ALBERT

Description: Nachdem die fossilen Früchte und Samen aus dem Ton von Reuver, von C. und E. M. Reid, und die Blätter von Laurent und Marty eingehendst studiert worden sind, schien es mir erwünscht auch die Hölzer, die in so grosser Zahl in den Tonschichten Vorkommen einer Untersuchung zu unterwerfen. In der Tongrube der Firma Teeuwen zu Reuver, die ein wenig jenseits der holländischdeutschen Grenze auf deutschem Gebiet am Nordrand des Icksbergs gelegen ist, wurden von mir am 16. September 1930 aus jeder der Schichten des damals blossgelegten Profils eine grosse Anzahl von Holzresten gesammelt. Die Anfertigung der Schnitte für microscopische Untersuchung war nicht schwer, ein gewöhnliches Rasiermesser genügte zu diesem Zweck. Nach einer Behandlung mit kochender Kalilauge gewannen sie sehr an Deutlichkeit und wurden mit einer Saffraninlösung gefärbt.

Description: 1. A review is given of some phenomena concerning pregnancy and parturition of the Cetacea, depending on data in literature and on observations made in Antarctic Blue and Fin Whales on board the f.f. \xe2\x80\x9cWillem Barendsz\xe2\x80\x9d (1946\xe2\x80\x941947). 2. In Mystacoceti the frequency of twins appears to be less than in man and the big domestic animals. The majority of twins is born by females that are longer than the average female in the period of greatest sexual activity. This does not mean, however, that just as in man most twins are born at an elder age than that corresponding with the maximum of sexual activity. It may also be possible that, just as in the big domestic animals, the ages correspond, but that twins are mostly produced by the physically stronger developed females. 3. In Odontoceti the left ovary shows a very distinct morphological and functional prevalence. The foetus is exclusively found in the left uterine cornu. In Mystacoceti there is a prevalence of about 60 % of the right ovary and the right uterine cornu with regard to ovulation and pregnancy. The above described phenomena have also been observed in other uniparous Mammals, whereas in multiparous Mammals no distinct prevalence of a special side has been found. Transference of an ovum from the ovary of one side to the cornu of the other has been observed twice in Cetacea. 4. In Mystacoceti the number of cephalic presentations of the foetus very distinctly increases during the last months of pregnancy, just as in man and the big domestic animals. There is a great possibility that, just as the other uniparous Mammals, which give birth to comparatively large infants, in Mystacoceti a very high percentage of the young is born in cephalic presentation. In Odontoceti, on the contrary, a great number of foetuses is apparently delivered in tail-presentation. With regard to their shape and dimensions, such a birth in tail-presentation must be considered as an unfavourable event. An attempt has been made to explain these facts with the aid of the peristaltic uterine contractions. If these contractions act in the same way as in other Mammals, it might be expected that most of the Cetacea should be born in tailpresentation. So it is highly possible that in Mystacoceti some other factors are responsible for the high percentage of cephalic presentations. 5. In Cetacea the relative length of the umbilical cord (in % of the length of the foetus) decreases markedly during the second part of pregnancy. At birth its length is about 40 % of the total length of the calf and 57 % of its snout-anus length. As compared with other Mammals the cord of the Cetacea is rather short and this fact may have some influence on the way in which the connection between mother and calf is broken. This may occur by rupture of the cord immediately after birth as in Ungulates. On the other hand it appears to be also possible that, just as in Primates, Carnivores and Chiropteres, the placenta and the cord stick to the baby for some time after birth. 6. Some congenital anomalies of foetal Cetacea are described. Abortus probably may occur during chasing of the big whales or when they are struck by the harpoon. A case of fibromyoma uteri is described in an old female Blue Whale that showed an abnormal lactation. Probably the fibromyoma had caused an abortus some months ago, this abortus causing the lactation.

Description: 1. Researches were made on 50 hearts of 9 different Pinnipeds, of the Hippopotamus and the Otter. 2. In the Northern Fur Seal, the Grey Seal, the Ringed Seal, the Bearded Seal, the Banded Seal and the Walrus the ductus arteriosus closes anatomically at the same time, or even earlier, than in terrestrial Mammals of comparable size. In contrast to those of the Common Seal the newborn pups of these animals remain on land for several weeks, or, if they swim during the first weeks after birth, they do so infrequently and they do not dive.\nThese data confirm the supposition that the retarded anatomical closure of the ductus arteriosus in Cetaceans and in the Common Seal may be explained by the fact that their newborns swim and dive immediately or almost immediately after birth. Shortage of oxygen (or eventually a rise of the pressure in the pulmonary artery) may cause a temporary re-opening of the ductus during the first period of postnatal life, which retards its anatomical closure. 3. All Pinnipeds show a more or less retarded closure of the foramen ovale during the first period of postnatal life. Obviously this phenomenon is not directly related with the closure of the ductus arteriosus. 4. In most Pinnipeds a ramification of the aortic arch was found, which corresponds with the situation in Man. Some animals show a more concentrated type, like the Dog and the Pig. The Walrus has a more dispersed type. 5. Data are given about the shape of the heart. This shape and the type of ramification of the aortic arch are highly correlated with the spatial relations in the thoracic cavity. 6. The ratio of the thickness of the right and left ventricular wall in Pinnipeds is the same as in terrestrial Mammals. This is the case in newborns as well as in young and adult animals.

Description: A.\nGENERAL REMARKS\nDuring three years 4500 reports of whales sighted from Netherlands ships were collected, bearing on approximately 11.000 individual animals. Most of the observations were made in the Atlantic and Indian Ocean. It was supposed that the species could be determined with a fairly high degree of reliability in the case of Humpback Whales, Sperm Whales and Right Whales. No distinction could be made between Blue, Fin, Sei and Bryde Whales. They were collected under the heading Rorquals. Catches of land stations and strandings of whales, however, indicate that in all areas, at least a part of these Rorquals must have been Blue or Fin Whales. Probably the majority of this part were Fin Whales.\nNevertheless it must he emphasized that the observations give no exact figures but only indications. It would be highly desirable if the results could be controlled by observations made by experienced whale biologists or gunners, especially in tropical and subtropical waters. We have the impression that for the time being no better results can be obtained with the present type of research. On the other hand, the fact that the generally known facts about the annual migration of the big whales were also clearly shown by this research, may be an indication for a certain degree of reliability of the research. The monthly number of animals of each species observed per 1000 hours steamed in daylight was plotted on charts in ten degrees squares. The reliability of the converted data is highest in the black dots.\nB. RORQUALS AND HUMPBACKS 1. Distribution The animals involved are not evenly distributed over the Oceans. There are big concentrations in certain areas, whereas in other areas practically no whales occur. Broadly outlined the highly populated areas coincide with the areas of greatest biological productivity of the sea, as shown by WALFORD (1958).\nIn the tropics and subtropics important areas with a great number of sighted whales are: the Caribbean, the North African west coast, the Atlantic coast of South Africa, the Arabian Sea, the Gulf of Aden, the Bay of Bengal, the Indonesian Archipelago and the African east coast between 30\xc2\xb0 S and 40\xc2\xb0 S. It could be demonstrated that in the Indian Ocean southern Rorquals migrate over the entire breadth of the Ocean south of 30\xc2\xb0 S. North of 30\xc2\xb0 they migrate only at the eastern and the western side, apparently in order to avoid the waters with low biological productivity in the central part of the Ocean.\nNo special relationship was found between the distribution or the migratory routes of the whales and the course of the big Ocean currents with regard to the locomotory aspect. There was a special relationship only in those cases where the big currents show a great biological productivity, as for example the Gulf Stream and the currents in the northern part of the Indian Ocean. 2. Migration, general remarks With regard to Rorquals in the North and South Atlantic, the Indian Ocean and the Pacific Ocean, as well as with regard to Humpbacks in the Atlantic and in the Indian Ocean it could be demonstrated that during the summer a part of the population does not migrate into Arctic or Antarctic waters, but that it stays in tropical, subtropical or temperate waters. In Humpbacks the phenomenon is less pronounced than in Rorquals. In Rorquals the phenomenon is not caused by observations of Sei or Bryde Whales only, because catches of land stations and strandings show that Blue and Fin Whales are present during the summer in the waters involved as well. The percentage of the stock of Blue and Fin Whales staying behind in warm and temperate waters is not known, but the authors have the impression that it is not unsignificant. The number of Rorquals staying behind during the summer appears to be larger in the North than in the South Atlantic, probably because Fin Whales in the North Atlantic feed on fish.\nThe phenomenon of staying behind of a part of the population confirms the assumption that estimations of the Antarctic population of Blue, Fin or Humpback Whales never bear on the total stock of the species involved. The phenomenon may also cause that the number of periods or laminations in baleen plates or ear plugs, used in determining the age of Whalebone Whales, is not a reliable indication for the actual age of the animals. The actual age may be higher than the number of periods, because the staying behind in warm waters causes irregularities in their formation.\nIt could, however, be demonstrated, that in most areas the majority of the populations showed the generally accepted type of annual migration. 3. Migration, Rorquals In the North Atlantic the principal northward migration of Rorquals takes place in April-July, the southward migration in September-November. In the South Atlantic the period of migration southward is September-December, that of the northward migration March-June.\nThe majority of the Rorqual population (which may be principally the Fin Whale population) lives in the North Atlantic during the northern winter between 0\xc2\xb0 and 40\xc2\xb0 N and during the northern summer between 30\xc2\xb0 N and the border of the pack ice. With regard to the South Atlantic these areas are: in the southern winter between 20\xc2\xb0 N and 50\xc2\xb0 S, in the southern summer between the equator and the pack ice, but mainly in Antarctic waters.\nThe northern and southern population apparently meet in the Caribbean, in waters off the North African west coast and probably also in the central part of the Ocean between 0\xc2\xb0 and 20\xc2\xb0 N.\nIn the Indian Ocean large concentrations of Rorquals have been encountered in the northern part of the Ocean during the southern summer, whereas the number of sightings during the southern winter is surprisingly small. During this season the majority of the Rorquals is concentrated in waters of Madagascar and off the Australian west coast. This suggests, that during the southern summer (northern winter) the northern part of the Ocean is populated by Rorquals coming from the northern part of the Pacific Ocean. Probably these whales enter the Indian Ocean by passing the waters of the Indonesian Archipelago and the straits between these waters and the Indian Ocean. This supposition is supported by the fact that in the northern part of the Indian Ocean calves have been sighted in almost equal monthly numbers during the whole year, whereas in the Atlantic Ocean seasonal peaks in the number of sightings have been demonstrated. On the other hand, the possibility of a local stock in the northern part of the Indian Ocean may not be excluded.\nAlthough a number of southern Rorquals certainly migrate into the northern part of the Ocean during the southern winter, the majority of the population probably live in this season between the equator (or 10\xc2\xb0 S) and 30\xc2\xb0 S. In the southern summer the majority of the population is found in Antarctic.\nIn the North Pacific Ocean the majority of the population is found during the northern summer between 20\xc2\xb0 N and the pack ice and in the northern winter between 10\xc2\xb0 N and 30\xc2\xb0 N. The South Pacific population apparently migrates northward during the southern winter up to about 10\xc2\xb0 N. 4. Migration, Humpbacks Humpbacks appear to migrate principally in coastal waters with the exception of the crossing part of the Gulf Stream in the North Atlantic (30\xc2\xb0 N to 50\xc2\xb0 N) where they are found during the northern winter over the entire breadth of the Ocean. In the northern part of the Indian Ocean they are spread over a large part of the Ocean as well.\nIn the North Atlantic the majority of the population is found during the northern summer between 30\xc2\xb0 N and 50\xc2\xb0 N, and during the northern winter between 40\xc2\xb0 N and 10\xc2\xb0 S (especially in the Caribbean and off the North African west coast). Probably all Humpbacks in the Caribbean belong to the northern stock, because the southern population appears to live during the southern winter between 30\xc2\xb0 S and 20\xc2\xb0 N at the African side of the Ocean, but between 30\xc2\xb0 S and the equator at the American side. During the southern summer they are found between 30\xc2\xb0 S and the pack ice (mostly in Antarctic waters). In former days the North Atlantic Humpback population probably lived further northward (in summer as well as in winter) than nowadays. This may be connected with changes in feeding conditions or with the general decrease of the stock.\nJust as has been shown with regard to Rorquals, a part of the North Pacific Humpback population seems to migrate into the northern part of the Indian Ocean during the northern winter. The southern population of the Indian Ocean lives during the southern winter between the continent and 30\xc2\xb0 S. During the southern summer the animals are found between 45\xc2\xb0 S and the border of the pack ice.\nThe northern and southern stocks of the Pacific Ocean meet in waters of the Indonesian Archipelago. At the eastern (American) side of the Ocean the northern population lives during the summer between 30\xc2\xb0 N and 50\xc2\xb0 N (or farther northward). During the winter they live between 10\xc2\xb0 N and 30\xc2\xb0 N. The southern stock appears to migrate as far to the North as 10\xc2\xb0 N. 5. Calves Sightings of calves of Rorquals (probably the majority of them being Fin Whales) in the Atlantic Ocean point to a peak in the number of births in December-January for the northern population and in May-June for the southern stock.\nNorth Atlantic Humpbacks appear to be born principally in the southern part of the North Atlantic in April, whereas births of the southern stocks apparently occur in tropical waters with a peak in September.\nC.\nSPERM WHALES\n1. North Atlantic Sperm Whales are always present in the North Atlantic between 10\xc2\xb0 S and 30\xc2\xb0 N, but on the African side the population appears to be much larger than on the American side. A great number of animals are sighted in the Gulf Stream during the summer. The northward migration starts in April, the animals return to the South in autumn. The majority of the females do not go farther to the North than 40\xc2\xb0 N (a minority probably up to 50\xc2\xb0 N). The males migrate into Arctic waters. During the northern winter the majority of the males and females apparently live between 10\xc2\xb0 S and 30\xc2\xb0 N (the American stock mostly in the Caribbean), but some males may stay behind in colder waters as far as 60\xc2\xb0 N. 2. South Atlantic Practically no sightings of Sperm Whales have been reported from the South American east coast, although these waters show a reasonable biological productivity and although a great number of Rorquals have been sighted there. In former days great numbers of Sperm Whales have been caught in these waters. During the summer the males migrate into Antarctic waters, the females migrate up to about 40\xc2\xb0 S. During the winter most of the animals live in tropical waters but some males and females are present up to 40\xc2\xb0 S. 3. Indian Ocean With regard to the Indian Ocean there is a very significant correlation between the distribution of Sperm Whales and the biological productivity of the sea. In the northern part of the Ocean there are many more Sperm Whales sighted during the northern winter than during the northern summer.\nThe general seasonal movements described with regard to the Atlantic and Pacific Oceans could not be demonstrated in the Indian Ocean. Apparently the Sperm Whales in this region show very special migratory movements which may be correlated with special conditions, caused by the fact that the Monsoon-stream in the northern part flows in an opposite direction in the two halves of the year. 4. Pacific Ocean Sperm Whales are encountered in the Indonesian Archipelago the whole year round. In the South Pacific they are not evenly distributed but apparently they are restricted to certain areas. The normal seasonal migratory movements could be demonstrated with regard to the South Pacific.\nD.\nOTHER SPECIES\n1. Little Piked Whales Fairly large numbers of this species were sighted throughout the whole year in tropical waters of all oceans. Large herds were also seen in the northern hemisphere. They show concentrations in areas with a high biological productivity of the sea. During the winter the majority of the animals apparently live in tropical and subtropical waters. During the spring and the autumn they show the usual migratory movements, just as Rorquals and Humpbacks. During the winter, however, some animals stay behind in northern waters, whereas during the summer there are some stragglers in warm waters.\nThe species has been observed in the northern part of the Indian Ocean during the northern winter. In the North Atlantic births take place in warm or temperate waters, probably from November to March. 2. Californian Grey Whales Sightings in the North Pacific were quite in accordance with the generally accepted opinion about the migration of this species. 3. Right Whales North Atlantic as well as Southern Right Whales have been reported. The majority of the animals do not migrate into waters between 20\xc2\xb0 N and 20\xc2\xb0 S, but there are indications that a few animals may also visit these tropical waters.\nWith regard to the North Atlantic no sightings have been reported from regions north of 50\xc2\xb0 N, whereas there was a large number of sightings between 20\xc2\xb0 N and 50\xc2\xb0 N during the northern summer.\nIn the Indian Ocean and in the Indian Archipelago two sightings were reported from waters between 10\xc2\xb0 N and 10\xc2\xb0 S. These observations, however, need further confirmation.

Description: Nachdem die fossilen Fr\xc3\xbcchte und Samen aus dem Ton von Reuver, von C. und E. M. Reid, und die Bl\xc3\xa4tter von Laurent und Marty eingehendst studiert worden sind, schien es mir erw\xc3\xbcnscht auch die H\xc3\xb6lzer, die in so grosser Zahl in den Tonschichten Vorkommen einer Untersuchung zu unterwerfen. In der Tongrube der Firma Teeuwen zu Reuver, die ein wenig jenseits der holl\xc3\xa4ndischdeutschen Grenze auf deutschem Gebiet am Nordrand des Icksbergs gelegen ist, wurden von mir am 16. September 1930 aus jeder der Schichten des damals blossgelegten Profils eine grosse Anzahl von Holzresten gesammelt.\nDie Anfertigung der Schnitte f\xc3\xbcr microscopische Untersuchung war nicht schwer, ein gew\xc3\xb6hnliches Rasiermesser gen\xc3\xbcgte zu diesem Zweck. Nach einer Behandlung mit kochender Kalilauge gewannen sie sehr an Deutlichkeit und wurden mit einer Saffraninl\xc3\xb6sung gef\xc3\xa4rbt.

Description: At the Royal Zoological Gardens \xe2\x80\x9cBlijdorp\xe2\x80\x9d at Rotterdam May 6th 1958 a Father David\xe2\x80\x99s Deer gave birth to a female calf. It was the first young of this three year old doe.\nDuring the days before the day of birth the doe was seen several times leaping upon the buck. Experience with other Ungulates has taught that such behaviour may be regarded as an indication that the time of delivery is approaching. On the day of birth the doe refused her food as has been observed in many other mammals. She lay down very often, gnashed her molar teeth and made movements with her head towards the belly and the udder, the ears flattened to the neck, as if she was suffering from cramps. One had the impression, however, that the expulsion of the young was kept back until the keeper went home and the door of the stable was closed. Apparently a great number of mammals prefer to give birth to their young in the quietest part of the space of 24 hours. For most mammals this is the evening or the night, but bats very frequently give birth during the day which is their time of rest (SLIJPER, 1959). A postponement of birth until all is quiet has frequently been observed in zoological gardens. In natural surroundings the Ringed Seal (Phoca hispida Schreb.) and other Seals are said to be able to postpone birth as long as 10 days if the weather is very bad (KRUMBIEGEL, 1947).

Description: INHALTS\xc3\x9c\nBERSICHT\nAbschnitt 1 Einleitung; Material .... 243\nI Strandung .... 243\nII Material .... 244\nAbschnitt 2 Nomenklatur; Geographische Verbreitung; Lebensweise .... 245\nI Nomenklatur .... 245\nII Geographische Verbreitung .... 246\nIII Lebensweise .... 249\nAbschnitt 3 \xc3\x84ussere Form; Wachstum .... 250\nI \xc3\x84ussere Form .... 250\nII Wachstum .... 251\nAbschnitt 4 Skelett .... 252\nI Wirbels\xc3\xa4ule .... 252\nII Rippen .... 255\nIII Sternum .... 255\nIV Becken .... 257\nV Vorderextremit\xc3\xa4t .... 258\nVI Sch\xc3\xa4del .... 260\nVII Z\xc3\xa4hne .... 260\nAbschnitt 5 Muskelsystem .... 262\nI Der M. scalenus .... 262\nII Die Bauchmuskeln .... 263\nA Anatomisches .... 263\n1 Allgemeine Lage- und Ursprungsverhaltnisse .... 263\n2 Die Rectusscheide .... 265