ALBERT

Description: The effects of prescribed burning and complete clearcutting on Populustremuloides and associated hardwoods and shrubs were compared for 8 years after commercial harvest of a 60-year-old P. tremuloides stand. Because of the lack of suitable burning weather, P. tremuloides suckers were 2 years old before the burn could be made. All suckers were killed by fire and new suckers were more numerous but less vigorous, probably because of heat damage to shallow sucker-producing roots, loss of nitrogen, and reduced root carbohydrate reserves. Although prescribed fire can effectively control residual hardwood overstories detrimental to P. tremuloides sucker growth and survival, the long term effect of fire on sucker growth is unknown. Fire can be used to prepare sites for P. tremuloides regeneration when other methods are unavailable or impractical. Burning should be done during the first dormant season following logging. Effort should be made to distribute slash uniformly to provide even burning conditions. Burning prescription guidelines are given.

Description: The pine-wood nematode, Bursaphelenchusxylophilus (Steiner and Buhrer), was most commonly extracted from Cerambycidae emerging from nematode-infested pines in Minnesota and Wisconsin during 1981 and 1982. The greatest number of nematodes were extracted from Monochamusscutellatus (Say) and Monochamuscarolinensis (Olivier). Low numbers of B. xylophilus were found in some buprestids but no nematodes were found in the curulionid and scolytid beetles examined. Two species of Cerambycidae, Monochamusmannorator (Kiby.) and M. scutellatus were associated with B. xylophilus from balsam fir in Minnesota. Bursaphelenchusxylophilus from insects associated with balsam fir were morphologically different from insects associated with pine. Dauer larvae of B. xylophilus were concentrated in the thoracic segments of M. scutellatus and Monochamusmutator (Lec.) examined. Bursaphelenchusxylophilus was transmitted to twigs during maturation feeding and to logs during oviposition by M. carolinensis, M. mutator, and M. scutellatus.

Description: Many trees in stands of Douglas fir, western hemlock and western red cedar on Vancouver Island were joined by functional grafts. In a partially cut stand, 45% of the stumps showed evidence of continued growth and half of these (23%) were still growing vigorously more than 22 years after logging. On experimentally detopped trees, growth extended several meters up the bole. Dominant trees usually supported the growth of the root system and lower boles of grafted suppressed trees.Translocation through grafts may partially explain the frequent stagnation and slow recovery of stands after thinning from above, and may be involved in the usually rapid increase of growth after thinning from below. It is probably a contributing factor in establishing dominance and determining mortality in overtopped trees. In species that graft freely, the use of silvicides in spacing and thinning treatments should be restricted to young stands before grafts are established.

Description: Alnusrubra Bong, dominates the first 65 – 80 years of a sere that is initiated naturally on the terraces of the Hoh River. Stands of 14, 24, and 65 years were studied to determine to what extent the Alnus stage enriched the nitrogen inventory of the site. Bare sandbars deposited by the river had a mean of 783 kg/ha nitrogen. Alnus communities caused an increase in the nitrogen inventory so that, by 65 years, total community nitrogen was 4659 kg/ha, soils held 3594 kg/ha N in the upper 45 cm, and Alnus trees held 942 kg/ha N. The nitrogen contents of the soil, Alnus wood, bark, and branches, grasses, total aboveground biomass, total belowground biomass, and sticks less than 1 cm diameter all showed significant increases from 14 to 65 years. The A. rubra stage is an important link in the nutrient inventory between unvegetated, recently deposited sandbars and the climax coniferous forests dominated by Tsugaheterophylla (Raf.) Sarg. and Piceasitchensis (Bong.) Carr.

Description: Flooding for 30 days induced several changes in Quercusmacrocarpa Michx. seedlings, with stomatal closure among the earliest responses. Stomata remained more closed in flooded than in unflooded plants during the entire experimental period. Leaf water potential was consistently higher in flooded than in unflooded plants. Other responses to flooding included acceleration of ethylene production by stems; formation of hypertrophied lenticels on submerged portions of stems; growth inhibition, with greatest reduction in roots; and formation of a few adventitious roots on submerged portions of the stem above the soil line. Some of the morphological responses to flooding, especially formation of hypertrophied lenticels, appeared to be associated with increased ethylene production. Quercusmacrocarpa seedlings adapted poorly to flooding as shown by failure of stomata to reopen after an early period of flooding and low capacity for production of adventitious roots. The much greater inhibition of root growth than shoot growth by flooding will reduce drought tolerance after floodwaters recede.

Description: Root elongation of greenhouse-grown Alaskan taiga tree seedlings increased with increasing root temperature in all six species examined and was most temperature sensitive in warm-adapted aspen (Populustremuloides Michx.). Root elongation was slower in fine than large roots and in black spruce (Piceamariana (Mill.) B.S.P.) was less temperature sensitive in fine than in large roots. Root elongation in the laboratory was slowest in black spruce, which has an inherently slow growth rate, and most rapid in poplar (Populusbalsamifera L.) and aspen, which grow more rapidly. In contrast, field root elongation rates tended to be highest in black spruce from cold wet sites, suggesting that site factors other than soil temperature (e.g., moisture) predominated over genetic differences among species in determining field root elongation rates. The seasonal pattern of root elongation was closely correlated with soil temperature and reached maximum rates in July for all tree species (except aspen medium-sized roots). Most roots of each species were in the top 20 cm of soil. However, root growth penetrated to greater depth in warm compared with cold sites. Root biomass in a 130-year black spruce forest (1230 g/m2) comprised only 15% of total tree biomass. Root biomass of 25-year aspen and 60-year poplar sites (517 and 5385 g/m2, respectively) comprised a greater proportion (57% in poplar) of total tree biomass than in spruce.

Description: The patterns of translocation of carbon in different-age tissues of four common moss species in a black spruce (Piceamariana (Mill.) B.S.P.) dominated forest near Fairbanks, Alaska, were studied by 14C labelling and carbohydrate analysis. A simple, in-vial combustion technique was developed for combustion of small (

Description: Water stress of subalpine conifer species may be measured with the pressure chamber after several hours of tissue storage. Tissue samples stored in cool, humid vials exhibited very little change in xylem pressure potential over a 4-h period. However, xylem pressure potential declined steadily when a source of water vapor was not available. Xylem pressure potentials of current-year and 1-year-old needles of lodgepole pine (Pinuscontorta var. latifolia Engelm.) were slightly lower than those of older needles.

Description: This paper discusses the occurrence of man-caused forest fires during the summer fire season in a section of northwestern Ontario. Fire occurrence is viewed as being a chance process and a stochastic model is developed to describe it. The results of this study indicate that a Poisson model with the average number of fires per day depending on the Fine Fuel Moisture Code is appropriate.

Description: The aboveground biomass of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and annual production over 7 years was studied in relation to thinning and nitrogen fertilization. Biomass yield of both treatments increased during the first 3–4 years then decreased for fertilization but not with thinning. Treatments doubled biomass production of individual trees over the study period when applied separately and quadrupled it when combined. Annual biomass production per unit of foliage (E) increased during the first 3–4 years, but was at or below control level after 7 years. The increased E accounted for 20, 37, and 27% of the stemwood dry matter response to thinning, fertilization and the combined treatments, respectively; the remainder was attributed to an increase in foliage biomass. Thinning, but not fertilization, influenced distribution of radial growth along the stem, increasing growth only below the top one-third of the stem. This pattern was related to crown development.

Description: In order to test the hypothesis that the deterioration of trembling aspen (Populustremuloides Michx.) is related to variations in climate, soil properties, and genotype, 59 trembling aspen clones were sampled in Michigan, Wisconsin, Minnesota, and Ontario. A longevity index (LI) was calculated by taking the difference between predicted basal area from normal yield tables and measured basal area for each clone. Correlations of environmental variables with LI indicate that aspen longevity decreases with increasing mean annual temperature. Under similar temperature regimes, aspen growing on xeric sites and on sites low in exchangeable Ca are most susceptible to early breakup. Since there were negligible differences in soil properties between nine pairs of adjacent deteriorating and relatively well stocked clones, we hypothesize that, under similar environmental conditions, variation in the timing of deterioration may be due to genotypic differences between clones.

Description: Carbohydrate reserves and root growth potential (RGP) of 2 + 0 Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) seedlings were monitored through a lifting season and during dark, cold storage. Concentrations of total nonstructural carbohydrate and extractable sugars in root and stem tissues remained relatively constant through winter, while foliar sugars showed a sharp midwinter peak at about 195 mg•g−1 dry weight. RGP was lowest in November and March and peaked in January. During storage at +2 and −1 °C, carbohydrates were depleted in all tissues through respiratory consumption. In contrast, RGP increased during the first 6 months in storage and then fell rapidly. The results do not support the view that changes in RGP are driven by changes in carbohydrate concentrations. Storage may affect frost hardiness and drought resistance through its effect on sugar concentrations.

Description: A nonlinear analytical model is developed to describe the relationship between average plant size (weight or volume) and stand density in single-aged, monospecific plant populations. The model gives estimates of the slope and intercept of the −3/2 power rule asymptote, the nature of the size–density trajectory, and such features as relative density at crown closure and the effects of soil type or site index. The model is tested by growing red alder (Alnusrubra Bong.) seedlings under greenhouse and lath house conditions at three initial spacings (8 × 8, 4 × 4, and 2 × 2 cm) and two soil types (river loam and alder forest soil) for 525 growth days. There are seven harvests, starting at crown closure. All size–density trajectories tend consistently towards the same single asymptote irrespective of initial spacing, soil type, or age. The asymptote slope and intercept are 1.46 and ca. 94 kg tree−1•m−2. The crown closure line is parallel to the asymptote at a relative density of 4.6 × 10−3. The model also adequately describes the size–density trajectories for 20–50-year-old red pines (Pinusresinosa Ait.) growing at six initial spacings. The asymptote slope and intercept are 1.6 and ca. 87 × 103 m3•tree−1•ha−1, respectively.

Description: The pipe model theory presents the idea that a unit weight of tree foliage is serviced by a specific cross-sectional area of conducting sapwood in the crown. Below the crown, a large fraction of the tree bole may be nonconducting tissue, so the sapwood area would have to be known to estimate foliage. We applied the pipe model theory to the analysis of several western coniferous species to learn whether the distribution of canopy leaf area could be accurately estimated from knowledge of the sapwood cross-sectional area at various heights, including breast height (1.37 m). Results are excellent, but taper in the conducting area must be considered when sapwood area is measured below the crown.

Description: The history of spruce budworm (Choristoneurafumiferana (Clem.)) outbreaks for the past 200 to 300 years, for nine regions in eastern Canada, indicates that outbreaks have occurred more frequently in the 20th century than previously. Regionally, 21 outbreaks took place in the past 80 years compared with 9 in the preceding 100 years. Earlier infestations were restricted to specific regions, but in the 20th century they have coalesced and increased in size, the outbreaks of 1910, 1940, and 1970 having covered 10, 25, and 55 million ha respectively. Reasons for the increase in frequency, extent, and severity of outbreaks appear mostly attributable to changes caused by man, in the forest ecosystem. Clear-cutting of pulpwood stands, fire protection, and use of pesticides against budworm favor fir–spruce stands, rendering the forest more prone to budworm attack. The manner and degree to which each of these practices has altered forest composition is discussed. In the future, most of these practices are expected to continue and their effects could intensify, especially in regions of recent application. Other practices, including large-scale planting of white spruce, could further increase the susceptibility of forest stands. Forest management, aimed at reducing the occurrence of extensive fir–spruce stands, has been advocated as a long-term solution to the budworm problem. The implementation of this measure at a time when man's actions result in the proliferation of fir presents a most serious challenge to forest managers.

Description: At 10 locations in Oregon and Washington, tree mortality resulted in dry-matter transfer of 1.5–4.5 Mg•ha−1•year−1 of boles and branches to the forest floor and 0.3–1.3 Mg•ha−1•year−1 of large-diameter roots directly to the mineral soil. The first value is about the same as that reported for leaf fall in similar stands; the second value generally is smaller than that reported for fine root turnover. Results are based on measurements by the U.S. Forest Service spanning 16–46 years and areas as large as 42 ha. Values based on intervals

Description: The resistance to a pulsed electric current decreased sharply as the tips of stainless steel electrodes entered discolored and decayed tissues in living deciduous trees. The apparatus delivering the pulsed electric current is described.

Description: Flooding of soil for 45 days severely inhibited growth of 115-day-old Pinusbanksiana Lamb, and P. resinosa Ait. seedlings, with significant effects apparent within 15 days after initiation of flooding. Both species adapted poorly to flooding of soil but P. banksiana was more adversely affected than P. resinosa as shown by earlier and more drastic reduction of growth in the former species. In both species flooding decreased the rate of height growth, production of secondary needles, dry weight increment, and relative growth rates of various plant parts. Root growth of both species was reduced more than shoot growth. The reduction in dry weight increment of root systems of both species reflected arrested branching and elongation of roots as well as decay of roots, mainly the very small nonwoody roots. Flooding slightly stimulated ethylene production by submerged stems.

Description: The presence of ectomycorrhizal and vesicular–arbuscular (VA) mycorrhizal fungi in soils from five sites in a mixed conifer zone in southwest Oregon, each consisting of a 1- to 1.5-year-old clear-cut adjacent to an undisturbed forest stand, was determined by bioassay with Pseudotsugamenziesii (Mirb.) Franco, Pinusponderosa Dougl. ex P. Laws & C. Laws, and Trifoliumsubterraneum L. 'Mt. Barker' as hosts grown at root zone temperatures ranging from 7.5 to 35 °C. Maximum formation of both ectomycorrhizae and VA mycorrhizae occurred at 18.5–24 °C in soils from all sites, and there were no significant qualitative or quantitative differences between disturbed (clear-cut) or undisturbed (forest) soils. Mycorrhiza formation was moderate even at the lowest temperature tested (7.5 °C) but was greatly reduced or prevented at or above 29.5 °C. Treatment of soil at 35 °C for 1 week did not appear to adversely affect viability of ectomycorrhizal fungus propagules, but young mycorrhizae subjected to the same treatment appeared to be severely injured. Thus the ability of native mycorrhizal fungi to grow at low soil temperatures is especially important as they may contribute to the survival of seedlings outplanted into climatic zones characterized by warm, dry summers following cool, wet winters and springs.

Description: Reduced levels of root activity were associated with winter injury to shoots of 1-year-old black spruce (Piceamariana (Mill.) B.S.P.) container seedlings.

Description: Techniques are described for field and laboratory extraction of small quantities of sugar maple (Acersaccharum Marsh.) xylem sap when pressure in the xylem is less than atmospheric. Accurate estimates of sap sugar concentration can be made with a hand refractometer most of the year. There is, however, appreciable within-tree variation in sap sugar concentration at any given time and within short periods of time.

Description: Conflicting reports of the occurrence of Cephaloascusfragrans Han. relate to the varied appearance and color of this mould upon wood and the fact it may be overgrown by other, more vigorous fungi. C. fragrans presents a problem in the commercial treatment of lumber, as it is not inhibited by formulations of chlorinated phenols presently used to prevent mould and sap stain.

Description: Progenies belonging to 49 open-pollinated families of paper birch (Betulapapyrifera Marsh.) representing 13 provenances were evaluated sequentially for rate of root elongation in solutions containing only calcium nitrate and solutions containing calcium nitrate + 120 ppm aluminum. Provenances, but not families within provenances, differed significantly in both rate of root elongation and apparent response to Al. Provenance mean tolerance indices (root growth in Al solutions/root growth in non-Al solutions) varied from 0.10 to 0.37. Exposure to 120 ppm Al caused decreased concentrations of Ca and Cu in the foliage, decreased or nearly unchanged concentrations of P and Mg, and increased concentrations of Al, B, and Mn. Compared with the more tolerant provenances, intolerant provenances exhibited generally greater increases in foliar Al concentration after exposure to Al. Differential tolerance to Al will be useful in improving the performance of planted trees on Al-toxic sites.

Description: A study was made on the movement of water from permeable-, partially-permeable-, and impermeable-walled seedling containers, filled with either peat or a peat and mineral soil mixture, into sand and loam planting soils. Irrespective of container type, there was rapid outward movement of from 15 to 39% of moisture initially stored in the containers to the surrounding soil within a few days of planting. Distinct wetting front patterns emanating from each type of container suggested that the path and eventual location of the moving moisture may be more critical for seedling survival than the amount moving into the surrounding planting soil. Radial moisture movement from the interface between the container and the soil to the soil surface from permeable- and partially-permeable-walled containers (Jiffy peat pot and Conwed tube) may aggravate evaporational losses. Elimination of radial moisture movement from an impermeable-walled container (Ontario tube) and limitation of outward flow from the open bottom to the root zone appear to be advantageous for seedling survival. Specifications for redesign of seedling containers for reforestation work are suggested.

Description: Three methods were compared for total sulphur analysis of tree foliage: magnesium nitrate ashing followed by turbidimetry, sodium hypobromite (NaOBr) oxidation followed by colorimetry, and an instrumental method using the Fisher S analyser, model 475. The latter method was found to be substantially better than the other methods with respect to accuracy and speed of analysis, and of satisfactory precision (average coefficient of variation (CV) was 2.5%). Dry ashing resulted in good precision (CV = 1.9%) but only recovered 76% of the reported total S in the United States National Bureau of Standards (NBS) orchard leaf standard. The NaOBr method recovered 88% of reported S from the NBS standard but showed poorer precision (CV = 4.5%). The Fisher analyser had the lowest operating cost per sample, largely owing to lower labour costs.

Description: Seasonal patterns of change in lipids, sugars, starch, labile (ethanol soluble) constituents, holocellulose, and lignin were studied in six forest-grown white oak (Quercusalba L.) trees. Contents of metabolically active constituents in leaves, twigs, branches, boles (upper and lower), and roots (support and small lateral) were used to construct whole-tree budgets of energy allocation. [14C]Sucrose was also concurrently supplied to the study trees to follow the fate and efficiency of utilization of food reserves. Results showed that white oak rapidly mobilized and replaced food reserves during the critical period of canopy generation in the spring. Starch was more important as a reserve food than lipids or sugar. Large fluctuations in starch in roots in spring and fall suggested a bimodal belowground growth pattern. Labile constituents showed the most pronounced seasonal changes and dominated the calculated whole-tree energy flux patterns. Rapid decline in labile compounds in early spring and a parallel increase in holocellulose suggested a possible pattern of mobilization and resupply of stored reserves associated with in cell wells. This possibility was supported by a concurrent shift of labile 14C to nonlabile 14C in tissues. Canopy generation was calculated to have cost ≤17.7 kg of glucose (1.6 g glucose/g of canopy) of which 13 kg appeared to have come from within the canopy.

Description: Total biomass of an aspen – mixed hardwood – spodosol ecosystem in northern Wisconsin, U.S.A., was 197 t/ha and net primary production was 11.5 t/ha per year. Populustremuloides Michx. accounted for 60% of the total biomass and 56% of the annual production and Acersaccharum Marsh, accounted for 25% of the biomass and 28% of the annual production. For all species combined, bole wood was 63% of the total biomass and bole bark was 12%. Bole wood was 33% and bole bark was 7% of the total production. Although crowns accounted for only 15% of the total biomass, they were responsible for 49% of net annual production. Using allometric equations from the literature, root biomass and production were calculated as being approximately 10% of the total biomass and of the annual production. The average rate of total production per unit leaf tissue was 5.7 g production/g leaf tissue for P. tremuloides and 3.7 g/g for A. saccharum.

Description: Water chemistry profiles of an 18-year-old forest ecosystem are compared with those of a 70- to 90-year-old forest ecosystem for a 9-month period. The younger ecosystem was dominated by Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and red alder (Alnusrubra Bong.) whereas western hemlock (Tsugaheterophylla (Raf.) Sarg.), Douglas-fir, and western red cedar (Thujaplicata Donn.) dominated the older ecosystem. Concentrations of nutrients and other chemicals were measured in throughfall, forest floor and mineral soil lecachates, saturated-zone water, and stream water. Concentrations of dissolved chemicals were much greater in the younger ecosystem than in the older ecosystem at intermediate stages in the profiles. However, stream water concentrations differed less between the two ecosystems for most of the chemicals investigated. Nitrate and silica were exceptional; stream water nitrate concentrations in the younger ecosystem averaged 16 times greater than those in the older ecosystem. This was probably a result of biological nitrogen fixation by red alder in the younger ecosystem, a process which would more than compensate for the higher nitrate losses. Silica concentrations in the younger ecosystem consistently exceeded levels in the older ecosystem by 40 to 100%. suggesting a possibility of a greater rate of mineral weathering in the younger ecosystem.Although nutrient concentrations were higher in the soil leachates of the younger ecosystem, these higher levels failed to persist through the saturated-zone water and stream water stages of the water chemistry profile. Consequently, the younger ecosystem appeared relatively more efficient at retaining dissolved nutrients than the older ecosystem. Stream water chemistry was relatively insensitive to the magnitudes of the differences in biogeochemical process rates of the two ecosystems.

Description: Seedlings of five half-sibs (seedlings that share a commom maternal parent) of Pinustaeda L. exposed to a bidirectional mechanical perturbation for 6 months exhibited a reduced rate of stem elongation and a reduction in the length of both juvenile leaves and needles. Of the five half-sibs tested, two different anatomical mechanisms of radial enlargement could be identified. Two half-sibs exhibited increased radial growth in the xylem parallel to the direction of the perturbation and no change in the axis perpendicular to the perturbation. Two exhibited a reduction in growth perpendicular to the perturbation with no change in the axis parallel to the perturbation. One half-sib appears to be intermediate between the two other pairs of half-sibs.An overall increase in extractable compounds was observed in the organic solvent fraction of mechanically perturbed seedlings. The amount of lignin appeared to remain fairly constant, and there was no change in the amounts of holocellulose or water extractable compounds.

Description: Above- and below-ground net primary production was estimated for 40-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stands growing on sites with apparently large differences in productivity potential. Aboveground net production was estimated from direct measurements of tree growth; belowground productivity was derived from data obtained by sorting live and dead roots from soil cores used in combination with measurements of root growth on observation windows.Aboveground net production was 13.7 t•ha−1 on the high productivity site and 7.3 t•ha−1 on the low productivity site. Belowground dry matter production on the high productivity site was 4.1 t•ha−1 compared with 8.1 t•ha−1 for the poorer site. On the more productive site, 8% of total stand dry matter production was in fine roots in contrast to over 36% on the poorer site. The difference in total net production (aboveground plus belowground) between the two sites was small (2.4 t•ha−1). Apparent differences in aboveground productivity may, to a large extent, result from the need for a greater investment in the fine roots on harsher sites.

Description: Low addition levels of high enrichment isotope (〉1% of the total nitrogen pool with 99 at.% excess 15N) were used to follow nitrogen movement through selected forest floor components of permafrost-free and permafrost-dominated black spruce ecosystems in subarctic Alaska. The nitrogen pool examined in this study was the total nitrogen pool. 15N was retained most effectively by the feather moss layer (Pleuroziumschreberi (BSG.) Mitt. and Hylocomiumsplendens (Hedw.) BSG.) on both black spruce sites. Twenty-eight months after isotope application the feather moss layer still contained over 90% of the 15N that could be recovered. The limited movement of 15N between feather moss layers and underlying forest floor horizons appeared to be slightly affected by climatological events. Differences in 15N movement patterns between permafrost-free and permafrost-dominated black spruce sites are discussed in terms of precipitation, soil temperature, and biological controls.

Description: Two parameters of growth rings from Engelmann spruce (Piceaengelmannii Parry) near Peyto lake, Alberta were studied and compared. Indices of maximum density of the latewood were derived from densitometric plots of X-ray negatives. Indices of ring widths were also obtained from the same specimens. Latewood density proved to be more useful for dendrochronological studies than ring width. The indices of density were significantly correlated with mean maximum air temperature and monthly runoff during August for three rivers in the region near Peyto lake. Analysis of latewood density as well as ring width improves the potential for dating tree-ring materials, and for using them to estimate past environment, especially for trees, such as the Peyto lake Engelmann spruce, with rings that do not vary greatly in width from one year to the next.

Description: A data screening algorithm (SCREEN) identified stand age, average stand diameter, timber type, soil type, aspect, habitat type, and elevation as forest stand or site characteristics strongly associated with root-disease occurrence in northern Idaho forests. The logistic regression model was used to predict root disease center occurrence. A second algorithm (RISK) was used to compute the estimates for the coefficients and to test different prediction equations. On wet aspects, highest probability of root disease centers was found on soils with lowest year-round moisture availability. The reverse was true on dry aspects. Increased slope was associated with increased root disease. Root disease center frequency was inversely related to elevation and directly related to occurrence of Pseudotsugamenziesii (Mirb.) Franco and Abiesgrandis (Dougl.) Lindl. Expression of disease centers was maximum at 60–100 years of stand age on all habitat types. A habitat-type–age interaction was observed in oldest aged stands.

Description: The mechanical and physiological support of a tree's crown is the principal function of its bole. A simple model for Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) of different crown classes explains much of the observed differences in stem form development. Sapwood cross-sectional area at any height on the bole of the tree was found to be related linearly to the amount of foliage above that point; however, in large trees the sapwood area needed to supply transpiring foliage with water is insufficient to provide mechanical support. The combination of sapwood and heartwood was found to provide the stem form that would be expected to ensure uniform resistance to bending by the wind.

Description: This paper describes the spatial pattern, expressed by Pielou's nonrandomness index, of trees within 13 sampled tracts from the major forest types of northern Ontario. Results indicate that: (a) the majority of natural coniferous or mixedwood stands have highly clustered patterns; (b) hardwood stands or the hardwood component of the mixedwood stands show nearly 'random' spatial patterns; and (c) uniform spacing in natural stands is very rare. Results also indicate that spatial patterns vary considerably during stand development for the various species group – size class combinations within a stand and between different forest types. The importance of spatial pattern and stand population dynamics is pointed out with regard to growth simulation modelling and mechanized harvesting and thinning studies.

Description: The effects of defoliation by western spruce budworm (Choristoneuraoccidentalis (Freeman)), on Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) radial growth at breast height and tree mortality are given. Four hundred and twenty trees were marked in an 81-year-old stand, and their defoliation levels were recorded annually from 1970 to 1980 in an outbreak that lasted from 1970 to 1974, inclusive. Forty-one trees were felled and dissected in 1977, 3 years after recovery began. The number of stems per hectare was reduced by 39.3% and basal area by 11.6% through mortality, most occurring among the small diameter, suppressed, and intermediate trees. Relationships were established between mortality and defoliation. Radial increments were examined, and the presence of four outbreaks during the life of the stand was detected. The combined effect of these infestations amounted to a loss of about 12% of the estimated potential diameter had not the insects been active. The most recent outbreak (1970–1974) caused a total of 10 years of subnormal growth, including 5 years due to defoliation and 5 years of recovery. The relationship between radial increment losses and defoliation intensity and duration is studied and quantified.

Description: Copper, nickel, and cobalt solutions were added to a variety of substrates to test the effects of these metals on the germination and radicle elongation of two deciduous and five coniferous woody species native to eastern North America. For species grown on filter paper, germination was little affected by metal concentrations of up to 100 ppm. Filter-paper tests showed that radicle elongation was reduced by 25% at concentrations that ranged from 1 to 5 ppm for Betulapapyrifera Marsh, to as much as 50 to 100 ppm for Pinusstrobus L. Concentrations causing similar effects on mineral and organic soil substrates were, respectively, 10 and 100 times greater than those for filter paper. For all species, toxicity followed the pattern Ni 〉 Cu 〉 Co for filter paper and [Formula: see text] for mineral and organic soils. Deciduous species were more readily damaged by these metals than were coniferous species.

Description: Stem eccentricity in second-growth western hemlock (Tsugaheterophylla (Raf.) Sarg.) from southern coastal British Columbia appears to be the rule rather than the exception. Although the difference was small, the average eccentricity ratios (ratios of minimum diameter inside bark (DIB) to maximum DIB) for two Lower Mainland sites were significantly larger than those of two Vancouver Island sites. The average eccentricity ratio of all 87 trees studied was 0.929, with a standard deviation of 0.0231. The angular rotation of the major axis of the stem cross section varied greatly, and was positively related to the average stem eccentricity ratio within 5-m log lengths for each stem. Average eccentricity of the whole stem was relatively independent of diameter at breast height (DBH), age, stem height, and eccentricity at breast height. It does not appear that a practical field technique could be devised for the rapid estimation of average stem eccentricity in western hemlock.

Description: The vegetative phenology of Piceaglauca (Moench) Voss and P. mariana (Mill.) B.S.P. was determined in 1979 based on four ramets selected from each of 14 clones of each species located at a seed orchard in northern Ontario. The time of flushing of the leaders and four lateral branches was determined by a qualitative index of bud and shoot development; time of growth cessation was scored as the date at which 95% of shoot growth was complete. The flushing of the vegetative buds of white spruce lasted for a 10-day period compared with only 6 days for black spruce. Although the average date of flushing for white spruce clones was 9 days ahead of the average date for black spruce clones, the latest white spruce clone flushed only 3 days before the earliest black spruce clones. As well, degree-day requirements for flushing were significantly different for clones within each of the two species. These results suggest that the selection of late-flushing white spruce trees for seed orchard stock has the potential to decrease spring frost damage in this species in northern Ontario. Date of budbreak was not correlated with date of growth cessation for either white or black spruce; however, early-flushing clones of black spruce produced significantly greater leader extension than late-flushing clones.

Description: Ocular and shoot-count defoliation estimation techniques were compared for accuracy and precision at the levels of branches and crown position within trees, whole trees, plots, and stands. A shoot-count estimate of defoliation for a midcrown branch had the best relative accuracy (±7%), while the best ocular estimate was for the whole tree by an experienced observer (accuracy ±12%). Ocular estimates were biased towards ovcrestimation at low defoliation levels (error 20–30% defoliation), and previous defoliation caused the ovcrestimation of current defoliation by 30–40% on trees that had been previously severely defoliated. Observers were found to be consistent in their rating, but biased by about ±10%; experienced observers were about 5–10% more accurate than inexperienced observers. Intertree variance in defoliation was greater than intratree variance or variance between plots in a stand. It was concluded that ocular estimation of defoliation is a viable technique that can give accuracy within the limits required for surveys and many research applications, if the influence of observer experience, observer bias, and previous defoliation arc recognized, and adjusted for when necessary.

Description: The photosynthetic capacity of different needle age-classes of black spruce (Piceamariana (Mill) B.S.P.) was studied using 14C labelling on whole branches at a site near Fairbanks, Alaska. Black spruce had highest photosynthetic rates in the 1-year-old tissue with the four most current age-classes showing similar high rates before declining with age. Older needles were found to maintain 40% of maximum photosynthetic rates after 13 seasons. Highest level of nitrogen was found in current needles with the first eight age-classes maintaining high levels before declining to 70% of maximum values in the oldest needles. Phosphorus levels were much higher in current tissue with a gradual decline after the 1-year-old tissue to a value of 55% of maximum values in the oldest tissue. Nutrient use efficiency was found to be highest in the 1-year-old tissue and declined with needle age. The loss of nutrient use efficiency may be due to the use of older tissue as aboveground nutrient stores, a decline in the efficiency of the photosynthetic apparatus of the needles, or an increase in the leaf specific weight. Black spruce has adapted to the low nutrient availability with low photosynthetic rates and low nutrient contents. Greater longevity of the needles would maximize the photosynthetic return per unit nitrogen invested in the needles.

Description: Twenty-four specimens of six boreal tree species were excavated: eight Pinusbanksiana Lamb., five Populustremuloides Michx., four Piceaglauca (Moench) Voss, three Piceamariana (Mill.) B.S.P., two Abiesbalsamea (L.) Mills., and two Lara laricina (Du Roi) K. Koch. Root systems were described, and percent biomass determined for above- and below-ground components for half these specimens. Tree ages ranged from 3 to 120 years. Horizontally spreading lateral roots dominated the root systems of all species and occurred within 3– 15 cm of the ground surface. Nutrient and moisture absorption appear to be the principal functions of lateral roots. Maximum vertical root growth occurred near the tree stump as: tap-, heart, sinker, and oblique lateral roots. Vertical root morphology is influenced by site conditions. In addition to providing mechanical support, sinker and taproots may be important adaptations for deep-water utilization on xeric sites. Our data suggest that roots may be important in boreal succession processes through two related mechanisms: (i) nutrient and water deprivation occurring through preemptive growth of tree roots from climax species over roots of serai species, and through interception by mosses; and (ii) niche partitioning occurring below ground between serai and climax, and among climax tree species, by vertical separation of the root systems.

Description: The impact of Hurricane David on the forests of Dominica was measured 40 days after the hurricane struck the island on August 29, 1979, with winds averaging 92 km/h which lasted 10.2 h. Sixteen 0.1-ha plots were studied. They were located on a variety of slope and exposure conditions, representing three life zones and 11 plant associations. Measurements included tree diameter at breast height (dbh), total height, wood volume, and percent canopy closure of stand. Species and damage conditions were noted. The hurricane caused severe damage, particularly to forests on the southern half of the island. Forty-two percent of the standing timber volume in the study areas was damaged at an average intensity of 27 m3/ha. About 5.1 × I06 timber trees were damaged in an area of 246 km2. "Complex" vegetation was more resistant to hurricane damage than "simpler" vegetation. The climax Dacryodesexcelsa Vahl associations suffered the least hurricane damage, while the palm brake suffered the most damage. Only one species (Richeriagrandis Vahl) exhibited significantly higher hurricane damage. Trees in larger diameter classes tended to uproot with more frequency than smaller diameter trees, which tended to snap and break. Regrowth of seedlings and saplings was rapid on wet areas of the damaged sites. We found more species on regenerating plots than on the undisturbed forest plots. The damage caused by Hurricane David is the most extensive reported to date anywhere, and damage to vegetation by hurricanes appears to be more extensive than damage by earthquakes and landslides. Owing to their high frequencies in the Caribbean, hurricanes induce cyclic plant successions.

Description: A probabilistic two-step method of analysis is described that overcomes the problems associated with using a single model to analyze data drawn from a two-state system. The two steps of the analysis are (i) to determine the probability that an observation with given characteristics falls in each of the two states of the system and (ii) to estimate the response of the dependent variable in each of the two states as a function of characteristics of observations. Estimating cull volume in standing trees provides an example. This analysis results in two component models that more closely describe the process of interest than is possible with any single model. Statistically sound approximations to confidence intervals and tests of hypotheses for the parameters of each of the component models may be computed.

Description: Growth of diameter of individual trees can be expressed as diameter increment or basal area increment. Little work has been done to determine which of these parameters is preferable for use in growth studies. This paper examines growth of trees in pure and mixed stands of even-aged, regrowth forest of Eucalyptusregnans, E. obliqua, and E. globulus, aged 6-80 years, thinned and unthinned, in southeastern Tasmania. Weighted least squares regression equations are developed to relate diameter and basal area increments, over 1- to 6-year increment periods, to initial tree diameter in 29 growth plots. The correlation between basal area increment and initial diameter was always greater than that between diameter increment and initial diameter in these relationships. Despite this, the precision of estimates of future diameter in each plot is shown to be the same whether diameter or basal area increment equations are used. Data from a thinning experiment in 38- to 85-year-old, even-aged Acersaccharum, Fraxinusamericana, and Prunusserotina stands in southern Ontario are also examined and similar results obtained. It is concluded that no apriori reason exists for expressing growth as diameter increment or basal area increment in studies of these types.

Description: The effects of fertilization and thinning of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) on the distribution of area increment along the bole were assessed using radial growth measurements 6 and 9 years after treatment. Within treatments, the average area increment per tree was linearly related to diameter at breast height, and this relationship was used to evaluate the effects of treatment on growth rate. Fertilization had the greatest effect on average area increment, and for a particular fertilization regime, thinning increased the response. Thinning modified the distribution of growth over the bole of all trees and increased butt flare, especially in smaller trees. The effect declined from the 4- to 6-year measurement period to the 7-to 9-year measurement period. Fertilization had no consistent effect on growth distribution. The regression methods used in this study provided a more sensitive measure of form changes than previous methods, were independent of size distribution, and facilitated extrapolations and evaluation of temporal trend.

Description: The annual amounts of N, P, K, Ca, and Mg in litter-fall, throughfall, and stemflow were measured in a 30-year-old jack pine (Pinusbanksiana Lamb.) stand on a coarse glacial outwash soil in northern Ontario. Litter from ground vegetation and from the pine overstory was estimated. The nutrient content of precipitation was measured and the quantity of nutrients in leaf wash determined.Tree litter was the most important source of N, P, Ca, and Mg for the forest floor (51–69% of the total depending on the element), whereas throughfall supplied most K (54% of the total). Ground vegetation litter contributed significant amounts of nutrients (7–23% of the total depending on the element) but stemflow added little (1–8% of the total). Potassium in throughfall was derived mainly from leaf wash whereas N, P, Ca, and Mg in throughfall were derived primarily from precipitation entering the ecosystem. This jack pine forest floor received an annual total of 30 kg/ha of N, 22 kg/ha of Ca, 19 kg/ha of K, 3 kg/ha of Mg, and 2 kg/ha of P from the processes studied. Most of the nutrients in these totals were returning to the forest floor from the vegetation.

Description: The response of a 40-year-old even-aged jack pine (Pinusbanksiana Lamb.) stand to nitrogen fertilization was studied in four experiments.Smothering of the ericaceous ground vegetation layer with straw led to increased tree growth rates and improved N uptake. The application of calcium nitrate, ammonium nitrate, and urea at 112 and 448 kg N/ha was followed by analysis of the composition of extractable humus N, together with foliar and increment analyses. After a 3-year response period, there was evidence for the superiority of the ammonium nitrate form of N fertilizer over urea. The applications of urea appear to have been associated with possible N volatilization losses, and also inadequate urease activity in the thin dry humus layer which led to delayed N uptake by the trees. Calcium nitrate, although not an efficient fertilizer, was associated with appreciable growth responses, even though increased foliar and extractable humus N concentrations were not maintained.An optimum nutrition experiment utilizing repeated fertilization indicated that sustained foliar nutrient regimes can be maintained.The study indicated that growth increases were associated with increased unit needle weights as well as increased foliar N concentrations. A graphical technique which analyzes treatment responses, unit needle weight, and nutrient concentration, was used to screen response data, without waiting the customary 5-year period for stem increment response.

Description: Organic content of the forest floor decreases for several years after clear-cutting, and then slowly recovers. Thickness, bulk density, organic matter, and nitrogen content of forest floors were measured for 13 northern hardwood stands in the White Mountains of New Hampshire. Stands ranged from 1 to about 100 years in age. Forest-floor thickness varied significantly with stand age, but bulk density, organic fraction, and nitrogen fraction were independent of age. Total organic content of the forest floor agreed very well with data from Covington's (W. W. Covington 1981. Ecology, 62: 41–48) study of the same area. Both studies indicated that mature forest floors have about 80 Mg organic matter•ha−1 and 1.9 Mg nitrogen•ha−1. Within 10 or 15 years after cutting, the organic matter content of the floor decreases to 50 Mg•ha−1, and its nitrogen content to 1.1 Mg•ha−1. The question whether the decrease is rapid and the minimum broad and flat, or if the decrease is gradual and the minimum sharp, cannot be answered. The subsequent increase to levels reached in mature forest requires about 50 years. Some of the initial decrease in organic matter and nitrogen content of the forest floor may be caused by organic decomposition and nitrogen leaching, but mechanical and chemical mixing of floor into mineral soil, during and after the harvest operation, may also be important. The difference is vital with respect to maintenance of long-term productivity.

Description: A variety of evergreen and deciduous forests in the taiga of interior Alaska were studied over a 5-year period to examine how the chemical quality of forest-floor organic matter affected its rate of decomposition and mineral cycling within and outside the tree vegetation. Litterbag and respiration studies were used to monitor decomposition. Natural forest-floor substrates and others altered by addition of N, P, and K fertilizer and glucose as a carbon source were studied in the laboratory and field for rates of weight loss and O2 consumption. Forest floors differing in C/N ratios, including those deficient in N, were used to measure substrate quality influences on seedling growth, nutrient content, and tannin content. Microbial (bacteria and fungi) biomass was measured across a range of forest types along with pH, base saturation total pool sizes of N and P, and annual mineralization of organic matter per square metre. Under identical moisture and temperature conditions average respiration rates in evergreen forest-floor L, F, and H substrates were 1.8, 2.8, and 2.0 times less than in the corresponding deciduous forest horizons, respectively. Birch L and F horizons had respiration rates 11.5 times higher than the corresponding black spruce layers. Weight losses in birch L, F, and H horizons were 6, 3, and 2 times higher, respectively, than in the corresponding black spruce substrates. Substrates had a quality-dependent decay rate which did not change when they were relocated within or between sites indicating that measured field climatic differences were not as influential on decay rates as substrate quality components. Fungal biomass was significantly correlated with the quantity of organic matter in all sites (n = 15, r = 0.62) but correlations were better for deciduous (n = 9, r = 0.89), and evergreen (n = 6, r = 0.82) forests separately. Strong correlations exist also between grams of organic matter decayed per square metre per year and fungal biomass (n = 13, r = 0.86), and fungal biomass and grams of N and P mineralized per square metre per year (n = 14, r = 0.95) and (n = 11, r = 0.94, respectively). Seedlings on mineral-deficient substrates produced more tannins than the controls, and seedlings on substrates with widening C/N ratios had successively less tissue with lower N content, and proportionally more roots. Nitrogen content of litter fall in increasingly nitrogen-poor forest floors was correspondingly lower. Nitrogen content of litter fall on N rich forest floors and N fertilized forest floors was proportionately higher. Nitrogen withdrawal in leaves at senescence was inversely correlated with grams N mineralized per square metre per year in forest floors. Fertilization did not influence microbial processes in the field, though lab studies indicated a negative influence of NH4, P, and K on microbial respiration. Glucose added in the laboratory and field markedly increased forest-floor microbial respiration. In vitro glucose-induced increases in respiration were not influenced by addition of ammonium nitrate and were significantly depressed by addition of P and K. In the field, fertilization had no effect on either glucose-induced respiration or microbial biomass.

Description: An old, merchantable, low-site class black spruce (Piceamariana (Mill.) B.S.P.) stand growing on a Lithic Humo-Ferric Podzol overlying a granite bedrock, and a younger but merchantable low-site class jack pine (Pinusbanksiana Lamb.) stand growing on a Ferro-Humic Podzol overlying a deep coarse sand near Baie Comeau, P.Q., were analysed for stand biomass and macronutrient contents of both stand and soil. The magnitude of the depletions of macronutrients from the site, in full-tree and tree-length methods of logging, are compared with their available and total quantities in the soil. The range of values from the literature for nutrient inputs are presented and discussed in relation to logging losses. The results suggest that full-tree logging in the dry jack pine stand could cause a severe loss of potentially mineralizable N supply; P, K, and Mg deficiency problems owing to export are not apparent, but a Ca balance problem is apparent. The implication is that full-tree logging should be avoided on such sites.

Description: Two perennial problems in the management of uneven-aged forests are considered: (i) determination of the optimal sustainable distribution of trees by diameter class, i.e. stand structure, for a given initial stocking level, and (ii) the optimal cutting schedule for the conversion of an irregular stand to a target structure. It is shown, using examples for northern hardwood stands in Wisconsin, that both problems can be solved via mathematical programming techniques. The programming approaches utilize a set of nonlinear equation models for stand table projections which consider the interdependence of size classes within the stand. To illustrate procedures, optimal stand structures are found for a case where initial stand basal area is constrained to specified levels and the objective is to maximize value growth over the cutting cycle. A conversion cutting schedule is then determined for a case in which the objective is maximization of present worth. It is emphasized that both the optimal distribution and conversion problems can be generalized to consider a broad range of objective functions, lengths of cutting cycle, and constraints on the growing stock.

Description: Using data on CO2 exchange, growth, and biochemical composition, an approach was developed to calculate the seasonal production, consumption, and surplus of photosynthate in shoots of loblolly pine (Pinustaeda L.). The amounts of CO2 released and of glucose substrate required in biosynthesis of shoot biomass over the growing season were also calculated. Individual growth flushes of different age-classes, as well as of an entire 1-year-old branch unit, ofPinustaeda were used as bases for calculations.Substrate requirements and CO2 production were relatively constant throughout the growing season. Within the whole 1-year-old branch unit, 63% of the photosynthate production was supplied by the two growth flushes of the previous year. Growth of the current year's shoot represented the major consumption of photosynthate, and this made up 66% of the total photosynthate consumed in the branch unit. Surplus of photosynthate, which would be available for translocation from the branch, was about 38% of the total photosynthate produced. Most of the photosynthate surplus was derived from the growth flushes of the previous year. Source and sink relationships, with regard to photosynthate partitioning in the 1-year-old branch unit over the growing season, were clarified and described.

Description: Fertilization and cultivation before and after planting greatly increased 9-year growth and yield of longleaf (Pinuspalustris), slash (P. elliottii), and loblolly pines (P. taeda). Intensive culture did not decrease specific gravity or fiber length of the juvenile wood. In loblolly pine, genetic variation was found in response to intensive culture.

Description: Seven units (about 2 ha each) of black spruce – feather moss forest were experimentally burned over a range of fuel moisture conditions during the summer of 1978. Surface woody fuels were sparse and the principal carrier fuel was the forest floor (largely mosses and their decomposition products). Forest floors after burning comprised a small-scale mosaic of unburned, scorched, lightly burned, moderately burned, and heavily burned (organic materials entirely consumed) conditions. Percentage of the unit area in the moderately and heavily burned condition ranged from 11.2 to 77.2% and percent decrease in forest-floor thickness varied from 27.4 to 63.1% in the seven units. Forest-floor consumption was most closely correlated with the moisture content of lower moss (01 horizon) and lower duff layers (022 horizon) at the time of burning. For the first 3 years after fire, biomass production was greater on heavily burned than on lightly burned sites (58 vs. 37 g/m2 on an annual basis). Heavily burned sites were completely dominated by the invading species Epilobiumangustifolium L., Ceratodonpurpureus (Hedw.) Brid., and Marchantiapolymorpha L., whereas lightly burned plots were occupied by sprouting species such as Calamagrostiscanadensis (Michx.) Beauv., Vacciniumuliginosum L., and Ledumgroenlandicum Oeder. Soil pH and amounts of total P and available P in the forest floor increased significantly as a result of burning; and in all cases, increases reached a maximum in moderately and heavily burned areas. Total N in the forest floor increased significantly in moderately burned, but decreased slightly in heavily burned areas. Total N and total P showed smaller increases in the surface mineral soil as a result of burning. Supplies of available P in the mineral soil increased almost 4-fold in moderately burned and over 16-fold in heavily burned areas.

Description: Straw was added over snow to smother ground vegetation (straw experiment) in sample plots in a 45-year-old jack pine (Pinusbanksiana Lamb.) stand in Quebec. Sample plots in the same stand were fertilized six times with N, P, and K in a 10-year period to maintain four foliar N regimes (optimum nutrition experiment). Smothering of ground vegetation with straw improved N nutrition and produced a sustained increase in tree growth. Sustained growth increases were obtained by repeated applications of 56 kg N/ha associated with 1.4% N foliar concentrations in current foliage. Gross volume increments of about 3 m3 • ha−1 • year−1 were sustained with these low applications. Repeated heavy N applications killed trees and reduced growth. Repeated additions of P and K with N did not produce appreciable differences in response from additions of N alone. Increment cores showed the annual development in growth as increasing over most of the 10-year period for low repeated N additions, and as increased and stable over the same period for the straw addition.

Description: White oak (Quercusalba L.) seedlings grown in a silt loam soil inoculated with Pisolithustinctorius (Pers.) Coker and Couch exhibited 80% ectomycorrhizal development after a 6-month establishment period whereas the noninoculated controls exhibited less than 5%.When water was withheld and soil water potential decreased as the soil dried, xylem pressure potential and leaf conductance of both the inoculated and noninoculated seedlings declined gradually. At the peak of the drying cycle, the inoculated seedlings exhibited mean prelight and midlight xylem pressure potential values which were 0.2 MPa and 0.15 MPa, respectively, more negative than the noninoculated seedlings. Although the inoculated seedlings had a mean leaf surface area 1.5 times larger than that of the noninoculated seedlings, there were no significant differences in leaf conductance at the peak of the dehydration cycle. Mean rates of root elongation were greater among the inoculated seedlings during the drying cycle. Following reirrigation of the soil, the inoculated seedlings yielded significantly less negative values of prelight and midlight xylem pressure potential and greater leaf conductance when compared to the noninoculated seedlings.

Description: Radial stem wood sections of fertilized (at age 14 years) and unfertilized loblolly pine (Pinustaecla L.) trees were analyzed 5 and 9 years after treatment to establish if xylem nitrogen (N) can be used for a diagnostic nitrogen test. The percentage of xylem nitrogen in wood formed since fertilization was greater in fertilized than in unfertilized trees. The 2 years of wood growth near the cambium had a higher percent N than older wood, but differences between treatments were less pronounced than in older wood. Concentrations of N for fertilized and unfertilized trees were nearly constant for 2- to 9-year-old wood indicating that a sampling technique to determine the N status of loblolly pine could be developed by increment coring.

Description: Allometric regressions for estimating red maple (Acerrubrum L.) biomass did not differ by stand age and site index. Significant differences were found only in terms of error variance, not in terms of regression slope or intercept. These trends suggest a single predictive model is valid for regional estimates. More attention should be given to regional sampling procedures and predictive relations that are valid for a wide range of stand and site conditions in developing weight tables and conducting biomass inventories.

Description: A factorial experiment was conducted in which Pinuscontorta var. latifolia and Piceaglauca (Moench) Voss seedlings were grown for 6 weeks under controlled environmental conditions with deficient and normal supplies of nitrate and ammonium. Free sugars (sucrose, fructose and glucose), starch (as glucose) and pectic sugars (galactose, arabinose, mannose and xylose) were quantitated. Effects of nitrogen form and nitrogen level, and their interactions, upon sugar contents and dry weights of the two species were examined. Growth responses to nitrogen occurred mainly in the shoots. For most parameters the two species reacted similarly to changes in nitrogen level and form, although the concentration of individual sugars differed between the species. In the shoots there were level × species interactions in the dry weight and the sum of free sugars plus starch. Plants grown under nitrogen deficiency were characterized by higher starch contents throughout and lower free sugars in the shoots. Supplying nitrate instead of ammonium lowered glucose levels, particularly in the shoots. Pectic sugar contents were unaffected by nitrogen nutrition. These compositional and growth responses to the nutritional regime implied changes in carbohydrate metabolism and physiological processes that are described by the C/N ratio of the plant.

Description: In 4 different years' experiments with loblolly (Pinustaeda L.), slash (Pinuselliottiielliottii Engelm.), and longleaf (Pinuspalustris Mill.) pine, timing of gibberellin A4/7 (GA4/7) treatments was critical for optimal promotion of pollen conebuds. Two or three biweekly treatments given at strategic times were equivalent to six given from May to August. July to August applications best promoted pollen conebuds. Using the cationic surfactant Aromox C/12, aqueous foliar sprays of 200 mg/L of GA4/7 were more effective than ethanolic topical bud treatments for pollen conebud induction in slash pine. Addition of naphthaleneacetic acid enhanced the GA4/7 effect in loblolly pine but diminished it in slash and longleaf pine.

Description: The relationships of root attributes and tree, stand, and site factors to root contact between second-growth Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) were studied in excavated root systems. Type 1 contacts (tenuous) were positively related to root length and inversely related to root volume. Type 2–3 contacts (with slight to severe root deformation) were positively related to number of roots. Type 4 contacts (apparent grafts) were positively related to root number and cross-sectional area. Differentiation among groups containing no contacts. Type 1 contacts only or all contact types showed that intertree distance, tree diameter, rooting depth, soil gravel content, and percent slope were significant in determining probability of root contact.

Description: Decomposition rates and changes in the nutrient content of needle and leaf litter were examined in Douglas-fir (Pseudotsugamenziesii Mirb. Franco), western hemlock (Tsugaheterophylla (Raf.) Sarg.), Pacific silver fir (Abiesamabilis (Dougl.) Forbes), and red alder (Alnusrubra Bong.) ecosystems in western Washington, U.S.A. Nylon litterbags (1-mm mesh) were placed in the stands in November and December 1974. Bags were collected after 3, 6, 12, and 24 months and weighed, except in the Pacific silver fir stand when bags were collected after 6, 9, 14, and 24 months. Litter was analyzed for C, N, P, K, Ca, Mg, Mn, lignin, and cellulose. Decomposition constants (k values) were determined. Fastest decomposition after 2 years occurred in red alder leaves, followed by Douglas-fir, western hemlock, and Pacific silver fir needles. There were significant differences in weight loss among species after 1 year, but no significant differences were evident after 2 years. Red alder leaves showed rapid weight loss in the 1st year but decomposed little in the 2nd year. Decomposition constants were highly positively correlated with minimum air temperatures and negatively correlated with C:N ratios. Low litter moisture tended to reduce decomposition in summer, particularly in the Pacific silver fir stand. Decomposition proceeded under snow in this ecosystem. The pattern of loss of elements from litterbags after 2 years varied from ecosystem to ecosystem, particularly for N. The following element mobility series resulted for the four ecosystems: red alder (K 〉 Mg 〉 Ca 〉 P 〉 N 〉 Mn), Douglas-fir (K 〉 P 〉 Ca 〉 Mg 〉 Mn 〉 N), western hemlock (K 〉 Ca 〉 Mg 〉 N 〉 Mn 〉 P), and Pacific silver fir (K 〉 Mg 〉 Ca 〉 Mn 〉 P 〉 N).

Description: The basic assumptions of regression analysis are recalled with special reference to the use of a logarithmic transformation. The limitations imposed on inference-making by failure to comply with these assumptions are discussed and ways to avoid the limitations indicated. A systematic bias of the order of 10 to 20% which is inherent in most, if not all, prior uses of the logarithmic equation to estimate plant biomass is noted as is the correction for the bias.

Description: Root and shoot biomass and mycorrhizal development were examined for white spruce (Piceaglauca (Moench) Voss) seedlings naturally regenerating in four floodplain communities in the boreal forest. Mean seedling biomass was highest in the open community and lowest in the spruce community. Seedlings growing in the open community had higher root:shoot ratios (0.50) compared with seedlings growing in the willow (0.34), alder (0.20), and spruce (0.24) communities. Essentially all short roots of spruce seedlings growing in all four communities were infected by mycorrhizal fungi throughout the growing season.

Description: Growth response of young, spaced balsam fir (Abiesbalsamea (L.) Mill.) to 3 successive years (1979–1981) of treatment with Bacillusthuringiensis Berliner for spruce budworm (Choristoneurafumiferana (Clem.)) control was examined in 20 plots on the Cape Breton Highlands, Nova Scotia. Defoliation commenced in 1976, 3 years before control operations began. Five plots were established both inside and outside the spray block, in areas that had been severely defoliated by budworm and in areas moderately defoliated. All plots in the severely defoliated area suffered heavy tree mortality, but mortality was significantly lower in the protected plots. In contrast, the moderately defoliated plots suffered virtually no tree mortality. The average volume increment of stem-analyzed trees from 1979 to 1981 was 0.63 dm3/tree for protected and 0.43 dm3/tree for unprotected trees in the severely defoliated area versus 4.15 dm3/tree for protected and 3.08 dm3/tree for unprotected trees in the moderately defoliated area. These volume increment values are equivalent to 8.8 m3/ha of growth between 1979 and 1981 for protected plots in the moderately defoliated area, compared with 6.1 m3/ha for unprotected plots. Therefore, a total gain of 2.7 m3/ha can be attributed to the 3 years of B. thuringiensis spraying. Long-term growth responses toB. thuringiensis spraying were not evaluated. Better results would be expected had protection started at the beginning of the budworm outbreak, instead of after 3 years of severe defoliation.

Description: Constructing a dynamic model of growth in fir stands (age 30–70 years), based on gross physiological structure of the stand, made possible reasonable qualitative and quantitative forecasts of growth loss due to defoliation by insects in a wide variety of situations. Analysis of the model, in comparison with the natural world, indicates that the key to understanding growth loss in fir stands subjected to defoliation by spruce budworm lies in more explicit knowledge of the mechanisms of new foliage production.

Description: Calculations are presented on the quantities of N, P, K, Mg and Ca in the soil and above-ground portions of two spruce-pulpwood stands on sites of average fertility in northern and southern Quebec. The magnitude of the depletions of these nutrients from the site, in full-tree and tree-length methods of logging, are compared with the available and total quantities of them in the soil. The ranges of values, from the literature, for the input of these nutrients in dust and precipitation, and the losses in leaching, are presented and discussed in relation to the logging losses.It is concluded that on both sites it is unlikely that full-tree logging will result in any reduction in growth, due to nutrient removal, during the second rotation of trees. However, nutrient depletion due to full-tree logging, particularly with respect to Ca, K and N, may require correction by means of fertilizers in forest ecosystems of marginal fertility. These are usually either dry sites with low reserves of organic matter and low exchange capacity or wet sites with excessive accumulations of organic matter. The need for further detailed studies of the nutrient cycle in different forest ecosystems is stressed.

Description: The Weibull distribution, [Formula: see text], summarized diameter, basal area, surface area, biomass, and crown profile distribution data well for several different ages of white and loblolly pine plantations. The data for diameter, basal area, surface area, and biomass were easily summarized by this one distribution in a theoretically consistent fashion. This is not possible with the normal and the gamma distributions, and the lognormal gives less satisfactory results. The distribution function should prove useful in modeling tree stands since only the parameter values need to be changed over time for the above variables. The change in these parameters may be a good way to characterize and interpret changes in stands over time.

Description: Seed size was examined as a possible explanation for variation in the size of containerized loblolly pine (Pinustaeda L.) seedlings. Under laboratory conditions of minimal environmental stress, larger seed were found to germinate more quickly and produce a larger germinant after 28 days of growth. In a similar experiment conducted under greenhouse growing conditions, large seed produced the largest seedlings. Size differences among seedlings in both experiments resulted from differences in the rate of germination unique to each seed size class. Consequently, seedling size and possibly uniformity of growth were considered to be a function of germination patterns which were strongly influenced by seed size and weight.

Description: A 24-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stand was treated with various levels and combinations of nitrogen fertilization and thinning. Over a period of 5–9 years after treatments, trees were sampled to determine effects on foliage quantity and sapwood characteristics at different stem heights together with their relationships. Sapwood width remained relatively constant up the stem where heartwood was present, but the number of annual rings it contained decreased with height. The sapwood width at breast height (bh) increased with stem diameter; treatments had little effect on percent sapwood at bh. The ratio of foliage mass to sapwood cross-sectional area changed for different portions of the crown and was lower when based on sapwood area at bh than at base of live crown. Significant linear relationships of foliage mass and area to sapwood area at bh were found, but relationships of foliage to basal area (bh) were just as close for all treatments; treatments significantly affected these relationships with control trees having the lowest regression slopes.

Description: The allocation of biomass and nutrients (N, P, K, Ca, Mg) was measured from August 1976 to September 1978 in a young, second-growth Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stand in the Oregon Coast Range. Tree biomass comprised 78–79% of the total standing crop of organic matter with the remainder allocated as follows: soil organic matter, 17%; forest floor, 4%; and fungi, 2%. Bole biomass accounted for 64–66% of the total tree standing crop; the remainder was apportioned among: nonmycorrhizal roots, 17–18%; branches, 7–8%; mycorrhizae, 6%; and foliage, 4%. Nutrient stocks in aboveground tree components exceeded those in belowground components by one to nine times. For all nutrients except Ca, roots and mycorrhizae contained larger stocks than either the forest floor or fungi; amounts of Ca in the forest floor and in fungi were twice those in roots and mycorrhizae. Return of organic matter to the soil by fine roots and mycorrhizae ranged from 84 to 78% of total tree return. About 73% of total net primary production was invested in growth and maintenance of roots and mycorrhizae. Return of N, P, and K to the soil by mycorrhizae comprised 83–87% of total tree return and 25–51% of Ca and Mg return. Return by mycorrhizae of N, P, and K was four to five times greater than that of roots, nearly equal for Ca, and three times less for Mg.

Description: Black alder, Alnusglutinosa (L.) Gaertn., seedlings were grown and kept well watered for 10 weeks, and then subjected to moisture stress conditioning for 5 subsequent weeks, where one-half of the seedlings were watered only when visibly wilted. The remaining seedlings (controls) were kept well watered. Moisture stress conditioning greatly reduced shoot, root, nodule, and total plant dry weight. The root–shoot ratio (grams/grams) of seedlings was significantly increased from 0.28 in the control seedlings to 0.33 in the water-stressed treatment. Acetylene reduction rates decreased only slightly in the range of water potentials between −0.50 and −1.29 MPa, then dropped rapidly below water potentials of −1.30 MPa. Moisture stress conditioning had no significant influence on this response. Although not significantly different, leaf and nodule osmotic potentials were consistently lower in the water-stressed plants.

Description: The relationships between foliage area and sapwood area between trees and within the crowns of 20 Piceasitchensis (Bong.) Carr., provenance Queen Charlotte Island, British Columbia (10 in a control plot and 10 in a plot fertilized with potassium and phosphorus 8 years before harvest) and 10 Pinuscontorta Dougl., provenance Ladysmith trees were examined using a physiological analysis based on Darcy's law. Foliage area index on the fertilized P. sitchensis plot was higher than on the control. The variation of foliage area density with depth in the canopies followed a normal distribution. Relationships between foliage area and sapwood basal area were linear but the slopes were different for the two species. There was no significant difference between the control and fertilized P. sitchensis trees. The relationship between foliage area and the product of sapwood area and permeability was linear and data from the three plots fell on the same line. Sapwood area, permeability, and their product decreased with depth through the crowns of the trees. Within the crowns, relationships between cumulative foliage area and sapwood area, and between cumulative foliage area and the product of sapwood area × permeability were different with species and treatment. A single linear relationship resulted when the product of cumulative foliage area above an internode × the internode length was plotted against sapwood area × permeability for the internode. This suggests that it is the drop in potential across a node and internode rather than the gradient of potential across the internode that is related to the flux of water through tree crowns. The data support the hypothesis that the relationship between foliage area and sapwood area depends on permeability of the sapwood and the local climate through its influence on transpiration rate, particularly via average water vapour pressure deficit of the air and stomatal conductance.

Description: A procedure is described for the separation of nine organic matter fractions from F and H horizons of forest floor materials formed under different vegetative cover. The extracts employed (and fractions obtained) were ethanol:benzene (A), dilute mineral acid (B), cold 0.1 N NaOH (H-1, F-1, F-2), and hot NaOH (H-2, F-3, F-4), with unextracted organic matter comprising the final fraction. Distribution between the fractions of carbon, and polysaccharides was investigated. Considerable variation was observed in the distribution of fractions among sites, and both between and within horizons. The significance of the fractions is discussed. It was concluded that the procedure was suitable for studying differences in distribution of organic fractions of forest floor materials, but required further modification for examination of the composition of fulvic fractions.

Description: The root density and depth patterns of four boreal forest age sequences were analyzed for successional trends. Rooting depths increased with age on sandy substrates which supported aspen (Populustremuloides Michx.) and jack pine (Pinusbanksiana Lamb.) communities. Rooting depth did not change in an aspen series on fine-textured substrates or in a black spruce (Piceamariana (Mill.) BSP) series growing on organic substrates. Plant communities growing on mineral soils showed a decrease in near-surface root densities and understory vascular plant cover with increasing age. Maximum rooting was deepest on sandy substrates and shallowest on organic deposits. Roots in all stands were concentrated near the ground surface. In most cases 50% of all roots were located within 15 cm of the forest floor. Root densities in this zone ranged from 11 000 to 30 000 roots m−2 of vertical surface. Densities were approximately 4300 roots m−2 for the overall rooting zone.

Description: Analysis of dead boles of Piceasitchensis (Bong.) Carr. and Tsugaheterophylla (Raf.) Sarg. in open- and closed-canopy forests of the Olympic Peninsula Washington, U.S.A., revealed that hemlock mortality in both forest types was due mainly to windthrow, whereas spruce typically died upright. The open forest contained 120 t/ha of dead bole wood; the closed forest contained 161 t/ha. Hemlock boles decayed more rapidly than the larger spruce boles, although both showed considerable variability. On a per-hectare basis, 146–223 kg of N, 147–197 kg of Ca. 39–61 kg of K, 18–29 kg of Mg, 6–14 kg of Na, and 17–29 kg of P were contained in dead boles of the open- and closed-canopy forests, respectively. Except for N and Mg, the nutrient concentrations of the wood were not significantly different after 33–68 years of bole decay. The N:P ratios increased with increasing decay for both species.

Description: Height and diameter growth to age three and the number of first year branches were analyzed for 25 cottonwood clones grown in six plantations in central Mississippi. Plantations were on two contrasting sites and planted in three consecutive years. Results showed small clone × planting year interactions and large clone × site interactions for all parameters. In the lower Mississippi Valley, therefore, emphasis should be placed on testing over a range of sites rather than replicating over time.

Description: The biomass of 76 jack pine (Pinusbanksiana Lamb.) trees (29 stands) and 72 red pine (P. resinosa Ait.) trees (28 stands) from throughout the northern Lake States was determined. All trees were from even-aged, unthinned plantations ranging from 20–61 years old; site indexes represented nearly the complete range for these species. Individual tree component weights (foliage, live branches, dead branches, stem wood, and stem bark) were regressed against dbh and tree height using the nonlinear form Bt = aDbHc. Stand biomass was also estimated with stand basal area and mean height of dominant and codominant trees using the equation form Bs = a + b(B) + c(Hs) or a + b(B)(Hs). The equations were tested in two additional stands of red pine and two of jack pine and by comparison with literature values. Individual tree equations were most accurate for estimating bole components and the total tree and less accurate for foliage and branches. The standard error of the estimate divided by mean weight ranged from 0.06 to 0.17 for bole components, from 0.21 to 0.28 for live crown components, and from 0.43 to 0.49 for dead branches. For all components, jack pine equations were slightly less precise than those for red pine. The individual tree equations appear to be applicable over a wide geographical area and usable for both natural stands and plantations. The equations appear to be valid for the majority of unthinned stands in the age range of 20–50 years. The stand equations, while less precise than individual tree estimates, should give reasonably accurate estimates of stand biomass components in most situations.

Description: Research was conducted to study (1) seasonal changes in tissue composition and (2) dynamic allocation of photosynthate to different biochemical fractions in growing shoots of the current and previous years of 15-year-old loblolly pine (Pinustaeda L.). As the growing season progressed, the shoots increased rapidly in total biomass; however, old needles formed in the previous year lost dry weight. The most striking biochemical change was a rapid decrease in starch content of old needles early in the growing season. Soluble sugars and organic acids in old needles also decreased slightly. Contents of biochemical constituents generally increased in current axes and needles as the organs grew and increased in weight, although there were decreases in organic acid contents of current axes early in the growing season. Formation of nonlabile major constituents (cellulose, hemicelluloses, lignin, and protein) in current-year shoots proceeded at different rates. These rates were interpreted in terms of aspects of shoot growth and development. Seasonal allocation of photosynthates into biochemical constituent fractions in current year growth flushes was calculated. Results suggested that photosynthates were allocated to constituents of different functional categories (metabolism, storage, structure, and protection) at different times of the growing season in such a way that structural growth was of first priority. Thus, allocation of photosynthates to structural growth early in the season enables branches to compete for favorable growing conditions.

Description: This paper is a comprehensive review of operational research studies in forest fire management during the years 1961 through 1981. It includes a brief discussion of fire management decision making, summaries of and comments regarding the practical merits of the work that has been done, and suggestions concerning future efforts in this field.

Description: Balsam woolly aphids were reared on young grand fir trees maintained in controlled environment or outdoors. Solutions of various nitrogen fertilizers were applied repeatedly to the trees and the effects on aphids and bark amino acids were studied. On trees fertilized with ammonium nitrate or a sequence of compounds, aphid population growth was less than on trees fertilized with urea or potassium ammonium nitrate. On urea-fertilized trees, populations multiplied 16.5 times in five generations, compared with a 5.7-fold increase on unfertilized trees and a 1.4-fold increase on ammonium nitrate-fertilized trees. These differences resulted from effects of these nutrients on aphid life history. Urea promoted aphid establishment and reproduction, whereas ammonium nitrate affected these processes, as well as survival, adversely. The highest establishment and survival rates resulted from potassium ammonium nitrate.These differences may be related to fertilizer-induced changes in the amino acid diet of the aphid. Thin layer electrophoresis and chromatography revealed that urea, ammonium nitrate, and calcium nitrate each increased arginine concentrations in the bark, the highest levels resulting from ammonium nitrate. Traces of phenylalanine and asparagine, found in other treatments, were absent from trees fertilized with ammonium nitrate during June.

Description: Retranslocation from leaves was investigated as a means of retaining nutrients in stands of Quercusprinus L. on two sites in southern Illinois, where wind rapidly moves litter downslope. Foliage samples were collected from late summer until leaf fall to describe the trends of leaf dry weight and nutrient concentration (N, K, P, Ca) changes. Free-falling rain and throughfall were collected to estimate foliar leaching. Foliar concentrations of N, K, and P decreased markedly during senescence while Ca concentrations increased. The pattern of concentration change was unique for each element, and the change in N concentration was closely correlated with change in leaf color. For the study sites as a whole, leaf dry weight decreased to 70% of the original value. Of 84.2 kg N/ha in green foliage, only 22.6% was returned to the site as litter. The canopy gained 0.3 kg N/ha (0.4%) from rainfall. The 78.5% N unaccounted for is attributed to retranslocation. Similarly, from 51.4 kg K/ha; 8.2, P; and 47.8, Ca in green foliage, 9.8, 1.3, and 3.3%, respectively, were removed by leaching; 27.4, 43.7, and 85.1% were returned to the site in litter. The remaining 63.0% K, 55.0% P, and 11.5% Ca unaccounted for is attributed to retranslocation. Retranslocation and leaching of nutrients was greater on the site of higher quality. But on both sites it appears that retranslocation is an important means of retaining and conserving N, K, and P countering the effect of annual litter removal.

Description: Mathematical growth analysis techniques were used to assess the effects of atmospheric carbon dioxide enrichment on growth and biomass partitioning of Liquidambarstyraciflua L. (sweetgum) and Pinustaeda L. (loblolly pine) seedlings. Plants were grown from seed under high (1000 μmol•m−2•s−1) and low (250 μmol•m−2•s−1) photosynthetic photon flux density at CO2 concentrations of 350, 675, and 1000 μL•L−1 for 84 or 112–113 days. Elevated atmospheric CO2 concentration significantly increased height, leaf area, basal stem diameter, and total dry weight of sweetgum seedlings grown under high irradiance and to a lesser extent under low irradiance. Increases in dry matter accumulation were associated with early CO2 enhancement of net assimilation rate, but increases in amount of leaf surface area contributed more towards maintenance of larger size as seedlings aged. For sweetgum seedlings in particular, reduction of growth by low irradiance under normal atmospheric CO2 was compensated for by growing plants with elevated CO2. In contrast, elevated CO2 concentration produced no significant increase in growth of loblolly pine seedlings.

Description: Seedlings of six Alaskan taiga tree species and one tall shrub were grown in sand at three phosphate levels. There was a positive correlation between the growth rate of a species at the high-phosphate level in sand culture and its productivity in the natural environment. Poplar (Populusbalsamifera L.), which had highest growth rate under high phosphate, was most sensitive to reduction in phosphate supply, followed by birch (Betulapapyrifera (Reg.) Fern, and Raup) and aspen (Populustremuloides Michx.), whereas growth of conifers (larch (Larixlaricina (Du Roi) K. Koch), white spruce (Piceaglauca (Moench) Voss), and black spruce (P. mariana (Mill.) B.S.P.)) from late successional sites was slow and unaffected by phosphate supply. Similarly, when birch and white spruce seedlings were transplanted into natural forest stands, the maximum growth rate of birch was greater than that of white spruce, but birch growth was curtailed more by unfavorable conditions than was that of white spruce. We conclude that a slow growth rate reduces nutrient requirement and therefore minimizes nutrient stress on infertile sites, whereas a rapid growth enables nutrient-demanding species to dominate fertile sites.

Description: Annual net N mineralization in the 0–10 cm mineral soil zone of nine forest stands on silt–loam soils was measured using a series of insitu soil incubations from April 1980 through April 1981. Differences in soil organic matter (SOM) dynamics among sites were shown with net N mineralization ranging from 0.54 to 2.10 mg N mineralized•g SOM−1•year−1. This variation was not related to percent N in SOM. Net N mineralization varied seasonally with maximum rates in June and very low rates in winter. Nitrification rates were constant from May through September despite fluctuations in soil ammonium pools. Nitrification was greater than 50% of annual net N mineralization at all sites. N uptake by vegetation, as estimated by net N mineralization plus mineral N inputs via precipitation, with minor corrections for mineralization below the incubation depth and for mineral N losses to groundwater, ranged from 40.3 to 119.2 kg N•ha−1•year−1. Annual leaf and needle litter production ranged from 2.12 to 4.17 Mg•ha−1•year−1 and was strongly correlated with N uptake (r = 0.938, P 

Description: The author describes a stochastic model of forest stand rotation which can be used to determine the optimal planned rotation interval for flammable forest stands. The model can also be used to estimate the value of fire management activities in terms of the potential enhanced value of timber production. The use of the model is illustrated by applying it to a simplified case of jack pine (Pinusbanksiana, Lamb.) management.

Description: Vegetation, forest productivity, and soils of 23 forest stands in the taiga of interior Alaska are described. The stands are arranged on an environmental gradient from an aspen (Populustremuloides Michx.) stand on a dry, steep south-facing bluff, to open black spruce (Piceamariana (Mill.) B.S.P.) stands underlain by permafrost on north-facing slopes. The coldest site is a mixed white spruce (Piceaglauca (Moench) Voss) and black spruce woodland at the treeline. Mesic upland sites are represented by successional stands of paper birch (Betulapapyrifera Marsh.) and aspen, and highly productive stands of white spruce. Several floodplain stands represent the successional sequence from productive balsam poplar (Populusbalsamifera L.) and white spruce to black spruce stands underlain by permafrost on the older terraces. The environmental gradient is described by using two soil factors: soil moisture and annual accumulated soil degree days (SDD), which range from 2217 SDD for the warmest aspen stand to 480 SDD for the coldest permafrost-dominated black spruce site. Soils vary from Alfie Cryochrepts on most of the mesic sites to Histic Pergelic Cryochrepts on the colder sites underlain by permafrost. A typical soil profile is described for each major forest type. A black spruce stand on permafrost has the lowest tree standing crop (15806 g•m−2) and annual productivity (56 g•m−2•year−1) whereas a mature white spruce stand has the largest tree standing crop (24 577 g•m−2) and an annual productivity of 540 g•m−2•year−1, but the successional balsam poplar stand on flood plain alluvium has the highest annual tree increment (952 g•m−2•year−1). The study supports the hypothesis that black spruce is a nutrient poor, unproductive forest type and that its low productivity is primarily the result of low soil temperature and high soil moisture.

Description: Biomass distribution and above- and below-ground net primary production were determined for 23- and 180-year-old Abiesamabilis (Dougl.) Forbes ecosystems growing at 1200-m elevation in the western Washington Cascade Range. Total organic matter accumulations were 427.0 t•ha−1 in the young stand, and 1247.1 t•ha−1 in the mature stand. Aboveground tree and detritus biomass were 49.0 t•ha−1 and 130.2 t•ha−1, respectively, in the young stand compared with 445.5 t•ha−1 and 389.4 t•ha−1 in the mature stand. Net primary production (NPP) was 18.3 t•ha−1 in the young stand and 16.8 t•ha−1 in the mature stand. Belowground dry matter production was 65% of total net production in the young stand and 73% of total net production in the mature stand. Conifer fine root production was 35.9% of NPP in the young and 66.4% of NPP in the mature stand. This apparent shift in fine root production as a proportion of NPP may be related to detritus accumulation.

Description: The effects of self-pollination in white spruce, Piceaglauca (Moench) Voss, were examined in 20 trees in three populations. The most drastic effect is the increased frequency of empty seeds which averages over 90%, and is 5 to 22 times higher than that from comparable cross-pollinations. The average numbers of embryonic lethal equivalents, although highly variable for individual trees, averaged 8.0, 9.1, and 12.9 for the three populations. Self-pollination also resulted in slightly lower germination and reduced survival. The total number of lethal equivalents affecting seed yield, germination, and survival to age 17 years is estimated as 12.6. The effect of selfing on growth becomes evident at age 2 years and increases to age 17 years when seedlings from self-pollinations averaged 44.5% shorter and 63.7% smaller diameter than seedlings from unrelated matings.

Description: An individual tree based stand growth model was analyzed, via dynamic programming, for thinning and final harvest strategies leading to maximum physical yields. Optimal density yield tables, indicating maximum yields and corresponding strategies, were constructed from the solution of the dynamic programming networks. Maximum yields averaged 25 – 30% higher than comparable yields reported previously for periodically thinned red pine plantations.

Description: The influence of mechanical and architectural properties of trees on growth rates, mortality rates, and relative probabilities of snapping and uprooting were examined on Barro Colorado Island, Republic of Panama. Of 310 fallen trees, 70% snapped, 25% uprooted, and 5% broke off at ground level. Stepwise discriminant analysis between snapped and uprooted trees indicated that of the variables measured, wood properties were the most important factors determining the type of death in trees. Uprooted trees tended to be larger, shorter for a given stem diameter, and to have denser, stiffer, and stronger wood than snapped trees. There were no significant differences between trees that snapped and trees that uprooted in the extent of buttress development or in the slope of the ground upon which they grew. Trees with low density wood grew faster in stem diameter than those with high density wood but also suffered higher mortality rates. After damage, many of the snapped trees sprouted; small trees sprouted more frequently than large trees. Sprouting is proposed as a means by which weak-wooded fast-growing trees partially compensate for being prone to snapping.

Description: Stomatal conductance was measured with porometers in two plots of Pinussylvestris L. with markedly different tree spacings (plot 1, 608 stems ha−1; plot 2, 3281 stems ha−1), and hourly rates of transpiration were calculated using the Penman–Monteith equation at intervals throughout one growing season. Stomatal conductance varied little in relation to height or age of foliage. There was a linear decrease in canopy conductance with increasing water vapour pressure deficit of the air. Transpiration rates on both plots increased during the summer (maximum 0.3 mm h−1); rates on plot 1 were always lower (ca. 0.7 times) than on plot 2. Needle water potentials were similar throughout the season and only slightly lower on plot 1 than on plot 2. The mean hydraulic resistance of the trees on plot 1 was 2.4 times that on plot 2. The results support a hypothesis that considers the changes in transpiration rate, conducting cross-sectional area, canopy leaf area, water potential, and hydraulic resistance following thinning as a set of homeostatic relationships.

Description: The degree of resistance to a pulsed electric current of wood in progressive stages of discoloration and decay in living trees was correlated primarily with the moisture and mineral contents of the wood. Below the fiber saturation point, the degree of resistance was correlated primarily with the amount of moisture while above the fiber saturation point resistance was correlated primarily with the concentration of mobile potassium and calcium ions. The apparatus used to produce the current and to measure the resistance to it was accurate in detecting and in indicating the stage of deterioration of discolored and decayed wood. The apparatus was tested and found to be reliable electrically within the range of resistances encountered in the trees studied.

Description: A population of saplings of basswood was studied to determine the effect of size of cutting and a 2-min basal dip in an aqueous solution of either indolebutyric acid (IBA), indoleacetic acid (IAA), or naphthaleneacetic acid (NAA) on root and shoot development of softwood cuttings grown under intermittent mist. Cuttings were 12–60 cm long, and treatments were: distilled water control; and aqueous solutions of auxin at levels of 1000, 5000, 10 000, and 20 000 p.p.m. Significant increases in the growth of roots were obtained with 5000 and 10 000 p.p.m. of IBA. Significant increases in root growth were also obtained with increasing sizes of cuttings.

Description: Jack pine (Pinusbanksiana Lamb.) seeds were sown in October, January, and March, and the seedlings were cultured under accelerated growth conditions in a greenhouse. At biweekly intervals, from May 15 to August 15, they were transplanted to a nearby nursery and sprayed with GA4/7 or GA4/7 + NAA. The following spring a fourfold increase in flowering was noted in trees receiving either of the GA4/7 treatments. Trees in the March sowing did not flower. The data suggest that the increased flowering was caused by GA4/7-mediated differentiation of lateral long-branch primordia into ovulate strobili.

Description: Sprouting of sugar maple (Acersaccharum Marsh.) was investigated in the first growing period following conventional, commercial clear-cutting in the Nashwaak Experimental Watershed Project, in New Brunswick. The number of sprouts and the height of the tallest sprout on 296 stumps from dormant-period cuts were recorded. A regression equation was developed from standing trees in adjacent uncut areas to predict dbh from diameter at stump height. The percentage of stumps which bore sprouts decreased with increasing dbh greater than 15 cm. The relationships between dbh and number of sprouts per stump, and between dbh and height of the tallest sprout per stump were bimodal with frequency peaks occurring at 10 and 65 cm. The number of clumps of sprouts per stump was relatively constant in all dbh classes except for an apparent increase in the 65-cm dbh class.

Description: Dispersal gradients of conidia of Sirococcusclavigignenti-juglandacearum Nair, Kostichka, and Kuntz, the causal agent of butternut canker, were studied in the forest for various rainfall dates. During rain, large numbers of conidia from branch cankers were carried in runoff water down the tree trunks of butternut (Juglanscinerea L.), causing infections that resulted in trunk cankers. Conidia also were dispersed in small droplets or aerosols and disseminated up to 40 m from a cankered tree during rainfall. Concentrations of conidia in the air decreased exponentially with increasing distances from the inoculum source. Dispersal gradients were influenced by stand composition and (or) size of the inoculum source, with steeper gradients associated with a dense forest canopy and a small area source of inoculum. Weather conditions, excluding the necessity for rain, had little influence in altering dispersal gradients within the same stand for different rainfall dates.

Description: Deer browsing and interference from forest weeds, particularly hayscented fern (Dennstaedtiapunctilobula (Michx.) Moore), New York fern (Thelypterisnoveboracensis L.), and short husk grass (Brachyelytrumerectum Schreb.), influence the establishment of Allegheny hardwood reproduction. We determined the independent interference by deer and weeds after a seed cut and a removal cut in a two-cut shelterwood sequence. Weeds, particularly the ferns, caused significant interference with germination, survival, and growth of desirable species following both cuttings. Deer browsing had no direct effect on desirable species because they did not grow enough to emerge from the herbaceous cover. Deer browsing did affect growth ofRubus, yellow and black birch (Betulaalleghaniensis Britt, and Betulalento L.), and pin cherry (Prunuspensylvanica L.) that grew above the herbaceous cover. Browsing of Rubus may be a serious problem in some stands because substantial reduction in fern and grass coverage occurred as the Rubus developed.

Description: The effect of two levels of thinning (zero and [Formula: see text] of basal area removed) and three levels of nitrogen fertilization (0,224, and 448 kg N/ha) on crown development of codominant, 24-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) trees was studied over a 5- to 7-year period. Thinning and heavy ferilization separately increased needle mass per tree after 7 years by 90% and, when combined, by 271%. Yearly needle production peaked 2–3 years after fertilization and resulted from an increase in needle size, needle number per shoot, and number of shoots produced. Maximum foliage mass per tree was reached 4–7 years after fertilization. Thinning effect on needle production was lower initially, but increased throughout the study period. A continuing foliage production in branches low in the crown contributed to thinning effect on foliage mass and crown size. Foliage distribution was affected most in the top half of the crown by fertilization and in the bottom half of the crown by thinning. Fertilization increased branch elongation at all crown heights, but thinning alone had no effect on crown width down to whorl 12.