Publication Date:
2024-06-19
Description:
In der vorliegenden Arbeit wird - basierend auf externmorphologischen Kriterien von Brachiopodengehäusen aus dem spongiolithischen süddeutschen Oberjura - eine ökologische Interpretation von Brachiopodenassoziationen vorgestellt und diese mit nicht verschwammten Vorkommen unter Verwendung verschiedener methodischer Ansätze verglichen. Aus insgesamt elf brachiopodenführenden Lokalitäten unterschiedlicher stratigraphischer und fazieller Niveaus wurden die generell bevorzugt auftretenden Wuchsformen ermittelt, zu Standardmorphotypen zusammengefaßt und mit digitalisierten Abbildungen aus vorhergehenden Bearbeitungen ökologisch verglichen.
[ …]
[ … Tabelle 1. Stratigraphie, Fazies und dominierende Externmorphologie der Brachiopodengehäuse (Standardmorphotyp) in den untersuchten Lokalitäten.]
Die Analysen ergaben, daß mit Hilfe des Verlaufs der dorsalen Umrißlinie, dem Verlauf der Frontalkommissur und untergeordnet der Foramengröße eine hinreichende Interpretation des Environments hinsichtlich Substratbeschaffenheit, Hydrodynamik und Bathymetrie erfolgen kann. Die wichtigsten Ergebnisse sind: a) Die Brachiopoden der süddeutschen Spongiolithfazies zeigen im Vergleich mit je einer Fauna aus einer schwach verschwammten Schichtfazies, einer Korallenfazies und einer oolithischen Mergelkalkfazies grundlegende Unterschiede in der äußeren Morphologie ihrer Gehäuse, b) In mergeldominierten Lokalitäten sind die Sphärizitätsindizes der Gehäuse deutlich höher als in der Korallenfazies, c) Reduzierte Werte des Dicken / Breiten - Verhältnisses scheinen ein Indiz für weichere Substrate zu sein, d) Im zeitlichen Verlauf (Oxfordium bis Unter-Tithonium) nehmen die Werte der Sphärizitätsindizes der Brachiopodengehäuse ab. e) Die Amplituden der Frontalkommissuren von Brachiopoden aus dem Moundbereich sind höher als diejenigen aus der Korallenfazies, f) Das Mengenverhältnis von terebratuliden zu rhynchonelliden Brachiopoden läßt Rückschlüsse auf das dominierende Substrat in den Moundbereichen zu. Ein quantitatives Übergewicht rhynchonellider Brachiopoden geht mit einem mergelreichen, stillen Tieferwasserenvironment einher, während die kalkdominierten Flachwasserbereiche mit erhöhter Hydrodynamik bevorzugt von Terebratuliden besiedelt werden, g) Untergeordnet wird ein Substratwechsel durch die Foramengröße und -position angezeigt. Große mesothyridide Foramina kennzeichnen meist Festsubstrate. Kleine hypothyridide Foramina sind eher für allomikritische Bereiche charakteristisch, h) Brachiopoden der koralligenen Fauna zeichnen sich durch Asymmetrien der Frontalkommissur (Rhynchonellida) und Asymmetrien in der dorsalen Umrißlinie aus (Terebratulida).
Die artikulaten Brachiopoden aus den untersuchten „Riff-Ablagerungen scheinen hinsichtlich ihrer Morphologie und faunistischen Komposition von den kontrollierenden Bildungsfaktoren (besonders Sedimentationsrate, Substratbeschaffenheit und Hydrodynamik) der Schwamm- und Korallenassoziationen, beeinflußt worden zu sein. Die in den jeweiligen Aufschlüssen dominierenden Brachiopoden-Morphotypen reflektieren wenigstens statistisch die Bildungsbedingungen.
Description:
Data on brachiopods in spongolites from the Upper Jurassic of southern Germany were collected in order to establish associations based on paleoenvironmental parameters and compare these results to similar fauna across a range of depositional paleoenvironments. Fossils were sampled and their dominant growth morphotypes identified from 11 localities representing different ages and facies of the Upper Jurassic in the Franconian and Swabian Alb. These growth morphotypes were compared to standard morphotypes and to biometric data of modern brachiopods gleaned from the literature. Through statistical analysis, it was found that the mode of commissure line and in part the diameter of the foraminae correlate well with substrate types and hydrodynamic to bathymetric paleoconditions. General results include: a) the external morphology of brachiopods from spongolites is significantly different from that of brachiopods from well-bedded limestone containing few to no sponges but corals and oolites, b) within a mostly marl substrate, the sphericity index of brachiopod shells is much larger than for shells in coral-bearing rocks, c) width to length ratios of shells are less for softer substrates indicating possible use as an index for substrate consistency, d) from the Lower Oxfordian to the Tithonian, the sphericity index of shells in general decreases, e) amplitudes of the frontal commissure line of brachiopods found within spongolite build-ups are much larger than those from coral-bearing rocks, f) Terebratulidae to Rhynchonellida biometric relations may be used as an index for judging the “quality” of the paleoenvironment; rhynchonellid brachiopods were dominantly found in marly, deeper paleoenvironments, whereas carbonate sediments deposited under shallow, high energy, water paleoconditions were colonized by terebratulid brachiopods, g) the size and position of foraminae are linked to the substrate type; brachiopods with larger mesothyrid foraminae are found associated with hard substrates while smaller hypothyrid foraminae in soft carbonate sediment areas, h) brachiopods from coral rocks are characterized by their asymmetric growth in form (Terebratulida) and of their frontal commissure (Rhychonellida).
In general, brachiopods from reefal deposits appeared to have been directly influenced morphologically by paleoecologic factors, such as sedimentation rate, substrate type, and water energy levels. The dominant standard morphotype of brachiopods appears to correlate in a statistically significant way with the interpreted depositional circumstances at each locality.
Due to here presented morphological data of brachiopodes from the Upper Jurassic of Southern Germany, associations of brachiopodes in spongiolites are interpreted palecologically and compared to similar faunae from other depositional environments. From 11 localities of different stratigraphical age and facies in the Franconian and Swabian Alb the dominant growth morphotypes are presented and abstracted to so-called „standard morphotypes“. The latter are compared to biometrical data of modern brachiopodes, mainly extracted from literature data. Interpreting the mathematical analyses it can be concluded that the dorsal outline, the mode of commissur line and partially the diameter of the foraminae can be used as indicator of substrates, hydrodynamical and paleobathymetrical conditions. The main results are: a) External morphology of brachiopodes from spongiolites exhibits strong differences in contrast to brachiopodes from well-bedded limestones with few or no sponges, to coral-bearing strata and to oolithic limestones, b) In localities, where marls have been predominantly deposited, the sphericity index is much larger than in the coral-bearing rocks, c) Reduced relations of width and length can be tentatively used as index for soft substrates, d) From the Lower Oxfordian to the Tithonian the sphericity index becomes smaller, e) Amplitudes of the frontal commissur line from brachiopodes of spongiolitic buildups are much higher than from brachiopodes in coral-bearing rocks, f) Quantitative relations from Terebratulidae to Rhynchonellida can be used as index for quality of substrates. Rhynchonellid brachiopodes are dominating marly, deeper environments, whereas carbona-tic sediments deposited under shallow water conditions with higher water energy are dominated by terebratulid brachiopodes. g) Changes of substrates are indicated by the size and position of foraminae. Larger mesothyrid foraminae characterize hard substrates, smaller hypothyrid foraminae are typically developed in allomicritic areas, h) Brachiopodes from coralligene rocks are characterized by their asymmetrical growth of form (Terebratulida) and of their frontal commissure (Rhynchonellida). Generally, brachiopodes from reefal deposits have been influenced morphologically by ecologically controlling factors such as the sedimentation rate, the substrate, and the water energy levels. The dominating standard morphotypes of brachiopodes in each locality reflect at least statistically the former depositional environments.
(Translation: Dr. Elizabeth H. Gierlowski-Kordesch, Department of Geological Sciences, Ohio University)
Description:
thesis
Description:
DFG, SUB Göttingen
Keywords:
ddc:560
;
Paläobiologie
;
Paläontologie
;
Brachiopoda
Language:
German
Type:
doc-type:book
Format:
118
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