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  • Sudan  (38)
  • aquatic Oligochaeta  (35)
  • Turbulence
  • Springer  (76)
  • Annual Reviews
  • Blackwell Publishing Ltd
  • 2020-2024
  • 2005-2009  (2)
  • 1990-1994
  • 1980-1984  (74)
  • 2009  (2)
  • 1984  (74)
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  • 2020-2024
  • 2005-2009  (2)
  • 1990-1994
  • 1980-1984  (74)
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  • 1
    Publication Date: 2022-05-25
    Description: © 2008 The Author. This is an open-access article distributed under the terms of the Creative Commons Attribution Noncommercial License. The definitive version was published in Environmental Fluid Mechanics 8 (2008): 551-560, doi:10.1007/s10652-008-9076-5.
    Description: Experiments are reviewed in which a two-layer salt-stratified tank of water was mixed by turbulence. The density profile began as a single step and evolved to a smooth mixed profile. The turbulence was generated by many excursions of a horizontally moving vertical rod with Richardson number Ri 〉 0.9 and Reynolds Number Re 〉 600. There was almost perfect collapse of all the profiles to one universal profile as a function of a similarity variable. We develop a theoretical model for a simple mixing law with a buoyancy flux that is a function of internal Richardson number Rii. A similarity equation is found. A flux law that increases with small Rii and decreases with large Rii is considered next. Since no analytical solution is known, the similarity concept is tested by numerically integrating the equations in space and time. With buoyancy flux monotonically increasing with internal Richardson number, the similarity approach is valid for a profile starting from a slightly smoothed step. However, a shock forms for a mixing law with higher initial Rii (so that buoyancy flux decreases with Richardson number) and the similarity approach is invalid for those initial conditions.
    Keywords: Turbulence ; Mixing ; Stratified ; Similarity solution ; Layered fluid
    Repository Name: Woods Hole Open Access Server
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  • 2
    Publication Date: 2022-05-25
    Description: © 2008 The Authors. This is an open-access article distributed under the terms of the Creative Commons Attribution Noncommercial License. The definitive version was published in Environmental Fluid Mechanics 8 (2008): 495-509, doi:10.1007/s10652-008-9107-2.
    Description: Estuarine turbulence is notable in that both the dissipation rate and the buoyancy frequency extend to much higher values than in other natural environments. The high dissipation rates lead to a distinct inertial subrange in the velocity and scalar spectra, which can be exploited for quantifying the turbulence quantities. However, high buoyancy frequencies lead to small Ozmidov scales, which require high sampling rates and small spatial aperture to resolve the turbulent fluxes. A set of observations in a highly stratified estuary demonstrate the effectiveness of a vessel-mounted turbulence array for resolving turbulent processes, and for relating the turbulence to the forcing by the Reynolds-averaged flow. The observations focus on the ebb, when most of the buoyancy flux occurs. Three stages of mixing are observed: (1) intermittent and localized but intense shear instability during the early ebb; (2) continuous and relatively homogeneous shear-induced mixing during the mid-ebb, and weakly stratified, boundary-layer mixing during the late ebb. The mixing efficiency as quantified by the flux Richardson number Rf was frequently observed to be higher than the canonical value of 0.15 from Osborn (J Phys Oceanogr 10:83–89, 1980). The high efficiency may be linked to the temporal–spatial evolution of shear instabilities.
    Description: The funding for this research was obtained from ONR Grant N00014-06-1-0292 and NSF Grant OCE-0729547.
    Keywords: Turbulence ; Estuaries ; Shear instability ; Buoyancy flux
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  • 3
    ISSN: 1573-5117
    Keywords: Sudan ; Schistosoma mansoni ; Biomphalaria pfeifferi ; miracidia ; sporocysts ; host-parasite relations ; laboratory studies ; decoy snails
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Laboratory experiments have been carried out to determine the susceptibility of Gezira Biomphalaria pfeifferi snails to S. mansoni miracidia and the relationship between miracidia and daughter sporocyst production at the 10–17 day development stage. The relationship between snail numbers, miracidia numbers and water volume has also been studied. Two non susceptible snails, Bulinus truncatus and Cleopatra bulimoides, both of which occur naturally in Gezira canals, were tested to see if they act as decoys for S. mansoni miracidia. The results showed that the B. pfeifferi are 100% susceptible to S. mansoni invasion, at least to the daughter sporocyst development stage. The more miracidia that penetrated the more daughter sporocysts were produced, however individual variation and overlap were great. When one miracidium was released to find one snail it succeeded in low water volumes (5 m, 50 ml), but failed in 5 litres. When 100 miracidia were released mortality of snails was high suggesting superinfection particularly when only one or five snails were available. Among survivors daughter sporocyst counts were very high. Cleopatra and Bulinus snails do have a decoy effect when present in large numbers. In their presence the number of infected snails was marginally reduced and the number of daughter sporocysts greatly reduced. However, if superinfection is reduced by decoy effect, it is conceivable that Biomphalaria may be protected by decoy snails in circumstances where miracidia counts are high.
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  • 4
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    Hydrobiologia 110 (1984), S. 241-245 
    ISSN: 1573-5117
    Keywords: Sudan ; pearl oyster ; feeding ; growth ; Red Sea ; Sudan
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Plankton samples and gut contents of P. margaritifera were analysed monthly from April 1972 to March 1973. Coscinodiscus sp. was the most ingested food by pearl oysters of all ages. However, food variety increased as the oyster grew older. Experiments confirmed the absence of food selectivity in P. margaritifera. Unlike the young ones, the adult showed reduced growth during summer (July–September), which coincides with its spawning season.
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  • 5
    ISSN: 1573-5117
    Keywords: Sudan ; Schistosoma mansoni ; Biomphalaria pfeifferi ; miracidia ; sporocysts ; transmission of schistosomiasis ; host-parasite relations ; ecology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The efficiency of S. mansoni miracidia in locating and infecting Biomphalaria pfeifferi in Gezira canals has been studied under field conditions. When S. mansoni eggs were introduced into clean stagnant water in small field channels, the miracidia hatched to infect 100% of 30 snails in cages at the release point. Fifteen metres upstream and downstream 13% of caged snails were infected but no infections were found in snails 20 m away. When eggs were released into the same canal in flowing water (8.3 cm · s−1), no infections were detected in any of the caged snails placed 0–100 m downstream. Releasing hatched miracidia instead of eggs resulted in infections in all cages at 5 m intervals from 0-100 m. The release of eggs into flowing water was likened to the method by which S. haematobium eggs are deposited during urination. The 0% infection suggests that eggs will be swept away from the point of contamination by the flow. Thus only urination into stagnant water will lead to heavy snail infection rates. When eggs were released into a small pond-like minor canal tail end snail infection rates were only 3%. This was probably due to the larger water volume, smaller number of caged snails, and the presence of vegetation and other fauna which may be decoys or predators. The results highlight how very high snail infection rates can be produced under ideal conditions but also show how large snail and miracidia numbers are required in natural situations.
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  • 6
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    Hydrobiologia 115 (1984), S. 25-36 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; annelids ; evolution
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The Haplotaxidae have all the characteristics to support the hypothesis that they are the living descendents of the stem forms from which all of the Oligochaeta Clitellata (Orders Lumbriculida, Haplotaxida, Lumbricida, Tubificida) can be derived. The Aphanoneura are distinct from the Clitellata and are raised to a separate Class. There is no evidence to support the view that the elaborate setae of many Tubificida are derived from a polychaete ancestry; both are held to be independent modifications to aquatic life derived from a simple burrowing protoannelid with lumbricine setae.
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  • 7
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    Hydrobiologia 110 (1984), S. 213-217 
    ISSN: 1573-5117
    Keywords: Sudan ; Penaeus ; food ; food selection ; chemoreception
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract A study of food and food selection of Penaeus monodon (Fabricius) on the Sudanese Red Sea coast was carried out. Juveniles feed mainly on algal material. Adults feed on crustacea, annelids, algae, mud and unidentified matter. Adults were opportunistic in their feeding behaviour, but their preference was towards animal protein. Antennae, mouth parts and fore-chelipeds were found to carry long, slender setae which are probably of sensory function. Some setae on the carpus of the endopodite were serrated and seem to be used in grooming. Numerous tufts of setae in the chela of the fore periopods may hold a gustatory function. The inner edges of the chela are lined with a row of peg-like setae, which are probably another site of reception.
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  • 8
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    Hydrobiologia 110 (1984), S. 57-59 
    ISSN: 1573-5117
    Keywords: Sudan ; algae ; Blue Nile ; primary productivity ; tropical rivers ; White Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Phytoplankton production and respiration in the Blue Nile and White Nile at Khartoum were measured during the period November 1969–January 1971 using the light and dark bottle technique. Maximum rates of production coincided with periods of maximum phytoplankton densities. In the Blue Nile gross production varied between 0.00 gCm−3d−1 during the flood season and 2.19 gCm−3d−1 (0.49 mgO2l−1h−1) during November 1969. In the White Nile the range was from 0.41 gCm−3d−1 (0.09 MgO2l−1h−1) in May to 3.74 gCm−3d−1 (0.83 MgO2l−1h−1) in November. The maximum rates of respiration in the Blue Nile and White Nile were 0.10 and 0.63 MgO2l−1h−1 respectively. The ratios net:gross production were generally higher in the White Nile than in the Blue Nile.
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  • 9
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    Hydrobiologia 110 (1984), S. 61-77 
    ISSN: 1573-5117
    Keywords: Sudan ; algae ; Nubian desert
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The algae contained in some plankton samples from several localities on four wadi in N.E. Sudan were examined. The observed algae amount to 260 taxa, of which the diatoms, representing 50% of the algal flora, are the most numerous. Other large groups are the blue-green algae (17%) and the green algae (24%) among which the desmids (6%) are rather unimportant.
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  • 10
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    Hydrobiologia 110 (1984), S. 109-111 
    ISSN: 1573-5117
    Keywords: Sudan ; mangroves ; Red Sea ; ecology
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract An ecological survey of the mangrove vegetation in the littoral salt marshes of the Red Sea coast of the Sudan was made during short periods between 1980 and 1982. Avicennia marina (Forsk) Vierh occurs in pure stands of different dimensions in four types of locations. In each location, the responses of Avicennia are expressed in the degree of compactness of aggregation, canopy, height and regeneration potential. Grazing by camels is the primary agent in degradation of accessible locations; salt pans and to a limited extend cutting contribute in part to the process of degradation. The badly damaged aggregations may benefit from protection through control of grazing by nomadic camel tribesmen.
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  • 11
    ISSN: 1573-5117
    Keywords: Sudan ; Mollusca ; Gastropoda ; fresh water snails ; trematode parasites ; schistosomiasis
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract An account is given of the aquatic gastropod molluscs obtained in a survey made to assess future ecological effects of the Jonglei Canal now under construction in the Sudd region of southern Sudan. A total of about 23 species of aquatic snail lives in this region, of which a number are utilised as intermediate hosts by trematode parasites infecting man, livestock and wild herbivores, causing the diseases schistosomiasis, fascioliasis and amphistomiasis. Public health and veterinary authorities should be alert to the possibility that the Jonglei Canal will increase snail populations and raise the prevalence of parasitic diseases.
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  • 12
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    Hydrobiologia 110 (1984), S. 273-279 
    ISSN: 1573-5117
    Keywords: Sudan ; Alestes ; growth ; feeding ; Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The food, feeding habits and growth of 0+ Alestes nurse (Rüppel) from the Blue Nile near Khartoum, was investigated. 0+ A. nurse were observed to change their food from phytoplankton and small zooplankton during their pre-metamorphosis stages, to larger zooplankton (Daphnia spp., Chydorus sp.) and insects (larvae, pupae and imagines) during their post-metamorphosis stage. Their growth in length and weight was characterized by two distinct stanzas; during the first stanza, the fish increased in total length at a rate of 0.196 mm mm−1 wk−1, and in wet weight at a rate of 0.430 g g−1 wk−1. These rates were reduced during the second stanza to 0.046 mm mm−1 and 0.080 g g−1 wk−1, respectively. The calculated value of the exponent b in their length-weight relationship suggested that their growth was allometric.
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  • 13
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    Hydrobiologia 110 (1984), S. 281-286 
    ISSN: 1573-5117
    Keywords: Sudan ; frontal bone ; age determination ; cyprinidae ; Jebel Aulia
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract In spite of the exuberance of recent approaches to age determination of fish, many ecologists are convinced that the most valid method is age reading from permanent marks or growth rings laid on skeletal parts. Of the known structures used, only the vertebrae are suitable and readable for a large variety of species and sizes. Examination of different skeletal structures of Labeo horie, a Cyprinid collected from Jebel Aulia reservoir, revealed that the frontal bone carried well-defined growth rings. The validity of these rings was checked against other direct and indirect methods of age determination. Results proved to be statistically valid, and the frontal bone offers an additional device of age determination of L. horie.
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  • 14
    ISSN: 1573-5117
    Keywords: Sudan ; abundance ; breeding ; sex-ration ; Tilapia ; Labeo ; White Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Seasonal abundance, maturation and sex-structure were studied in populations of T. nilotica and L. niloticus in the Jebel Aulia reservoir and in the White Nile below the dam. L. niloticus showed a single annual breeding cycle from mid-July to December and was most abundant in October in the lake, and in early November in the river. T. nilotica showed two distinct annual seasons of abundance: summer (April to June) and autumn (November to October). Spawning activity was highest in summer (April to June) and autumn (late July to September). The sex-ratio varied inversely with age (size); males were predominant in populations of older fish, perhaps because of their greater endurance to adverse conditions as evidenced by their better survival in hypoxic water.
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  • 15
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    Hydrobiologia 110 (1984), S. 305-314 
    ISSN: 1573-5117
    Keywords: Sudan ; fish species ; selectivity ; abundance ; distribution ; fish production
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Developing lake conditions in Lake Nubia were found to offer good commercial fisheries potential. This prospect initiated fishery research since 1967. In this study, 26 fish species belonging to 10 families were recorded. Compared with previous studies the total number has decreased and some of the species that were commercially important during the early years of the lake formation, are now negligible in occurrence (Distichodus, Citharinus, Bagrus). The species widely distributed in the lake are Labeo niloticus, Lates niloticus, Sarotherodon niloticus, Hydrocynus forskallii, Alestes baremose and Eutropius niloticus. Their highest production occurs during two periods: February to May and July to September. The dominant fish species feed on different items in a balanced equilibrium with the ecological and biological conditions.
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  • 16
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    Hydrobiologia 110 (1984), S. 295-304 
    ISSN: 1573-5117
    Keywords: Sudan ; Clarias ; breeding ; maturation ; hormones ; growth ; cultivation
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The seasonal abundance, population structure, reproduction, feeding, respiratory adaptations and tolerance were studied in the catfish Clarias lazera (Cuv. & Val.) in the reservoir of Jebel Aulia on the White Nile, and in the Green-Belt canals near Khartoum. Continuous monitoring of the ripe, spent and spawning females and immature fish in the population revealed that C. lazera breed synchronously in the autumn in the reservoir and rather sporadically in the canals. Medium-sized females were more fecund than old and young females. Administration of pituitary extract enhanced maturation, induced spawning in mature fish, stimulated body growth, especially in maturing specimens, and produced, as a side-effect, a diuretic response. Mature fish depended on pulmonary respiration for about 45% of the total oxygen requirement. C. lazera fed preferentially on its fry and on decayed proteinaceous food. The suitability of this species for economic cultivation is discussed.
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  • 17
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    Hydrobiologia 110 (1984), S. 315-317 
    ISSN: 1573-5117
    Keywords: Sudan ; Lake Nubia ; fisheries
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The gill nets presently used in Lake Nubia fishing were subjected to a comparative study in regard to their relative efficiency. The results obtained indicate that Mesh 6 netted with Twine 4 is the most effective net, followed by Meshes 2, 4, 8 and 10 respectively. It has also been shown that, although individual nets undergo seasonal fluctuations in CUE, this does not, substantially affect their order of merit. The role of water characteristics on fish population distribution and subsequenct changes in the CUE was also noted.
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  • 18
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    Hydrobiologia 110 (1984), S. 319-325 
    ISSN: 1573-5117
    Keywords: Sudan ; Pisces ; Cyprinidae ; Barbus bynni ; breeding ; Jebel Aulia Reservoir
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Barbus bynni begins to mature at Age IV. Ripening of gonads of mature fish starts in May when water temperature approaches the annual maximum. However, the spawning season coincides with the onset of the flood season in July. These facts, as well as the cyclic growth of the gonads, show that B. bynni spawns once a year. Fecundity varies with size of fish and gonads. However, this levels off in the middle size group. At this age the fecundity was estimated to be 1 424 693 eggs.
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  • 19
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    Hydrobiologia 110 (1984), S. 327-332 
    ISSN: 1573-5117
    Keywords: Sudan ; Pisces ; Cyprinidae ; food and feeding
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Basic knowledge on the feeding ecology of one of the common and commercially important fish species in Jebel Aulia Reservoir is provided. The structure of the feeding apparatus indicates that Labeo niloticus is a bottom feeder, depending on soft and decayed vegetation, organic debris and whatever small organisms found within. However, juveniles and fry are prone to explore all layers and depths of the river selectively for plankton. There is little evidence of seasonal selection of food. Changes in diet quality are governed by the availability of type of food. Variability of feeding activity is connected with climate and breeding season.
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  • 20
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    Hydrobiologia 110 (1984), S. 333-338 
    ISSN: 1573-5117
    Keywords: Sudan ; fish ; Characidae ; growth of rings ; back-calculation ; aging structure
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The scales, opercula and vertebrae from 69 Hydrocyon forskali and 63 Alestes dentex were investigated for their reliability as aging structures. In both species the scales were unreliable due to their regenerative property, in spite of the clarity of their growth rings. Back-calculated lengths from the first growth ring laid on opercula and vertebrae resulted in values that coincided with the annual growth of the two species. In H. forskali, the mean back-calculated length from the two bones were 16.2 ± 5.9 and 16.8 ± 4.4 respectively, and for A. dentex, 11.1 ± 1.7 and 12.8 ± 1.8, respectively. Predictive equations for bone radius from fish length were written for both species. In this study, reliability was maximal between the observed and back-calculated lengths for both species (r 〉 0.95).
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  • 21
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    Hydrobiologia 110 (1984), S. 351-363 
    ISSN: 1573-5117
    Keywords: Sudan ; Polypterus ; aerial ; branchial ; respiration ; hypoxia ; hypercarbia
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The respiratory behaviour and partitioning of O2 uptake between air and water were investigated in Polypterus genegalus using continuous-flow and two-phase respirometers and lung gas replacement techniques P. senegalus rarely resorts to aerial respiration under normal conditions. Partitioning of O2 consumption depends on the activity and age of fish and the availability of aquatic oxygen. Immature fish (12–22 g) cannot utilize aerial O2 but older fish exhibit age-dependent reliance on aerial respiration in hypoxic and hypercarbic waters. Pulmonary respiration accounts for 50% of the total requirement at aquatic O2 concentrations of about 3.5 mg · l−1 (or CO2 of about 5%) and fish rely exclusively on aerial respiration at O2 concentrations of less than 2.5 mg · l−1. Branchial respiration is initially stimulated by hypercarbia (CO2: 0.5–0.8%) but increased hypercarbia (CO2 − 1%) greatly depresses (by over 90%) brancial respiration and initiates (CO2: 0.5%) and sustains pulmonary respiration.
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  • 22
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    Hydrobiologia 110 (1984), S. 339-349 
    ISSN: 1573-5117
    Keywords: Sudan ; Protopterus ; Polypterus ; Clarias ; ATP ; GTP ; development ; hypoxia
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The ATP and GTP contents of erythrocytes of Protopterus annectens, Polypterus senegalus and Clarias lazera were assessed in relation to maturation and hypoxia. The steady state ‘normal’ concentrations of ATP were: 446, 316 and 475 µM/100 ml and those of GTP were 286, 93 and 85 µM/100 ml in Protopterus, Polypterus and Clarias, respectively. The level of ATP decreased and that of GTP increased with increments in body growth, especially during development of the pulmonary organs. The concentration of GTP also increased progressively with hypoxia (O2: 7.4 to 3.6 mg/l); the level ATP remained almost unaltered. The ratio of GTP increased markedly with increased hypoxia particularly in the adaptive respiratory range of aquatic oxygen concentrations (6.5–4.8 mg/l) when the fish were forced to depart from branchial respiration and became reliant on air-breathing.
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  • 23
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    Hydrobiologia 110 (1984), S. 1-13 
    ISSN: 1573-5117
    Keywords: Sudan ; Nile ; hydrology ; control ; utilization
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The Nile is a vital resource of north-eastern Africa. After a description of its basin, and its hydrology,the different phases in the process of man gaining control on the river are discussed. Presently, the Nile is used for irrigation, the generation of hydroelectric power, navigation, and, of particular importance to limnologists, fisheries. Future developments in the management of the Nile are summarized.
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  • 24
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    Hydrobiologia 110 (1984), S. 15-19 
    ISSN: 1573-5117
    Keywords: Sudan ; RSC programme ; tailings ; impacts on environment
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The unique Red Sea environment has up to now been relatively unaffected by polluting consequences of Man's activities, except for evidence of oil along the coasts and reefs from transiting ships, harbours, industrial and human wastes with growing population and industrialization. The discovery, in the mid-sixties, of deposits of metalliferous muds, rich in heavy metals, and the probable mining and processing of these was expected to add to the risk of pollution. The Saudi-Sudanese Red Sea Commission, entrusted with the development and exploration of these non-living resources, has already set out an environmental study programme, more or less directly related to the conservation of the living marine resources. Within this framework, the studies have focused on the assessment and magnitude of possible risks for the environment resulting from a tailings disposal in the Atlantis 11 Deep over an extended period of time, and on the development of methods to minimize these risks. The results obtained so far, indicate that a well-controlled tailings disposal below 1 000 metre water depth would keep the environmental impact of such an operation in acceptable dimensions. But it is hoped that the forthcoming Pilot Mining Operations will be capable of clarifying some uncertainties through further computer modelling, monitoring a full scale tailings disposal test, using realistic input conditions and evaluation of the ecosystem.
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  • 25
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    Hydrobiologia 110 (1984), S. 33-44 
    ISSN: 1573-5117
    Keywords: Sudan ; lakes ; fauna ; distribution ; comparative survey
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Lakes Kundi and Keilak show large seasonal variations in area, but have permanent dry season residual areas rich in aquatic macrophytes. The faunas of the two lakes are similar, with L. Kundi (130 species recorded) being richer than L. Keilak (91 species). The fauna of L. Kundi includes the protomonad Rhipidodendron huxleyi, which has not been previously recorded in Africa. In L. Keilak we found Neozonocypris mirabilis, previously known only from L. Tanganika.
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  • 26
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    Hydrobiologia 110 (1984), S. 21-32 
    ISSN: 1573-5117
    Keywords: Sudan ; Blue Nile ; chemical composition ; tropical rivers ; White Nile
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    Topics: Biology
    Notes: Abstract Fortnightly measurements of physical and chemical variables were made at two locations on the Blue and White Niles near Khartoum from August 1968 to December 1970. Variables analysed from each river were: temperature, pH, total residue, current velocity, oxygen, alkalinity, phosphate, nitrate, ammonia, silica, sulphate, iron, calcium, magnesium, sodium, potassium and oxidizable organic matter. The seasonal variations of these factors in the two Niles are compared and the interrelationships existing between some of them are discussed. Comparisons with earlier studies on the Nile and with some tropical rivers are made. In the Blue Nile, the amounts of suspended matter and nutrients are largely dependent upon the flood regime. Nitrate, phosphate, iron, oxidizable organic matter and total residue increase considerably in the Blue Nile when the river is in flood (peaks: 1 880 µg NO3-N l−1; 0.31 mg Fe l−1; 3 842 mg total residue · l−1). In the White Nile, concentrations of nitrate, phosphate, iron, oxidizable organic matter and total residue attain their peaks during the rainy season (270 µg NO3-N l−1; 163 tag PO4-P l−1; 0.46 mg Fe · l−1; 502 mg total residue · l−1). In both rivers, alkalinity, calcium, sodium and potassium tend to increase during the dry season while declining in the rainy season. Silica is depleted at certain times of the year, yet relatively high concentrations are maintained throughout the year and were not expected to limit growth of diatoms. Fall in silica concentrations, unlike nitrate, phosphate and iron, was always followed by a rapid restoration of a high level. Silica and magnesium showed no response to changes in discharge rates.
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    Hydrobiologia 115 (1984), S. 127-129 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; biological quality indices ; large rivers ; pollution ; fine sediments
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    Notes: Abstract This study emphasizes the usefulness of Oligochaeta communities as descriptors of pollution in the fine sediments of large rivers. Two indices of biological quality of fine bottoms are proposed and are related to chemical parameters of the water. Tentative proposals are made for an empirical classification of polluted fine sediments.
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    Hydrobiologia 115 (1984), S. 37-44 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Tubificidae ; phylogeny
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    Topics: Biology
    Notes: Abstract A tentative phylogeny of the oligochaete family Tubificidae, with emphasis on the marine representatives, is presented. The scheme is based on the morphology and arrangements of prostate glands and the setal patterns. The rhyacodriline, more or less diffuse prostates are regarded as a primitive stage in prostate evolution, preceded only by the aprostate condition assumed for the ancestor of the family. An early split of the subfamily Rhyacodrilinae supposedly led to (1) a marine branch, from which evolved the highly diverse, exclusively marine subfamilies Phallodrilinae and Limnodriloidinae, and (2) a freshwater branch, which later divided into the Telmatodrilinae, Tubificinae and Aulodrilinae. The marine subfamilies invariably lack hair setae, whereas about half of the species within the other, freshwater subfamilies possess such setae in their dorsal bundles. Some marine genera, such as Monopylephorus (Rhyacodrilinae), Tubificoides and Clitellio (both Tubificinae) are regarded as recent off-shoots from the main freshwater stock. The families Naididae and Opistocystidae are considered likely to have evolved from rhyacodriline Tubificidae, whereas Phreodrilidae, the fourth family within the suborder Tubificina, is regarded as a sister group to the Tubificidae.
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    Hydrobiologia 115 (1984), S. 45-50 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; oligochaete taxonomy ; Enchytraeidae ; Grania
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    Topics: Biology
    Notes: Abstract The structure of the penial bulb and male efferent duct system of Grania species may be used in addition to setal pattern and spermathecal shape to distinguish species. Six penial bulb types are distinguished: (1) a simple, small, glandular bulb surrounding the male pore; (2) a small, glandular bulb, with a large, associated, dorso-medial gland mass; (3) a small glandular bulb, medial to the male pore, with an elongate male bursa (the aglandular sac), the vas deferens exitting directly into the invaginated male pore; (4) a glandular bulb with an aglandular sac and a small, cuticular stylet embedded in the bulb, extending from the ectal end of the vas deferens; (5) a glandular bulb and an aglandular sac with a long stylet extending from the vas deferens, through the bulb into the sac; and (6) glandular bulb reduced or absent, with or without an aglandular sac; with a long stylet and other prominent modifications, usually muscular, of the vas deferens. The details of the male duct structure were consistent within specimens grouped on the basis of setal distribution and shape and detailed spermathecal structure. Diverse male duct patterns are found within the polytypic species G. macrochaeta and G. postclitellochaeta. The positions of the spermathecal and male pores in their respective segments are distinctive for some species.
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    Hydrobiologia 115 (1984), S. 51-54 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Capilloventer ; Capilloventridae ; new family ; new genus ; Tubificida
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    Topics: Biology
    Notes: Abstract Specimens of a previously undescribed oligochaete from the Bay of Rio de Janeiro, Brazil were determined to belong to a new family of microdiles within the order Tubificida, suborder Enchytraeina. The specimens are characterized by dorsal and ventral hair setae, spermathecae in VII, testes in XI, ovaries in XII, and a clitellum. The number of primitive characters present in the new species indicate that it is a primitive member of the order that diverged early from the evolutionary line leading to the Enchytraeidae.
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  • 31
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    Keywords: aquatic Oligochaeta ; marine Enchytraeidae ; recognition of immature stages ; epidermal glands ; vital staining ; Neutral Red
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    Notes: Abstract Epidermal glands in Enchytraeidae are arranged in several transverse rows in each segment. These glands are more or less inconspicuous. It is possible to show up the gland cells by vital staining with Neutral Red (dilution 1:25 000 to 1:200 000 in tap- or sea-water, pH 6.0–6.8). Studies using this staining method have shown that there is a distinct pattern of epidermal glands for each species of marine Enchytraeidae investigated. The possibility of distinguishing immature specimens of different marine enchytraeid species, found in one particular locality, is demonstrated.
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    Hydrobiologia 115 (1984), S. 59-64 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; spermatozoa ; spermatogenesis ; cell differentiation
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    Topics: Biology
    Notes: Abstract Spermiogenesis in Oligochaeta occurs with peculiar modalities, common to all the species studied in this group. Gonial cells, produced in the testes, drop into coelomatic cavities (usually seminal vesicles) where they undergo a series of mitotic divisions, without cytodieresis, and finally meiosis. In the seminal vesicles, a series of ‘morulae’ is present, composed of 2, 4, 8, 18, 32, ... cells. The number of spermatids produced is variable, but is constant for each species. The process of spermiohistogenesis involves many steps, including: nuclear shaping, chromatin condensation, production of an acrosome, reduction of the number of mitochondria. The present knowledge of the mechanisms of cell differentiation, and the problem of the presence of two different sperm lines in some species is discussed.
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    Hydrobiologia 115 (1984), S. 65-69 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Tubificidae ; life-cycle ; karyology
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    Notes: Abstract Data on the life-cycle of a population of Branchiura sowerbyi Beddard in a water-lily tank at the Botanical Garden in Padua are reported. The breeding period is from April to July, after which the reproductive system is partially resorbed (August–September) and reformed later in the autumn. The karyology of the species was also studied, and revealed 38 mitotic chromosomes in the gonia, and 19 bivalents in the primary spermatocytes and in the primary oocytes.
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    Hydrobiologia 115 (1984), S. 71-81 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; nutrition ; sediment ; histophysiology
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    Notes: Abstract Lumbricillus lineatus selectively ingests masses of organic and inorganic interstitial particles from a sand-clay substratum in the upper littoral zone. Particle-masses are ingested, passed along the esophagus and into the anterior intestine where the pH becomes acid. A- and C-esterases, acid β-galactosidase, acid phosphatase and β-N-acetylglucosaminidase are present in the epithelium, while the rotating food masses are surrounded by a membrane of sulphated, acid glycoprotein. These enzymes, with the exception of acid phosphatase and the addition of aminopeptidase M, are also present in the epithelia of the mid and posterior intestinal regions where the pH is alkaline. The cells in the ventral wall of the mid intestinal region contain high concentrations of alkaline phosphatase, acid β-galactosidase and β-N-acetylglucosaminidase. The food consists of absorbed organics and bacteria with absorption and intracellular digestion occurring along the intestine, particularly in the mid ventral region.
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  • 35
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    Keywords: aquatic Oligochaeta ; gutless oligochaetes ; structure ; ecology ; physiology
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    Notes: Abstract Phallodrilus leukodermatus is not only characterized by the complete absence of mouth, gut, anus and nephridia, but also by an exceptional dermal ultrastructure which is associated with gram-negative bacteria. The vertical distribution of the worms from Bermudian carbonate sands is also unusual in attaining population maximum at oligoxic or anoxic depths around the redox discontinuity (RPD) layer, where extremely high concentrations of amino acids and sugars are to be recorded. Based on results from current ecophysiological and ultrastructural studies, an interpretation of the unique biology of the worms is attempted.
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    Hydrobiologia 115 (1984), S. 91-95 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; asexual reproduction ; genetical-ecological mechanisms
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    Notes: Abstract A great variety of asexual reproductive modes are known among aquatic oligochaetes. The main types of these modes are shortly described. Based upon observations on natural populations, the possible genetical and ecological implications of asexual reproduction are discussed. The following points are emphasized: (1) The often expressed expectations of a strong predominance of one particularly adaptive genotype is not born out. (2) In most cases, a number of genetically distinct clones are present in each population, and they show a strong differential distribution in heterogeneous environments, indicating an effective exploitation of the available resources. (3) Most cases of asexual propagation are reproductive strategies of their own and not escape mechanisms. (4) The mechanisms underlying asexual propagation are complex and involve many aspects of the life history. The great variety of types among aquatic oligochaetes offer particularly useful models for the study of these problems.
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    Hydrobiologia 115 (1984), S. 97-99 
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    Keywords: aquatic Oligochaeta ; Tubificidae ; parthenogenesis ; experiment
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    Notes: Abstract The ability of tubificids to reproduce parthenogenetically following or in place of bisexual reproduction has been proved. During parthenogenesis the spermatogenesis ceases at an early stage of sexual cell development, which, together with some peculiarities of the structure of the sexual system, rules out the possibility of self-fertilization in the worms. For two years of cultivation the life cycles of 3 parthenogenetic generations in Tubifex tubifex and 2 in Limnodrilus hoffmeisteri were traced.
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    Hydrobiologia 115 (1984), S. 101-104 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; life-cycle ; aquarial cultures
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    Notes: Abstract The maximum lifetime of species with exclusively sexual reproduction covers 5–15 years or even more in aquaria, although most specimens die consecutively earlier. The zooids of paratomic species live some weeks or months, their clones usually less than a year, rarely for some years. In the species with facultative architomy (fragmentation) the clonal age also exceeds that of the individual, both lasting for several years. The actual lifetime of worms in nature may be shorter than in cultures.
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    Hydrobiologia 115 (1984), S. 105-107 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Naididae ; growth ; experimental
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    Notes: Abstract The peculiarities of the growth in weight and length of Stylaria lacustris (L.) on the basis of observations in experimental vessels are considered. The growth of this species fits a parabolic curve. The equations relating weight to absolute growth rate as well as weight to duration of life are given.
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    Hydrobiologia 115 (1984), S. 109-112 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Psammoryctides barbatus ; growth ; cocoon production
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    Notes: Abstract The tubificid Psammoryctides barbatus is common in the sediments of English chalk streams. The species is primarily a spring and summer breeder with the most intense period of reproduction from April to June throughout its distribution in the river. Recruitment of juveniles takes place chiefly from June to August and most of the worms attain maturity in the following spring. In culture at 15 °C worms had attained 38 mm within 90 days of hatching and showed early signs of maturity. Growth rates and rates of cocoon production in culture are presented.
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    Hydrobiologia 115 (1984), S. 121-126 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; eutrophication ; oxygen demand ; population dynamics ; zoobenthos of lakes
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    Topics: Biology
    Notes: Abstract The paper attempts to combine the low oxygen content of the hypolimnion during stratification and the oxygen uptake of zoobenthos. Data of declining oxygen content in the hypolimnion and critical limits of respiration are combined for Chironomus anthracinus, Potamothrix hammoniensis and three species of Pisidium, P. casertanum, P. subtruncatum and P. henslowanum. The respiratory adaptation to low oxygen content influences both growth and population dynamics of the different species. The results have important bearing on eutrophication of the Lake Esrom ecosystem and temperate eutrophic lakes in general as well as the composition of profundal zoobenthos and its population dynamics.
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  • 42
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    Keywords: aquatic Oligochaeta ; Tubificidae ; experimental ; density regulation ; bionomic strategies
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    Notes: Abstract Laboratory research on the tubificids Psammoryctides barbatus and Spirosperma ferox was done. Embryonic development time, growth, time required to attain the first cocoon laying and egg production were estimated at different temperatures and population densities. The results allow us to demonstrate some intrinsic density regulation mechanisms in the profundal tubificid communities and substantiate the hypothesis that the succession from oligotrophic to eutrophic species in the profundal of lakes undergoing eutrophication is mainly based on their biotic characteristics.
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    Hydrobiologia 115 (1984), S. 131-138 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; benthos ; eutrophication ; indicator communities ; lake ; tubificid
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    Topics: Biology
    Notes: Abstract In 1978–80, oligochaete communities of meso-eutrophic Lake Léman (Lake of Geneva) were compared to those of mesotrophic Lake Neuchâtel. Worm species were classified into three groups corresponding to their increasing tolerance to eutrophication: (1) oligotrophic species, mostly Peloscolex velutinus, Stylodrilus heringianus; (2) mesotrophic species, mostly Potamothrix vejdovskyi, P. bedoti; (3) eutrophic species, mostly Potamothrix hammoniensis, P. heuscheri, Tubifex tubifex. In both lakes, eutrophic species constituted the bulk of the communities in terms of absolute abundance. However, relative abundance of mesotrophic and eutrophic species was higher in Lake Léman; oligotrophic species were more important in Lake Neuchâtel. These data confirmed the trophic classification of lakes based on chemical parameters. The number of zero values, which perturbated statistical analysis, was reduced by using species groupings instead of isolated species. Thus, making the lakes more comparable even if different species were present in each one. Relative density values based on all samples were distributed among 4 density classes for the 3 species groupings. The 12 resulting frequencies described the community structure expressed in terms of eutrophication. Furthermore, these frequencies may be used for comparison of eutrophication levels in several lakes.
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  • 44
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    Keywords: aquatic Oligochaeta ; pollution ; indicators ; toxicity
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    Notes: Abstract A series of recent studies have been completed by the authors involving: 1) determining the lethal tolerances of 12 oligochaete species classified (from ecological studies) as tolerant, moderately tolerant and intolerant to selected chemical toxicants and environmental factors under defined bioassay conditions with and without sediment; 2) determining lethal tolerances of candidate species to toxicants in combination with a range of abiotic factors; 3) measuring respiratory stress imposed by exposure to individual and combined sublethal concentrations of toxicants and environmental factors; and, 4) determining differences in lethal tolerance and respiratory stress between individual and mixed species. Surprisingly few previous studies have been done in this area considering the importance of oligochaetes as field pollution indicators. The results of the above major studies coupled with histopathological work are reviewed. Data from these studies substantiate the present use of oligochaete species assemblages as indicators of organic pollution and suggest their use in the laboratory for toxicant screening tests. The range of responses of different oligochaete species to individual and combined stress is complex, particularly in mixed species, which provides useful indications of specific stress factors. The application of these experimental laboratory studies to field situations is described.
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    Hydrobiologia 115 (1984), S. 153-158 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; long term changes ; population dynamics ; polychaetes ; recruitment
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    Topics: Biology
    Notes: Abstract Weekly sampling over a two year period from a muddy sand bottom in the polyhaline York River, Virginia, U.S.A., clearly identified the pattern of recruitment and survival of the dominant annelid species. Three intermingled recruitment strategies and two survival patterns were observed, ranging from the classic opportunistic life style of mass recruitment over short time periods followed by mass mortality to prolonged recruitment with lower mortality. Qualitatively the annelid assemblage was very similar from year to year with most of the changes being quantitative. Oligochaetes, Tubificoides spp., were the most stable and characteristic members of the annelid assemblage.
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    Hydrobiologia 115 (1984), S. 145-152 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; benthos ; principal component analysis
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    Topics: Biology
    Notes: Abstract The application of principal component analysis to two types of habitat (the benthos of macrophytes and of central river bed) enabled us to single out some of the factors that affect the dynamics and the structure of the oligochaete population and its various reactions to environmental conditions. As regards macrophytes, the distribution of the variables on the basis of the first component is correlated, to a certain extent, with a seasonal factor without any significant differences among sites. The largest population is most closely correlated with the summer months. In fact, we found that the Naididae and Tubificidae species generally develop in larger numbers at higher temperatures. For the Tubificidae, we could detect a precise seasonal cycle. In the central river bed habitat, the first component was correlated with the river discharge, which determines the granulometric characteristics of the sediment; we noticed a correlation among the sites that have the same characteristics, regardless of sampling site or date. The species which correlate most closely among themselves are the Tubificidae Limnodrilus hoffmeisteri, Tubifex tubifex, L. udekemianus and L. profundicola, which are very characteristic of environments that contain abundant organic matter. The second component is correlated with temperature, and hence with the availability of oxygen, which determines the presence and the abundance of more sensitive species.
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    Hydrobiologia 115 (1984), S. 159-170 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Enchytraeidae ; environmental parameters ; ordination techniques
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    Topics: Biology
    Notes: Abstract The Enchytraeidae are essentially terrestrial oligochaetes but many species have marked aquatic tendencies. Over two thirds of recorded Irish species were found in soils which were submerged or frequently flooded and 35% showed a distinct preference for these conditions. Relatively few species were living in soils subject to drought. Red blood was present in 28 species, all but one from soils with more than 55% water. Cognettia sphagnetorum and C. glandulosa developed red blood in very wet conditions. In a survey of Irish wetlands, samples were taken from bog, heath, marsh, fen, margins of lakes and rivers, and saltmarsh. The influence of various environmental parameters was determined using ordination techniques. Magnesium and pH were found to be the most important factors. A high level of magnesium distinguished coastal sites and pH 5.2 separated two clusters representing acid peat and marsh-fen-aquatic sites. Groups of indicator species characterized each of the three clusters. The ecological distribution of the indicator species is described, and their usefulness in classifying enchytraeid communities is discussed.
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    Hydrobiologia 115 (1984), S. 171-174 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; diversity ; respiration ; production
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    Topics: Biology
    Notes: Abstract Data are presented concerning an investigation into the diversity, abundance, production and respiration of the oligochaete fauna in the eutrophic, polymictic Lake Zbechy and in a melioration canal in the Wielkopolska Region, an area of intensive agriculture. It was found that in the canal the average biomass of oligochaetes was about four times higher than in the lake. Oligochaetes expend 1.1–4 times more energy in respiration than in tissue production. Species diversity and species number are positively correlated, while the correlation between diversity and abundance is negative.
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    Hydrobiologia 115 (1984), S. 175-182 
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    Keywords: aquatic Oligochaeta ; biotypology ; French upper Rhône
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    Notes: Abstract Within a framework of interdisciplinary research, a biotypological trial of the epigean and underground fauna (Oligochaeta and Aphanoneura) of the alluvial plain of the French upper Rhône is proposed, according to the partition of this hydrographic complex into superficial and subterranean functional sets.
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    Hydrobiologia 115 (1984), S. 183-186 
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    Keywords: aquatic Oligochaeta ; water quality ; bioindication
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    Topics: Biology
    Notes: Abstract Using as examples water bodies in different geographical zones (Lake Ladoga, the River Sukhona, shallow-water ponds of the North Caucasus) the known methods of water quality evaluation by means of oligochaetes are considered. There is no unique universal method of pollution bioindication in. this way. In water bodies of all types the mass development of oligochaetes and the reduction of their species diversity are indications of. (1) large quantities of organic matter; (2) favourable oxygen regime; (3) absence or insignificant quantity of heavy metal solids, petroleum substances and agricultural chemicals; (4) intensive self-purification of the water body.
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    Hydrobiologia 115 (1984), S. 187-190 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Lumbricomorpha ; biology of aquatic earthworms ; breeding of aquatic earthworms
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    Topics: Biology
    Notes: Abstract Twelve species of aquatic earthworms have been found in European inland waters where they occupy different environments. Two species live exclusively in the sapropel, that is in the black fetid asphyctic mud, while four species can be found either in the sapropel or in the more oxygenated gyttja. The remaining species prefer this type of soil, but Eisenia spelaea in Italy lives only within submerged litter in mountain streams. The biology of these earthworms is discussed and the possibility of their utilization by man is considered.
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  • 52
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    Keywords: aquatic Oligochaeta ; Naididae ; Tubificidae ; Israel & Sinai distribution ; habitat characteristics
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    Notes: Abstract Nineteen species of Naididae and fourteen species of Tubificidae were found in collections from inland waters in Israel and Sinai. The local distribution patterns of the majority of these species were determined. The ranges of several physical and chemical variables characterizing the habitats of each species were defined. A correlation was found between the distribution patterns of some species and the following variables: salinity, temperature, dissolved oxygen, current velocities and stability of the habitats.
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    Hydrobiologia 115 (1984), S. 191-196 
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    Keywords: aquatic Oligochaeta ; Tubificidae ; zoogeography
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    Topics: Biology
    Notes: Abstract The specific and generic diversities of the marine Tubificidae (Annelida, Oligochaeta) of the NE Pacific are compared to those of the NE and NW Atlantic as well as to those of Heron Island, Australia. Diversity in the NE Pacific is relatively high when compared to that of the NE Atlantic. The Tubificidae of the NW Atlantic (limited to the eastern coast of the USA) show two distinct zoogeographic regions: Florida to Cape Hatteras; Cape Hatteras to Massachusetts. Diversity, both in terms of the number of species and number of genera, is approximately the same in these two regions, and is similar to that of both the NE Pacific and Heron Island. Evidence suggests that the widespread marine species, in particular Tubificoides pseudogaster, have a range of morphotypes across their distributions. The apparent wide distributions of these species may be due to a taxonomy unable to resolve the differences between the morphotypes. The tubificid oligochaete fauna of the NE Atlantic appears impoverished compared to the other regions examined. The NE Pacific, NW Atlantic, and Heron Island regions are not dominated by one group of species while the NE Atlantic fauna is dominated by Tubificoides benedeni and Clitellio arenarius.
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  • 54
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    Keywords: aquatic Oligochaeta ; Tubificidae ; Enchytraeidae ; zoogeography
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    Notes: Abstract On the basis of material collected along the Ligurian, Tyrrhenian, Ionic and Adriatic coasts of Italy, a tentative list of 26 species (18 tubificids, 8 enchytraeids) of marine Oligochaeta is presented. Most of the species are new to science, and it can therefore be concluded that there is a very high diversity of oligochaetes in the Mediterranean Sea.
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  • 55
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; distribution according to sediment type
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    Topics: Biology
    Notes: Abstract This paper aims to describe the ecology and determine the distribution of Oligochaeta according to sediment type, depth and other environmental factors in 21 lakes of the National Park of the Lithuanian SSR. Diagrams are included showing the distribution of the lakes according to the index of density and biomass of Oligochaeta in the trophic structure of biocenosis of the zoobenthos. Light is thrown on the problem of sediment composition as a significant factor in the distribution and development of aquatic Oligochaeta.
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    Hydrobiologia 115 (1984), S. 215-222 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; fenland ; cluster analysis ; ecological groups
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    Topics: Biology
    Notes: Abstract A synoptic study of the aquatic ecosystems in the Dutch province of Overijssel revealed the presence of 33 oligochaete taxa in the fendland area of N.W. Overijssel. The material was collected at 101 sampling sites in 1981. Oligochaetes were grouped by means of a normal and inverse cluster analysis. Although most oligochaetes are quite biquitous, differences in occurrence and abundance were observed and related to minor differences in environmental parameters.
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    Hydrobiologia 115 (1984), S. 223-230 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; biogeography ; ecology
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    Notes: Abstract The profundal benthic fauna of 63 Spanish reservoirs was studied during 1972–75. Samples were taken in the deepest part of each reservoir with a modified Van Veen grab. The fauna was dominated by tubificid worms and chironomids. Sixteen species of Oligochaeta were found, two of them new for the Spanish fauna (Potamothrix heuscheri and Haber pyrenaicus). This was the first time H. pyrenaicus was found since the species was described in 1974 from the Pyrenees. Four species were frequent in the reservoirs (Limnodrilus hoffmeisteri, Tubifex tubifex, L. claparedeianus and Dero digitata) but only the first two were abundant as mature individuals. Another interesting fact is the high abundance of immature animals in all samples, accounting for more than 80% of the total individuals examined. The geographical distribution of all species in Spain and their relationship with the typology of Spanish reservoirs are discussed. There seems to be a correlation between the throphic status of the reservoirs and the relative proportion of L. hoffmeisteri as compared to T. tubifex, the number of individuals of the first species increasing with water eutrophication.
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    Hydrobiologia 115 (1984), S. 15-17 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Lumbriculidae ; phylogeny
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    Topics: Biology
    Notes: Abstract The rudimentary atria, and the posterior sperm funnels and sperm ducts, which occur in some species of the Lumbriculidae, are discussed. It is shown that the posterior location of funnels and sperm ducts is the result of a forward shift of the atria, which refutes Stephenson's supposition that the Lumbriculidae is the most archaic family of the present-day Oligochaeta.
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    Hydrobiologia 115 (1984), S. 3-13 
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; oligochaete spermatozoa ; acrosome ; electron microscopy ; phenetics ; phylogeny
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    Topics: Biology
    Notes: Abstract Spermatozoal ultrastructure of nine oligochaete families has been examined for congruence with phylogenetic and taxonomic systems for the Oligochaeta based on general morphology, particularly the holomorphological hennigian analysis of Jamieson (1978a, 1980, 1983). Estimation of congruence has been made following phenetic and cladistic (phylogenetic) analysis. Correspondence, in phenograms and phylograms, of sperm types with taxonomic and phylogenetic groupings previously recognized is generally good. Departure from this rule in the similarity of the phreodrilid sperm to that of the Lumbricina suggests a corresponding alteration of fertilization biology in the phreodrilids. The results indicate that the Haplotaxidae lie at the base of the opisthopores though they do not unequivocally contraindicate acceptance of a Haplotaxis like form as a stem form of the Haplotaxida (opisthopores and Haplotaxidae) and Tubificida. An even more basal position for prosopores, now represented by the Lumbriculida, cannot yet be dismissed.
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  • 60
    ISSN: 1573-5117
    Keywords: aquatic Oligochaeta ; Tubificinae ; phylogeny ; taxonomy ; chitinous penis sheaths
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    Topics: Biology
    Notes: Abstract Setae and chitinous penis sheaths are the main characters used to distinguish genera and species of the subfamily Tubificinae. Genital setae and penis sheaths are of functional importance to facilitate copulation. Similar structures in different genera may be homologous or products of parallel evolution (homoiologous). Form and dimensions of the penis sheaths of many tubificine species are very variable. Transspecific overlap of quantitative species characters can make the determination of specimens in the genus Limnodrilus difficult. The configuration of the distal ends of the penis sheaths is an important character to distinguish Limnodrilus species. The definition of the intraspecific variability of this morphological character is problematic.
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    Hydrobiologia 110 (1984), S. 47-55 
    ISSN: 1573-5117
    Keywords: Sudan ; algae ; Blue Nile ; plankton ecology ; tropical rivers ; White Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The variation in numbers of the major planktonic algae in the surface waters of the Blue Nile and White Nile were followed for 29 months (August 1968–December 1970). In terms of biomass, the phytoplankton was dominated by diatoms and blue-green algae; other groups were of minor importance. High phytoplankton densities occurred during winter in both rivers, and during summer in the Blue Nile, with values in the latter river greater than in the former. Melosira granulata and its variety angustissima and Anabaena flos-aquae var. intermedia f. spiroides were the most important taxa in the two Niles. Melosira distans and Attheya zachariasi, which are reported for the first time in the present study, preponderated at certain times in the Blue Nile. Microcystisflos-aqua which had been a minor component of the phytoplankton in both rivers during the early 1950s, showed a profuse growth in the Blue Nile. Lyngbya limnetica, Anabaenopsis cunningtonii and A. tanganyikae, which constituted important components of the phytoplankton in the early 1950s, either disappeared or maintained themselves in reduced numbers during the present study. This change in the algal flora of the two Niles may be attributed to the construction of the Roseres dam across the Blue Nile in 1966 and to the invasion of the White Nile by Eichhornia crassipes in 1957.
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    Hydrobiologia 110 (1984), S. 79-90 
    ISSN: 1573-5117
    Keywords: Sudan ; swamp vegetation ; Sudd
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    Topics: Biology
    Notes: Abstract This paper describes vegetation in the nothern Sudd area of the Sudan. A visit by boat was made in April 1978 along the White Nile from Malakal to Lake No; to the inflow regions of the Bahr el Jebel, the Bahr el Zeraf and to the Bahr el Ghazal. Only the vegetation along the fringe of the rivers was observed as plants were tall and dense and it was not possible to penetrate into the inner swamps. The outer edge of the vegetation was fringed by extensive mats of free-floating Eichhornia crassipes which was easily dislodged and formed floating islands to be carried downstream. The main rivers, excluding the Bahr el Ghazal, were too deep and turbid to support beds of submerged species but plants such as Ceratophyllum demersum could be found amongst floating plants. Vossia cuspidata usually produced a broad zone to the landward of Eichhornia. It dominated the shallow water with a floating raft of stems growing out from the shore over the water surface. Massive stands of Cyperus papyrus and/or Phragmites karka, over 5 metres tall, dominated the vegetation behind and sometimes excluded the Vossia zone. The papyrus could form dense, floating rafts of rhizomes at the water's edge, but Phragmites always remained firmly rooted to the bottom. Floating papyrus and Vossia would become detached from the main beds and drifted downstream with Eichhornia. Creepers were common amongst papyrus but less common amongst Phragmites where the vegetation was generally more dense. Large stands of Typa domingensis were observed behind the fringe vegetation. The vegetation of the Bahr el Ghazal, in marked contrast to the rest, was stunted, and reasons for this are suggested. Species diversity was greater and the less turbid waters allowed considerable development of floating-leaved and submerged species. Estimates of fresh weights and heights of dominant vegetation types were attained but methods were very laborious and alternative ways have been suggested. The fresh weight of Eichhornia from a one metre square quadrat in the White Nile was around 70 kg. The shoots of Phragmites weighed 18.5 kg and Cyperus papyrus, 11.0 kg m−2.
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    Hydrobiologia 113 (1984), S. 93-98 
    ISSN: 1573-5117
    Keywords: tropical zooplankton ; swamps ; Sudan
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    Topics: Biology
    Notes: Abstract Samples from the main Nile channel yield a poor zooplankton, deficient in species and heavily laden with organic debris. Samples from side arms and floodplain lakes yield a richer zooplankton, often containing 15–20 species of rotifers and ten or more species of Crustacea. When the floodplain lakes are heavily vegetated the number of species increases. For the Crustacea the index of diversity (α) normally varies between 0.6 and 2.4, but in a side arm of the Bahr el Zeraf it reaches 6.1 and in Lake Ambadi on the Bahr el Ghazal it reaches 8.4. The diversity of the rotifers is somewhat higher, normally ranging between 2 and 4.5, but reaching 23 in Lake Ambadi. The zooplankton of Lake Ambadi is the most distinctive and contains several species not found at other stations. The reasons for the peculiar zooplankton of Lake Ambadi are discussed.
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    Hydrobiologia 110 (1984), S. 99-108 
    ISSN: 1573-5117
    Keywords: Sudan ; water hyacinth ; nutrients ; P, N, Fe ; growth ; White Nile
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    Topics: Biology
    Notes: Abstract The possibility that the stunted growth of the water hyacinth in Bahr el Ghazal river in Sudan is influenced by nutrient elements is considered. Greenhouse experiments were carried out to determine the effects of deficiency and mineral nutrient additions on the growth of this plant. The water hyacinth was found to grow at a wide range of nutrient levels. Maximum growth was recorded at 21 mg l−1 N, 62 mg l−1 P, and 0.60 mg l−1 Fe.
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    ISSN: 1573-5117
    Keywords: Sudan ; water hyacinth ; biological control ; weevils
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    Topics: Biology
    Notes: Abstract Water hyacinth Eichhornia crassipes (Mart.) Solms. is an aquatic weed that infests most of the White Nile system in the Sudan. Serious economical and ecological problems are caused by this weed. The two weevils Neochetina eichhorniae and Neochetina bruchi were imported and released in an attempted biological control against the weed. The adults of these weevils attack the plant and feed by removing tissues from the leaf pseudolamina and petioles. The larvae tunnel inside the petioles and the crown. The optimum temperature for feeding and development of both species is 25° C. Results obtained from stocking hyacinth plants with adults and larvae of both species separately revealed that N. bruchi is more efficient in checking the growth of the plant. The progeny of a pair of N. bruchi and N. eichhorniae reared separately on 41 hyacinth plants for a period of 61 days (one generation period) reduced their population growth by 25.4% and 12.7% respectively. The progeny of both species in a mixed culture reduced the growth of the plants by 22.5% in the same period, while in the control the population of the plants increased 136.6%.
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    Hydrobiologia 110 (1984), S. 113-130 
    ISSN: 1573-5117
    Keywords: Sudan ; rotifers ; taxonomy ; biogeography ; Africa
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    Topics: Biology
    Notes: Abstract A collection of rotifers contained in samples from the Blue, White and joint Niles in the Sudan, the Red Sea Hills, and Jebel Marra mountains is studied, and the previous literature on the rotifera of Sudan is reviewed. A total of 145 rotifer taxa are now known from this country. Ecological and distributional notes on selected species are added. Besides cosmopolitan, pantropical, and tropical species, a small but significant fraction of the fauna is of northern origin, possibly reflecting climatic conditions of the late-Pleistocene.
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    Hydrobiologia 110 (1984), S. 131-134 
    ISSN: 1573-5117
    Keywords: Sudan ; amphipoda ; Bogidiellidae ; systematic ; ground water fauna
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    Topics: Biology
    Notes: Abstract Description of Bogidiella nubica, from interstitial waters in the Sudan. It is the first Bogidiella species known for the African continent, except for the Mediterranean area.
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    Hydrobiologia 110 (1984), S. 135-136 
    ISSN: 1573-5117
    Keywords: Sudan ; Nile ; Acari ; Halacaridae
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    Topics: Biology
    Notes: Abstract Limnohalacarus africanus and L. fontinalis are recorded from Lake No. These are respectively the third and second records of these species, and represent considerable extensions of their known ranges.
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    Hydrobiologia 110 (1984), S. 163-169 
    ISSN: 1573-5117
    Keywords: Sudan ; Cladocera ; biogeography ; taxonomy ; Sudan ; Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Twenty species of Cladocera are reported from the Nile, where lacustrine species dominate, and from Jebel Marra and the Red Sea Hills, where chydorids dominate. The community found in the Red Sea Hills is more typically desertic than that of Jebel Marra, which appears closely related to the fauna of the West and Central African Sahel.
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    Hydrobiologia 110 (1984), S. 171-175 
    ISSN: 1573-5117
    Keywords: Sudan ; Crustacea ; Bathynellacea ; Nile
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    Topics: Biology
    Notes: Abstract Nilobathynella predynastica n.g., n.sp. from Abri, Nubia, is the first representative of the Tribus Cteniobathynellini in the Nile Valley. Its relationship with the other Cteniobathynellini, as well as its biogeographical significance, are briefly discussed.
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  • 71
    ISSN: 1573-5117
    Keywords: Sudan ; freshwater ostracods ; Africa ; taxonomy ; zoogeography
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    Topics: Biology
    Notes: Abstract A collection of freshwater ostracods from the Sudan is studied and 18 species are recorded. Paracypretta amati sp.n. is new to science. A redescription of the holotype female of Paracypretta aratra (Brady, 1904) (not recorded in this collection) and a provisional key to the species of Paracypretta are given. Heterocypris giesbrechtii (G. W. Müller, 1898) is re-described; Heterocypris sobrinus (Masi, 1932) is placed in the synonymy of H. fretensis; Hemicypris intermedia (Lindroth, 1953) is recorded for the first time since its original description and is redescribed, while Cyprinotus largereticulatus Rome, 1969 is transferred to Hemicypris. The zoogeographical composition of this fauna is briefly discussed.
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    Hydrobiologia 110 (1984), S. 177-180 
    ISSN: 1573-5117
    Keywords: Sudan ; Nubia ; groundwater ; Copepoda ; Nitocrella
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    Topics: Biology
    Notes: Abstract Description of Parapseudoleptomesochra attirei from a well in a dry wadi bed at At Tire, Nubia. Its closest relative is found in a semidesert area in Iran. The chorological implications of this finding are briefly discussed.
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    Hydrobiologia 110 (1984), S. 181-190 
    ISSN: 1573-5117
    Keywords: Sudan ; Odonata ; Nile ; Red Sea Hills ; biogeography ; deserts
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract Twenty-one species of dragonflies are recorded from the Nile and the Red Sea Hills in the Sudan. The majority are species of africotropical origin, but wide-ranging and tolerant of strongly fluctuating environmental conditions. A small but significant fraction is restricted to the Nile Valley, and another fraction, found only in the Red Sea Hills, is composed of Eremian species of Palaearctic origin. Besides adults, the last-instar larvae of Pseudogrion niloticum and of Paragomphus pumilio are described and figured.
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    Hydrobiologia 110 (1984), S. 191-212 
    ISSN: 1573-5117
    Keywords: Sudan ; Copepoda ; Calanoida ; taxonomy ; biogeography ; Africa ; Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract At least 11 or 12 calanoid taxa occur in the Nile system. One species is typical of the river and of Lake Chad (Th. galebi); two are restricted to Lake Victoria (Th. galeboides, T. stuhlmanni), but both might be only subspecies to more widespread species. One is restricted to Lakes Edward and George (T. worthingtoni) but is a little known species. No calanoids are on record from Lakes Albert and Kyoga, while the species reported from Lake Turkana (T. banforanus) is out of range, and almost certainly represents an erroneous record. Among the remaining species, four are East-African, ranging from the southern tip of the continent to the Ethiopian plateau and the Nile valley (P. schultzei, Th. mixtus, T. kraepelini, T. cf orientalis), while two are Sahelian species that span Africa from east to west (M. mauretanicus, T. processifer et ssp.). Two new synonyms are introduced. One new subspecies (T. processifier friedae) is described from the Ethiopian plateau. It is suggested that Tropodiaptomus orientalis (Brady, 1886), the type species of its genus, should not be considered a nomen dubium, but should be redescribed on its type female(s) and on topotypical males from Sri Lanka.
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    Hydrobiologia 110 (1984), S. 219-225 
    ISSN: 1573-5117
    Keywords: Sudan ; Bivalvia ; Corbicula ; ring formation ; White Nile
    Source: Springer Online Journal Archives 1860-2000
    Topics: Biology
    Notes: Abstract The growth of C. consobrina of the White Nile near Khartoum was studied during March–December, 1977 from material collected from fish guts every two months. It was found that the growth season extends from February to July in correlation with the rise in temperature. Food was abundant and the concentration of the dissolved calcium in the river water was also increasing. During the flood season from July to October, the clams entered a resting state as a result of the annual siltation and consequently growth was arrested. Growth rings were found to be formed as a result of the siltation-induced resting state. There is evidence that little growth occurred during the cooler months from October to December. The development of the false rings on the shells is possibly a result of the changes in the water levels. Analysis of the growth of the adult clams showed that there is a clear size-time relation; there was close agreement between the calculated maximum sizes and the actual data collected from the field. Other examples from Sudanese inland waters were also discussed. The role of the annual hydrological cycles in regulating the populations activities and hence the growth of the individuals was found to be important.
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    Annals of biomedical engineering 12 (1984), S. 357-369 
    ISSN: 1573-9686
    Keywords: Oscillatory flow ; High frequency ventilation ; Unsteady pipe flow ; Turbulence ; Pulsatile flow ; Transition
    Source: Springer Online Journal Archives 1860-2000
    Topics: Medicine , Technology
    Notes: Abstract Turbulence during pulsatile flow has been suggested as a possible mechanism to enchance the transport of gases during high-frequency ventilation. Experimental studies on oscillatory flow in straight, circular tubes have identified three types of flow: (a) laminar; (b) conditionally turbulent, in which high-frequency disturbances occur during the decelerating phase of the flow cycle but relaminarize by the beginning of the subsequent accelerating phase; and (c) fully turbulent flow, in which disturbances occur throughout the flow cycle. Fully turbulent flow has been observed only when a mean flow is present, and only laminar or conditionally turbulent flow has been observed for purely oscillatory flow. A critical Reynolds number based on the Stokes layer can be defined, and transition Reynolds numbers between 400 and 550 have been experimentally determined for purely oscillatory flow in a circular tube, although lower values are expected for physiological flows. There are some indications that the structure of oscillating turbulent flow is similar to steady turbulent flow, and preliminary work in our laboratory shows that the spectral content of flows during high-frequency ventilation is similar to that in steady turbulent flow.
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