ALBERT

Description: https://www.researchgate.net/publication/230891291_The_Orbital_Theory_of_Pleistocene_Climate_Support_frim_a_Revised_Chronology_of_the_Marine_d18O_Record

Description: Summary Holocene sediments of the North Lagoon, Bermuda, were studied with shallow seismic reflection profiles (200 km CSP-survey, UNIBOOM-system) and vibration coring (40 sediment cores, pneumatic vibration corer, Meischner et al., 1981). Seismic Stratigraphy Four seismic sequences are distinguishable by seismic stratigraphy. All seismic sequences correspond to depositional sequences built up during high sea levels in interglacial times. The seismic sequences are separated by unconformities which are often strongly reflective and correspond to emersion planes during glacial phases. The upper sequence (sequence 4) is related to Holocene sediments. The pre-Holocene bedrock is divided into three different seismic sequences (Kuhn et al., 1981): Sequence 1: oldest Pleistocene sequence (pre-Sangamon sea-level highstands), upper boundary with levelled relief (lower boundary not discernible), composed of strongly cemented carbonate sediments, forms the bedrock below Three Hill Shoals Sequence 2: Sangamon (125 ky sea-level highstand), distinct surface morphology, forms the bedrock of a large area below Holocene sediments, Holocene reefs grew up on elevations of the sequence 2 surface, the Holocene reef rim was developed on an elevated rim of sequence 2 Sequence 3: youngest Pleistocene sequence (Sangamon, 105 and 85 ky sealevel highstands lower than recent), deposited mainly in depressions of the bedrock deeper than -15 m below recent Mean Sea Level, levelling the older relief, peat sedimentation in places The distribution of recent reef areas and lagoonal basins is strongly controlled by pre-Holocene topography and geology of the bedrock. During the Holocene approx. 1050 x 106 m3 of carbonate sediments were deposited in the North Lagoon (290 km2) and approx. 1350 x 106 m3 in the reef rim area (170 km2). Sedimentology There are no larger oscillations of the Holocene sea level identifiable in the sedimentological record. The pre-Holocene topography was gradually drowned during the Holocene sea-level rise. At first, the depositional depressions were separated and landlocked. Fresh water peat marshes, fresh water ponds, marine ponds and bays were formed. With rising sea level, the land barriers were more and more eroded, drowned and lost their influence on the back-barrier sedimentation area. Autochthonous and allochthonous peat, lime gyttja and carbonate mud are a typical transgressive back-barrier sediment sequence. After destruction of the barrier, the depositional milieu changed from restricted marine to normal marine, open lagoonal. Sea-grass sediments and nearly mud-free carbonate sand were deposited in shallow water in an exposed environment. Hydrodynamic energy decreases with increasing water depth in the lagoonal basin. A more densely growing reef rim and intralagoonal reef growth added to the protection of the deeper lagoonal floors. Fine-grained sediments were deposited in this environment. They are distributed over a large area of the North Lagoon and form the top of the transgressive lagoonal sediment sequence. Holocene reefs mainly developed on rises of the pre-Holocene surface. In the early Holocene, solid reef build-ups were able to keep up with the rapid rise of sea level. Sand pockets in the reefs were left behind and filled up mainly in the later Holocene. The percentage of fine-grained sediments, produced and resuspended in the reef rim and deposited in the near lagoonal back-reef zone, increased during the Holocene. Two models of Holocene sedimentation in a depression and on an elevation of the pre-Holocene surface illustrate the dependence of vertical facies gradation on pre-Holocene topography. Trends of the mostly polymodal grain-size distributions of the Holocene sediments are a coarsening-upward in the back-barrier and a fining-upward in the lagoonal sediment sequences. Change in the composition of the molluscan fauna in the Holocene sediments (particle size 〉 2000 µm) is an Indication for fades changes. Gastropods are abundant in the basal backbarrier sediments. Bivalves are rare and their diversity 1s low. Sea-grass sediments contain Codakia orbicularis and Astraea phoebia shells. In the sheltered lagoonal environment shell fragments 〉 2000 µm become rare, common species are Gouldia cerina, Pitar fulminata and Finella sp. (approx. 1000 µm). Fine-grained reef-rim derived sediments differ from lagoonal sediments by a higher percentage of Homotrema rubrum fragments and Alcyonaria spicules.

Description: Neohattoria Kamim. is a monotypic genus of the Jubulaceae (= Frullaniaceae) with a single species, N. herzogii (Hatt.) Kamim., known from central to northern Japan and the southern part of the Kurile Islands. The present genus was segregated from Frullania by Kamimura (1961; sub. nom. Hattoria Kamim. nom. illeg., non Schust., 1961) on the basis of the branching type, the shape of the first leaf and underleaf on branch, the total lack of secondary pigmentation, the uniform cell structure of the stem in cross section, and the strongly toothed leaf lobes. The generic concept of Neohattoria was greatly expanded by Schuster (1970), who included eight species and classified them into two subgenera, subgen. Neohattoria (with a single species) and subgen. Microfrullania Schust. (with seven species); however, Hattori et al. (1972) transferred all species of subgen. Microfrullania to a newly segregated genus Schusterella Hatt. et al., thus retaining the monotypic status of Neohattoria. As already described and illustrated by Hattori (1955), Kamimura (1961), Mizutani (1961), Ladyzhenskaja (1963), Schuster (1970), and Hattori et al. (1972), Neohattoria herzogii is closely related to species of both Jubula and Frullania. Regarding the taxonomic desposition of Neohattoria, Mizutani (1961) and Mizutani & Hattori (1969) placed it with Jubula in a subfamily Jubuloideae of Lejeuneaceae and Hattori et al. placed it in Jubulaceae (s. lat.). But, Kamimura (1961), Schuster (1970, 1979), and Guercke (1978) placed it more close to Frullania, e.g. in a subfamily Frullanioideae of Jubulaceae (s. lat.); more recently, Asakawa et al. (1979b), admitting three distinct families, Jubulaceae, Frullaniaceae, and Lejeuneaceae, placed Neohattoria and Jubula in the Jubulaceae (s. str.) but Frullania and Schusterella in the Frullaniaceae.

Description: The originally monotypic eastern Malaysian genus Schiffneriolejeunea Verdoorn 1933 has now become a widespread, pantropical group of about fifteen species by the inclusion of species from the genus Ptychocoleus Trev. nom. illeg. Six species are known from Asia, three of which constitute the sect. Saccatae (Verdoorn) Gradst. & Terken comb. nov. These are the widespread Schiffneriolejeunea tumida (Nees) Gradst., the eastern Malaysian S. cumingiana (Mont.) Gradst. and S. nymannii (Steph.) Gradst. & Terken comb. nov. Schiffneriolejeunea tumida is a rather polymorphic species in which two not sharply defined varieties may be distinguished: S. tumida var. tumida with more or less involuted leaf margins, and S. tumida var. haskarliana (Gott.) Gradst. & Terken comb. nov. with plane margins.

Description: A phytosociological survey based on methods of the Zürich-Montpellier School was carried out in the páramo vegetation of the Cordillera Oriental, Colombia. The study area covers about 10,000 and comprises the páramo between the Nevado de Sumapaz (3°55'N, 4250 m), the Sierra Nevada del Cocuy (6°25'N, 5493 m) and the Páramo del Almorzadero (7°N, 4375 m). The páramo vegetation was studied along various altitudinal transects from the upper forest line (3000-3500 m) up to the lower limit of the snowcap (4800 m). A general description of the study area includes data on geology, geomorphology, soils, climate, flora, phytogeography, morphological characters of the vegetation, fauna and landuse. The evolution and Quaternary history of páramo vegetation and climate is reviewed, incorporating the first data from the Lateglacial and Holocene of the Páramo de Sumapaz. The general altitudinal zonation of the páramo vegetation was studied and is presented for both the dry and the humid side of the Cordillera. The zonal and azonal plant communities are described including their physiognomy, composition and syntaxonomy, habitat and distribution. Eighty five syntaxa from the rank of variant to that of the class are newly described, 17 of which are provisional. The vegetation is not ranked syntaxonomically yet, but described on the basis of preliminary tables. A number of azonal communities, part of them of lesser extent, are described in a similar way. The páramo vegetation is primarily determined by the tropical diurnal high mountain climate. The diversity of the páramo vegetation is related to temperature (altitudinal gradient) and to humidity (dry and wet climate). The presence of zonal bunchgrass páramo, bamboo-bunchgrass páramo or bamboo páramo mainly depends on the complex interrelation between these factors. Finally a synthesis is provided on ecology, morphology and phytogeography of the páramo vegetation of the study area.

Description: Isoëtes Palmeri with a distribution in the High Andes from the Páramo of Venezuela to the Páramo of Ecuador is described as a new taxon, and dedicated to the then American specialist of the genus, Thomas Chalkley Palmer (1860-1934). The new species belongs to the tropical-Andeanaustral-antarctic section Laeves, described as new here as well. The publication of the new species had to be anticipated to the projected monographic treatment of the South-American representatives of the genus Isoëtes, as A.M. Cleef, Utrecht intends to base a new association, the Isoëtetum Palmeri on this new taxon, observed and collected by him at many instances within the Colombian Páramo between 1971 and 1980 in the context of the preparation of his doctoral thesis now under way.

Description: Until recently relatively little attention has been paid to the study of chromosomes in liverworts. The first substantial contributions were made by Heitz (1927, 1928) and Lorbeer (1934). In the second half of this century chromosome studies on liverworts were mainly carried out in Europe (e.g. Fritsch 1972; Newton 1977, 1979) and Japan (e.g. Tatuno 1959; Segawa 1965a, b, c; Inoue 1968). Inoue (in Koponen 1979) reports that until now 28% of all bryophyte species in Japan have been investigated as to their chromosome complement. A comprehensive, but rather outdated, survey of chromosome numbers in Hepaticae and Anthocerotae was given by Berrie (1960). Work on a new, updated survey is now underway (Fritsch, in prep.). In the present article results are presented of a cytotaxonomic investigation of European species of the genera Aneura and Riccardia (Aneuraceae). Most specimens were gathered in the Netherlands, but some chromosome counts based on French and German plants are also included.

Description: ANDERSON, J.A.R., A checklist of the trees of Sarawak, 364 pp. (1983, Dewan Bahasa dan Pustaka Cawangan Sarawak, for Forest Department, Kuching, Sarawak). Cloth Mal$ 15.00. When Dr. Anderson retired from the Forest Department in 1973 he left the manuscript of this checklist for publication. Unfortunately publication was delayed for 10 years. It contains data on over 2500 arboreous plant species. The text consists mainly of two parts: the first is a list of vernacular names with their scientific equivalents, the second is a list of plant names alphabetically arranged by family. Each species is concisely annotated with its vernacular name(s), maximum diameter, ecology, frequency, soils, etc. Species names have been coded: the first two figures are for the family, the next two for the genus and the last two for the species. A list is given of the trees of the peat-swamp forests of which Anderson was a great expert. A small draw-back is that the literature of the last ten years has not been included. Nevertheless this is a most helpful book. — C.G.G.J. van Steenis.

Description: Small evergreen trees, shrubs or lianas; two genera ( Cansjera and Opilia) are known to be root-parasites. Leaves distichous, simple, usually extremely variable in form and size, entire, exstipulate, pinnately veined; dried leaves mostly finely tubercled by cystoliths located in the mesophyll. Inflorescences axillary or cauliflorous, panicle-like, racemose, umbellate (in Africa) or spicate; bracts narrowly ovate or scale-like, in Opilia peltate, often early caducous. Flowers small, (3—) 4—5) (—6)-merous, mainly bisexual, sometimes unisexual and plants then dioecious ( Gjellerupia, Melientha, and Agonandra) or gynodioecious (Champereia). Perianth with valvate, free or sometimes partly united tepals (in ♀ flowers of Gjellerupia wanting). Stamens as many as and opposite to the tepals (in ♀ flowers only small staminodes); anthers introrse, 2-celled, longitudinally dehiscent. Disk intrastaminal, lobed (lobes alternating with the stamens), annular, or cupular. Ovary superior, 1-celled; style short or none, stigma entire or shallowly lobed. Ovule 1, pendulous from the apex of a central placenta, anatropous, unitegmic and tenuinucellar. Fruit drupaceous, pericarp rather thin, mesocarp ± fleshy-juicy, endocarp woody or crustaceous. Seed large, conform to the drupe, without testa; hilum basal, often in a funnel-shaped cavity. Embryo terete, embedded in rich, oily endosperm, nearly as long as the seed or shorter, with 3—4 linear cotyledons, radicle often very short. Distribution. There are 9 genera with about 30 spp., widespread in the tropics. Rhopalopilia is restricted to Africa and Madagascar, Agonandra to South and Central America. In Malesia: 7 genera, 5 of these only known from the eastern Old World (1 endemic: Gjellerupia in New Guinea); Opilia and Urobotrya occur also in tropical Africa.

Description: Monoecious, medium-sized to very large trees (rarely shrubby in very exposed situations). Either four independent cotyledons or two fused pairs (which may be retained in the seed after germination). The growing point of foliage shoots quite distinct between the two genera, being just a few highly reduced leaves in Araucaria and a highly organized bud formed of overlapping scales in Agathis. The leaves vary from scales or needles to broad leathery forms with many parallel veins sometimes on the same plant at different stages of growth. Pollen produced in cylindrical cones from one to as much as twenty cm long with numerous pedunculate spirally placed microsporophylls each with several to many pendent elongated pollen sacs attached to the lower side of an enlarged shieldlike apex which also projects apically more or less overlapping the adjacent microsporophylls. Pollen cones solitary, terminal or lateral, on branches separate from those bearing seed cones, subtended by a cluster of more or less modified leaves in the form of scales, deciduous when mature. Pollen globular, without ‘wings’. Seeds produced in large, well-formed cones which disintegrate when mature, dispensing the seeds in most cases with the help of wing-like structures; the seed cone terminal on a robust shoot or peduncle with more or less modified leaves that change in a brief transition zone at the base of the cone into cone bracts, formed of numerous spirally-placed bract complexes, usually maturing in the second year. Individual seed cone bract leathery or woody and fused with the fertile scale which bears one large inverted seed on its upper surface. Distribution. The 40 species in two genera are well represented in Malesia (13 spp.) and extend eastward and southward into Fiji, New Caledonia (18 spp.), Australia, and New Zealand, with 2 spp. also in the cooler parts of South America, giving the family a distinct Antarctic relationship. Only one species of Araucaria (in South America) occurs completely outside of the tropics, while the majority of the species in the family belong in the lowland tropics and others grow in the tropical highlands.

Description: Many botanists must have wondered why as yet no volume of Flora Malesiana was dedicated to the outstanding botanist Carl Ludwig Blume, undisputed pioneer in planning the compilation of a ‘Flora Malesiana’. The writing of this Dedication would have been greatly facilitated if a full biography of BLUME had been existent, but none is available; there is not even a bibliography of his works. Only recently, in 1979, two biographical attempts were made, by J. MACLEAN and by A. DEN OUDEN, but only for the period 1820-1832; together with other biographical and obituary notes they are here assembled in Appendix B. I have also compiled a bibliography: Appendix A.²

Description: Erect or straggling herbs, shrubs or trees, sometimes monoecious or dioecious, the herbs sometimes rhizomatous; branches sometimes jointed at the nodes, sometimes without vessels ( Sarcandra). Leaves simple, decussate or sometimes whorled in fours, serrate, crenate or dentate, the teeth often thickened at the apex, penninerved, usually petiolate; petioles more or less connected at the base at least by a transverse line or connate into a distinct sheath; in Ascarina often alternating with leafless internodes which have the petiolar sheath; stipules minute to fairly conspicuous, subulate, borne on the petiole bases or sheath, occasionally pectinate. Flowers much reduced, without perianth, fully unisexual or essentially bisexual with the reduced anther-bearing organ adnate to the side of the ovary; arranged in spicate, paniculate, or capitate axillary or terminal inflorescences. — Male flowers bracteate or not, apparently consisting of 1—5 stamens, or in Hedyosmum consisting of numerous anthers in a cone-like structure; if 3 then the whole forming a fused 3-lobed organ sometimes enveloping the female flower by its edges, the central anther with 2 or aborted loculi and the laterals with single loculi, simply lobed or with connectives slightly to considerably produced so that the whole organ is 3-fingered; if with only 2 anther locelli then these on either side of a thickened filament plus connective. — Female flowers naked or enclosed by a cupular bract, the perianth adnate to the ovary, often minutely or shortly dentate at the apex and the ovary thus inferior; ovary 1-locular; stigma sessile or style short; truncate, 2-lipped, depressed or subcapitate (or horseshoe-shaped in one species), rarely linear or clavate. Ovule solitary, orthotropous, pendulous, bitegmic and crassinucellate. Drupes fleshy, small, ovoid or globose, sometimes more or less 3-sided in Hedyosmum, free or united into a mass by the bracts; endocarp hardened and crustaceous. Seeds subglobose, exarillate, with copious fleshy or oily endosperm and minute embryo, the cotyledons divaricate or scarcely formed. Distribution. Four genera with about 80 species. Since VESTER’S (1940) small-scale map the family (Ascarina) has been found in Madagascar. It is mainly tropical but Ascarina extends south to North Island of New Zealand (fig. 6) and Chloranthus and Sarcandra extend north to Japan, China, Korea and the eastern U.S.S.R. (Ussuri).

Description: Trees or shrubs (or rarely suffrutices outside Malesia). Leaves simple, alternate, often coriaceous, glabrous or with an indumentum on undersurface, margin entire; petioles often with 2 lateral glands. Stipules 2, minute and caducous to large and persistent, usually linear-lanceolate. Inflorescence racemose, paniculate or cymose; flowers bracteate and usually bibracteolate; bracts and bracteoles small and caducous or larger and enclosing flower or groups of flowers and persistent. Flowers actinomorphic to zygomorphic, hermaphrodite or rarely polygamous, markedly perigynous. Receptacle campanulate to cylindrical or rarely flattened cupuliforum, often gibbous at base; calyx lobes 5, imbricate, often unequal, erect or reflexed. Petals 5 (absent in some Neotropical species), inserted on margin of disk, commonly unequal, imbricate, deciduous, rarely clawed. Stamens indefinite, 2—60 (to 300 in Neotropics), inserted on margin of the disk, in a complete circle or unilateral, all fertile or some without anthers and often reduced to small tooth-like staminodes; filaments filiform, free or ligulately connate, short and included to long and far exserted; anthers small, 2-locular, longitudinally dehiscent, glabrous or rarely pubescent. Ovary basically of three carpels but usually with only one developed, the other two aborted or vestigial, variously attached to (the base, middle or mouth of) receptacle, usually sessile or with short gynophore, pubescent or villous; ovary unilocular with two ovules or bilocular with one ovule in each locule. Ovules erect, with micropyle at base (epitropous). Style filiform, basally attached; stigma 3-lobed or truncate. Fruit a fleshy or dry drupe of varied size, interior often densely hairy; endocarp much varied, thick or thin, fibrous or bony, often with a special mechanism for seedling escape. Seed erect, exalbuminous, the testa membraneous; cotyledons amygdaloid, plano-convex, fleshy, sometimes ruminate. Germination hypogeal with the first leaves opposite or alternate or epigeal with opposite first leaves. An extensive review of the generic limits of the family has been published: G.T. PRANCE & F. WHITE, The genera of Chrysobalanaceae: a study in practical and theoretical taxonomy and its relevance to evolutionary biology, Phil. Trans. Roy. Soc. London 320 (1988) 1—184. This contains full details of taxonomic history, morphology, anatomy, pollen, ecology and distribution of the family. A condensed version of these subjects is given here. Details of the Neotropical members of the family are given in: G.T. PRANCE, Chrysobalanaceae, Flora Neotropica 9 (1972) 1—410. The African members of the family were treated in: F. WHITE, The taxonomy, ecology and chorology of African Chrysobalanaceae (excluding Acioa), Bull. Jard. Bot. Nat. Belg. 46 (1976) 265—350.

Description: Perennial herbs, more commonly woody at the base, undershrubs or shrubs, erect, scrambling or scandent, sometimes high lianas. Rhizome not rarely tuberous. Branches often slightly swollen and jointed at nodes. Hairs simple, uni- or multicellular, short ones often with a hooked apex. Leaves simple, spiral or alternate, petioled (without an abscission zone), exstipulate; midrib usually prominent beneath, elevated or flat above; nervation commonly palmate, or pinnate, nerves often obliquely extending towards the margin. Flowers bisexual, actinomorphic or zygomorphic, solitary, fasciculate, or in axillary or cauligerous, racemose, paniculate or cymose inflorescences, usually only one or two flowers open at a time; bracts present and often persistent; pedicel often hardly distinct from the ovary. Calyx petaloid, gamosepalous, 3- (or 6-) lobed or 1-lipped; lobes valvate or induplicate. Petals (in Mal.) absent. Disk (?) 0, rarely present (e.g. a few Thottea spp.). Stamens 6 (4 or 5 in some extra-Mal. Aristolochia spp.) or 6—c. 36 (—46), in 1 whorl or in 2 (3 or 4) whorls (Thottea); filaments free or slightly mutually united at the base, and/or almost completely adnate to the style column to form a gynostemium; anthers free (Thottea) or dorsally united with the style column (Aristolochia), each consisting of 2 thecae with 4 pollen sacs, extrorse, rarely introrse (extra-Mal. spp.), dehiscing longitudinally. Ovary inferior (rarely half-inferior in extra-Mal. genera), 4—6-carpellate, 4—6-celled, syncarpous (or ± apocarpous in extra-Mal. Saruma); placentae parietal (distinct when young, then intruding and connivent axially, thus often seemingly axile); ovules usually many, anatropous, in 1 or 2 vertical rows in each locule of the ovary, horizontal or pendulous; style-column 3—many-lobed, sometimes some of the lobes redivided; stigmas or stigmatic tissue apical, lateral, or on the surface of style lobes. Fruits capsular or siliquiform (follicular or cocci in extra- Mal. genera), 4—6-celled; dehiscing apically towards the base (basipetal, e.g. Thottea) or basally towards the apex (acropetal, e.g. most Aristolochia); septicidal, rarely septifragal (some extra-Mai. Aristolochia) or bursting irregularly (extra-Mal. Asarum); rarely indehiscent (W. African Pararistolochia). Seeds many in each locule (1-seeded in extra-Mal. Euglypha), often coated with remains of placental tissue (membranous when dry), horizontal or pendulous, variously shaped; ovate, deltoid or triangular, flat, convex-concave, or longitudinally curved, or oblong (and triangular in cross-section), rugose, finely verrucose, or smooth, immarginate (Thottea; Aristolochia, p.p.) or winged (Aristolochia, p.p.); albumen fleshy, copious; embryo minute, cotyledons two, distinct. Distribution. There are 7 genera, Aristolochia worldwide, Asarum over the northern hemisphere, Thottea in continental Southeast Asia and Malesia, Pararistolochia in tropical Africa, and 3 monotypic genera, viz. Saruma in China, Holostylis and Euglypha in South America. As to number of species, Aristolochia is by far the largest with some 300 spp., largely concentrated in the New World, especially in Central and South America, in Malesia with 28 spp.; Asarum (incl Hexastylis and Heterotropa) with possibly some 70 spp. in northern temperate regions, Thottea with 26 spp., of which 22 in Malesia, and Pararistolochia with 12 spp. in West Africa.

Description: Type material of Tulasnella cystidiophora Höhn. & Litsch. has been studied. The species is characterized by often moniliform gloeocystidia and clamp-less hyphae (at least in the subhymenium).

Description: During an ecological study of fungi of the tidal mudflats in Kuwait, a Sporothrix species has been recorded twice, in 1977 and 1980. It differs from other species of the genus (de Hoog, 1974, 1978) in several characters and is here described as a new species. A comparison with similar species of the genus is added.

Description: A new species, Alstonia undulifolia Kochummen & Wong, is described from the Malay Peninsula. Two sections of the genus occur in the Malay Peninsula, Alstonia sect. Monuraspermum Mon. and Alstonia sect. Alstonia, the latter being the correct name for what was previously known as sect. Pala (Adr. Juss.) Benth. Various characteristics, including growth architecture, are examined for their usefulness in distinguishing these two sections of the genus. In comparing A. angustiloba Miq. and A. pneumatophora Berger, both of which have not been properly differentiated by characteristics of the reproductive organs, A. pneumatophora var. petiolata Mon. is reduced to synonymy under A. angustiloba. A key to the seven species of Alstonia native to the Malay Peninsula is provided.

Description: Among the collections of Knema acquired by the Rijksherbarium since the publication of my new account of the genus Knema, in Blumea 25, 1979: 321 — 478, a few specimens caused problems with the identification, and at closer examination these yielded facts of interest which are published here. Some specimens represented stages not yet known, for instance fruits, or male flowers, while other specimens meant a significant range extension of the species. Two new species and one new subspecies are described. For easy reference, the sequence and numbers of the species presently treated correspond with the numbers as used in the account of 1979. The new species bear the number of the species after which they appear in the general key of 1979, with the addition ‘-bis’.

Description: The early development (ontogeny) of the carpels of 20 species belonging to 8 apocarpous families was investigated with the scanning electron microscope. The results indicate that on the floral apex a circular or a convex meristem develops into an obliquely ascidiate primordium by unequal growth of its periphery. By further unequal growth it develops into a young carpel. The terminal mouth of a cup becomes the lateral cleft of a carpel. The different forms of the young carpels in different species are defined by the varying degree of development of the adaxial region of the initial meristem and/or its margin on the side of the floral apex. This hypothesis is theoretically evaluated.

Description: The stomata as occurring on the fronds of the sporophytes of a large number of Polypodiaceae s.s. (Filicales) are investigated. A number of different stomatal types is recognised, (newly) described, and their ontogeny investigated. The different types of stomata are discussed in relation to their possible significance for tracing phylogenetic relationships in the Polypodiaceae following a cladistic analysis.

Description: Revision of the Malesian species of the genus Steganthera, which centres in New Guinea; precursor to treatment in Flora Malesiana. There are 16 species accepted; 5 are described as new, 12 names are reduced, 3 are excluded and 9 are imperfectly known.

Description: In a recent thesis B.S. Fey (Zürich) has developed a new theory about the origin of the cupule in Fagaceae. He has concluded that the appendages (spines, lamellae, etc.) on the outside of the cupule are regularly arranged and that they reflect a condensation (concrescence) of a dichasial flower system, so that cupule and fruit(s) form together the representation of one ancestral inflorescence; the cupular appendages would then largely represent the bracts of the ancestral inflorescence. This stands in contrast with former opinions, in which the cupule was interpreted as of separate vegetative origin from the nut(s) which was (were) the remain (s) of the inflorescence.

Description: Recent studies in Sabah and Sarawak have demonstrated the presence of an undescribed species of Podocarpus.

Description: A world-revision of Arthraxon Beauv. ( Gramineae) is presented. Three wide-spread species, A. hispidus (Thunb.) Makino, A. lanceolatus (Roxb.) Hochst., and A. lancifolius (Trin.) Hochst. are very variable and have caused the description of a great number of taxa, most of which are here reduced to synonomy. There are now 7 species and 9 varieties; for 6 of the latter new combinations are proposed. No new taxa are described.

Description: Pholidota kinabaluensis is transferred to the new monotypic genus Entomophobia. Coelogyne phaiostele, C. ridleyana, and Pholidota triloba are identical and transferred to the new genus Geesinkorchis, that also comprises the new species G. alaticallosa. The monotypic genus Sigmatochilus is reduced to Chelonistele, in which C. dentifera and C. lurida var. grandiflora are described as new. Chelonistele crassifolia is regarded as a variety of C. sulphurea.

Description: The genera Hunteria and Lepiniopsis of the family Apocynaceae are in Malesia represented by one species each. Distribution and ecology are cited in full.

Description: Five new species of Rafflesia (Rafflesiaceae) are described, while attention is drawn to a sixth, possibly also new one. A key to all recognized species is given.

Description: The morphology and leaf anatomy of Myxopyrum is described and a key to the species is given. Of the 15 species previously described four species and two subspecies are recognised: M. nervosum Bl. (synonyms M. horsfieldii, M. zippelii) with one subspecies coriaceum (Bl.) Kiew (synonym M. ellipticum), M. ovatum Hill (synonyms M. macrolobum, M. cordatum, M. philippinensis), M. pierrei Gagnep. (synonym M. hainanense) and M. smilacifolium Bl. (synonym M. serrulatum) with one subspecies confertum (Kerr) Kiew. Myxopyrum enerve Steen. is Chionanthus enerve (Steen.) Kiew. Descriptions for the extra-Malesian species, M. smilacifolium, is given.

Description: In subgenus Malachobatus twenty Malesian species are recognized, one of them ( Rubus moluccanus L.) with four varieties. Synonymy, descriptions, habitat notes, etc. are given. New names: R. moluccanus L. var. discolor (Bl.) Kalkm. and var. angulosus Kalkm. A key is given to the Malesian species.

Description: In Southeast Asia (excluding India) 44 taxa are recognized, 39 species, of which four are newly described ( I. kerrii, I. luzoniensis, I. emmae, and one unnamed species A, which will be treated by Nguyen Van Thuan, Paris), four subspecies, one of which is new (I. sootepensis subsp. acutifolia) and three are new combinations ( (I. suffruticosa subsp. guatemalensis, I. trifoliata subsp. unifoliata, I. trita subsp. scabra) ), and one variety which is a new combination I. spicata var. siamensis). A key, descriptions and full synonymy are given as well as 4 distribution maps and 5 figures.

Description: Owing to their limited possibilities for either active or passive dispersal, their association with the soil habitat, their vulnerability towards a dry atmosphere, and, in fact, on account of their general ecology and ethology, Diplopoda among arthropods are surely one of the most important classes in relation to the study of historical biogeography. For the class as a whole the sea appears to be an unsuperable barrier as is proved by the almost complete absence of endemic taxa on oceanic islands. In many cases lowland plains also act as severe obstacles against the dispersal of millipedes. The presence or absence of diplopods on islands or continents, therefore, may give a strong argument in favour or against any supposed former land connection. The long geographical isolation of the Australian continent and the absence of endemic higher taxa seems to imply that most, if not all, of its diplopod fauna dates from the time this continent was solidly attached to other southern continents, i.e. the Mesozoic. Subsequent penetration of fauna elements from the north or northwest seems utterly unlikely, although perhaps not entirely impossible.

Description: Recent investigations of the distribution of trace elements in metamorphic index minerals of metapelites have revealed, that the plurifacial character of the Hercynian metamorphism in this area is confirmed by the distribution of Yttrium in Hercynian garnets of the metamorphic series.

Description: The formation of thick piles of flysch-like sediments needs the existence of narrowed seas, active denouement of neighbouring continents, and generalized marginal subsidence. These conditions are present during the initial and final stages of Wilson’s perceptive cycle. In this context, the Late Precambrian flysch of the Iberian Massif must be related to the initial rifting, whilst the Culm of southwestern Iberia was accumulated during an episode of Upper Palaeozoic subduction that remained active after the impingement of Iberia against North America. Culm sediments shed from the uplifted collision zone and fed into a remnant ocean that remained at the nonsutured southern border of Iberia. This model of synorogenic flysch formation has been described elsewhere for similar plate arrangements. On other grounds this model provides a framework that explains the different structural and magmatic trends of the Ossa-Morena Zone (near the active margin) in the context of the rest of the Massif (basement reactivation). In addition to this, it seems to support a partly primary origin for the Iberian arc versus a secondary origin.

Description: Five halacarid species, found in the mesopsammal of Caribbean Islands, are described, viz. Halacarellus tropicalis n. sp., Copidognathus grandiosus n. sp., Agaue arubaensis n. sp., Scaptognathus ornatus n. sp., and Limnohalacarus cultellatus Viets, 1940. H. tropicalis is the first member of the genus Halacarellus reported from tropical beaches.

Description: Seven springs in the Middle Atlas and five in the Rif have been studied. These show a great diversity of crenal habitats: water temperature ranges from 8.7° to 21°C, and the flow from 1 l/s to 1,800 l/s. Based on hydrologic and thermic characteristics, a spring typology is provided. The invertebrate community consists of 60 species, among which 4, found in the Rif, are new to science: Protonemura sp. (Plecoptera), Obuchovia sp. (Diptera, Simuliidae), Rhyacophila fonticola n. sp., and Philopotamus ketama n. sp. (Trichoptera). The new Trichoptera are both described. Two rare endemic species (the planarian Acromyadenium maroccanum and the coleopteran Elmis atlantis) have been found in a cold-water spring in the Middle Atlas; two black-fly species ( Cnetha carthusiensis and Simulium lamachei), new to North Africa, have been collected in a cold-water spring in the Rif. The cold-water spring community shows a high rate of endemism. Seven endemic cold-stenothermous species constitute a most characteristic crenon fauna in northern Morocco. The fauna of warmer springs (18° ≤ temp. ≤ 21°C) contains potamophilous and thermophilous species, a few of them belonging to the Ethiopian fauna. A comparative study of spring and rhithric communities of Morocco shows that, in the Middle Atlas and the Rif, cold-water springs became refugia for cold-stenothermous, west-palaearctic species; in the past, these species occupied a larger territory which has been reduced after recent climatic and hydrologic changes.

Description: Larvae of the crabs Menippe mercenaria Say (Menippidae), Panopeus herbstii Milne-Edwards, Neopanope sayi Smith (Xanthidae), Sesarma cinereum Bosc (Grapsidae), and Libinia emerginata Leach (Majidae) were reared in the laboratory. Starvation periods different in length and timing within the first zoeal stage were studied as to their effects on later development and survival rate. After 1-3 days of initial feeding, most larvae had accumulated enough reserves to reach the second stage, independently of further food availability. The development of the survivors was delayed in the following stages, and their later mortality rate was higher than the fed controls. Starvation periods commencing directly after hatching of the larvae exert far stronger negative effects than those beginning later. All observations suggest a particularly sensitive phase in the beginning of larval life in brachyurans. When initial starvation periods exceed the point-of-no-return (PNR), the larvae will die later, even if feeding begins long before the energy reserves are depleted. Temporary lack of suitable prey may be an ecological factor controlling the survival of crab larvae as effectively as physical factors.

Description: Resistance to starvation in early larval stages of six species of brachyuran crabs representing four families was observed at various constant temperatures. In the optimal temperature range of 25-30°C for these warm temperate crab larvae, survival time of starved zoeae was longer than the development duration time in fed zoeae, while at lower temperatures the relationship of these two duration periods became inversed. This response pattern is found in larvae of the mud crab Rhithropanopeus harrisii and is considered to be typical for warm temperature brachyuran larvae. It indicates that reserved utilization is strongly controlled by temperature, but not to the same degree as development.

Description: Rates of food uptake were measured for individually reared larvae of the spider crab Hyas araneus L. feeding on freshly hatched Artemia nauplii at constant 12 degree C. Feeding rates (FR) of crab larvae were given as number of nauplii and amounts of dry weight, carbon, nitrogen, hydrogen, and energy (estimated from C) consumed per day. In both zoeal stages FR increased during postmoult and intermoult, remained high during early and intermediate premoult, and decreased again during late premoult. No clear pattern was found in the course of daily FR of the megalopa. Gross growth efficiencies (K sub(1)) showed a dramatic decrease from postmoult to early premoult (〉 60 to 〈 20%) in both zoeal stages. Daily consumption expressed as % body weight also decreased significantly in these instars. Average daily FR were highest in the zoea II, lowest in the megalopa, and intermediate in the zoea I. Since development of the megalopa took the longest time, the total amount of food consumed by this instar was equal to consumption in both zoeal stages combined.

Description: Duration of development in the larval and early juvenile stages H. coarctatus was studied in relation to temperature, and compared at extreme (18 and 6 °C) than at intermediate (9 to 15 °C) temperatures. The results were used to estimate the duration of development from hatching to the third crab stage in the field. Settling and metamorphosis was predicted to occur mainly during June. Biomass increased exponentially during larval development. Juvenile growth was also exponential and was maximum at 9 degree C, and minimum at 18 and 6 °C.

Description: Statistically significant differences were found in development duration of Hyas araneus L. larvae hatching on different days from the same egg batch. Larvae from different females show a decreasing trend in development time the later they hatch during the season. This trend was found in all larval instars; it was particularly apparent in the megalopa. Development durations in the 2 zoeal stages are positively correlated with each other, i.e. individuals developing slower than the average in the first larval instar tend to delay moulting also in the second instar. There are negative correlations between larval development time in all stages and the size of juvenile crabs, i.e. weak individuals tend to develop more slowly and to become smaller juveniles than the average. These larvae show lower accumulation rates of biomass already during the first zoeal stage. Larval development rates (at 12 °C) were not clearly affected by the temperature prevailing during previous embryonic development, but embryos incubated at higher temperatures tended to become smaller crabs.

Description: Exuvial losses in relation to late premoult matter and energy, and in relation to growth achieved during each instar, were studied in laboratory-reared larvae and early juveniles of the decapod H. araneus (L.). Changes of composition during development were measured in the complete body and in the exuvia from hatching through the second crab stage. Rates of exuvial loss increased during development in all parameters measured. They were generally highest in inorganic carbon and lowest in N. six to 7% of late premolte energy was lost by moulting zoeae, i.e. 9 to 13% of the energy produced during these stages. The megalopa lost 13%, and juveniles 19 to 20% of their LPRM energy ( similar to 29 to 41% of growth). During complete larval development of H. araneus a total of 18% of produced energy was lost at ecdysis. The same amount had been reported in the literature for larval development of 3 other decapod species.

Description: The influence of continuous and differential transitory starvation on the moult cycle and morphogenesis of H. araneus L. larvae was studied in laboratory experiments. Larvae starved from hatching (zoea I) or from moulting to later instars (zoea II, megalopa) develop, independently of food supply, to Stage C (intermoult). Postmoult Stages (A and B) and parts of intermoult are completed by utilizing internal body reserves under such conditions but cuticle formation is terminated at an advanced but incomplete stage within intermoult. In the zoea-II instar there is morphogenesis in appendages (pereiopod and pleopod buds) during continuous starvation. This supports the hypothesis that moult cycle and morphogenesis may be partly independent processes which are normally synchronized.

Description: Early diagenetic ultrastructural alterations of benthic foraminifers of the genera Elphidium and Ophtalmina from the shallow water sediments of the Kiel Bight were investigated by scanning electron microscopy. Pure solution patterns were deduced from supplementary experiments.Several carbonate destroying processes can be specified by ultrastructural patterns of the shell surfaces. Based on these patterns three zones are established, each showing different mechanisms of shell fragmentation: 1) zone of abrasion, 2) zone of disintegration, 3) zone of corrosion. This zonation depends on the water depth and is caused primarily by water agitation and by undersaturation of the bottom water with respect to carbonate.