ALBERT

Description: New taxa and combinations are published here in anticipation of the revision of the Rutaceae-Pilocarpinae to be published in the near future (thesis, and in Flora Neotropica). Two new combinations of species excluded from subtribe Pilocarpinae are added.

Description: Oil bodies 7-12 in upper leaf cells, 10-20 in elongated basal leaf cells; globose to ellipsoid, 3-7(-10)x3-5 μm; colourless, coarsely segmented, consisting of c. 15-30 aggregated droplets (Colombia, Boyacá, páramos NW of Belén, Cabeceras Q. El Toral, 3765 m, Cleef 2292e; Ecuador, páramos de El Angel, 17 km. S. of Tulcán, 3350 m, Gradstein, Lanier & Weber s.n.). The presence of segmented oil bodies in Colura patagonica is remarkable because previous studies of living Colura (from Japan) reported homogeneous oil bodies (cf. Schuster & Hattori 1964; Inoue 1974).

Description: On 27 May 1976 Peter Arnold Florschütz, bryologist, died at the age of only 53 at De Bilt, Netherlands. Only six weeks prior he had been hospitalized as a result of kidney cancer. His untimely death came as totally unexpected and shocking news to his friends and colleagues all over the world, many of whom had seen him in excellent health the year before at the Botanical Congress in Leningrad. He was a lector of botany and curator of the cryptogamic herbarium at the Institute for Systematic Botany and acting director of the Botanical Gardens of the University of Utrecht, the same institution where he had studied biology from 1941 to 1949. In his professional capacity he had held positions at the Institute for Systematic Botany from 1946 until 1949 as student-assistant and from 1949 on as staff member. Initially under the directorship of his teacher in plant systematics Professor A. A. Pulle, and from 1948 until 1970 under Professor J. Lanjouw’s leadership, the “Flora of Suriname” was being tackled by the staff of the institute. Thus, as a young graduate student, Florschütz was assigned the revision of the mosses of Suriname; a comprehensive and difficult task, because in those post-war years there was a vacuum in European exotic bryology. The heydays, with Herzog in Germany, Brotherus in Scandinavia, Dixon in Great Britain and Thériot and Camus in France were over. At the beginning, Florschütz was entirely dependent on Brotherus’ treatment of the world’s mosses in Engler and Prantl, “Die natürlichen Pflanzenfamilien.” In those years he had run over the leaves of this book for weeks on end in a typical posture, like he used to tell: folded in a chair, book on his lap.

Description: Campylopus introflexus, a new neophyte in western Europe, occurs throughout the Netherlands. After its first appearance in 1961, it is now a common moss. It grows as a pioneer on acid, well-drained places. The differences with C. pilifer are summarized. The occurrence of the latter in the Netherlands could not be affirmed.

Description: Campylopus galapagensis J.-P. Frahm & Sipman spec. nov. is described. It is closely related to C. pilifer Brid., from which it differs mainly by the presence of substereids in the ventral layer of the costa. It is endemic on the Galapagos Islands, where it occurs frequently from sea level to the highest summits at 1500 m.

Description: Tortula grandiretis Broth., differing from T. muralis Hedw. mainly in the larger, quite smooth lamina cells, is reported from three localities in the SW-Netherlands, where it occurred on open, sandy or clayey, brackish soil on recently enclosed mud flats or salt-marshes. It is also reported from one locality in Turkey. It was formerly known only from Turkestan (U.S.S.R.).

Description: The structure of the wood of the genera Castilla, Helicostylis, Maquira, Naucleopsis, Olmedia, Perebea and Pseudolmedia, considered to belong in the Olmedieae (cf. Berg 1972) is described. The diversity in anatomical structure between the genera is small, and it is hard to distinguish Maquira, Perebea and Pseudolmedia from each other. Castilla can be recognized by its thinwalled and wide-lumined fibres, Helicostylis by its parenchyma distribution, Naucleopsis (usually) by its more numerous vessels with a smaller diameter. A more marked difference is shown by the monotypic genus Olmedia with apotracheal banded parenchyma instead of the paratracheal aliform to confluent-banded parenchyma of the other genera. Septate fibres, which are characteristic for the other genera – some species of Helicostylis excepted – are nearly completely absent in Olmedia. This structural difference is considered as an argument in favour of the exclusion of Olmedia from the tribe Olmedieae (Berg 1977).

Description: The new checklist of Dutch liverworts comprises 126 species, 1 subspecies and 5 varieties. Since 1962 seven liverwort species have been added to the flora: Barbilophozia hatcheri, Calypogeia muellerana, Cephalozia pleniceps, Fossombronia incurva, Haplomitrium hookeri, Lophozia perssonii and Plagiochila porelloides. Of twelve species presumed occurrence in the Netherlands needs verification. Nomenclature follows Grolle’s “Verzeichnis der Lebermoose Europas” (Feddes Repert. 87: 171-279. 1976), except for Isopaches, Leiocolea and Microlejeunea, which are maintained as genera and Phaeoceros carolinianus, Cephalozia lammersiana, Chiloscyphus pallescens, Lophozia silvicola and Lophocolea cuspidata. , which are treated as intraspecific taxa.

Description: The new Amazonian liverwort genus Verdoornianthus is considered to be a specialized derivative of the widespread tropical genus Archilejeunea. Differences are the absence of innovations, the dull, suberect leaves, the tristratose rhizoid pad and the larger size of the lobule of the female bracts in Verdoornianthus. There are two species, V. marsupiifolius (Spruce) comb. nov. (Lejeunea marsupiifolia Spruce) from the upstream part of the Rio Negro and V. griffinii sp. nov. from Manaus.

Description: In continuation of de Ruiter’s treatment of Myrianthus and Musanga (Bull. Jard. Bot. Nat. Belg. 46: 471-510.1976), the present paper gives a revision of the African representatives of 17 genera of the Moraceae. The area studied not only consists of the African Continent, but also includes Madagascar, the Comoro Islands, the Mascarenes, the Seychelles, and the Aldabra Islands. Several new combinations are made: Antiaris toxicaria ssp. africana (Engl.) C.C. Berg, A. toxicaría ssp. africana var. usambarensis” (Engl.) C.C. Berg, A. toxicaria ssp. macrophylla (R.Br.) C.C. Berg, A. toxicaría ssp. madagascariensis (H. Perrier) C.C. Berg, A. toxicaria ssp. humbertii (Léandri) C.C. Berg, Broussonetia greveana (Baillon) C.C. Berg, Treculia africana ssp. madagascarica (N.E.Br.) C.C. Berg, and T. africana ssp. madagascarica var. sambiranensis (Léandri) C. C. Berg. Many names are brought into synonymy. Besides revising taxa, the present study aims to fill a gap in our knowledge between Asian Moraceae (studied by Corner, whose studies resulted in a new classification of the family) and the neotropical Moraceae, a subject of study by the present author. Therefore discussions about classification of the family and relationships of African Moraceae with moraceous taxa elsewhere are an essential part of the present paper.

Description: From April 1974 to October 1975 the author conducted field work on the Galápagos Islands for a vegetation study of Santa Cruz and Volcán Alcedo, Isabela. Plants were collected on other islands as well. Thirty-five taxa are new for the archipelago. When determining the material, I found some changes in nomenclature to be necessary. The first set of the collection is in U while a duplicate set will be deposited in CAS. A representative set will be deposited in an Ecuadorian Herbarium. The sequence of the taxa in the Flora of the Galapagos Islands (Wiggins & Porter 1971) is followed.

Description: This year Prof. Dr. F.P. Jonker, Frits as he is known among his friends, will retire from the formal academic life at the State University of Utrecht: a long and busy life of 49 years, devoted to teaching, administration, and scientific research. Looking back on all these years, one realises the important contributions that Jonker has made to botanical science in general and to palaeobotany in particular, both in The Netherlands and abroad, as well as the impact he has exerted on his surroundings, culminating in the vigorous activities of the Laboratory of Palaeobotany and Palynology at Utrecht. To describe Jonker’s life history is indeed to describe the history of his laboratory. To understand the significance of Jonker and the character of the “lab”, we have to trace his life from its very beginnings at the town of Almelo in the eastern Netherlands, where he was born in 1912. His father and mother were teachers and both liked (wild) flowers. Thus both an intellectual and botanical background were already part of his life at a very young age. Soon Jonker joined a group of boy-scouts, where he combined his love for the outdoors with his interest in nature. In high school the biology teacher was Dr. J. Van Beusekom, an Utrecht botanist, who was at the same time scout-master of the scout group. In these formative years, “de Beus” was a decisive factor in influencing Jonker’s career. It was largely because of Van Beusekom that Jonker went to Utrecht University as a student.

Description: El 25 de mayo de 1976 falleció inesperadamente, a la edad de 53 años, Peter Arnold Florschütz, eminente briólogo y profesor de Botánica Sistemática en Utrecht (Holanda). Era bien conocido por sus estudios de los musgos de Surinam. Fue coauter del "Index Muscorum”, miembro de la comisión de la Flora Neotrópica y tesorero del IAPT. Durante sus últimos 10 años estudiaba, junto con la señora Florschütz, los musgos de los Andes colombianos. En 1972 visitó muchas zonas de páramos y selvas andinas, especialmente en los alrededores de Bogotá (Cundinamarca), la Sierra Nevada del Cocuy (Boyacá, Arauca) y el Nevado del Ruiz (Caldas). En 1975 tuvo la oportunidad de visitar nuevamente algunos páramos cercanos a la capital colombiana. Sus colecciones de 1972 y 1975 (con cerca de 1.000 números) se conservan en Bogotá (COL) con duplicados en Utrecht (U).

Description: BAKHUIZEN VAN DEN BRINK Jr, R.C., A synoptical key to the genera of the Rubiaceae of Thailand. Thai Forest Bull. (Bot.) 9 (1976) 15-55. Key of the bracketed type, often leading to flowers as well as fruits, with built-in descriptions of c. 6-12 lines; diagnostic characters are marked. Number of genera 68, incl. 3 introductions and 5 genera not recorded but possibly occurring in Thailand (mostly dependent on delimitation); Craib in 1932-34 has 71. Schumann’s system of 1891 is largely upheld, although no subdivision is here given, and some surprising changes in delimitation occur (e.g. in Keenania, Mycetia, Myrioneuron), which means that many new combinations must be floating around on herbarium sheets. Caution is in order where e.g. on p. 49 Mitragyna seems to have a new section Paradina with a supposedly basal placenta, or where Gardenia is authorized L. on p. 35 but authorized L. emend. Bakh.f. on p. 32. A comparison with Thonner’s keys reveals that Bakhuizen’s key works slower. His generic descriptions are true ’mines of information’ – mining requires a lot of backtracking before all characters can be compared. Desirable as it would be to extend a work like this to all Malesia, it would be better to abandon the Backer-way of keying, and instead describe all genera clearly, and prepare a multiple key as worked out by Leenhouts. Some synonyms are given (Notodontia yes, Quiducia and Symphyllarion no), nomina conservanda indicated, no references, no species. Several critical notes are added. — C.E. Ridsdale & M.J.

Description: Portraits of botanists who worked on the Ryukyu Islands, 80 in number, most Japanese, a few Americans, were published in the book by S. Hatusima, Flora of the Ryukyus, p. 56-75 (1971). Baas Becking, L. G. M. A meticulous bibliography, of the former Professor of Experimental Botany at Leiden and later Director of the Bogor Botanic Gardens, was prepared by J. Westenberg, 20 p. (North Holland Publishing Company, Amsterdam, 1977).

Description: Brunonia is the title of a journal that will replace the Contributions from Herbarium Australiense (last no. 17, 1976). Subscriptions Aust. $ 4. annual, Herbarium Australiense, P.O. Box 1600, Canberra City, A.C.T. 2601, Australia. Nature Malaysiana, published quarterly by Tropical Press, 64A Jl. Bukit Bintang, Kuala Lumpur, Malaysia, started in July 1976. The price is Mal. $ 2.50 a copy. This first number, size 28 by 20 cm, containing 40 pages of text and some pages of ads, is devoted to ’our natural heritage’. It is full of showy photographs all in colour, with high quality popular texts on snakes, malaria parasites, spiders, wild orchids, mantis, frogs and elephants. Execution is very good. The journal seems aimed at the general educated public, well suited for display in airline offices, dentist’s waiting rooms, the reading table in an embassy, etc. where is surely will make life more pleasant, and set people’s minds in the proper direction.

Description: Austrobaileya replaces the Contributions from the Queensland Herbarium, and was devised to accommodate also shorter taxonomic notes. The Contributions amount to 20 numbers, with one article each; a cumulative index of names is in no 20, p. 73-88. In format and execution Austrobaileya resembles its predecessor but the useful page heads should be retained. Volume 1 number 1 (1977) was received in March 1978. It carries 9 papers on 74 pages, and a map with subdivisions of Queensland on the back flap. Frequency and price are unknown. Editor: L. Pedley, Queensland Herbarium, Meiers Road, Indooroopilly, Qld. 4068, Australia. Brunonia replaces Contributions from Herbarium Australiense or rather seems a continuation of it in the same scope under a new name, and paged through per volume. The first issue appeared on 24 February 1978, it has 129 pages, carrying 11 papers. It will be ”issued at irregular intervals”. Subscription is A$ 10 per annum. Editor is B.J. Walby, CSIRO, Box 89, East Melbourne, Vic. 3002, Australia.

Description: Localizing specimens and mapping localities has always been a tedious and time-consuming task for which much depends on the data mentioned on the labels. It has been found a blessing if collectors mention on labels the latitude and longitude. If this is given in an exact way it comprises degrees and minutes, e.g. 6° 45’ S, 141° 30’ E. If no dot-map is provided this appears to be a slightly clumsy formula in print and the question arises whether such exact figures are really needed. In scanning a geographical map the minutes will hardly mean something unless one uses local small-scale maps, as one minute is only a little more than 2 km in the terrain. In Pretoria only the degrees are given, joined into one figure, preceding the collector / after the locality. This simplification is, I think, practical and useful.

Description: On 3 September 1977, Dr. H.P. Nooteboom (L) went to Ceylon for 2 months to collect additional material of Symplocaceae for ’A revised Flora of Ceylon’. Although this project was due to end by September 1977, it appeared to have been extended for another year. The genus Symplocos, with about 20 taxa, is found in the wet zone (in the mountains of the central part, in the mossy forest up to 2400 m, descending to sea-level in the everwet primary forest in the SW. part of the island). Some species also occur in the secondary forest in the same region, one species is found in the whole island, in a variety of vegetation types, but mostly in secondary forest and shrubbery. Dr. Nooteboom could collect material of all the taxa, sometimes in many individuals, which revealed the difficult patterns of variability. Besides he made also general collections (Nooteboom 3036—3420). The weather was extremely bad; heavy rains caused inundations and landslides. Therefore the total number of collections was limited. Labelling and distribution is still going on.

Description: Flora Malesiana series i volume 8 instalment 2, pages 31-300, came from the press in December 1977*. It contains the Ulmaceae by E. Soepadmo: 6 genera, 27 species; the Iridaceae by D.J.L. Geerinck: 6 genera, 7 species; the Cornaceae by K.M. Matthew: 1 genus Mastixia with 10 species; the Onagraceae by P.H. Raven: 2 genera, 14 species; the Bignoniaceae by C.G.G. J. van Steenis: 15 genera, 31 species + in concise treatment 23 ornamental species; the Crypteroniaceae by R.J. van Beusekom-Osinga: 3 genera, 8 species; the Symplocaceae by H.P. Nooteboom: 1 genus Symplocos, 58 species; the Lentibulariaceae by P. Taylor: 1 genus Utricularia, 22 species. Volume 8 instalment 3 is in proof. It contains the Labiatae and Anacardiaceae, as well as some Addenda, the Dedication to F.A.W. Miquel, and the Index, since volume 8 will then be completed.

Description: The preparation of botanical drawings is a craft in its own right, and furthermore, draughtsmen are human beings. Even these simple truths are trodden down by the taxonomist who during a final hour hands the draughtsman a bundle of specimens and some hasty indications. Naturally the result is anguish and confusion. Let us therefore add some observations to improve the situation. First: a botanical artist looks at plants with a different eye from the taxonomist – that’s why he is an artist and not a scientist. Fortunately, some overlap exists, where the two can meet.

Description: The entries have been split into five categories: a) Algae – b) Fungi & Lichens — c) Bryophytes — d) Pteridophytes — e) Spermatophytes & General subjects. — Books have been marked with an asterisk.

Description: Dr. J.A.R. Anderson, former conservator of Forests, Kuching, now consultant forester and ecologist, new address: 30 Greenhill Gardens, Edinburgh EH10 4BP, U.K. His Far East address: c/o Room 432, 4th floor, Katong Shopping Centre, Singapore 15. Dr. P.S. Ashton of Aberdeen spent months in Kuala Lumpur, during the second half of 1975, principally to teach economic and forest botany at the University of Malaya.

Description: Some of the species and names of hydnoid fungi treated in Furukawa’s work are discussed. Hydnum albidum is recorded in Europe for the first time. Further finds of some interesting species are reported. Auriscalpium barbatum (Western Australia) and Steccherinum peruvianum (Peru) are described as new species. A key to the species of Auriscalpium is given.

Description: Subramanian’s concept of true and false chains of phialoconidia is rejected and replaced by a distinction between connected and disconnected chains. In connected conidial chains the primary conidial wall is strongly thickened at both ends and a connective is formed. This criterion allows the distinction between trichocomaceous or eurotiaceous (connected) and sphaeriaceous (disconnected) catenulate phialoconidia. The ultrastructure of conidiogenesis is described. On the basis of this criterion, the species of the Acremonium diversisporum series as well as the anamorph of Sagenoma viride Stolk & Orr with connected chains are transferred from Acremonium to the new genus Sagenomella to which four new species are added.

Description: The author describes two new subspecies of Rubus, viz. R. schlechtendalii subsp. subcentreuropus Beek and R. glandulosus subsp. picearum Beek.

Description: In the course of a study of coprophilous fungi collected in the Zoological Garden in Delhi, an ascomycete belonging to the genus Achaetomium was isolated. It differs from previously described species by larger ascospores and almost colourless ascomata with a wide apical opening.

Description: Identities are given for all 35 combinations published in Anplectrum A. Gray ( Melastomataceae) with additional notes on some of the taxa involved. Two new combinations are proposed in Creochiton Bl. and one in Dissochaeta Bl.

Description: A new monotypic genus Tamaricaria Qaiser & Ali of Tamaricaceae is described with a new combination i.e. Tamaricaria elegans (Royle) Qaiser & Ali.

Description: 1. The interrelations between the genera together constituting the Dimocarpus group in the tribe Nephelieae are represented in a scheme. In this scheme are added the main characters that are thought to be of phylogenetic importance. 2. A neotype.is proposed for Cubilia cubili (Blanco) Adelb., the single species of its genus. To its distribution can be added the eastern half of Borneo, incl. also the Island of P. Laut. Mention is made of a geographic clinal variation in a few macromorphological characters. 3. Lilchi is considered to comprise only one species, L. chinensis Sonn., which is subdivided into three subspecies: subsp. chinensis, the commonly grown form, cultivated for thousands of years already, apparently adapted (by nature or partly by selection by man?) to a monsoon climate, if actually wild probably originating from northern Indo China; subsp. philippinensis (Radlk.) Leenh., a wild form closely related to subsp. chinensis, known from the Philippines and New Guinea; and subsp. javensis Leenh., strikingly different from both other forms, known only as a cultivated fruit tree from southern Indo China and Java, apparently adapted to an everwet tropical climate. For subsp. philippinensis a lectotype is proposed. 4. Pometia, though macromorphologically distinctly derived and, moreover, palynologically apparently very exclusive in the alliance under discussion, seems clearly connected with Dimocarpus, the central genus in the group.

Description: Description of a new species, Diploglottis bracteata Leenh., from Queensland, Australia. Reduction of Euphorianthus (E. Malesia) to Diploglottis (NE. Australia). Discussion of the occurrence of actinomorphic and zygomorphic flowers in the Sapindaceae in nearly all tribes and even within ten genera. Discussion of the systematic position of Diploglottis bracteata: this species seems distinctly allied to and more derived than the New Caledonian genus Storthocalyx, and thus may belong to an old element of the Queensland flora allied with that of New Caledonia. On the other hand, D. bracteata is within Diploglottis closest to the East Malesian species, whereas the further Australian species are distinctly more derived. They may belong to a younger element in the Queensland flora of Malesian derivation.

Description: Herbae erectae caespitosae 1.5—2.0 m altae; rhizoma suberectum 1—2 cm diametro. Folia multa apice rhizomatis spiratim aggregata; petioli usque ad 1 m longi basi vaginantes, sparsim pubescentes; laminae ellipticae oblongo-lanceolatae glabrae 18—35 cm longae, 8—15 cm latae, basi acutae vel cuneatae, apice breviter acuminatae, infra subvirides, nervis lateralibus multis parallelis, nervulis multis scalariformibus inter nervos subtiliter transverse reticulatis; costa basi incrassata gradatim angustata et a nervis lateralibus haud distincta. Pedunculus principalis terminalis usque ad 1.5 m altus ad apicem folio uno inflorescentia unaque, raro folium deest; folium pedunculare foliis primariis simile petiolo ad medium usque vaginante parce piloso, apicem versus pubescente. Inflorescentia laxe paniculata e spicis spiculisque constans, in axillam vaginae folii pedunculati, pedunculo ad 15 cm longo, bractea basi vaginante lineari-lanceolata puberula ad 10 cm longa; panicula puberula, 2—10-ramosa usque ad 15 cm longa et 10 cm lata; spicae 2—10, longitudine variantes, 3—8 cm longae usque ad 4 cm distantes e axillis bractearum principaliarum spiratim dispositae in rachibus principalibus; bracteae principaliae ovati-lanceolatae ± 2 cm longae, ± 1 cm latae, extus puberulae margine hirtae, cito ad medium usque, tandem basem versus in fibras multas secedentes; spiculae multae e axillis bractearum primariarum subspiraliter in spicis dispositae, usque ad 2 cm longae; bracteae primariac ovatae obtusae mucronatae ± 1.5 cm longae, ± 1 cm latae, extus puberulae margine hirtae bracteis principalibus similes. Flores albi in cymis parorum 1—3 vel plurorum; omnis par e axilla prophylli, flos unus paris omnis primo aperiens, ceterus invicem; prophyllum externum magnum dorsaliter 2-carinatum ovato-lanceolatum margine incurvum ± 14 mm longum et ± 5 mm latum, alis carinarum fimbriatis; prophylla interiora dorsaliter 3-carinata parva; pedicelli omnis paris basi connati ± 1 mm longi. Calycis lobi usque ad basim discreti, lineari-lanceolati extra hirti,± 1.2 cm longi, ± 2 mm lati. Corolla basi tubularis, tubo ± 7 mm longo lobis oblongis ovatis erectis obtusis ± 8 mm longis. Staminodia 4 in tubum connata tubum corollae adnata; lobi staminodiorum extemorum subaequales; lobus magnus obovatus obtusus marginibus recurvatis, ± 1 cm longus, ± 8 mm latus; lobus parvus subspathulatus recurvatus, ± 7 mm longus, ± 3 mm latus; staminodium interioris carnosum cucullatum, lobo laterali apicem styli includenti, ± 6 mm longum et ± 3 mm latum; staminodium interiorissimum cucullatum, staminodium extimum magnum omnino adnatum apicem stigmatis includens, infra stigma callo carnoso puberulo instructum, lobo ad laterem stigmatis auriculato. Stamen fertile 1-cellulare appendice subulata ± 1 mm longa uno latere instructum; filamenti segmentum librum ± 3 mm longum; anthera oblonga apiculata lutea ± 2 mm longa. Ovarium ±2 mm longum dense aureo villosum; stylus tubum staminalem adnatus ± 1 cm longus segmento libro cucullato ± 4 mm longo; stigma truncata irregulariter lobata. Fructus 3-lobatus sparsim breviter hirtus rubiginosus ± 9 mm longus, ± 7 mm latus; semina 3 oblonga subtrigona lurida ± 7 mm longa et ± 4 mm diametro arillis 2 lobatis albis lobis lineari-acuminatis ± 3 mm longis. Erect caespitose herbs, 1.5—2.0 m tall; rhizomes suberect, 1—2 cm thick. Leaves many, spirally crowded at apex of rhizome; petiole sheathing at base, up to 1 m long, sparsely pubescent; leaf-blade elliptic to oblong-lanceolate, acute to cuneate at base, shortly acuminate at apex, 18—35 cm long, 8—15 cm wide, glabrous, whitish green beneath; lateral nerves many, parallel, with many fine scalariform transversely reticulate nervules in between; midrib thickened towards base, gradually narrowed and indistinct from lateral nerves towards apex. Main peduncle terminal, up to 1.5 m tall, bearing one leaf and inflorescence at apex, rarely without leaf; peduncular leaf similar to primary leaves; petiole up to 15 cm long, sheathing at basal half, scattered hairy, pubescent towards apex. Inflorescence Inflorescence a lax panicle of spikes and spikelets, arising from axil of peduncular leafsheath with up to 15 cm long stalk, subtended by up to 10 cm long, linear-lanceolate, sheathing, puberulous bract at base; panicle up to 15 cm long, up to 10 cm wide, 2—10- branched, puberulous; spikes 2—10, of various length, 3—8 cm long, arranged at distances of up to 4 cm apart in axils of spirally arranged bracts on main rachis; bracts ovate-lanceolate, ± 2 cm long, ± 1 cm wide, puberulous outside, hairy at margins, soon splitting up in upper half into many fibres, later up to base; spikelets many, subspirally arranged on spikes in axils of primary bracts, up to 2 cm long; primary bracts ovate, obtuse, mucronate, ± 1.5 cm long, ± 1 cm wide, puberulous outside and along margins, similar to main bracts. Flowers white, in cymes of 1—3 or more pairs; each cymule (pair of flowers) in axil of a prophyll, one flower in each pair opening first, the other next; outer prophyll large, 2-keeled at back, ovate-lanceolate, ±14 mm long, ± 5 mm wide, incurved at margins, minutely fringed on wings of keels; inner prophylls 3-keeled at back, smaller; pedicels of each pair of flowers united at base, ± 1 mm long. Calyx-lobes free up to base, linear-lanceolate, ± 1.2 cm long, ± 2 mm wide, hairy outside. Corolla-tube ± 7 mm long; lobes oblong, ovate, obtuse, ± 8 mm long, erect. Staminodes 4. united into a tube and adnate to the corolla-tube; lobes of outer staminodes subequal; larger lobe obovate, obtuse, with recurved margins, ± 1 cm long, ± 8 mm wide; smaller lobe subspathulate, recurved, ± 7 mm long, ± 3 mm wide; fleshy inner staminode hood-shaped with a lateral lobe enclosing style apex, ± 6 mm long, ± 3 mm wide; innermost staminode hooded, entirely adnate to outer large staminode, enclosing lip of stigma, with a fleshy puberulous callus below stigma and an auricular lobe at side of stigma. Fertile stamen 1-celled, with a thin, subulate, ± 1 mm long appendage on one side; free portion of filament ± 3 mm long; anther ± 2 mm long, oblong, apiculate, yellow. Ovary ± 2 mm long, densely golden hairy outside; style adnate to staminodial tube, ± 1 cm long; free portion curved, hooded, ± 4 mm long; stigma irregularly lobed, truncate. Fruit 3-lobed, ± 9 mm long, ± 7 mm wide, sparsely short hairy, reddish-brown; seeds 3, oblong, subtrigonous, ± 7 mm long, ± 4 mm thick, dull brown; arils 2, lobed, white; lobes linear-acuminate, ± 3 mm long.

Description: The Star Mountains of New Guinea are situated at the geographic center of the Island of New Guinea extending on both sides of the Indonesian-Papua New Guinea border. Access to these mountains from either side of the border which divides the island is relatively difficult and as a result few collections have come from the area. A Dutch expedition traveled to the western Star Mountains in 1959, but ran into various difficulties and as a result did little collecting above 1500 m. In 1975 an expedition sponsored jointly by the Division of Botany, Lae, and the Rijksherbarium, Leiden, returned to the Star Mountains collecting extensively throughout the eastern half of the range. The results of this expedition include the first extensive collections of material from the higher altitudes within the Star Mountains. Material collected for the Division of Botany, Lae, by J. R. Croft and G. S. Hope while on the 1975 expedition is surprisingly rich in species of Rhododendron. I was asked by Mr. Croft to examine the Lae material prior to its distribution. The collections contain representatives of several poorly known species of the genus, at least one new plant record for Papua New Guinea, Rhododendron rubrobracteatum Sleumer, and the new taxon described below.

Description: Tristania R. Brown, Aiton’s Hortus Kewensis (2nd Ed.) 4 (1812) 417, was established with three species — T. neriifolia, T. laurina, and T. conferta. A number of other species have since been added to the genus and a recent study (Wilson, 1971) has shown that the three original species belong to three different groups and further that these groups are sufficiently different to warrant their separation at the generic level. All of the New Caledonian species belong to the Tristania laurina group. It has not yet been decided which of the groups should retain the original generic name, but if the T. laurina group is not selected the name Tristaniopsis Brongniart et Gris, Bull. Soc. Bot. Fr. 10 (1863) 371, would become available for it. Six species are currently recognised in New Caledonia where they mostly grow at low elevations in scrub and forest on ultrabasic rocks. Species of the same group are found in Australia, New Guinea, Borneo, and probably elsewhere in Malesia.

Description: The pollen morphology of all 11 species of the genus Mischocarpus is studied. All species possess basically the same syntricolpate pollen type. Transitions to the tricolpate type were observed rarely. Within the syntricolpate type, subtypes could be established. For a few species a rather wide range of variability in some characters is described. Pollen morphology correlates with macromorphology as well as with geography, thus supporting the results, based on macromorphological evidence, concerning infrageneric structure and relationships of Mischocarpus.

Description: At the age of 85, Herman Johannes Lam died at his house on the 15th of February, 1977. From 1933 to 1962 he was director of the Rijksherbarium and although the day of his retirement lies some 15 years behind us now, he is still remembered in our institute for his pleasant personal qualities. The Rijksherbarium as it is today we owe for a large part to his vision and work during the 29 years of his directorate. He broadened the basis of the institute’s research but kept intact its specialization; he succeeded in obtaining valuable collections; he started a programme of botanical expeditions; he provided a home for the Flora Malesiana, to mention some of his accomplishments. When he came to Leiden after a 14 years’ career in the Herbarium at Buitenzorg (now Bogor, Indonesia) he found a small and rather sleepy institute. Through the years of poverty before and during the war, and through the years of prosperity afterwards, he transformed this into a large herbarium which was (and still is, I hope) very much alive and active in many fields.

Description: In this fourth report on the taxonomy of Ulothrix Kützing a new classification of the marine and brackish-water species in western Europe is proposed. Comparative studies on field collections, uni-algal cultures, herbarium collections and sections prepared for electron microscopy lead to the recognition of three marine species, viz. Ulothrix speciosa (Carmichael ex Harvey in Hooker) Kützing, U. flacca (Dillwyn) Thuret in Le Jolis, and U. palusalsa Lokhorst (nov. nom.), and two brackish-water species, viz. U. implexa (Kützing) Kützing and U. subflaccida Wille. The vegetative anatomy, the life history, the fine structure of the vegetative thallus and the distributional pattern in nature are amply discussed. Salient, reliable characters proved to be, e.g., the nature and construction of the cell wall, the texture of the cell wall’s surface, the fine structure of the pyrenoid, the developmental stages of germinating zoospores, the coalescense of filaments, the shape of the gametangial filament, and the limited variation of the number of zoospores and gametes. A brief discussion is given of the ecological status of the individual species. In addition there is a brief comment on the taxonomic affinity of Ulothrix with the morphologically related genus Urospora Fries and on the phyletic relationship of Ulothrix with the progenitors of the higher land plants. The reproductive behaviour of the species under different photo periods in culture appeared to be correlated with the seasonal periodicity expressed by the algae in nature.

Description: There has been a notable divergence of opinion in the recent literature concerning the number of species of dammar (Agathis) that might occur in the general area of the Moluccas. Even early literature presents a tangled nomenclatural trail. Perhaps the difficulty in obtaining good representative collections from these huge rainforest emergents may explain the general lack of careful diagnostic descriptions that bedevils most contributions. Among the hundreds of specimens I have been able to study, however, I have found enough data to support a clear conclusion. The important dammar tree was among those described in the early work by Rumphius (1741) that dealt with Ambon. Meijer Drees (1940) reports that natives in the Moluccas recognize two types of dammar, the ‘white dammar’ damar putih) with abundant resin production and the ‘brown dammar’ damar merah) with poor resin production (the ‘white’ or clear resin does turn brown upon aging about a year). Presumably, Rumphius, who spoke of abundant resin, had in mind the ‘white dammar’ when he referred to this tree as Dammara alba.

Description: A world-wide revision of the tribe Naucleeae with a general discussion of the systematic position and affinities of the tribe and the genera. The generic concepts have been modified and 21 genera are recognized (Ochreinauclea, Ludekia, Diyaminauclea, Khasiaclunea, Adinauclea, Sinoadina, Pertusadina, and Haldina being new), which are placed in three subtribes, Anthocephalinae, Naucleinae, and Adininae, sublrib. now There are keys to the subtribes, genera, and species, followed by descriptions of the Asiatic and Malesian genera. The Asiatic species are described and accompanied by complete synonymy, but the Malesian species are treated in an abbreviated form. Three new species are described: Myrmeconauclea stipulacea, Ludekia borneensis, and Pertusadina malaccensis.

Description: The species related to Medinilla myrtiformis (Naud.) Triana are described and a key is given. A variety of M. monantha Merr. is recognized as a distinct species, while the typical form is considered a synonym of M. myrtiformis. M. neglecta Nayar is reduced to M. rubrifructus Ohwi.

Description: The type species of tie genus Polyscias (P. pinnata J. R. & G. Forst.) is closely related to a small number of Pacific and Indo-Malayan species, several of which have long been in cultivation. This group of species have a distinctive facies but can be defined most readily by the elongated sheathing leaf-base. The genus has usually been extended beyond this group to include other pinnate-leaved members of sub-family Schefflereae in which the pedicel is articulated below the flower. There has been uncertainty whether to restrict the genus to species in which the style arms are free or also to include species with connate styles. In his treatment of the New Guinea species, Harms (Bot. Jahrb. 56,1920: 374—414) does include some species with connate styles within Polyscias though also retaining the genus Kissodendron, a genus distinguished from Polyscias mainly by the united style arms. Bernardi (Candollea 26, 1971: 13—89) resolved this difficulty by uniting Kissodendron, and also Palmervandenbroekia, with Polyscias and this treatment is followed here. As thus defined, the genus comprises rather diverse elements, and these have been given sectional status.

Description: A list of the species and subspecies, including synonyms, of the tipulid subfamily Ctenophorinae is provided. References to the literature are nearly complete. The distribution of the species is indicated by abbreviations and figures referring to the geographic regions and subregions. A survey of the distribution of the 5 genera is given separately.

Description: 1. Samples of benthic invertebrate fauna were collected in – a ditch, situated in a protected area in the peaty area near the Maarsseveense plassen (prov. Utrecht), – four ditches in the neighbourhood of Castricum (N-Holland), not far from the dune district, – two ditches with water of a high chloride-content on Texel (N- Holland). 2. The faunistic results, with the additional information of some physical and chemical data on the different sampling points, made it possible to draw some conclusions concerning – the importance of the factor salinity for a faunistically oriented typology of ditches, – the importance of the group Hydracarina for a classification of ditch-biocenoses, – the influence of artificial current, caused by the inlet of water, on the invertebrate fauna of ditches and on the faunistical evaluation of the pollution-level in ditches.

Description: An annotated list of the brachyuran (12) and anomuran (1) tree-climbing crabs of Trinidad (West Indies) is presented (see Table 1 for species names). Some of the species mentioned (e.g. Aratus pisonii, Goniopsis cruentata) are well-known treeclimbers, in others (e.g. Sesarma roberti, S. ricordi) this peculiar behaviour is recorded for the first time. Some data on the diet and the locomotion of climbing Grapsidae are given. Aratus was found to feed mainly on algae and decayed wood, not on mangrove leaves. A synopsis of pertinent data from literature (Table 2) yielded 30 further species names of tree-climbing crabs. Like in Trinidad, most of these belong to systematic groups comprising many or only semiterrestrial species. The trees ascended are mangroves in about half of all cases. Motives, grades, phyletic routes, predispositions and consequences of the habit of tree-climbing are discussed.

Description: The study of some newly collected material from the West Indies may justify a fourth paper on Caribbean Tenebrionidae in these “Studies”. Thanks to dr. P. WAGENAAR HUMMELINCK’S collecting work, the Tenebrionid fauna of the Antilles and the adjacent South American mainland shores may be considered to be pretty well known — at least as far as the Melasomes are concerned. Thus zoogeographical conclusions — though not differing essentially from those published in 1962 — appear to have a rather solid basis. Unfortunately much less is known about planticolous Tenebrionids, which anyhow are relatively less interesting for zoogeographical purposes, than the geophilous ones. We also had the privilege of consulting the collections of the I.N.R.A. at Guadeloupe (see MARCUZZI & D’AGUILAR 1971) which considerably increased our knowledge of the Tenebrionid fauna of that and neighbouring islands. Several specimens on hand at the Institute of Marine Biology, Mayagüez, proved extremely useful for obtaining a better knowledge of the Tenebrionid fauna of the old, sedimentary island of Puerto Rico. In a few single cases material from other sources (British Museum, Museum G. Frey and the private collection of the author) has been used.

Description: Some twenty-five years have passed since short descriptions were published of marine and saltpond habitats sampled in the Caribbean during three zoological collecting trips made by the author in 1930, 1936/37 and 1948/49 (these Studies, vol. 4, no. 17, 1953). Sampling of the shallow coastal waters of the Caribbean was continued in 1955, 1963/64, 1967, 1968, 1970 and 1973, during six visits the main purpose of which was not always the study of the marine fauna. Although collecting was done single-handed and rather incidentally, with no other equipment than a knife, fine-meshed nets, formaldehyde and alcohol, the material collected proved to be sufficiently valuable for scientific purposes to justify the publication of a list of the new marine localities. In this paper the descriptions of the “Marine Habitats”, published in 1953 (p. 56-58) are included, but those of the “Salt Pond Habitats” (p. 69-77) are only referred to.

Description: During a study still in progress on the fauna of phreatic waters of various West Indian islands, a number of isopods of the family Microparasellidae were obtained. Although material from some 20 Caribbean islands was examined, only the island of Bonaire yielded microparasellid isopods so far. These animals are described in the present paper. Up to now, the only other West Indian records for the family are those of COINEAU & BOTOSANEANU, 1973, from Cuba. The methods employed are the same as in my previous study (STOCK, 1976a) on the Antillean Thermosbaenacea. All chlorinities have been determined with the aid of an E.E.L. electric chlorinimeter.

Description: In 1973, 1974 and 1975 I visited St. Maarten, Netherlands Antilles. During these visits I spent much time observing birds on this island, while short excursions were also made to the neighbouring islands Saba and St. Eustatius. The periods of our visits were: ST. MARTIN: 7-19 April, 30 April-3 May and 11-19 May 1973, 1-7 and 16-27 February, 2-11 and 18-31 December 1974, 6-13 January 1975. SABA: 4-8 May 1973. ST. EUSTATIUS: 8-10 May 1973, 8-15 February and 12-17 December 1974.

Description: Since publication of “The Amphibia of Trinidad” (KENNY 1969) some minor errors have been drawn to the attention of the author. Also, it has been possible to do some additional field observations which have resulted in a new record for Trinidad and the extension of the distribution of two species. This short paper summarises this new information.

Description: Rosacea flaccida, a new prayine siphonophore, is described from specimens collected by SCUBA divers in the upper 30m of the subtropical and temperate North Atlantic Ocean. The new species has stoutly cylindrical, flaccid nectophores and delicate flattened bracts. The nectophores are morphologically similar to those of R. plicata sensu Bigelow, 1911 and R. cymbiformis (delle Chiaje, 1822) having a deep hydroecial groove and meandering lateral radial canals in the nectosac. In one of the nectophores there is a slight dorsal prolongation of the somatocyst at its apical end into the mesoglea. The eudoxid bracts are distinctive, being flattened dorsoventrally and divided, on the proximal side of the stem, into two lobes which are twisted at an angle of approximately 90° to the lobe on the distal side. Right and left longitudinal bracteal canals are well developed. The origin of the dorsal bracteal canal from the right longitudinal canal differs from that in the other Rosacea species but resembles the configuration found in the bracts of species of the genus Praya.

Description: Epifaunal community development on plexiglas panels was observed over one year. The course ofcolonization is described, and some data are presented on autecology, reproduction, and growth rate ofparticular species. About three months after initial settlement, conditions of coexistence in a mixed barnacle-ascidian community began to change increasingly due to heavy competition for space. The colonial species Botryllus schlosseri proved to be potentially dominant. Shortly before it attained monopolization by replacing barnacles (mainly Elminius modestus), a major physical disturbance eliminated the fast growing ascidian. The roles of physical factors, of biological interactions, and of historical events in community development are discussed in context with succession theory and other concepts evolved more recently. It is concluded that succession-like processes can occur in subtidal fouling communities, but there the existence of a globally stable climax is unlikely. Generally the concept of multistable points seems to be better applicable to marine ecosystems than that of succession in the classical sense.

Description: The quantitative distribution of some important benthic invertebrates in a shallow inshore area of Kiel Bay (Western Baltic Sea) is described. This region is partly polluted by domestic sewage from the municipality of Kiel. Three groups of species are distinguished with reference to their densities and other population parameters in the different subareas: Progressive species of the 1st and 2nd order indicate a high and moderate to slight degree of pollution respectively by high population numbers. Regressive species are adverse indicators, absent or occurring in exceptionally low numbers in affected areas. In many cases the examination must be concentrated on sand bottom, because a series of species normally dwelling on aufwuchsbuild up dense populations in sand, if it is organically enriched. The polychaetes Capitella capitata, Nereisdiversicolor and Polydora ligni are regarded as progressive species of the first order. The amphipods Corophium insidiosum and Gammarus salinus, the mussel Mytilus edulis, the polychaete Pygospio elegans, the snail Hydrobia ulvae, and some other invertebrates belong to the group of indicators for slight organicpollution. The main regressive species are the amphipod Bathyporeia sarsi and the mite Copidognathus fabriciusi.

Description: In-situ studies on sublittoral soft bottom macrofauna (depth: 14-16 m) employing the underwater laboratory (UWL) Helgoland were carried out. Sets of samples were compared for small-scale local and short-term changes in species richness, faunal abundance, numerical dominance, diversity, evenness, homogeneity, and similarity. It could be shown that minor differences in sediment quality can cause conspicuous heterogeneity within a small sampling area (diameter = 100 m). Both spatfall and mortality of benthic invertebrates can change the faunal structure within a short period (two months). The degree of change varies between species and thus at stations harbouring different faunal assemblages as well.

Description: In-situ investigations on the life of the common sea star (Asterias rubens L.) were carried out in 1976, employing the Underwater Laboratory 'Helgoland' in Lübeck Bay (Western Baltic Sea). The abundance of A. rubens amounted to 2-31 m SUP--2 on sediment (fine sand), and to 324-809 m SUP--2 on mobile algal carpets drifting over the bottom. Actual population parameters (abundance, size class distribution) areinfluenced by both substrate quality and drifting. Stomach investigations revealed prey-size selectivity: small sea stars feed mainly on the snail Hydrobia ulvae when living on the sediment, but on mussel brood (Mytilus edulis) in the phytal. The principal food items of larger sea stars are the sand-dwelling clam Macoma baltica and the phytal-living isopod Idotea baltica respectively. A.rubens is very adaptive to the food availability; the diversity of its diet corresponds to the species diversity found in its environment. A change of biotope during active or passive migrations causes switching. The sea star is able to catch motile animals and to dig outinfaunal clams. It exhibits a diurnal feeding pattern related to light periodicity; the activity decreases at night. The average frequency of feeding is highly dependent on predator body size; it declines with growth. In situ-experiments indicate an exponential relationship between the feeding duration upon M. baltica and the quotient of clam size to logarithm of sea-star size. An approach is made toward a rough estimate of macrofauna consumption by A. rubens on sediment. The sea star seems to be an important predator and thus a competitor of demersal fishes on soft bottoms of the western Baltic Sea.

Description: Aspects of the life cycle of the cumacean D. rathkei, the most important food item of demersal fishes in the Western Baltic Sea, were investigated from the Underwater Laboratory (UWL) 'Helgoland' in Lübeck Bay. In Aug the population density was 10.1 ±3.6 individuals 100 cm Super(-2) in fine-medium sand (Md = 292 µm), and 3.6 +- 2.4 individuals in coarser medium sand (Md = 470 µm). In both substrata, abundance values decreased until Oct, at each station to a different extent. Particular attention was paid to vertical migrations of D.rathkei. Plankton catches and numerous direct observations provided evidence that the pelagic phase, observed at night, is always combined with ecdysis. During short periods of high swarming andmoulting activities, the population structure changed considerably: juveniles of the 3rd stage dominated inearly Aug; thereafter they were replaced by 'prematures', the last juvenile stage. The first mature individualswere observed in Oct. Some data are given on growth rate.

Description: Abscission of anthers was found in most Cecropia species studied for a taxonomic treatment of this genus for the Amazon region of Brazil*. Cecropia is a common genus in the neotropics. It comprises about 80 species. Most of them are arborescent pioneer plants and common in more or less open (secondary) vegetation.

Description: The hydrophilous plant communities of Western Crete described and classified in this paper belong to the following classes: Potametea, Adiantetea, Phragmitetea, Molinio- Juncetea, Juncetea maritimi, Isoeto-Nanojuncetea and Alno-Populetea. Two new alliances and five new associations are described: Brachypodio-Holoschoenion, Dorycnio- Rumicion conqlomeratae. Dorycnio-Caricetum otrubae, Dorycnio-Cladietum marisci and Caricetum creticae (all Molinio-Juncetea), Acrocladio-Adiantetum (Adiantetea), and Juncetum subulato-maritimi (Juncetea maritimi).

Description: Eight Stereocaulon (Lichenes) species have been found in the Netherlands. Of these S. paschale (L.) Fr. and S. tomentosum Fr. have been observed only in the last century, in heather and woodland on sand. S. condensatum Hoffm. is found on old drifted sands in the interior, where it occurs now and is not rare locally. S. saxatile occurs on the same sites but is not common. The other, epilitic species were discovered in this century. S. dactylophyllum Flk. and S. evolutum Graewe grow on a few erratic boulders. S. pileatum Ach. and S. vesuvianum Pers. have been found several times recently, on artificial substrates. S. condensatum of the sands shows a somewhat aberrant form; the primary phyllocladia are usually less flattened, and often develop into branched structures of several mm length, see fig. 2, a. A key, brief descriptions and data on chemistry of the indigenous species are given.

Description: A syntaxonomic survey is given from the heathland communities in the Netherlands. The almost natural heathlands on coastal dunes belong to the Carici arenariae – Empetretum (Empetrion nigri) from dry or humid habitat and to the Empetro – Ericetum (Ericion tetralicis) from wet dune valleys. The antropogenous inland heaths belong to the Ericetum tetralicis (Ericion tetralicis) on wet peaty soils and to the Genisto – Callunetum and Vaccinio-Callunetum (Calluno-Genistion pilesoe) on humid to dry sandy soils. The latter association is confined to hilly districts with high precipitation. Ten new subassociations are described.

Description: A preliminary account is given of the parenchyma-like tangential bands, as seen on the transverse surface of Miconia species. The bands consist of parenchyma strands, fusiform parenchyma cells and fibres. These fibres differ from the fibres of the ground tissue in wall thickness, lumen diameter and sometimes in pit size. In the bands intermediate forms between parenchyma cells and fibres occur. The functional relationship between the elements forming the parenchyma-like bands is discussed. The systematic value of this phenomenon is still uncertain. In other genera of the Melastomataceae, however, as well as in other families of the Myrtales it has also been observed.

Description: The distribution of male, female and sporulating plants of Thamnobryum alopecurum( Hedw.) Nieuwl. [= Thamnium alopecurum (Hedw.) Schimp.] in the Netherlands is given in an attempt to clarify the problem, of why this species sporulates so seldom and in which environment sporophytes are formed most frequently. In places with air with a constantly high humidity the plants bear the largest numbers of inflorescences and in such places female and male plants also grow more strongly intermingled. In places with (periodically) drier air the plants remain mostly sterile or nearly so and develop into large sprouting systems with little contact between each other; even „moss balls” can occur then. Another important point is that in some localities only males or only females occur and in some one sex is very rare. The length of the seta of Thamnobryum alopecurum appears to be rather variable (10-30 mm). though within one specimen remarkably constant: mostly there is only 2-3 mm difference between the longest and the shortest seta per plant. Populations occur containing both plants with long setae and plants with short setae but without intermediate specimens. Elsewhere a continuous range has been found.

Description: In Note I it is shown that the delimitation of the sections Bryoidium, Pachylomidium, Aloma and Semilimbidium subsect. Bryolimbidium is very vague. Consequently they are united. The resulting section contains the type species of the genus Fissidens and the correct name for it is, therefore, Fissidens. In Note II F. bambergeri Schimp. ex Milde, F. firmus Linb. ex Roth, F. herzogii Ruthe in Herzog, F. canariensis Bryhn and F. bilewskyi Pot. Varde are reduced to synonymy under F. minutulus Sull. F. minutulus is described, figured and characterized. Besides, it is compared with F. ovatifolius Ruthe. F. pusillus (Wils.) Milde, F. minutulus Sull. and F. viridulus (Swartz) Wahl, are compared and found to be three distinct species. F. subimmarginatus Phil, is reduced to synonymy under F. exiguus Sull.

Description: Fossil pollen grains from the Quaternary of Colombia, formerly provisionally indicated as “Valeriana” stenophylla Killip, have now been identified as those of the Andean genus Lysipomia H.B.K. (Campanulaceae). In the genus Lysipomia s.l. (fide McVaugh) two considerably different pollen types are found: That of the Lysipomia s.s. and the Rhizocephalum type. The former was probably derived from the latter.

Description: Oil bodies in leaves variable in number, (2-)3-8(-13) per cell; subglobose to, more rarely, ellipsoid, 4-8(-12)x4—6 μm; colourless, rather coarsely to finely granulosepapillose, becoming homogeneous upon degeneration (Colombia, Cundinamarca, Cleef 78b, 3410b, 397b; ibid., Cauca, Cleef 679). Variation in number and morphology of the oil bodies apparently is characteristic for Lophocoleaceae, since it has been found not only in this and the other species of Lophocolea reported here but also in European material of Lophocolea and Chiloscyphus (unpubl. obs.).

Description: On 31 May 1976 Drs. A.J. Kostermans and C.E. Ridsdale, both of Leiden, set out on a 10 weeks botanical expedition to the Western Ghats in S. India as a summer holiday tour, partly financed by contributions from British Government Parliamentary Grant In Aid, administered by the Royal Society, London and the Treub Maatschappij, Utrecht. This support is most gratefully acknowledged. Part of the expenses were paid by the botanists themselves. Here follows their itinerary. The aim was to collect specimens of this area of the world where one of the earlier works on botany appeared: Rheede van Drakenstein, Hortus Malabaricus, 1678-1703. The area is greatly under-collected and many of the species are known only from a single collection. The area was studied by Bourdillon who was Conservator of Forests for over 30 years, but despite many efforts on his part he was unable to refind some 10% of the plants reported to occur in the area by Beddome and other previous workers. The later botanists collected in the hill stations and in the dry deciduous forest, the tropical evergreen forest, particularly its trees, being neglected.

Description: The Herbarium is arranged according to Christensen’s Index Filicum (1905). I examined all the Old World specimens in Dryopteris and Aspidium, both in the general herbarium and in the separate herbarium for China and Japan. It appears that the genera segregated by Christensen from Dryopteris (e.g. Pteridrys) in the Third Supplement to the Index (1954) have not been segregated in this herbarium. Type specimens have been removed by Dr. A. Bobrov and are in a separate series arranged alphabetically under basionyms. Selection of types was done rather hastily, and many of the specimens need critical study. For example, most Wallich specimens are filed as types of the names in Wallich’s catalogue, with no check as to whether the names have been subsequently validated. An additional complication is the fact that (especially in Thelypteridaceae) Wallich in several cases included specimens of two or three species under one number.

Description: The taxonomic study of this genus is, and always has been, severely hampered by the almost universal incompleteness of the material available in the herbarium. The vast majority of herbarium specimens, at least of the terrestrial species which comprise some 70% of the genus, consist of inflorescences alone, i.e. without vegetative parts. The aquatic species fare little better and although specimens are often more complete they are usually badly prepared so that vital characters such as the specifically characteristic branching of the foliar organs is completely obscured. Furthermore, in view of the very delicate nature of both the floral and vegetative parts, especially the complex and often very small traps, drying is rarely as satisfactory as preservation in liquid. This latter method, which has been increasingly used in recent years, can however produce relatively useless specimens if the incorrect liquid preservative is used. Having personally collected some 25% of the 180 known species both in Europe and the tropics of Africa and America the author feels qualified to offer some advice on the preparation of specimens of this genus for taxonomic study. 1) Dried specimens — Aquatic species should be put into a suitable receptacle (at least 0.5 m square) full of clean water about 15-20 cm deep. Gentle agitation will cause the vegetative parts to assume their natural position. A sheet of paper, either thin and suitably supported on a rigid sheet of metal or thicker and unsupported, is then carefully introduced beneath the floating plant and very gradually raised out of the water so that the plant eventually lies on the paper with all of its parts in their natural positions. The whole should then be dried as rapidly as possible in a ventilated press. Additional separate inflorescences (and infructescences) may of course be dried normally. Terrestrial species often have no very obvious vegetative parts as they are usually beneath the substrate. However it is the experience of the author that they are in fact almost always present and may be found by carefully removing a small piece of the substrate with the inflorescence. Gentle agitation in water may wash away the sand, soil or mud and these vegetative parts are again best displayed by ’floating out’ as for the aquatic species. If, as is frequently the case, the substrate is bound together by filamentous algae or the subterranean parts of other plants, the vegetative parts are difficult to separate and display, at least in the field, and in such cases the whole should be pressed.

Description: Index to J.J. Smith: Enumeration of the Orchidaceae of Sumatra (publ. in Fedde, Repert. 32, 1933, 130-386). — 32 pp. stenciled. Gratis. Index to C.L. Blume: Museum Botanicum Lugduno-Batavum vol. 2 (1856-1857). — 24 pp. offset. Gratis.

Description: Rhizome creeping or low-climbing (Bolbitis) or climbing ( Lomariopsis, etc.) or epiphytic (Elaphoglossum), dorsiventral, with a broad ventral vascular strand which supplies the roots and one or more dorsal strands (the fronds in two or more longitudinal rows, according to the number of strands); stipes jointed to rhizome (Teratophyllum, Elaphoglossum) or not, containing several separate vascular strands; scales peltate or pseudopeltate, clathrate or not; no elongate unicellular hairs. Rhizomes of young plants always with one dorsal meristele, this condition persisting to the adult plant in Teratophyllum and many species of Elaphoglossum. Fronds simple ( Elaphoglossum, Bolbitis spp.), pinnate (all but Elaphoglossum) or bipinnate ( Teratophyllum and Lomagramma spp.), the pinnae on fronds of Lomariopsis, Teratophyllum and Lomagramma jointed to the rachis, terminal unjointed lamina present in Lomariopsis; distinctive bathyphylls, usually more dissected than acrophylls, present in genera with climbing rhizomes (least distinctive in Lomariopsis); veins free ( Teratophyllum, Lomariopsis, most Elaphoglossum, some Bolbitis) or uniting near the margin ( Elaphoglossum spp.) or in several series of areoles with (most species of Bolbitis) or without ( Lomagramma; Bolbitis p.p.) free veins in the areoles. Fertile fronds with reduced lamina, covered beneath (rarely also above) with sporangia (except Thysanosoria, where sori are at ends of veins only), a special vascular supply for the sporangia variously developed or not; spores with perispore (except Lomagramma). Genera. Bolbitis SCHOTT, Lomariopsis FÉE, Lomagramma J.SM., Teratophyllum METT., Thysanosoria GEPP, Elaphoglossum J.SM.; also Peltapteris LINK (Rhipidopteris FÉE ex SCHOTT) and Microstaphyla PRESL, small genera of tropical America and St Flelena, allied to Elaphoglossum and not dealt with in the present work.

Description: Newly discovered mycorrhizal relationships of boletes with Sapotaceae and Nyctaginaceae in the Neotropics are discussed. The eight neotropical species of Phylloporus are keyed out and three described. Fistulinella Henn. is transferred to the Strobilomycetaceae. Phylloporus manausensis Sing. and P. sect. Manausenses Sing., P. leucomycelinus Sing., Xerocomus amazonicus Sing., X. radicicola Sing. & Araujo, Tylopilus arenarius Sing., T. potamogeton Sing., T. sect. Potamogetones Sing. sect. nov., Fistulinella campinaranae Sing. and Porphyrellus rionegrensis Sing. & Araujo. are new taxa.

Description: Until recently the mycoflora of the West Frisian Islands has been rather neglected. In the last few years, however, members of both the Rijksherbarium and the Netherlands’ Mycological Society (N.M.V.) have been collecting more frequently on these islands. The results of their fieldtrips are promising for the future as many interesting and rare fungi have been found. In a previous paper (Noordeloos, 1975) I described a rare species of the genus Marasmiellus Murr., viz. M. caespitosus (Pat.) Sing. from the Island of Texel¹. I now want to introduce, from the same island, another member of the same genus.

Description: Neosprucea kuhlmannii Sleumer, Lilloa 23 (1950) 248.

Description: Cerastium papuanum Schltr. ex Mattfeld (1929) and C. keysseri Schltr. ex Mattfeld (1929) are united under the name C. papuanum Schlechter ex Mattfeld emend. Möschl, as they form one series of reduction, which starts with macropetalous and pentamerous tall-growing forms (subsp. phaenops Mattf., at about 1770 m and above) and ends with micropetalous and tetramerous low-growing cushions [subsp. keysseri (Schltr. ex Mattf.) Möschl] in the regions around the summits up to 4700 m in New Guinea. A glandular form of this species (var. eciliatum f. glandulosum Moschl) is described for the first time. Plants with few-flowered inflorescences with foliaceous bracts are described as var. dispersiflorum Möschl. A table shows petals, placentas, and other organs of this species for the first time. The possible relationship with C. octandrum Hochstetter ampl. Moschl in the peak regions of tropical Africa is suggested. Mention is also made of Uredo morobensis Cummins, which is the only parasite so far known.

Description: The new species Roureopsis confudens Leenh. is described from Thailand

Description: Mischocarpus Blume, Bijdr. (1825) 238, nom. cons.; Rumphia 3 (1849) 166; Radlk., Pfl. R. Heft 98 (1933) 1288—1310. — Cupania § Mischocarpus Miq., Fl. Ind. Bat. I, 2 (1859) 566. — Type: M. sundaicus Bl. Pedicellia Lour., Fl. Coch. (1790) 655, nom. rejic. (see under dubious names). — Type: P. oppositifolia Lour. Mischocodon Radlk., Bot. jahrb. 50 (1913) 79; Pfl. R. Heft 98 (1933) 1327—1328. — Type: M. reticulatus Radlk. Shrubs or, sometimes large, trees, sometimes with a slender unbranched stem. Buttresses sometimes present (M. largifolius). Indumentum rather dense to sometimes very sparse, consisting of mostly appressed, short to long, brownish to ferruginous hairs; no glandular scales. Twigs brownish to reddish-brown to greyish. Axillary buds just above or, mostly in ramiflorous species, up to 6 mm above the base of the petiole. Leaves spirally arranged, paripinnate, the leaflets accrescent in size towards the top, 1—6-jugate, without stipules; petiole ± semi-terete, sometimes dorsiventrally flattened. Leaflets alternate to subopposite, petioluled, ratio 1.5—5(—8), widest below, in, or above the middle, sometimes curved downwards (in the herbarium showing a folded base and apex and sometimes an undulate or folded margin), sometimes bullate, pergamentaceous to coriaceous, when dry above mostly greyish-green, sometimes smooth and shiny, beneath mostly brownish-green, not papillose, above glabrous or hairy on midrib and nerves, beneath glabrous or hairy mainly on midrib, nerves and along the margin, between the nerves very sparsely appressedly short-hairy, often glabrescent; domatia often present in axils of main nerves; base equalsided, rarely slightly oblique, rounded or acute to blunt, decurrent; margin entire, flat or sometimes revolute; apex rounded or acute to blunt, mostly shortly mucronate, or ± acuminate or retuse or rarely emarginate; acumen rounded or acute, mostly slightly retuse; midrib above prominent to sunken, rounded or angular, sometimes carinate, beneath prominent, in cross-section about semi-circular, sometimes nearly completely circular ( M. grandissimus), slightly angular to the base; nerves not or sometimes indistinctly connected in the lower 0.5—0.75, in the upper part connected, about straight to rather strongly curved; intercalated veins present, sometimes indistinct; veins and veinlets nearly always forming a very regular reticulate pattern, dense; nerves, veins, and veinlets ± prominent on both faces, beneath stronger so than above, veinlets inconspicuous beneath. Inflorescences pseudoterminal, axillary, and ramiflorous (probably also cauliflorous in a few species), composed of one or more thyrsoid axes, these nearly always branched, erect to spreading, mostly slightly grooved, with stalked or sometimes sessile cymules, glabrous to densely hairy; cymules 1—7(—10)-flowered; pedicels 1—3(—5) mm; bracts triangular to lanceolate, sometimes subulate, outside glabrous or hairy, inside mostly glabrous. Flowers unisexual, probably mostly monoecious (dioecious in M. reticulatus?). Calyx spreading or cup-shaped, early expanding, 5 (rarely 6)-merous, connate for up to 65%, membranaceous to subcoriaceous, sometimes somewhat fleshy; lobes subequal, sometimes slightly imbricate at the base, triangular to ovate, outside variably hairy, inside glabrous or hairy, often only a row of hairs near the base sometimes hidden by the disk; apex acute, sometimes acuminate. Petals 0—5, from minute up to slightly longer than the calyx, apert, unguiculate or not, variably hairy, mostly on claw, base of plate, and auricles; plate elliptic to ovate, sometimes triangular or rhomboid; apex sometimes lobed; 2 auricles or scales mostly present, without crest. Disk complete or sometimes interrupted, annular or cup-shaped, sometimes surrounding base of stamens and confluent with pistil, glabrous or short-hairy. Stamens (5—)8(—9), exserted (sometimes rather long); filament thread-like, glabrous or appressedly to patent-hairy, more densely so to the base; anther basifixed, base and apex emarginate; connective sometimes with a lighter coloured wart at the top; thecae about ellipsoid, glabrous or sparsely hairy, smooth or papillose (most distinct when not yet exserted), dehiscence lateral or latero-introrse. Pistil 3-(rarely 2- or 4-) celled, glabrous or appressedly short-bairy; ovary stiped or almost sessile, about ellipsoid- to obovoid-triangular; style apical, shorter to slightly longer than ovary, the upper part either split in 3 ± recurved stigmatic lobes or almost undivided, bearing 3 stigmatic lines (M. exangulatus); ovules 1 per cell, apotropous, anatropous, ascending, base collar-like, surrounding micropyle and funiculus. Pistillode small, densely hairy to subglabrous. Infructescences sometimes with accrescent axes and pedicels; calyx present, sometimes accrescent, mostly glabrescent; disk present, not accrescent. Fruit nearly always distinctly stiped (in M. paradoxus only up to 1 mm), not lobed, the cells about equally developed but the ovules abortive in (1) 2 cells, loculicidal, up to 3.5 cm long, reddish when ripe, glabrous or hairy; stipe empty, 3-celled, cylindrical near the base, distally becoming triangular; seed bearing part triangular to rounded in cross-section, with elliptic to obovate valves, apiculate; valves thin to almost woody, mostly shrivelled after dehiscence; pericarp slightly fleshy; endocarp sclerenchymatic, either complete, lining valves (except for M. exangulatus also lining stipe) and distal parts of the septa, or incomplete, only along the sutures (see fig. 1g and 1h); septa membranaceous, at least in the proximal half; endocarp and septa glabrous or variably hairy. Seed sometimes pendulous by the appendix of the arillode, globose to ellipsoid; hilum adaxial, basal; testa shining, chestnutbrown, finally (nearly) completely covered by a thin-fleshy, translucent, bluish or yellow to orange arillode which is attached around hilum and micropyle; arillode nearly always (except M. paradoxus) with an appendix abaxial of the hilum and micropyle, descending into the stipe; cotyledons equal or not, folded or not; suture between cotyledons either transverse and straight, or curved (see fig. 1e and 1f). Plumule glabrous or variably hairy (not constant within one species).

Description: In the genus Phylacium 2 species are recognized. Special attention is paid to the morphology of the inflorescence; full descriptions are given with plates and a map, showing the distribution of both species.

Description: In 1969 I published, together with P. G. Heinsbroek, a paper on the anatomy of the stamens of Victoria amazonica. The flowers used in that study came from plants which were cultivated in the green-house of the Leiden Botanic Garden. Because the possibility could not be excluded that the structures then observed were partly the effect of greenhouse conditions, I subsequently took the opportunity to study flowers of well developed plants, which were cultivated in the open air under the tropical conditions of the Botanic Garden at Bogor*). The results were exactly the same. Apart from the set of central vascular bundles, normal in laminar structures, there proved to be a peripheral sheath of bundles consisting of abaxial bundles, terminating half way up the stamen, as well as adaxial bundles. All bundles run parallel and the central ones branch upwards into the fertile region of the stamen. The adaxial bundles anastomose at the lower end of the pollen sacs and one of the resulting anastomoses pursues its course in the middle between the thecae. In literature the last mentioned vascular bundle had been named the ‘auxiliary vein’ (Moseley, 1958). Its position is opposed to the normal median vein, and its xylem pole is inverted. This vein played a role in diverse morphological opinions on the flat stamens. American authors (Eames, 1961), who advocated the primitiveness of laminar stamen structure, disposed of the auxiliary vein by considering it as an insignificant vein. Schneider (1976) thinks the peripheral bundle system is explicable in functional terms. On the other hand, the discovery of the opposed median auxiliary vein was welcomed by authors like Leinfellner (1956), who thought, mainly on the ground of teratology, that the stamens are diplophyllous structures, that is consist of a dorsal and a ventral blade fused medianly. By this view the existence of apparently homogeneous laminar stamens in Ranales had been difficult to explain. However, now the auxiliary vein could be considered as the median vascular bundle of the fused ventral blade, as requested by the theory. Meeuse (1972) took up the suggestion brought forward in our paper of 1969, namely that the stamen vasculature consisted of a bract component (the abaxial bundles) and a flattened axis component (the central and adaxial bundles), in analogy with the Coniferous female cone scale. According to Meeuse this is proof of his thesis that the structure of the stamen in the Ranales is a bract amalgamated with an axial system. However, careful comparison with the results of Cécile Lemoine-Sebastian (a.o. 1972) show that the vascular patterns are different from a situation as described above.

Description: The pollen morphology of the nine taxa within the genus Pometia, the three subspecies of Litchi chinensis, and the species Cubilia cubili and Otonephelium stipulaceum (Sapindaceae — Nephelieae) was studied. Three pollentypes were found: 1) the Pometia pollentype, which occurs in all nine taxa of the genus; 2) the Litchi pollentype, which can be divided in two subtypes: a) characteristic for Litchi chinensis subsp. chinensis. subsp. phillippinensis, and Otonephelium and, as could be concluded from a provisional investigation, also for Xerospermum and Nephelium, and b) characteristic for Litchi chinensis subsp. javensis; 3) the Cubilia pollentype. Possible evolutionary trends in the tribe Nephelieae are discussed

Description: A comprehensive study of floral and vegetative anatomy of the monotypic New Caledonian genus Paracryphia Baker was initiated in an attempt to help clarify the evolutionary relationships of the genus. Detailed descriptions of leaf, axis, nodal, wood, floral, pollen, and fruit morphology and anatomy are presented. In general, most vegetative characters are distinctly primitive whereas those of the reproductive organs are regarded as advanced or specialized. The present study confirms the opinion that Paracryphia merits familial status. All previously suggested relationships of the genus are rejected in favor of a view that envisions Paracryphia as an independent and early divergence from the Thealean, Ericalean, and/or Celastralean lines of evolution. This view is based on similarities of Paracryphia with Sphenostemonaceae, Actinidiaceae and Theaceae in a number of characters.

Description: La faune isopodologique littorale des Antilles est peu connue et les études systématiques sont rares. Nous pouvons signaler, parmi les principaux travaux, ceux de MOORE (1901) et de MENZIES & GLYNN (1968), tous les deux sur Porto Rico; celui de RICHARDSON (1912) sur la faune terrestre et marine de Jamaïque et celui de GLYNN (1970) sur les Sphaeromidae de l’île Margarita (Vénézuéla). Les études de GLYNN (1968) sur les isopodes associés aux chitons de Porto Rico et celui de MENZIES (1957) sur les Limnoriidae de l’Amérique du Nord et Centrale sont aussi des contributions importantes. La faune isopodologique des Bermudes, elle, a été étudiée par RICHARDSON en 1902.

Description: Several rocktypes and their metamorphic mineral growth are described from an area on the western border of the Sesia-Lanzo Zone in the Orco Valley (NW Italy). It is argued that in some rocks (garnet-rich gneisses and micaceous gneisses) pre-Alpine metamorphic minerals are present, in other rocks (carbonate-bearing schists, albite-chlorite gneisses) such minerals are rare or absent. For the latter rocks it is therefore difficult to establish whether they are strongly retrograded Alpine basement rocks, or rocks belonging to the suite of ophiolitic schistes lustrés. The two possibilities are discussed.

Description: So long as the lithologic nature of the cliff and the physico-chemical nature of the sea water remain similar, variation of cliff morphology is largely a function of water turbulence (degree of exposure). Different cliff profiles (without, or with one or two notches; without or with surf platforms) are all part of a continuous range of variation.

Description: Bijlage 2, bij het doctoraalverslag van E.J. van Nieukerken

Description: Verslag van een doctoraalonderwerp van 12 maanden bij de vakgroep Oecologie, Rijksuniversiteit Leiden, 1974-1977

Description: This paper is the second (Florschütz & Florschütz-de Waard 1974) in the series of reports on cryptogams of Colombia, especially the high Andean bryophytes and lichens, in the framework of recent phytosociological and ecological studies in the area by A. M. Cleef and T. van der Hammen (Amsterdam) and his collaborators. The aim of these studies is to prepare comprehensive descriptions of the various Andean biota of Colombia as a basis for an evaluation of the biological diversity of the Andean environment of Colombia. This paper deals with the Hepaticae or liverworts and focuses on the structure of the oil bodies in the liverwort cell and the geographical and ecological distribution of the species. There are few papers on tropical Andean liverworts. The classical work by Spruce (1884-85) included liverworts of the Ecuadorian and Peruvian Andes. A large number of Andean taxa are treated in Fulford’s Manual of the leafy liverworts of Latin America (Fulford 1963, 1966, 1968, 1976) and the latter has been our main source of data on distribution of species. Some ecological notes on tropical Andean taxa are found in the works of Herzog (1934, 1955), Robinson (1967) and Winkler (1976), who for the first time presented a synthesis of liverwort species distribution in relation to Andean vegetation typology in a limited area in Northern Colombia.

Description: The present paper is the result of a study of Dorstenia mainly made on behalf of the Floras of Cameroun and Gabon. It presents, as a precursor to the treatment in those Floras, some general remarks on the genus Dorstenia, a key to the species, a survey of the taxa distinguished, new combinations and new taxa, the synonymy (as far as based on study of types), and distribution.