ALBERT

Description: Locating fuel treatments with scarce resources is an important consideration in landscape-level fuel management. This paper developed a mixed integer programming (MIP) model for allocating fuel treatments across a landscape based on spatial information for fire ignition risk, conditional probabilities of fire spread between raster cells, fire intensity levels, and values at risk. The fire ignition risk in each raster cell is defined as the probability of fire burning a cell because of the ignition within that cell. The conditional probability that fire would spread between adjacent cells A and B is defined as the probability of a fire spreading into cell B after burning in cell A. This model locates fuel treatments by using a fire risk distribution map calculated through fire simulation models. Fire risk is assumed to accumulate across a landscape following major wind directions and the MIP model locates fuel treatments to efficiently break this pattern of fire risk accumulation. Fuel treatment resources are scarce and such scarcity is introduced through a budget constraint. A test case is designed based on a portion of the landscape (15 552 ha) within the Southern Sierra fire planning unit to demonstrate the data requirements, solution process, and model results. Fuel treatment schedules, based upon single and dual wind directions, are compared.

Description: This paper examines the problem of harvest capacity planning at a tactical level. Annual capacity planning allows planners to determine the number of contractors to hire per period throughout the year and to define the duration of their contracts. In practice, this process usually involves the analysis of historical data regarding the operational use of capacity and aggregated demand forecast, the output of which then serves to plan harvest operations. Although this form of hierarchical planning reduces the complexity of the task, the decomposition into subproblems that must be successively resolved can lead to infeasibility or poor use of harvesting capacity. The specific problem addressed here resides in how one can consider the operational impact of harvesting decisions taken at the tactical level to ensure a plan’s feasibility at the operational level. We present a tactical planning process based on Schneeweiss’ generic hierarchical modeling approach. A computational experiment demonstrates how a tactical planning process is influenced by the input of the operational level anticipation model. The anticipation approach we propose appears to be a valid method to better integrate key operational-level decisions into tactical plans.

Description: Spatial tree data are required for the development of spatially explicit models and for the estimation of summary statistics such as Ripley’s K function. Such data are rare and expensive to gather. This paper presents an efficient method of synthesizing spatial tree point patterns from nearest neighbour summary statistics (NNSS) sampled in small circular subwindows, which uses a stochastic optimization technique based on simulated annealing and conditional simulation. This nonparametric method was tested by comparing tree point patterns, reconstructed from sample data, with the original woodland patterns of three structurally different tree populations. Analysis and validation show that complex spatial woodland structures, including long-range tree interactions, can be successfully reconstructed from NNSS despite the limited range of the subwindows and statistics. The influence of the NNSS varies depending on the woodland under study. In some cases, the sampling results can be improved by reconstruction. Furthermore, it is clearly shown that it is possible to estimate second-order characteristics such as Ripley’s K function from small circular subwindows through the reconstruction technique. The results offer new opportunities for adding value to woodland surveys by making raw data available for further work such as growth projections, visualization, and modelling.

Description: The relative importance of fire and flooding on the population dynamics of eastern white-cedar ( Thuja occidentalis L.) and black ash ( Fraxinus nigra Marsh.) was evaluated in eight old-growth riparian stands of southwestern boreal Quebec, Canada. Rising water levels and decreasing fire frequency since the end of the Little Ice Age (ca. 1850) were expected to have favoured an inland migration of the riparian forest fringe, with the flood-tolerant black ash colonizing the lower parts of the shore terraces and eastern white-cedar the upper parts. Black ash was found to be restricted to the riparian zone (

Description: A forest plot with a clustered spatial pattern of tree locations was used to investigate the impacts of different kernel functions (fixed vs. adaptive) and different sizes of bandwidth on model fitting, model performance, and spatial characteristics of the geographically weighted regression (GWR) coefficient estimates and model residuals. Our results indicated that (i) the GWR models with smaller bandwidths fit the data better, yielded smaller model residuals across tree sizes, significantly reduced spatial autocorrelation and heterogeneity for model residuals, and generated better spatial patterns for model residuals; however, smaller bandwidth sizes produced a high level of coefficient variability; (ii) the GWR models based on the fixed spatial kernel function produced smoother spatial distributions for the model coefficients than those based on the adaptive kernel function; and (iii) the GWR cross-validation or Akaike’s information criterion (AIC) optimization process may not produce an “optimal” bandwidth for model fitting and performance. It was evident that the selection of spatial kernel function and bandwidth has a strong impact on the descriptive and predictive power of GWR models.

Description: The rate of spread of Armillariaostoyae (Romagnesi) Herink was calculated from observations over 20 years on plots in two Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) plantations in the southern interior of British Columbia. Average rates of spread in active disease centers ranged from 0.7 to 1.3 m/year.

Description: Using a mail questionnaire targeted at 500 softwood sawmills in the United States and Canada, firm innovativeness was assessed using three methods: (1) current technology, (2) self-evaluation, and (3) a new scale — the propensity to create and adopt scale. The results of these three methods were then compared to assess the performance of each method. Additionally, the relationship between firm innovativeness and financial performance was examined. Based on responses from 85 sawmills (19% adjusted response rate), the results show that both the self-evaluated and the propensity to create and adopt measures differentiate between mills with high and low levels of innovativeness. The composite of the propensity to create and adopt scale shows higher reliability (Chronbach’s α = 0.97) than the self-evaluated scale (Chronbach’s α = 0.68). Significant relationships between sawmill performance and each of the three measures of innovativeness were seen, with the propensity to create and adopt scale generally having the strongest positive relationships. Current technology was significantly related to sales growth, but not gross profit.

Description: Thinning and thinning followed by prescribed fire are common management practices intended to restore historic conditions in low-elevation ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) forests of the northern Rocky Mountains. While these treatments generally ameliorate the physiology and growth of residual trees, treatment-specific effects on reproductive output are not known. We examined reproductive output of second-growth ponderosa pine in western Montana 9 years after the application of four treatments: thinning, thinning followed by spring prescribed fire, thinning followed by fall prescribed fire, and unthinned control stands. Field and greenhouse observations indicated that reproductive traits vary depending on the specific management treatment. Cone production was significantly higher in trees from all actively managed stands relative to control trees. Trees subjected to prescribed fire produced cones with higher numbers of filled seeds than trees in unburned treatments. Seed mass, percentage germination, and seedling biomass were significantly lower for seeds from trees in spring burn treatments relative to all others and were generally higher in trees from fall burn treatments. We show for the first time that thinning and prescribed-burning treatments can influence reproductive output in ponderosa pine.

Description: Airborne laser scanning (lidar) technology is increasingly being applied in forest ecosystem surveys. This research note proposes a design-based approach for the lidar-assisted estimation of forest standing volume when ground surveys are performed by means of fixed-area plots. The lidar measurement of the height of the upper canopy (digital crown model) is performed for the whole study area, and the resulting pixel heights are adopted as auxiliary information to couple with the standing volume acquired on the ground by means of sample plots. The ratio estimator for the total volume of the forest is derived in a complete design-based framework together with an unbiased estimator of its sampling variance and the corresponding confidence interval. The proposed procedure has been tested in Bosco della Fontana, a lowland forest in Northern Italy, obtaining a 95% confidence interval for the total volume, which is approximately 2/3 smaller than that obtained by solely using information arising from field plots.

Description: An approach for including biological diversity in calculations of multiobjective forest planning is described. This makes it possible to consider biodiversity as a decision objective among multiple objectives in tactical forest planning. Biodiversity is operationalized by decomposing it into measurable environmental components that describe its dimensions. The relative importance of the components is assessed via pairwise comparisons. The relationship between the value of the component and the overall biodiversity is estimated and described for each component as a subpriority function. The relationship can be nonlinear. A biodiversity index is calculated for each alternative forest plan based on the components and their weighting, the subpriority functions, and the predicted qualities of the forest area when implementing the plans. The approach is illustrated by a case study. Using this approach, the relationship between biodiversity and the other objective variables, as well as the effects of the components of biodiversity, can be analyzed numerically.

Description: Natural stands and a 3-year-old plantation of red alder (Alnusrubra Bong.) trees were used to study the incidence of leaning stems, the level of growth stresses and tension wood formation, and the ability of the stems to right themselves to vertical. Overall, 10% of the 512 trees in 10 natural stands leaned 〉22°. The largest diameter trees on the steepest slopes leaned most. Most (61%) of the trees curved upward, showing a righting response. For samples without tension wood, growth stress levels on the upper side of leaning stems, but not on the lateral or lower sides, were positively correlated with lean angles above 6°. These leaning stems had a significant righting response without tension wood. Tension wood formation was variable at leans from 9° to 26° both within and among trees, but was correlated with eccentric growth rings. We measured stem recovery in the year-old stem of 3-year-old trees bent to angles of 0–37.5°. During the 5-month experiment all stems righted to near vertical. Tension wood formed on the upper side in stems bent 〉6°, but reversed to the lower side before reaching vertical in 22 of 30 trees.

Description: We analyzed spatial patterns of overstory trees in late-successional Abies amabilis (Dougl. ex Loud.) Dougl. ex J. Forbes forests and late-successional Pseudotsuga menziesii (Mirb.) Franco forests to establish reference spatial patterns for restoration thinning treatments, and to determine whether thinning treatments with minimum intertree spacing rules result in spatial patterns characteristic of late-successional forests. On average, 32.7% of overstory trees in Abies plots and 26.3% of overstory trees in Pseudotsuga plots occurred as members of multitree clusters (groups of trees in which trees are spaced within a specified minimum distance of each other) at a distance of 3.0 and 4.0 m, respectively. Multitree clusters occurred throughout the three Abies plots; the distribution of multitree clusters within the two Pseudotsuga plots was variable. Spatial patterns of overstory trees in late-successional forests were significantly different from those created by simulated restoration thinning treatments. Restoration thinning treatments that release both individual trees and multitree clusters promote characteristic late-successional tree spatial patterns at the within-patch scale (

Description: This paper uses hedonic regression techniques to analyze timber bid transactions in central Georgia. Softwood stumpage prices from pay-as-cut transactions are regressed against timber sale and stand characteristics. We identify observable factors that are statistically associated with the volatility of pine sawtimber stumpage prices in the market. The remaining price volatility, defined as market risk, characterizes undiversifiable price volatility in the market. Isolating market risk in this way has implications for relative price risk across predefined timber markets. Applications of this these techniques suggest that analyzing market price variability with total measures alone, such as standard deviation, may provide false senses of timber price risk.

Description: Height observations H1and H2present on the right- and left-hand sides of site index models, respectively. The error terms associated with H1and H2, along with parameter estimate errors, affect the estimate of the site index. Projection error variance (PEV), in a projection from A1to A2, consisted of four components associated with H1, H2, the covariance of H1and H2, and the parameter estimate errors. In this study, behaviors of these components were investigated via simulations on the basis of six equations derived from the Lundqvist–Kerf and the Hossfeld IV functions. Simulation results showed that projection interval, projection direction, and selected site-dependent parameter influenced PEV and its components. PEVs of backward and forward projections with the same projection interval lengths were remarkably different if the underlying model was anamorphic. With increasing projection interval length, the PEV of forward projections monotonically increased to a certain value, whereas the PEV of backward projections decreased to zero after reaching a maximum.

Description: Growth modeling of forests at the individual tree and stand levels is a highly refined procedure for many forest types. A method to incorporate predictions from such models into a forest inventory system is developed. Variance components from the actual measurements and from the predicted measurements are used to estimate the variance of the combined predicted value. The only assumption required to justify this method is that the model estimate has a bias that does not change from one time period to the next. The estimation procedure proposed here can also incorporate remotely sensed information via a regression estimator.

Description: We describe the development of a statistical model of spatial variation in the area burned by lightning-caused forest fires across the province of Ontario. We partitioned Ontario’s fire region into 35 compartments, each of which is relatively homogeneous with respect to its vegetation, weather, and the level of fire protection it receives. We used linear regression and spatial autoregressive models to relate the average annual area burned in a compartment to its vegetation, weather, and level of protection attributes. We also examined the relationship between burned area and the level of protection in two areas that are relatively homogeneous with respect to vegetation and weather. We found a statistically significant relationship between the average annual fraction of the area of a compartment burned by lightning-caused forest fires and its vegetation, weather, and the level of fire suppression effort it receives.

Description: Phenolic metabolites are frequently implicated in chemical defense mechanisms against pathogens in woody plants. However, tree breeding programmes for resistance to pathogens and practical tree-protection applications based on these compounds seem to be scarce. To identify gaps in our current knowledge of this subject, we explored some of the recent literature on the involvement of phenolic metabolites in the resistance of northern forest trees (Pinus, Picea, Betula, Populus, and Salix spp.) to pathogens. Although it is evident that the phenolic metabolism of trees is often activated by pathogen attacks, few studies have convincingly established that this induction is due to a specific defense response that is capable of stopping the invading pathogen. The role of constitutive phenolics in the resistance of trees to pathogens has also remained unclear. In future studies, the importance of phenolics in oxidative stress, cell homeostasis and tolerance, and the spatial and temporal localization of phenolics in relation to invading pathogens should be more carefully acknowledged. Possibilities for future studies using advanced methods (e.g., metabolic profiling, confocal laser scanning microscopy, and use of modified tree genotypes) are discussed.

Description: Due to the scarcity of old-growth forests in much of Europe, there is little quantitative information on disturbance processes that influence forest dynamics. However, this information is crucial for forest management that tries to emulate patterns and processes in natural forests. We quantified the gap disturbance regime in an old-growth forest dominated by European beech ( Fagus sylvatica L.) and silver fir ( Abies alba Miller) in the Dinaric Mountains of Bosnia and Herzegovina. We sampled 87 gaps in four stands using line-intercept sampling. The percentages of forest area in canopy gaps and expanded gaps ranged from 12% to 17.2% and 35.5% to 39.7%, respectively. Although many of the gaps were small (1000 m2 with numerous gapmakers made up a disproportionate amount of the total gap area. More than half the gaps had more than one gapmaker and were often in separate decay classes, indicating gaps had expanded over time during separate disturbance events. Furthermore, 51% of all gapmakers were uprooted or wind-snapped, whereas only 22% died standing. These results suggest that wind disturbance plays an important role in creating intermediate to large canopy openings through both gap formation and gap expansion processes.

Description: European beech ( Fagus sylvatica L.) is increasingly managed by close-to-nature principles, mimicking the gap dynamics of seminatural forests. The prime aim of this study was to analyse natural regeneration reliability under favourable conditions in newly formed gaps. A total of 12 gaps were created by felling three canopy trees for each gap: six gaps in each of the two winters 1996–1997 and 1997–1998. One-half of the gaps were fenced against deer. We recorded advance regeneration density (1997), regeneration density and height (1997–2002), relative light intensity (1997–2002), and volumetric soil moisture content (1997–2002). We also studied the effect of year of establishment, fenced versus unfenced, and position within gap on regeneration. Three or 4 years after gap formation, most gaps had nearly closed. Response of European beech, European ash ( Fraxinus excelsior L. ), and sycamore maple ( Acer pseudoplatanus L.) regeneration to gap formation was limited, and few seedlings were added to the advance regeneration pool during the study period. Other factors, such as relative light intensity, soil moisture, fencing, year of establishment, and position within gaps, all had rather low effects. Thus, the presence of advance regeneration appeared to be a key factor in explaining regeneration patterns in artificially created gaps.

Description: Increased forest productivity has been obtained by improving resource availability through water and nutrient amendments. However, more stress-tolerant species that have robust site requirements do not respond consistently to irrigation. An important factor contributing to robust site requirements may be the distribution of biomass belowground, yet available information is limited. We examined the accumulation and distribution of above- and below-ground biomass in sweetgum ( Liquidambar styraciflua L.) and loblolly pine ( Pinus taeda L.) stands receiving irrigation and fertilization. Mean annual aboveground production after 4 years ranged from 2.4 to 5.1 Mg·ha–1·year–1 for sweetgum and from 5.0 to 6.9 Mg·ha–1·year–1 for pine. Sweetgum responded positively to irrigation and fertilization with an additive response to irrigation + fertilization. Pine only responded to fertilization. Sweetgum root mass fraction (RMF) increased with fertilization at 2 years and decreased with fertilization at 4 years. There were no detectable treatment differences in loblolly pine RMF. Development explained from 67% to 98% of variation in shoot versus root allometry for ephemeral and perennial tissues, fertilization explained no more than 5% of the variation in for either species, and irrigation did not explain any. We conclude that shifts in allocation from roots to shoots do not explain nutrient-induced growth stimulations.

Description: We examined the impacts of a severe, regional drought (2001–2002) on trembling aspen ( Populus tremuloides Michx.) forests in the western Canadian interior. A total of 150 plots were established in 25 study areas as part of a regional-scale study (CIPHA). Aspen health and mortality were assessed annually during 2000–2005, and changes in stem biomass were estimated using tree-ring analysis and plot-based measurements. Net mean increment in living biomass for all plots was 2.2 t·ha–1·year–1 during 2000–2002 but subsequently decreased to near zero. This collapse was driven by a more than two-fold increase in stem mortality and a 30% decrease in regional stem growth during and following the drought. The analysis showed that spatial variation in aspen productivity and biomass across the region was positively related to multiyear mean values of a climate moisture index and mineral soil silt content but was negatively related to levels of insect defoliation and wood-boring insects. In contrast, mortality and dieback was best correlated with minimum annual climate moisture index, which provided a measure of short-term drought severity. The results support previous studies showing that aspen forests are moisture limited in this region, which poses concerns for the future under a changing climate.

Description: Trends in gap dynamics among pole, mature, and old-growth northern hardwood stands were investigated on eight sites in the Porcupine Mountains of western upper Michigan. Recent gaps (created between 1981 and 1992) were identified using permanent plot records of tree mortality, while older gaps (1940–1981) were identified using stand reconstruction techniques. Although canopy gaps were somewhat more numerous in pole and mature stands, gaps were

Description: Controlled, localized heating was applied to the trunks of young Aleppo pines (Pinushalepensis Mill.) during the winter. Three levels of burning were applied to the circumference of the trunks (zero, partial, or total), and the effects on tree physiology and growth were monitored during the 7 months following heating. Effects of the heat treatment on tree physiology were assessed by measuring predawn water potential, sap flux density, microvariations in trunk diameter, electrical resistance of the cambium, and gas exchanges from the needles (photosynthesis and stomatal conductance). Morphological response measurements included height and radial growth of the trees and morphological characteristics of the needles. The amount of chlorotic foliage was also measured. Aleppo pine was highly resistant to trunk injury, surviving when 80% of the trunk circumference was destroyed by fire. The behaviour of partially burnt pines was not significantly different from that of controls, for all variables studied. Trees whose trunks were totally burnt died from 1 to 5 months after treatment. Sap flux density of totally burnt trees was already significantly lower than that of other trees in the first week following the treatment. A decrease in net photosynthesis and stomatal conductance in the still surviving totally burnt trees was also recorded during the first measurements made in July. Treatment differences in predawn water potential, microvariations in trunk diameter, and electrical resistance of the cambium only appeared a few days before death of the totally burnt trees.

Description: The mechanisms governing tree mortality in surface fires are poorly understood, owing in large part to the absence of a process-based framework for defining and evaluating these mechanisms. This paper begins the development of such a framework by deriving a first-order process model of tree mortality in surface fires (intensities less than approximately 2500 kW·m–1). A buoyant line-source plume model is used to drive heat transfer models of vascular cambium and vegetative bud necroses, which are linked to tree mortality using an allometrically-based sapwood area budget. Model predictions are illustrated for white spruce ( Picea glauca ), lodgepole pine ( Pinus contorta ), and trembling aspen ( Populus tremuloides ) and are compared with independent mortality data for Engelmann spruce ( Picea engelmannii Parry ex Engelm.) and Pinus contorta Dougl. Results help define first-order mortality mechanisms and suggest second-order mortality mechanisms that should be incorporated into future modeling efforts.

Description: Mean annual acorn crop size of oaks in central coastal California correlated with weather for four of five species. For the deciduous Quercuslobata and Quercusdouglasii, larger crops were produced in years when April temperatures were warmest, while for the evergreen Quercusagrifalia and Quercuschrysolepis, crops were larger when there was more rain one and two years prior to the year when acorns were produced. No correlations were found between weather and mean annual acorn production by Quercuskelloggii. Within years, individual Q. douglasii that flowered during periods of greater solar radiation produced more acorns, and during 1 of 5 years individual Q. lobata and Q. agrifolia produced larger crops when they flowered during warm, dry periods. These results indicate that conditions favorable for pollination and fertilization strongly enhance mean annual acorn production in Q. lobata and Q. douglasii and may have an effect on differences in acorn crop size among individual Q. lobata, Q. douglasii, and Q. agrifolia. Mean annual acorn crops of these three species were significantly concordant among three sites in central coastal California 300 km apart. Weather conditions were also significantly correlated, especially annual rainfall and mean April temperatures. Thus, the environmental factors that have the strongest influence on mean annual acorn production within a site are also those that are the most geographically concordant. However, we found no relationship between environmental factors and differences in mean annual acorn production by these same three species at the three sites.

Description: The purpose of this study is to investigate the suitability of the Birnbaum–Saunders distribution to model diameter at breast height (DBH) distributions of near-natural complex structure silver fir ( Abies alba Mill.) – European beech ( Fagus sylvatica L.) forests. The investigations were carried out in Świętokrzyski National Park, situated in Central Poland. To estimate the parameters of the Birnbaum–Saunders distribution, three methods were used: the maximum likelihood method (MLE) and the mean–mean estimator, the modified moment method (MME), and the graphical method (GME). The empirical DBH distributions in near-natural fir–beech stands, which arose according to the model taking into account the overlapping of fir mortality and beech regeneration, were generally conformed to the Birnbaum–Saunders distribution. In such forests, the Birnbaum–Saunders distribution approximated the empirical DBH distributions more precisely than the Weibull and gamma distributions.

Description: We sampled second-growth forests ranging in age from 28 to 98 years and compared them with old-growth forests to quantify rates of terrestrial vegetation recovery following harvesting on the northcentral coast of British Columbia. Species richness approximately doubles, while Simpson’s index of diversity increases from 0.81 to 0.91 from young to old forests. Nonmetric multidimensional scaling ordinations showed differentiation, with some overlap, of old-growth and second-growth forests and a fairly strong correlation of stand age with plot scores, driven by plant species presence and cover. Vegetation succession following logging disturbance is driven primarily by predisturbance species composition; most species found in the young forests are present in old forests and the higher species richness typical of old growth is largely due to the establishment of additional cryptogam and herb species of low cover and constancy. Significantly higher cover of shrub, herb, and bryophyte species differentiates old forests from second-growth forests. Forests 41–100 years old average 63%–73% similarity (depending on site type) to old-growth forests based on species presence–absence and 53%–58% similarity based on species cover. The scarcity of western redcedar ( Thuja plicata Donn ex D. Don) in second-growth stands is of particular concern because of the high ecological, cultural, and economic importance of this tree species.

Description: Buffer zones are an important component of forest-management strategies and are thought to reduce the impact of nutrients released after harvesting on water quality. Conceptually, steep slopes have shorter water residence times than shallow slopes, have a reduced capacity to moderate water quality, and therefore, require wider buffers. Carbon and N concentrations in riparian zone shallow soil water at 30 cm depth and lake water were measured on shallow and steep slopes at the Esker Lakes Research Area in northeastern Ontario to determine if nutrient concentrations were correlated to catchment terrain attributes. Field measured slope, slope class obtained from a triangular irregular network model, and upslope contributing area and topographic index calculated from a digital elevation model were calculated for each sampling location. Modeled terrain properties, including those currently used during forest-management planning, were not significantly correlated with soil water N and C concentrations, whereas only dissolved organic carbon levels were significantly greater on field measured steep slopes. Forest species composition and soil N levels were positively correlated with soil water N concentrations. These results from the undisturbed boreal ecosystem highlight the potential limitation of using only catchment slope as a tool for prescribing riparian buffers during harvesting when considering terrestrial nutrient export.

Description: In forest breeding programmes, growth has typically been used as a selection trait of prime importance in Norway spruce ( Picea abies (L.) Karst.), whereas less attention has been given to the wood and fibre characteristics. In the above context, we investigated phenotypic relationships between different fibre properties and growth and wood density traits in 20 cloned Norway spruce based on a clonal trial established in the 1970s in southeastern Finland. We found that fibre width showed, on average (2.9%), the lowest phenotypic variation followed by fibre wall thickness (3.4%), coarseness (5.5%), and fibre length (8.1%). All of the phenotypic correlations between the fibre properties were also positive (p 〈 0.05), ranging from moderate to strong, suggesting that selection for one trait could simultaneously affect the other traits. The phenotypic correlations, on average, were quite weak but positive between growth and fibre properties and slightly negative or weak positive between wood density and different fibre properties (p 〈 0.05). Individually, some of the clones showed negative correlation between growth traits and fibre length. As a result, selection for fibre properties alone could also reduce overall stem volume (or stem mass) and would not directly indicate wood density traits and vice versa.

Description: Total amount and vertical distribution of foliage represent important aspects of forest stand structure and its influence on dry matter productivity, forest microclimate, watershed properties, and habitat structure. Variation in foliage distribution was analyzed on trees and plots in a series of even-aged Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stands scheduled for management under a wide range of silvicultural regimes. Branch-level foliage mass and foliage area equations were developed from a sample of 138 branches. These equations were applied to 27 trees on which the diameter and height of all live primary branches were measured, allowing estimation of both the total amount of foliage and its vertical distribution. A β-distribution was fitted to data describing the vertical distribution of foliage on each tree, and the resulting parameter estimates were modelled as functions of tree height, diameter at breast height, crown length, and relative height in the stand. Foliage area distribution tended to be shifted downward relative to foliage mass because of the expected increase in specific leaf area with depth into the crown. Similarly, the relative foliage distribution in terms of both mass and area was shifted downward as the tree became more dominant, or as relative height in the stand increased. In contrast, foliage on trees of similar relative height was shifted upward in response to the lower stand densities imposed by precommercial thinning. On the stand level, relative vertical distribution of foliage in the canopy was more peaked than would be implied by assuming a constant leaf area/sapwood area ratio throughout the composite tree crowns. Between-stand variation in vertical foliage distribution was dictated by differences in stand top height, height to crown base, and number of trees per hectare.

Description: A study was established between 1959 and 1961 to study the long-term responses of balsam fir ( Abies balsamea (L.) Mill.) and red spruce ( Picea rubens Sarg.) to precommercial thinning. Three nominal spacings of 4 ft (1.2 m), 6 ft (1.8 m), and 8 ft (2.4 m) were compared with an unthinned control in a randomized complete block design with five replicates. At the time of thinning, natural regeneration averaged 16 years of age, 8 years after harvest. Although thinning had minimal effect on gross total volume production over a 42 to 44 year observation period, actual spacings between 2.1 and 2.5 m produced an average of 360 m3·ha–1gross merchantable volume (GMV), representing a 21% gain over unthinned stands. The same spacings produced quadratic mean diameters of 21 and 23 cm, respectively, compared with 18 cm in the unthinned stands. These size increases translated to individual stem volume gains of 33% and 62%, significantly reducing the age at which thinned stands would meet a specified minimum requirement for merchantability or habitat. The mean annual increment of GMV ranged from 6 m3·ha–1·year–1in unthinned stands, to more than 7 m3·ha·–1·year–1in the thinned stands, and had not yet culminated an average of 50 years postharvest.

Description: Height, weight, root volume, shoot length, needle length and weight, and total chlorophyll content were measured for both control and tomato mosaic Tobamovirus (ToMV)-inoculated red spruce (Picearubens Sarg.) seedlings in each of five, 3-month growth periods. Mean bud-break rating was determined for each seedling in growth period 5. In January 1995 the freezing tolerance of control and ToMV-infected seedlings was determined. Approximately 85% of the inoculated seedlings became infected with ToMV, as determined by enzyme-linked immunosorbent assay of young root tissue. Reduction in seedling height, weight, root volume, bud break, and mean shoot length of infected seedlings was observed when compared with control seedlings. The rate of increase over time in height, weight, and root volume was lower (54, 57, and 52%, respectively) in infected seedlings compared with control seedlings. Additionally, needles of infected seedlings were less susceptible to freezing damage than those of control seedlings. However, virus infection had no effect on the freezing tolerance of roots.

Description: We measured soil density and tree growth after wet-season, ground-based yarding on fine-textured soils at three clear-cut sites. Four treatment conditions were sampled on or near four skid trails (replicates): nontilled and tilled primary skid trails, and adjacent slash-treated areas; the fourth treatment was secondary skid trails at two locations and a logged-only control at the third location. The 16 treatment plots were split into 4 subplots, each randomly assigned to a species–stock type and planted with 30 seedlings. Tree data through year 8 after planting were analyzed as a randomized block, split-plot design. Compared with nontrail areas, bulk density in the 0- to 8-cm depth on primary skid trails after logging averaged 41–52% greater. Eight years later, bulk density in the 0- to 30-cm depth of primary skid trails still exceeded that outside trails by about 20%, yet tree survival was similar except for western hemlock (Tsugaheterophylla (Raf.) Sarg.) having poorer survival on nontilled trails at one location. Average tree height and volume of Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco var. menziesii), Sitka spruce (Piceasitchensis (Bong.) Carr.), and hemlock (except height at one location) did not differ among treatments. Observed differences among treatment means were small. Power analyses indicated that 20% or larger differences in tree height or volume were detectable with 30–95% power.

Description: not available

Description: Decline index (indicator of crown condition) data from 102 forest plots (approximately 10 000 trees) during 1986–2004 were compiled to derive survival models for south-central Ontario, Canada. The dominant species was sugar maple ( Acer saccharum Marsh.) with approximately 75% occurrence (n = 7640). The predictor variables for sugar maple survivorship included the decline index of 1 or 2 years prior to the beginning of the modelled period and ecological region (Algoma, Georgian Bay, Huron–Ontario, and Upper St. Lawrence). The observed crown condition of sugar maple improved significantly over the study period; in contrast, short-term mortality rate did not improve. The risk of sugar maple mortality could be predicted from decline index data for a single year indicating that the risk of tree death increases with higher decline index values (declining crown condition). Moreover, using 2 years of decline index data indicated that the risk of tree death also increased with the length of consecutive time individual trees have higher decline index values. Trees in the Algoma region, which represent the northern limit of sugar maple distribution in Ontario, were significantly more likely to die than trees in Huron–Ontario region.

Description: Using an extensive Douglas-fir data set from southwest Oregon, we examined the (1) performance and suitability of selected prediction strategies, (2) contribution of relative position and stand-density measures in improving tree height (h) prediction values, and (3) effect of different subsampling designs to fill in missing h values in a new stand using a regional nonlinear model. Nonlinear mixed-effects models (NMEM) substantially improved the accuracy and precision of height prediction over the conventional nonlinear fixed-effects model (NFEM) that assumes the observations are independent, particularly when a few trees are subsampled for height. The predictive performance of a correction factor on a NFEM with relative position and stand-density measures was comparable to that of a NMEM when four or more trees were subsampled for height. When two or more heights were randomly subsampled, the NMEM efficiently explained the differences in the height–diameter relationship because of the variations in relative position of trees and stand density without having to incorporate them into the model. When only one height was subsampled, selecting the largest diameter tree in the stand would result in a lower predicted root mean square error (RMSE) than randomly selecting the height, regardless of the model form or fitting strategy used.

Description: Past years of research indicate that halogenation of organic matter is a general occurrence in soil. As this is a virgin research field, little is known about, for example, the possible relation to the turnover of organic matter, influence of environmental parameters, or ecological role. The aim of the present paper was to study the influence of fertilization on in situ formation of organically bound halogens and its possible relation to decomposition of organic matter. Total amount of organically bound halogens was determined in nutrient-rich and nutrient-poor spruce litter incubated up to 4 years in a fertilized plot and a control plot. It was found that the concentration of organically bound halogens increased throughout the incubation period, and the increase was significantly larger in the control than in the fertilized plot. This resulted in an accumulated increase in the control plot that was approximately twice as large (ca. 200 μg Cl/g dry mass) as that in the fertilized plot at the end of the incubation period. The changes in absolute amounts were complexly related to litter mass loss. This strongly indicates that there are multiple underlying processes and that halogens are both incorporated into and released from organic matter during decomposition. Calculated on a daily basis, the rate at which the amounts of organohalogens changed showed a seasonal pattern, with a pronounced increase during late summer and autumn. After 1 to several years, a net decrease was observed, particularly in the summer period. This decrease appeared earlier and was more pronounced in the fertilized plot than in the control plot, strongly indicating that mineralization of organically bound halogens, i.e., release of inorganic halides, was enhanced by fertilization. In the litter incubated in the control plot, the absolute amounts increased in relation to lignin degradation. In combination with previous findings, the results of the present study suggest that formation of organically bound halogens is conducted by organisms responsible for lignin degradation. Furthermore, the findings indicate that fertilization may favour organisms that degrade organically bound halogens at the expense of organisms that cause the formation of such compounds.

Description: Witness tree data from 1780–1856 for the Monongahela National Forest in eastern West Virginia were analyzed with respect to physiographic unit (Ridge and Valley versus Allegheny Mountains) and landform, and compared with present-day forest composition. Contingency table analysis and standardized residuals were used to quantify the preference or avoidance of common tree species with various landforms. Pre-European settlement forests in the Ridge and Valley were dominated by mixed oak (Quercusalba L., Quercusprinus L., Quercusvelutina Lam. and Quercusrubra L.), Pinus spp., Castaneadentata (Marsh.) Borkh., and Carya on ridge sites and Q. alba, Acersaccharum Marsh., Pinus, Tiliaamericana L., and Tsugacanadensis (L.) Carr. on valley floors. The original forests in the Allegheny Mountains were dominated by Fagusgrandifolia Ehrh., T. canadensis, A. saccharum, Acerrubrum L., Betula spp., and Pinus, with predominantly Fagus–Tsuga–Pinus forests on mountain tops and Tsuga–Acer–Betula forests on valley floors. Compared with the presettlement era, present-day forests on both physiographic units lack overstory C. dentata and have decreased Pinus and (or) Q. alba. Species that have increased substantially following Euro-American settlement include Q. prinus, Q. rubra, Quercuscoccinea Muenchh., and A. rubrum in the Ridge and Valley and Prunusserotina Ehrh., A. rubrum, and Betula spp. in the Allegheny Mountains. These dramatic changes in forest composition were attributed to the chestnut blight (caused by Endothiaparasitica (Murrill) P.J. Anderson & H.W. Anderson), widespread logging, intensive wildfires, and more recently, fire exclusion.

Description: Using results from the field and laboratory, we give a detailed explanation of an important and commonly observed ecological phenomenon, fire scarring of trees. Four key observations associated with the formation of fires scars are explained using fluid dynamics and heat transfer processes. When a fire passes by a tree, its height increases on the tree's leeward side because of the occurrence of two leeward vortices. The flame height increases in the vortices because the turbulent mixing of fuel and air is suppressed. The flow of gaseous fuel in the vortices becomes greater than the rate of mixing with the air, and hence there is an increased height along which combustion can occur. Fire scars are found only on the leeward side of trees because the vortices increase the residence time of the flame on the leeward side of the tree compared with the residence time of the flame on the tree's windward side. This leads to differential heating around the tree bole. Small trees rarely have fire scars because their cambium is usually completely killed by a passing fire and (or) their foliage is killed by crown scorch. Fire scars are usually triangular in shape; wider at the base and decreasing in width with height. The triangular shape is a result of the temperature isotherms within the standing leeward flame, which are triangular in shape.

Description: Seedling stands of loblolly pine (Pinustaeda L.) and sweetgum (Liquidambarstyraciflua L.) were grown in monoculture or mixed stands for two growing cycles in controlled-environment chambers. Treatments consisted of ambient (408 ppm) and elevated (806 ppm) CO2, concentrations, water-stressed and well-watered conditions, and low (20 kg N/ha) and high (215 kg N/ha) nitrogen application rates. Photosynthesis rates were measured under ambient and elevated cuvette CO2 concentrations for both whole stands and individual seedlings from these stands. Significant interactions between CO2 and water suggested that elevated CO2 concentration compensated for low water availability in individually measured loblolly pine and in whole seedling stands regardless of stand type. Expressing photosynthesis on a soil area versus a leaf-mass basis influenced the photosynthetic rankings of the three stand types relative to one another. Net photosynthetic rates per unit leaf mass were 390 and 880% higher in individually measured seedlings than in whole monoculture stands for loblolly pine and sweetgum, respectively. Lower photosynthetic contributions from lower canopy leaves in whole seedling stands compared with the upper canopy leaves used in individual-seedling measurements were thought to be responsible for lower photosynthetic rates in seedling stands. These results suggest that photosynthetic response is influenced by canopy dynamics that are unaccounted for by individual-seedling measurements of photosynthesis. Differences in photosynthetic response between loblolly pine and sweetgum stands and individuals are thought to be largely due to species-specific differences in canopy light extinction characteristics.

Description: We studied growth, crown architecture, and specific leaf area acclimation of a shade-intolerant species, Pinuscontorta Dougl. ex Loud. var. latifolia Engelm., and a moderately shade-tolerant species, Pseudotsugamenziesii var. glauca (Beissn.) Franco, using naturally regenerated saplings along a wide range of light conditions. Study sites were located within a dry-summer, cool-temperate climate represented by the Dry Cool Interior Douglas-fir biogeoclimatic subzone near Williams Lake, British Columbia, Canada. In an open light environment, Pinuscontorta had a greater growth increment in both terminal and lateral shoots than did Pseudotsugamenziesii. With decreasing light availability (i) terminal increment, mean lateral increment, and total lateral increment decreased in both species (although Pinuscontorta reduced lateral growth significantly faster than Pseudotsugamenziesii); (ii) the ratio of mean lateral to terminal increment in Pinuscontorta did not change, but significantly increased in Pseudotsugamenziesii; (iii) the ratio of total lateral to terminal increment in Pinuscontorta decreased, but increased in Pseudotsugamenziesii; and (iv) specific leaf area in both species increased (in Pseudotsugamenziesii, it was always higher and increased marginally faster than in Pinuscontorta). We concluded that the less shade-tolerant Pinuscontorta was less plastic in crown architecture and specific leaf area than the more shade-tolerant Pseudotsugamenziesii.

Description: The purpose of the study was to investigate fine-root growth in gaps created for beech (Fagussylvatica L.) regeneration. Fine-root growth was measured using the ingrowth core technique. Measurements were carried out in gaps 30 m in diameter, which were either untreated or treated with lime, and in a mature beech stand. Ingrowth core experiments showed that growth of beech fine roots in gap centres was negligible during the 2nd and 3rd year after gap creation, indicating that although fine roots from stumps stayed alive long after trees were cut, they did not grow. It also indicated that trees surrounding gaps did not effectively grow fine roots that reached 10 m into the gap centre. At the edge of unlimed gaps (5 m away from the stems), fine-root growth was one-third that of the mature stand. In the stand the amount of live fine roots in ingrowth cores (390 g•m−2) had attained the standing crop level after 16 months. In limed gaps, where herbaceous vegetation had established, herbaceous root growth was 800–970 g•m−2 after 16 months. Neither fine-root growth nor aboveground biomass of herbaceous plants was substantial in untreated gaps. The slow recovery of biomass production in unlimed gaps showed that the resistance of this beech forest to nutrient losses following disturbance is low.

Description: Effective fire suspension in Fennoscandian boreal forests has caused a number of species to become threatened. To compensate for the negative ecological impacts of fire elimination, prescribed burning of forests as a restoration method has been introduced recently. We studied the effects of controlled burning on assemblages of wood-decaying polypores (Basidiomycota), including red-listed species, in a large-scale field experiment in Finland. A total of 24 forest sites were included in the factorial study design with two factors: logging and burning. The presence of polypore fruiting bodies was documented 1 year before the treatments, and 1 and 4 years after the treatments. Over 11 000 observations of 104 species of polypores were made. Change in the fungal species composition due to logging and burning was clear after 4 years. At the species level, the responses to logging and fire varied depending on the species. Treatments increased fruiting of pioneer decayers; however, most red-listed species seemed to suffer. Thus, prescribed burning does not offer immediate benefits for most red-listed species. In unlogged forests, the restorative effects of fire are likely to be seen later as the death and decay processes of trees continue and provide more resources for polypores.

Description: We used General Land Office survey data (1860–1890) and interpreted aerial photography from the 1930s, 1970s, and 1990s to quantify forest disturbance frequency and spatial patterns for four time periods in the Mixed Forest Province of Minnesota. The study region included eight subsections within the Mixed Forest Province of Minnesota’s Ecological Classification System. Presettlement disturbance and spatial pattern estimates varied across the eight subsections indicating a strong relationship to soil and landform characteristics. Land surveyors primarily recorded higher severity disturbances that resulted in significant tree mortality. The 1900–1940 era was characterized by a short-term increase in fire frequency that was relatively uniform across the study region, in contrast to the variability of the presettlement (1860–1890) landscape. In the postsettlement period (1940–1995), timber harvest replaced fire as the dominant disturbance factor. Similar management practices among subsections created similar harvest rates throughout the study region. These management practices imposed a more homogeneous pattern dominated by small (10–25 ha) patches. Management practices now have a greater influence than natural processes in the generation of landscape pattern in the Mixed Forest Province of Minnesota. Information on presettlement forest conditions and subsequent changes can be used by land managers to restore spatial pattern variability in managed forest landscapes.

Description: Fine root production and turnover were studied in hardwood and coniferous taiga forests using three methods. (1) Using soil cores, fine root production ranged from 1574 ± 76 kg•ha−1•year−1 in the upland white spruce (Piceaglauca (Moench) Voss) stand to 4386 ± 322 kg•ha−1•year−1 in the floodplain balsam poplar (Populusbalsamifera L.) stand, accounting for 49% of total production for coniferous stands and 32% of total production for deciduous stands. Fine root turnover rates were higher in floodplain (0.90 ± 0.06 year−1) stands than in upland (0.42 ± 0.10 year−1) stands. Across all sites, the ratio of fine root turnover to litter fall averaged 2.2 for biomass and 2.8 for N. Both values were higher in floodplain stands than in upland stands, and in coniferous stands than in deciduous stands. (2) The C budget method showed that C allocation to fine roots varied from 150 to 425 g C•m−2•year−1 and suggested that soil respiration was more dependent on C derived from roots than from aboveground inputs. The C allocation ratio (C to roots: C to litter fall) was inversely correlated with litter-fall C and varied from 0.3 to 69.5; there was a tendency for higher proportional belowground allocation in coniferous stands than in deciduous stands and the highest levels were at the earliest successional sites. (3) Estimates of apparent N uptake (Nu), N allocation to fine roots, and fine root production based on N budget calculations showed that annual aboveground N increments exceeded Nu estimates at half the sites, indicating that the method failed to account for large amounts of N acquired by plants. This suggests that plant and (or) mycorrhizal uptake of soil organic N may be more significant to ecosystem N cycling than mineral N turnover by the soil microbial biomass.

Description: Leaf physiology and stem growth were assessed in loblolly pine ( Pinus taeda L.) in response to 10 to 11 years of treatment with weed control (W), weed control plus irrigation (WI), weed control plus irrigation and fertigation (WIF), or weed control plus irrigation, fertigation, and pest control (WIFP) to determine whether increased resource availability can push productivity of loblolly pine closer to its biological growth potential expressed in favorable, exotic environments. Maximum basal area and stem biomass were 41 m2·ha–1 and 172 Mg·ha–1, respectively, in response to fertigation. Stemwood biomass production was positively and linearly related to basal area. Belowground woody biomass was highest in the WIF and WIFP treatments and averaged 50 Mg·ha–1, but the W and WI treatments exploited a greater area of soil with low-density coarse roots. Fertigation increased foliar nitrogen concentration and foliage biomass, but treatment had no effect on leaf physiological parameters or growth efficiency. Comparison with growth rates reported for loblolly pine in Hawaii revealed that loblolly pine grown in its native range can produce the high yields observed in exotic environments when stands are below maximum carrying capacity.

Description: Light is a key resource controlling tree regeneration in the understory of closed-canopy old-growth forests. To evaluate the distribution of understory light environments at a landscape scale, we used stratified random sampling in a 500-ha stand of Costa Rican tropical rain forest. Fifteen 100 m long transects were placed using random coordinates within two soil–geomorphology units (flat alluvial terraces and dissected ridge-slope-swale terrain). At 2.5-m intervals we measured canopy height and slope angle, classified topographic position, and took canopy photographs with a fish-eye lens at 1 and 3 m above the ground (and at 0.6 and 5 m height at five stations per transect). Photographs were analyzed for global site factor (GSF), which is analogous to the percentage of full sun radiation reaching a point. Canopy height and GSF at 1 and 3 m above the ground were significantly autocorrelated (Moran's I) at 2.5-m intervals. The autocorrelation rapidly declined at greater intervals, reaching nonsignificance at ca. 20 m. Both canopy height and GSF at 3 m height had a weak tendency for negative autocorrelations at intervals of 25–50 m. Median canopy height (615 stations) was 23 m (range 0–37). Gaps (canopy height ≤2 m, Brokaw 1982) were only 1.5% of sample points. Gaps were more frequent on steep slopes than on terraces, ridgetops, swales, and gentle slopes. Canopy height varied significantly across this topographic gradient. At all four heights (0.6, 1, 3, and 5 m) median GSF was ≤2.4%. GSF values 〉8% accounted for only 3% of the total sample (N = 1380). GSF was only weakly negatively correlated with canopy height and the relation was not monotonie. Under canopies 13-19 m tall, nearly all GSF values were ≤5%. Higher GSFs were more frequent under both shorter and taller canopies. Given the observed variance in GSF and canopy height, 100–200 points separated by intervals of ≥20 m are necessary to measure the forest-wide means of these variables to ±10%. We discuss implications of these results for current approaches to modeling understory light based on canopy characteristics. We compared the random background of light environments from 1–3 m above the ground at La Selva with those occupied by saplings of pioneer and nonpioneer tree species. The two pioneers (Cecropia spp.) occurred in microsites significantly brighter than random sites, while sapling microsites of all five nonpioneer species were significantly darker than random. Comparing the landscape-scale distribution of key resources with species' actual distributions at similar scales offers a quantitative method for assessing plant life histories within and among forests.

Description: This study contributes to the Pinus pinaster Ait. breeding programme, which is reaching the third generation by adding information on wood quality of 46 open-pollinated families from a progeny trial located in Leiria, Portugal, that originated from seed collected in a clonal seed orchard. A total of 552 seventeen-year-old trees were sampled at 2 m height. Trends were studied from the pith outward in variance components and narrow-sense heritability (h2) of wood density components and ring-width characteristics as well as genetic correlations between cambial ages. Mean ring density (RD), minimum density (MND), maximum density (MXD), earlywood density (EWD), latewood density (LWD), earlywood width, latewood width, ring width, latewood percentage, and heterogeneity index were determined using X-ray densitometry procedures. RD had higher genetic control (h2 = 0.63), and heritability values of earlywood components (h2MND = 0.54, h2EWD = 0.60) exceeded those of latewood components (h2MXD = 0.34, h2LWD = 0.26). Heritabilities increased with ring number from pith for almost all wood density components, and there were high age–age genetic correlations for wood density traits (rg 〉 0.98).

Description: Persistence of tree species in a habitat depends on their ability to avoid and respond to disturbance-related damage. Responses to stem damage vary among species but typically include bark wound closure and prevention of xylem decay spread. These responses are associated with anatomical, structural, and physiological traits. This study explores how xylem (vessel size and (or) abundance, parenchyma abundance, ray width, and wood density) and phloem (bark thickness, proportion of live inner bark, ray width and (or) dilation, inter-ray distance, and tissue density) traits relate to responses to stem damage in seven species from the Bolivian Amazon. Rates of bark wound closure and radial xylem decay penetration were compared 2 years after experimental damage. A species that closed bark wounds rapidly (100% in Chorisia speciosa A. St.-Hil.) was not efficient at constraining xylem radial decay spread (1.7 mm). The opposite was true for Pseudolmedia laevis (Ruiz & Pav.) J.F. Macbr., a species that closed wounds slowly (30%) but efficiently controlled decay spread (0.5 mm). The relationship between anatomical and (or) structural traits and damage response variables revealed that species with favorable traits for rapid wound closure (e.g., widely dilating rays) had traits that favored xylem decay spread (e.g., low wood density). It is plausible that this apparent trade-off is based on physiological and phylogenetic constraints.

Description: Two differential forms of growth equations, called the power decline, or PD form, and the exponential decline, or ED form, generate classic growth equations (such as the logistic, Chapman–Richards, Korf) and many other integral forms. Having a full range of these integral solutions allows us to classify them, establish requirements to their parameters, and relate these parameters and initial values (starting age and tree size). Comparisons with data confirm theoretical results. Some applications of the results are discussed.

Description: The Gus Pearson Natural Area is one of the longest maintained individual-based forest-monitoring sites in the world. It is an 800 × 400 m plot established in 1908 within pristine ponderosa pine (Pinusponderosa Dougl. ex Laws, van scopulorum) forest near Flagstaff, Arizona. I quantified decadal-scale growth trends of individual trees and of the entire stand from timber inventories repeated at 10-year intervals between 1920 and 1990. A mixed linear model for longitudinal data was used to test significance of temporal trends. Stand density increased throughout the 20th century because of successful regeneration pulses, active fire control, and no tree cutting. Aboveground growth rates of the stand as a whole did not change significantly from 1920 to 1990, but individual growth rates declined. Mixed linear model results indicated that decadal basal area increment of large pines declined more than that of small pines. In 1920–1930 large pines were growing faster than small pines, but by 1980–1990 large pines were growing slower than small pines. Since the number of small pines multiplied and the number of large pines changed slightly, I inferred that competition for resources was more detrimental for large pines than for small pines.

Description: It is critical to understand how the public prioritizes multiple forestry values when establishing objectives for sustainable forest management. While this is a complex and difficult task, a necessary step is to elicit a broad range of public opinions in forest planning to ensure that decisions serve the needs of various forest stakeholders and society at large. This study seeks to understand how six forest dependent communities in British Columbia prioritize a number of attributes associated with sustainable forest management by using a simple survey-based measurement tool, the Thurstone scale. The results suggest that ecological attributes are a higher priority for survey respondents followed by quality of life, global warming, and economic considerations. This paper explores some of the ramifications of the priorities for sustainable forest management measured in these six communities as well as implications for using the Thurstone scale in processes like Public Advisory Groups.

Description: We conducted a greenhouse experiment to determine how differences among tree species in allocation of biomass to roots versus shoots affected their responses to different combinations of light, water, and soil nutrients. Across a full range of light levels, we were specifically interested in examining the sensitivity of tree seedlings to additional reductions in aboveground growth due to soil resource limitation, and the relative sensitivity of seedlings to water versus nutrient stress under different light regimes. The four tree species used in our experiment included two species that are the most common initial tree invaders of abandoned agricultural lands (old fields) in the Hudson Valley (red maple (Acerrubrum L.) and white pine (Pinusstrobus L.)) and two species that are less frequently found in old fields, but that are dominant forest species, and are noted for their tolerance of either light stress or water stress (sugar maple (Acersaccharum Marsh.) and red oak (Quercusrubra L.), respectively). At the lowest light levels (2% of full sun), there was no effect of variation in soil resource availability on shoot growth of any of the four species. At the 9% full sun light level, red maple seedlings showed clear evidence of simultaneous limitation by light, water, and nitrogen. At higher light levels (20–100% of full sun), all four species responded to variation in at least one of the two soil resources. The four species showed two contrasting patterns of allocation of biomass to roots. Red maple and white pine responded to an increase in soil resource availability by reducing relative allocation to roots and increasing aboveground growth. Sugar maple and red oak had much more conservative root allocation patterns: root allocation was high (58–75% of added biomass allocated to roots) and did not vary in response to soil resource availability. Allocation to roots was affected more strongly by variation in soil nitrogen availability than it was by soil moisture availability.

Description: We studied longleaf pine (Pinuspalustris Mill.) ecosystems to determine causes and rates of overstory mortality, size of canopy disturbances, and the effects of disturbance on canopy structure. Further, we used redundancy analysis to relate variation in characteristics of mortality across a landscape to site and stand variables. We analyzed mortality that occurred from 1990 to 1994 in 70 second-growth plots that spanned a range of site conditions and stand structures, and in five large disturbances that occurred outside the random sample of plots. Half of pine mortality over 5 years in the 70 plots was from unknown causes. Lightning was the primary identifiable cause of mortality, followed by suppression and wind. Lightning mortality was most frequent on xeric sites, while windthrow was common on wet–mesic sites. Suppression mortality was frequent on wet–mesic sites and in higher density stands. Five-year mortality rates averaged 2.3 trees/ha, or 1.9% of original density. Most mortality consisted of single trees. Large disturbances (mostly from lightning) of 15–30 trees were rare, occurring once per 1000 ha in 5 years. Variation in amount of mortality and size of disturbance were unrelated to soil or stand structural characteristics. Low mortality rates from small-scale disturbances result in slow canopy turnover. These results indicate that large openings sufficient for natural regeneration of longleaf pine develop slowly in the absence of hurricanes. Silvicultural options for longleaf pine can be designed to mimic the canopy structure that results from natural canopy disturbances, which leave many live trees standing. Such options may be desirable if a goal of silviculture is to increase structural complexity in stands managed for timber.

Description: Modeling height in the context of site index involves two processes: (i) estimating height at base age (site index), given height at some other age, and (ii) estimating height at some desired age given height at base age. When these two processes are modeled by individual functions for each process, height is assumed to be measured without error when on the right-hand side of the equation, but possessing error when on the left-hand side of the equation. This assumption causes a bias in the parameters of the site index curves. The bias in the estimated parameters is evident in the magnitude of the curves and in the shape of the curves, the typical change of shape of the curves from low site index to high site index is underestimated. An ad hoc method is presented that solves the problem. The ad hoc method is conceptually related to methods used in linear regression and to maximum likelihood estimation in a measurement errors problem context. A general measurement errors model is presented as an alternative. The measurement errors model requires an independent estimate of measurement error.

Description: Japanese stone pine (Pinuspumila Regel) is a dominant species characteristic of alpine zones of high mountains. Eighteen natural populations of P. pumila were studied in an effort to determine the extent and distribution of genetic diversity. The extent of genetic diversity within this species was high (HT = 0.271), and the genetic differentiation among populations was also high (GST = 0.170) compared with those of other conifers. In previous studies of P. pumila in Russia, the genetic variation within the species was also high, but the genetic differentiation among populations was low. We infer that this difference originates from differences in geographic distribution and ecological differences between the two countries. The genetic variation within each population tended, as a whole, to be smaller within marginal southern populations than within northern populations. Genetic relationships among populations reflect the geographic locations, as shown by unweighted pair-group method with arithmetic means and neighbor-joining phylogenetic trees.

Description: The effect of prechilling on the germination response to temperature was investigated by germinating prechilled (at 5 °C) and unchilled seeds of Betulapendula Roth (silver birch) at temperatures of 10, 15, 20, 25, and 30 °C in darkness. Unchilled and prechilled seeds produced 50% germination at temperatures of 26.7 and 15.3 °C, respectively. Two experiments were organized to study the effects of a range of prechilling temperatures (0–12 °C) on release of dormancy. The level of dormancy in the seeds was tested at 12 °C using 6- and 18-h photoperiods. The optimal prechilling temperatures were between 1.0 and 3.3 °C. As the prechilling temperature increased, the effect of prechilling decreased so that the upper effective limit was between 8.7 and 12.0 °C. The effect of prechilling time on dormancy release was studied in an experiment where seeds prechilled at 2.4, 5.5, and 12.4 °C for 6, 12, 21, and 42 days were germinated at 12 °C using 6- and 18-h photoperiods. The germination probability (in logit scale) was directly related to the logarithm of the number of prechilling days. In seeds prechilled at 2.4 °C the requirement for a long photoperiod for germination was removed in 21 days, whereas 42 days were needed in seeds prechilled at 5.5 °C.

Description: This study provides a comprehensive set of functions for predicting biomass for Common beech ( Fagus sylvatica L.) in Central Europe for all major tree compartments. The equations are based on data of stem, branch, timber, brushwood (wood with diameter below 5 or 7 cm), foliage, root, and total aboveground biomass of 443 trees from 13 studies. We used nonlinear mixed-effects models to assess the contribution of fixed effects (tree dimensions, site descriptors), random effects (grouping according to studies), and residual variance to the total variance and to obtain realistic estimates of uncertainty of biomass on an aggregated level. Candidate models differed in their basic form, the description of the variance, and inclusion of various combinations of additional fixed and random effects and were compared using the Akaike information criterion. Model performance increased most when accounting for between-study differences in the variability of biomass predictions. Further, performance increased with the inclusion of age, site index, and altitude as predictor variables. We show that neglecting variance partitioning and the fact that prediction errors of trees are not independent with respect to their predictor variables will lead to a significant underestimation of prediction variance.

Description: A midrotation commercial thinning experiment conducted in two independent spruce plantations compared several tree selection strategies as both early (age 19 or 24 years) single and double (again at age 30 or 34 years) entries over a 16 year period. A delayed (age 30 or 32 years) single-entry thinning and unthinned portions of the plantations served as references for the other treatments. With the exception of 50% systematic row thinnings, removal intensities were held relatively constant at about 40% of stand basal area. All of the thinning treatments satisfied the objectives of focusing diameter and volume growth on a reduced number of stems and making merchantable volume available during the rotation. Early single-entry thinning allowed 30–50 m3/ha to be harvested, increasing quadratic mean diameter and mean merchantable volume per stem at the end of the observation period by 10% and 24%, respectively, over unthinned stands. A second thinning removed an additional 48–64 m3/ha and increased diameter and volume gains to 25% and 71%, respectively. Only marginal differences were observed between the tree-selection strategies, and there were no overall gross total or gross merchantable volume gains or losses associated any of the thinning treatments relative to the unthinned plantations.

Description: A percent stocking change model was developed for lodgepole pine ( Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) in Alberta based on spatially mapped permanent sample plot data. Percent stocking was defined as the percentage of 10 m2 subplots occupied by at least one tree with a minimum height of 1.3 m. The difference equation technique was employed to fit the model. Three model forms were examined and the logistic function was chosen as the final model. Site index was found to be a significant predictor and incorporated into the model. Analyses revealed that the model had correlated, but homoskedastic errors and the correlated errors were modeled by spherical covariance structure using NLINMIX macro in SAS. A percent stocking index, defined as the percent stocking at 50 years total age, was introduced and derived from the developed model. The percent stocking model had both forward and backward projection capabilities. It was demonstrated, both on model fitting and validation data, that the model adequately portrayed the percent stocking dynamics of lodgepole pine stands in Alberta. The model also provided an important basis for creating linkages between reforestation survey results and future yield, which is crucial for sustainable forest management.

Description: The emergence of forest ecosystem management presents new information challenges for forest managers. Shifting views of the forest from primarily one as a production system for wood fibre to an ecosystem with spatially and temporally complex interrelationships is changing the demand for information about the forest. These new information needs are characterized by greater complexity, limited availability of mechanistic hypotheses, and a paucity of data. Empirical and process modelling approaches have evolved in forest management to solve different problems, and debate about the two approaches has existed for some time. Which approach to forest modelling will best be able to meet the challenges of ecosystem management? Empirical models seek principally to describe the statistical relationships among data with limited regard to an object's internal structure, rules, or behaviour. In contrast, process models seek primarily to describe data using key mechanisms or processes that determine an object's internal structure, rules, and behaviour. In addition, mechanisms included in process models are general enough that they can maintain some degree of relevance for new objects or conditions (mechanism constancy), while empirical models tend not to be tied to any specific mechanism, so that derived model parameters must remain constant (parameter constancy) for new objects or conditions. Based on these differences, we argue that process models offer significant advantages over empirical models for increasing our understanding of and predicting forest (a tree, a stand, a landscape) behaviour. Process models are, therefore, more likely to meet the information challenges presented by ecosystem management.

Description: The effects of geographic information system (GIS) data conversion on several polygon-and landscape-level indices were evaluated by using a GIS vegetation coverage from eastern Oregon, U.S.A. A vector–raster–vector conversion process was used to examine changes in GIS data. This process is widely used for data input (digital scanning of vector maps) and somewhat less widely used for data conversion (output of GIS data to specific formats). Most measures were sensitive to the grid cell size used in the conversion process. At the polygon level, using the conversion process with grid cell sizes of 3.05, 6.10, and 10 m produced relatively small changes to the original polygons in terms of ln(polygon area), ln(polygon perimeter), and 1/(fractal dimension). When grid cell size increased to 20 and 30 m, however, polygons were significantly different (p 

Description: Measurements of frost damage and bud-break phenology were made during the spring of 1993 and 1994 on planted white spruce (Piceaglauca (Moench) Voss) seedlings and on trembling aspen (Populustremuloides Michx.) suckers in combination with measurements of minimum air temperature and long-wave radiation in forest openings of different sizes and configurations (1.5-ha clearcut, 9 and 18 m wide strips, 9 and 18 m diameter circular openings and intact forest) near Chapleau, Ontario. The average minimum air temperature during early summer (May–June) decreased linearly as the sky view factor of the forest openings increased. The average difference in minimum air temperature between the forest and clearcut opening was nearly 3 °C for the period, and during clear night sky conditions, this temperature difference approached 6 °C. The difference in minimum temperature between the forest and clearcut during cloudy night sky conditions was slightly greater than 1 °C. Differences in minimum air temperature between the forest and each of the openings increased with decreasing incoming long-wave radiation. The largest differences occurred between the forest and the clearcut with low incoming long-wave radiation (cloudless nights). Minimum air temperatures in the 9-m strip and 9-m circle differed little from minimums in the forest under all long-wave radiation regimes. Moderation of air temperature on clear nights would be best achieved with sky view factors of less than 0.3, implying a maximum strip width of 0.6 tree heights and a maximum patch diameter of 2.2 tree heights. Damage to white spruce seedlings after a frost in the spring of 1993 increased with increasing sky view factor. Nearly 75% of the seedlings in the clear-cut had medium or heavy frost damage, while only 2% of the seedlings in the 9-m circle and forest were frost damaged. Opening size and configuration had little influence on the timing of bud break in either the white spruce seedlings or the trembling aspen suckers.

Description: The Johnson system of distributions was used to fit both diameter and height data collected from 226 sample plots located in the ponderosa pine (Pinusponderosa P. Laws, ex C. Laws.) and mixed-conifer forest types in New Mexico and Arizona. To estimate the Johnson system parameters, five methods (namely the four-percentile method, Knoebel–Burkhart method, mode method, maximum likelihood estimation method, and a new method developed in the study, linear regression method) were compared and evaluated. For these sample plots, the linear regression method was superior for estimating parameters of SB distributions for both diameter and height.

Description: We studied responses of Acari, especially oribatid mites, to prescribed low-intensity fire in an east side pine site in the southern Cascade Range in California. We compared oribatid population and assemblage responses to prescribed fire in stands that had been selectively logged to enhance old growth characteristics, in logged stands to minimize old growth characteristics, and in undisturbed forest reference stands. Low-intensity prescribed fire altered habitat characteristics within the organic layer of forest soil. Acarine populations declined following prescribed fire, and oribatid losses accounted for two thirds of that decline. Individual oribatid species responded differently to prescribed fire, with a few populations increasing after fire but most declining. The prescribed fire also altered oribatid assemblages, reducing species richness and species diversity and modifying assemblage dominance relationships. We also identified several oribatid taxa that were potential indicator species of fire effects upon forest soil fauna. Finally, our results suggested that oribatid responses to fire were intensified by stand alteration and especially by removal of old growth structural characteristics. Decline in oribatid abundance, species richness and diversity, and loss of equilibrium dominance relationships was greatest in the low structural diversity plots.

Description: Foliar macronutrient (N, P, K, Ca, Mg) concentration, content, retranslocation, internal balance, and variation with crown location and foliage age were quantified in 11-year-old loblolly pine (Pinustaeda L.) trees grown in two soil nitrogen regimes. Concentrations of N, P, and K decreased with foliage age, while the concentration of Ca increased. Nutrient content of new foliage increased with the elongation of foliage. Estimates of retranslocation were 75%, 73%, 83%, 28%, and less than 5% for N, P, K, Mg, and Ca, respectively. Increases in nutrient content of new foliage were associated with decreases in nutrient content of old foliage, suggesting transfer of nutrients to developing foliage. Concentrations and contents of P and K were greatest in the upper crown. Differences in N concentration among crown locations were small, although N content increased in the upper portion of the crown. Calcium and Mg concentrations decreased with crown height, but their contents generally did not differ among crown locations. Nitrogen fertilization increased N and Ca concentrations and fascicle weight, decreased P and Mg concentrations, and had no significant effect on K concentration. For the purposes of determining nutrient deficiency, our results suggest that foliar samples should be collected from 1-year-old foliage in the midcrown position during the rapid growth period for new shoots.

Description: We evaluated geographic variations in mean fire return intervals and postfire forest succession within a 66 497 km2 land area located in the eastern Quebec boreal forest. Fire return intervals were calculated using a time since last fire map for 1800–2000, and forest dynamics were studied by superimposing 3204 forest inventory plots onto the fire map. Mean fire return interval proved significantly shorter in the western part of the study area, at 270 years, compared with the eastern part, where it was probably more than 500 years. The two main tree species in the study area were balsam fir ( Abies balsamea (L.) P. Mill.) and black spruce ( Picea mariana (Mill.) BSP). Balsam fir abundance increased progressively as a function of time since fire, whereas black spruce abundance increased during the first 90 years after fire and then declined. Balsam fir was significantly more abundant in the southeastern portion of the study area, which we attribute to the combined limitations imposed by temperature along the north–south axis and by fire along the east–west axis. Large forest patches (i.e., ≥200 km2) dominated by early successional tree species, within a matrix of irregular black spruce – balsam fir mixtures, are an important feature of preindustrial forest landscapes in this region.

Description: The intrinsic variability of volatile organic compound emissions and photosynthetic parameters in response to two drying cycles was investigated on Quercus suber L. saplings originating from three Tunisian populations native to contrasting climates. Emissions mainly included monoterpenes plus traces of sesquiterpenes and methylsalicylate that steadily decreased during the experiment unrelated to treatments. Instead, monoterpene emissions increased by 30% during a first moderate drought and remained enhanced after rewatering, while CO2–H2O gas exchange was reduced by 30%. A second severe drought decreased monoterpene emissions to about two third of controls, strongly inhibited gas exchange and photochemical efficiency, and caused a partial loss of chlorophyll. Rewatering led to only partial recovery of emissions and photosynthetic parameters of stressed plants whose biomasses and leaf nitrogen and chlorophyll contents were lower than in control plants. There was no clear difference among populations in volatile organic compound emissions or photosynthetic parameters. However, the population inhabiting the most arid region had intrinsically smaller leaves with lower specific leaf masses and higher drought-related leaf losses than populations native to less arid climates. These results indicate an adaptive shift from stress-tolerating towards stress-avoiding ecotypes under arid climates involving mainly morphological adaptations without apparent differentiation in volatile organic compound production, probably because of its high phenological plasticity.

Description: Preindustrial forest composition for 〉180 000 km2 throughout central and northeastern Ontario was recreated from Ontario Crown land survey notes (1816–1955) and compared with existing forest composition derived from current Forest Resource Inventories (1998–2009) in each of Site Regions 3E, 4E, and 5E. A validation analysis was performed using the Forest Resource Inventory data to test the assumption that sampling the land survey tree species composition along township boundaries is adequate in describing the composition of the whole forest. The majority of tree species in each of the three site regions validated successfully. A binary logistic regression model allowed birch genera to be classified at the species level to aid in the interpretation of survey notes. All analyses showed significant reductions in conifers (especially red pine ( Pinus resinosa Ait.), white pine ( Pinus strobus L.), and eastern larch ( Larix laricina (Du Roi) K. Koch)) and significant increases in maple ( Acer spp.), oak ( Quercus spp.), white birch ( Betula papyrifera Marsh.), and poplar ( Populus spp.).

Description: This study compared the performance of Eucalyptus camaldulensis Dehnh. planted at four spacings (2.0 m × 2.0 m, 2.5 m × 2.5 m, 3.0 m × 3.0 m, and 3.5 m × 3.5 m) at three experimental saline sites in Punjab province of Pakistan over 5 years and assessed the impact of these planting densities on soil salinity amelioration. Tree response was assessed by measuring survival, height, and diameter at breast height (DBH) of trees at 1, 2, and 5 years as well as wood volume after 5 years. Wood volume per hectare, height, and DBH were greater at sites I and II than at site III. The 3.0 m × 3.0 m spacing resulted in taller trees, but DBH and wood volume were greater with the 3.5 m × 3.5 m spacing. The study confirmed that initial tree spacing or density has a significant impact on the subsequent height, DBH, and wood volume of E. camaldulensis plants. The study also showed that broad spacing could be a better option in saline environments. There was a definite reduction in soil salinity (electrical conductivity, ECe) at five soil depths after 5 years of tree growth at all sites and spacing treatments, with reductions varying from 46% to 47% at site I, 31% to 52% at site II, and 25% to 58% at site III. Soil ECewas generally higher in surface (0–15 cm) soils.

Description: Episodic stand-replacing wildfire is a significant disturbance in mesic and moist Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco) forests of the Pacific Northwest. We studied 24 forest stands with known fire histories in the western Cascade Range in Oregon to evaluate long-term impacts of stand-replacing wildfire on carbon (C) and nitrogen (N) pools and dynamics within the forest floor (FF, Oe and Oa horizons) and the mineral soil (0–10 cm). Twelve of our stands burned approximately 150 years ago (“young”), and the other 12 burned approximately 550 years ago (“old”). Forest floor mean C and N pools were significantly greater in old stands than young stands (N pools: 1823 ± 132 kg·ha–1vs. 1450 ± 98 kg·ha–1; C pools: 62 980 ± 5403 kg·ha–1vs. 49 032 ± 2965 kg·ha–1, mean ± SE) as a result of significant differences in FF mass. Forest floor C and N concentrations and C/N ratios did not differ by time since fire, yet potential N mineralization rates were significantly higher in FF of old sites. Old and young mineral soils did not differ significantly in pools, concentrations, C/N ratios, or cycling rates. Our results suggest that C and N are sequestered in FF of Pacific Northwest Douglas-fir forests over long (∼400 year) intervals, but that shorter fire return intervals may prevent that accumulation.

Description: The decay resistance of black locust ( Robinia pseudoacacia L.) heartwood was evaluated on standardized test specimens taken from mature and juvenile wood in 25 trees distributed over five sites in Wallonia, Belgium. The percentage mass losses caused by the wood-destroying fungi Coriolus versicolor CTB 863 A and Coniophora puteana BAM Ebw.15 were evaluated according to Cen/ts 15083-1 (Cen/ts 15083-1. 2005. Durability of wood and wood-based products — determination of the natural durability of solid wood against wood-destroying fungi, test methods — Part 1: Basidiomycetes. European Comm. for Standardization, Brussels). Black locust is classified as a “very durable” wood (class 1), while the juvenile heartwood (located near the pith) is classified in classes 1 to 2. The statistical analysis does not reveal any significant differences among the sites, suggesting that the growth environment of the trees has no effect on their natural durability. However, statistical analysis shows very highly significant differences among trees and between the two radial positions in the tree, with mature wood being more resistant than juvenile wood. Using such a naturally very durable species could reduce reliance on preservative treatment and the use of tropical species generally recommended for outdoor woodwork.

Description: Tree species can differ in their responses to resource availability during the critical phase of establishment, which could influence forest dynamics. In Mediterranean forests, most of the attention has focused on the effects of shade and summer drought on seedling survival, but little is known about the effect of autumn to spring rains on earlier stages of recruitment. A sowing experiment was set up along natural light and water gradients with three co-occurring oak species ( Quercus suber L. (cork oak), Quercus canariensis Willd. (Algerian oak), and Quercus pyrenaica Willd. (Pyrenean oak)) that show limited natural regeneration in southern Spain. Recruitment stages were monitored for 1 year. Models of seed germination, seedling emergence, and seedling survival as well as of overall recruitment patterns were developed as functions of light, soil moisture, and soil compaction. The influence of intraspecific variation in seed mass and emergence time were also tested. Excess soil water levels during the winter reduced germination and emergence and lengthened time to emergence (in waterlogged open areas), which in turn decreased seedling survival during the dry season. Seedlings from larger seeds were more likely to germinate and emerge. The results suggest that temporal and spatial variability of soil water content, mediated by emergence time and seed size, play a crucial role in the regeneration dynamics of Mediterranean oak forests.

Description: Perpendicular distance sampling (PDS) has emerged as a compelling alternative to line intersect sampling (LIS) for the inventory of forest fuels and other downed woody materials (DWM), particularly where the aggregate volume of DWM is of primary interest. This article develops a selection protocol and design-unbiased estimators for a new probability proportional-to-volume sampling strategy, termed line intersect distance sampling (LIDS). LIDS combines the distance sampling protocol of PDS with the transect sampling protocol of LIS and provides unbiased estimates of aggregate DWM volume from counts of selected logs or log fragments. Simulations indicate that LIDS along multidirectional (e.g., Y-shaped) transects should perform similarly to PDS in terms of sampling error; however, it remains unclear how LIDS and PDS compare with LIS, especially when interest is attached to multiple DWM population parameters. It is argued that LIDS will be most useful in reducing implementation errors, particularly detection errors, relative to PDS under limited visibility field conditions.

Description: This 2-year study examined the effect of fertilizers on tree growth and foliar nutrition in a Populus trichocarpa Torr. & A. Gray × Populus deltoides Bartr. ex Marsh. plantation located in southwestern Québec. The treatments included a control that did not receive N or P fertilizer, inorganic NP fertilizers, organic fertilizers applied at 65–70 kg N·ha–1, and organic fertilizers applied at 130–140 kg N·ha–1. Fertilized trees were taller and had larger diameters than control trees. Three methods were used to diagnose limiting nutrients and nutrient imbalances, and compare the nutrient supply from different fertilizer sources. The critical value approach and the compositional nutrient diagnosis methods found below-optimum N and P concentrations, sufficient K and Mg concentrations, and an excessive Ca concentration in foliage. Vector analysis compared the N nutrition in foliage from fertilized trees and the control trees. The compositional nutrient diagnosis r2(nutrient imbalance index) was negatively correlated with annual tree growth in height (r = –0.46, P 〈 0.05) and diameter (r = –0.59, P 〈 0.05), meaning that trees with a greater nutrient imbalance grew less in height and diameter than trees with balanced foliar nutrition. Of these diagnostic methods, compositional nutrient diagnosis holds promise for identifying nutrient limitations and predicting growth responses to fertilization in hybrid poplar plantations.

Description: Genetic diversity within populations is an important component of adaptive evolution, and recent research has demonstrated that genetic variation within plant populations can have important ecological effects. In this study, we investigate quantitative-genetic variation in several traits within a quaking aspen ( Populus tremuloides Michx.) population. A common garden experiment was planted with replicates of 13 aspen genotypes collected from wet and dry sites within a population in southern Utah, USA. Ten growth, leaf, physiological, and structural traits were measured. There were significant, heritable phenotypic differences among genotypes in every measured trait and differences in 4 of the 10 traits among genotypes originating from wet and dry collection sites. The data were compared with other published studies, showing that aspen heritability (H2) estimates and coefficients of genetic variation (CVG) were comparable or higher than other Populus species and hybrid F1 Populus genotypes, indicating a large amount of quantitative-genetic variation in aspen.

Description: This study presents a new, simple, and efficient method for estimating microsite light availability in the understory of a forest. The percentage of above-canopy photosynthetic photon flux density (%PPFD) transmitted above 16 microsites in the understory of a mixed conifer–broadleaf forest was measured every minute between 07:00 and 19:00 for both a completely overcast and a cloudless day. Instantaneous measures of %PPFD were also taken at different times on 3 overcast days. The instantaneous measures of %PPFD were strongly and directly related (P 〈 0.001) among themselves and with mean daily %PPFD values. These results demonstrate the usefulness of using an instantaneous measure of %PPFD taken under overcast sky conditions for estimating the mean daily %PPFD at any microsite under a forest canopy.

Description: Newly transplanted, 3-year-old bare-root white spruce (Piceaglauca (Moench) Voss) seedlings were grown in three levels of absolute humidity difference (AHD), under water-stressed and well-watered conditions in growth chambers. Stomatal conductance to CO2 as well as net assimilation were more than twice as high in the low AHD treatment (6.8 μg H2O•cm−3) as in the high AHD treatment (21.0 μg H2O•cm−3). Transpiration rates were uniform among humidity treatments, but in the low AHD treatment water use efficiency was more than double that of the high AHD treatment. Water use efficiency was greatest in the low AHD conditions, in the water-stressed seedlings, and during the time immediately after planting. There were no differences in the number of new roots produced among humidity treatments, despite the different levels of photosynthesis. In the stress treatment, bud flush of seedlings in the high AHD treatment occurred 3 days later than in the low AHD treatment. A field study on the effects of varying aspen (Populustremuloides Michx.) canopies on humidity levels showed that AHD under partial and full canopies was lower than that of the clearcut. Results suggest that silvicultural treatments that promote higher humidity levels on planting sites should improve white spruce seedling photosynthesis immediately after planting.

Description: Innovativeness can help companies differentiate themselves, with the ultimate goal of securing survival and improving performance. Modern theories in organizational behavior look at innovation as something that starts with individual creativity but that is also affected by the work environment. Using one broad industry sector, the US forest products industry, this study attempts to integrate into a unifying model the concepts of work climate, innovativeness, and firm performance using structural equation modeling. Results support the proposed theoretical model, with some modifications, finding a positive and significant relationship among all factors. Having innovation as a core part of a company’s strategy and fostering a climate for innovation positively affects the degree of innovativeness and performance of a company. This is especially true for secondary or value-added wood products manufacturers. A climate for innovation is characterized by high levels of autonomy and encouragement, team cohesion, openness to change and risk taking, and sufficient resources available to people. Lack of a validation sample suggests treating the model as tentative until further testing.

Description: The coupling of synoptic scale weather conditions with local scale weather and fuel conditions was examined for 2551 fires and 1 537 624 lightning strikes for the May through August fire seasons in 1988, 1989, 1992, and 1993 in Alberta and Saskatchewan. The probability of lightning fire occurrence (number of fires/number of strikes) is near zero until the Fine Fuel Moisture Code reaches 87 (moisture content of 14% dry weight), after which the probability increases rapidly. Duff Moisture and Drought Codes show less clear increases. In all cases, the probability of fire occurrence was low (the number of strikes greatly exceeded the number of forest fires), suggesting that lightning fire ignition coupled with early spread to detection was an uncommon event. This low probability of fire occurrence even at low fuel moisture may be a result of the arrangement and continuity of fuels in the boreal and subalpine forests. The literature suggests a higher probability of lightning-ignited fires in qualitatively different fuels, e.g., grasslands. The higher probability of fire at lower fuel moistures occurred primarily when high pressure dominated (positive 50-kPa anomaly) for at least 3 days and less than 1.5 mm precipitation occurred. The highest number of lightning strikes and largest number of fires also occurred when high pressure dominated. The high lightning numbers during high pressure systems were logistically related to increasing atmospheric instability (K-index).

Description: Among the current environmental concerns for forests of the eastern United States is nitrogen (N) saturation, a result of excessive inputs of N associated with acidic deposition. We studied nutrient responses on N-treated and untreated watersheds of the Fernow Experimental Forest, West Virginia, to test for evidence of N saturation on the treated watershed. The watersheds were WS7 (23-year-old even-aged control), WS4 (mature mixed-aged control), and WS3 (23-year-old even-aged treatment). WS3 has received aerial applications of (NH4)2SO4 from 1989 to the present (a total of 4 years for the study period) at 3 × ambient inputs of N and S (54 and 61 kg•ha−1•year−1, respectively). Base-flow stream samples were collected weekly from each watershed and analyzed for NO3− and Ca2+. Mineral soil was incubated in situ, placed in bags, and buried about every 30 days during the growing season in each of seven sample plots within each watershed. Moist samples of soil from the bags were analyzed for extractable NH4+ and NO3−. In addition, forest floor material and leaves of an herbaceous species (Violarotundifolia Michx.) from each plot were analyzed for N and other nutrients. Violarotundifolia was present on all 21 plots and used as an additional indicator of N availability and soil fertility. Foliage tissue was sampled from overstory tree species (Liriodendrontulipifera L., Prunusserotina Ehrh., Betulalenta L., and Acerrubrum L.) from WS3 and WS7 and analyzed for nutrient content. Results from the 1993 growing season showed few, if any, differences among watersheds for (1) N content and C/N ratio of the mineral soil and forest floor and (2) relative proportion of NH4+ and NO3− produced in the buried bags. Nitrification rates were equally high in soils of all watersheds; N concentrations were significantly higher in foliage tissue of overstory tree species and of V. rotundifolia in the treatment versus control watersheds; plant tissue Ca was significantly lower for the treatment watershed than for the control watersheds. Our results support the conclusions of earlier studies that high amounts of ambient N deposition have brought about N saturation on untreated watersheds at the Fernow Experimental Forest. This is suggested by minimal differences among watersheds in N mineralization and nitrification and soil and forest floor N. However, aggravated N saturation on our treated watershed can be seen in differences in plant tissue nutrients among watersheds and streamflow data, indicating increased losses of NO3− with accompanying losses of Ca2+ in response to further N additions to a N-saturated system.

Description: Partial cutting is increasingly applied in European temperate oak-dominated forests for biofuel harvesting, and to counteract succession in protected stands. Effects on biodiversity of these measures need to be carefully evaluated, and species-rich but neglected taxa such as fungi should be considered. We studied the effects of partial cutting on fungal fruiting bodies on woody debris. In 21 closed canopy forests rich in large oaks in Sweden, on average 25%–30% of the basal area was cut. Fruiting bodies were counted and some were collected in treated and control plots before and after treatment. We found 334 basidiomycete and 47 ascomycete species. Species richness of basidiomycetes declined significantly more in treated plots (on average 26%) than in control plots (on average 13%) between seasons. Species richness of ascomycetes increased by 17% in control plots and decreased by 2% in treated plots. Total species richness was significantly reduced on fine woody debris (1–10 cm in diameter), but not on coarse woody debris (〉10 cm). Overall species composition did not change significantly as a result of partial cutting, but red-listed species tended to decrease more in treated plots. We suggest that approximately 30% of the stands should not be thinned, and dead stems and fallen branches should not be removed, to favor saproxylic fungi and their associated fauna.

Description: We designed and developed an internet mapping application to collect data on the locations of forest landscape values across a 2.4 million hectare study area in the province of Alberta, Canada. Four communities in the study area were surveyed and 8053 point locations were mapped for 10 different value types. Importance weights of landscape values were determined through a ranking exercise. Nearest-neighbour and second-order spatial point pattern analysis (K functions) suggested that all value types were significantly clustered across the study area. Recreational, wilderness, existence, and biological diversity values exhibited maximum clustering at larger spatial scales in comparison with educational, economic, historic or cultural, and spiritual values. Maximum clustering was positively related to mean road density and negatively related to mean distance to water, which suggests that landscape features influence the spatial pattern of values by acting as focal points or attractors for values. An applied use of the data for values hotspot detection and community protection zoning in forest fire management planning is presented.

Description: Severe droughts have the potential of inducing transient shifts in forest canopy composition by altering species-specific adult tree mortality patterns. However, permanent vegetation change will occur only if tree recruitment patterns are also affected. Here, we analyze how a massive mortality event triggered by the 1998–1999 drought affected adult and sapling mortality and recruitment in a mixed Nothofagus dombeyi (Mirb.) Blume – Austrocedrus chilensis (D. Don) Flor. et Boult. forests of northern Patagonia. Comparing drought-induced and tree-fall gaps, we assessed changes in forest composition, microenvironments, and seedling density and survival of both species. Drought-kill disturbance shifted species composition of both canopy and sapling cohorts in favour of A. chilensis. Drought gaps were characterized by a shadier and more xeric environment, affecting the recruitment pattern of N. dombeyi seedlings. The seedling cohort was composed mostly of A. chilensis, and its survival was always higher than that of N. dombeyi. Additionally, A. chilensis seedlings showed higher plasticity than N. dombeyi seedlings, increasing its root to shoot ratios in drought gaps. The results suggest that extreme drought itself is a strong driving force in forest dynamics, with important imprints on forest landscapes. Future climate-change scenarios, projecting an increased in frequency and severity of droughts, alert us about expected long-term compositional shifts in many forest ecosystems.

Description: Survival of live retention trees is a key issue for stand-scale applications of natural-disturbance-based silviculture. We explored the survival of 3255 trees in 102 cut areas (mean size 2.3 ha) in Estonia for 6 years, focusing on spatial variation and preadaptation of the trees. Altogether, 35% of the trees died during the study period, contributing 4.4 m3 of downed dead trunks and 1 m3 of standing dead trees per hectare. The annual mortality rates declined over time. The main survival determinants were tree species (higher for hardwood deciduous trees), diameter (species dependent), position relative to forest edge (higher for trees near current or former forest edges), retention density (positive), and exposure (negative). The results suggest that (1) green-tree retention can effectively increase the abundance of large shade-tolerant trees, but it is equally important for producing deadwood; (2) larger individuals, former interior-forest trees near current forest edges, and preadapted trees in open conditions should be preferably retained; (3) there is no obvious necessity to modify tree-retention techniques for tree survival according to geographical region or forest site type.

Description: In the Georgia Piedmont (U.S.A.), size, abundance, and species diversity of trees were quantified in a plantation of Pinustaeda L. 12 years after various methods and intensities of site preparation. In clear-cut only versus site-prepared plots, greater hardwood abundance (27% vs. 8% of the total basal area) and size (8.6 vs. 7.4 m in height) were associated with reduced pine volume (73 vs. 123 m3/ha) and increased Simpson and Shannon diversity indices. Tree-species richness was greater in plots where residual trees from clear-cutting had been removed with a chainsaw versus large machinery (10 vs. 7 species). With increasing site-preparation intensity, reductions in basal area of volunteer pines coincided with proportionate increases (R2 = 0.80) in basal area of planted pines. As a result of this compensatory effect, total volume of all pines varied little (122–134 m3/ha) among site-preparation intensities. Research results suggest that site-preparation treatments can be selected to facilitate the development of a variety of stand structures, including those that favor evenness (clear-cut only) or richness (manual cutting) of tree species, low-cost production of pine fiber (manual cutting), and stand uniformity for management of pine sawtimber (mechanical and herbicide).

Description: The effect of thinning on the relationship of wood quality traits measured on standing trees (dynamic modulus of elasticity (MOE) and outerwood density) and traits measured on logs or short clear specimens was determined using data collected from radiata pine ( Pinus radiata D. Don) trees growing in 22 unthinned and 16 thinned plots of harvest age trees in New South Wales, Australia. Stiffness showed a linear decrease along the stem. Trees growing on thinned sites were, on average, 3% lower in stiffness at each height in the stem. MOE measured on short clear specimens was moderately related to standing tree MOE (R2 = 0.62) and outerwood density (R2 = 0.56) but less well related to MOE of the adjacent log (R2 = 0.30). Standing tree MOE was a better predictor of whole stem MOE for the thinned sites (R2 = 0.60) than for the unthinned sites (R2 = 0.31). Stiffness and density appear to follow different patterns of variation and results for density may not be extrapolated to stiffness. Outerwood density was a very poor predictor of mean whole stem stiffness (R2 = 0.14). Overall, the acoustic tool, TreeTap, was a better predictor of whole stem stiffness than outerwood density, particularly for the thinned sites.

Description: Following release, mature trees can acclimatively shift aboveground growth allocation from height to diameter at breast height (DBH) growth to adjust their stem form (reducing height/diameter ratio (HDR)) to better withstand increased wind stress. The purpose of this study was to determine if, over a 6-year period, 7- to 12-year-old eastern white pines ( Pinus strobus L.) also responded to different levels of partial release with (i) stem-form adjustments (reduced HDR) through growth-allocation shifts from height to DBH growth and (ii) lower HDR values (shorter heights) than for unreleased trees of the same DBH (“acclimative stem-form development hypothesis”). Over the 6-year postrelease period, juvenile white pine seedlings exhibited unsynchronized height and DBH growth response patterns that depended on their prerelease height growth. Height growth of faster growing white pines was temporarily reduced. Concurrently, DBH growth was enhanced with increasing release intensity. HDR reductions followed a fairly narrow and predictable trajectory, but HDR values of released trees were not lower than those of unreleased trees of the same DBH after 6 years. Juvenile white pine seedlings appear to maintain a balance among aboveground tree parts to enhance future tree stability, which may be an adaptive trait for moderately shade-tolerant species.

Description: To shed light on the currently increasing proportion of Fagus grandifolia Ehrh. saplings in the tolerant hardwood forests of Quebec, we studied 48 Acer saccharum Marsh. dominated stands with contrasting histories of canopy disturbance: old commercial clear-cutting (CC), old fire (F), and either one or two partial cuts (1PC and 2PC). Our results indicated that higher densities of both F. grandifolia and A. saccharum saplings were associated with partial cutting histories (1PC and 2PC) than with severe canopy disturbance (CC and F). The density of F. grandifolia saplings was not related to any soil or stand characteristics in stands with a history of severe canopy disturbance. However, in stands with a history of partial canopy disturbances, the relative density of F. grandifolia saplings as compared with A. saccharum was related to soil C/N ratio and the presence of F. grandifolia overstory trees, whereas the absolute density showed a negative relationship with stand basal area. Therefore, it appears that partial canopy disturbances favored the regeneration of F. grandifolia relative to A. saccharum, whereas severe canopy disturbances may have provided an advantage to A. saccharum. We suggest that the presence of a light threshold can explain this shift in sapling performance between these two species.

Description: Effect of interannual climate variations on radial growth was compared among jack pine ( Pinus banksiana Lamb.) of diverse geographical origins in a 41-year-old common-garden experiment in Petawawa, Ontario. Provenance experiments established from seeds transferred from different parts of a species range (from the northern United States to northern Canada) to the same environment might be considered as a simulation model of climate change and a shift of climate zones. The following questions are addressed: Did the response of growth to interannual climate variations differ among the provenances transferred within the experimental site? What climatic factors affect interannual growth variations of jack pine provenances? Tree-ring chronologies for 16 populations were developed for the period 1970–2004. The best climate predictors of radial growth were precipitation of June and March of the current year and precipitation of December of the previous year. Although, climatic factors affecting growth were similar between the provenances, absolute radial growth was proportional to the growth potential of the provenances. We conclude that variability due to seeds origins is not a significant source of variation for dendroclimatic studies of jack pine. Increased frequency of summer droughts might result in a growth decrease of jack pine.

Description: Boreal forest disturbance regimes have changed during the past century, in turn changing regeneration and stand dynamics of these forests. This is occurring at the same time that the forest industry is emphasizing operations that take advantage of pre-established natural regeneration. This study has therefore investigated the effect of various ecological factors on the abundance of Abiesbalsamea (L.) Mill., Piceaglauca (Moench) Voss, Piceamariana (Mill.) BSP, Thujaoccidentalis L., Betulapapyrifera Marsh., and Populustremuloides Michx. natural regeneration in some boreal stands in the Abitibi region of Quebec. Abiotic ecological site classification variables were found to be poorly correlated with seedling densities for most species. Parent trees, as a seed source, and stand type, for its influence on the quality of the seedbed, were strongly correlated with abundant conifer regeneration. As expected, time since fire was positively correlated with seedling abundance for late successional species, whereas it was not related to the abundance of early successional species. Post-fire disturbances, such as those due to spruce budworm (Choristoneurafumiferana (Clem.)) outbreaks, accounted for the variations in seedling densities observed with respect to stand age. The presence of budworm-caused gaps was positively correlated with regeneration abundance, whereas shrub competition, which was greatest in large gaps, was negatively correlated with advance regeneration density. Forest management based on abundance of pre-established regeneration should focus mainly on mixed stands because seedling density is very low in hardwood stands and because of the strong competition found in large gaps of resinous forests. The low number of seedlings observed, especially in coniferous stands, may limit the effectiveness of operations that take advantage of advance regeneration. The large variability observed among site and stand types limits their operational use in predicting seedling densities.

Description: We used tree-ring chronologies from sugar maple (Acersaccharum Marsh.) stands showing various degrees of dieback (i.e., 16 sugar maple chronologies from healthy trees and 11 from damaged trees), distributed throughout the species range in southern Quebec, to analyze the spatial extent and timing of the recent sugar maple decline. Furthermore, six tree-ring chronologies of American beech (Fagusgrandifolia Ehrh.) from six damaged sugar maple stands were used to compare for differential responses associated with factors such as insect defoliation (from the forest tent caterpillar, Malacosomadisstria Hbn., for which American beech is a nonpreferred species), drought, and other climatic events. It was found that several small-scale drought-induced disturbances occurred repetitively over the last 100 years in the western part of the species range in southern Quebec. Most sugar maple chronologies from stands located west, north, and south of Québec City also show extreme narrow tree rings, indicating the incidence of three large and deep growth depressions from the early to mid-1910s, mid-1950s, and early 1980s. The factors explaining the large growth depression of dominant sugar maple of the early 1980s, in the region where the 1980s maple decline was the most severe, are likely associated with the synergistic influence of drought and insect defoliators. The recovery of sugar maple stands from the 1980s growth decline emphasizes the positive responsiveness of the robust native trees to frequent natural disturbances. The 1980s maple decline corresponds to the category of natural disturbances affecting stand dynamics by the combination of events such as drought and insect infestations, and possibly (but to a minor extent) winter thaw-frost, which has yet to be demonstrated, rather than by anthropogenic pollution.

Description: We describe the decision support system RuttOpt, which is developed for scheduling logging trucks in the forest industry. The system is made up of a number of modules. One module is the Swedish road database NVDB, which consists of detailed information of all of the roads in Sweden. This also includes a tool to compute distances between locations. A second module is an optimization routine that finds a schedule, i.e., set of routes for all trucks. This is based on a two-phase algorithm where linear programming and a standard tabu search method are used. A third module is a database storing all relevant information. At the center of the system is a user interface where information and results can be viewed on maps, Gantt schedules, and result reports. The RuttOpt system has been used in a number of case studies and we describe four of these. The case studies have been made in both forest companies and hauling companies. The cases range from 10 to 110 trucks and with a planning horizon ranging between 1 and 5 days. The results show that the system can be used to solve large case studies and that the potential savings are in the range 5%–30%.

Description: We reduced early winter snowpack in four experimental plots at the Hubbard Brook Experimental Forest in New Hamphire for 2 years to examine the mechanisms of root injury associated with soil freezing. Three lines of evidence suggested that direct cellular damage, rather than physical damage associated with frost heaving, was the principal mechanism of root injury: (i) decreases in root vitality were not greater on sites with more frost heaving, (ii) in situ freezing damage was confined to first- and second-order roots in the organic horizons rather than entire root systems, and (iii) tensile strength of fine roots was not significantly compromised by experimental stretching to simulate ice lens formation. Although significant differences in the intensity of soil freezing (depth, rate, and minimum temperature) were observed across the plots, no clear effects of soil freezing intensity on root injury were observed. Snow manipulation had no effect on mycorrhizal colonization of sugar maple ( Acer saccharum Marsh.) roots. A significant increase in root growth was observed in the second summer after treatments, coincident with a significant pulse of soil nitrate leaching. Through their effects on fine roots, soil freezing events could play an important role in forest ecosystem dynamics in a changing climate.

Description: Forest transportation planning problems (FTPP) have evolved from considering only the financial aspects of timber management to more holistic problems that also consider the environmental impacts of roads. These additional requirements have introduced side constraints, making FTPP larger and more complex. Mixed-integer programming (MIP) has been used to solve FTPP, but its application has been limited by the difficulty of solving large, real-world problems within a reasonable time. To overcome this limitation of MIP, we applied the ant colony optimization (ACO) metaheuristic to develop an ACO-based heuristic algorithm that efficiently solves large and complex forest transportation problems with side constraints. Three hypothetical FTPP were created to test the performance of the ACO algorithm. The environmental impact of forest roads represented by sediment yields was incorporated into the economic analysis of roads as a side constraint. Four different levels of sediment constraints were analyzed for each problem. The solutions from the ACO algorithm were compared with those obtained from a commercially available MIP solver. The ACO solutions were equal to or slightly worse than the MIP solution, but the ACO algorithm took only a fraction of the computation time that was required by the MIP solver.