ALBERT

Description: A new species of the genus Artocella van Achterberg is described from Spain. On the basis of its substantial sexual dimorphisim, the differences between the two previously known species of this Turanian-Mediterranean genus, which had each been described from specimens of only one (and differing) sex, are reassessed.

Description: Dasypoda intermedia spec. nov. from Iran is described. Its description fills a gap of our knowledge of the East Mediterranean fauna of the genus Dasypoda. The West Mediterranean Dasypoda species are well known but the eastern species lack convincing records. Moreover, D. intermedia spec. nov. is a very interesting species from a phylogenetic point of view. It shares some characters common to subgenera Dasypoda s. str. and Megadasypoda Michez, 2004, which provide further evidence for the close relationship of both subgenera.

Description: Thirty-one species of the family Platystictidae of the Philippines are revised, i.e. all species recognised, excluding the species of the Drepanosticta halterata-group. The following new taxa are described: 16 species in Drepanosticta Laidlaw: D. acuta spec. nov., D. aurita spec. nov., D. centrosaurus spec. nov., D. clados spec. nov., D. flavomaculata spec. nov., D. furcata spec. nov., D. hermes spec. nov., D. krios spec. nov., D. luzonica spec. nov., D. malleus spec. nov., D. myzouris spec. nov., D. paruatia spec. nov., D. pistor spec. nov., D. quadricornu spec. nov., D. rhamphis spec. nov., D. trachelocele spec. nov., two in Protosticta Selys, viz. P. lepteca spec. nov. and P. plicata spec. nov., and three in Sulcosticta gen. nov., viz. S. striata spec. nov., S. pallida spec. nov. and S. viticula spec. nov. The status of eleven previously described nominal taxa is established. One, D. septima Needham & Gyger, is doubtfully considered a synonym of D. mylitta Cowley. Based on a preliminary phylogenetic analysis, the species of Drepanosticta are divided into informal species groups. Most species of the Philippines have affinities to species of Sulawesi, the Moluccas and New Guinea. Several species confined to Palawan have sister-group relationships with species from Borneo. The affinities of various other species confined to the Sulu archipelago, are unsettled as yet. The species of Platystictidae here assigned to Protosticta Selys are presumably not closely related to the type species, P. simplicinervis Selys from Sulawesi. However, a better placement has to await a more detailed phylogenetic study of the family. For three species the new genus Sulcosticta gen. nov. is erected. These species are closely allied based on the structure of the appendages, but should have been assigned to different genera if based on the present generic definitions. Many species here described have small distributional ranges, a common phenomenon in Platystictidae. Since most forests in the Philippines are heavily under threat or have already disappeared in the last fifty years, several taxa described in this paper should be considered under threat of immediate extinction.

Description: In this part the remaining 78 species of the genus Pepsis, belonging to ten species-groups, are described and figured, and their phylogenetics and biogeography are discussed. 14 of the species are described as new: P. achterbergi spec. nov., P. adonta spec. nov., P. boharti spec. nov., P. caliente spec. nov., P. dayi spec. nov., P. esmeralda spec. nov., P. ianthoides spec. nov., P. jamaicensis spec. nov., P. krombeini spec. nov., P. martini spec. nov., P. multichroma spec. nov., P. nanoides spec. nov., P. wahisi spec. nov., and P. willinki spec. nov. Three species-names, P. infuscata Spinola, 1841, P. lampas Lucas, 1895, and P. thoreyi Dahlbom, 1845, are recalled from synonymy. The following 293 names are newly synonymized (the valid names are listed first): P. atalanta Mocsáry, 1885 = P. nitens Mocsáry, 1894, P. mocsaryi Lucas, 1895; P. inclyta Lepeletier, 1845 = P. mutabilis Lepeletier, 1845, P. vagabunda Lepeletier, 1845, P. cupripennis Taschenberg, 1869, P. violaceipennis Mocsáry, 1885, P. clotho Mocsáry, 1888, P. spengeli Mocsáry, 1888, P. sickmanni Mocsáry, 1888, P. nireus Mocsáry, 1894, P. atrovirens Lucas, 1895, P. cerastes Lucas, 1895, P. pygidialis Brèthes, 1908, P. guaranitica Brèthes, 1908, P. parca Lucas, 1919, P. atahualpa Banks, 1946, opimicornis, Haupt, 1952, atropos, Haupt, 1952, azurea Haupt, 1952; crassicornis Mocsáry, 1885 = P. sappho Brèthes, 1908, P. nitocris Brèthes, 1908, P. vivida Brèthes, 1908, P. arechavaletai Brèthes, 1908, P. lynchii Brèthes, 1908, P. operosa Brèthes, 1908, P. ataraqua Banks, 1946, P. splendida Haupt, 1952; P. sommeri Dahlbom, 1845 = P. azteca Cameron, 1893; P. xanthocera Dahlbom, 1843 = P. nigrescens Smith, 1855, P. fulgidipennis Mocsáry, 1885, P. juno Brèthes, 1908, P. ismare Banks, 1946, P. nigroprasina Haupt, 1952; P. seifferti Lucas, 1895 = P. cornuta Lucas, 1895, P. moebiusi Lucas, 1895, P. stygia Lucas, 1895; P. luteicornis Fabricius, 1804 = P. strenua Erichson, 1848, P. tinctipennis Smith, 1873, P. citreicornis Mocsáry, 1894, P. venosa Banks, 1945, P. alector Banks, 1946; P. asteria Mocsáry, 1894 = P. luridicornis Brèthes, 1926; P. convexa Lucas, 1895 = P. humeralis Brèthes, 1914; P. helvolicornis Lucas, 1895 = P. bahiae Brèthes, 1914; P. vitripennis Smith, 1855 = P. obscura Lepeletier, 1845, P. amabilis Mocsáry, 1885, P. centralis Cameron, 1893, P. margarete Lucas, 1895, P. venezuelae Kaye, 1913, P. aeneipennis Banks, 1946, P. helenae Haupt, 1952, P. coeruleoviridis Haupt, 1952; P. fumipennis Smith, 1855 = P. pallidicornis Mocsáry, 1885; P. amyntas Mocsáry, 1885 = P. vicina Lucas, 1895, P. clarinervis Brèthes, 1908, P. amyntoides Lucas, 1919, P. eurydice Lucas, 1919; P. dimidiata Fabricius, 1804 = P. vittigera Lucas, 1897, P. argentina Brèthes, 1908, P. sanctaeannae Brèthes, 1908, P. virgo Brèthes, 1908, P. externa Brèthes, 1908, P. transversa Brèthes, 1908, P. cordubensis Brèthes, 1908, P. banghaasi Lucas, 1919; P. menechma Lepeletier, 1845 = P. elegans Lepeletier, 1845, P. dubitata Cresson, 1867, P. prismatica Smith, 1855, P. advena Mocsáry, 1885, cinctipennis Mocsáry, 1885, P. guatemalensis Cameron, 1893, P. nestor Mocsáry, 1894, P. nigricornis Mocsáry, 1894, P. auranticornis Lucas, 1895, P. fruhstorferi Lucas, 1895, P. concolor Lucas, 1895, P. cerberus Lucas, 1895, P. euchroma Lucas, 1895, P. nigrocincta Lucas, 1895, P. mordax Lucas, 1895, P. inermis Fox, 1898, P. roberti Brèthes, 1908, P. janira Brèthes, 1908, P. cultrata Brèthes, 1908, P. novitia Banks, 1921; P. decipiens Lucas, 1895 = P. similis Lucas, 1895; P. minarum Brèthes, 1914 = P. pulchra Brèthes, 1914; P. basifusca Lucas, 1895 = P. angustimarginata Viereck, 1908; P. chrysoptera Burmeister, 1872 = P. exigua Lucas, 1895, P. smaragdinula Lucas, 1895, P. nebulosa Lucas, 1895, P. karschi Lucas, 1895, P. anisitsii Brèthes, 1908, P. indistincta Brèthes, 1908, P. dimidiatipennis Brèthes, 1908, P. chloroptera Brèthes, 1908, P. culta Brèthes, 1908, P. recta Brèthes, 1908, P. tornowii Brèthes, 1908, P. schrottkyi Brèthes, 1908, P. itinerata Brèthes, 1908, P. miniata Brèthes, 1908, P. spegazzinii Brèthes, 1908, P. paulistana Brèthes, 1914, P. chloe Brèthes, 1914, P. coronaria Brèthes, 1914, P. semilucana Haupt, 1952, P. bruneipes Haupt, 1952, P. brachynotus Haupt, 1952, P. diagonalis Haupt, 1952, P. discrepans Haupt, 1952; P. elongata Lepeletier, 1845 = P. purpurascens Smith, 1855, P. fuscipennis Smith, 1873, P. longula Banks, 1946; P. australis Saussure, 1867 = P. centaurus Lucas, 1897; P. cyanescens Lepeletier, 1845 = P. micans Mocsáry, 1885, P. jucunda Mocsáry, 1885, P. balloui Banks, 1946, P. diversa Haupt, 1952; P. lampas Lucas, 1895 = P. venturii Schrottky, 1902; P. nitida Lepeletier, 1845 = P. lucidula Smith, 1855, P. vaualba Smith, 1855, P. pruinosa Mocsáry, 1894, P. cylindrica Lucas, 1895, P. andina Brèthes, 1908, P. dilatata Brèthes, 1908, P. holmbergi Brèthes, 1908, P. concava Brèthes, 1908, P. ephebus Brèthes, 1908, P. vaga Brèthes, 1908, P. fuscobasalis Brèthes, 1908, P. cordata Brèthes, 1914, P. impatiens Brèthes, 1914, P. tricolor Brèthes, 1914, P. joergenseni Brèthes, 1914, P. cleone Brèthes, 1914, P. dorsata Brèthes, 1914, P. aretheas Brèthes, 1914, P. lassonis Lucas, 1819, P. consors Banks, 1946, P. interrupta Banks, 1946, P. analis Haupt, 1952; P. seladonica Dahlbom, 1843 = P. deuteroleuca Smith, 1855, P. kohli Lucas, 1895, P. venezolana Brèthes, 1908, P. burmeisteri Brèthes, 1908; P. cybele Banks, 1945 = P. weberi Banks, 1946; P. thoreyi Dahlbom, 1845 = P. lurida Lucas, 1895, P. euterpe Brèthes, 1908; P. flavescens Lucas, 1895 = P. periphetes Lucas, 1895, P. limbatella Brèthes, 1908, P. discoidalis Brèthes, 1914, P. limbatica Brèthes, 1914, P. militaris Brèthes, 1914, P. cavillatrix Haupt, 1952, P. arcuata Haupt, 1952, P. recterugosa Haupt, 1952, P. adversatrix Haupt, 1952; P. nigricans Lucas, 1895 = P. troglodytes Brèthes, 1908; P. montezuma Smith, 1855 = P. quitonensis Packard, 1869, P. sibylla Mocsáry, 1885, P. circe Mocsáry, 1885, P. occidentalis Cameron, 1893, P. peruanus Lucas, 1895, P. fulva Lucas, 1895, P. nessus Lucas, 1895, P. fusca Lucas, 1895, P. andicola Cameron, 1903, P. chilloensis Cameron, 1903, P. patagonica Brèthes, 1908, P. fasciculata Brèthes, 1908, P. pisoensis Strand, 1911, P. pacifica Brèthes, 1914, P. huascar Banks, 1946; P. completa Smith, 1855 = P. quichua Brèthes, 1908, P. comes Banks, 1946; P. smaragdina Dahlbom, 1843 = P. thunbergi Dahlbom, 1843, P. lara Mocsáry, 1888, P. satrapes Lucas, 1895, P. nupta Lucas, 1895, P. erynnis Lucas, 1895, P. fraterna Lucas, 1895, P. diabolus Lucas, 1895, P. mystica Lucas, 1895, P. thalia Brèthes, 1908, P. brasiliensis Brèthes, 1908, P. pallida Brèthes, 1908, P. iheringi Brèthes, 1908, P. dromeda Brèthes, 1908, P. sepultrix Lucas, 1919, P. strickeri Lucas, 1919; P. discolor Taschenberg, 1869 = P. sinnis Lucas, 1895, P. jujuyensis Brèthes, 1908, P. modesta Brèthes, 1908, P. comparata Brèthes, 1908, P. neutra Brèthes, 1908, P. terebrans Brèthes, 1908, P. procera Haupt, 1952, P. plaumanni Haupt, 1952, P. ogloblini Haupt, 1952, P. deletrix Haupt, 1952; P. limbata Guérin, 1831 = P. richteri Brèthes, 1908, P. polita Brèthes, 1908, P. limbella Haupt, 1952, P. artemis Haupt, 1952; P. basalis Mocsáry, 1885 = P. erdmanni Lucas, 1895, P. basinigra Haupt, 1952; P. infuscata Spinola, 1841 = P. niobe Mocsáry, 1885, P. sagana Mocsáry, 1894, P. incerta Banks, 1946; P. hyalinipennis Mocsáry, 1885 = P. subruficornis Haupt, 1952; P. festiva Fabricius, 1804 = P. pulchella Lepeletier, 1845, P. solitaria Smith, 1879, P. gallardoi Brèthes, 1908, P. hora Brèthes, 1914, P. amok Lucas, 1919, P. riojaneirensis Lucas, 1919; P. gracilis Lepeletier, 1845 = P. diana Mocsáry, 1885, P. hecate Mocsáry, 1885, P. spathulifera Lucas, 1895, P. sphinx Lucas, 1895, P. ierensis Banks, 1945, P. alceste Banks, 1946, P. scalaris Haupt, 1952; P. mildei Stål, 1857 = P. charon Mocsáry, 1885, P. cyanoptera Lucas, 1895, P. dryas Lucas, 1919; P. filiola Brèthes, 1914 = P. denserugosa Haupt, 1952; P. ruficornis Fabricius, 1804 = P. saphirus Palisot de Beauvois, 1805, P. violacea Mocsáry, 1885, P. hexamita Lucas, 1895, P. omniviolacea Haupt, 1952; P. brunneicornis Lucas, 1895 = P. glabripennis Lucas, 1895; P. purpurea Smith, 1873 = P. pan Mocsáry, 1885, P. parthenope Mocsáry, 1885, P. sagax Lucas, 1895, P. clypeata Brèthes, 1914, P. consimilis Banks, 1946, P. laconia Banks, 1946; P. viridisetosa Spinola, 1841 = P. eximia Smith, 1873; P. viridis Lepeletier, 1845 = P. errans Lepeletier, 1845, P. chlorotica Mocsáry, 1885, P. excelsa Lucas, 1895, P. selene Lucas, 1895, P. fimbriata Lucas, 1895, P. calypso Brèthes, 1908, P. fluminensis Brèthes, 1908, P. argentinicus Strand, 1910, P. mimetica Brèthes, 1914, P. garbei Brèthes, 1914, P. erecta Brèthes, 1914, P. tandilensis Brèthes, 1914, P. meridionalis Brèthes, 1914, P. minor Lucas, 1919, P. basifulgens Lucas, 1919, P. nebulosipennis Lucas, 1919, P. purpurea Lucas, 1919, P. koerberi Lucas, 1919, P. inimicissima Lucas, 1919, P. debilitans Lucas, 1919, P. itapaca Banks, 1946; P. aciculata Taschenberg, 1869 = P. nero Lucas, 1895; P. atripennis Fabricius, 1804 = P. flavilis Brèthes, 1908; P. ianthina Erichson, 1848 = P. fulvicornis Mocsáry, 1885, P. sirene Lucas, 1895, P. balboae Lucas, 1919, P. herodes Lucas, 1919, P. curti Lucas, 1919; P. nana Mocsáry, 1885 = P. mapiriensis Lucas, 1919, P. vinciens Lucas, 1919, P. ilione Banks, 1946, P. moesta Banks, 1946, P. orestes Banks, 1946, P. amautas Banks, 1946, P. inaequalis Haupt, 1952; P. hirtiventris Banks, 1946 = P. viridaurea Haupt, 1952, P. aequalis Haupt, 1952; P. auriguttata Burmeister, 1872 = P. aurimacula Mocsáry, 1885, P. flavicornis Mocsáry, 1894, P. guttata Lucas, 1895, P. incendiaria Lucas, 1895, P. pubiventris Lucas, 1895, P. planifrons Lucas, 1895, P. lestes Lucas, 1895, P. villosa Brèthes, 1908; P. sabina Mocsáry, 1885 = P. astioles Banks, 1946; and P. purpureipes Packard, 1869 = P. chlorana Mocsáry, 1885, P. antennalis Cameron, 1893, P. sulcifrons Cameron, 1903, P. carinata Brèthes, 1914, P. equatoriana Brèthes, 1914, P. angusta Banks, 1946. Keys to all forms are given. The mimicry-groups of P. atripennis Fabricius, 1804, and P. completa Smith, 1855, are defined and described and a comparative account of mimicry based on all four mimicrygroups in Pepsis is given. Lists of excluded species (with their current taxonomic placement and depository where ascertained), unplaced names, and a nomen nudum are given.