ALBERT

Description: The sedimentation regime off Northwest Africa is shaped by: (1) structur~al factors. which result in generallv low relief on land. shelf widths between 40 and more than 120 km. and av-erage sfope inclinations between 10 30' and 30; (2) land climates. which contral the delivery of terrigenous particles to the margin: (3) water movements including boundary currents and upwelling; and (4) the post- Pleistocene sea level rise. This chapter combines published and new results arising from research into the sedimentation processes off Northwest Africa. and emphasizes particularly the activities of the Kiel marine geological group during the past few years. Reviews of cruise activities and results were given in Closs et al. (1969) (Meteor cruise 8. 1967. off Morocco) . Seibold (1972) (Meteor cmise 25 . 1971. off Sahara to Central Senegal). Seibold and Hinz (1976) (Meteor cmise 39,1975 . and Valdivia cruise 10. 1975, from Morocco to South Senegal), and Waiden et al. (1974) (Meteor cmise 30, 1973, off Sierra Leone). Some of these cmises were used for pre- or post-site surveys for the Deep-Sea Drilling Project, or to add undisturbed Quaternary cores to the Glomar Challenger cores (leg 41, ] 975; Lancelot, et al .• 1978); leg 47 A, Arthur er al .• 1979; Lutze et al., 1979). We have concentrated our geological investigations on a number of standard profiles from the shelf to the upper continental rise as given in Figure 1. The manuscript was finished May 1979.

Description: https://www.researchgate.net/publication/230891291_The_Orbital_Theory_of_Pleistocene_Climate_Support_frim_a_Revised_Chronology_of_the_Marine_d18O_Record

Description: Summary Holocene sediments of the North Lagoon, Bermuda, were studied with shallow seismic reflection profiles (200 km CSP-survey, UNIBOOM-system) and vibration coring (40 sediment cores, pneumatic vibration corer, Meischner et al., 1981). Seismic Stratigraphy Four seismic sequences are distinguishable by seismic stratigraphy. All seismic sequences correspond to depositional sequences built up during high sea levels in interglacial times. The seismic sequences are separated by unconformities which are often strongly reflective and correspond to emersion planes during glacial phases. The upper sequence (sequence 4) is related to Holocene sediments. The pre-Holocene bedrock is divided into three different seismic sequences (Kuhn et al., 1981): Sequence 1: oldest Pleistocene sequence (pre-Sangamon sea-level highstands), upper boundary with levelled relief (lower boundary not discernible), composed of strongly cemented carbonate sediments, forms the bedrock below Three Hill Shoals Sequence 2: Sangamon (125 ky sea-level highstand), distinct surface morphology, forms the bedrock of a large area below Holocene sediments, Holocene reefs grew up on elevations of the sequence 2 surface, the Holocene reef rim was developed on an elevated rim of sequence 2 Sequence 3: youngest Pleistocene sequence (Sangamon, 105 and 85 ky sealevel highstands lower than recent), deposited mainly in depressions of the bedrock deeper than -15 m below recent Mean Sea Level, levelling the older relief, peat sedimentation in places The distribution of recent reef areas and lagoonal basins is strongly controlled by pre-Holocene topography and geology of the bedrock. During the Holocene approx. 1050 x 106 m3 of carbonate sediments were deposited in the North Lagoon (290 km2) and approx. 1350 x 106 m3 in the reef rim area (170 km2). Sedimentology There are no larger oscillations of the Holocene sea level identifiable in the sedimentological record. The pre-Holocene topography was gradually drowned during the Holocene sea-level rise. At first, the depositional depressions were separated and landlocked. Fresh water peat marshes, fresh water ponds, marine ponds and bays were formed. With rising sea level, the land barriers were more and more eroded, drowned and lost their influence on the back-barrier sedimentation area. Autochthonous and allochthonous peat, lime gyttja and carbonate mud are a typical transgressive back-barrier sediment sequence. After destruction of the barrier, the depositional milieu changed from restricted marine to normal marine, open lagoonal. Sea-grass sediments and nearly mud-free carbonate sand were deposited in shallow water in an exposed environment. Hydrodynamic energy decreases with increasing water depth in the lagoonal basin. A more densely growing reef rim and intralagoonal reef growth added to the protection of the deeper lagoonal floors. Fine-grained sediments were deposited in this environment. They are distributed over a large area of the North Lagoon and form the top of the transgressive lagoonal sediment sequence. Holocene reefs mainly developed on rises of the pre-Holocene surface. In the early Holocene, solid reef build-ups were able to keep up with the rapid rise of sea level. Sand pockets in the reefs were left behind and filled up mainly in the later Holocene. The percentage of fine-grained sediments, produced and resuspended in the reef rim and deposited in the near lagoonal back-reef zone, increased during the Holocene. Two models of Holocene sedimentation in a depression and on an elevation of the pre-Holocene surface illustrate the dependence of vertical facies gradation on pre-Holocene topography. Trends of the mostly polymodal grain-size distributions of the Holocene sediments are a coarsening-upward in the back-barrier and a fining-upward in the lagoonal sediment sequences. Change in the composition of the molluscan fauna in the Holocene sediments (particle size 〉 2000 µm) is an Indication for fades changes. Gastropods are abundant in the basal backbarrier sediments. Bivalves are rare and their diversity 1s low. Sea-grass sediments contain Codakia orbicularis and Astraea phoebia shells. In the sheltered lagoonal environment shell fragments 〉 2000 µm become rare, common species are Gouldia cerina, Pitar fulminata and Finella sp. (approx. 1000 µm). Fine-grained reef-rim derived sediments differ from lagoonal sediments by a higher percentage of Homotrema rubrum fragments and Alcyonaria spicules.

Description: Twelve species of terricolous microlichens from the Angmagssalik District, Southeast Greenland, are reported: Caloplaca friesii, C. livida, Lecanora boligera, Lecidea oligotropha and Leciophysma arctophila, which are new to the lichen flora of Greenland, Rinodina conradi, which is new to the eastcoast, and Baeomyces roseus, B. rufus, Buellia geophila, B. punctata, Caloplaca tornoensis and Mycoblastus tornoensis, new to Southeast Greenland. In a discussion of the greenlandic distribution, unpublished records from the herbarium of Copenhagen (C) are incorporated. Notes on the habitats are given and the pertinent phytosociological units indicated. Some morphological and anatomical characters are commented upon briefly.

Description: The 16 recognized species of Sorocea are listed with their synonyms and distribution. Two new taxa are described: S. steinbachii C.C. Berg and S. hirtella Mildbread ssp. oligotricha Akkermans & Berg. Three new combinations are made: S. muriculata Miquel ssp. uaupensis (Baillon) C.C. Berg, S. trophoides W. Burger ssp. rhodorachis (Cuatrecasas) C.C. Berg, and S. sprucei (Baillon) Macbride ssp. saxicola (Hassler) C.C. Berg. A key to the species is presented.

Description: The bogs of S. E. Groningen are part of the great peat-marshes extending from S. E. Drente as far as N.W. Germany inclusive. So far as the territory of Westerwolde is concerned, people have begun digging off very early. According to the map by Krayenhoff in 1816 nearly the whole peat-marsh westward from the line Blijham—Termaarsch had already been reclaimed, only a few parts still being covered with the original peat-layer (cf. map, fig. 1). The digging off east of the above line commences at the beginning of the 19th century on the borderland of Groningen and Drente. Borings were performed in three places and the samples pollenanalytically and stratigraphically examined.

Description: Neohattoria Kamim. is a monotypic genus of the Jubulaceae (= Frullaniaceae) with a single species, N. herzogii (Hatt.) Kamim., known from central to northern Japan and the southern part of the Kurile Islands. The present genus was segregated from Frullania by Kamimura (1961; sub. nom. Hattoria Kamim. nom. illeg., non Schust., 1961) on the basis of the branching type, the shape of the first leaf and underleaf on branch, the total lack of secondary pigmentation, the uniform cell structure of the stem in cross section, and the strongly toothed leaf lobes. The generic concept of Neohattoria was greatly expanded by Schuster (1970), who included eight species and classified them into two subgenera, subgen. Neohattoria (with a single species) and subgen. Microfrullania Schust. (with seven species); however, Hattori et al. (1972) transferred all species of subgen. Microfrullania to a newly segregated genus Schusterella Hatt. et al., thus retaining the monotypic status of Neohattoria. As already described and illustrated by Hattori (1955), Kamimura (1961), Mizutani (1961), Ladyzhenskaja (1963), Schuster (1970), and Hattori et al. (1972), Neohattoria herzogii is closely related to species of both Jubula and Frullania. Regarding the taxonomic desposition of Neohattoria, Mizutani (1961) and Mizutani & Hattori (1969) placed it with Jubula in a subfamily Jubuloideae of Lejeuneaceae and Hattori et al. placed it in Jubulaceae (s. lat.). But, Kamimura (1961), Schuster (1970, 1979), and Guercke (1978) placed it more close to Frullania, e.g. in a subfamily Frullanioideae of Jubulaceae (s. lat.); more recently, Asakawa et al. (1979b), admitting three distinct families, Jubulaceae, Frullaniaceae, and Lejeuneaceae, placed Neohattoria and Jubula in the Jubulaceae (s. str.) but Frullania and Schusterella in the Frullaniaceae.

Description: E sectione Peltaea, Pavoniae speciosae H.B.K. proxima, sed forma folorium, indumento, involucri phyllis peltatis diversa. Suffrutex, caule minute stellato-piloso glabrescente, linea singula pilis simplicibus longioribus vestita in primo internodio ramulorum lateralium adaxiale notato. Folia breviter petiolata, petiolis tomentellis 2—4 mm longis, oblongo-elliptica, elliptica vel ellipticolanceolata, 3—5 cm longa, 1.25—1.5 cm lata trinervia basi acuta vel obtusa, superiora 5-nervia, basi subcordata, acutissima vel subacuminata, margine regulariter serrato-dentata, supra minute stellato-pilosa, oculo nudo glabra, infra dense sed minute stellatotomentella. Flores in axillis foliorum vel in apice ramulorum 2—3-glomeratis, bracteis ovato-triangularibus suffulti, plerumque subsessiles, interdum usque ad 4 mm pedicellati. Involucri phylla fere io linearia birta uniserialia, basi paullo connata, apice lamina foliacea peltata, id est supra basin affixa, anguste elliptica hirta, basi rotundata, apice acuta, appendiculata, 4 mm longa. Calyx cupuliformis, ultra medium incisus, 4—9 mm longus, lobis acutis hirtis, nervis trinis conspicuis, binis intermediis brevibus vel nullis. Petala 2.5—3 cm longa, teste collectore roseo-rubra, sicca rosea, basi atropurpurea. Stamina et styli more generis. Carpella 4 mm longa, mutica, dorso costa perpendiculari instructa, transverse nervosa, dense pubescentia.

Description: Culmi robusti, foliati. Folia lata, linearia, trinervia. Inflorescentia corymboso-paniculata, multispiculata. Spiculae (”spicae” multorum auctorum) parvae, multiflorae. Flores hermaphroditi (”spiculae androgynae” auctorum) perianthio utriculiformi, compresso, vix carinato, staminibus (”floribus masculinis monandris” auctorum) tribus, binis lateralibus tertio anteriore, ovario (”flore foemineo terminali nudo” auctorum) rostrato, basi angustato, haud stipitato, styli ramis ternis. Nux tri-costata, rugulosa. Generi Hypolytro L. C. Rich. proxima, a quo differt styli ramis tribus et nuce tri-costata. A Thoracostachyo et Paramapania, quibuscum stigmatum numero convenit, et structura florum et perianthio connato et nucis forma longe diversa, faciliter dignoscenda. Mapaniae potius affinis, sed ab omnibus speciebus huius generis inflorescentia a plerisque etiam perianthio connato discrepat.

Description: The genus Trymatococcus has been published in 1838 by Poeppig and Endlicher in Nova Genera ac Spec. Plant II. p. 30, and the genus was based on the species T. amazonicus. In 1876 Baillon added the species T. africanus to the genus. This gave a peculiar distribution for a genus with two species only: one in the Amazone region and one in West Africa. Later on several new species from Africa were described: three by Engler (T. kamerunianus, dorstenioides, and Conrauanus), one by De Wildeman (T. Gilletii) and one by Pellegrin (T. oligogyna). In 1922 (Archivos do Jardim Botanico Rio de Janeiro vol III. p. 22) Ducke described a second species from Amazonian Brazil (T. paraensis) and said in the notes to this new species that Lanessania turbinata Baill. should be transferred to the genus Trymatococcus and published a new combination (T. turbinatus Ducke). In 1925 (Archives IV. p. I) he emphasized his statements Trymatococcus and published a new combination (T. turbinatus as well as turbinatus and amazonicus have the stamens erect in the bud and not inflexed as was described in the former publications. He also emphasized that the place of Trymatococcus in the system has to be changed and the genus has to take the place taken up to this moment by Lanessania. Among the material of the Moraceae from Surinam which I am studying for the Flora of Surinam, I found also a Trymatococcus species. By the study of this genus I was struck by the peculiar geographic distribution of the genus, which fully supported my observations on the Euphorbiaceae (cf. Lanjouw, The Euphorbiaceae of Surinam pp. 70—84). For the preparation of a map of this distribution I studied the african species and after a careful examination I noted a number of important differences between the african species and the american ones. Part of these differences were never noticed before and no attention has ever been given to these facts. The first error in this case was made by Baillon. Most probably he had not seen T. amazonicus Poepp. et Endl. when he described his T. africanus. This is still more striking as he described in the same paper his genus Lanessania based on L. turbinata, which is a true Trymatococcus species. It is very curious that it was not possible for Baillonto observe his mistake because in his Histoire des Plantes (vol. VI. p. 199) he states „filamentis aestivatione inflexis vel nunc suberectis”. One can not understand why he did not observe that at least one of the species of Trymatococcus is the same as his genus Lanessania. After Baillon’s publication, we could say that we had got two type species, one american (Tr. amazonicus Poepp. et Endl.) and one african (Tr. africanus Baill.). Apparently Engler did not study exactly Tr. amazonicus Poepp. et Endl. when he described his new species though he states (Monogr. Afr. Pfl. fam. I. Morac. p. 28); ”Ein besonders auffallender Unterschied im Bau der Blüte und Frucht is nicht zu constatieren; bei der amerikanischen Art sind die männlichen Blüten dreimännig mit dreiteiliger Blütenhülle, bei den afrikanischen Arten sind sie zweimännig”. Likewise Ducke knew apparently only the american species when he pointed out the new place for this genus in the family. By these reasons only it is explained how confusion has crept into this genus. I have studied many specimens of Trymatococcus from the following herbaria: Berlin-Dahlem, British Museum (Natural History Museum), Kew, Leiden, Paris and Utrecht. I wish to express mv sincere thanks to the directors for their hospitality or fore sending the material on loan.

Description: El género Plagiochila (hepatica) esta representada en las Islas Galapagos por ocho (8) especies diferentes: P. bursata (Desv.) Lindenbg., P. galapagona Inoue, P gradsteinii Inoue, P. guilleminiana Mont., P. inouei Grolle, P. scabrifolia Inoue, P. spinifera Ångstr. y P. subplana Lindenbg. El endemismo en este género es más alto que en otros géneros de las hepaticas, con cinco (5) especies que comienzan a conocerse solamente de los Galapagos ( P. galapagona, gradsteinii, scabrifolia, inouei, y spinifera). Las otras tres (3) son comunes y ampliamente distribuidas a lo largo de la America tropical. La mayoría de las especies estan restringidas a las zonas altas-húmedas de vegetación de las Islas Galapagos (matorrales de Zanthoxylum, Miconia y pampa) excepto P. guilleminiana muy común, la cual puede presentarse en la zona seca de transición de bosque. La más amplia variación de Plagiochila ha sido vista en Isabela (Cerro Azul), San Cristobal y Santa Cruz.

Description: The species Polypodium banaense C.Chr. is transferred to Crypsinus. The recognition of a genus Phymatopteris Pic. Ser. (= Phymatopsis J.Sm.) separate from Crypsinus is discussed.

Description: The six species of Curtia, including a hitherto undescribed species published here, as well as the monotypic genus Hockinia can be distinguished from each other by the seed coat structure. The anticlinal walls and the cuticle provide the most useful information. Curtia tenuifolia appears to be a complex species, but subsp. tenella can be readily separated from this complex by the seed coat structure. Heterostyly has been found in C. tenuifolia subsp. tenuifolia, C. obtusifolia, and Hockinia montana, but differences in seed coat structure can not be correlated with long-, short-, and equal-styled flowers. The differences in seed coat structure, the length of the seeds, and the number of cells per seed plead for maintaining Hockinia (closely related to Curtia) as a distinct genus. One new species of Curtia is described and a new combination is made.

Description: SETTEN, A. K. van & KOEK-NOORMAN, J.: Studies in Annonaceae. VI. A leafanatomical survey of genera of Annonaceae in the Neotropics. — Bot. Jahrb. Syst. 108: 17—50. 1986. — ISSN 0006-8152. Within the scope of the multidisciplinary research project on systematics of Annonaceae, a survey of the leafanatomical features and their distribution in the neotropical Annonaceae is presented. The studied specimens form a rather homogeneous group, as may appear from the family description given here. A detailed study of the leafanatomical features reveals, that differences are mainly found in the indument, the position and contents of the idioblasts, the structure of the primary vein, the type of crystals in the epidermal cells, and the type of sclereids. Based on character states, phenetic similarities and differences are discussed and compared with the classifications of FRIES (1959) and WALKER (1971).

Description: One new species of Dorstenia from Brazil is described: D. carautae C.C. Berg, and four new combinations are made: D. cayapia Vellozo subsp. asaroides (Hooker) C.C. Berg, D. cayapia Vellozo subsp. paraguariensis (Hassler) C.C. Berg, D. cayapia Vellozo subsp. vitifolia (Gardner) C.C. Berg, and D. ramosa (Desvaux) Carauta, Valente & Sucre subsp. dolichocaula (Pilger) C.C. Berg. A list of and a key to the 22 Dorstenia species distinguished in south-eastern tropical America are presented, together with synonymy and distributional data.

Description: Recently a multidisciplinary investigation program on the systematics of Annonaceae was started at Utrecht with special emphasis on the Neotropics. This project will be carried out largely within the framework of the UNESCO-project Flora Neotronica. The first goal is to provide a modern classification of the family as a whole, the second is the publication of a series of monographs for Flora Neotropica. The project has been planned and started in close consultation with leading botanists on the Neotropical flora. The Annonaceae are a family of pantropical distribution with between 2000 and 2500 species in ca. 130 genera as presently understood. In the Neotropics the family is represented by ca. 750 species and 35 genera. It is a family of trees, shrubs, and lianas. Its place is within the order of the Magnoliales and its supposedly closest relative is the family of the Myristicaceae. The Annonaceae, although generally considered primitive in many features, nevertheless offer a number of specialized features as well This makes it a promising object using various kinds of comparative morphological, karyological, and anatomical data. Besides, many species are of medicinal or commercial value, such as various species of Annona and Rollinia, the fruits of which are commonly eaten in most countries of Central America and South America; the Soursop (Annona muricata) is widely cultivated throughout the tropics.

Description: Es handelt sich hier um ein grosses, zusammenhängendes Hochmoorgebiet, das sich nord-süd über 20 km, ost-west über 10 km ausdehnt. Im Süden und Westen ist es grösstenteils abgetorft. Die besonders im Zentrum und Osten noch erhaltenen Teile sind durch die intensive Trockenlegung meist verheidet; stellenweise, so in den „Engbertsdijkvenen”, wo grosse Flächen heute wenig entwässert sind, findet sich eine lebende Sphagnumdecke (Taf. III). (Lit. 5). Das Moor liegt auf pleistozänem Untergrunde (Fluvioglazial der Riss-Eiszeit und Niederterrasse der Würmeiszeit); im Osten und Westen stosst es an diluviale Rücken; im Nordwesten bildet die Niederterrasse der Vechte die Grenze. Im Südosten und Osten schliesst sich eine ausgedehnte Versumpfungszone an, während sich im Westen zwischen den Hügeln isolierte, ähnliche Bildungen vorfinden. Es handelt sich hier wahrscheinlich um ein Entwässerungsgebiet des Hochmoores. Ein prae-rissglazialer mit nördlichen Erratica bestreuter Rücken dringt vom Osten her, parallel dem Vechtetal, ungefähr bis in die Mitte, in das Moor vor. Für eine ausführliche Angabe der geologischen Verhältnisse verweisen wir auf die „Geologische Kaart van Nederland” vom „Rijks Geologische Dienst” (Blätter Almeloo I und II; Koevorden III und IV). Wir sammelten eine Anzahl Probenreihen. Die angeführten Analysen beziehen sich auf eine süd-nord gerichtete Profillinie im östlichen Teil des Gebietes (Paterswal 1 u. 2, Engbertsdijk, Bruine Haar) und ein Punktprofil im Nordwesten (Boerendijk), nahe dem Vechtetal.

Description: It is to be hoped, that the genus Pandanophyllum Hassk. never will revive, for it is based on a bad generic description and two nomina nuda, P. palustre Hassk. (Harassas tjaai) and P. humile Hassk., the first of which is supposed to indicate Mapania palustris (Steud.) Vill., while the other name has brought about much confusion, as it has been used for Hypolytrum humile (Steud.) Boeck. as well as for Mapania humilis (Miq., partly) Vill. The first validly published description of Pandanophyllum humile Hassk. nomen nudum in Cat. Pl. Hort. Bot. Bog. 1844, p. 297 has been given by Steudel in his Synopsis II (1855), p. 134 and is based upon a specimen collected in Java by Zollinger (n. 1511, Brit. Mus., Paris), belonging to the genus Hypolytrum. So this is the type-specimen of H. humile (Steud.) Boeck. in Linnaea XXXVII (1871—1873), p. 128. Bentham and Hooker, however, accepting the interpretation of Kurz in Journ. As. Soc. of Bengal XXXVIII, part 2 (1869), p. 82 and the revised opinion of Miquel in his Ill. Fl. Arch. Ind. (1871), p. 61, included both species in their section Pandanophyllum of Mapania (Gen. Pl. III, 1883, p. 1056). A quarter of a century later C. B. Clarke divided Benth. and Hooker’s section into two subgenera, viz. Pandanophyllum, including Mapania humilis Vill. and Halostemma (Wall.), including Mapania palustris (Steud.) Vill. Consequently our present section Pandanophyllum sensu Clarke probably excludes both species, which originally belonged to it. One might be inclined to rectify the mistake by changing the name of Halostemma into Pandanophyllum and coining a new name for the other subgenus, but the principal difficulty, caused by the ambiguity of Hasskarl’s generic description can not be solved in this manner. This description calls for a bifid style (perhaps referring to Hypolytrum humile Boeck.) and 3—5 spikelets (not appropriate to Mapania palustris Vill., highly improbable as to Mapania humilis Vill. and Hypolytrum humile Boeck.). The only way out of the difficulty is to reject the name Pandanophyllum as a nomen dubium in the sense of the rules of nomenclature (art. 63) and to rename the subgenus Pandanophyllum Benth. et Hook., sensu Clarke. I propose the name Pandanoscirpus.

Description: A new species of Asterophorum, A. mennegae, is described from the Sipaliwini Savanne (Suriname). The position of Asterophorum within the family is briefly reviewed, and a key to the 2 species is given.

Description: The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.

Description: As Prof. Bremekamp has dealt with the genus Pleiocraterium from the taxonomic point of view, I intend to supplement his exposition here with some observations on the ecology of these remarkable additions to the Malaysian mountain flora. Some of these observations have been included already in a general report on the results of the Losir expedition published in Dutch. As a further illustration I am giving two photographs taken from one of the two Sumatran species in its natural habitat. Altitude. Both species were found on the highest parts of the mountains only, viz. Pl. gentianifolium just below the summit of Mt Goh Lembuh, and Pl. sumatranum between our camp at the base of the central Peak of Mt Losir at c. 3250 m. and the summit of the latter at 3460 m. These two mountains lie rather far apart: Mt Losir is the highest top of the Barisan Range proper, whereas Mt Goh Lembuh is a more isolated mountain, rising c. 50 km. NNE of Mt Losir and separated from the latter by a wide depression. The two mountains also differ geologically.

Description: Nanocyperion communities (s.l.) are considered here as “warp-and-woof” communities; the Nanocyperion components are described separately as synusiae. On the Netherlands Frisian Islands, four main synusiae have been recognized. Raunkiaer life form spectra show few differences between the communities. Life strategy spectra of the Nanocyperion synusiae, based on systems for phanerogams (modified after Bakker 1966) and bryophytes, yield the clearest patterns. A comparison of the ecology of the communities and an interpretation of the spectra in terms of avoidance of stress or competition suggest that inundations and standing crop of the communities are the main factors determining the distribution of the synusiae. Winter inundations overrule the influence of differences in productivity level, which becomes prominent in drier situations.

Description: La forme est le phénomène de la vie le plus important. Aussi on pourrait croire que toute étude biologique devait commencer par la forme. En effet aucune fonction n’est imaginable indépendante de la forme, tandis qu’on peut étudier la forme indépendemment de la fonction, par exemple à des objets morts. Cependant depuis Sachs le botaniste moderne est tellement possédé par les conceptions matérialistes et mécaniques, qu’il veut aussi expliquer causalement les formes organiques en oubliant que, même si toutes les formes sont matérielles, cela ne veut pas nécessairement dire que les lois physiques et chimiques qui dominent la matière sont capables d’expliquer la forme, c.à.d. l’organisation des êtres vivants. A l’aide de briques on peut bâtir des bâtiments les plus divers, mais on peut aussi bien construire ces mêmes bâtiments de bois ou de pierre naturelle: le matériel employé n’explique pas le projet de l’architecte. Ce n’est qu’en le contemplant et en le comparant à d’autres qu’on arrive à mieux le comprendre (von Veh, p. 139). La forme („type” ou „idée” dans la conception platonique) est indépendante de la matière. Elle est ce qui reste. C’est par la forme que passe le courant de la cause et de l’effet, comme l’eau passe par un endroit clair d’une rivière (Carus). La forme présente un des problèmes les plus difficiles de la biologie. Le physiologue et le morphologue (deux extrêmes psychologiques) commencent pour ainsi dire aux deux extrémités de la nature, chacun à sa manière (Troll, Meyer), l’un avec sa méthode physique et chimique, l’autre avec sa méthode comparative. Au domaine du premier appartient tout ce qui est dynamique: le métabolisme et la croissance, au domaine du second ce qui est statique: la forme. Que la feuille est la partie principale de la plante, sur cela les physiologues et les morphologues sont d’accord. Le premier la considère comme un organe qui a pour fonctions principales la CO2-assimilation et l’évaporation. Depuis Goethe le second considère tous les appendices de la tige, aussi bien les sépales que les pétales ainsi que les organes sexuels comme des feuilles métamorphosées. Même, sous l’impression de la phyllotaxie des frères Bravais, Nees d’Esenbeck croyait que „la plante n’est rien d’autre qu’une unité de feuilles reliées entre-elles par un ordre défini”. C’est pourquoi on peut aisément considérer la morphologie de la feuille comme le problème central de toute la morphologie. Il est intéressant de se rendre compte comment dans le courant des temps on a essayé d’approcher ce problème de divers côtés. Cela pourrait apporter quelque lumière sur les différentes tendances de l’étude scientifique et sur les manières de penser qui sont caractéristiques pour les différentes périodes.

Description: Notwithstanding the large amount of work spent by several botanists on this family, taxonomy does not appear very satisfactory, and a general agreement on generic limits has not yet been reached. The result has been a perplexing number of generic and sectional names. The present author apologizes for his adding to the number of interpretations. This study of American Sapotaceae, primarily undertaken in connection with the Flora of Surinam, could not have been completed without the generous loan of specimens by the herbaria at Brussels [B], Berlin—Dahlem [D], Kew [K], and Leyden [L]. In 1934 the author paid a short visit to the herbaria at Brussels [B] and at Paris [P]. The collections of this family at Paris are of special interest owing to the fact that they contain the material studied by Baillon, Pierre and Dubard, and bear numerous notes and analytical drawings, especially by Pierre, attached to the sheets. A number of British Guiana Sapotaceae from the Kew Herbarium was received for determination shortly afterwards. The author feels greatly indebted to the directors of the above mentioned Herbaria for their kind help, and particularly to Prof. Dr. A. Pulle, Utrecht, under whose direction this study was undertaken.

Description: The genus Pausandra Radlk. belongs to the Tribe Cluytieae of the Euphorbiaceae. It was described by Radlkofer in 1870 in Flora LIII pp. 79—95. The genus is based on Thouinia Morisiana of Casaretto. In his paper Radlkofer discussed at length that this species does not belong to the Sapindaceous genus Thouinia, but represents a new genus of the Euphorbiaceae. As at that time female flowers were unknown Radlkofer stated that the systematic position of the new genus was still doubtful, but that most probably it should belong to a new subtribe of the Jatropheae. Two new species were described in the genus in 1873 by Baillon, P. Trianae Baill. based on Pogonophora Trianae Müll. Arg. which was published in 1864, and P. Martinii Baill. based on very young material and erroneously described by Baillon as being 3-merous, as will be discussed below. He placed the genus in the affinity of Argithamnia Sw., which is certainly not right as this genus is quite different both in habit and in flowercharacters. A fourth species was added by Müller Arg. in 1874 in Flora Brasiliensis XI. II., where he inserted the genus in the same group as was suggested by Radlkofer. No more species had been described when Pax published in 1911 his monograph of the Tribe Cluytieae Pax in Engler, Das Pflanzenreich IV. 147. III. He inserted the genus Pausandra Radlk, with the genera Givotia Griff, and Ricinodendron Müll. Arg. in a new subtribe Ricinodendrinae Pax. I think that this is the right position for the genus, though it could be placed in a separate subtribe for its penninerved, glanduliferous leaves and the capsular fruits. It was a pity that Pax published this monograph without studying the original material. He now copied Baillon’s bad descriptions and the lack of a thorough study on the genus caused the publication of several superfluous species in recent years. P. quadriglandulosa Pax et K. Hoffm. and P. extorris Standley described in 1919 and 1929 are the same as P. Trianae (Müll. Arg.) Baill. P. flagellorhachis Lanj. is identic with P. Martinii Baill., while it was proved that the latter species is not trimerous. P. integrifolia Lanj. could not be maintained in the genus. Only the two new species published by Ducke in 1925 were truly new ones. Moreover three new species were recognized in the recent collections made by Krukoff in Brazil. It is for all these reasons that it seemed to me highly desirable to give a new treatment of this genus. Perhaps several of the old and new species can be united, as one can find often only small differences, but for the present I think it advisable to keep them separate. Pausandra Radlk, has been described to be dioecious, but recently it has been proved in some species that they are monoecious, so it is probable that most of them are under special cicumstances.

Description: Among the most remarkable finds made by Dr. van Steenis in the higher parts of the mountains of North Sumatra are a number of cushion plants. Two of these he recognized as Rubiaceae nearly related to Hedyotis verticillaris W. et A., a species occurring in similar habitats in the Nilgiri Hills, India, and in Ceylon. Hesitating, however, to express a definite opinion on their taxonomic position, he sent the material to me for further investigation. As I had occupied myself already for some time with the genus Hedyotis L. and its allies, this investigation offered me a Wellcome opportunity to test some of the principles which I had laid down for the subdivision of this group. Apart from the characters of the fruit I lay stress on the position of the inflorescence and on the form of the stipules. The name Hedyotis itself I wish to restrict to H. fruticosa L. and its nearest allies, i.e. to those species that are provided with terminal inflorescences, an ovary not distinctly produced beyond the insertion of the calyx, and fairly large drupes with apically and ventrally dehiscent pyrenes: to a group, therefore, which roughly agrees with Hedyotis section Diplophragma W. et A.

Description: Recent study of the copious material of Melastomaceae conserved in the Botanisch Museum en Herbarium at Utrecht has shown the existence of several undescribed species in Surinam and has given new ideas on the taxonomic status of a few other species. These results are presented below, in advance of the treatment of the family in the „Flora of Surinam”. Ernestia Pullei Gleason, sp. nov. Suffruticosa 4 dm. alta. Caulis purpureo-brunneus 4-angulatus dense glanduloso-pubescens, internodiis 10—15 mm. longis. Petioli graciles 5—10 mm. longi glanduloso-villosi. Laminae tenues ovatae usque ad 25 mm. longae 17 mm. latae acutae minutissime serrulatae basi cordulatae 5-nerviae, supra sparse minuteque glanduloso-pilosae, subtus dense cinereo-tomentellae. Paniculae magnae terminales ramosae 8—12 cm. longae multiflorae glanduloso-polisae, bracteis minimis oblongis. Florum 4- merorum non bene conservatorum structura difficiliter et fortasse non rite observanda. Hypanthium tubuloso-campanulatum 8-costatum dense glanduloso-pilosum. Sepala erecta triangularia acuta sparse glandulosa 1.6 mm. longa. Petala non visa. Stamina valde dimorpha. Filamenta glabra erecta gracilia 3-7 mm. longa. Antherae lineari-subulatae, staminum episepalorum horizontales 4.2 mm. longae, connectivo subtereti in semicirculum 1.5 mm. diam. curvato et supra insertionem filamenti in appendices 2 V-forme connatas dilatato, ad angulam externam appendicum inserto; appendicibus in angulo interno ad filamentum affixis, triangulari-subulatis 3.2 mm. longis, infra filamentum attenuatis in calcaria filiformia et interdum calcaribus similibus lateralibus 1 vel 2 ornatis; antherae staminum epipetalorum erectae 3.3 mm. longae, connectivo ad angulam 90° deflexo 1 mm. longo, infra insertionem filamenti calcaria 2 lineari-subulata erecta 1.7 mm. longa gerente. Ovarium superum, teste cl. Pulle in schedis 3-loculare, sed in uno dissecto distinctissime 4-loculare; stylo stigmateque non visis; seminibus cochleatis.

Description: Some months ago the first author published in his Studies in Moraceae II (Rec. trav. bot. néerl. XXXIII, 1936, pp. 254—276) a synopsis of the genus Clarisia R. & P. The second author traced in the Berlin Herbarium a specimen of this genus which had been described in 1821 as Excoecaria ilicifolia Spreng. As this species is identic with Clarisia strepitans (Fr. Allem.) Lanj., the name of the latter species has to be changed. As in addition some interesting specimens were kindly sent to Utrecht for determination by the Herbaria at Berlin-Dahlem (D), Geneva (G) and the Arnold Arboretum, Jamaica Plain (A), it seemed desirable to publish these notes.

Description: The Colombian representatives of the lichen family Parmeliaceae with linear lobes and marginal cilia have been revised. A key is given and morphology, chemistry and distribution are treated of 12 species in three genera: Cetrariastrum Sipm. gen. nov, with C. andense (Kärnef.) Sipm. comb. nov., C. dubitans Sipm. spec. nov. and C. equadoriense (Sant.) Sipm. comb. nov., Everniastrum with E. catawbiense (Degel.) Hale, E. cirrhatum (Fr.) Hale, E. columbiense (Zahlbr.) Hale, E. fragile Sipm. spec. nov., E. planum Sipm. spec. nov., E. sorocheilum (Vain.) Hale and E. vexans (Zahlbr.) Hale, and Parmelina cleefii Sipm. spec. nov. and P. swinscowii (Hale) Hale.

Description: A phytosociological survey based on methods of the Zürich-Montpellier School was carried out in the páramo vegetation of the Cordillera Oriental, Colombia. The study area covers about 10,000 and comprises the páramo between the Nevado de Sumapaz (3°55'N, 4250 m), the Sierra Nevada del Cocuy (6°25'N, 5493 m) and the Páramo del Almorzadero (7°N, 4375 m). The páramo vegetation was studied along various altitudinal transects from the upper forest line (3000-3500 m) up to the lower limit of the snowcap (4800 m). A general description of the study area includes data on geology, geomorphology, soils, climate, flora, phytogeography, morphological characters of the vegetation, fauna and landuse. The evolution and Quaternary history of páramo vegetation and climate is reviewed, incorporating the first data from the Lateglacial and Holocene of the Páramo de Sumapaz. The general altitudinal zonation of the páramo vegetation was studied and is presented for both the dry and the humid side of the Cordillera. The zonal and azonal plant communities are described including their physiognomy, composition and syntaxonomy, habitat and distribution. Eighty five syntaxa from the rank of variant to that of the class are newly described, 17 of which are provisional. The vegetation is not ranked syntaxonomically yet, but described on the basis of preliminary tables. A number of azonal communities, part of them of lesser extent, are described in a similar way. The páramo vegetation is primarily determined by the tropical diurnal high mountain climate. The diversity of the páramo vegetation is related to temperature (altitudinal gradient) and to humidity (dry and wet climate). The presence of zonal bunchgrass páramo, bamboo-bunchgrass páramo or bamboo páramo mainly depends on the complex interrelation between these factors. Finally a synthesis is provided on ecology, morphology and phytogeography of the páramo vegetation of the study area.

Description: Endlicheria Nees (non Presl) in Linnaea 8 (1833), p. 37; id., Progr. (1833), p. 16; id., Syst. (1836), p. 365; Endl., Gen. (1837), p. 321; id., Ench. (1841), p. 197; Dietrich, Synops. Pl. 2 (1840), p. 1332, 1350; Spach, Hist. nat. Végét. X (1841), p. 473; Steudel, Nomencl. ed. 2 (1841), p. 554; Meissn., Gen. I (1836—43), p. 326, II, p. 238; Orbigny, Dict. univ. (1846), p. 259; Lindl., Veg. kgd. (1846), p. 537; Meissn. in D.C., Prodr. XV, 1 (1864), p. 172; id. in Fl. Bras. V, 2 (1866), p. 281; Baillon, Hist. II (1870), p. 480 in adnot.; Pfeiffer, Nomencl. (1873), p. 1201; Benth. in Benth. & Hook., Gen. III (1880), p. 153; Durand, Index Gen. (1888), p. 348 sub Aydendron; Mez in Jahrb. Bot. Gart. Berl. V (1889), p. 111; Pax in Engl.-Prantl, Pfl. Fam. III, 2 (1889), p. 122; dalla Torre & Harms, Gen. (1900—07), p. 178 sub Aniba; Post & Kuntze, Lexicon (1904), p. 197; Lemée, Dict. 2 (1929), p. 857; Benoist in Arch. Bot. V (1931), p. 63; Kostermans in Meded. Bot. Mus. Utrecht 25 (1936), p. 41; id. in Pulle, F1. Surin. 2 (1936), p. 327. – Goeppertia Nees, Syst, l.c., p. 354, 365 (non alibi nec aliis); Endl., Gen., l.c., p. 321, n. 2051; id., Ench., l.c., p. 197; Dietrich, l.c., p. 1332, 1350; Spach., l.c., p. 473; Steudel, l.c., p. 697; Reichb., Nomencl. (1861), p. 70, n. 2659; Meissn., Gen. I, p. 326, II, p. 238; Orbigny, l.c., p. 259; Lindl., l.c., p. 537; Meissn. in D.C., l.c., p. 172; id. in Fl. Bras., l.c., p. 281; Baillon, l.c., p. 480; Pfeiffer, l.c., p. 1473; Benth., l.c., p. 153; Durand, l.c., p. 348 sub Aydendron; Mez, l.c.; Pax, l.c., p. 122; dalla Torre & Harms, l.c., p. 178 sub Aniba; Post & Kuntze, l.c., p. 253; Kosterm. in Meded., l.c. – Schauera Nees in Lindley, Nat. Syst. ed. 2 (1836), p. 202 in adnot. (non aliis nec alibi); Endl., l.c., p. 321; id., Ench., p. 197; Meissn., Gen. II, l.c., p. 238; Orbigny, l.c., p. 259; Lindl., Veg. kgd., l.c., p. 537; Benth., l.c., p. 153; Durand, l.c., p. 348 sub Aydendron; Mez, l.c.; Pfeiffer, l.c., p. 1071; dalla Torre & Harms, l.c., p. 178 sub Aniba; Post & Kuntze, l.c., p. 503; Lemée, l.c., p. 1006. – Schaueria Nees ex Meissn. in D.C., l.c., p. 172; id. in Fl. Bras., l.c., p. 281 (non aliis); Baillon, l.c., p. 480; Pax, l.c., p. 122. – Ampelodaphne Meissn. in D.C., l.c., p. 81; id. in Fl. Bras, l.c., p. 167; Baillon, l.c., p. 473; Pfeiffer, l.c., p. 1071; Benth., l.c., p. 153; Durand, l.c., p. 348 sub Aydendron; Pax, l.c., p. 122; dalla Torre & Harms, l.c., p. 178 n. 2812; Post & Kuntze, l.c., p. 24; Lemée, Dict., l.c., p. 210; Kosterm. in Meded., l.c. – Aydendron Griseb. (non Nees), p.p. in Fl. Brit. W. Ind. isl. (1860), p. 284; Benth., l.c., p. 153; Mez, l.c. – Huberodaphne Ducke in Arch. Jard. Rio de Janeiro 4 (1925), p. 191; Lemèe, Dict., l.c., 3 (1931), p. 661. Type species: Endlicheria hirsuta Nees.

Description: Nineteen species of Stereocaulon are treated from the northern Andes, mainly from Colombia. Descriptions and keys are given, with notes on the north-Andean distribution and ecology. Seven species are new for the Colombian flora, viz. St. atlanticum, St. claviceps, St. corticatulum (chem. strain with atranorin and perlatolic acid), St. delisei, St. microcarpum, St. pachycephalum and St. pomiferum. St. crambidiocephalum is reported for the first time from Costa Rica, as is St. didymicum from Venezuela, and St. delisei is reported for the first time from the New World (Colombia and Costa Rica). St. cornutum Müll. Arg. is reduced to synonymy under St. pityrizans Nyl.

Description: The wood and leaf anatomy of representatives of the 9 genera of the Opiliaceae are described in detail. It is possible to separate the genera on the base of both wood- and leaf anatomical characters. Herein the presence of cystoliths of varying shape and size is important. Some comments on the taxonomy and possible phylogeny of the familiy are given.

Description: Isoëtes Palmeri with a distribution in the High Andes from the Páramo of Venezuela to the Páramo of Ecuador is described as a new taxon, and dedicated to the then American specialist of the genus, Thomas Chalkley Palmer (1860-1934). The new species belongs to the tropical-Andeanaustral-antarctic section Laeves, described as new here as well. The publication of the new species had to be anticipated to the projected monographic treatment of the South-American representatives of the genus Isoëtes, as A.M. Cleef, Utrecht intends to base a new association, the Isoëtetum Palmeri on this new taxon, observed and collected by him at many instances within the Colombian Páramo between 1971 and 1980 in the context of the preparation of his doctoral thesis now under way.

Description: Anaueria Kosterm. in Chronica Botanica IV, 1 (1938), p. 14. Arbores brasilienses foliis sub-oppositis. Flores hermaphroditi ex-involucrati paniculati; tepalis sex tribus exterioribus minoribus. Stamina novem quorum sex exteriora fertilia filamentis in annulum ovarium cingentem connatis antheris liberis bilocellatis sub-introrsis; tria interiora sterilia staminodialia sub-aequilonga. Ovarium subglobosum tubo planiusculo insertum, stylo obtuso brevi stigmate inconspicuo. Staminodia seriei quartae nulla. Bacca magna ellipsoidea pedicello vix elongate cylindrico tepalis non incrassatis persistentibus insidens.

Description: Gradstein et al. (1982) propose to conserve four generic names of Lejeuneaceae: Lopholejeunea (Spruce) Schiffn., Acrolejeunea (Spruce) Schiffn., Trachylejeunea (Spruce) Schiffn. and Taxilejeunea (Spruce) Schiffn., each of which was introduced as a subgeneric name in Lejeunea by Spruce (1884), and subsequently raised to generic rank by Schiffner in his treatment of the Hepaticae in Engler-Prantl (preprint 1893) [see proposals to conserve 675-678 see p. 746]. Although Spruce (l.c.) used for his Lejeunea species a binary nomenclature by combining subgeneric names with specific epithets, it is clear (e.g. text, index) that the binomina are meant as Lejeunea combinations and they are considered as such by most authors (see Gradstein et al. for further details). Before 1893, however, the Sprucean subgeneric names were used in various papers by F. Stephani in a “seeming” generic rank; indeed Stephani now and then referred to them as “genus.” A chronological survey of a number of relevant papers by Stephani, mainly those published in Hedwigia, was given by Bonner et al. (1961), in conjunction with a brief discussion of the subject of this paper. These authors were the first to realize that on the basis of Art. 42 ICBN some generic names in Lejeuneaceae, e.g. Taxilejeunea and Trachylejeunea, can be considered as validly published by Stephani in Hedwigia 28, 1889. Later on Grolle (1979) demonstrated valid publication of monotypic new Lejeuneaceae genera by Stephani in the Bot. Gaz. 15, 1890, e.g. Lopho-Lejeunea and Acro-Lejeunea. For an evaluation of the status of Lopho- Lejeunea Steph., Acro-Lejeunea Steph., Trachylejeunea Steph. and Taxilejeunea Steph., one might consider these names against the background of the entire context of Stephani’s work on Lejeuneaceae until 1893. As the survey of Stephani’s papers in Bonner et al. is rather incomplete, and as there are several points of divergence in opinion, a new analysis of Stephani’s relevant papers (before Sep 1893) is presented below.

Description: Zu meiner Bearbeitung des surinamischen Materials der Gentianaceae für die von Pulle herausgegebene „Flora of Surinam” gehören nog einige kritische Bemerkungen. Ich muszte z.B. in einigen Fällen von der von Gilg in Engler und Prantl, Nat. Pflanzenfamilien gegebenen Einteilung der Gattungen und deren Umgrenzung abweichen. Auch stellte es sich heraus, dasz sich unter dem Material eine neue Art befand, deren Beschreibung und Abbildung unten folgen.

Description: Juniperus macropoda Boiss. Fl. Orient. V (1884) p. 709; Hooker Fl. Br. Ind. V (1890) p. 647. Umlung (Thalam-buti valley) 4200 m, 28 July no. 58. Big shrubs.

Description: In his introductory statements to 'The Symposium on the Phylogeny and Classification of the Filicopsida' which was held in London, 1972, HOLTTUM, when dealing with 'dubious groups of relationships which would particularly repay investigation', mentioned the Polypodiaceae first (HOLTTUM, 1973: 6). Talking about Polypodiaceae the present authors deal with the Polypodiaceae sensu stricto only, thus excluding the Cheiropleuriaceae, Dipteridaceae, Grammitidaceae, and also the Loxogrammaceae, taxa which were formerly (or are still) included in the Polypodiaceae sensu lato. As delineated in this way, this almost exclusively pantropical family consists of about 600 species and an indefinite number of genera.

Description: De in deze Jubileumserie van de „Mededeelingen van het Botanisch Museum en Herbarium te Utrecht” opgenomen artikelen zijn door de schrijvers ingezonden om Prof. Pulle, ter gelegenheid van zijn zilveren jubileum als hoogleeraar, hun waardeering te toonen. Een kort woord over den jubilaris moge hier als inleiding van deze bijdragen volgen. Op 10 Januari 1878, op den dag dat in verschillende plaatsen den Ioosten sterfdag van Linnaeus werd herdacht, werd August Adriaan Pulle te Arnhem geboren.

Description: Since the appearance of my „Notes on the Rubiaceae of Surinam” (in Rec. d. Trav. bot. néerl. XXXI, 1934, 248; also in Meded. Bot. Mus. Herb. Utrecht no. 11, 1934) a number of species and varieties new to the flora of that country have come to light. The majority have been collected by Mr. Rombouts during the 1935/36 expedition of the Boundary Commission who is surveying at present the border in the southern part of the colony; they were found along the River Corantyne and in the savannahs in the south-western part. One species was secured by Dr. Lanjouw, and has been mentioned already in his „Additions to Pulle’s Flora of Surinam I” (in Rec. d. Trav. bot. Néerl. XXXII, 1935, 258) and one, represented by a rather poor fruiting specimen collected years ago by the Forestry Bureau, was found among material provisionally consigned to another family. New to the flora of Surinam are the following twelve species: Alseis longifolia Ducke var. pentamera Brem. n. var., Sabicea cinerea Aubl., S. Romboutsii Brem. n. spec., S. surinamensis Brem. n. spec., Tocoyena surinamensis Brem. n. spec., Thieleodoxa nitidula Brem. n. spec., Guettarda Spruceana Müll. Arg., Psychotria Romboutsii Brem. n. spec., Declieuxia fruticosa (Willd. ex R. et S.) Kuntze, Diodia pulchristipula Brem. n. spec., Spermacoce guianensis Brem. n. spec, and Borreria verticillata (L.) G. F. W. Mey (the B. verticillata of the Flora of Surinam IV, 287 proved to be B. suaveolens G. F. W. Mey., under which name it had been recorded already by Miquel), and one variety: Sipanea pratensis Aubl. var. glaberrima Brem. n. var. Four of the ten genera to which these species belong, namely Alseis, Thieleodoxa, Declieuxia and Spermacoce, are also new to the flora of Surinam. Seven species and two varieties are entirely new, and will be described below. Before entering on this part of my task I will make a few remarks however on two of the species known already from elsewhere, namely on Guettarda Spruceana Müll. Arg. and on Borreria verticillata (L.) G. F. W. Mey, and on a third species, Coccocypselum guyanense (Aubl.) K. Sch., which is known since long from Surinam, but of which Mr. Rombouts collected a specimen differing somewhat from the older Surinam findings.

Description: The new species Coussapoa manuënsis C.C. Berg is described.

Description: Until recently relatively little attention has been paid to the study of chromosomes in liverworts. The first substantial contributions were made by Heitz (1927, 1928) and Lorbeer (1934). In the second half of this century chromosome studies on liverworts were mainly carried out in Europe (e.g. Fritsch 1972; Newton 1977, 1979) and Japan (e.g. Tatuno 1959; Segawa 1965a, b, c; Inoue 1968). Inoue (in Koponen 1979) reports that until now 28% of all bryophyte species in Japan have been investigated as to their chromosome complement. A comprehensive, but rather outdated, survey of chromosome numbers in Hepaticae and Anthocerotae was given by Berrie (1960). Work on a new, updated survey is now underway (Fritsch, in prep.). In the present article results are presented of a cytotaxonomic investigation of European species of the genera Aneura and Riccardia (Aneuraceae). Most specimens were gathered in the Netherlands, but some chromosome counts based on French and German plants are also included.

Description: During 1981 the Botanical Survey of India had again collections made. We list them in the same manner as on pages 3559-3560. In Andaman & Nicobar Is.: Great Nicobar, 300 specimens. In Andhra Pradesh: Anantagiri, Endrika Hills, Ganganaju-medugula, Paderu, 1590. In Arunachal Pradesh: Ganganagar, Hapoli, Naharlagan, Namdapha Biosphere Reserve of Tirap Distr., Tamer Road, Tiruli of Subansiri Distr., Ziro, 1054. In West Bengal: areas of Jalpaiguri, Bankura and Midnapur Districts, places of Bangaon, Tantulia and Basirhat of 24-Parganas Districts, Jaldapara Reserve, Totopara, &c., 2240. In Gujrat: Lalpur and vicinity, 1090. In Karnataka: vicinity of S. Karnataka River-Mulla Periyar and catchment areas, 500. In Kerala: Alleppey, Anathode, Cannanore, Devicolam, Kakki, Kasargod, Kokharjam, Munnar Peermade, Muzhiyar, Pachakanam, Pamba Dam areas, Peruvanzuzhi, Ponnambala Medu, Sabarigiri, 4150. In Madhya Pradesh; areas of Panna Distr., 800. In Maharashtra: Bhimsankar, Janar, Purandar, 985. In Meghalaya: Cherrapunjee, Nongapoh, Sunnapahar of Khasi Hills, Jowai, Jorain of Saintea Hills, Tura of Garo Hills Distr., 3500. In Nagaland: areas of Mekokchung, Tuensang, Wokha, Zunbebato Districts, 500. In Rajasthan: Jaisalmer and areas of Barmer Distr., 1000. In Sikkim: Burtuk Busty, Chakung, Changu, Chuten, Enchy Monastery, below Honuman Top, Jorethang, Lower Bustak, Ranipal, Reumtek, Sang Ratepani, Sinchey, Singtham East, Soren, Suntale forests, Tadong, 4800. In Tamil-Nadu: Kannayakumari, Sethur Hills, Srivilliputhur R.F., 2090. In Uttar Pradesh: Agra-Khal, Ballaieri, Chamoli Chakrata, Dudhwa Nat. Park, Govana, Khan-Khaliadha, Mussoorie, Pam Vali-Kantha, Panwali, Parbagi, Rajkhark, Saharshradhara, 2500.

Description: Because of their fleshy nature, thin leaves and membranous sepals and petals, Impatiens tend to make particularly poor herbarium specimens. If dried while still attached to the leafy part of the plant the flowers generally become badly crumpled and brittle. In such a state their more important characters become unrecognisable, and it is rarely possible to restore them to any useful degree. The leaves may also become badly crushed especially if they are not pressed absolutely flat. The collectors’ time may thus be completely wasted.

Description: IUCN, says the paper Categories, Objectives and Criteria for Protected Areas, ”is dedicated to the wise use of the Earth’s natural resources and to the maintenance of the Planet’s natural diversity.” What to think of the sequence? Use first, maintain second? And this comes from the International Union for the Conservation of Nature and Natural Resources? ”The World National Parks Congress, taking place in Bali, Indonesia, October 11-22, 1982, will provide case studies from around the world to illustrate how the various categories of protected areas are meeting the needs of countries of all economic, social, cultural, and political backgrounds,” writes J.A. McNeely, the secretary of the Commission on National Parks and Protected Areas, in a special issue of the Swedish journal Ambio (11: 237. 1982). ”No longer just playgrounds for vacationers and means for conserving natural heritage, protected areas have become an inseparable part of the modern human ecosystem.”

Description: Dr. M.M.J. van Balgooy and his companions on the Celebes Expedition, Dr. E. Hennipman, Mr. G.J. de Joncheere and Dr. E.F. de Vogel left Leiden on 5 April 1979, visited the SING and BO-Herbaria on the way. In Celebes visit was paid to Hasanudin University at Ujung Pandang (olim Makassar), in Bali to the Botanical Garden at Bedugul. In the course of August they returned to Holland. See also Exploration. The Botanical Survey of India kindly sent the following list of changes: D.K. Banerjee: to the Industrial Section of the Indian Museum at Calcutta; N. Bhargava: to the Northern Circle, Dehra Dun; U.C. Bhattacharyya: Deputy Director, Northern Circle, Dehra Dun; B.N. Chakraborty: Assistant curator, Industrial Section, Indian Museum, Calcutta; U. Chatterjee: Botanist, Eastern Circle, Shillong; Mrs. Dr. S.J. Das: Botanist, Eastern Circle, Shillong; P.K. Hajra: to HQ, Howrah; B. Krishna: to HQ, Howrah; Ram Lall: Botanist, Central Circle, Allahabad; C.L. Malhotra: to Northern Circle, Dehra Dun; P.C. Pant: to Northern Circle, Dehra Dun; B.B. Pramanick: Botanist, CAL-Herbarium, Howrah; M.K.V. Rao: to Andaman Circle, Port Blair; Dr. G.P. Roy: to Central Circle, Allahabad; B.D. Sharma: Deputy Director, Western Circle, Poona; Dr. R.C. Srivastava: Systematic Botanist, Eastern Circle, Shillong; C.R. Tarafder: Botanist, CAL-Herbarium, Howrah. Proficiat to all!

Description: The entries have been split into five categories: a) Algae — b) Fungi & Lichens — c) Bryophytes — d) Pteridophytes — e) Spermatophytes & General subjects. — Books have been marked with an asterisk. The SEM-observation of plant material normally requires dehydrated, dry specimens coated with carbon or metal. Unfortunately, the standard drying methods (including the critical-point-drying-technique) often cause shrinking and deformation of the specimen surface; therefore, SEMstudies on plant ontogeny are rather difficult, material- and time-consuming. Experiments using deep-frozen specimens have been carried out in England and in the USA, but have proved not satisfying.

Description: Mrs. Delia D. Adefuin, Museum Research Assistant, Manila, is pursuing her M.S. in Botany degree. She is currently the Secretary of the Fern Society of the Philippines. She is working on the Fern Flora of Metro Manila and is preparing the manuscript of a pictorial encyclopedia which will include descriptions of species and horticultural recommendations. Miss Barbro Axelius (S) collected and studied Xanthophytum and Lerchea (Rubiaceae) in Sarawak, Kalimantan and Sumatra, August 1982- February 1983.

Description: Tropical Botany in Aberdeen University. This was started by Professor J.W.H. Trail, who held the chair from 1877 to 1919, and travelled in the Amazon Valley (1873-75) mainly collecting cryptogams and studying palms. He was succeeded by Prof. W.G. Craib (1920-33) who was never in the tropics but devoted his work to the Flora of Siam, based on the collections of A.F.G. Kerr, and assisted by Miss E.C. Barnett. After a considerable lag, tropical botany was revived by the energetic efforts of Dr. P. Ashton as lecturer in systematics and ecology of the eastern tropics, establishing ties with Malayan colleges in teaching and research. This is at present perpetuated by two lecturers, Dr. K. Jong and Dr. M.D. Swaine, the latter’s experience lying largely in the tropics of West Africa. In addition Dr. N.M. Pritchard, Dr. J.B. Kenworthy and Dr. G. Hadley have been on secondment to the University of Malaya, while Dr. I. Alexander made research visits to India, Ghana and Peru. Over the years the Department has provided undergraduate and research training to innumerable students from many different tropical countries, some of which attained responsible posts, e.g. Prof. E. Soepadmo. Important courses in tropical biology are given, not available elsewhere in the U.K. (started 1973). The benefits for Aberdeen students is important: amongst others they led to expeditions to various parts of the tropics, recently to Sabah and to the Ivory Coast. Royal Society Tropical Rain Forest Collaborative Research Programme. Arising out of a feasibility study by Dr. T.C. Whitmore and P.F. Cockburn, the theme ’Recovery of tropical rain forest after disturbance’ was adopted as the initial basis of the programme. Possible territories for the research include Sabah and the Philippines. Detailed plans for a 5-year project are being prepared in consultation with colleagues in Southeast Asia.

Description: ANDERSON, J.A.R., A checklist of the trees of Sarawak, 364 pp. (1983, Dewan Bahasa dan Pustaka Cawangan Sarawak, for Forest Department, Kuching, Sarawak). Cloth Mal$ 15.00. When Dr. Anderson retired from the Forest Department in 1973 he left the manuscript of this checklist for publication. Unfortunately publication was delayed for 10 years. It contains data on over 2500 arboreous plant species. The text consists mainly of two parts: the first is a list of vernacular names with their scientific equivalents, the second is a list of plant names alphabetically arranged by family. Each species is concisely annotated with its vernacular name(s), maximum diameter, ecology, frequency, soils, etc. Species names have been coded: the first two figures are for the family, the next two for the genus and the last two for the species. A list is given of the trees of the peat-swamp forests of which Anderson was a great expert. A small draw-back is that the literature of the last ten years has not been included. Nevertheless this is a most helpful book. — C.G.G.J. van Steenis.

Description: Loss of species is the key issue of conservation. Contrary to misuse of land which is visible to anybody with eyes to see, the issue of extinction is sly, treacherous, and open to clear perception only for experts. It touches on quality, and reaches far out in time: hard things to grasp for non-biologists. Thus an extra responsibility devolves on those who are in a position to know and to speak. The value of the genetic resource base has been set forth in e.g. the book by O.H. Frankel & E. Bennett, Genetic resources in plants (1970), and in the BIOTROP symposium edited by J.T. Williams e.a., South East Asian plant genetic resources (1975); Myers adds many striking facts: half the prescriptions in the U.S.A. contain a drug of natural origin. The cardiac drug reserpine, from Rauvolfia, costs $ 1.25 per gram to synthesize, $ 0.75 from natural sources. The anti-polio vaccin was developed in experiments in chimpanzees. The Amerindians in Amazonia know 750 medicinal plant species. Now the possibility of massive destruction of tropical forests — where most species are located — casts some frightening shadows on the future. The question how to cope with the threat appears to be connected with human ethics and the international order. Consequently, most publications on the subject suffer from a partial lack of maturity: don’t look to Myers for ethics, nor to the Routleys for biology. It seems therefore advisable that on the part of all disciplines a common fund of knowledge and insight be built up. In my efforts, great stimulation was received from correspondence with Dr. Willem Meijer (Botany, University of Kentucky, Lexington, Ky. 40506, U.S.A.), who in his disinterested manner never fails to come up with things true and shocking.

Description: It is with the utmost regret that we announce the sudden and quite unexpected death of Dr. Marius Jacobs, editor of this Bulletin. See the obituary on page 3869. He was co-editor of the Flora Malesiana Bulletin for nr. 17 (1962) to nr. 22 (1968) and took full responsibility onwards of nr. 27 (1974). He showed great ability in enlarging its scope and we have many letters in our archives expressing appreciation and admiration for the lively and informative style in which he edited the Bulletin. I had to take over the editorial work for this number at short notice, but I was greatly helped by a number of Rijksherbarium colleagues, which help is gratefully acknowledged. In this way the delay has been kept to a minimum. It is, however, possible that some news items etc. have not been printed and that information submitted to Dr. Jacobs has not been entered due to this sudden change of editorship. I offer my apologies if this has happened and hope that (if still relevant) the news will again be forwarded to the new editor, Dr. J.F. Veldkamp, Rijksherbarium, who will take over starting next number.

Description: Apocynaceae wanted — pickled. Mary E. Fallen, Systematische Botanik, Zollikerstrasse 107, CH-8008 Zürich, Switzerland, who has done considerable morphological work on development of the reproductive organs in Apocynaceae, has been frustrated in her many efforts to obtain suitable material of Lepinia and Lepiniopsis. Ample information on both can be found in Pacific Plant Areas 3, Blumea Suppl. 5 (1966) 112-113, with map and description. The very oddly shaped fruit of Lepinia (W. Pacific) has been depicted in Blumea 11 (1962) 302, Van Steenis’s paper on the Land Bridge Theory. The one of Lepiniopsis (E. Malesia) seems to be buoyant. Also material of Anechites (Central America) is needed; it may be closely related to Condylocarpon. Any stages of flowers can be used, from tiny green buds at initiation up through anthesis, as well as fruiting stages. They should be pickled in FAA. Expenses of handling and postage will gladly be refunded. Vials with the liquid can be provided. Thanks on her behalf!

Description: Small evergreen trees, shrubs or lianas; two genera ( Cansjera and Opilia) are known to be root-parasites. Leaves distichous, simple, usually extremely variable in form and size, entire, exstipulate, pinnately veined; dried leaves mostly finely tubercled by cystoliths located in the mesophyll. Inflorescences axillary or cauliflorous, panicle-like, racemose, umbellate (in Africa) or spicate; bracts narrowly ovate or scale-like, in Opilia peltate, often early caducous. Flowers small, (3—) 4—5) (—6)-merous, mainly bisexual, sometimes unisexual and plants then dioecious ( Gjellerupia, Melientha, and Agonandra) or gynodioecious (Champereia). Perianth with valvate, free or sometimes partly united tepals (in ♀ flowers of Gjellerupia wanting). Stamens as many as and opposite to the tepals (in ♀ flowers only small staminodes); anthers introrse, 2-celled, longitudinally dehiscent. Disk intrastaminal, lobed (lobes alternating with the stamens), annular, or cupular. Ovary superior, 1-celled; style short or none, stigma entire or shallowly lobed. Ovule 1, pendulous from the apex of a central placenta, anatropous, unitegmic and tenuinucellar. Fruit drupaceous, pericarp rather thin, mesocarp ± fleshy-juicy, endocarp woody or crustaceous. Seed large, conform to the drupe, without testa; hilum basal, often in a funnel-shaped cavity. Embryo terete, embedded in rich, oily endosperm, nearly as long as the seed or shorter, with 3—4 linear cotyledons, radicle often very short. Distribution. There are 9 genera with about 30 spp., widespread in the tropics. Rhopalopilia is restricted to Africa and Madagascar, Agonandra to South and Central America. In Malesia: 7 genera, 5 of these only known from the eastern Old World (1 endemic: Gjellerupia in New Guinea); Opilia and Urobotrya occur also in tropical Africa.

Description: Monoecious, medium-sized to very large trees (rarely shrubby in very exposed situations). Either four independent cotyledons or two fused pairs (which may be retained in the seed after germination). The growing point of foliage shoots quite distinct between the two genera, being just a few highly reduced leaves in Araucaria and a highly organized bud formed of overlapping scales in Agathis. The leaves vary from scales or needles to broad leathery forms with many parallel veins sometimes on the same plant at different stages of growth. Pollen produced in cylindrical cones from one to as much as twenty cm long with numerous pedunculate spirally placed microsporophylls each with several to many pendent elongated pollen sacs attached to the lower side of an enlarged shieldlike apex which also projects apically more or less overlapping the adjacent microsporophylls. Pollen cones solitary, terminal or lateral, on branches separate from those bearing seed cones, subtended by a cluster of more or less modified leaves in the form of scales, deciduous when mature. Pollen globular, without ‘wings’. Seeds produced in large, well-formed cones which disintegrate when mature, dispensing the seeds in most cases with the help of wing-like structures; the seed cone terminal on a robust shoot or peduncle with more or less modified leaves that change in a brief transition zone at the base of the cone into cone bracts, formed of numerous spirally-placed bract complexes, usually maturing in the second year. Individual seed cone bract leathery or woody and fused with the fertile scale which bears one large inverted seed on its upper surface. Distribution. The 40 species in two genera are well represented in Malesia (13 spp.) and extend eastward and southward into Fiji, New Caledonia (18 spp.), Australia, and New Zealand, with 2 spp. also in the cooler parts of South America, giving the family a distinct Antarctic relationship. Only one species of Araucaria (in South America) occurs completely outside of the tropics, while the majority of the species in the family belong in the lowland tropics and others grow in the tropical highlands.

Description: The Botanical Survey of India continued to make collections during 1982, with the following results: Andaman & Nicobar Is.: Shola Bag, Mt Harriet, Jirkathang, Poona Nallah, Saddle peak, Diglipur, Rutland I. & Little I., 2875 specimens. Arunachal Pradesh: Various areas of Kameng Distr., Subansiri Distr., 9750 specimens. Assam: Garampani, 60 specimens. West Bengal: Darjeeling, Jalpaiguri, Hollong, Jaldapara, Chilapata, Salkumar, Daidaighat, Barasat, 2665 specimens. Bihar: Madhuban, Nimiaghat, Paresnath Hills, 315 specimens. Dehra Dun: Chakrata, Missoori, 325 specimens. Gujrat: Catchment and submergence areas of Sipu Reservoir Project, 1505 specimens. Kerala: Trichur, Idduki, Silent Valley, Valra Reserve Forest, Cannanore, Trivandrum, 3770 specimens. Madhya Pradesh: Kanha National Park, Chhodarpur Distr., 1190 specimens. Maharashtra: Areas of Jalgaon Distr. and Buldhana Distr., 4390 specimens. Manipur: Cherrapunjee, Mawphlong, Sorharim, areas of West Khasi Hills Districts, 2000 specimens. Rajasthan: Bharatpur, Desert National Park, 1605 specimens. Sikkim: Rangpo, Singtham, Bumbing, Manuring, Duga, Pandam, Takchi, Meli, areas of Gangtok, Chungtham, Lachi, Thanga, Panthang, 2590 specimens. Uttar Pradesh: Gori & Kali Valley, Chittoragarh Distr., 500 specimens.

Description: Many botanists must have wondered why as yet no volume of Flora Malesiana was dedicated to the outstanding botanist Carl Ludwig Blume, undisputed pioneer in planning the compilation of a ‘Flora Malesiana’. The writing of this Dedication would have been greatly facilitated if a full biography of BLUME had been existent, but none is available; there is not even a bibliography of his works. Only recently, in 1979, two biographical attempts were made, by J. MACLEAN and by A. DEN OUDEN, but only for the period 1820-1832; together with other biographical and obituary notes they are here assembled in Appendix B. I have also compiled a bibliography: Appendix A.²

Description: Erect or straggling herbs, shrubs or trees, sometimes monoecious or dioecious, the herbs sometimes rhizomatous; branches sometimes jointed at the nodes, sometimes without vessels ( Sarcandra). Leaves simple, decussate or sometimes whorled in fours, serrate, crenate or dentate, the teeth often thickened at the apex, penninerved, usually petiolate; petioles more or less connected at the base at least by a transverse line or connate into a distinct sheath; in Ascarina often alternating with leafless internodes which have the petiolar sheath; stipules minute to fairly conspicuous, subulate, borne on the petiole bases or sheath, occasionally pectinate. Flowers much reduced, without perianth, fully unisexual or essentially bisexual with the reduced anther-bearing organ adnate to the side of the ovary; arranged in spicate, paniculate, or capitate axillary or terminal inflorescences. — Male flowers bracteate or not, apparently consisting of 1—5 stamens, or in Hedyosmum consisting of numerous anthers in a cone-like structure; if 3 then the whole forming a fused 3-lobed organ sometimes enveloping the female flower by its edges, the central anther with 2 or aborted loculi and the laterals with single loculi, simply lobed or with connectives slightly to considerably produced so that the whole organ is 3-fingered; if with only 2 anther locelli then these on either side of a thickened filament plus connective. — Female flowers naked or enclosed by a cupular bract, the perianth adnate to the ovary, often minutely or shortly dentate at the apex and the ovary thus inferior; ovary 1-locular; stigma sessile or style short; truncate, 2-lipped, depressed or subcapitate (or horseshoe-shaped in one species), rarely linear or clavate. Ovule solitary, orthotropous, pendulous, bitegmic and crassinucellate. Drupes fleshy, small, ovoid or globose, sometimes more or less 3-sided in Hedyosmum, free or united into a mass by the bracts; endocarp hardened and crustaceous. Seeds subglobose, exarillate, with copious fleshy or oily endosperm and minute embryo, the cotyledons divaricate or scarcely formed. Distribution. Four genera with about 80 species. Since VESTER’S (1940) small-scale map the family (Ascarina) has been found in Madagascar. It is mainly tropical but Ascarina extends south to North Island of New Zealand (fig. 6) and Chloranthus and Sarcandra extend north to Japan, China, Korea and the eastern U.S.S.R. (Ussuri).

Description: Trees or shrubs (or rarely suffrutices outside Malesia). Leaves simple, alternate, often coriaceous, glabrous or with an indumentum on undersurface, margin entire; petioles often with 2 lateral glands. Stipules 2, minute and caducous to large and persistent, usually linear-lanceolate. Inflorescence racemose, paniculate or cymose; flowers bracteate and usually bibracteolate; bracts and bracteoles small and caducous or larger and enclosing flower or groups of flowers and persistent. Flowers actinomorphic to zygomorphic, hermaphrodite or rarely polygamous, markedly perigynous. Receptacle campanulate to cylindrical or rarely flattened cupuliforum, often gibbous at base; calyx lobes 5, imbricate, often unequal, erect or reflexed. Petals 5 (absent in some Neotropical species), inserted on margin of disk, commonly unequal, imbricate, deciduous, rarely clawed. Stamens indefinite, 2—60 (to 300 in Neotropics), inserted on margin of the disk, in a complete circle or unilateral, all fertile or some without anthers and often reduced to small tooth-like staminodes; filaments filiform, free or ligulately connate, short and included to long and far exserted; anthers small, 2-locular, longitudinally dehiscent, glabrous or rarely pubescent. Ovary basically of three carpels but usually with only one developed, the other two aborted or vestigial, variously attached to (the base, middle or mouth of) receptacle, usually sessile or with short gynophore, pubescent or villous; ovary unilocular with two ovules or bilocular with one ovule in each locule. Ovules erect, with micropyle at base (epitropous). Style filiform, basally attached; stigma 3-lobed or truncate. Fruit a fleshy or dry drupe of varied size, interior often densely hairy; endocarp much varied, thick or thin, fibrous or bony, often with a special mechanism for seedling escape. Seed erect, exalbuminous, the testa membraneous; cotyledons amygdaloid, plano-convex, fleshy, sometimes ruminate. Germination hypogeal with the first leaves opposite or alternate or epigeal with opposite first leaves. An extensive review of the generic limits of the family has been published: G.T. PRANCE & F. WHITE, The genera of Chrysobalanaceae: a study in practical and theoretical taxonomy and its relevance to evolutionary biology, Phil. Trans. Roy. Soc. London 320 (1988) 1—184. This contains full details of taxonomic history, morphology, anatomy, pollen, ecology and distribution of the family. A condensed version of these subjects is given here. Details of the Neotropical members of the family are given in: G.T. PRANCE, Chrysobalanaceae, Flora Neotropica 9 (1972) 1—410. The African members of the family were treated in: F. WHITE, The taxonomy, ecology and chorology of African Chrysobalanaceae (excluding Acioa), Bull. Jard. Bot. Nat. Belg. 46 (1976) 265—350.

Description: Perennial herbs, more commonly woody at the base, undershrubs or shrubs, erect, scrambling or scandent, sometimes high lianas. Rhizome not rarely tuberous. Branches often slightly swollen and jointed at nodes. Hairs simple, uni- or multicellular, short ones often with a hooked apex. Leaves simple, spiral or alternate, petioled (without an abscission zone), exstipulate; midrib usually prominent beneath, elevated or flat above; nervation commonly palmate, or pinnate, nerves often obliquely extending towards the margin. Flowers bisexual, actinomorphic or zygomorphic, solitary, fasciculate, or in axillary or cauligerous, racemose, paniculate or cymose inflorescences, usually only one or two flowers open at a time; bracts present and often persistent; pedicel often hardly distinct from the ovary. Calyx petaloid, gamosepalous, 3- (or 6-) lobed or 1-lipped; lobes valvate or induplicate. Petals (in Mal.) absent. Disk (?) 0, rarely present (e.g. a few Thottea spp.). Stamens 6 (4 or 5 in some extra-Mal. Aristolochia spp.) or 6—c. 36 (—46), in 1 whorl or in 2 (3 or 4) whorls (Thottea); filaments free or slightly mutually united at the base, and/or almost completely adnate to the style column to form a gynostemium; anthers free (Thottea) or dorsally united with the style column (Aristolochia), each consisting of 2 thecae with 4 pollen sacs, extrorse, rarely introrse (extra-Mal. spp.), dehiscing longitudinally. Ovary inferior (rarely half-inferior in extra-Mal. genera), 4—6-carpellate, 4—6-celled, syncarpous (or ± apocarpous in extra-Mal. Saruma); placentae parietal (distinct when young, then intruding and connivent axially, thus often seemingly axile); ovules usually many, anatropous, in 1 or 2 vertical rows in each locule of the ovary, horizontal or pendulous; style-column 3—many-lobed, sometimes some of the lobes redivided; stigmas or stigmatic tissue apical, lateral, or on the surface of style lobes. Fruits capsular or siliquiform (follicular or cocci in extra- Mal. genera), 4—6-celled; dehiscing apically towards the base (basipetal, e.g. Thottea) or basally towards the apex (acropetal, e.g. most Aristolochia); septicidal, rarely septifragal (some extra-Mai. Aristolochia) or bursting irregularly (extra-Mal. Asarum); rarely indehiscent (W. African Pararistolochia). Seeds many in each locule (1-seeded in extra-Mal. Euglypha), often coated with remains of placental tissue (membranous when dry), horizontal or pendulous, variously shaped; ovate, deltoid or triangular, flat, convex-concave, or longitudinally curved, or oblong (and triangular in cross-section), rugose, finely verrucose, or smooth, immarginate (Thottea; Aristolochia, p.p.) or winged (Aristolochia, p.p.); albumen fleshy, copious; embryo minute, cotyledons two, distinct. Distribution. There are 7 genera, Aristolochia worldwide, Asarum over the northern hemisphere, Thottea in continental Southeast Asia and Malesia, Pararistolochia in tropical Africa, and 3 monotypic genera, viz. Saruma in China, Holostylis and Euglypha in South America. As to number of species, Aristolochia is by far the largest with some 300 spp., largely concentrated in the New World, especially in Central and South America, in Malesia with 28 spp.; Asarum (incl Hexastylis and Heterotropa) with possibly some 70 spp. in northern temperate regions, Thottea with 26 spp., of which 22 in Malesia, and Pararistolochia with 12 spp. in West Africa.

Description: The ultrastructure of the spore walls of Beenakia dacostae (Beenakiaceae, Gomphales) has been studied. Spore walls are mainly composed of a distinct episporium and a thick, dark, ornamented ectosporium. The general structure is identical with that of other members of the Gomphales, such as Gomphus and Ramaria

Description: Type material of Tulasnella cystidiophora Höhn. & Litsch. has been studied. The species is characterized by often moniliform gloeocystidia and clamp-less hyphae (at least in the subhymenium).

Description: Several aspects of the sexuality in Mucorales are discussed. It is stated that neither heterothallism nor homothallism are absolute conditions and that a continuum exists between zygospores and azygospores. Mating type switching as known in ascomycetous yeasts would explain several up to now inexplicable phenomena.

Description: During an ecological study of fungi of the tidal mudflats in Kuwait, a Sporothrix species has been recorded twice, in 1977 and 1980. It differs from other species of the genus (de Hoog, 1974, 1978) in several characters and is here described as a new species. A comparison with similar species of the genus is added.

Description: J. J. Smith was born June 29th, 1867, at Antwerp, where his father was the director of the Netherlands’ Railway Post Office. In 1872 the family moved to Utrecht and in 1875 to Amsterdam. Smith spent his school days in the capital. His leisure hours were occupied by growing and sketching plants and tending such animals as mice and keeping an aquarium and a terrarium. His 10th birthday was celebrated by the establishment of a private herbarium, the first plant inserted being Bellis perennis. His years at secondary school were greatly influenced by the then teacher of Natural History, Dr J. C. Costerus, who advised Smith to look for a position in horticulture. Horticultural schools being not yet ”en vogue“, Smith got his education in this field at the Horticulturist’s Messrs Groenewegen & Co., Amsterdam. In these years the Orchids began to impress him and Smith spent his few free hours in making pictures of flowering species. The connection with Dr Costerus was continued. Together they looked after their herbaria and later on started to study teratologica, found in the Groenewegen gardens and greenhouses, a field in which both would publish several valuable papers later on. After having been working for his firm for 3½ years, Smith went to Kew where he stayed one year and afterwards to Brussels for completing his horticultural knowledge and skill. At Brussels he was working one year in the famous Orchid nursery of Messrs Linden, and then another year at the ”Jardin Botanique“.

Description: Pendant une tournée du chalutier ”De Lanessan“ de l’Institut Océanographique de Nhatrang (Annam) vers le récif Tizard¹) en avril 1936, une collection d’algues marines a été constituée, provenant des îlots Itu-Aba, Sand Caye et Nam Yit. La situation de ces îlots est environ 10° de latitude Nord et 114° de longitude Est. Qu’il me soit permis de remercier M. R. Serène de l’Institut Océanographique de l’Indochine à Cauda par Nhatrang, qui m’a confié l’étude de cette collection.

Description: In continuation of a previous publication by Lam, in which meiomery and pleiomery in male flowers of Canarium decumanum were described, the same phenomenon is now discussed concerning the fruits of C. Mehenbethene (176 of one single tree) and C. commune (1126 fruits mixed from more than one tree). An investigation of the material gave the following results: 1. C. commune and C. Mehenbethene are closely related; the latter may prove to be a polyploid of the former. Their areas are partly overlapping, but C. commune has its centre in the Moluccas, C. Mehenbethene in New Guinea and W. Polynesia. 2. A tendency to reduce the number of ovules and carpels in the ovary is assumed. By means of a statistical method (”phase index“) the position of either species in the phases of this regression is indicated. 3. From this, it is concluded that C. Mehenbethene represents a more advanced phase than C. commune and that therefore an eastward migration must be accepted. This agrees with other facts stated earlier, both in the Burseraceae and in other plant families of western origin. 4. In Canarium commune pleiomery is found in 2.3% of the fruits, meiomery in 0.45%, which agrees fairly well with the figures found earlier for the corolla and the androeceum of the male flowers of C. decumanum (0.9% and 0.3% respectively). 5. The desirability is expressed to investigate the following points: a. the ontogeny and the fertilization of ovaries and ovules in Canarium. b. cytological relations between related trees in the tropics, especially as far as they may supply indications towards migration tracks (cf. the work of Hagerup on Vaccinium [Hereditas 18, 1933]). c. the ”phase index“ of a number of related Canarium species. d. the exact distribution of some of the phases mentioned along those migration tracks which are both geologically and biogeographically supported (e.g. Sunda centre—Philippines, Philippines—Moluccas—New Guinea, New Guinea—Moluccas—Central Celebes, Malay Peninsula—Sumatra—Java, etc.).

Description: Im Jahre 1907 wurde ich als Botaniker der Gouvernements China-Plantagen in Tjinjiroean bei Pengalengan, West-Java, angestellt, wo ich bis 1916 arbeitete. Tjinjiroean liegt etwa 1566 m über dem Meere und hat ein sehr feuchtes Klima. Es war sehr interessant nachzugehen, welche aus der Ebene von Java und aus Europa eingeführten Pflanzen dort wachsen würden. Was würde der Einfluss des Klimas, der Meereshöhe, der Temperatur, u.s.w. auf die Pflanzen sein? In Tjinjiroean fand ich sogleich viele eingeführte Pflanzen, welche dort üppig wuchsen. In den Chinaplantagen fand ich Georginen und Tropaeolum majus L. verwildert; in meinem Garten blühte Richardia africana Kunth reichlich, bildete Früchte, welche wieder zahlreiche Pflanzen lieferten. Nur einige interessante Pflanzen werde ich hier weiter erwähnen.

Description: Een man, die zich nimmer op den voorgrond stelde en wiens naam toch in de geheele botanische wereld bekend is, moet wel heel wat in die wereld hebben gepresteerd. Zoo’n man is Dr J. J. Smith, die op 29 Juni 1937 zijn 70sten verjaardag viert. Zeventig jaar te worden is op zichzelf beschouwd geen verdienste, maar het geeft vrienden en vereerders zulk een mooie gelegenheid den jubilaris eens te toonen, hoe zeer men zijn werk waardeert!

Description: Het lijkt mij niet mogelijk een juisten indruk te krijgen van de beteekenis van J. J. Smith’s phytographisch werk voor den huidigen kweeker, zonder de belangrijkste phasen in de geschiedenis der Orchidophilie in Europa kort te schetsen, die aan dit werk zijn voorafgegaan. Deze geschiedenis heeft zich practisch geheel in Engeland afgespeeld. Dit machtige rijk, in zijn gouden eeuw onbetwist heerscher ter zee, had ter behartiging van zijne overzeesche belangen de beschikking over een kolossale handelsvloot. De bemanningen der schepen voerden van heinde en verre allerlei rariteiten mede, ook levende planten en dieren. Op deze wijze kwamen in de laatste helft der achttiende eeuw de eerste exotische Orchideeën binnen uit gebieden, die niet al te ver van Engeland af lagen: Jamaica, de Bahama-eilanden, Trinidad.

Description: Het is mij een bijzonder voorrecht, om uit het leven van Dr Smith eenige bijzonderheden te mogen vertellen, waarvan enkele wellicht minder algemeen bekend zijn. Deels heb ik de gegevens geput uit datgene wat van zijn levensloop bekend is, deels stammen ze uit mijn persoonlijk contact met Dr Smith, en de gelegenheid van dit jubileum lijkt mij bij uitstek geschikt om dezen te bescheiden werker in het licht te plaatsen waarin hij behoort te staan. In de beginjaren van mijn loopbaan als Hortulanus van ’s Lands Plantentuin was Dr Smith voor mij de groote vraagbaak, was hij de man die met zijn groote liefde voor en zijn uitgebreide kennis van den Plantentuin mij als het ware heeft ingewerkt en opgeleid.

Description: Some collections which I received recently, contain interesting addenda to former studies of the paleotropical Frullaniaceae (cf. especially “De Frullaniaceis VII”, Ann. Bryol. Suppl. Vol. I, 1930) and Lejeuneaceae Holostipae (esp. “De Frullaniaceis XVII”, Ann. Bryol. Suppl. Vol. IV, 1934).

Description: Durch die extremen Existenzbedingungen, welche die Mangroven als: Formation bestimmen, sollte man glauben, dass die ökologischen Probleme, die sich in dieser Hinsicht zeigen, leicht gelöst werden könnten, um so mehr, weil diese Pflanzengenossenschaft relativ arm an Arten ist. Dass dies jedoch nicht der Fall ist, möge hier im Hinblick auf die Verbreitung der Lumnitzera-Arten im Malaiischen Archipel mit einigen Beispielen gezeigt werden. Im Jahre 1924 habe ich kurz auf die Verbreitung von 3 Lumnitzera- Arten im genannten Gebiet hingewiesen ¹). Meine Absicht war, speziell auf die unerklärliche Erscheinung aufmerksam zu machen, dass L. littorea (Jack) Voigt das Küstengebiet rund um die Java See, im Gegensatz zu L. racemosa Willd., vermeidet, obwohl beide Arten nicht nur in, sondern auch ausserhalb des Malaiischen Archipels vorkommen, ja selbst zusammen in ziemlicher Nähe angetroffen werden. Bevor wir diese Erscheinung noch einmal näher betrachten, möchte ich an der Hand von beigefügter Karte (Fig. 1) das gesamte Verbreitungsgebiet nachgehen. Dieses Gebiet liegt nahezu vollkommen innerhalb der Wendekreise der alten Welt ²): Die Mangroven, wozu Lumnitzera gehört, finden als selbständige Waldoder Gebüschformation ihre natürliche Begrenzung ungefähr auf den gleichen Breiten. Nur L. racemosa überschreitet grade an 2 Stellen die- Wendekreise: An der Ostküste von Afrika streckt sie sich südlich vom Steinbrockkreis bis in die Mangroven bei Durban aus, während sie nördlich vom Wendekreis des Krebses noch in dem Riu Kiu (Lu Tschu) Archipel, nördlich von Formosa vorkommt.

Description: De publicatie van dit deel is mogelijk gemaakt door den geldelijken steun van vele van Smith’s vrienden, wier handteekeningen zijn vereenigd in een album, dat hem is aangeboden tezamen met dit Jubileum-Supplement van „Blumea” en de speciale aflevering van het „Bulletin du Jardin botanique de Buitenzorg”. Het oude Menangkabausche echte gouddraadweefsel uit Kota Gadang, dat heeft gediend voor de banden van het album en van de voor Dr Smith bestemde exemplaren van „Blumea” en het „Bulletin”, dankt het Comité ad hoc aan Dr E. R. Jacobson te Bandoeng.

Description: A new species, Alstonia undulifolia Kochummen & Wong, is described from the Malay Peninsula. Two sections of the genus occur in the Malay Peninsula, Alstonia sect. Monuraspermum Mon. and Alstonia sect. Alstonia, the latter being the correct name for what was previously known as sect. Pala (Adr. Juss.) Benth. Various characteristics, including growth architecture, are examined for their usefulness in distinguishing these two sections of the genus. In comparing A. angustiloba Miq. and A. pneumatophora Berger, both of which have not been properly differentiated by characteristics of the reproductive organs, A. pneumatophora var. petiolata Mon. is reduced to synonymy under A. angustiloba. A key to the seven species of Alstonia native to the Malay Peninsula is provided.

Description: ROXBURGH described in his Flora in the year 1820 a very curious annual grass and placed it in the genus Eleusine as E. verticillata ROXB.. This grass has spikelets which agree in many characters with those of the genus Eleusine, especially as to the rugose grain with a caducous pericarp, but differing from Eleusine in the up to 20-flowered spikelets and in the lemmas with a three-cuspidate summit. The many-flowered spikelets give the plant more the habit of an Eragrostis and under this genus a specimen was mentioned by WALLICH in his Catalogue. There are in the characters of the spikelets many other differences with the genus Eleusine and with Eragrostis. KUNTH and STEUDEL, indeed placed the plant under Leptochloa and there are still other opinions about this plant. An advancement in this matter was the opinion of LINDLEY, who published in the year 1836 a new genus Acrachne WIGHT et ARN., in the second edition of his ”Natural System of Botany“, p. 381, based upon ROXBURGH’s Eleusine verticillata, The name Acrachne was already given by WIGHT et ARNOTT as Acrachne eleusinoides, a nomen in WIGHT, Cat. no. 1760. This name was placed by STEUDEL in the year 1854 under E. verticillata ROXB., a name also accepted by NEES. The name Acrachne, although based upon a species which was validly published, was, however, not described by LINDLEY and the combination A. verticillata was not made by LINDLEY. At that time the genus Acrachne was therefore not valid.

Description: Among the collections of Knema acquired by the Rijksherbarium since the publication of my new account of the genus Knema, in Blumea 25, 1979: 321 — 478, a few specimens caused problems with the identification, and at closer examination these yielded facts of interest which are published here. Some specimens represented stages not yet known, for instance fruits, or male flowers, while other specimens meant a significant range extension of the species. Two new species and one new subspecies are described. For easy reference, the sequence and numbers of the species presently treated correspond with the numbers as used in the account of 1979. The new species bear the number of the species after which they appear in the general key of 1979, with the addition ‘-bis’.

Description: This extensive collection, famous among algologists both of the Old and the New World, forms part of the collections of the National Herbarium (Rijksherbarium) Leiden since 1934. About fifty years ago it was started by Mrs. Dr. A. A. WEBER-VAN BOSSE (1852—hodie), an enthusiastic pupil of HUGO DE VRIES. The colonies of Nostoc, living in the ditches round about the Dutch village of Doom, evoked her admiration, which was the primary cause of an intense study in the freshwater as well as in the marine Algae. In the harbour of Den Helder North Sea Algae were collected; by collecting Algae on trips to the French Atlantic Coasts and several times to Norway (1883—1885) and further on a South African journey (1894—1895) the herbarium grew, as it did by the Malaysian specimens collected in Java, Celebes, etc. (1888—1889). During this Malaysian tour Mrs. WEBER worked in Tjibodas, where she described the new genus Phytophysa. In Sumatra (West Coast, Lake of Manindjau) she discovered in collaboration with her husband, MAX WEBER, a new case of symbiosis between Algae and Sponges.

Description: The early development (ontogeny) of the carpels of 20 species belonging to 8 apocarpous families was investigated with the scanning electron microscope. The results indicate that on the floral apex a circular or a convex meristem develops into an obliquely ascidiate primordium by unequal growth of its periphery. By further unequal growth it develops into a young carpel. The terminal mouth of a cup becomes the lateral cleft of a carpel. The different forms of the young carpels in different species are defined by the varying degree of development of the adaxial region of the initial meristem and/or its margin on the side of the floral apex. This hypothesis is theoretically evaluated.

Description: Thanks to the kind cooperation of Dr. ROBERT PILGER, Director of the Botanical Gardens and Museums at Berlin-Dahlem, I have recently had the privilege of studying and photographing a unique specimen belonging to that institution, which bears the words „Schizostachyum Blumii nobis”, in the hand of NEES, the author of the species. Although there are no data on the sheet to indicate its source, or the date of the determination, this presumably represents NEES’S type³) of this species (which is the type species of the genus). At any rate, the available evidence 4) points to that conclusion, and the specimen agrees in all respects with NEES’ description of the genus and of the type species (NEES, 1829, pp. 534—5). Since the original characterizations are so brief and, since those parts referring to the spikelets are so difficult to interpret, I present here a full description 5) of the rather fragmentary type specimen.