ALBERT

Notes: Experiments were carried out on seven sites in Fermanagh to investigate the yield loss caused by leatherjackets in grassland. Populations on the sites ranged from 65,000 to 865,000 ha−1 and significant damage was recorded from all sites except the least populated. Greater yield increases were obtained by controlling leatherjackets in September when compared with control in March. Regression models were fitted and it was found that 125,000 leatherjackets ha−1 in March caused a yield loss of 50 kg herbage dry matter (DM) ha−1 by mid-May. Based on these figures the average yield loss to leatherjackets in Northern Ireland, from 1965 to 1982, was 208 kg DM ha−1. Controlling leatherjackets in September, rather than March, increased the potential avoidable yield loss by a factor of 2.72. The average yield loss at first silage cut is therefore 566 kg DM ha−1. On average, 100,000 ha grassland in Northern Ireland may be suffering an annual loss of 1t herbage DM ha−1.

Notes: The utilization of sown and indigenous plant species was studied in three experiments in which plots were stocked with similar live weights of sheep and goats. In the first experiment the animals grazed plots containing 0–5 ha of rush (Juncus effusus)- infested reseeded pasture and 0–5 ha of unimproved blanket bog. The second and third experiments took place on old rush-infested improved pasture; in one experiment two levels of herbage mass of grass were provided while in the other the rushes were cut in spring or remained uncut.The goats grazed the rushes readily in all three experiments. Reduction in herbage mass of grass increased utilization of rushes by goats although these animals still grazed rushes when grass supply was plentiful. Sheep scarcely grazed J. effusus even when hard-pressed by shortage of grass. Both sheep and goats grazed J. acutiflorus.In Experiment I sheep utilized reseeded pasture more heavily than did goats. Sheep grazed similar proportions of the leaves of grass and clover while goats grazed a lower proportion of clover leaves as compared with grass. Among the grasses sheep discriminated in favour of Lolium perenne whereas goats did not. On the blanket bog vegetation, Eriophorum vaginatum and Calluna vulgaris were grazed both earlier in the season and more heavily by goats than by sheep.The relationships between sward structure and grazing height in accounting for differences in species selection by sheep and goats are briefly discussed. The possibility of using goals to control coarse weeds in hill pasture and for strategic grazing to manipulate floristic composition is outlined.

Notes: The performances of a rising-plate meter and a single-probe electronic capacitance meter were compared for estimating the herbage mass of irrigated pastures both before and after grazing by dairy cows. Mean CV for the rising plate meter and the single-probe meter were 12.7% and 13.3% respectively when herbage mass was measured before grazing, and 21.8% and 15.4% respectively immediately after grazing. These coefficients of variation indicate that while the performance of the two meters was similar when herbage mass was measured before grazing, the single-probe meter was a more precise instrument for estimating herbage mass after grazing.Trampling of herbage prior to measurement had a major effect on the regressions of both meters; the regression intercept was increased by 2800 kg dry matter (DM) ha−1 for the rising-plate meter and by 2580 kg DM ha−1 for the single-probe meter. The slopes of the graphs, however, remained unchanged.It was concluded that while the single-probe meter performed better than the rising-plate meter for estimating the mass of post-grazing herbage when trampling was negligible, the large effect that trampling had on the post-grazing calibration regressions precluded both instruments from general use in dairy cattle research. This indicates that another technique is required in circumstances in which pastures are heavily trampled.

Notes: Only a very limited number of species, including red fescue (Festuca rubra), perennial ryegrass (Lolium perenne), timothy (Phleum protense) and white clover (Trifolium repens) are considered as suitable for wet hill land. Detailed comparison of red fescue, perennial ryegrass and timothy on contrasting acid brown earth and stagnogley soils from 1966 to 1969 at Pant-y-dŵr Hill Centre (305 m OD) highlighted the advantages of red fescue in annual DM production and its more even distribution over a longer growing season but also indicated drawbacks of slow initial establishment and lower feed quality. In vitro DOMD of red fescue averaged 645 g kg−1 over the year compared with 690 g kg−1 for perennial ryegrass and timothy. Assessment of a narrow range of species under surface seeding conditions on a stagnogley soil from 1971 to 1975 confirmed the value of red fescue in DM production and persistence. The results are discussed in relation to the characteristics required in herbage species for hill land improvement.

Notes: High-yielding grass-legume mixtures play an important role in forage-animal systems but finding compatible and adapted species can sometimes be difficult. The objective of this study was to examine productivity of perennial ryegrass and orchardgrass in pure stands with N and in mixtures with legumes.Broadcast sowings were made on conventionally prepared seedbeds in August 1979 and May 1980 on a Hagerstown silt loam soil (fine, mixed mesic Typic Hapludalf). When sown alone orchardgrass cv. Pennlate and perennial ryegrass cv. Reveille received rates of N ranging from 0 to 448 kg ha−1 a−1. The grasses were also grown in mixtures with alfalfa cv. Arc, red clover cv. Arlington, or birds foot trefoil cv. Viking. The same legumes were sown alone. Four legume seed rates were used in both the pure legume and mixed stands.Orchardgrass-N swards were more productive than ryegrass-N swards over 3 years. More dry matter (DM) was harvested from pure stands of

Notes: Fungicide applied to plots of perennial ryegrass cv. S24 in the autumn of 1981 significantly increased tiller number and total dry matter accumulation in February and March of 1982. However, fungicide application in the autumn of 1982 failed to increase tiller number or dry matter accumulation in the spring of 1983. It is suggested that fungicide application in autumn 1981 prevented the death of tillers caused by fungal attack after snow covered the plots in December 1981.

Notes: A mathematical model of grass production and utilization on a dairy farm is described. Using the model an assessment has been made of the extent to which financial uncertainty arising from year-to-year variability in grass yields, coupled with a preference among farmers for minimizing risks, may explain the relatively low stocking rates and observed nitrogen usages on many dairy farms in England and Wales. The degree of risk has been equated with the probability of profits in a particular year being less than those required to cover the consumption needs and short-term borrowing requirements of the farmer. The results of the analysis indicate that a strategy of minimizing risks may lead to a significantly lower stocking rate than one of maximizing profits. Thus, considerations of risk may lead to stocking rates which are suboptimal from the viewpoint of economic and biological efficiency. However, there is no evidence to suggest that the comparatively low average usage of nitrogen on dairy farms in the UK is determined by risk considerations. On the contrary, increasing nitrogen usage lowers the apparent financial risk at a given stocking rate.

Notes: The herbage dry matter (DM) harvested, N response and herbage quality of monocultures and mixtures of Holcus lanatus (local ecotype) and Lolium perenne (cv. S24) were compared in a field experiment. The species were seeded in the proportions 100:0, 75:25, 50:50, 25:75 and 0: 100; three rates of N were applied (150, 300 and 450 kg ha−1) and five or six cuts were taken in each of four harvest years. H. lanatus established poorly in the seeding year and in the first full harvest year L. perenne contributed 1, 54, 76, 89 and 99% of the total herbage DM harvested from the five sward types (mean of N treatments). Only a small decline in the L. perenne content of the swards occurred during the experiment.Total amounts of herbage harvested were significantly lower for H. lanatus monoculture than for the other sward types in three of the four harvest years However, L perenne monoculture yielded significantly more total DM than mixtures with H lanatus in only one of the four harvest years. The five sward types did not differ overall in response to N fertilization. Nitrogen, phosphate and potassium content of the herbage was greatest for H. lanatus monoculture and decreased with increasing proportions ofL perenne present in the mixtures.Thus the presence of a high proportion of H. lanatus in mixture with L. perenne cv. S24 had

Notes: Two experimental populations of Italian ryegrass selected for high and low magnesium content were established as swards, harvested fresh daily and offered to sheep. Chemical analysis of the crops showed a 44% higher Mg and a 22% higher Ca concentration in the high-Mg selection but no significant differences between the selections for the other minerals.Blood and rumen contents were sampled throughout the feeding period and voluntary intake, digestibility and mineral balances for Na, K, Mg, Ca and P were determined. Magnesium intake, apparent availability and retention were significantly greater with the high-Mg selection. There were no significant differences in the plasma-mineral concentrations and all were within the normal range. Rumen fluid from animals fed the high-Mg selection had a higher proportion of soluble Mg and a higher Na:K ratio in the supernatant liquor.

Notes: A perennial ryegrass sward was managed by continuous stocking with sheep (April–September) for 4 successive years after sowing. The sward was grazed to maintain a leaf (lamina) area index (LAI) close to 1.0. Areas of the sward were released from grazing on three occasions: once during summer in the third year after sowing, and twice during spring and summer in the fourth year after sowing. There were marked changes in the structure and physiology of the continuously stocked sward following release from grazing. After several successive years of continuous stocking, the sward comprised a large population of small tillers and the small LAI resulted in consistently low rates of photosynthesis. Following release from grazing, photosynthesis increased markedly as the LAI increased but this change was associated with the loss of a large proportion of the population of tillers. There were seasonal differences in the pattern of changes in photosynthesis and tiller numbers following release from grazing which were not apparent under continuous stocking. The changes in the structure and physiology of the sward following release from grazing suggest that the net accumulation of herbage in areas of sward from which the animals are excluded, for instance using cages, may be an unreliable estimate of production under continuous stocking.

Notes: A randomized block experiment was conducted to compare unwilted and wilted grass silages and the effects of the feed additive monensin sodium on the silage intake and performance of finishing beef cattle. Two regrowths from a predominantly perennial ryegrass (cv. S24) sward were ensiled either without wilting or after field wilting for 3 d (dry matter (DM) concentrations 161 and 266 g kg−1 respectively). Both silages were treated with formic acid (2·6 and 30 litre t−1 respectively) and were well preserved. The silages were offered ad libitum to forty-eight Charolais-cross cattle (thirty-two steers and sixteen heifers, mean initial live weight 351 kg) for 145 d. All animals received 2·2 kg concentrates per head daily and half of those on each silage treatment received in addition 200 mg monensin sodium per head daily. Silage DM intake was 5.04, 504. 5·48, 5·63 ± 0.134 kg d−l; fasted liveweight gain was 0·69, 0·77. 0·64 and 0·73 ± 0.033 kg d−l and carcass gain was 0·47, 0·50, 0·40 and 0·45 ± 0·020 kg d−1 for the unwilted silage without and with monensin and the wilted silage without and with monensin respectively. It is concluded that wilting grass of low DM concentration for 3 d prior to ensiling reduced the performance of finishing beef cattle below that obtained from well-preserved unwilted silage in spite of a higher DM intake being achieved with the wilted silage. The inclusion of monensin sodium in a silage-based diet increased performance without significantly affecting feed intake.

Notes: Formic acid, sodium diacetate, propionic acid and two of its ammonium salts, ammonium isobutyrate and tributyl phosphate, were examined for their antimicrobial effects with cultures of micro-organisms typical of the microflora of moist hay. The assays, which were conducted at pH 5, 6, 7 and 8, revealed that all potential preservatives discriminated against actinomycetes in particular and were least effective against fungi. In addition, it was apparent that antimicrobial properties were enhanced under slightly acid conditions and for this reason the use of mixtures of organic acids and salts in commercial hay preservatives is justified. Tributyl phosphate not only had the strongest antimicrobial properties of the compounds tested but was also the least affected by pH.

Notes: Yields of grass were measured on three dairy farms in 1981. Farm 1, on free-draining loamy soil over chalk, produced 10.3 t DM ha−1 using 336 kg N ha−1. Farm 2, on poorly-drained loam over Oxford clay, produced 12.8 t DM ha−1 using 354 kg N ha−1. Farm 3, on loam over clay with some free-draining sandy loam, produced 9.5 t DM ha−1 with 169 kg N ha−1. Allowing for differences in soil nitrogen and summer rainfall these yields were quite close to expectations from experimental results, despite considerable poaching damage to some swards.Animal production records were kept and all livestock were weighed every 6 months, so as to calculate the utilized metabolizable energy (UME) output from grassland on each farm. These were 73, 62 and 59 GJ ha−1 on farms 1, 2 and 3 respectively.The UME output, expressed as a proportion of the yield of grass (converted to ME), was taken as the ‘efficiency of utilization’ of grass. This was 70% on farm 1,44% on farm 2 and 58% on farm 3. Results are presented separately for the summer and winter 6 months. Utilization by grazing appeared to be markedly reduced under very wet soil conditions. Major losses occurred in the conservation of grass.The results begin to establish a valuable base of data from commercial farms.

Notes: The establishment and growth of white clover (cv. Grasslands Huia) was investigated in two experiments with seeds sown in slots cut in old permanent pasture in early May 1982. In the first experiment, fully protected from pests and diseases, either 50 (low), 100 (medium) or 200 (high) seeds were sown evenly per metre length of slot; 100 seeds were also sown in a clumped pattern unevenly. In the second experiment, the effects of various combinations of molluscicide, insecticide and fungicide were investigated.Nine weeks after sowing there were 27, 37, 34 and 62 seedlings m−1 on the low, medium even, medium uneven and high seed-rate treatments, representing percentage establishments of 54, 37, 34 and 31 respectively. By September, however, there were no significant differences between the dry weights m−l of any of the treatments. In the second experiment, damage by slugs occurred early in treatments without molluscicide and caused a mean reduction of 70% in seedling establishment. As a result significantly smaller dry weights m−1 of clover were recorded in September in these than in other treatments. Failure to protect against insects resulted in smaller seedlings.The results indicated that variations in seed rate within the range investigated were not critical but that failure to protect against pests decreased percentage establishment and subsequent growth.

Notes: Masham ewes were stocked at 12(L), 16(M) or 20 (H) ewes ha−1, with 1·8 lambs per ewe on average, on two blocks (A and B) in both 1976 and 1977. Silage was offered at pasture to the ewes in early lactation and as stocking rate increased from 12 to 16 and then 20 ewes ha−1, 47, 73 and 100 kg DM per ewe was consumed respectively. Silage was cut mainly in the autumn and 179, 100 and 9 kg per ewe was made for L, M and H respectively. More variation between treatments was measured in net herbage accumulation in summer than spring and 11·4, 10·0 and 9·7t DM ha −1 was grown over the whole season for L, M and H respectively as a mean of both years.High intakes of herbage OM were measured with values up to 40 g per kg live weight for BL lambs in 1976. Rapid lamb growth was achieved. There were effects of stocking rate on lamb performance and for L, M and H the respective growth rates to 4 weeks were 266, 248 and 247 g d−1; growth rates to sale were 274, 263 and 252 g d−1; days to sale were 124, 126 and 129; mean carcass weights were 17·7, 17·1 and 16·7 kg. Most ewes lost weight in early lactation, especially AH and BH in 1976, but there were no stocking rate effects on ewe weight change in 1977.

Notes: Measurements of rates of growth and senescence of leaf lamina per tiller and of changes in tiller population densities were made in three experiments designed to investigate the influence of sward slate on leaf turnover and net production under continuous stocking.In each experiment initially uniform swards were fenced to provide four plots on which animal numbers were adjusted twice weekly to give a series of swards maintained as nearly as possible in a steady state with respect to sward surface height (range 1.1–6.4 cm) and herbage mass (range 440–2690 kg OM ha−1). Two experiments were carried out in July–September on vegetative swards and one in May–June on a reproductive sward. Measurements were begun 3–7 weeks after treatments were started and were repeated weekly during 3–4 week measurement periods.In all three experiments the rate of lamina growth per tiller increased linearly with an increase in sward surface height and herbage mass. In the two experiments conducted in July–August this relationship was partially offset by a linear increase in the rate of senescence per tiller but net production per tiller also increased linearly in relation to sward height and mass. In the experiment conducted in May–June the rates of growth and senescence per tiller increased in parallel so that net production per tiller showed no relationship with sward condition.Tiller population densities in the July–August experiments were highest in swards maintained between 2 and 3 cm surface height and declined in swards maintained above and below this height. In the experiment in May–June tiller numbers were similar in all treatments prior to the summer solstice but diverged in a manner similar to the other experiments later in the year.The rate of lamina growth per unit area increased in a manner consistent with an asymptotic relationship and the rate of senescence increased linearly with increasing sward height and mass in all three experiments. Net production per unit area was reduced on swards below about 2.5 cm in height but was insensitive to variation in sward surface height between 2.5 and 6.0 cm (approximately 1000–2500 kg OM ha−1 herbage mass).The effectiveness of adjustments in tiller numbers and production per tiller and of changes in the balance between growth and senescence as mechanisms of sward homeostatis, together with their implications in the scope for manipulating herbage production by grazing management are discussed briefly.

Notes: There was a three-fold variation in epicuticular wax content between populations of white clover and a five-fold variation between populations of red clover. The mean for all populations of red clovers was twice that of white. There was also a two- to three-fold variation within each population.The epicuticular wax content of white and red clovers grown in the field showed an increase with age. Thin layer chromatography showed no qualitative differences in the epicuticular waxes extracted either between populations or with sampling date.The higher levels of epicuticular wax found in some clovers were considered to be of possible significance as a source of antifoaming compounds in bloat-inducing pastures.

Notes: Differences in ease of digestion of bundle sheath cell walls of leaf blades of Panicum spp. are shown to be associated with differences in bundle sheath cell wall structure, including the presence or absence of a suberized lamella. These structural differences are correlated with photo-synthetic type (viz. C3, intermediate C3/C4, and the C4 types, PCK, NADP-ME and NAD-ME), as are mesophyll: bundle sheath area ratios.

Notes: The construction of a transducer and the use of a recording system to measure jaw movements of grazing sheep is described, The time spent grazing, ruminating and idling may be measured by this equipment and the data produced automatically processed using a microprocessor which determines each minute whether the animal is grazing, ruminating or idling and also summarizes these data to give total times. The accuracy of this technique in relation to manual observation is discussed.

Notes: The effect of two heights of cutting (5 and 8 cm) on the dry matter harvested and persistency of a range of different grass species and varieties were monitored over a 3-year period (1976 to 1978). Dry matter harvested was measured in the second and third harvest year and persistence of sown grass was assessed at the end of the first and third harvest year. The grasses were managed under a frequent cutting system, i.e. simulated grazing. Results showed that at the low cutting height tetraploid hybrid ryegrasses and diploid Italian ryegrasses in the third harvest year gave lower annual dry matter harvests of sown grass and were less persistent than at the high cutting height, whilst, in contrast, perennial ryegrasses gave higher annual harvests of sown grass but persistency was unaffected. In general at the low cutting height varieties of cocksfoot, meadow fescue and tall fescue gave comparatively higher annual harvests of sown grass in the second harvest year but lower yields of sown grass in the third harvest year. A notable exception was Cambria cocksfoot which in the second harvest year gave higher yields of sown grass and was more persistent at the low cutting height.However, at 5 cm cutting height, the proportions of weed grasses (mainly Poa spp.) in swards sown to tetraploid hybrid ryegrasses and diploid Italian ryegrasses were generally greater, particularly in mid-season.

Notes: Laminae of Lolium perenne and Phleum pratense were tested mechanically and the results analysed using engineering composite theory and fracture mechanics. As the lamina dries the fibres double in stiffness. The intervening cells show a sevenfold increase in stiffness below 20% water content (based on dry weight). Work to fracture across the veins is almost independent of water content, but below 50% water content the mode of fracture changes. It is predicted that total fibre content will not affect hay shatter; total protein content may affect it by altering the water-binding properties.

Notes: Grass silages made from first-harvest perennial ryegrass in mid-May and early-June and termed high-D and low-D respectively had mean DOMD values of 694 and 633 g kg−1 and were offered ad libitum to twelve Ayrshire cows in a 16-week feeding experiment using a cyclic changeover design. The silages were supplemented with a concentrate containing 363g CP per kg DM at daily rates of 1.5, 3.0 and 4.5 kg per cow. The average daily intakes of silage DM were 12.6 and 11.1 kg per cow on the high- and low-D silage treatments respectively, and were not significantly different on the three concentrate treatments. The mean daily milk yields were 171, 18.6 and 21.0 kg per cow on the low-D silage treatments, and 19.6, 21.2 and 22.8 kg per cow on the high-D silage treatments at the 1.5, 3.0 and 4.5 kg rates of concentrate feeding respectively. Fat and lactose concentrations in the milk were not affected significantly by treatment, whereas the CP and SNF concentrations increased progressively and significantly as the rate of supplementary feeding increased. It is concluded that a high-protein concentrate supplement allows silage to make the maximum contribution in the diet, and gives a high response in terms of milk yield and composition.

Notes: Grass which had grown without interruption from late April was cut as for hay in early July in each of 2 years. Fifteen dates of removal of the cut herbage were compared ranging, at 1-d intervals, from the day of cutting to 14 d after cutting. Sward regrowth was studied.A delay of 5 d in removing cut herbage reduced herbage dry matter harvested during the remainder of the growing season by 9% and a delay of 10 d reduced yield by 16%. Delaying the removal of cut herbage reduced the density of grass tillers considerably in the short term, but the effect had worn off in 8 to 12 weeks. The presence of cut herbage reduced the rate of extension of leaf blades considerably. Delaying the removal of cut herbage reduced the area per leaf blade and the weight per unit area and these effects persisted for several weeks. Grass growing under cut herbage had a less negative leaf water potential and a rather cooler but much darker environment in which to grow than uncovered grass.

Notes: Book reviews in this article: Milk from Grass Edited by C. Thomas and J. W. O. Young Nitrogen Fixation in Legumes Edited by J. M. Vincent Herbage Intake Handbook Edited by J. D. Leaver Man's Proper Study By Richard J. Colyer Introduction to Sheep Fanning By R. G. Johnston

Notes: Information about the grazing behaviour and the return of dung to pasture by lactating dairy cows was collected from stocking rate experiments in northern Victoria. Grazing behaviour was observed for 24-h periods in mid-summer in two years. Grazing time increased as herbage allowance decreased to about 32 kg DM cow−1 d−1 but as herbage allowance decreased further grazing time also decreased. A herbage allowance of 32 kg DM cow−1 d−1 corresponded to a stocking rate of about 5.5 cows ha−1. This effect of herbage allowance on grazing time may have been confounded by herbage mass, however. Rumination time of the cows increased by 003 h for each kg increase in herbage allowance while resting time was not affected by treatment.The effects of stocking rate on some of the characteristics of faecal output were measured for a 3-d period in mid-summer. The number of pats deposited per cow daily declined by 0.66 for each unit increase in stocking rate. The fresh weight of dung also declined as stocking rate increased by 0.16 kg per unit of stocking rate. While the values for the amounts of dung deposited on the pasture by the cows at the lower stocking rates are similar to many of those reported in the literature, this study has quantified the way in which stocking rate may influence this in one instance. Furthermore, provided that some measure of herbage intake is made when measurements of dung excretion are performed, it is suggested that estimates of in vivo digestibility can be obtained.

Notes: Two red clover (Trifolium pratense) cultivars, Red Head (tetraploid) and Kuhn (diploid), were sown at a seed rate of 13 kg ha−1 either alone or in mixture with Italian ryegrass (Lolium multiflorum) cv. RvP sown at seed rates of 10, 15, 20 or 30 kg ha−1. RvP was also sown alone at a seed rate of 30 kg ha−1 and received nil or 300 kg ha−1 fertilizer a−1 fertilizer N. All plots were established using the barley cultivar Midas sown at a seed rate of 100 kg ha−1 as a nurse crop.Neither clover cultivar nor ryegrass seed rate significantly influenced either dry matter harvested or botanical composition over the 3 harvest years. On average over all years the grass-clover mixtures produced 75% of the yield of the N-fertilized RvP, 125% of the clover monocultures and 225% of the unfertilized RvP. The red clover contribution to the total dry matter harvested of the mixtures averaged 45–60%. The dry matter concentrations of the mixtures were considerably higher than those of the pure clover stands. In the third year yields were markedly reduced in comparison with those in the first and second years.It was concluded that Italian ryegrass can be a suitable companion grass for red clover. Its superior yielding capacity over other grasses such as perennial ryegrass or timothy under a conservation management can be coupled to advantage with red clover to give a sward which Is essentially stable, at least over a 2- to 3-year cropping period, although giving slightly reduced yields in the third year. Italian ryegrass-red clover mixtures, without the use of fertilizer N, can produce high DM yields of good quality herbage.

Notes: Red clover cv. Hungaropoly was slot-seeded into a perennial ryegrass-dominant sward in April 1979. Glyphosate and paraquat were applied separately as bandsprays each at two doses and at two band widths. Control plots were either slot-seeded without a herbicide bandspray or received ±150 kg N ha−1 a−1. Red clover establishment was assessed and amounts of dry matter (DM) and total N accumulated were measured at two harvests in 1979 and three harvests in 1980. Bandspraying increased seedling vigour and development and resulted in the eventual replacement of 1 t grass DM ha−1 by an equivalent amount of red clover. Of the treatment variables investigated, bandspray width had the greatest influence on red clover establishment and productivity. The slot-seeded area, meaned for all treatments, produced a total of 6.40 and 13.16 t DM ha−1 in 1979 and 1980. This was estimated to be equivalent to the all-grass sward receiving 112 kg N ha−1 a−1 during the second year of the experiment or 238 kg N ha−1 over the 2 years when measured in terms of N yield. Slot-seeding overcomes several of the problems associated with conventional establishment of red clover.

Notes: Two experiments were conducted to determine the effect of pre-harvest treatment by formic acid spray on the moisture concentration (MC) of perennial ryegrass (Lolium perenne) in vegetative growth. In Experiment 1 aqueous solutions containing 0.080, 0.148 and 0.259 kg formic acid per litre were applied to a perennial ryegrass sward (fresh matter yield 21.0 t ha−1) at a constant volume of 41.2 litres t−1to give active ingredient application rates of 3.5, 6.0 and 11.5 kg t−1 respectively. In Experiment 2 similar solutions were applied to a perennial ryegrass sward (fresh matter yield of 6.9 t ha−1) at 35.2, 19.0 and 10.9 litres t−1 respectively to give a constant formic acid application of 2.8 kg t−1. The changes in MC and chemical composition after treatment with acid were compared with those in untreated plotsIn both experiments treatment with acid significantly reduced the MC (P 〈 0.001) within 5 h of treatment from 4.4 to 3.0 kg per kg of dry matter (DM) in Experiment 1 and from 3.0 to 2.6 kg per kg DM in Experiment 2. There was no effect of level of acid applied in Experiment 1 or of the volume of water applied with the low level of acid in Experiment 2 on the changes in MC of acid-treated plots. There was however a significant (P 〈 0.00l) difference between the MC of plots treated with acid and those untreated which was maintained in Experiment I for up to 9 d even during rainfall.There were no differences between the chemical compositions of samples from acid-treated plots in either Experiment I or 2 or between untreated and acid-treated plots in Experiment 2. However there were significant differences between untreated and acid-treated plots in Experiment 1 for the content of nitrogen (N), water-soluble carbohydrate (WSC) and digestible organic matter (DOM). There were changes in the content of N, WSC and DOM with time in both experiments. In Experiment I there was an interaction between sampling time and treatment for both N and WSC but not for DOM. Compared with untreated plots, treatment with acid at all levels accelerated the loss of WSC and delayed the fall in N content. There were no such interactions in Experiment 2.

Notes: Data are presented for the fourth (1979) and fifth (1980) harvest years of a trial in which four levels of N fertilizer (0, 30, 60 and 90 kg N ha−1) were applied each spring to swards with grass (perennial ryegrass cv. Barlenna) and each one of four cultivars of while clover (Blanca, Sabeda, Olwen and S100) or grass alone. Results from the first three years have been published.Dry matter (DM) harvested dropped from the fourth to fifth years over all swards by 1 −5 to 2 0 t ha−1 but response to N was maintained (17.4 and 24.4 kg DM (kgN)−l in response to 90 kg N ha−1 in 1979 and 1980 respectively) despite reductions in summer yields relative to unfertilized swards. Blanca swards produced significantly less DM harvested than all other cultivars in 1979 and all cultivars in 1980 except for Olwen. Clover DM harvested continued to fall from 1977, the mean for the unfertilized treatment in 1980 being 55% of that in 1979. Up to 1980 Sabeda swards produced more clover DM harvested than Blanca swards. Linear regressions between annual clover content at zero N and at each spring N level for each sward type over 5 harvest years were very highly significant. It is concluded that all cultivars used responded similarly to spring N. Further work to develop a method for predicting the effect of spring N on clover content of given swards is required.

Notes: Preliminary experiments were conducted to assess the winter hardiness of representative cultivars or populations of Festuca rubra, Holcus lanatus and Agrostis spp. in comparison with a hardy (Premo) and a susceptible (Grasslands Ruanui) cultivar of Lolium perenne. Two controlled-environment tests and a field experiment including two cutting and two fertilizer managements are reported.Grasslands Ruanui andH lanatus suffered most damage in all tests. In the controlled-environment experiments, Premo was at least as hardy as the F. rubra and Agrostis cultivars but in the field Premo was more susceptible. Spring growth of Premo was affected more by the cutting and fertilizer managements applied the previous summer and autumn than was spring growth of F. rubra or Agrostis spp.It was concluded that, whereas use off. F. rubra or Agrostis spp. should not be limited by fears of winter damage, poor winter hardiness may limit the value of H. lanatus for hill and upland reseeding.

Notes: Patterns of defoliation in perennial ryegrass-dominated swards were examined in two experiments. In Experiment I measurements were made on four swards after a single overnight grazing. In Experiment 2 observations were made repeatedly on marked tillers in two continuously stocked swards which were grazed down over a 24-d period. In both experiments the chance of defoliation was related to lamina height both within and between age classes. Lamina angle was measured in Experiment 2 and within age classes the chance of defoliation was also independent of lamina angle. In Experiment 1 tillers most frequently had both of the two youngest laminae grazed; in Experiment 2 tillers with only lamina 1 grazed occurred most frequently, but defoliation of the two youngest laminae was again common.The different frequencies of grazing of the different classes of laminae is interpreted as being a function of their frequency of occurrence within the grazed layer at the top of the sward. The pseudostem apparently restricted the depth of the grazed layer in the very short swards of Experiment 2.Grazing behaviour in relation to sward structure and the patterns of defoliation and their implications for herbage production are discussed.

Notes: Perennial ryegrass and white clover were grown in boxes with either no competition, root competition only, shoot competition only, or both root and shoot competition between them, Boxes received either no nitrogen or 200 kg N ha−1 as a split application. The experiment was harvested at 8, 12, 16 and 20 weeks after sowing. Perennial ryegrass was more competitive than white clover throughout the experiment and its overall competitive ability increased with Lime. The effects of root competition were greater than those of shoot competition at the first harvest, with both forms of competition having similar affects at the second harvest, while at the third and final harvests shoot competition had greater effects than root competition. At the later harvests N application increased the overall competitive ability of perennial ryegrass, relative to white clover, mainly owing to the increase in its shoot competitive ability.

Notes: Inflorescence-bearing tillers of Italian ryegrass (Lolium multiflorum cv. RvP) were dried in controlled temperature, relative humidity and air speed conditions chosen to be within the range likely to occur in a hay swath drying in the field. After drying for increasing intervals the tillers were separated into inflorescence, exposed flowering stem, leaf-laminae, leaf-sheath and flowering stem enclosed by leaf-sheath and the drying rates of these components in addition to that of whole tillers were measured. There were large differences in drying rate between components, but the differences were similar over the range of drying conditions employed. The drying rate of the inflorescence as well as that of stem enclosed by leaf-sheath was found to be very low and that of leaf-laminae and leaf-sheath very high.

Notes: Book reviews in this article: Genetic Resources of Forage Plants Edited by J. G. McIvor and R. A. Bray Nutritional Limits to Animal Production from Pastures Edited by J. B. Hacker Herbivory. The Dynamics of Animal-Plant Interactions By Michael J. Crawley

Notes: Forty hill farms in Northern Ireland were surveyed to obtain information on stocking rates and output and to identify factors affecting output. The mean farm area of 194 ha was composed of 155 ha hill land and 39 ha inbye land with a mean stocking rate of 0–7 cow-equivalents ha−1.On the average farm most of the grassland (88%) was over 20 years old. The preferred species content of the swards averaged 22% over the whole farm and 42% in cut swards. This latter result may be attributed to the relatively high level of fertilizer N applied to these swards (131 kg ha−1) compared to the mean application rate over the whole farm of only 29 kg N ha−1 and 93 kg ha−1 applied to the inbye land.Output data calculated as utilized metabolizable energy (UME) per unit area for the whole farm and separately for hill and inbye components showed that although only 21% of the farm area was inbye land almost half the annual total metabolizable energy requirements of stock had to be met by grazing inbye. The capacity of herbage to meet stock energy demands was much lower on the hill with supplementary feed having to be provided over the winter.Calculated output from inbye land was 39 GJ ha−1 compared to 7 GJ ha−1 from the hill and 15 GJ ha−1 from the whole farm. UME output of the whole farm was negatively correlated with farm size (r=−0 55) and positively correlated with stocking rate (r = 0 89), applied N level (r = 0 63)

Notes: Forage-harvested crops of perennial ryegrass (200 g DM kg−1) and red clover (163 g DM kg−1) were inoculated with cultures of Streptococcus durans str. 1024, Lactobacillus acidophilus sir. 2356 and Lactobacillus plantarum str. 6, and 1:1 mixtures of the Streptococcus and each of the lactobacilli, at rates of 106 and 106 (g fresh weight)−1 and ensiled in laboratory silos for up to 128 d. The possible influence of these bacteria on fermentation was examined in relation to formic acid at rates of application of 0 and 2.3 g(kg fresh weight)−1, None of the cultures or culture mixtures, either at the low or high rate of application, had any notable influence on microbial development, the rate of acidification or promotion of the homolactic fermentation; they tended to exacerbate loss of insoluble nitrogen and deamination. All silages were well-preserved with no butyric acid being detected, in spite of pH increase during storage. Only formic acid had any consistent and desirable effect on preservation.

Notes: Book reviews in this article: Grass: its Production and Utilization Edited by W. Holmes Fream's Agriculture Edited by C. R. W. Spedding

Notes: The results or estimations of in vitro digestibility of three standard herbage samples, analysed weekly over about 48 weeks per year, have been examined using time-series methods of Box and Jenkins to separate seasonal and non-seasonal trends. The study includes data from March 1971 to February 1981. The dominant feature of variability is the seasonal component with a cycle of 52 weeks; no other significant cycles of longer periods were detected. The model of Box and Jenkins (1976) was found to be the most appropriate in describing the three series. A decrease in the in vitro digestibility of the low standard, which is 10 years old, occurred over the last year or so, although the reasons for this apparent storage effect are unknown.

Notes: Salts of formic, acetic and propionic acids and glutaraldehyde were subjected to an assay with representatives of the major groups of silage micro-organisms at pH 4·0, 4·5, 5·0 and 6·0. The antimicrobial spectra of the salts suggested that they are most effective against the least desirable components of the bacterial flora of silage, coliforms, Clostridia and bacilli, and would create conditions for a desirable fermentation in practice. In many instances the activity of the salts was better than the corresponding free acid and it was postulated that, if supplemented with acid, aerobic stability of silage might also be improved. Glutaraldehyde, whilst being less selective as an antimicrobial agent, possesses properties which would encourage a lactic-acid fermentation in silage.

Notes: Previous results with red clover (Trifolium pratense) leaflets, leaf petioles and stems indicated that the epidermis continued to impose a barrier to water loss at low water contents. Further experiments with red clover leaf petioles and stems suggest that cuticular wax is the epidermal component concerned and that treatments which remove or merely modify this will also increase drying rate. However, when treatments to reduce epidermal resistance were applied to pseudostems and flowering stems of Italian ryegrass (Lolium multiflorum) the initial increase in drying rate produced was not sustained at low water contents. The pseudostems consisted of three layers of leaf material rolled one within the other and the true flowering stems were surrounded by one leaf sheath. Their pattern of response to treatments suggested that these affected only the outer sheath, which produced an initial acceleration in drying rate, but that this was not sustained after the outer sheath was dry because the lower untreated layers dried in a similar way to untreated controls.The results suggest the possibility of devising practical treatments to increase drying rate at low water contents with crops containing a high proportion of leaf or stem not surrounded by leaf sheath. Devising such treatments for crops with a high proportion of grass pseudostem or flowering stem surrounded by leaf sheath will be considerably more difficult.

Notes: Awned seeds of four grasses were submitted to humidity changes on dry and damp soil to see how far the awns assisted in distribution and seed burial. The results indicate that awns on their own are probably of limited importance. In another study, seeds of seven species were soaked in water with high or low oxygen levels to determine whether dormancy could be induced. Waterlogging delayed germination briefly but did not induce dormancy except in Avena fatua. Oxygen level proved unimportant. The effect of temperature on germination and dormancy was examined in seeds of Agrostis capillars. Seeds of this species germinated best at fluctuating temperatures with a minimum difference of 10°C between day and night temperatures. Constant or slightly fluctuating temperatures over 20°C or under 10°C induced dormancy in a small percentage of seeds.

Notes: Changes in the population of stem-boring Dipterous larvae (including Oscinella frit) were examined weekly for a year on plots of perennial ryegrass (Lolium perenne cv. S24). The herbage net regrowth rate of these plots and of plots receiving the pesticide, phorate, was also monitored. Herbage net regrowth rates were greatest in April and October. At the same times the differences in net regrowth between treated and untreated plots were greatest, thus indicating the periods of the greatest loss of production due to pest damage to the sward.Populations of stem-boring larvae fell below 100 m−2 for only 4 weeks in the year and were maximal in autumn and early winter. Larval mortality exceeded 50% during the winter period. A correlation was found between larval activity and sward damage.

Notes: Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha−1) and high (22 sheep ha−1) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward 〈 lightly grazed sward 〈 heavily grazed sward 〈 repeatedly cut sward.When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.

Notes: A preliminary field experiment on a sward cut six times during 1978 indicated that total grass herbage harvested was negatively related to the number of Rumex obtusifolius plants present; densities of five to ten plants m−2 resulted in reductions of up to 30%, although the total quantity of herbage harvested (including Rumex) may have remained constant. However, R. obtusifolius densities did not significantly affect grass yield during the first half of the growing season. A negative relationship between R. obtusifolius ground cover and grass yield, particularly from late July onwards, was shown in a second field experiment during 1979 and 1980 using a 3-cut management. It was estimated that to produce a 20% reduction in grass growth in July-August and October, R. obtusifolius ground covers of 30% and 20% respectively were needed. Sixty per cent of the total herbage harvested in 1980 consisted of R. obtusifolius. These results indicate the damage that can be caused by this species and suggest that control measures should be taken before early summer.

Notes: A total of 104 cultivars of six agricultural grass species were compared under controlled conditions for their relative susceptibility to the common soil-borne fungus Fusarium culmorum (W.G.Sm.) Sacc. A number of the perennial ryegrass (Lolium perenne) cultivars showed a high degree of resistance, their emergence being little reduced under the stringent test conditions. The emergence of others was, however, severely affected. Italian ryegrass (L. multiflorum) cultivars generally had a high level of resistance, the outstanding exception being cv. Leda Daehnfeldt which proved very sensitive. None of the cultivars of cocksfoot (Dactylis glomerata), timothy (Phleum pratense), meadow fescue (Festuca pratensis) or tall fescue (F. arundinaceae) which were tested had more than a moderate level of resistance and most were highly susceptible.The vigour of ryegrass seedlings was, in most cases, closely correlated with percentage emergence but was less closely correlated in the more susceptible species. These often had a very low emergence but those seedlings which did establish were frequently of a size similar to that of healthy plants.Treatment of grass seed with a fungicide mixture of benomyl (15% a.i.) plus captan (60% a.i.) provided excellent protection against F. culmorum on even the most susceptible cultivars.Drazoxolon (40% a.i.) was, however, much less effective.

Notes: In field trials in 1979–81, perennial ryegrass cultivars Royal and Majestic (amenity) and Morenne (agricultural) produced maximum seed yields at levels of applied N ranging from 40 kg ha−1 to 160 kg ha−1. Available soil N levels were estimated at 55 kg ha−1; hence maximum seed yields were obtained at total nitrogen levels of 95–135 kg ha−1 in Royal, 95–215 kg ha−1 in Morenne, and 175–215 kg ha−1 in Majestic.Applied N at rates above 120 kg ha−1 either reduced or did not significantly increase seed yield, decreased seed numbers per unit area and decreased spikelets per tiller and seeds per spikelet.The use of a growth regulator increased seed yields by preventing lodging, but did so irrespective of nitrogen application rate. It is suggested that failure to increase seed yield at high N rates is a result not of poor pollination because of lodging, but seed abortion as a consequence of competition for assimilate supply by secondary vegetative tillers.

Notes: Swards were developed from plants of contrasting populations of Dactylis glomerata L. selected for high and low dry-matter (DM) production in monocultures and mixed cultures grown during 1970 and 1971. DM production of these mono-genotype swards was assessed during 1974. Those swards developed from the divergent selections from monocultures maintained a significant difference between the high and low selections, whereas those developed from selections out of mixed cultures showed no significant difference in yield. These results are consistent with earlier reports that there is no positive relationship between competitive ability in mixed cultures and vigour in pure stands. The implications of the physiological basis of these responses in terms of grass breeding are discussed.

Notes: White clover swards were grown in a freely ventilated glasshouse during August and September 1979. The uninterrupted growth of these swards, which were grown in Perlite with either nitrate or biologically fixed nitrogen, was studied from 5 weeks after sowing until peak dry weights of the live sward components occurred. The dry weight of leaf laminae, petioles, stolons, roots, nodules and dead material was determined weekly together with live leaf numbers and leaf lamina area. The pattern of dry matter accumulation of the yield components of the swards (live leaf laminae plus petioles) was characterized by an initial period of more or less exponential growth up to 6 weeks from sowing dominated by leaf lamina development. This was followed by a linear phase, when petiole weight increased rapidly and a final slowing down period terminated by peak yields of live DM 12 weeks after sowing.The weight of stolons, roots and nodules increased slowly during the period dominated by leaf lamina growth compared with the rapid increase thereafter, which resulted in a two- to three-fold increase in the weight of stolons, roots and nodules by the time the weights peaked. It is assumed that after the leaf area index of the sward had reached a plateau there was a limit to the assimilate supply to the sward components at a stage when they were all apparently competing for photosynthates. There was a progressive slowing down in the rate of dry matter accumu-

Notes: The appearance of successive leaves on labelled tillers of S24 perennial ryegrass, and their subsequent death, were monitored for 1 year on cut plots on which four levels of N fertilizer were applied.A new leaf appeared about once every 15 d from the end of April until the beginning of September, and about once every 52 d from the end of October to the middle of February. Rate of appearance changed rapidly during September and October, and again from mid-February until the end of April. The response of leaf appearance rate to temperature changed early in the year; the effect of overwintering is discussed.Leaf lifespan was 50–60 d in summer and 120–140 d in autumn and early winter, and was influenced by conditions either during expansion of the leaf or shortly afterwards.The results are discussed in relation to the construction of a model to describe changes in the herbage available to animals grazing a grass sward.

Notes: A mathematical model which simulates the potential production and utilization of herbage on perennial ryegrass and grass-white clover swards in the context of an 18-month beef enterprise is described. Using the model the potential costs and benefits of replacing grass by grass-clover mixtures have been determined. Potentially grass-clover swards receiving no fertilizer N are shown to have annual DM yields equivalent to ryegrass swards receiving at least 300 kg N ha −1. Apart from the benefits of reduced fertilizer N usage there appear to be indirect benefits related to a lower concentrate requirement per head. In consequence, from an economic viewpoint grass-clover swards potentially compare very favourably with heavily fertilized grass swards, even if account is taken of the opportunity costs of lower stocking rates associated with grass-clover systems. However, this conclusion awaits experimental confirmation.

Notes: Grass silage made in late May from S24 perennial ryegrass was offered ad libitum to eight Ayshire cows in a 16-week feeding experiment. The silage had a DM concentration of 244 g kg−1, contained 163 g crude protein (kg DM)−1 with a ruminal degradability of 0.77 and had an in vitro DOMD concentration of 678 g kg−1. In addition, four concentrates each containing 167 g soya-bean meal kg−1 were consumed at a mean daily rate of 6.43 kg DM per cow. The soya-bean meal was either untreated, or ‘protected’ by formalin and mixed in the following proportions, 100:0; 66:34; 34:66; and 0:100 respectively, in the four concentrates. The daily intakes of silage DM were not significantly different on the four treatments and averaged 90 kg DM per cow, giving a mean total daily DM intake of 32.4 g kg−1 live weight. The milk yields were not significantly different on the four treatments and averaged 23.9 kg −1 The treatments had small and non-significant effects on milk composition and live weight. It is concluded that with a high-digestibility, well-preserved grass silage of satisfactory protein content the inclusion of ‘protected’ protein in the supplementary concentrate had no beneficial effects on milk production.

Notes: Earlier studies have concluded that for a dairy enterprise the most profitable conservation strategy, where cutting and grazing areas are separate, is generally associated with relatively low-digestibility silage. The aim of this study has been to investigate whether this conclusion remains true where cutting and grazing are integrated. A mathematical model of grass production and utilization by a dairy enterprise has been developed from information collected from a variety of experimental sources. It was found that cutting strategies aimed at making low-digestibility silage could not automatically be regarded as the most profitable. In good grass-growing conditions strategies resulting in low-digestibility silage did appear to offer the highest gross margins per unit area, but under average and poor growth conditions the greatest profits were associated with high-digestibility silage strategies. In particular, it was noted that the timing and frequency of conservation cuts influenced the efficiency with which grass could be utilized on the grazing areas and this fact rather than yield of conserved material influenced the comparative biological and economic efficiency of the different cutting strategies.

Notes: A study was conducted to determine the influence of herbage mass under continuous stocking management with sheep on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in a mixed-species sward of Lolium perenne. Poa annua and Trifolium repens. Plots were maintained in as nearly constant herbage mass as possible at 500, 700, 1000 and 1700 kg organic matter (OM) ha−1 by continuous but variable stocking with sheep for the period May to July inclusive. Estimates of G, S and NP were obtained for each species from repeated measurements over two 2-week periods on individually identified grass tillers or clover stolons. Estimates of species population densities enabled estimates of G. S and NP per unit area to be made.Rates of G were higher and of S were lower for Lolium tillers and Trifolium stolons than for Poa tillers. The laminae of Poa tillers occupied inferior positions in the canopy of the swards maintained at higher herbage masses and were less accessible to the grazing animals, so that Poa tissue was less well utilized.The total population densities of tillers and stolons declined at herbage masses both above and below 700 kg OM ha−1. The rate of G for the three species combined increased with herbage

Notes: The most likely reasons for seeding failures of sod-seeded lucerne (Medicago sativa L.) were determined from eight field experiments, where the effects of a range of pesticides on the establishment of and damage to lucerne were compared. The experiments were conducted during 1979–80 at four sites near Ithaca, N.Y., USA. Pesticides evaluated were: methyl bromide (MeBr), carbofuran, acylalanine (1979, 1980); fonofos, diazinon, methiocarb, metaldehyde and combinations of carbofuran with methiocarb or metaldehyde (1980). Seeded areas were pretreated with glyphosate for vegetation control.In 1979 seedling losses were decreased by carbofuran and methyl bromide whereas in 1980 seedling losses were decreased by methyl bromide at two sites and by methiocarb at all sites. Methiocarb was the only pesticide to reduce defoliation damage. The effect of combining methiocarb and metaldehyde each with carbofuran tended to be additive in terms of reducing seedling numbers and damage, and increasing yield. Treatment effects on lucerne yield were still apparent up to 12 months after seeding. Based on the consistent results of methiocarb, it was concluded that molluscs were probably the main factor limiting the establishment of sod-seeded lucerne but an arthropod component also appeared to be involved.

Notes: Book Reviews in this article: Silage in Milk Production By A. J. Corrall, H. D. St C. Neal and J. M. Wilkinson Sward Measurement Handbook Edited by J. Hodgson, R. D. Baker, Alison Davies, A. S. Laidlaw and J. D. Leaver Managing Agricultural Systems By G. E. Dalton

Notes: Faeces of cattle, pronghorn and rabbit were collected from the Central Plains Experimental Range, Colorado, 6, 18 and 30 months after being excreted. The faecal material was placed in moistened potting soil and seedling emergence observed. Bouteloua gracilis (blue grama grass) and Sporobolus cryptandrus (sand drop seed), emerged in abundance from cattle dung. Emergence of B. gracilis decreased with increasing period of exposure before collection, but emergence of S. cryptandrus increased with exposure. Forbs dominated the seedling populations in pronghorn dung, and few seedling populations of any species emerged from rabbit dung. The results are discussed in relation to the effect of large herbivores on the establishment of species from seed in rangeland.

Notes: The effect of temperature on the nutritive value of Macroptilium atropurpureum cv. Siratro and line 10–105 was determined in a controlled environment at day/night temperatures of 24/18, 30/24 and 36/30°C, and a 13-h daylength. Estimated dry matter digestibility, cell wall and lignin concentrations of leaf, stem and stubble were measured in two 5-week regrowth cuts.Siratro and line 10–105 were similar in digestibility and responded similarly to temperature with whole tops, stem and stubble decreasing by an average of 0.0023, 0.0026 and 0.0056 units of digestibility for each 1°C rise in growth temperature. These changes reflect the increase in cell wall and lignin concentrations with higher temperature.The digestibility of young, just-expanded leaves was increased by increased growth temperature. This effect may be explained by the faster rate of leaf expansion at high temperature which reduced cell wall and lignin concentrations at this stage of leaf development. However, for the bulk leaf fraction digestibility was unaffected by temperature. Leafiness declined at high temperatures.

Notes: Small plots of permanent pasture containing 50%Agrostis capillaris-Festuca rubra were defoliated frequently or infrequently during the growing seasons in 1978–80 and fertilized with 133, 125 and 125 kg N, P2O5 and K2O respectively ha−1 a −1 or not fertilized during 1978–81. Yields of herbage were assessed during 1979–80 and botanical composition assessed on three occasions in 1978, once in April 1979 and once in 1982.Fertilizer application increased annual dry matter (DM) harvested from frequently cut plots by 40%, 53% and 65% in 1978, 1979 and 1980 respectively, and on infrequently cut plots it doubled hay DM harvested in 1978 and increased total DM harvested by 126% and 186% in 1979 and 1980 respectively. Infrequent cutting and fertilizer both decreased the proportion of A. capillaris but only fertilizer decreased the proportion of F. rubra. Fertilizer greatly increased the proportion of Holcus lanatus especially where plots had been cut frequently and of Alopecurus pratensis where they had been cut infrequently. It decreased the proportion of Luzula campestris. Infrequent cutting, especially with fertilizer, discouraged Cerastium fontanum ssp. glabrescens and Trifolium repens but encouraged Rumex acetosa.

Notes: Data on sheep live weights, herbage standing crop and herbage quality are presented from a 3-year grazing management study comparing set stocking, four-paddock rotational grazing and four-paddock forward rotational grazing systems. In both rotational grazing and forward rotational grazing systems, sheep were sequentially moved from one paddock to the next every 5 days. Lambs were weaned at 12–15 weeks of age in the forward rotationally grazed system and from then on grazed one paddock ahead of the ewes. Herbage mass was consistently greater on the rotationally grazed system than on the set-stocked system, averaging 38.4% more herbage in 1978. 32.8% more in 1979 and 52.7% more in 1980. No differences were observed in ewe live weights at the end of the grazing year between the rotationally grazed and the set stocked systems. Live weights of rotationally grazed lambs were superior to set-stocked lambs only when herbage allowance was low; otherwise no difference existed between these two systems. The forward rotationally grazed lambs generally had lower liveweights than did the rotationally grazed lambs, perhaps due to weaning stress.

Notes: Comparisons between some aspects of grazing and ruminating behaviour by sheep recorded automatically and by manual observation are reported.Behaviour was recorded manually and automatically on four swards with surface heights of 3, 6, 9 and 12 cm. The automatic recording method gave significantly higher biting rates (78 v 67 ± 2·4 bites min−1) than manual recording. Possible reasons for these differences are discussed. For recordings during ruminating there was no significant difference between the two recording methods.As sward surface height increased biting rate decreased by 4·6 (± 0.65) and 2·9 (± 0·87) bites min−1 cm−1, and masticating rate increased by 4·5 (± 1·13) and 3·7 (± 0·95) chews min−1 cm−1 for the manual and automatic recording methods respectively.It is concluded that the automatic recording system gives higher rates for jaw movements during grazing than manual recording but there is little difference between the two methods for activities recorded during ruminating.

Notes: An experiment was carried out in which simulated swards of ryegrass (cv. S23) were grown in boxes. In the first instance the swards were cut at weekly intervals to maintain five levels of leaf area index (LAI) from LAI 1 to 4–5 in simulation of continuous grazing. Measurements were made of growth, senescence and net growth rate and of net canopy photosynthesis at constant irradiance. The results showed that the swards adapted to the defoliation regimes mainly by changes in tiller population density and pseudostem length. When the swards had equilibrated to the cutting regime growth rate increased with LAI but, since tiller density and the partitioning of growth between herbage harvested and that lost by sensecence also changed with LAI, net growth rate was constant over the LAI range 2–4·5. Maximum weight of herbage harvested was obtained between LAI 2 and 3.After 10 weeks of weekly cutting all the swards were cut back to LAI 1 and allowed to regrow. Growth rate showed almost no response to the previous culling treatments. The relationship of net canopy photosynthesis to LAI was linear for the frequently defoliated swards and curvilinear for regrowing swards. The reasons for this difference were examined.

Notes: White clover seedlings were defoliated when either two, four or six leaves had opened. There were four defoliation treatments, involving removal of younger laminae, older laminae, all laminae or all laminae and petioles, plus a control (no defoliation).Leaf removal reduced area of subsequently emerging leaves when measured as they became fully opened, though some compensatory expansion occurred after this. Petiole length was also reduced considerably. The magnitude of these effects varied according to severity of the treatment: removal or older laminae had little effect, removal of younger laminae had a similar effect to removing all laminae, but the effect was considerably increased by removing petioles in addition to laminae. The youngest plants showed the greatest reductions in leaf size following leaf removal. Defoliation had little effect on the rate of development of subsequent leaves.The most severe defoliation treatments reduced plant dry matter but younger seedlings appeared to have a capacity for recovery equal to or greater than that of older seedlings.It is concluded that white clover seedlings have a considerable ability to recover from leaf removal, especially if only laminae of old leaves are removed, but the growth reduction following removal of petioles as well as laminae appears to be especially severe.

Notes: Experiments were carried out over three seasons in which swedes were sown by precision drilling after ploughing and conventional cultivation or by direct drilling with two types of drill at two row widths with and without shallow cultivation. The swedes were sown in June or early July following a first silage cut and sward desiccation with paraquat. Precision drilling with inter-row cultivation and singling gave a uniform weed-free stand and had the highest mean yield of over 8 t DM ha−1 Where hoeing was omitted yields were reduced due to weed competition. Yields following direct drilling were lower than with precision drilling due to uneven plant establishment, weed competition and grass regrowth. A shallow rotavation prior to direct drilling improved uniformity and yields and reduced grass regrowth. Direct drilling produced high numbers of small roots. The highest root yields were associated with sowing in early June and harvesting in December or January. Overall yields from the silage cut plus direct drilled swedes and silage cut plus precision drilled swedes were 12 and 13 t DM ha−1respectively compared with about 14 t DM ha−1expected from a grass sward.

Notes: Botanical composition of swards can markedly affect their productivity, forage quality, and acceptance by animals. The objective of this research was to determine the amounts of dry matter (DM) and N harvested in the different botanical components of swards where perennial ryegrass and orchardgrass were sown alone and fertilized with N, or sown in mixture with legumes.Broadcast sowings were made on conventionally prepared seedbeds in August 1979 and May 1980 on a Hagerstown silt loam soil (fine, mixed mesic Typic Hapludalf). When seeded alone, orchardgrass cv. Pennlate and perennial ryegrass cv. Reveille received rates of N ranging from 0 to 448 kg ha−1 a−1. The grasses were also grown in mixtures with alfalfa cv. Arc, red clover cv. Arlington, or birdsfoot trefoil cv. Viking. The same legumes were sown alone. Four legume seed rates were used in both the pure legume and mixed stands.Legume DM harvests were greater in mixtures with perennial ryegrass than in legume-orchard-grass swards. An exception occurred in the first year as birdsfoot trefoil harvested DM was lower in trefoil-ryegrass swards than in trefoil orchardgrass mixtures, probably owing to vigorous competition from ryegrass in the seedling stage. As legume seed rates were increased, both the legume and grass components of mixed swards produced more DM and N. Grass DM and N harvested in orchardgrass mixtures exceeded those of ryegrass mixtures for the 3-year period.Sward weediness, most prevalent in pure legume stands planted at low seed rates, was sharply reduced by including either grass with the legumes. Volunteer legumes contributed to DM harvested only when the grasses were sown alone and not fertilized with N.Species differences in crude protein (CP) concentration were significant. Birdsfoot trefoil had the highest CP of the legumes, averaging 241 g (kg DM)−1. Alfalfa and red clover averaged 219 and 208 g CP (kg DM)−1 respectively. Perennial ryegrass herbage had a higher CP concentration than that of orchardgrass in both pure and mixed swards. Grass CP increased significantly in response to fertilizer N and to higher legume seed rates.

Notes: Four cocksfoot varieties were harvested at three different growth stages and offered ad libitum to yearling Clun wethers in three 24-d feeding trials indoors over a 4-month period. A further cocksfoot variety conserved in bulk was offered concurrently as a control feed.The concentration of digestible organic matter in the dry matter (DOMD) and voluntary intake were measured in each feeding period. There was no significant change in the measured DOMD values between the different feeding periods. Voluntary intake of digestible organic matter (DOMI) increased over the 4-month period for all four varieties and for the control feed. The increase in intake was greatest for the most digestible material, i.e. first-cut harvests. Differences in DOMI were statistically significant between feeding periods for all varieties but the differences could be eliminated by correcting intakes with values from the control feed. It is concluded that comparisons of grass varieties involving intake measurements must either be made in the same feeding period or a control feed included and results corrected according to the control response.

Notes: Records of grassland productivity were kept for 2 years on 136 lowland dairy farms and 131 lowland beef farms. The grassland on each farm was classified according to the proportion of sown (preferred) species. The farms were then ranked on this basis and divided into quarters:On dairy farms the lowest quarter had 31% sown species and utilized metabolizable energy (UME) output from grass of 40.8 GJ ha−1, whereas the highest quarter had 70% sown species and UME output of 45.6 GJ ha−1. Dairy farmers in the lowest quarter used least fertilizer N and differences in output were no greater than would be expected from differences in N input. On beef farms the lowest quarter had 28% sown species and UME output of 32.3 GJ ha−1, and the highest quarter had 63% sown species and UME output of 45.9 GJ ha−1On these farms differences in fertilizer N could not completely account for differences in output.It is concluded that on beef farms botanical composition of grassland is a good indicator of its productivity but is not necessarily a prime cause of differences in output. On dairy farms there is little evidence of a direct benefit resulting from a high proportion of sown species. On such farms the need for deliberate manipulation of the botanical composition of grassland by reseeding or chemical means is questioned.

Notes: Twenty-one strains of lactic acid bacteria were subjected to a range of tests based on established criteria to ascertain their suitability for use as inoculants for silage. The tests included a monitor of their growth in the presence of representatives of the microflora of forage crops with which they might have to compete for substrate, and their ability to effect rapid acidification and achieve a low pH in conditions which simulate low and high ensiling potential. In addition, their ability to produce acid from sugars and their action against organic acids found in forage crops, together with their tolerance of acid, elevated temperatures and conditions of low moisture availability (e.g. in wilted material) and possible proteolytic and amylolytic activities were examined. None or these cultures satisfied all of these criteria. However, three strains (Streptococcus durans str. 1024, Lactobacillus acidophilus, str. 2356 and Lactobacillus plantarum str. 6) had a greater potential than the others.

Notes: Three grass silages made in sunny weather in early July from second-harvest perennial ryegrass were compared in a 16-week feeding experiment with twelve Ayrshire cows. The silages were either unwilted or wilted with and without conditioning, and had mean dry matter (DM) concentrations of 201, 261 and 272 g kg−1, and in vitro DOMD concentrations of 650, 669 and 672 g kg−1 DM respectively. All the silages had formic acid (‘Add-F’) applied at a rate of 2.6 litres t−1 and were offered ad libitum plus 6 kg concentrates per cow per d. The daily intakes of silage DM were 905 kg per cow on the unwilted treatment and 9.86 and 9.65 kg on the wilted treatments with and without conditioning respectively. Daily milk yields were 171, 17.6 and 17.4 kg per cow on the unwilted, and wilted with and without conditioning treatments respectively and were not significantly different. Fat concentrations in the milk were not affected significantly by treatment, whereas the crude protein and solids-not-fat concentrations were significantly higher on the wilted than on the unwilted treatment. The efficiency of utilization of metabolizable energy for lactation was 6–7% lower with the wilted than with the unwilted silages and it is concluded that the unwilted silage was superior to the wilted silages as a feed for dairy cows.

Notes: Three diploid—Sabtoron, Violetta and Essex—and three tetraploid—Hungaropoly, Teroba and Red Head—red clover cultivars were sown alone and with each of three companion grasses—timothy (S48), tall fescue (S170) and perennial ryegrass (S24). The productivity and persistence of the red clover cultivars were compared. Results obtained over the first four harvest years were presented in an earlier paper (McBratney, 1981) and this paper presents results from the fifth and sixth (final) harvest years.The low yield obtained in the fifth and sixth year from Essex under each seeding treatment indicated a deterioration in its persistency or vigour or both. In the fifth year the remaining two diploids, Sabtoron and Violetta, were as highly productive and persistent as the three tetraploids but in the sixth year the decrease in productivity was greater in the diploids than in the tetraploids. In the fifth year Teroba, the highest yielder, produced 9 8 t total DM ha−1 over all seeding treatments of which 73% was clover DM; in the sixth year Teroba was again highest yielding, producing 9.3 t ha−l with 67% clover content. Tall fescue was the most consistent companion grass in the mixed swards, in the sixth year contributing 53% grass DM to the total 8 8 t DM harvested ha−1.The trial demonstrates the potential of a number of red clover cultivars grown either pure or with a suitable perennial grass to achieve and maintain a high output of herbage of good quality over a period of 6 years without any application of fertilizer N. The practical agricultural implications of using red clover are discussed.

Notes: Leaf sheaths of two cultivars of perennial ryegrass were prepared by chemical oxidation so that the silica bodies in the tissues could be studied. The cultivar Fortis, which is relatively resistant to stem-borer attack, contained many-intercostal silica bodies in the leaf sheath but the more susceptible cultivar S24 contained few. If the pattern of silica bodies affects levels of stem-borer infestation, it may be possible to breed for this characteristic.

Notes: Ammonia and sodium hydroxide (NaOH) have been used to improve the dry matter (DM) digestibility of roughages or to conserve them. These treatments have had incidental effects on pH alone or both the pH and nitrogen contents of the roughages. in the present study, the effects of pH and NH4OH on the in vitro dry-matter digestibility (IVDMD) of eight roughages—four legumes and four grasses—were investigated. The pH levels of approximately 5, 7, 9 and 12 were achieved by adding appropriate amounts of NH4OH or NaOH and H2SO4. Nitrogen levels of approximately 20, 30, 40 and 50 g kg−1 on a DM basis were obtained by adding NH4OH. Results of a multiple regression showed that increasing the pH caused a linear increment in IVDMD and that the added NH4OH reduced the increment of IVDMD. The effects of both pH and N on IVDMD were found to be specific to the individual roughage; the response of grasses to both pH and NH3 was greater than that of legumes

Notes: In view of the revived interest in sainfoin, an economic assessment of the prospects for an expansion of its area in Britain has been attempted. It is estimated that potentially it could be grown on 950 × 103 ha of the agricultural area of England and Wales, though it is unlikely in the near future that it would exceed 20 × 103 ha, even if economic conditions were favourable. Comparison of the energy and protein yields of sainfoin, red clover and lucerne under conservation management indicates that one of the attractions of sainfoin is its high protein content. However, an assessment of the costs of production and utilization of sainfoin indicate that, for it to be a more widely used legume, sustainable dry matter (DM) yields would need to be increased by 35% to about 11.5 × 103 kg DM ha−1. At the same time, although growing sainfoin as a mixture with grass appears to improve the economic attractiveness of the crop, silage production costs (kg DM)−1 are still about 15% lower on pure grass swards. However, trials in the UK, Canada and Rumania have shown that, compared to current UK levels, sainfoin yields can be significantly increased.

Notes: The relationship between net canopy photosynthesis (Pnc) measured at 400 J m−2 s-1 and leaf area index (LAI) was determined on ryegrass-dominant swards over a range of her bage masses in five grazing experiments. In three experiments the swards were continuously stocked either by sheep or by cattle to maintain a number of herbage heights and hence LAI values. In two experiments the swards were intermittently grazed with 21 -d regrowth intervals. On the continuously stocked swards measurements were made over nine periods each of 7 to 21 d in length. On the intermittently grazed swards measurements were made over six periods while the herbage was being grazed down from a high to a low LAI and over three periods during the regrowth phase.On all the swards where measurements were made while grazing was in progress, the relation of Pnc to LAI was linear. The photosynthetic rate was greater on the continuously grazed than on the intermittently grazed swards up to LAI 35 and in two experiments to LAI 5. The relationship of Pnc to LAI of swards in the regrowth phase was predominantly curvilinear and such that over much of the LAI range they had a greater photosynthetic potential than the same swards being grazed down from high to low mass. Daily rates of CO2 uptake and growth in the field were calculated for one continuously stocked experiment. Both were linearly related to LAI in the range LAI 2–4·5. The agronomic implications of the results are discussed.

Notes: This second paper concerned with effects of prolonged spring defoliations on two early perennial ryegrass varieties (Cropper and RvP Hay Pasture), an intermediate perennial ryegrass (Talbot) and an early cocksfoot (Roskilde) examines the effects of different closing dates on ear emergence, weekly DOMD (in vitro) and conservation yields. The grasses were mown fortnightly to simulate sheep grazing from January until three closing dates, 19 April, 3 May or 17 May, in 1978 and 1979. A set of plots was left undefoliated. All plots were sampled weekly from 2 or 3 weeks after 17 May until the end of June or early July.Prolonged mowing resulted in a small delaying effect on 50% ear emergence of the ryegrasses. The maximum delay was 3 d with the latest closing date. Under this treatment, ear emergence of the cocksfoot was delayed by an average of 12 d but by only 0–4 d under the earlier closing dates. Delay in date of closing caused a significant delay in the time to reach a given DOMD but the effect was largely confined to the latest closing date and was greatest for the cocksfoot. The time when 670 g kg−1 DOMD was reached was delayed in the ryegrasses by no more than 5 d, except for a 12-d delay in the intermediate ryegrass in 1978 following the late closing date. Under this treatment the delay for the cocksfoot was 8 d in both years. The early ryegrasses produced stemmy regrowths.

Notes: For the 2 years 1977 and 1978 Masham ewes grazed a perennial ryegrass (Lolium perenne cv. S24) pasture from the beginning of April until the end of September each year. Twelve ewes were allocated to each of the three stocking rate treatments, 12 (L), 16 (M) and 20 (H) ewes per ha. Each treatment was rotationally grazed around six paddocks with a forward creep for the lambs. Conservation cuts were taken from each treatment in late May and were generally sufficient to cover a 3-month winter feeding period for treatments L and M but were always insufficient for treatment H. Lambing took place at pasture during the second half of May. No concentrates were fed to the ewes during late pregnancy or lactation but lamb birth weights (4–8 kg) and lamb growth rates over the first 4 weeks (240 g d−l) were satisfactory. Organic matter intakes (OMI)of grass by the pregnant ewes(1816,1844 g OMI ewe−1) were not affected by the stocking rate. Intake of grass by the lactating ewes was affected by year and by stocking rate. The overall growth rate of lambs was higher on treatment L than either M or H but decreased on all treatments during the latter half of the grazing season, resulting in 43% of L and only 3% of M and H lambs being fit for slaughter by the end of September. The main features of this time of lambing are low costs and the production of predominantly store lambs.

Notes: A trial was carried out to investigate whether or not sheep grazing at stocking rates of 22 (low) and 44 (high) ha−1 select red clover from a red clover-perennial ryegrass sward, The distribution of botanical and morphological components was assessed in consecutive 5-cm layers through the canopy inside and outside exclosure cages moved every 2 weeks for 6 weeks. The difference between inside and outside cages for a given layer was an indication of the amount of each component consumed from that position in the sward, and the corresponding difference for clover content was considered an indication of selection.Evidence of selection was observed over the third and fourth weeks of grazing, especially in the high stocking rate treatment. Laminae of red clover were selected in preference to other plant parts and were grazed throughout the depth of the canopy in contrast to grass. The basal layer at the high stocking rate during weeks 5 and 6 was not grazed suggesting that the crown of red clover may be protected from severe overgrazing by accumulation of dead matter.It was concluded that the low herbage density of red clover-grass swards allowed the occurrence of selection which was most pronounced when the sward was being grazed down relatively quickly.

Notes: An attempt was made to determine monetary values for grass which might be used by researchers to assess the likely economic implications of their findings. It is shown that the value of extra grass production will vary considerably depending on the use made of it. At 1982 price levels, estimated values for grass in the field ranged from 1–4 to 10–6 pence kg−1 DM, with a modal value around 3–5 pence kg−1 DM. At the same time, the seasonal distribution of the extra grass production is important since the value of extra grass at different points in the growing season may vary markedly. The way in which these values for grass may be used is illustrated, using as an example weed control during sward establishment. Finally, the sensitivity of investment returns to changes in the value placed on grass is discussed. Extreme caution is necessary in drawing general conclusions about benefits and costs from an evaluation based on a specific value for grass.

Notes: In three separate feeding experiments using a total of twenty-six individually-housed Ayrshire cows, three wilted silages made from Blanca white clover were offered ad libitum with either different supplements or different proportions of grass silage. The clover silages contained 680 g white clover kg−1 on a DM basis, and had a mean DM concentration of 263 g kg−1 with 231 g CP kg−1 DM and 91 g ammonia-N kg−1 N. The pH values averaged 4·16 and the DOMD concentrations 611 g kg−1. In experiment 1 the daily intake of clover silage given alone was 15.2 kg DM per cow, i.e. 30·1 g kg−1 live weight, and decreased by 0·76 kg DM kg−1 barley DM and by 0·66 kg DM kg−1barley plus soybean meal DM when these feeds were offered as supplements. Milk yield and fat concentration were higher on the supplement treatments than on the clover silage-only treatment. In experiments 2 and 3 the intakes of silage and total DM increased as the weight of clover in the diet increased from 0 to 700 g kg−1 with parallel increases in milk yield. The effects on milk composition were small and generally non-significant. Although white clover silages with excellent fermentations were made, it is concluded that the main role of white clover in a silage system will be in mixed swards with grass to reduce the input of fertilizer N and to increase the voluntary intake of silage.