach to test a series of factors hypothesised to influence the use of root disease centres by mule deer. Our results show that mule deer use root disease centres less than control forests as well as negatively respond to the deeper snow packs found in root disease centres, especially in late winter. Our cameras also detected higher vertebrate diversity in root disease centres. We suggest that forest policy-makers should acknowledge heterogeneous habitat features such as root disease centres within ungulate winter ranges and consider adjusting estimates of habitat capability for deer based on our findings.

Description: Determining forest volume increment, the potential of wood production in natural forests, is a complex issue but is of fundamental importance to sustainable forest management. Determining potential volume increment through growth and yield models is necessary for proper management and future prediction of forest characteristics (diameter, height, volume, etc.). Various methods have been used to determine the productive capacity and amount of acceptable harvest in a forest, and each has advantages and disadvantages. One of these methods involves the artificial neural network techniques, which can be effective in natural resource management due to its flexibility and potentially high accuracy in prediction. This research was conducted in the Ramsar forests of the Mazandaran Province of Iran. Volume increment was estimated using both an artificial neural network and regression methods, and these were directly compared with the actual increment of 20 one-hectare permanent sample plots. A sensitivity analysis for inputs was employed to determine which had the most effect in predicting increment. The actual average annual volume increment of beech was 4.52 m3ha−1 yr−1, the increment was predicted to be 4.35 and 4.02 m3ha−1 yr−1 through the best models developed using an artificial neural network and using regression, respectively. The results showed that an estimate of increment can be predicted relatively well using the artificial neural network method, and that the artificial neural network method is able to estimate the increment with higher accuracy than traditional regression models. The sensitivity analysis showed that the standing volume at the beginning of the measurement period and the diameter of trees had the greatest impact on the variation of volume increment.

Description: Efficient subsampling designs reduce forest inventory costs by focusing sampling efforts on more variable forest attributes. Sector subsampling is an efficient and accurate alternative to big basal area factor (big BAF) sampling to estimate the mean basal area to biomass ratio. In this study, we apply sector subsampling of spherical images to estimate aboveground biomass and compare our image-based estimates with field data collected from three early spacing trials on western Newfoundland Island in eastern Canada. The results show that sector subsampling of spherical images produced increased sampling errors of 0.3–3.4 per cent with only about 60 trees measured across 30 spherical images compared with about 4000 trees measured in the field. Photo-derived basal area was underestimated because of occluded trees; however, we implemented an additional level of subsampling, collecting field-based basal area counts, to correct for bias due to occluded trees. We applied Bruce’s formula for standard error estimation to our three-level hierarchical subsampling scheme and showed that Bruce’s formula is generalizable to any dimension of hierarchical subsampling. Spherical images are easily and quickly captured in the field using a consumer-grade 360° camera and sector subsampling, including all individual tree measurements, were obtained using a custom-developed python software package. The system is an efficient and accurate photo-based alternative to field-based big BAF subsampling.

Description: There is increasing interest across Europe in adopting forest management strategies, which promote species and structural diversity through the use of irregular silvicultural systems, an approach often described as continuous cover forestry (CCF). However, there is little information about the proportion of CCF practised across the continent or about the knowledge gaps and other obstacles that limit its use. A survey of respondents in 33 European countries sought to address these issues. The results indicated that the silvicultural systems associated with CCF were single stem, group selection and irregular shelterwood. Rotational forest management (RFM) was more frequent than CCF in about 66 per cent of countries, whereas in 25 per cent the reverse was true. We estimated that between 22 and 30 per cent of European forests are managed through CCF, although good data are lacking. The main knowledge gaps were: uncertainties arising from climate change (e.g. appropriate species choice, carbon storage in CCF), using CCF to increase forest resilience, deployment of mechanized harvesting systems, lack of knowledge about CCF amongst professional foresters and better information on economic implications of this approach. Major obstacles included: little awareness of CCF amongst forest owners, limited competence in CCF within the forestry profession and a scarcity of skilled forest workers to implement this approach, high ungulate populations damaging natural regeneration, a sawmilling sector geared to processing medium-sized logs, subsidy regimes favouring practices associated with RFM and a lack of experience in transforming plantation forests to more diverse structures. Better information on the use of different silvicultural systems is essential to allow policymakers and other stakeholders to monitor progress in diversifying forests. Establishment of a continental network of long-term operational trials (e.g. expanding the existing Association Futaie Irrégulière network) would improve professional understanding of CCF, would demonstrate this approach to forest owners and other stakeholders and could provide a valuable platform for supporting research.

Description: The translation of genetic gains into economic gains is important for evaluating the impact of using genetically improved forest reproductive material (FRM) in the forest sector. However, few studies based on European forests have been published to date. Here, we analyse the stand-level wood production and financial performance of planting genetically improved FRM in even-aged planted forests focusing on four European case studies with advanced breeding programme material and different management contexts: Scots pine (Pinus sylvestris L.) in southern Finland, central Sweden and central France, and maritime pine (Pinus pinaster Ait.) in southwestern France. The growth of improved stands was simulated using species-specific growth models by incorporating two levels of expected genetic gains (present and next generations of seed orchards, varying from 7 to 40 per cent depending on the breeding programme) into the estimated mean annual volume increment over a rotation (m3 ha−1 yr−1). For each level of genetic gain, we tested the plantation of improved FRM managed with two silvicultural scenarios (maintaining the standard baseline rotation and thinning regime vs shorter rotation through the earlier achievement of the recommended felling criteria) in comparison with the plantation of the reference unimproved material (absence of genetic gain) managed according to the standard silvicultural regime. The use of improved FRM resulted in a larger financial performance in terms of soil expectation value (SEV € ha−1, discount rate 3 per cent) than planting unimproved reference material in all case studies and silvicultural scenarios for different wood price contexts (SEV gain from +20 to +190 per cent depending on the genetic and silvicultural context). The challenges associated with the economic assessment of realized gains from genetically improved FRM are discussed. We argue that silvicultural guidelines should be adapted to the use of improved FRM in order to gain better financial performance and flexible silvicultural response of planted forests to future environmental and socio-economic changes.

Description: Wild deer exert strong top–down control on forest composition by browsing on palatable trees, and these effects are exacerbated as red, fallow, and roe deer populations increase in northern temperate forests. However, the relationship between deer abundance and plant recruitment remains poorly documented. Here, we combined camera trap and vegetation plot data to quantify the shape of the relationship between habitat utilization by deer (red, fallow and roe deer) with different components of the forest understory in ten sites distributed across a temperate mixed forest in the Veluwe, the Netherlands. The list of forest attributes included the density, richness and diversity of saplings, the proportion of conifers to broadleaves, the number of browsed broadleaves and conifers, the forest basal area, the understory cover of shrubs, moss and bare soil and the depth of the litter layer. When applying General Linear Mixed Models to those camera trap data, six of the eleven forest variables were related to the utilization level (UL) by deer. With increasing UL by fallow deer, there was a decrease in sapling species richness (β = −0.26). With increasing UL by red deer, there was a decrease in litter depth (β = −0.14) and an increase in browsed broadleaf stems (β = 0.40). With increasing UL by roe deer, there was a decrease in species richness of sapling plants (β = −0.26), Shannon diversity (β = −0.11) and shrub cover (β = −0.36), whereas there was an increase in stem density (β = 0.06). When combining all deer species into one guild, a negative relation was found between UL by deer and sapling richness (β = −0.21), diversity (β = −0.09) and litter depth (β = −0.14) and a positive relation with sampling density (β = 0.04). The relationship between UL by deer and different forest attributes followed a linear log10 shape. When back transforming, the response was curvilinear with large changes at a low UL and small changes when the UL was high. Yet, the exact shape of the curve varied according to (a) biotic factors specific to each study location. The conservation of temperate forests can benefit from limiting the size of deer populations and the time deer spend browsing in forest patches with high biodiversity value.

Description: A prerequisite for successful afforestation and reforestation is understanding the quality of a site before establishing a specific tree species. Ecograms have been widely applied to determine the suitability of different sites for different species by a simple assessment of nutrient and water availability. Their graphical representation of suitability into classes allows ecograms to be easily understood. However, ecograms have generally been mapped for small areas only and their validity has rarely been documented. The aim of this study is to map ecograms for five tree species across Denmark and validate the maps using forest stand production data. For this purpose, we classified the landscape into six nutrient classes and nine water classes based on four variables to generate the ecogram maps. Based on these classes, the generated maps depict if a tree species is unsuitable, suitable or optimal for a specific site. The absolute average misclassification for nutrient and water supply was 1 and 2 classes, respectively. Stepwise linear regression was implemented to determine if the four variables used to create the ecograms were able to predict production as observed from forest experiment and management planning data obtained from across the country. All five species used the full model to explain variation in production. However, the average production values per ecogram growing condition were not significantly different for all species. The range of $R$2 for the five species was 0.05–0.32, indicating that one ecogram template might not work for all species. The high-resolution national ecogram maps incorporate large-scale variables important for tree growth and will be beneficial when selecting new land for afforestation. The simplicity of ecograms allows for easy interpretation, meaning that foresters can quickly determine which regions of a landscape are suitable, saving time and resources.

Description: Bottomland hardwoods forests (BLHW) are increasingly subject to active management for water quality improvement, streambank stabilization, to mitigate for urban expansion and to improve habitat for wildlife. Since the majority of remaining BLHW are degraded, restoration attempts are becoming commonplace within the Western Gulf Coastal Plain (WGCP) of the US. However, restoration success in terms of obtaining high stem densities of desirable species has been mixed, with managers observing survival rates

Description: Productive Norway spruce forests in mountainous growing areas are expected to be affected by increasing climate-driven disturbances, which will most likely result in a reduction of their economic performance. This study seeks to compare the potential of currently discussed management options aiming to maintain the economic performance on typical sites of spruce under a changing climate. For this purpose, we used a combined approach of tree mortality simulation and portfolio optimization to assess the management options: (1) including a deciduous species, (2) establishing mixed stands, (3) shortening the rotation periods, (4) forest protection measures focusing on bark beetles and (5) introducing another highly productive species. As a novel feature, we also compared the effect of combined management strategies on the return and risk of stand management. In our simulation and optimization approach, the inclusion of an alternative, highly productive tree species, such as Douglas fir, into the tree species portfolio outcompeted the forest protection measure of intensive bark beetle monitoring with timely sanitation fellings in terms of balancing risks and returns. The adaptation gain of introducing Douglas fir was further enhanced when combined with additional management options such as adapted rotation periods. The resulting synergies were able to overcompensate for the adverse economic effects of climate change. On the contrary, including beech did not only reduce the risks but also the return in a magnitude which was clearly outcompeted by Douglas fir, even with the assumption of high mortality risks for Douglas fir. These results prove the importance of a proactive tree species selection in relation to more reactive forest protection measures for climate change adaptation. They also advise a careful consideration of expected market demands when selecting an additional species for the portfolio.

Description: The pattern of change in carbon (C) accumulation with forest ageing can vary greatly amongst different forest types. Documenting how C accumulates in various forest ecosystems in the absence of logging makes it possible to predict what would be the outcome of extending forest rotations or in dedicating more land to conservation on C storage. This study was conducted in boreal balsam fir (Abies balsamea (L.) Mill.) forests of Quebec, in eastern Canada. We compared carbon stocks in forest pools (aboveground (live) biomass, deadwood, FH horizon and mineral soil) of mature (70 years after harvest) vs old-growth stands (stands with no signs or history of human disturbance). Total ecosystem C stocks were not significantly different between mature and old-growth stands. However, as mature stands transition to old-growth stage, there appears to be a shift of C from live biomass pools towards deadwood and soil FH horizons. Coarse woody debris in old-growth stands were also found to be at more advanced stages of decay. The variability of C stocks was also high amongst old-growth stands; however, there was no obvious difference in structural diversity between mature and old-growth stands. Results suggest that ageing balsam fir stands through the lengthening of forest rotations (e.g. past the maturity age of 70 years) or by placing them under conservation, while not creating an important C sink, still contributes to maintain large forest C stocks across landscapes. Preserving or increasing the presence of old-growth forests is an important aspect of ecosystem-based forest management; our study concludes that it could also be compatible with sustainable forest carbon management.

Description: Climate change is predicted to increase temperature and seasonal temperature variance in Great Britain (GB). Sitka spruce (Picea sitchensis (Bong.) Carr) is the most important tree species used in commercial plantations throughout Europe and GB. Frosts that occur outside the winter dormancy period can negatively affect trees, since they happen after dehardening. Damage can be especially severe at bud burst, before emerging needles mature and form protective barriers. Here, we modelled the impact of climate change on frost sensitivity in Sitka spruce with temperature data from five climate projections. The UKCP09 climate model HadRm3 uses emission scenario SRESA1B for the years 2020–2099. The global and downscaled versions of the UKCP18 HadGem3 model use the emissions scenario RCP 8.5. The global model CMCC-CM uses the RCP 4.5 and RCP 8.5 emissions scenarios. The predictions based on these models were compared with results from gridded historical data for the period 1960–2015. Three indicators that assessed the frost sensitivity of Sitka spruce were explored: the total number of frosts between the onset of dehardening and the end of summer, which use three different temperature thresholds (Index 10°C, 1–3°C, 1–5°C); the total number of frosts after bud burst (Index 2); the number of days with minimum temperatures below the resistance level (backlashes) during the hardening–dehardening period (September–August) (Index 3). The indices were validated with historical data for frost damage across GB, and Index 1–3°C, Index 1–5°C and Index 3 were shown to be significantly correlated. The frequency of all frosts and backlashes is expected to decrease with climate change, especially under higher emissions scenarios. Post-bud burst frosts have been historically very rare in GB and remain so with climate change. Downscaled regional climate models detect geographic variability within GB and improve prediction of overall trends in frost damage in comparison to global climate change models for GB.

Description: Quality information on forest resources is fundamental for sustainable forest management. Manual aerial photointerpretation is used as a cost-effective source of data for forest inventories; however, the process of photointerpretation is inherently subjective and is often undertaken by multiple photointerpreters for a given forest management area. In contrast, airborne laser scanning (ALS) data enable characterization of forest structure in a systematic fashion with quantifiable levels of accuracy and precision that often exceed required targets and standards. However, the gains associated with the use of new technologies for forest inventory are difficult to measure because the quality of existing photointepreted inventories have rarely been quantified. Using ALS data as reference, the objective of this study was to quantify the precision of photointerpreted estimates of forest stand height and canopy cover (CC). We examined forest inventories from three study sites in three different forest regions of Canada. Each of the study sites was located within a different provincial jurisdiction with unique photointerpretation standards and forest ecosystems. Stand-level estimates of forest height and cover were compared to reference estimates generated from the ALS data. Overall, our results indicated that precision was greater for photointerpreted estimates of height, with a relative standard deviation ranging from 22 per cent to 29 per cent among our three sites, compared to estimates for CC, with precision ranging from 28 per cent to 59 per cent. While the relationship between photointerpreted estimates of height and ALS estimates of height were generally linear and consistent for all study sites, relationships for CC were non-linear. We found that precision for both stand height and cover varied by dominant species, inventory stand structure, age, and ALS canopy complexity, and that in the majority of cases, the differences between the photointerpreted estimate and the ALS estimate were statistically significant. It is also noted that the variability in photointerpretation precision as a function of the aforementioned factors was not consistent among our three study sites, indicating that site-specific forest conditions and photointerpretation procedures influence the precision of photointerpreted estimates. The influence of local forest conditions and interpretation procedures are therefore important considerations when seeking to quantify the potential relative gains in precision, which may be afforded by technologies such as ALS for forest inventory programs. Moreover, approaches to improve consistency in photointerpreted estimates of cover would be useful for operational inventory programs.

Description: Simplified forest structures following even-age management have been associated with the loss of biodiversity, which may be avoided through disturbance-inspired silviculture. Here, we ask how much do gap characteristics in a managed old-growth differ from those in unmanaged old-growth subject only to natural dynamics? In this study, we compared important characteristics of gaps (e.g. canopy gap fraction, distribution of gap sizes) and gapmakers (e.g. size classes, frequency, decay classes) between a managed and an adjacent unmanaged old-growth Oriental beech (Fagus orientalis Lipsky) compartment in the Keladarsht region of northern Iran 10 years after a single harvest entry using single-tree selection. Canopy openings 〉100 m2 with visible remnants of gapmakers (i.e. stumps) were included in this study. Gap characteristics of both compartments were within typical ranges for old-growth beech. Nonetheless, small but potentially important differences between the two areas were observed. In the managed compartment, harvesting poor quality trees with structural defects and typical diameters at breast height 〉52.5 cm plus natural mortality resulted in 102 canopy gaps (1–6 gapmakers, averaging 3.5 gaps/ha, gap fraction 9.8 per cent) compared with 59 natural canopy gaps (1–7 gapmakers, averaging 2.6 gaps/ha, gap fraction 13.7 per cent) in the unmanaged compartment. In both compartments, medium-sized gaps (200–500 m2) were most prevalent. In the managed compartment, 60 per cent of gapmakers were large or very large (typically cut) compared with 39 per cent in the unmanaged compartment where large trees typically snapped and became snags. Uprooting, particularly of small and medium sized gapmakers, was less common in the managed than the unmanaged compartment. Our results indicate that even one single-tree selection harvest may lead to a short-term divergence in stand structure compared with the unmanaged forest. While such managed forests may no longer be considered as old-growth, divergences in canopy gap characteristics indicate that a more nuanced harvesting scheme that includes cutting some larger gaps may more closely mimic the canopy dynamics of this old-growth forest.

Description: The European and Mediterranean Plant Protection Organization (EPPO) real-time PCR protocol for the detection of ash dieback (Hymenoscyphus fraxineus (T. Kowalski) Barak, Queloz, Hosoya) was optimized for diagnostics on ash (Fraxinus excelsior L.) fruits. Ash fruits were collected from a range of sites in Britain, from trees with various levels of infection. To assess the potential pathway and the presence of H. fraxineus, tissue components of ash fruits as well as ash flowers were tested for the presence of H. fraxineus DNA and RNA. The fungus was detected on ash fruits from trees on all five sampled sites including symptomless, lightly and highly infected sites. DNA of H. fraxineus was detected on the pericarps of all fruit lots and on seeds from fruit lots that included damaged and empty fruits, but not on embryos or flowers. RNA of H. fraxineus was never detected on any of the samples or sample types, indicating that H. fraxineus was only detected in an inactive form (i.e. dormant or dead). The absence of RNA from the pathogen suggests that only spores of H. fraxineus are present on ash fruits. A double hot-water treatment was evaluated as a control measure for the eradication of H. fraxineus on ash fruits; these were treated before and after warm stratification. Large proportions of ash fruits survived the double hot-water treatments, with a 60.4 ± 28.4 per cent germination success rate depending on crop year. Double hot-water treatment of ash fruits proved to be an efficient, low cost and low-tech approach for the eradication of H. fraxineus on ash fruits.

Description: Many non-wood forest products (NWFPs) such as mushrooms and berries are collected and consumed in Europe; but both national statistical and scientific data on this topic are reported only for a limited number of countries, products and case-study areas. Without an adequate quantitative basis, their importance as source of food and income, their links to recreation and cultural heritage, are all under-valued in forest-focused and forest-related policies. In this study, we aimed to address this gap by assessing the consumption and collection of NWFPs through a statistically representative survey in 28 European countries with over 17 000 respondents. Our results show that 90 per cent of households consume NWFPs and about one-quarter collects them. The consumption and collection rates, as well as the number of collected products and their contribution to household income, increase from the west to east of Europe. The vast majority of collected products are consumed fresh. Households with higher income consume a more diverse range of NWFPs, especially in Western Europe. The relation between income and collection is more ambiguous, but there is some indication that the collection rate is higher than average among higher-income households in North and Western Europe and among lower-income households in Eastern and South-Eastern Europe. Households for which NWFP collection is the main income source are predominantly located in Eastern Europe, and they focus their activities on few key products. Our results also identify recreational, hobby and professional collectors whose characteristics vary across socio-economic variables and geographical gradient. Recreational collectors in Western and Southern Europe collect 8 kg of NWFPs from five different products, while recreational collectors in Central-Eastern and North-Baltic Europe collect about four times more from 10 different products. Hobby collectors collect ~100 kg of NWFP per year and professional collectors half a ton, where both groups focus on 8–12 different products. Professional collectors are predominantly located in Eastern and South-Eastern Europe. We end the study by pointing to future research directions and with a series of policy recommendations on how NWFPs could be addressed along the geographical, income and urban–rural gradient with respect to their role in forest recreation as a food and income source.

Description: Tree neighbourhood conditions vary greatly through time for individual trees and throughout a forest at any given time. Therefore, many trees experience very different neighbourhood conditions to the stand mean. Although changes in stand density or species composition are often recommended to mitigate the effects of global change on forest functioning, these recommendations are often based on stand means and lack guidance on acceptable variability around the means or acceptable ranges in neighbourhood conditions. This study examined how the spatial and temporal variability in tree neighbourhood conditions varies between species and with stand structural characteristics. A trees neighbourhood, defined by all trees within a 10-m radius, was quantified in terms of basal area GN, relative height rhN (tree height divided by mean height of all trees in neighbourhood), species proportion PropN and species richness RN. Weibull functions were used to describe the frequency distributions of GN, rhN, PropN and RN within the stands. Equations were developed to describe the Weibull shape, scale and location parameters as functions of the species, stand basal area, mean tree diameter and management (even-aged vs single-tree selection forests). All of these variables significantly influenced the frequency distributions of neighbourhood characteristics. These equations can be used to show the proportion of trees experiencing significantly higher or lower values of a given characteristic than the stand mean or other threshold values relevant for management objectives. Single-tree selection forests had much greater ranges in neighbourhoods. There were also large temporal changes in tree neighbourhoods, especially in single-tree selection forests. Large trees had not experienced better neighbourhood conditions than smaller trees of approximately the same age, when considered over the long-term. Given that foresters typically consider tree neighbourhoods when marking trees for thinning, and that trees respond to tree neighbourhood conditions rather than to mean stand conditions, stand-level recommendations may be easier to transfer into practice and to interpret, when accompanied by neighbourhood-level information that indicates the distribution of neighbourhoods that actually exist within a stand.

Description: Field trials containing 8–10 sycamore seed sources were established on ‘farm woodland’ sites in 1992 to identify the most productive and adapted seed sources for use in lowland Great Britain. Early results from these trials found little difference among provenances for growth traits. To guide the next steps of sycamore improvement in the British Isles, three of the five original trials have been revisited and assessed for variation in growth and form at 27 years, which is between one-third and half rotation age. There were large differences in growth among sites but differences among provenances were small or non-significant and showed no interaction across sites. Spring phenology evaluation from one of the trials demonstrated that timing of bud burst was negatively associated with latitude which was the only trait providing any evidence of adaptive differentiation among provenances. However, the effect was small and driven by much later flushing of French and German seed sources than British seed sources. Given the lack of clear provenance differentiation and qualitatively similar rankings across sites, we found no justification for subdividing Britain into breeding zones for sycamore at this stage, nor for excluding material from mainland Europe in planned progeny tests.

Description: White birch (Betula platyphylla Sukaczev) and costata birch (Betula costata Trautv.) are valuable hardwood tree species growing in northeast China. Several taper studies have analysed birch species in the countries harbouring the boreal forests. However, this study presents an initial attempt to develop stem taper models using the fixed- and mixed-effects modelling for white birch and costata birch in Xiaoxing’an Mountains, northeast China. Ten commonly used taper models were evaluated by using 228 destructively sampled trees of both tree species comprising of 4582 diameter and height measurements. The performance of these models was tested in predicting diameter at any height, total volume and merchantable volume (10 and 20 cm top diameters). We incorporated a second-order continuous-time error structure to adjust the inherent autocorrelation in the data. The segmented model of Clark best predicted the diameter and total or merchantable volume when the upper stem diameter at 5.3 m was available. When diameter measurements at 5.3 m were not available, the models of Kozak and Max and Burkhart were superior to other models for white birch and costata birch, respectively. After model comparison, the best model of Clark was refitted as the NLME model.

Description: Reforestation in the British Isles (UK and Ireland) has been dominated with the use of an exotic conifer tree species, Sitka spruce (Picea sitchensis [Bong.] Carr.). Sitka breeding in the UK was developed from a single provenance, the Haida Gwaii Islands (Canada), which is both well suited to the British climate and highly susceptible to the white pine weevil (Pissodes strobi L.) in its native range. We examined variation and heritability of insect resistance related traits and assessed potential trade-offs with tree growth in 50 full-sibling families and 13 clonally replicated genotypes growing in the UK. We measured bark levels of three terpenes (dehydroabietic acid, (+)-3-carene and terpinolene) shown to confer resistance to the white pine weevil in Sitka spruce’s native range, on the principle that these defence compounds may also contribute to pest resistance in the UK. We compared our results with published findings from the native range and also used individuals from a Haida Gwaii seed lot grown in the UK for comparison of terpene levels. Dehydroabietic acid content in the UK breeding population was similar to populations from resistant native populations, but (+)-3-carene and terpinolene levels were relatively low. Narrow sense heritability for dehydroabietic acid, (+)-3-carene and terpinolene was estimated as 0.20, 0.93 and 0.98, respectively from the full-sib data, and this evidence of genetic variance was supported by estimates of broad sense heritability from the smaller clonal study. Terpene content was found to be positively correlated to growth traits. The heritability estimates and genetic correlations indicate that selective breeding should be effective in raising levels in the UK breeding population of the three candidate terpenes implicated in weevil resistance. However, low levels observed indicate that other provenances from the native range may produce greater short-term improvements for two of the terpenes.

Description: Airborne laser scanning (ALS) is one of the most innovative remote sensing tools with a recognized important utility for characterizing forest stands. Currently, the most common ALS-based method applied in the estimation of forest stand characteristics is the area-based approach (ABA). The aim of this study was to analyse how three ABA methods affect growing stock volume (GSV) estimates at the sample plot and forest stand levels. We examined (1) an ABA with point cloud metrics, (2) an ABA with canopy height model (CHM) metrics and (3) an ABA with aggregated individual tree CHM-based metrics. What is more, three different modelling techniques: multiple linear regression, boosted regression trees and random forest, were applied to all ABA methods, which yielded a total of nine combinations to report. An important element of this work is also the empirical verification of the methods for estimating the GSV error for individual forest stand. All nine combinations of the ABA methods and different modelling techniques yielded very similar predictions of GSV for both sample plots and forest stands. The root mean squared error (RMSE) of estimated GSV ranged from 75 to 85 m3 ha−1 (RMSE% = 20.5–23.4 per cent) and from 57 to 64 m3 ha−1 (RMSE% = 16.4–18.3 per cent) for plots and stands, respectively. As a result of the research, it can be concluded that GSV modelling with the use of different ALS processing approaches and statistical methods leads to very similar results. Therefore, the choice of a GSV prediction method may be more determined by the availability of data and competences than by the requirement to use a particular method.

Description: The health of Corymbia calophylla (marri), a keystone tree species in the native forests of southwest Western Australia, has been in decline for the past few decades. Phytophthora root disease and waterlogging have often been cited as contributing to this decline. Traditional methods (i.e., field surveys and sampling) of mapping Phytophthora root infection in the field are time-consuming and expensive; thus, the potential of reflectance spectroscopy to characterize marri response to Phytophthora and waterlogging stress was investigated. Twelve-month old marri plants were infected with either P. cinnamomi or P. multivora in two glasshouse trials and waterlogged for 24 h each fortnight. Spectral measurements with a portable high-resolution spectroradiometer were taken weekly. Plant biophysical measurements were taken at harvest time. Normalized difference spectral index (NDSI) was calculated for every combination of reflectance values between 400 and 2500 nm for all time points, correlated with the treatment effects and displayed as heat maps. Narrowband vegetation indices (VIs), utilizing different wavelengths of the electromagnetic spectrum, were also calculated from the spectral data. The Phytophthora treatments did not cause significant differences with the biophysical measurements in both trials. In the second trial, the waterlogging treatment significantly lowered plant top dry weight (P = 0.016) and diameter (P = 0.044). Reflectance values plotted against wavelength displayed differences between treatments as well as a seasonal trend. The NDSI heat maps indicated that the Phytophthora and waterlogging treatment effects were strongest correlated with bandwidths in the visible and near-infrared portions of the electromagnetic spectrum (538–558 nm and 701–709 nm). Six of the VIs (normalized difference nitrogen index 2, anthocyanin reflectance index 1, photochemical reflectance index, Carter index 1, Vogelman index 3 and water band index) were able to track the biochemical changes in the leaves over the 10 weeks, confirming the seasonal trend. The interaction effect between P. cinnamomi, waterlogging and elapsed time in the first trial was significant for water band index (P = 0.010). This study demonstrates that reflectance spectroscopy holds promise for characterizing marri response but more work needs to be done to identify the optimum wavelengths for identifying Phytophthora and waterlogging stress with marri.

Description: There is a strong policy move in Britain to improve forest resilience to climate change by increasing stand structural and species diversity. Although currently little used in Britain, the technique of underplanting allows regeneration and diversification of stands while avoiding some of the disadvantages of clearfelling. Two experiments were examined: (1) the growth and survival of five underplanted conifer species of differing shade tolerance in a shelterwood and (2) compared performance of underplanted and open-grown Douglas-fir seedlings on restocking sites. Underplanted Sitka spruce, Norway spruce, noble fir, European silver fir and Douglas-fir were all able to survive and grow. However increased exposure following overstorey removal resulted in some damage and ‘socketing’, especially to taller seedlings, particularly Douglas-fir. This may be linked to poor root development when growing under an overstorey. Microclimate conditions on some underplanted sites were more sheltered from extreme climatic conditions, and in some cases this improved survival of Douglas-fir seedlings. However, seedling growth rates were reduced compared with those on open sites probably due to lower light levels. Underplanting may help to improve establishment success of some species, particularly in exposed areas. However, the shelter benefits of underplanting must be carefully balanced against the trade-off with lower light, and underplanting is likely to be more successful where low canopy density is maintained.

Description: The recent emergence of Olive Quick Decline Syndrome in Italy, caused by Xylella fastidiosa, has drawn attention to the risks posed by this vector-borne bacterium to important crops in Europe (especially fruit trees and grapevine). Comparatively very little is known on actual and potential impacts of this pathogen in forests, in the native (North American) and introduced (European) regions, respectively. The present review aims to address important questions related to the threat posed by X. fastidiosa to European forests, such as the following: What are the symptoms, hosts and impact of bacterial leaf scorch caused by X. fastidiosa on trees in North America? Which forest tree species have been found infected in the introduction area in Europe? How does X. fastidiosa cause disease in susceptible hosts? Are there any X. fastidiosa genotypes (subspecies and sequence types) specifically associated with forest trees? How is X. fastidiosa transmitted? What are the known and potential vectors for forest trees? How does vector ecology affect disease? Is the distribution of X. fastidiosa, especially the strains associated with trees, restricted by climatic factors? Is disease risk for trees different in forest ecosystems as compared with urban settings? We conclude by pointing to important knowledge gaps related to all these questions and strongly advocate for more research about the Xylella-forest pathosystems, in both North America and Europe.

Description: Wood removed in Southwestern US forest restoration treatments currently has limited markets and thus low value. One important property of wood in structural products is its stiffness (measured as modulus of elasticity), which is known to vary systematically within trees. Directly measuring wood stiffness is expensive, time consuming and destructive. Therefore, we tested samples of ponderosa pine (Pinus ponderosa var. scopulorum Engelm.) from northern Arizona destructively in bending and also non-destructively using acoustic velocity (AV) methods. In total, we tested multiple pith-to-bark small clear (2.54 × 2.54 × 40.64 cm) samples from up to four heights in 103 trees. We first measured the standing-tree AV of sample trees, then the AV of small clear samples, and finally measured wood stiffness using three point static bending tests. We found that a Michaelis–Menten curve was a good fit to the radial profile of wood stiffness, with a steep increase outward from the pith that approached an asymptote. The AV of small clear samples, coupled with measured volumetric density values, approximated the static modulus of elasticity values with high accuracy (r2 = 0.86). At the stand level, a model predicting standing tree AV from tree morphology fit the data well (r2 = 0.77). Results indicate that southwestern ponderosa pine contains outerwood with relatively high stiffness that could be suitable for structural products. However, when assessed using wood stiffness, the trees take a long time to reach maturity (~50 years) and thus the corewood proportion is large. AV measurements are a good way to assess variability within and among stands and thus could be employed to segregate the resource by expected stiffness values. Segregation could help identify stands not suitable for structural uses and direct them toward more appropriate products.

Description: In UK forestry, the synthetic pyrethroid insecticides alpha-cypermethrin and cypermethrin have been used for many years to provide protection for young trees planted on restock sites from damage by the large pine weevil, Hylobius abietis L. However, concerns over the toxicity of these insecticides to aquatic life if misused have led to a search for alternative forms of protection. This paper describes a detailed programme of efficacy experiments undertaken between 2009 and 2015 to find replacements for these products. Over 50 combinations of chemical and non-chemical approaches were tested on 16 different sites. Of the alternative synthetic insecticides tested, applications of 0.037 g a.i. stem−1 acetamiprid provided high levels of protection from Hylobius browsing damage on young Sitka spruce (Picea sitchensis (Bong.) Carrière) trees, without causing any phytotoxic symptoms, and gave comparable levels of protection to those achievable using alpha-cypermethrin or cypermethrin. Acetamiprid is less toxic to aquatic life than alpha-cypermethrin or cypermethrin and has not been linked to bee decline. Applications of 0.0129 g a.i. stem−1 chlorantraniliprole also showed promise, and this relatively low toxicity non-neonicotinoid insecticide merits further study. Although imidacloprid and thiacloprid also provided good levels of protection, their use in forests is not now permitted due to concerns over their potential impacts on bees and drinking water, respectively. Whilst the natural product insecticide spinosad, and the entomopathogenic fungal control agent Metarhizium anisopliae (Metschn.) Sorokin, gave only limited protection in our work, they may have some future potential if methods of deployment can be improved. Other chemical and non-chemical approaches tested, but found to be largely ineffective in UK conditions, included the natural product insecticides azadirachtin, maltodextrin and pyrethrins, the synthetic insecticides lambda-cyhalothrin and spirotetramat and a wide range of repellents, flexible stem coatings and physical barrier products. However, we conclude that physical barrier sleeves such as MultiPro® and BioSleeve® may have a limited role as a partial substitute for the use of insecticides in the UK in some circumstances, but only if on-site populations of Hylobius are predicted to be low.

Description: To increase forest resilience to global change, forest managers are often directing forest stands towards a broader diversity of tree species. The small-leaved lime (Tilia cordata Mill.), a rare and scattered species in northwestern Europe, is a promising candidate for this purpose. Its life traits suggest a high resilience to climate change and a favourable impact on forest ecosystem services. This study used a dendroecological approach to assess how lime tree radial growth had responded to the past climatic change. First, 120 lime trees from nine sites were selected in southern Belgium based on criteria adapted to the rareness of the species. Chronology quality was assessed and resulting tree-ring series were validated at site and region levels. Second, a range of dendrochronological methods was used to analyze the changes over time in the variability and long-term trends of lime tree growth and their relation to climate during the period 1955–2016. Last, behaviour of lime trees was compared with that of beech from the same region and time period. For this purpose, the same methodology was applied to an additional beech tree-ring dataset (149 trees from 13 sites). Beech is the climax tree species of the region, but is known to be drought-sensitive and has shown weaknesses in the current climate. The quality of our tree-ring series attests that dendroecological investigation using rare and scattered species is possible, opening the way to further analysis on other such lesser-known forest tree species. The analysis showed that the small-leaved lime had been resilient to the past climatic change in multiple ways. Lime growth increased during the preceding decades despite an increased frequency and intensity of stressful climatic events. Lime growth quickly recovered in the years following the stresses. The growth–climate relationships were either stable over time or had a positive evolution. The behaviour of lime contrasted strongly with that of beech. Lime performed better than beech in every analysis. Small-leaved lime is thus a serious candidate for addressing climate change challenges in the region. It should be considered by forest managers planning to improve the sustainability and resilience of their forests, in particular in vulnerable beech stands.

Description: The restoration of forest structure and function is increasingly being used in boreal forests in order to halt the loss of biodiversity. Often ecological restoration is aimed at increasing the volume of dead and dying trees to enhance the biodiversity of deadwood-dependent organisms, but it may also increase population sizes of pest bark beetle species, even several years following restoration. Herein, we used a large-scale restoration experiment in Northern Sweden to assess the 5 years post-restoration effects of restorative gap cutting and prescribed burning on the populations of a set of economically harmful pest bark beetles (Ips typographus, Polygraphus poligraphus, Tomicus piniperda and Pityogenes chalcographus) and the most important predators of bark beetles, Thanasimus spp. In addition, we assessed the effects of forest stand characteristics at stand and landscape scale on the abundance of I. typographus. Five years post-restoration, gap-cut stands supported the highest abundances of P. poligraphus and contained the highest count of spruce trees newly attacked by bark beetles. By contrast, prescribed burning generally sustained the lowest abundances of pest bark beetles, especially I. typographus and P. poligraphus, and the highest abundance of their natural predators Thanasimus spp. The population abundance of I. typographus was also positively affected by the area of clear cuts within a 500 m radius from the stand. In conclusion, prescribed burning appears to be a safer method for ecological restoration than gap cutting in the long-term. According to our results, a risk of a local bark beetle outbreak still remains 5 years following the initiation of ecological restoration treatments on spruce-dominated mature gap-cut stands.

Description: ‘Analog’ forestry is a novel silvicultural approach for reforestation of degraded land that seeks to establish trees that are analogous in structure and ecological function to the original climax or subclimax vegetation of the region where they are planted. Analog forestry aims to strengthen rural communities, both socially and economically, with tree plantings of high biodiversity, which provide commercial products and resiliency, while avoiding agrochemicals and fossil fuels. The latter links this strategy to climate change mitigation and adaptation strategies, but there has been little analysis comparing the potential for above- and below-ground carbon storage in analog forests, as compared with more typical reforestation efforts via single-species plantations. We present the results of a study to estimate carbon sequestration levels, above- and below-ground, from an experimental analog forest system and a teak (Tectona grandis L.f.) plantation system, in comparison with a degraded pasture system. The study found that the analog and teak plantation forest systems stored higher quantities of total carbon (178 and 141 t C ha−1, respectively), when compared with a degraded pasture system (124 t C ha−1). However, the teak plantation had decreased soil carbon stocks, relative to the degraded pasture. The analog forest had the best balance of both above- and below-ground carbon stored over the 17-year study period and provided a more diverse array of timber and non-timber forest products when compared with the teak plantation. This suggests that analog forestry could be a viable social–ecological approach to carbon storage and reforestation in the study region and other places with large areas of degraded pasture and a good understanding of the structure and function of the original vegetation.

Description: Loblolly pine is a major fibre source for the pulp and paper industry. Here we developed the first nonlinear models to predict the within-tree and regional variation of tracheid length and width for planted loblolly pine. Data were obtained from macerated tracheids and near-infrared spectroscopy calibration models from trees sampled in 109 stands across the southeastern United States. The fixed effects for the final tracheid length model, which included cambial age, height of disk within tree, and physiographic region, explained 71 percent of the variation with root mean square error (RMSE) of 0.28 mm, while the fixed effects for the final tracheid width model explained 57 percent of the variation with RMSE of 1.4 μm. There was significant variation in tracheid properties across the growing regions. Tree maps showing within-tree variability in tracheid properties were produced. Five simulated scenarios were compared using the models developed, with mean tracheid dimensions calculated on a whole-tree basis at a first and second thinnings, and at final harvest. Also from the final harvest, the tops of trees, and outerwood chips produced during lumber manufacturing were also simulated. For the whole tree scenarios, both mean tracheid length and width increased with age, increasing from 2.24 mm and 40.5 μm (age 12), to 2.51 mm and 41.3 μm (age 18), and to 2.73 and 41.8 μm at age 25, respectively. The tops of the trees at age 25 had a mean tracheid length of 2.46 mm and a mean width of 41.0 μm, while the chips had a mean tracheid length of 3.13 mm and a mean width of 42.5 μm. Due to the models representing samples collected from across the southeastern United States, and their relatively high precision, they are suitable for incorporation into growth and yield systems allowing for prediction of tracheid properties.

Description: Light detection and ranging (LiDAR) is used to estimate tree, stand and forest characteristics across large geographic areas. In most analyses, several independent LiDAR-based allometric equations are built to predict various forest attributes. When each forest attribute is estimated independently, there is potential for predictions of forest attributes that are not mathematically or biologically consistent. Combined allometric equations can be considered a system of equations describing the stand structure. Mathematically compatible and biologically meaningful estimates can be derived by estimating key structural variables and solving for other components, rather than estimating each forest attribute separately and independently. In this study, we propose the development of a system of allometric equations describing the relationship between volume per unit area, Lorey’s average height, basal area, quadratic mean diameter (QMD) and density. The system of allometric equations is derived from extensive field data. Key structural attributes are predicted using LiDAR metrics, and the remaining structural variables are solved for using the system of allometric equations. Predictions of structural attributes from the system of allometric equations are compared with predictions from independent LiDAR-derived prediction equations. Results showed that applying the systems approach can provide reasonable and compatible estimates with lower required sample sizes, especially when multiple attributes need to be considered simultaneously. Testing the portability of the systems approach in more complex stand structures and across different LiDAR acquisitions will be required in the future.

Description: Forests of the world constitute one-third of the total land area and are critical for e.g. carbon balance, biodiversity, water supply and as source for bio-based products. Although the terrain within forest land has a great impact on accessibility, there is a lack of knowledge about the distribution of its variation in slope. The aim was to address that knowledge gap and create a globally consistent dataset of the distribution and area of forest land within different slope classes. A Geographic Information System (GIS) analysis was performed using the open-source QGIS, GDAL and R software. The core of the analysis was a digital elevation model and a forest cover mask, both with a final resolution of 90 m. The total forest area according to the forest mask was 4.15 billion hectares whereof 82 per cent was on slope  30°. Out of the major forestry countries, China had the largest proportion of forest steeper than 15° followed by Chile and India. A sensitivity analysis with 20 m resolution resulted in increased steep areas by 1 per cent point in flat Sweden and by 11 per cent points in steep Austria. In addition to country-specific and aggregated results of slope distribution and forest area, a global raster dataset is also made freely available to cover user-specific areas that are not necessarily demarcated by country borders. Apart from predicting the regional possibilities for different harvesting equipment, which was the original idea behind this study, the results can be used to relate geographical forest variables to slope. The results could also be used in strategic forest fire fighting and large-scale planning of forest conservation and management.

Description: Forest harvesting operations with heavy machinery can lead to significant soil rutting. Risks of rutting depend on the soil bearing capacity which has considerable spatial and temporal variability. Trafficability prediction is required in the selection of suitable operation sites for a given time window and conditions, and for on-site route optimization during the operation. Integrative tools are necessary to plan and carry out forest operations with minimal negative ecological and economic impacts. This study demonstrates a trafficability prediction framework that utilizes a spatial hydrological model and a wide range of spatial data. Trafficability was approached by producing a rut depth prediction map at a 16 × 16 m grid resolution, based on the outputs of a general linear mixed model developed using field data from Southern Finland, modelled daily soil moisture, spatial forest inventory and topography data, along with field measured rolling resistance and information on the mass transported through the grid cells. Dynamic rut depth prediction maps were produced by accounting for changing weather conditions through hydrological modelling. We also demonstrated a generalization of the rolling resistance coefficient, measured with harvester CAN-bus channel data. Future steps towards a nationwide prediction framework based on continuous data flow, process-based modelling and machine learning are discussed.

Description: In forest inventory, traditional ground-based resource assessments are often expensive and time-consuming forcing managers to reduce sample sizes to meet budgetary and logistical constraints. Small area estimation (SAE) is a class of statistical estimators that uses a combination of traditional survey data and linearly related auxiliary information to improve estimate precision. These techniques have been shown to improve the precision of stand-level inventory estimates in loblolly pine plantations using lidar height percentiles and thinning status as covariates. In this study, the effects of reduced lidar point-cloud densities and lower digital elevation model (DEM) spatial resolutions were investigated for total planted volume estimates using area-level SAE models. In the managed Piedmont pine plantation conditions evaluated, lower lidar point-cloud densities and DEM spatial resolutions were found to have minimal effects on estimates and precision. The results of this study are promising to those interested in incorporating SAE methods into forest inventory programs.

Description: We explore the implications of managing forests for the dual purpose of sequestering carbon and producing timber, using a model of the forest sector that includes a Hartman-based representation of forest owners’ behaviour as well as heterogeneity in environmental conditions. We focus on France, where recent policies aim at increasing the carbon sink and where the diversity of forests makes an analysis of spatial dynamics relevant, and we use recent estimates of the shadow price of carbon consistent with the country’s climate commitments. Results suggest that forests may sequester up to 550 MtCO2eq by 2100, driven by changes in harvest levels and species choice, whilst rotation lengths increase overall. A spatial analysis reveals a high spatial variability for these trends, highlighting the importance of considering the local context. Changes in investment patterns affect the spatial distribution of forest cover types: by the end of the century, a majority of regions comprise a larger share of older, multiple-species and mixed-structure forests. Whilst such an evolution may present benefits in terms of biodiversity, ecosystem services provision and resilience, it raises questions regarding the adequacy of such developments with current forest policy, which also aims at increasing harvest levels. An overall mitigation strategy for the forest sector would likely include incentives to energy and material substitution in downstream industries, which we did not consider and may interact with sequestration incentives.

Description: Invertebrate forest pests and pathogens can cause considerable economic losses and modern patterns of trade have facilitated the international movement of pest species on an unprecedented level. This upsurge in trade has increased the pathways available to high risk species, facilitating entry and potential establishment in nations where they were previously absent. To support policy and pest prioritization, pest risk analyses are conducted to decide ‘if’ and ‘how’ pests should be regulated in order to prevent entry or establishment; however, they cannot be carried out for every potential pest. This paper utilizes a hierarchical clustering (HC) approach to analyse distribution data for pests of Sitka spruce (Picea sitchensis (Bong.) Carr.) in order to identify species of high risk to Ireland, as well as potential source regions of these pests. The presence and absence of almost a 1000 pests across 386 regions globally are clustered based on their similarity of pest assemblages, to provide an objective examination of the highest risk pests to Irish forestry. Regional clusters were produced for each taxon analysed including the Coleoptera, Diptera, Hemiptera, Hymenoptera, Nematoda, Lepidoptera and the Fungi. The results produced by the HC analysis were interpreted with regard to biological realism and climate. Biologically meaningful clusters were produced for each of the groups, except for the Diptera and Nematoda, and each of the species analysed were ranked within their group by a quantitative risk index specific to the island of Ireland. The impact of uncertainty in the distribution data is also examined, in order to assess its influence over the final groupings produced. The outputs from this analysis suggest that the highest risk pests for Ireland’s Sitka spruce plantations will originate from within Europe. Ultimately, Ireland could benefit from seeking regulation for some of the higher ranking pests identified in this analysis. This analysis provides the first of its type for Sitka spruce, as well as its application in Ireland. It also serves to highlight the potential utility of HC as a ‘first approach’ to assessing the risk posed by alien species to hitherto novel regions.

Description: Regular biomass estimations for natural and plantation forests are important to support sustainable forestry and to calculate carbon-related statistics. The application of remote sensing data to estimate biomass of forests has been amply demonstrated but there is still space for increasing the efficiency of current approaches. Here, we investigated the influence of field plot and sample sizes on the accuracy of random forest models trained with information derived from Pléiades very high resolution (VHR) stereo images applied to plantation forests in an arid environment. We collected field data at 311 locations with three different plot area sizes (100, 300 and 500 m2). In two experiments, we demonstrate how plot and sample sizes influence the accuracy of biomass estimation models. In the first experiment, we compared model accuracies obtained with varying plot sizes but constant number of samples. In the second experiment, we fixed the total area to be sampled to account for the additional effort to collect large field plots. Our results for the first experiment show that model performance metrics Spearman’s r, RMSErel and RMSEnor improve from 0.61, 0.70 and 0.36 at a sample size of 24–0.79, 0.51 and 0.15 at a sample size of 192, respectively. In the second experiment, highest accuracies were obtained with a plot size of 100 m2 (most samples) with Spearman’s r = 0.77, RMSErel = 0.59 and RMSEnor = 0.15. Results from an analysis of variance type-II suggest that the overall most important factors to explain model performance metrics for our biomass models is sample size. Our results suggest no clear advantage for any plot size to reach accurate biomass estimates using VHR stereo imagery in plantations. This is an important finding, which partly contradicts the suggestions of earlier studies but requires validation for other forest types and remote sensing data types (e.g. LiDAR).

Description: Prosopis chilensis (Molina) Stuntz (Leguminosae) is a valuable native species in Argentina included in the Prosopis Management Programme. Natural provenances show important height and shape differentiation throughout their distribution in the Monte Desert. The availability of progeny trials provides an opportunity to quantify genetic differentiation among provenances and test the relative importance of demographic vs adaptive processes on morphological variation. We quantified both genetic and quantitative differentiation of neutral markers and five economically important traits, respectively, among four provenances in a provenance-progeny trial. We aimed to quantify the genetic basis of variations in height, basal diameter, tree shape, spine length and biomass. Two hypotheses were tested: (1) economically important traits have significant heritability, and (2) the phenotypic variation among provenances is the result of local adaptation to particular environmental conditions. Our results indicate that most morphological variation was found among individuals within families (~95 per cent). The h2 estimates were heterogeneous among traits and ranged from low (0 for number of stems) to moderate (0.22 and 0.28 for spine length and biomass, respectively). Variance among families (~5 per cent) was evenly distributed within and among provenances. Morphological differentiation among provenances was low, but significant, and could be attributed mainly to individuals from Villa Unión. Based on molecular markers, genetic differentiation among provenances was low and significant (FST = 0.03; P = 5 × 10−4) but was able to differentiate the groups from Villa Unión, Fiambalá and Mogna-Chilecito. Neutrality tests were conducted using the FST–QST test and DJSOST and δGREGORIUS alternative coefficients of differentiation. Neutrality tests yielded no evidence of local adaptation and were rather consistent in showing a trend toward stabilizing selection, particularly for spine length. The selection strategy for breeding programmes should depend on the trait to be improved and should consider both provenance and familiar information. Considering an intra-familiar ranking is encouraged in order to maximize the genetic gain. Additionally, in order to recover germplasm provenance diversity, based on morphological and microsatellite results, our recommendation would be to include seeds from individuals from at least the Villa Unión, Fiambalá and Mogna-Chilecito areas.

Description: To investigate enemy pressure across the range of the pine processionary moth (Thaumetopoea pityocampa, Den. & Schiff.) as it spreads northwards in Europe because of climate change, a survey of 206 egg batches from historical and newly colonized areas at both the westernmost and the easternmost ends of the front was carried out in 2016–2018. Out of them, 97 egg batches were collected from nine locations in three regions in France and 109 from four localities in two regions in Bulgaria, both within historical and newly colonized ranges of the pest. The average number of eggs per batch collected in Bulgaria (226.6 ± 43.2) was higher than that in France (194.3 ± 50.1). However, the hatching percentage was higher in French samples, varying from 69.8 to 95.7, vs 49.8 to 85.2 per cent in Bulgarian samples. Four primary parasitoids (Ooencyrtus pityocampae, Baryscapus servadeii, Anastatus bifasciatus, Trichogramma sp.) and a hyperparasitoid (Baryscapus transversalis, found only in two regions of the historical range in Bulgaria) were identified. The oligophagous species B. servadeii was present at all sites within the historical range of the pest. In newly colonized areas, the impact of primary parasitoids on the host was distinctly low, suggesting that they lag behind the range expansion of pine processionary moth. In France, the most abundant species in the T. pityocampa parasitoid complex were B. servadeii in Ré Island and Orléans (97.3 and 87.4 per cent, respectively), and Trichogramma sp. (99.7 per cent) in a newly colonized locality in Fréhel. Ooencyrtus pityocampae prevailed in three of four Bulgarian localities (72.1 per cent in Sandanski, 89.7 per cent in Maglizh and 65.7 per cent in Sladak kladenets), whereas B. servadeii was the most abundant in Gega (75.4 per cent). Mortality of B. servadeii and O. pityocampae caused by the hyperparasitoid B. transversalis amounted to 4.8–6.2 per cent. The impact of predators on the pine processionary moth in the egg stage was negligible at most sites, reaching 12.5 per cent in only one site (Maglizh).

Description: Widely distributed in Quebec, balsam fir (Abies balsamea (L.) Mill.) is highly vulnerable to wind damage. Recently, there has been a trend in forest management to increase the use of partial cuttings in naturally regenerating stands, leaving the remnant trees at increased risk of wind damage. In order to limit wind damage after partial cuttings, it is therefore important to find silvicultural practices that minimize the risk of wind damage in these fir stands. Our main objective was to find balsam fir-specific values of parameters to integrate into the wind risk model ForestGALES, in order to simulate the impact of different types of commercial thinning on wind damage risk, and to determine which practice potentially minimizes the risk in a naturally regenerated stand. An anemometer placed at canopy height and strain gauges attached to the trunks of balsam firs allowed us to measure the wind-induced bending moments experienced by a sample of balsam fir trees. This enabled the calculation of the turning moment coefficients specific to each of the trees in order to compare them with the ForestGALES model predictions and to adapt the model for balsam fir stands. The model was tested first with only tree diameter and height as input variables to calculate the turning moment coefficient, then with the addition of a competition index, and finally with the addition of crown dimensions. Wind climate parameters for prediction of the probability of damage were calculated using the Wind Atlas Analysis and Application Program airflow model. The model with the highest accuracy was then used to simulate two types of thinning and determine the impact on wind damage risk for each tree in the stand. According to the model’s predictions, thinning from below has a reduced risk of wind damage compared with thinning from above.

Description: An understanding of the relationship between volume increment and stand density (basal area, stand density index, etc.) is of utmost importance for properly managing stand density to achieve specific management objectives. There are two main approaches to analyse growth–density relationships. The first relates volume increment to stand density through a basic relationship, which can vary with site productivity, age, and potentially incorporates treatment effects. The second is to relate the volume increment and density of thinned experimental plots relative to that of an unthinned experimental plot on the same site. Using a dataset of 229 thinned and unthinned experimental plots of Norway spruce, a growth model is developed describing the relationship between gross or net volume increment and basal area. The models indicate that gross volume increases with increasing basal area up to 50 m2 and thereafter becomes constant out to the maximum basal area. Alternatively, net volume increment was maximized at a basal area of 43 m2 and decreased with further increases in basal area. However, the models indicated a wide range where net volume increment was essentially constant, varying by less than 1 m3 ha−1 year−1. An analysis of different thinning scenarios indicated that the relative relationship between volume increment and stand density was dynamic and changed over the course of a rotation.

Description: Substantial growth losses in Douglas-fir can be the result of Swiss needle cast, a foliar disease caused by the fungus Nothophaeocryptopus gaeumannii. Although the disease is native to western North America, it is expected to become increasingly problematic in regions where climates become warmer in winter and wetter in spring, such as in coastal British Columbia (BC), Canada. Previous research suggests that tolerance to this disease is under partial genetic control. We therefore aim to screen for tolerance and resistance to the disease in the tree breeding population of coastal Douglas-fir (Pseudotsuga menziesii var. menziesii) in BC. We evaluated if early screening for resistance or tolerance to this disease is possible. We worked with 32 families grown for 18-years on two full-sibling genetic field trial sites representing different climates. We assessed 〉900 trees for disease signs and symptoms from 2017 to 2019. Needle retention was assessed in the field. In the laboratory, the proportion of stomata occluded with pseudothecia was measured, deoxyribonucleic acid (DNA) was extracted, and the proportion of fungal DNA in the needles was assessed using quantitative polymerase chain reaction. Strong differences were observed among families for needle retention, stomatal occlusion and fungal load. The milder and wetter site closer to the coast was generally more severely affected. Families showed rank changes between the two sites for all response variables. Higher needle retention was correlated with increased tree volume. No correlation was found between the proportion of stomata occluded with pseudothecia, fungal DNA load and needle retention. These results indicate that a more complex pathology is involved in causing needle loss. We conclude that screening for Swiss needle cast tolerance in the coastal BC Douglas-fir breeding population is possible if needle retention can be assessed and that area-specific deployment strategies may be needed given family rank changes in different environments.

Description: Higher temperatures and drought are key aspects of global change with the potential to alter the distribution and severity of many arthropod pests in forest systems. Scale insects (Hemiptera: Coccoidea) infest many tree species and are among the most important pests of trees in urban and rural forests, plantations and other forest systems. Infestations of native or exotic scale insects can kill or sicken trees with economic and ecosystem-wide consequences. Warming can have direct effects on the life history, fitness and population dynamics of many scale insect species by increasing development rate, survival or fecundity. These direct benefits can increase the geographic distribution of scale insects and their consequences for tree health. Warming and drought can affect scale insects indirectly by altering the quality of their host trees. Additive or interactive effects of warming and drought can change tree quality in such a way that it increases scale insect fitness and population growth. However, the effects are species- and context-dependent with some scale insect species negatively affected by drought-induced changes in tree quality. Warming and drought are often coincident in urban forests and predicted to co-occur in many parts of the world under climate change scenarios. The individual and interactive effects of these factors require further research to inform predictions and management of scale insect pests. Warming also indirectly affects scale insects by altering interactions with natural enemies. This includes changes in natural enemy phenology, community composition and abundance. In addition, warming can alter scale insect phenology or voltinism causing asynchrony with natural enemies or population growth too rapid for natural enemies to suppress. Direct and indirect effects of warming and drought on scale insects can increase the potential for some exotic species to become established and for some native species to become invasive. Unfortunately, much research on scale insects is confined to a few particularly important native or exotic pests which limits our ability to predict the effects of warming on many current or potential pests. More research is required to understand how warming and drought affect scale insects, scale insect management and the forest systems they inhabit.

Description: Smartphones with their capability to receive Global Navigation Satellite Systems (GNSS) signals can be currently considered the most common devices used for positioning tasks, including forestry applications. This study focuses on possible improvements related to two crucial changes implemented into Android smartphone positioning in the last 3 years – dual-frequency (L1/L5) GNSS receivers and the possibility of recording raw GNSS data. The study comprises three experiments: (1) real-time measurements of individual points, (2) real-time recording of trajectories, and (3) post-processing of raw GNSS data provided by the smartphone receiver. The real-time tests were conducted using final positions provided by the internal receiver, i.e. without further processing or averaging. The test on individual points has proven that the Xiaomi Mi8 smartphone with a multi-constellation, dual-frequency receiver was the only device whose accuracy was not significantly different from single-frequency mapping-grade receiver under any conditions. The horizontal accuracy of most devices was lower during leaf-on season (root mean square errors between 5.41 and 12.55 m) than during leaf-off season (4.10–11.44 m), and the accuracy was significantly better under open-area conditions (1.72–4.51 m) for all tested devices when compared with forest conditions. Results of the second experiment with track recording suggest that smartphone receivers are better suited for dynamic applications – the mean shift between reference and measured trajectories varied from 1.23 to 5.98 m under leaf-on conditions. Post-processing of the raw GNSS data in the third experiment brought very variable results. We achieved centimetre-level accuracy under open-area conditions; however, in forest, the accuracies varied from meters to tens of meters. Observed loss of the signal strength in the forest represented ~20 per cent of the open-area value. Overall, the multi-constellation, dual-frequency receiver provided more robust and accurate positional solutions compared with single-frequency smartphones. Applicability of the raw GNSS data must be further studied especially in forests, as the provided data are highly susceptible to multipath and other GNSS adverse effects.

Description: National forest inventories, in which trees are often mapped within the plots, provide a tool for the quantification of large-scale forest structure since they cover all forest areas. Many National Forest Inventories follow a nested design in order to reduce the sampling effort for smaller trees. We propose and test a methodology that allows the spatial pattern of trees, species mingling and size differentiation to be characterized using the nearest neighbour indices and second-order moment functions from nested plot data. The nearest neighbour indices and second-order moment functions for the actual distribution are compared with simulations of the appropriate null model: spatial randomness for spatial pattern characterization or spatial independence for species mingling and size differentiation. The proposed method consists of constraining the null model to fit the nested plot design. For the purposes of the study, we simulated 120 plots and used 26 real plots located in pure and mixed stands in Central Spain, for which a complete census with detailed information about trees was available. The nested design used in the Spanish National Forest Inventory (SNFI) plots was simulated to test the performance, taking the complete census as reference. Despite of the limited accuracy for some structural measures, the proposed method based on nested design data performed better for most of the nearest neighbour indices and second-order moment functions than the strategy currently used in the SNFI for structure assessment in a subsample of SNFI plots, consisting of mapping the 20 trees closest to the plot centre. Nearest neighbour indices provided greater accuracy for species mingling assessment than second-order moment functions, whereas the opposite occurred when describing spatial pattern and size differentiation. The methodology proposed provides the first insight into the characterization of forest structure in nested designs although more evaluations are required for different forest types.

Description: Oak-dominated forests, economically and ecologically valuable ecosystems, are widely distributed in China. These oak-dominated forests are now generally degraded coppice forests, and are of relatively low quality. Climate change has been shown to affect forest growth, tree mortality, and recruitment, but available forest growth models are lacking to study climate effects. In this study, a climate-sensitive, transition-matrix growth model (CM) was developed for uneven-aged, mixed-species oak forests using data collected from 253 sample plots from the 8th (2010) and 9th (2015) Chinese National Forest Inventory in Shanxi Province, China. To investigate robustness of the model, we also produced a variable transition model that did not consider climate change (NCM), and fixed parameter transition matrix model (FM), using the same data. Short-term and long-term predictive performance of CM, NCM, and FM were compared. Results indicated that for short-term prediction (5 years), there was almost no significant difference among the three predictive models, though CM exhibited slightly better performance. In contrast, for long-term prediction (100 years), CM, under the three representative concentration pathways (RCPs), i.e. RCP2.6, RCP4.5 and RCP8.5, indicated rather different dynamics that were more reliable because climate factors were considered which could significantly influence forest dynamics, especially in long-term prediction intervals. The CM model provides a framework for the management of mixed-species oak forests in the context of climate change.

Description: Tree crown volume is a fundamental tree characteristic. It correlates to forest biomass production and most relevant ecosystem and environmental functions, such as carbon sequestration and air pollution reduction. When researching these relationships, it is necessary to clearly define and then quantify tree crown variables in a both accurate and operational manner. In this paper, we review the reported literature on the assessment of tree crown volume. First, we compile the varying definitions of crown volume and other tree crown variables that may be used as inputs to quantify crown volume. Then, we examine the data sources for quantifying these variables, including field measurements, terrestrial photographs, aerial photographs and laser scanning. Furthermore, we compare the published approaches on translating these crown variable measurements into tree crown volume. These approaches include the approximation of simple geometric solids, approaches of computational geometry and voxelization. We also compare the reported accuracies and major challenges of these approaches. From this literature review, the reader may craft a suitable approach for the assessment of crown volume.

Description: In this study, for five sites around the world, we look at the effects of different model types and variable selection approaches on forest yield modelling performances in an area-based approach (ABA). We compared ordinary least squares regression (OLS), k-nearest neighbours (kNN) and random forest (RF). Our objective was to test if there are systematic differences in accuracy between OLS, kNN and RF in ABA predictions of growing stock volume. The analyses are based on a 5-fold cross-validation at five study sites: an eucalyptus plantation, a temperate forest and three different boreal forests. Two completely independent validation datasets were also available for two of the boreal sites. For the kNN, we evaluated multiple measures of distance including Euclidean, Mahalanobis, most similar neighbour (MSN) and an RF-based distance metric. The variable selection approaches we examined included a heuristic approach (for OLS, kNN and RF), exhaustive search among all combinations (OLS only) and all variables together (RF only). Performances varied by model type and variable selection approaches among sites. OLS and RF had similar accuracies and were more efficient than any of the kNN variants. Variable selection did not affect RF performance. Heuristic and exhaustive variable selection performed similarly for OLS. kNN fared the poorest amongst model types, and kNN with RF distance was prone to overfitting when compared with a validation dataset. Additional caution is therefore required when building kNN models for volume prediction though ABA, being preferable instead to opt for models based on OLS with some variable selection, or RF with all variables together.

Description: In this paper, we present new dynamic top height models for 10 major species in Great Britain. These models for pure, even-aged stands replace the current, static top height-age curves published in the 1980s, which are no longer reliable in the light of the new data that have been added over the past decades. We followed the generalized algebraic difference approach (GADA) to build the new models and specifically focused on models derived from the Hessfeld IV and Chapman–Richards height-age base equations. We selected a multi-asymptote and polymorphic GADA version of the Chapman–Richards equation for all the species, with the anamorphic functional form as a special case derived from the non-significance of one of the global parameters. The new models are dynamic, accurate and robust, which will permit unbiased yield predictions from current top height-dependent yield simulators in Great Britain.

Description: Understanding the impact of management on the dispersal potential of forest tree species is pivotal in the context of global change, given the implications of gene flow on species evolution. We aimed to determine the effect of logging on gene flow distances in two Nothofagus species from temperate Patagonian forests having high ecological relevance and wood quality. Therefore, a total of 778 individuals (mature trees and saplings) of Nothofagus alpina and N. obliqua, from a single plot managed 20 years ago (2.85 hectares), were mapped and genotyped at polymorphic nuclear microsatellite loci. Historical estimates of gene dispersal distance (based on fine-scale spatial genetic structure) and contemporary estimates of seed and pollen dispersal (based on spatially explicit mating models) were obtained. The results indicated restricted gene flow (gene distance ≤ 45 m, both pollen and seed), no selfing and significant seed and pollen immigration from trees located outside the studied plot but in the close surrounding area. The size of trees (diameter at breast height and height) was significantly associated with female and/or male fertility. The significant fine-scale spatial genetic structure was consistent with the restricted seed and pollen dispersal. Moreover, both estimates of gene dispersal (historical and contemporary) gave congruent results. This suggests that the recent history of logging within the study area has not significantly influenced on patterns of gene flow, which can be explained by the silviculture applied to the stand. The residual tree density maintained species composition, and the homogeneous spatial distribution of trees allowed the maintenance of gene dispersal. The short dispersal distance estimated for these two species has several implications both for understanding the evolution of the species and for defining management, conservation and restoration actions. Future replication of this study in other Nothofagus Patagonian forests would be helpful to validate our conclusions.

Description: Europe has a history rich in examples of successful and problematic introductions of trees with a native origin outside of Europe (non-native trees, NNT). Many international legal frameworks such as treaties and conventions and also the European Union have responded to the global concern about potential negative impacts of NNT that may become invasive in natural ecosystems. It is, however, national and regional legislation in particular that affects current and future management decisions in the forest sector and shapes the landscapes of Europe. We identified all relevant legal instruments regulating NNT, the different legal approaches and the regulatory intensity in 40 European countries (no microstates). Information on hard and effective soft law instruments were collected by means of a targeted questionnaire and consultation of international and national legislation information systems and databases. In total, 335 relevant legal instruments were in place in June/July 2019 to regulate the use of NNT in the investigated 116 geopolitical legal units (countries as well as sub-national regions with their own legislation). Countries and regions were empirically categorized according to ad hoc-defined legislation indicators. These indicators pay respect to the general bans on the introduction of non-native species, the generally allowed and prohibited NNT, approval mechanisms and specific areas or cases where NNT are restricted or prohibited. Our study revealed a very diverse landscape of legal frameworks across Europe, with a large variety of approaches to regulating NNT being pursued and the intensity of restriction ranging from very few restrictions on species choice and plantation surface area to the complete banning of NNT from forests. The main conclusion is that there is a clear need for more co-ordinated, science-based policies both at the local and international levels to enhance the advantages of NNT and mitigate potential negative effects.

Description: The mixed-conifer forests in California’s Sierra Nevada include species from several genera (Pinus, Abies, Pseudotsuga, Calocedrus and Sequoiadendron). These forests have complex disturbance regimes dominated by low to moderate severity fire that often resulted in patchy spatial patterns and multiaged stands. Leaf area index (LAI) describes the total leaf surface area per unit area in a forest community and is related to wood and biomass production and ecosystem values such as water usage, water yields and carbon sequestration. LAI can also serve as a representation of growing space occupancy and the basis for stocking control, including in multiaged stands. Nine study sites were sampled with 22–37 0.05 ha plots per study site to estimate LAI and other metrics. LAI was highest in study sites with greater proportions of shade tolerant Abies and Calocedrus species and on higher productivity sites. Recent drought-related mortality has reduced stocking and LAI. The combination of fire suppression and timber harvest over the past century has resulted in stands with higher densities, and greater proportions of shade tolerant species. Managing these structures to restore their presettlement character will involve reducing overall stocking, increasing proportions of intolerant species and increasing fine-scale heterogeneity. LAI allocation—allocating leaf area to age classes, species or canopy strata—can be used to design new structures that resemble presettlement structures and are resilient to disturbances.

Description: Forests are dynamic ecosystems, subject to both natural and anthropogenic agents of change. Wildfire, harvesting and other human activities alter the tree-covered area present in forests. From national and international reporting perspectives, forests include areas currently treed, as well as those disturbed forest areas that are not currently treed but will be, given time for regeneration and the advancement of natural successional processes. As a consequence, forest area can be depicted at a particular point in time, informed by a retrospective temporal context. Using time series of Landsat imagery, annual land cover maps can be generated that are informed by knowledge of past disturbance history (such as wildfire and harvesting). In this research, we use over three decades of annual land cover data generated from Landsat time series to generate a spatially explicit estimate of the forest area of Canada in 2010. We demonstrate how land cover and disturbance information can be combined to map the area of ‘forest’, as defined by the Food and Agricultural Organization of the United Nations (FAO), within Canada’s 650 Mha of forested ecozones. Following this approach, we estimated Canada’s total forest area in 2010 to be 354.5 Mha. This estimate includes 324.5 Mha of current forest cover in 2010, plus an additional 33.2 Mha (or 9.4 per cent) of temporally informed forest area where tree cover had been temporarily lost due to fire or harvest, less 3.3 Mha that were removed to meet a definitional minimum size (0.5 ha) for contiguous forest area. Using Canada’s National Forest Inventory (NFI) as an independent reference source, the spatial agreement between the two estimates of forest area was ~84 per cent overall. Aspatially, the total area of the Landsat-derived estimate of 2010 forest area and the NFI baseline estimates differed by only 3 per cent, with notable regional differences in the wetland-dominated Hudson Plains Ecozone. Satellite-derived time series land cover and change information enable spatially explicit depictions of forest area (distinct from representations of forest cover) in a robust and transparent fashion, producing information of value to science, management and reporting information needs.

Description: Since the 1990s, forest resource maps and forest variable estimates for small areas have been produced by combining national forest inventory (NFI) field plot data, optical satellite images and numerical map data. A non-parametric $k$-NN method has frequently been employed. In Finland, such multi-source NFI (MS-NFI) forest variable estimates for municipalities have been produced eight times. A relatively large variation has been observed between subsequent estimates. In this study, a large-scale evaluation of small-area estimates from an MS-NFI conducted in 2013 was carried out in comparison with pure NFI field data-based estimates and error estimates. The proportion of municipalities with significant differences was larger than expected, e.g. over 10% for the mean volume, which indicates systematic error in the small-area estimates. A multi-temporal data fusion combining MS-NFI estimators from three time points—2011, 2013 and 2015—was tested as a means to improve single time point MS-NFI estimates of the mean volumes of growing stock and of tree species groups. A generalized least squares (GLS) technique and unweighted averaging were tested. The improvement was small but consistent when validated against the NFI field data-based estimates for the municipalities. The unweighted averaging worked nearly as well as a GLS estimator.

Description: We fertilized a Norway spruce (Picea abies (L.) Karst.) stand on rich mineral soil with 3 t ha−1 of wood ash (ASH), 150 kg ha−1 of nitrogen (N) or a combination of wood ash and nitrogen (ASH + N), in addition to unfertilized control plots. After five growing seasons, we remeasured the trees and took core samples. Current- and previous-year needles were sampled and analyzed for total nitrogen and carbon, low-molecular weight phenolics and condensed tannins. Annual volume increment and standing volume were significantly higher in the ASH + N treatment than in control plots after 5 years. N gave a significant positive effect on basal area growth in the third year, after which the effect diminished. The ASH + N treated trees, on the other hand, showed an increasing basal area growth trend throughout the period. ASH reduced the total concentration of low-molecular weight phenolic compounds significantly in current-year needles. Phenolic acids increased under both ASH and ASH + N, while flavonoids decreased significantly under the same treatments compared to N. By including annual growth rate before fertilization in the analyses, the effect of N-treatment on flavonoids was positive only in trees with higher growth rates, and in those trees the concentration was higher than in both ASH-treated plots and controls. An acetophenone, constituting more than half of the total low-molecular weight phenolics concentration, was strongly reduced under all fertilization treatments. These results demonstrate that in addition to effects on tree growth, fertilization of the forest floor also has a strong influence on other metabolic processes of trees, with potential implications for ecosystem functioning.

Description: The visual assessment of tree vigour before partial cutting is a key element of the long-term sustainability of managed hardwood forests. Several classification systems have been developed and applied to perform this task during the tree marking process. They segregate trees in different vigour classes based on the presence and severity of stem and crown defects. Yet, the relevance of using these defects to assess tree vigour has rarely been empirically validated. In this study, we analysed the relationships between quantitative vigour indicators and a full range of individual defects using 336 sugar maple and 84 yellow birch trees. Among the tested defects, the tree crown density showed the best ability to identify slow growing, non-vigorous sugar maple trees, regardless of their size. However, none of the stem-related defects, such as bark aspect, the presence of cambial necrosis or conks and stromata were strongly related to our quantitative vigour indicators. For yellow birch, none of the stem defects and crown conditions were found to be related to our vigour indicators. We conclude that, pending longitudinal studies that will provide a full empirical validation of classification systems, crown density should be used to assess recent growth and vigour of sugar maple trees and guide tree marking.

Description: Transforming even-aged coniferous stands into multi-aged ones is attracting growing interest in Europe. However, applying this silvicultural treatment, maintaining a continuous cover and relying on natural regeneration require a deep understanding of the factors driving interspecific competition in the understorey. In particular, knowledge of species-specific response to different light conditions is needed to plan silvicultural treatments and forecast long-term stand composition. In this context, we assessed regeneration (±10–400 cm in height) and light conditions (±1–40 per cent of transmittance) in nine coniferous stands with ranging stand age (±20–120 years) and species composition (Norway spruce, Douglas fir, larch, silver fir and western hemlock) in Belgium. We then modelled interspecific differences in regeneration height growth to forecast the outcome of interspecific competition in different light conditions. Controlling understorey light seems an efficient way to control the interspecific competition, but with some limits, and taking into account sapling size. Maintaining low light conditions (transmittance 〈 15 per cent) probably reduces interspecific competition as it allows small saplings (height 

Description: Most of the current understanding of the orientation and communication of jewel beetles arose from research on the Asian emerald ash borer (EAB), Agrilus planipennis, which has become one of the most destructive invasive forest insect pests in history following its introduction to North America and European Russia. From a European perspective, a number of jewel beetles have a high invasive risk similar to that of the emerald ash borer, including the potential threat of the bronze birch borer Agrilus anxius, the goldspotted oak borer Agrilus auroguttatus, and the twolined chestnut borer Agrilus bilineatus. Native jewel beetles expanding their geographic range include the cypress jewel beetle Ovalisia festiva and the black-banded oak borer Coraebus florentinus. Other native species are increasing in their importance, including the flathead oak borer Coraebus undatus, the two-spotted oak borer Agrilus biguttatus, the flatheaded beech borer Agrilus viridis and Agrilus cuprescens. Commonly used prism and multi-funnel trap designs and other promising experimental trap designs have been tested and compared in the US and in Europe. One factor considered has been colouration, typically purple and green. Another is olfactory attraction, both to plant volatiles and extracts such as (Z)-3-hexenol, Manuka oil, Phoebe oil and Cubeb oil, and also to pheromones such as (Z)-3-lactone, for emerald ash borer. Field observations have been made of mating and host-finding behaviours of oak buprestids based upon visual stimuli in North America and Europe. By using pinned dead EAB models, visual mating approaches have been observed by males of Agrilus biguttatus, Agrilus sulcicollis and Agrilus angustulus, which is a behaviour similar to that previously observed in EAB. Green plastic-covered branch-traps significantly out-performed other trap designs and caught more Agrilus jewel beetles if an artificial visual decoy that copies a beetle body was included. A higher fidelity decoy offered the same distinctive light-scattering pattern as real resting EAB females and elicited the full sequence of stereotypical male mating flight behaviour of EAB and A. biguttatus from up to 1 m away. An optimization of visual, olfactory and other possible stimuli has likely not yet been achieved. More sophisticated trap designs could lead to more sensitive detection capabilities with increased selectivity.

Description: Six experiments were established between 1955 and 1962 in different parts of northern and western Britain which used replicated randomized block designs to compare the performance of two species 50:50 mixtures with pure stands of the component species. The species involved were variously lodgepole pine (Pinus contorta Dougl.), Japanese larch (Larix kaempferi Lamb. Carr.), Scots pine (Pinus sylvestris L.), silver birch (Betula pendula Roth.), Sitka spruce (Picea sitchensis Bong. Carr.) and western hemlock (Tsuga heterophylla Raf. Sarg.). The first four species are light demanding, while Sitka spruce is of intermediate shade tolerance and western hemlock is very shade tolerant: only Scots pine and silver birch are native to Great Britain. In three experiments (Bickley, Ceannacroc, Hambleton), the mixtures were of two light-demanding species, while at the other three sites, the mixture tested contained species of different shade tolerance. The experiments were followed for around 50 years, similar to a full rotation of even-aged conifer stands in Britain. Five experiments showed a tendency for one species to dominate in mixture, possibly reflecting differences in the shade tolerance or other functional traits of the component species. In the three experiments, the basal area of the mixtures at the last assessment was significantly higher than predicted based on the performance of the pure stands (i.e. the mixture ‘overyielded’). In two of these cases, the mixture had had a higher basal area than found in the more productive pure stand indicating ‘transgressive overyielding’. Significant basal area differences were generally more evident at the later assessment date. The exception was in a Scots pine: western hemlock mixture where greater overyielding at the earlier date indicated a nursing (‘facilitation’) effect. In the remaining experiments, the performance of the mixture conformed to predictions from the growth of the component species in pure stands. Taken overall, the results suggest that functional traits can be used to interpret the performance of mixtures but prediction of the outcome will require better understanding of the interplay between species and site characteristics plus the influence of silvicultural interventions.

Description: The European Food Safety Authority (EFSA), by request of the European Commission, develops pest survey cards for pests of relevance for the European Union (EU) member states, summarizing key biological, epidemiological and diagnostic information relevant for the detection and identification of these pests by inspectors and laboratory technicians in the EU member states. For three pilot pests, including emerald ash borer (Agrilus planipennis), detailed guidelines are being prepared for the survey planners in the EU member states. Interaction with experts on the relevant organisms and the member states is needed before and after implementation of the surveys to ensure they are fit for purpose and can be harmonized across the EU. An important feature of the survey cards is the identification of risk factors, to focus the surveys on the most likely areas to find the pest if it is present and thus being able to apply a risk-based surveillance. Since 2014, ash wood and bark (from countries where A. planipennis is known to occur) are subjected to specific requirements laid down in Council Directive 2000/29/EC, the beetle is unlikely to enter the EU via this pathway. However, it cannot fully be excluded that introductions have happened before these requirements came into force, without being detected until now. In addition, the beetle could already be present in new third countries without being noticed yet and thus not regulated. Furthermore, firewood from countries adjacent to Russia (Belarus, Ukraine) is not restricted. The beetle could also hitch-hike to the EU by various means of transport, in particular via highways and railroads. Given the above, surveys should focus on these areas.

Description: Process-based models are increasingly applied for simulating long-term forest developments in order to capture climate change impacts and to investigate suitable management responses. Regarding dimensional development, however, allometric relations such as the height/diameter ratio, branch and coarse root fractions or the dependency of crown dimension on stem diameter often do not account for environmental influences. While this may be appropriate for even-aged, monospecific forests, serious biases can be expected if stand density or forest structure changes rapidly. Such events occur in particular when forests experience disturbances such as intensive thinning or during early development stages of planted or naturally regenerated trees. We therefore suggest a calculation of allometric relationships that depends primarily on neighbourhood competition. Respective equations have been implemented into a physiology-based ecosystem model that considers asymmetric competition by explicit simulation of resource acquisition and depletion per canopy layer. The new implementation has been tested at two sites in Germany where beech (Fagus sylvatica) saplings have either been planted below a shelterwood of old spruces (Picea abies) or grown under clear-cut conditions. We show that the modified model is able to realistically describe tree development in response to stand density changes and is able to represent regeneration growth beneath a gradually decreasing overstorey of mature trees. In particular, the model could represent the faster crown size development in saplings until full ground coverage is established and a faster height growth afterwards. The effect enhances leaf area and thus assimilation per tree and increases carbon availability for stem growth at early development stages. Finally, the necessity to consider dynamic allometric relations with respect to climate change impacts is discussed, and further improvements are suggested.

Description: The threats posed by the buprestid beetles emerald ash borer (Agrilus planipennis Fairmaire) and bronze birch borer (Agrilus anxius Gory) have been the subject of considerable research, primarily to develop methods for detection and management of the pests. PREPSYS, a Euphresco project, has worked with collaborators globally to assess the ‘state of the art’ for the two insect pests and to identify those measures that would best prepare Europe for potential invasion by the pests, especially emerald ash borer which is now in the western part of Russia and in eastern Ukraine. Building on an excellent exchange of knowledge and discussion at the OECD-sponsored international conference held in Vienna in October 2018, the concept of a European Toolbox to increase preparedness for dealing with the pests has been developed. This includes key components including surveillance, direct intervention, use of natural enemies and increased awareness of the problems associated with the pests. Collaboration is essential in delivering and refining the European Toolbox.

Description: We evaluated the impacts of juvenile stand management on wood production based on a nationwide scenario analysis. Measured plot-level information of the Finnish 11th National Forest Inventory (NFI) from juvenile stands formed the initial data for the analysis. To represent a total of 2.5 million hectares of productive forest land available for wood production, 6900 sample plots were selected. We predicted the development of the sample-plot stands for the next 100 years using the Motti stand simulator. We examined the impacts of juvenile stand management, including early cleaning and precommercial thinning, with three scenarios of different intensities of early-stage management (intensive, no management and business-as-usual). The results showed the benefits of juvenile stand management: earlier harvesting revenues, larger total removals, more valuable wood in the removals, and additional benefits that are associated with the first commercial thinning (i.e. no need for clearing before thinning, a larger stem size of removed trees). The main downside of juvenile stand management was the cost of the early cleaning and precommercial thinning. According to the results, a €560 million increase in costs would result in a €1700 million increase in stumpage earnings in Finnish forests during the next 100-year period. The 34 million m3 increase in the sawlog yield together with 28 million m3 decrease in the pulpwood yield also demonstrated the nationwide potential for increasing valuable sawlog production by intensive management of juvenile stands.

Description: Felled trees with tops and branches are transported to the landing with a grapple skidder in conventional ground-based whole-tree (WT) harvesting. This method has greater potential to damage advance regeneration than those in which trees are processed at-stump. Hybrid tree-length (Hyb TL) harvesting using an stroke-boom delimber for in-woods processing might be a feasible alternative, but little is known about the production economics of this method. An experimental strip-cutting study was conducted in central Maine, US in the winter of 2018 to: (1) evaluate and compare operational productivity and costs of ground-based Hyb TL and WT methods; (2) identify factors influencing productivity of at-stump and at-landing log processing; and (3) calculate best management practice (BMP) implementation costs in WT harvesting. Time-motion data were recorded for operational phases such as felling, extraction, processing, sorting and loading; machine rates were calculated to determine productivity and costs of operations. Total cost of Hyb TL (US $17.01 m−3) was lower than that of WT ($18.38 m−3). Processing cost was lower at-stump than at-landing ($2.66 and $2.73 m−3 for Hyb TL and WT, respectively). This is likely due to fewer logs handled per cycle at-landing (1.2 logs per turn) compared to the number handled per cycle at-stump (1.4 logs per turn). Sensitivity analysis showed that a 30-m increase in average in-woods distance travelled by the delimber would result in a 41 per cent increase in the processing cost. Cost of BMP implementation in WT was $2.25 m−3 or $59.2 per productive machine hour. Results suggest that it is feasible to apply Hyb TL method in an industrial harvesting operation, though distance of in-woods delimber movement influences processing costs. Insights from this study will help forest managers and loggers efficiently plan and execute harvesting operations.

Description: Early detection of infestation by the emerald ash borer (EAB), Agrilus planipennis is extremely difficult; hence developing additional methods is desirable. We built on the successful use of canine scent detection for the invasive long-horned beetles Anoplophora glabripennis and Anoplophora chinensis and trained six dogs in detection of EAB. A first test series was performed to evaluate detection accuracy of five of these dogs. Seven different experimental settings were tested under single blind conditions: (1) forest nursery, (2) piles of firewood, (3) firewood on the ground, (4) ash logs on the ground, (5) old urban ash trees, (6) urban forest with ash trees and (7) natural forest with ash trees. In total, 214 positive samples were presented to the dogs, out of which 20 remained undetected. The experiments ascertained sensitivity (correct positives of all positives) ranging from 73.3 to 100 percent and specificity (correct negatives of all negatives) from 88.9 to 99.8 percent in the tested settings. This initial study demonstrates that trained dogs are able to detect EAB scent from sources such as larval galleries in bark/wood, frass, living or dead larvae or dead dry beetles. The numbers of tested dogs and test series were limited, and further studies are needed to confirm the initial results. However, the preliminary findings demonstrate the potential of the method particularly for inspection of wood or plants at entry points.

Description: Methods for the three-dimensional (3D) reconstruction of forest trees have been suggested for data from active and passive sensors. Laser scanner technologies have become popular in the last few years, despite their high costs. Since the improvements in photogrammetric algorithms (e.g. structure from motion—SfM), photographs have become a new low-cost source of 3D point clouds. In this study, we use images captured by a smartphone camera to calculate dense point clouds of a forest plot using SfM. Eighteen point clouds were produced by changing the densification parameters (Image scale, Point density, Minimum number of matches) in order to investigate their influence on the quality of the point clouds produced. In order to estimate diameter at breast height (d.b.h.) and stem volumes, we developed an automatic method that extracts the stems from the point cloud and then models them with cylinders. The results show that Image scale is the most influential parameter in terms of identifying and extracting trees from the point clouds. The best performance with cylinder modelling from point clouds compared to field data had an RMSE of 1.9 cm and 0.094 m3, for d.b.h. and volume, respectively. Thus, for forest management and planning purposes, it is possible to use our photogrammetric and modelling methods to measure d.b.h., stem volume and possibly other forest inventory metrics, rapidly and without felling trees. The proposed methodology significantly reduces working time in the field, using ‘non-professional’ instruments and automating estimates of dendrometric parameters.

Description: Emerald ash borer (EAB), an invasive forest insect from Asia, has impacted vast areas in the United States and European Russia. To proactively understand the social impact of an EAB invasion in Europe and the USA, this study analyzed visitors’ preferences and preference heterogeneity for EAB-impacted forest scenarios in Vienna, Austria (n = 510) and Minneapolis, USA (n = 307). An image-based discrete choice experiment with latent-class analysis among on-site completed questionnaires in Vienna indicated four different visitor segments based on trade-offs among biophysical, social and viewscape elements. Within the forested environment, two segments placed greater importance on (bio)physical attributes and two on social aspects. Although all segments preferred a non-impacted ash forest, only one of the four identified the attribute describing EAB impacts and forest management as the most important attribute. Rather, visitor numbers and background viewscapes were more important than EAB impact and management to differentiate landscape preferences for three of the four segments. Differences in preferences were found between the Vienna and Minneapolis samples. Vienna respondents showed a higher preference for more natural conditions, disliked more the initial stage of EAB impact and placed more importance on background viewscapes and visitor numbers. Forest managers and greenspace planners need to consider the entirety of the forested condition, social and visual, for effective management and address that visitors differ in their preferences for all of these conditions.

Description: Collaboration has been the key to success for urban forest management in Colorado, not only collaboration amongst agencies at all levels of government but also in engaging industry allies, coordinating education and outreach efforts and in fostering community support. A unique interagency team, the Emerging Pests in Colorado (EPIC) Workgroup, was formed in 2009 to address the immediate threat from Thousand Cankers Disease (TCD) of walnut and to plan for the arrival of other invasive urban forest pests to Colorado. When the emerald ash borer (Agrilus planipennis Fairmaire) (EAB) was detected in Boulder, Colorado in 2013, it marked the westernmost occurrence of EAB in the US, threatening millions of planted and naturalized ash trees representing over 25 percent of the tree canopy throughout Colorado’s urban and riparian forests. The detection in Boulder prompted the development of a second multi-agency group, the Colorado EAB Response Team (CORT). The preparedness and established working relationships between stakeholders and responsible authorities allowed for a quick, decisive and unified response. We review as a case study: (1) the formation and history of collaborative interagency groups in Colorado; (2) how the interagency collaborative planning and post-detection EAB response have supported community forestry programmes throughout the state; (3) development of the post-detection EAB management plan and economics behind the strategy in Boulder, Colorado; and (4) the proactive EAB planning and outreach efforts underway in Denver, Colorado.

Description: Harvesting practices in temperate mixedwoods of eastern North America have a history of diameter-limit cuts, which have often resulted in degraded residual stands. In this study, we examined the factors influencing stand basal area (BA) and tree diameter growth in previously high-graded mixedwood forests, to understand which stands are more likely to recover from high-grading. Over 15 years, we monitored tree growth, recruitment and stem quality of 532 sample plots that were located in high-graded stands of Quebec’s mixedwood forest. We found that diameter growth rates were positively correlated with precipitation-related variables for balsam fir (Abies balsamea) and for yellow birch (Betulla alleghaniensis) but opposing trends for temperature-related variables were found. Conversely to balsam fir, yellow birch growth was positively correlated to temperature variables. Our results also show that BA growth was greater for plots with a larger acceptable growing stock (AGS: trees with potential sawlog production) and that the increase in AGS was greater for plots with larger amount of conifer BA. These result highlights the importance to maintain a proportion of conifer trees in these mixed stands. Moreover, the significant effect of asymmetric competition in our study underscores the relevance of considering the spatial distribution when choosing crop trees.

Description: European ash (Fraxinus excelsior L.) is a highly valued native species in the UK and elsewhere. The rapid spread and predicted impact of ash dieback (caused by Hymenoscyphus fraxineus), has led researchers to explore a range of responses from introducing non-native ash to use of Genetic Modification (GM) technologies to develop disease resistance. To support decisions over which approaches should receive further investment to encourage adoption, it is necessary to understand what is acceptable to forest practitioners. Interviews with 22 forest advisors and managers revealed ash is highly valued for the multiple benefits it provides but past experiences with pest and diseases have encouraged a more cautious (‘wait and see’) attitude to approaches for dealing with ash dieback. These practitioners showed a strong interest in the concept of resistant ash but emphasized that any ‘new’ varieties should have similar characteristics, retain genetic diversity and be able to withstand future pests. There was limited support for planting non-native ash and use of GM methods and greater support for traditional breeding, though views varied dependent upon objectives and could evolve with experience countering the disease. There were concerns about the time required to produce resistant varieties and more information was needed about the costs, benefits and impacts of the different approaches. These findings point to the need for continued communication and engagement with stakeholders in the refinement of responses to dieback and any development of resistant ash.

Description: In this study, we investigate the relationship between the mean wind-induced load on an open-grown oak tree and the mean wind speed. The estimation of the wind load is based on bending moment observations on the stem, using strain gauges installed close to the base of the tree. The measurement campaign included periods where the crown was both with and without foliage. A new post-processing method to compensate for time- and temperature-dependent drifts of long-term strain gauge measurements is presented. The time-scale, at which the method is applied, is determined from the observed high coherence between the wind speed and the induced bending moments. The analysis shows that the wind load on the tree is proportional to the wind speed raised to an exponent of 1.60–1.68 and 1.90–2.01 in the case of a crown with and without leaves, respectively. The results indicate that the presence of foliage increases the wind-induced load by a factor of 2–3. The impact of the wind speed range on these results is discussed and compared to the measured wind deficit in the lee of the tree. The application of this method can potentially be extended to other tree mounted sensors.

Description: Mixed mountain forests of European beech (Fagus sylvatica L.), Norway spruce (Picea abies (L.) Karst), and silver fir (Abies alba Mill.) cover a total area of more than 10 million hectares in Europe. Due to altitudinal zoning, these forests are particularly vulnerable to climate change. However, as little is known about the long-term development of the productivity and the adaptation and mitigation potential of these forest systems in Europe, reliable information on productivity is required for sustainable forest management. Using generalized additive mixed models this study investigated 60 long-term experimental plots and provides information about the productivity of mixed mountain forests across a variety of European mountain areas in a standardized way for the first time. The average periodic annual volume increment (PAI) of these forests amounts to 9.3 m3ha−1y−1. Despite a significant increase in annual mean temperature the PAI has not changed significantly over the last 30 years. However, at the species level, we found significant changes in the growth dynamics. While beech had a PAI of 8.2 m3ha−1y−1 over the entire period (1980–2010), the PAI of spruce dropped significantly from 14.2 to 10.8 m3ha−1y−1, and the PAI of fir rose significantly from 7.2 to 11.3 m3ha−1y−1. Consequently, we observed stable stand volume increments in relation to climate change.

Description: A wealth of recent research has improved our understanding of the structure, growth and yield of mixed-species stands. However, appropriate quantitative concepts for their silvicultural regulation remain scarce. Due to the species-specific stand densities, growing area requirements and potential over-density, the density and mixing regulation in mixed stands is much more intricate than in monospecific stands. Here, we introduce the species-specific coefficients: density equivalence coefficients (DEC), for density equivalence; and density modification coefficient (DMC), for density modification in mixed species stands. DEC is suitable for the conversion of the stand density and growing area requirement of one species into that of another species. DMC estimates the modification of maximum stand density by tree species mixing using as reference the maximum stand density of one of the species. First, we introduce the theoretical concept of these coefficients. Second, we derive the mean values of these coefficients based on long-term experiments using different mixtures of European beech. Third, we apply DEC and DMC for flexible regulation of the stand density and mixing proportion. Thus, silvicultural regulation of monospecific stands and mixed-species stands forms a continuum, where monospecific stands represent an extreme case of mixed-species stands. Lastly, we discuss the advantages and limitations of these concepts. Future directions comprise the inclusion of additional species, their integration in guidelines and simulation models, and their establishment for the quantitative regulation of experimental plots and the practical implementation in forest stands.

Description: Tree growth is expected to be responsive to climatic drivers across a spectrum of temporal scales, ranging from yearly growth to daily water use and photosynthesis. Automatic dendrometers offer the potential to provide continuous high-resolution measurements of tree radius changes. The signal recorded contains three components: (1) a long-term seasonal growth component, (2) a mid-term component representing swelling after rainfall and subsequent drying and (3) daily cycles of water-uptake related to tree transpiration. For 91 trees at 4 sites (Picea abies: 58, Pinus cembra: 17, Fagus sylvatica: 14, Pinus sylvestris: 2) monitored in Austria between 2012 and 2015, we simultaneously modelled these three processes using a hierarchical nonlinear mixed-effects model represented by two logistic growth curves. The focus was on the mid-term and daily component, and therefore long-term growth that is typically modelled by including tree size, competition or site variables was represented by random effects only. Both mid-term and short-term components were species-specific. In general, P. cembra and F. sylvatica were less sensitive to climate variables than P. abies. For all species, the mid-term component was best represented using a 14-day moving average difference between rainfall and potential evapotranspiration, a 24-h moving average of precipitation and its 1–3 days lags, a 24-h moving average temperature and its 1–3 days lags. The daily cycles of water uptake were best related to hourly humidity and its 3-h lag, and interactions with the 14-day moving average difference between rainfall and potential evapotranspiration accounted for attenuating cycles after rainy events and increasing cycles in dry periods.

Description: This paper presents a spatially explicit methodology for integrated forest management and wood supply chain optimization over time in the context of a Finnish forest strategy anticipating new investments and renewal of business in the wood processing industry. The Finnish MELA simulator was used to generate multiple treatment schedules over time at the management unit level – each treatment schedule providing unique estimates of extracted wood volumes by different assortment categories for each time period. The J linear programming (LP) software was used to analyse different regional forest strategies in terms of wood supply and transportation costs to multiple market destinations. The analysis revealed clear differences both in wood flows and forest resources between strategies maximizing wood supply or optimizing wood supply to market destinations. In addition, the wood flows appeared responsive to new mill and increased demand. Further, the changes in factory price had a heavier impact on pulpwood supply than on sawlog supply. The same methodology can be applied for analysing the impact of new factories on wood flows from forest to factories and between factories or to support forest enterprises in planning their wood supply over multiple time periods and multiple destinations.

Description: The Asian chestnut gall wasp (ACGW), Dryocosmus kuriphilus (Hymenoptera: Cynipidae), is an invasive pest that affects chestnut trees. The first record of this pest in Spain was in 2012, in Catalonia, and it is now distributed in virtually every chestnut growing area in the country. In this study, we present an overview of parasitoid recruitment by ACGW in Catalonia over a 4-year period (2013–2016) comparing parasitoid communities attacking galls on oak and chestnut trees at the same sites. A total of 22 species of native parasitoids that normally attack oak cynipids emerged from ACGW galls, together with the non-native Torymus sinensis. The most abundant species were Bootanomyia dorsalis, Torymus flavipes, Ormyrus pomaceus and Eupelmus urozonus. The study also found that 20 of the 22 native parasitoids in ACGW galls were also reared from galls of 13 species of oak gall inducer (12 Cynipidae, 1 Cecidomyiidae).

Description: This study addresses the use of multiple sources of auxiliary data from unmanned aerial vehicles (UAVs) and airborne laser scanning (ALS) data for inference on key biophysical parameters in small forest properties (5–300 ha). We compared the precision of the estimates using plot data alone under a design-based inference with model-based estimates that include plot data and the following four types of auxiliary data: (1) terrain-independent variables from UAV photogrammetric data (UAV-SfM); (2) variables obtained from UAV photogrammetric data normalized using external terrain data (UAV-SfMDTM); (3) UAV-LS and (4) ALS data. The inclusion of remotely sensed data increased the precision of DB estimates by factors of 1.5–2.2. The optimal data sources for top height, stem density, basal area and total stem volume were: UAV-LS, UAV-SfM, UAV-SfMDTM and UAV-SfMDTM. We conclude that the use of UAV data can increase the precision of stand-level estimates even under intensive field sampling conditions.

Description: We present a recruitment model for pure and mixed beech and oak stands in Belgium, the first empirical model for this forest type in this geographical area. Data from the Wallonia National Forest Inventory were used to fit the model. We adopted a zero-inflated formulation where model parameters governing species’ behaviour were simultaneously fitted. Plot random effects specific to each species were included, the simultaneous fit allowing them to correlate. Model predictions proved accurate and corresponded to current ecological knowledge about the regeneration dynamics of this kind of mixture. While our model could potentially be used to complement the existing beech and oak growth models for this region of Europe, our results also show that beech recruits tend to dominate regardless of the oak share in the overstorey composition and the stand stocking. This confirms that the beech–oak mixture may not be stable under the conditions of the study area and current management aimed at promoting continuous forest cover.

Description: Loblolly pine (Pinus taeda L.) is one of the most widely planted tree species globally. As the reliability of estimating forest characteristics such as volume, biomass and carbon becomes more important, the necessary resources available for assessment are often insufficient to meet desired confidence levels. Small area estimation (SAE) methods were investigated for their potential to improve the precision of volume estimates in loblolly pine plantations aged 9–43. Area-level SAE models that included lidar height percentiles and stand thinning status as auxiliary information were developed to test whether precision gains could be achieved. Models that utilized both forms of auxiliary data provided larger gains in precision compared to using lidar alone. Unit-level SAE models were found to offer additional gains compared with area-level models in some cases; however, area-level models that incorporated both lidar and thinning status performed nearly as well or better. Despite their potential gains in precision, unit-level models are more difficult to apply in practice due to the need for highly accurate, spatially defined sample units and the inability to incorporate certain area-level covariates. The results of this study are of interest to those looking to reduce the uncertainty of stand parameter estimates. With improved estimate precision, managers, stakeholders and policy makers can have more confidence in resource assessments for informed decisions.

Description: The southernmost portion of the Andes in South America hosts Nothofagaceae forests that form ectomycorrhizal (EcM) associations. We compiled all the published reports of EcM taxa from these woodlands, based on fruit-body collections and molecularly identified root tips. This resulted in 87 peer-reviewed research papers dealing with EcM associations from Nothofagaceae forests published over the past 62 years. Based on these papers the EcM richness and its association with Nothofagaceae species was analyzed. Represented by 45 genera (34 Basidiomycetes and 11 Ascomycetes), 479 EcM taxa have been recorded, plus an additional 46 EcM taxa which are potentially EcM. Cortinarius was the most frequent and diverse genus, with 64.9% of total species. Nothofagus dombeyi had the highest number of unique EcM species, followed by N. pumilio, N. antarctica and Lophozonia obliqua. The EcM community associated with Nothofagus species, except N. nitida, showed higher similarities among themselves, than with Lophozonia and Fuscospora species. Beta diversity indexes showed EcM Nothofagus species have 29–42% niche overlap, while Nothofagus—Lophozonia only had 1–16%. The assessment of the accuracy of the EcM diversity, host specificity and community structure deserve further studies combining phylogenetic analysis based on broad ecological surveys. On the basis of pre-selected criteria Austropaxillus statuum, Cortinarius fragilis, Cortinarius xiphidipus and Hallingea purpurea are proposed as candidates for nursery spore inoculations, and should be subject to scientific evaluation to determine costs and benefits.

Description: Modern silviculture is based on simulating natural disturbance regimes on the one side and reducing their impact on the other side. Disturbances modify light regimes of forests along complex spatial patterns, and species participating in the ecosystem tend to have unique adaptations to the multitude of light conditions found in forests. We studied the height growth reaction of saplings from eight native North American tree species to different light availabilities in a Sierra Nevada mixed conifer forest. Large height growth differences occurred between species. Height growth increased for most species with increasing light availability. However, black oak but also incense cedar and tanoak reached an optimal value or saturation point of light availability at intermediate light levels, above which their growth stayed the same or slightly declined on average. A variety of stand conditions, representing the range from early to late successional stages, are beneficial to maintain tree species diversity across stands. Such a gradient of conditions can be achieved by diversifying silvicultural regimes and using regeneration approaches that mimic a range of disturbance intensities. The results add to knowledge on the species’ ecology by specifying their height growth in dependence of a quantified light environment. The quantification of the light environment can directly be linked to % canopy cover and a range of regeneration methods, respectively. With this, the results may help achieve a more precise and target-oriented management of Sierra Nevada mixed conifer forests.

Description: Knowledge of the spatial distribution of aboveground biomass (AGB) is crucial to guide forest conservation and management to maintain carbon stocks. LiDAR has been highly successful for this purpose, but has limited availability. Very-high resolution (