ALBERT

Description: The northeastern boreal forest of Quebec is characterized by a humid climate. Consequently, fires are less frequent and small-scale disturbances play an important role in forest dynamics. Natural mortality and nonfire disturbances such as insect outbreaks and windthrow lead to gap-driven processes. Changes in structure and species composition can result from gap dynamics. The objectives of this study were to characterize gaps and examine patterns of species replacement in gaps in old conifer stands. Line intersect sampling was used to sample stands dominated by balsam fir (Abies balsamea (L.) Mill.) and (or) black spruce (Picea mariana (Mill.) BSP). Results show that 54% of the forest was in expanded gaps and that canopy gaps are relatively small, since 87% of them were smaller than 100 m2. The majority (94%) of the openings were caused by the mortality of less than 10 gap makers. Replacement probabilities show self-replacement of A. balsamea in Abies stands and of P. mariana in Picea stands. However, in Abies-Picea stands, there seems to be a reciprocal replacement of the two species. These results provide knowledge of the disturbance dynamics of the region as a basis for development of silvicultural practices that preserve the structural components of older forest stands.

Description: Spatial patterns, rates, and temporal variation of standing-tree mortality were studied in unmanaged boreal old-growth forests of northeastern Quebec. The study was carried out by sampling living and dead trees within 15 transects (400 m long, 40 m wide). The transects lay in stands that were classified according to their species composition in three types: dominated by black spruce, Picea mariana (Mill.) BSP; mixed P. mariana and balsam fir, Abies balsamea (L.) Mill.; and dominated by A. balsamea. Spatial patterns were analysed using Ripley's K function. The year of death was cross-dated using 190 sample discs extracted from dead standing A. balsamea and P. mariana to assess the rates and temporal variation of mortality. The spatial patterns of standing dead trees in P. mariana stands were predominantly clustered. The spatial patterns of large dead trees (〉19 cm diameter at breast height (1.3 m height; DBH)) in mixed and A. balsamea-dominated stands were mainly random, with few stands showing clustered patterns. Small dead trees (9–19 cm DBH) in these stands were generally more clustered than larger trees. Tree mortality varied from year to year, though some mortality was observed in all the studied stand types for almost every year. Standing trees that had recently died accounted for 62%, 48%, and 51% of overall mortality in P. mariana-dominated, mixed, and A. balsamea-dominated stands, respectively. The results of this study indicate that mortality of standing trees outside of episodic mortality events (such as insect outbreaks) is an important process in the creation of structural complexity and habitat diversity in these stands.