ALBERT

Notes: Measurements of rates of growth and senescence of leaf lamina per tiller and of changes in tiller population densities were made in three experiments designed to investigate the influence of sward slate on leaf turnover and net production under continuous stocking.In each experiment initially uniform swards were fenced to provide four plots on which animal numbers were adjusted twice weekly to give a series of swards maintained as nearly as possible in a steady state with respect to sward surface height (range 1.1–6.4 cm) and herbage mass (range 440–2690 kg OM ha−1). Two experiments were carried out in July–September on vegetative swards and one in May–June on a reproductive sward. Measurements were begun 3–7 weeks after treatments were started and were repeated weekly during 3–4 week measurement periods.In all three experiments the rate of lamina growth per tiller increased linearly with an increase in sward surface height and herbage mass. In the two experiments conducted in July–August this relationship was partially offset by a linear increase in the rate of senescence per tiller but net production per tiller also increased linearly in relation to sward height and mass. In the experiment conducted in May–June the rates of growth and senescence per tiller increased in parallel so that net production per tiller showed no relationship with sward condition.Tiller population densities in the July–August experiments were highest in swards maintained between 2 and 3 cm surface height and declined in swards maintained above and below this height. In the experiment in May–June tiller numbers were similar in all treatments prior to the summer solstice but diverged in a manner similar to the other experiments later in the year.The rate of lamina growth per unit area increased in a manner consistent with an asymptotic relationship and the rate of senescence increased linearly with increasing sward height and mass in all three experiments. Net production per unit area was reduced on swards below about 2.5 cm in height but was insensitive to variation in sward surface height between 2.5 and 6.0 cm (approximately 1000–2500 kg OM ha−1 herbage mass).The effectiveness of adjustments in tiller numbers and production per tiller and of changes in the balance between growth and senescence as mechanisms of sward homeostatis, together with their implications in the scope for manipulating herbage production by grazing management are discussed briefly.

Notes: There was a three-fold variation in epicuticular wax content between populations of white clover and a five-fold variation between populations of red clover. The mean for all populations of red clovers was twice that of white. There was also a two- to three-fold variation within each population.The epicuticular wax content of white and red clovers grown in the field showed an increase with age. Thin layer chromatography showed no qualitative differences in the epicuticular waxes extracted either between populations or with sampling date.The higher levels of epicuticular wax found in some clovers were considered to be of possible significance as a source of antifoaming compounds in bloat-inducing pastures.

Notes: Differences in ease of digestion of bundle sheath cell walls of leaf blades of Panicum spp. are shown to be associated with differences in bundle sheath cell wall structure, including the presence or absence of a suberized lamella. These structural differences are correlated with photo-synthetic type (viz. C3, intermediate C3/C4, and the C4 types, PCK, NADP-ME and NAD-ME), as are mesophyll: bundle sheath area ratios.

Notes: The construction of a transducer and the use of a recording system to measure jaw movements of grazing sheep is described, The time spent grazing, ruminating and idling may be measured by this equipment and the data produced automatically processed using a microprocessor which determines each minute whether the animal is grazing, ruminating or idling and also summarizes these data to give total times. The accuracy of this technique in relation to manual observation is discussed.

Notes: The effect of two heights of cutting (5 and 8 cm) on the dry matter harvested and persistency of a range of different grass species and varieties were monitored over a 3-year period (1976 to 1978). Dry matter harvested was measured in the second and third harvest year and persistence of sown grass was assessed at the end of the first and third harvest year. The grasses were managed under a frequent cutting system, i.e. simulated grazing. Results showed that at the low cutting height tetraploid hybrid ryegrasses and diploid Italian ryegrasses in the third harvest year gave lower annual dry matter harvests of sown grass and were less persistent than at the high cutting height, whilst, in contrast, perennial ryegrasses gave higher annual harvests of sown grass but persistency was unaffected. In general at the low cutting height varieties of cocksfoot, meadow fescue and tall fescue gave comparatively higher annual harvests of sown grass in the second harvest year but lower yields of sown grass in the third harvest year. A notable exception was Cambria cocksfoot which in the second harvest year gave higher yields of sown grass and was more persistent at the low cutting height.However, at 5 cm cutting height, the proportions of weed grasses (mainly Poa spp.) in swards sown to tetraploid hybrid ryegrasses and diploid Italian ryegrasses were generally greater, particularly in mid-season.

Notes: Laminae of Lolium perenne and Phleum pratense were tested mechanically and the results analysed using engineering composite theory and fracture mechanics. As the lamina dries the fibres double in stiffness. The intervening cells show a sevenfold increase in stiffness below 20% water content (based on dry weight). Work to fracture across the veins is almost independent of water content, but below 50% water content the mode of fracture changes. It is predicted that total fibre content will not affect hay shatter; total protein content may affect it by altering the water-binding properties.

Notes: Grass silages made from first-harvest perennial ryegrass in mid-May and early-June and termed high-D and low-D respectively had mean DOMD values of 694 and 633 g kg−1 and were offered ad libitum to twelve Ayrshire cows in a 16-week feeding experiment using a cyclic changeover design. The silages were supplemented with a concentrate containing 363g CP per kg DM at daily rates of 1.5, 3.0 and 4.5 kg per cow. The average daily intakes of silage DM were 12.6 and 11.1 kg per cow on the high- and low-D silage treatments respectively, and were not significantly different on the three concentrate treatments. The mean daily milk yields were 171, 18.6 and 21.0 kg per cow on the low-D silage treatments, and 19.6, 21.2 and 22.8 kg per cow on the high-D silage treatments at the 1.5, 3.0 and 4.5 kg rates of concentrate feeding respectively. Fat and lactose concentrations in the milk were not affected significantly by treatment, whereas the CP and SNF concentrations increased progressively and significantly as the rate of supplementary feeding increased. It is concluded that a high-protein concentrate supplement allows silage to make the maximum contribution in the diet, and gives a high response in terms of milk yield and composition.

Notes: Grass which had grown without interruption from late April was cut as for hay in early July in each of 2 years. Fifteen dates of removal of the cut herbage were compared ranging, at 1-d intervals, from the day of cutting to 14 d after cutting. Sward regrowth was studied.A delay of 5 d in removing cut herbage reduced herbage dry matter harvested during the remainder of the growing season by 9% and a delay of 10 d reduced yield by 16%. Delaying the removal of cut herbage reduced the density of grass tillers considerably in the short term, but the effect had worn off in 8 to 12 weeks. The presence of cut herbage reduced the rate of extension of leaf blades considerably. Delaying the removal of cut herbage reduced the area per leaf blade and the weight per unit area and these effects persisted for several weeks. Grass growing under cut herbage had a less negative leaf water potential and a rather cooler but much darker environment in which to grow than uncovered grass.

Notes: Book reviews in this article: Milk from Grass Edited by C. Thomas and J. W. O. Young Nitrogen Fixation in Legumes Edited by J. M. Vincent Herbage Intake Handbook Edited by J. D. Leaver Man's Proper Study By Richard J. Colyer Introduction to Sheep Fanning By R. G. Johnston

Notes: Information about the grazing behaviour and the return of dung to pasture by lactating dairy cows was collected from stocking rate experiments in northern Victoria. Grazing behaviour was observed for 24-h periods in mid-summer in two years. Grazing time increased as herbage allowance decreased to about 32 kg DM cow−1 d−1 but as herbage allowance decreased further grazing time also decreased. A herbage allowance of 32 kg DM cow−1 d−1 corresponded to a stocking rate of about 5.5 cows ha−1. This effect of herbage allowance on grazing time may have been confounded by herbage mass, however. Rumination time of the cows increased by 003 h for each kg increase in herbage allowance while resting time was not affected by treatment.The effects of stocking rate on some of the characteristics of faecal output were measured for a 3-d period in mid-summer. The number of pats deposited per cow daily declined by 0.66 for each unit increase in stocking rate. The fresh weight of dung also declined as stocking rate increased by 0.16 kg per unit of stocking rate. While the values for the amounts of dung deposited on the pasture by the cows at the lower stocking rates are similar to many of those reported in the literature, this study has quantified the way in which stocking rate may influence this in one instance. Furthermore, provided that some measure of herbage intake is made when measurements of dung excretion are performed, it is suggested that estimates of in vivo digestibility can be obtained.

Notes: Two red clover (Trifolium pratense) cultivars, Red Head (tetraploid) and Kuhn (diploid), were sown at a seed rate of 13 kg ha−1 either alone or in mixture with Italian ryegrass (Lolium multiflorum) cv. RvP sown at seed rates of 10, 15, 20 or 30 kg ha−1. RvP was also sown alone at a seed rate of 30 kg ha−1 and received nil or 300 kg ha−1 fertilizer a−1 fertilizer N. All plots were established using the barley cultivar Midas sown at a seed rate of 100 kg ha−1 as a nurse crop.Neither clover cultivar nor ryegrass seed rate significantly influenced either dry matter harvested or botanical composition over the 3 harvest years. On average over all years the grass-clover mixtures produced 75% of the yield of the N-fertilized RvP, 125% of the clover monocultures and 225% of the unfertilized RvP. The red clover contribution to the total dry matter harvested of the mixtures averaged 45–60%. The dry matter concentrations of the mixtures were considerably higher than those of the pure clover stands. In the third year yields were markedly reduced in comparison with those in the first and second years.It was concluded that Italian ryegrass can be a suitable companion grass for red clover. Its superior yielding capacity over other grasses such as perennial ryegrass or timothy under a conservation management can be coupled to advantage with red clover to give a sward which Is essentially stable, at least over a 2- to 3-year cropping period, although giving slightly reduced yields in the third year. Italian ryegrass-red clover mixtures, without the use of fertilizer N, can produce high DM yields of good quality herbage.

Notes: Red clover cv. Hungaropoly was slot-seeded into a perennial ryegrass-dominant sward in April 1979. Glyphosate and paraquat were applied separately as bandsprays each at two doses and at two band widths. Control plots were either slot-seeded without a herbicide bandspray or received ±150 kg N ha−1 a−1. Red clover establishment was assessed and amounts of dry matter (DM) and total N accumulated were measured at two harvests in 1979 and three harvests in 1980. Bandspraying increased seedling vigour and development and resulted in the eventual replacement of 1 t grass DM ha−1 by an equivalent amount of red clover. Of the treatment variables investigated, bandspray width had the greatest influence on red clover establishment and productivity. The slot-seeded area, meaned for all treatments, produced a total of 6.40 and 13.16 t DM ha−1 in 1979 and 1980. This was estimated to be equivalent to the all-grass sward receiving 112 kg N ha−1 a−1 during the second year of the experiment or 238 kg N ha−1 over the 2 years when measured in terms of N yield. Slot-seeding overcomes several of the problems associated with conventional establishment of red clover.

Notes: Two experiments were conducted to determine the effect of pre-harvest treatment by formic acid spray on the moisture concentration (MC) of perennial ryegrass (Lolium perenne) in vegetative growth. In Experiment 1 aqueous solutions containing 0.080, 0.148 and 0.259 kg formic acid per litre were applied to a perennial ryegrass sward (fresh matter yield 21.0 t ha−1) at a constant volume of 41.2 litres t−1to give active ingredient application rates of 3.5, 6.0 and 11.5 kg t−1 respectively. In Experiment 2 similar solutions were applied to a perennial ryegrass sward (fresh matter yield of 6.9 t ha−1) at 35.2, 19.0 and 10.9 litres t−1 respectively to give a constant formic acid application of 2.8 kg t−1. The changes in MC and chemical composition after treatment with acid were compared with those in untreated plotsIn both experiments treatment with acid significantly reduced the MC (P 〈 0.001) within 5 h of treatment from 4.4 to 3.0 kg per kg of dry matter (DM) in Experiment 1 and from 3.0 to 2.6 kg per kg DM in Experiment 2. There was no effect of level of acid applied in Experiment 1 or of the volume of water applied with the low level of acid in Experiment 2 on the changes in MC of acid-treated plots. There was however a significant (P 〈 0.00l) difference between the MC of plots treated with acid and those untreated which was maintained in Experiment I for up to 9 d even during rainfall.There were no differences between the chemical compositions of samples from acid-treated plots in either Experiment I or 2 or between untreated and acid-treated plots in Experiment 2. However there were significant differences between untreated and acid-treated plots in Experiment 1 for the content of nitrogen (N), water-soluble carbohydrate (WSC) and digestible organic matter (DOM). There were changes in the content of N, WSC and DOM with time in both experiments. In Experiment I there was an interaction between sampling time and treatment for both N and WSC but not for DOM. Compared with untreated plots, treatment with acid at all levels accelerated the loss of WSC and delayed the fall in N content. There were no such interactions in Experiment 2.

Notes: The main object of the experiment was to assess the effect of the relative proportion of non-protein nitrogen (NPN) to total nitrogen in silage on digestion in the sheep. Four unwilted perennial ryegrass silages were made with the addition of formic acid at 0, 2·2, 4·2 and 5·2 litres t-1 to provide foods with NPN proportions reducing from 0·26 to 0·20 of the total N. The digestion of the silages was studied in a 4 × 4 Latin Square experiment with sheep cannulated in the rumen, proximal duodenum and terminal ileum.Results for organic matter (OM), cellulose and N showed no major difference between silages in their digestion in the rumen, small intestine and caecum and colon, though small differences (P 〈0·25) in rumen fermentation pattern and in the proportion of digestible OM disappearing in the small intestine were observed. Concentrations of ammonia N in the rumen and rates of rumen bacterial protein synthesis did not differ significantly between silages and there were no treatment effects on the passage of individual amino acids to the small intestine. The results indicate that the proportions of NPN to total N in the silages examined had little influence on the efficiency of silage N utilization in the rumen or on the passage of undegraded dietary protein to the small intestine.

Notes: Two experiments were carried out to determine whether the differences in intake and feeding value previously shown between two grass cultivars when offered to ruminants as chopped artificially dehydrated (dried) material could also be demonstrated when the grasses were offered in other forms. Two cultivars of tetraploid Italian ryegrass (Lolium multiflorum), Sabalan and Tetila, were established in the same field in 1975. In 1976 they were grazed and conserved (two cuts of primary growth) as dried material or as silage.The composition of the herbage selected at pasture and conserved showed higher concentrations of normal detergent fibre and acid detergent fibre in the dry matter for Tetila than for Sabalan, but differences between cultivars in digestibility were small. In both grazing and winter feeding trials differences in voluntary intake between the two cultivars were not significant, but at pasture young beef cattle spent less time grazing and tended to spend more time ruminating on Tetila than Sabalan. There was no measure of animal performance at pasture but liveweight gain was 15% higher for Sabalan than Tetila when both were offered to young beef cattle as the sole feed of dried grass or of silage. The voluntary intakes of the three forms of feed were very similar, which in part reflected a similarity in digestibility. However, gains were lower for calves given silage than those given dried grass. This may have been due to a lower efficiency in the utilization of the nitrogeneous components of silage for tissue growth than those of dried grass.

Notes: Four grass silages, all made in mid-July from second-harvest perennial ryegrass swards, were compared in a 16-week feeding experiment with twelve Ayrshire cows. Two silages were unwilted and two wilted. All the silages received formic acid (‘Add-F’) at the rate of 3 litres t-1 either with formalin at the rate of 1 litre t-1 or without formalin. The unwilted and wilted silages had mean dry matter (DM) concentrations of 200 and 243 g kg-1, and in vitro D-values of 0·293 and 0·272 respectively. The silages were offered ad libitum plus 6 kg concentrates per cow per day. The daily intakes of unwilted and wilted silage DM were 10·2 and 9·2 kg per cow respectively on the formic acid treatment, and 10·2 and 9·2 kg on the formic acid + formalin treatment. The mean daily milk yield on the unwilted silage treatments was 19·2 kg per cow which was significantly higher than the yield of 17·2 kg per cow on the wilted silage treatments. The formalin had no significant effect on milk yield. The four silage treatments had small and non-significant effects on milk composition. It is concluded that the unwilted silages, which had excellent fermentation characteristics, were superior to the wilted silages as a feed for dairy cows.

Notes: Book review in this article Forage Evaluation: Concepts and Techniques Edited by J. L. Wheeler and R. D. Mochrie Improved Feeding of Cattle and Sheep By P. N. Wilson and T. D. A. Brigstocke Nitrogen Cycling in West African Ecosystems Edited by T. Rosswall Land Evaluation By S. G. McRae and C. P. Burnham

Notes: Three field experiments showed that perennial ryegrass can be successfully slot-seeded into an existing Festuca-Agrostis sward. Ryegrass survival was satisfactory in all cases but was improved on a low fertility site by the application of fertilizer N, P and K in the slot at sowing.The effect of varying inter-row spacing from 37·2 to 15 cm was measured in a 3-year experiment. In year one, involving seven cuts, slot-seeding increased total herbage dry matter harvested by a mean value of 17%; inter-row distances of 22·2-30 cm gave the optimal combination of ryegrass + old sward herbage. Differences in yield between inter-row spacings declined in the subsequent two years, as the rows of ryegrass thickened. Ryegrass digestibility (measured only in year two, from six cuts) was higher than that of the old sward; total metabolizable energy harvested from 15-cm rows was 20% higher than that from unsown controls. Increasing N input from 200 to 400 kg ha-1, starting in year two, only produced a significant increase in total herbage harvested in year three but proved effective in increasing the ryegrass contribution from the wider spaced rows.

Notes: In three separate feeding experiments using a total of thirty individually-housed Ayrshire cows three silages made from perennial ryegrass were given ad libitum together with supplements of four different hays in the long form. The in vitro D-values of the silages ranged from 0·298 to 0·283, and the hays from 0·280 to 0·200. The daily intake of hay DM varied from 0·2 to 4·2 kg per cow and was given either without or with a daily maximum of 2·2 kg concentrate DM containing 379–527 g CP per kg DM. On average, 1 kg hay DM decreased silage intake by 0·24 kg DM with a range of 0·21–1·20 kg. The hay supplements had only small and non-significant effects on total DM intake, milk yield and milk composition, but increased the daily intake of drinking water. In three behavioural studies, the eating and ruminating times expressed as min per kg DM did not differ significantly between the various supplement treatments. It is concluded that hay has only a marginal value as a supplement for grass silage, although the hay could serve as a useful ‘buffer’ feed if the amount of silage was limited.

Notes: Two areas of an early-heading perennial ryegrass cv. Cropper were harvested by either a precision-chop or a flail harvester at around 50% ear emergence (15 May 1978) and 14d later (29 May). Formic acid (85%) was applied at the rate of 2·2 litres t-1. Mature crossbred wethers were used in a 2 × 2 factorial design to determine the effect of stage of maturity and method of harvesting (chop length) on the in vivo digestibilities of formic acid-treated grass in experiment 1 and formic acid silage in experiment 2. Apparent digestibility coefficients were determined at a fixed level of feeding for both grass and silage and at ad libitum access to feed for silage only.There were no significant differences in the concentrations of crude protein (CP), ether extract (EE), acid-detergent fibre (ADF) or acid-detergent lignin (ADL) in grass or silages of differing chop lengths but the later cut forages had significantly higher ADF and ADL concentrations and lower CP concentrations than the early-cut forages. The ADF and ADL concentrations were also higher in the silages than in the corresponding grasses.In general, the fermentation characteristics of precision-chopped silage were better than for the corresponding flail-cut silage but date of harvest was a more important determinant of quality and the late, flail-cut silage had the highest butyrate and ammonia N concentrations and the highest pH (411) of any treatment. There was a significantly higher intake of precision-chopped as compared with flail-cut silage with both the early-and the late-cut silage but there were no significant differences attributable to stage of maturity (i.e. date of harvest) or significant interaction between chop length and maturity. The slightly increased intake of early harvested, precision-chopped silage as compared with late precision-chopped silage was not significant.Dry matter digestibility (DMD) of the grass decreased at a rate of 0004 units d-1 post 50% ear emergence. The results of experiment 2 indicated a decrease of 0·207 units d-1 in silage fed at a similar level. The late-cut silage (DMD 0·292, mean of both harvesting treatments) thus had a significantly lower digestibility than the corresponding grass (mean DMD 0·247). Chop length had a variable influence on the DMD of both grass and silage fed at a fixed level but treatment differences were non-significant. However, a trend towards higher digestibility of flail-cut as compared with precision-chopped silage was apparent and this became statistically significant when the animals were allowed ad libitum access to feed. This may be a response to the generally lower intake of flail-cut silage.

Notes: Book review in this article Weed and Pasture Management in South Africa Edited by N. M. Tainton Collecting and testing tropical forage plants Edited by R. J. Clements and D. G. Cameron Herbage Intake by Grazing Dairy Cows By J. A. C. Meijs

Notes: Two experiments were carried out to evaluate propionic acid-treated hay as a feed for sheep in which eighteen ewes (experiment 1) and eighteen ewe lambs (experiment 2) were fed ad libitum on hay only. At feeding, the propionic acid-treated hays had higher D-values and water-soluble carbohydrate concentrations, but a lower dry matter concentration, than the untreated hays. The voluntary feed intake and liveweight gain of the sheep fed on two acid-treated hays and on an untreated hay in experiment 1 were similar. In the second experiment an acid-treated moist hay was eaten in greater amounts by the sheep and liveweight gains were greater than on the corresponding untreated hay, but were not significantly different from those of sheep fed on field-cured hay.

Notes: Six varieties of white clover, each grown with perennial ryegrass, four intervals between cuts and two levels of applied nitrogen in all combinations, were compared in a field experiment during the first 27 months after sowing. Information about yields, crop fractions, heights and ryegrass tillers has been presented in an earlier paper (Wilman and Asiegbu, 1982). The present paper is concerned with the more detailed studies of white clover, which help to explain the yield results and contribute to the understanding of the response of this species to management when grown in competition with grass.Increasing the interval between harvests increased the length of clover stolon per unit area of ground and increased stolon diameter, petiole length, weight per leaf and number of leaves harvested as a proportion of the number present in the sward while only slightly reducing the rate of leaf emergence, helping to explain the positive effect of increasing the interval on clover yield noted in the earlier paper. During regrowth, successive leaves had longer petioles and the length of individual petioles increased beyond the stage at which the leaflets were fully opened. Weight per leaf in clover increased considerably from April to June and declined to below the April value by October. It was shown that weight per leaf can be greatly increased by increasing the interval between harvests without reducing the number of leaves harvested per unit area per year. The stolon length measurements provided some support for the view that medium large-leaved varieties of white clover can with advantage be defoliated rather less frequently than small-leaved varieties. Stolon length was less adversely affected by applied N in the medium large- than in the small-leaved varieties. The small-leaved varieties had thinner stolons than the medium large-leaved varieties but about twice the stolon length when no N was applied, and a relatively high proportion of leaves which escaped defoliation. The application of N reduced stolon diameter, increased petiole length and had little or no effect on weight per clover leaf.

Notes: Data from twenty-two comparisons carried out at ADAS Experimental Husbandary Farms are used to compare untreated and formic acid-treated silages. Additive treatment led to an improved fermentation in some crops, particularly those of low DM concentration (〈262 g kg-1). Where this occurred there were associated benefits in silage digestibility (+0·234 units), intake (+16%) and the growth rate of young cattle (+0·28 kg d-1). Where the fermentation of the untreated silage was good, both digestibility and animal performance associated with treated and untreated silages were similar. It is suggested that the justification for using formic acid in a commercial situation is thus restricted to occasions where the untreated crop would be liable to develop a clostridial fermentation. These may be when crops contain less than 35 g water-soluble carbohydrate kg-1.

Notes: The change in structure of continuously grazed versus infrequently cut swards of perennial ryegrass (Lolium perenne L), cv. S23, was investigated during their first full harvest year. Measurements were made from early May until late September. The intensity of stocking by sheep in the grazed sward was adjusted in an attempt to maintain a high level of radiation interception and the cut sward was harvested at approximately monthly intervals.The herbage mass, lamina area index and radiation interception of the cut sward varied in a cyclic pattern between harvests but in the grazed sward these parameters showed considerably less variation, although they all increased early in the season and then declined later. The proportion of dead material above ground increased throughout the season in both sward types but was more marked in the grazed sward.There were major differences between the grazed and cut swards in the number of tillers per unit ground area; the difference became more marked throughout the season and by September the tiller densities in the grazed and cut swards were 3·204 m-2 and 6·203 m-2 respectively. Divergence in tiller density was associated with differences in specific stem weight and leaf area per tiller.Rates of appearance and death of leaves on tillers in the grazed sward were determined. During May, leaf appearance exceeded leaf death but this was reversed in June. During the rest of the season as a new leaf appeared on a tiller so the oldest leaf died.

Notes: A rising-plate meter was used in a double sampling technique to measure the herbage mass of rotationally grazed perennial ryegrass-white clover swards over a period of 2 years. The meter was calibrated by developing a linear regression between meter reading and herbage dry matter mass as measured by cutting 0·2-m2 quadrats to ground level.There was a strong relationship between meter reading and herbage mass, and correlation coefficients were consistently 0·2 or above. The calibration regression was normally constant for extended periods, especially over the winter and spring. The relationship was more variable over the summer but appeared to follow a pattern that was to some extent repeatable between years. The slope of the regression (kg DM ha-1 cm-1) was 312 and 267 in the two winters and reached values of 800 and 452 in the two summers. The calibration relationship was adequately described by a linear model over the winter and spring but there was a tendency for a curved relationship in mid-to late summer.The standard pooled regression found with winter ryegrass-clover swards was not applicable to the more erect prairie grass, and the standard regression overestimated slightly the yield of heavily grazed swards. There was no evidence of a difference in relationship between irrigated and non-irrigated swards over the summer.The individual meter readings could be used to develop a useful diagrammatic picture of the changes that occurred in sward structure as the pastures were subjected to different managements and this could be used to illustrate problem areas in pasture management.The meter was useful in overcoming the problem of variability of herbage mass within paddocks and could give precise estimates of herbage mass, especially when it was possible to use a pooled regression encompassing a large number of calibration cuts. In these cases the meter could be used to detect differences of about 8%.

Notes: Awned seeds of four grasses were submitted to humidity changes on dry and damp soil to see how far the awns assisted in distribution and seed burial. The results indicate that awns on their own are probably of limited importance. In another study, seeds of seven species were soaked in water with high or low oxygen levels to determine whether dormancy could be induced. Waterlogging delayed germination briefly but did not induce dormancy except in Avena fatua. Oxygen level proved unimportant. The effect of temperature on germination and dormancy was examined in seeds of Agrostis capillars. Seeds of this species germinated best at fluctuating temperatures with a minimum difference of 10°C between day and night temperatures. Constant or slightly fluctuating temperatures over 20°C or under 10°C induced dormancy in a small percentage of seeds.

Notes: Changes in the population of stem-boring Dipterous larvae (including Oscinella frit) were examined weekly for a year on plots of perennial ryegrass (Lolium perenne cv. S24). The herbage net regrowth rate of these plots and of plots receiving the pesticide, phorate, was also monitored. Herbage net regrowth rates were greatest in April and October. At the same times the differences in net regrowth between treated and untreated plots were greatest, thus indicating the periods of the greatest loss of production due to pest damage to the sward.Populations of stem-boring larvae fell below 100 m−2 for only 4 weeks in the year and were maximal in autumn and early winter. Larval mortality exceeded 50% during the winter period. A correlation was found between larval activity and sward damage.

Notes: Swards of Phalaris aquatica-Trifolium subterraneum were subjected to four defoliation treatments—zero, low (11 sheep ha−1) and high (22 sheep ha−1) stocking rates, and weekly cutting. At high stocking rate the annual grass Hordeum leporinum dominated while clover was dominant at low and zero stocking rates. Weekly cutting suppressed species other than clover and so failed to simulate grazing.There were similarities in net herbage production between zero and lightly grazed swards and between heavily grazed and repeatedly cut swards. Net herbage production decreased in the order undisturbed sward 〈 lightly grazed sward 〈 heavily grazed sward 〈 repeatedly cut sward.When sheep grazed swards where herbage mass was low their daily consumption of herbage, and therefore liveweight change, depended on their recent grazing experience. Sheep accustomed to swards where herbage mass was low ate more because they grazed for much longer each day than unaccustomed sheep, although they selected a diet of similar digestibility.

Notes: A preliminary field experiment on a sward cut six times during 1978 indicated that total grass herbage harvested was negatively related to the number of Rumex obtusifolius plants present; densities of five to ten plants m−2 resulted in reductions of up to 30%, although the total quantity of herbage harvested (including Rumex) may have remained constant. However, R. obtusifolius densities did not significantly affect grass yield during the first half of the growing season. A negative relationship between R. obtusifolius ground cover and grass yield, particularly from late July onwards, was shown in a second field experiment during 1979 and 1980 using a 3-cut management. It was estimated that to produce a 20% reduction in grass growth in July-August and October, R. obtusifolius ground covers of 30% and 20% respectively were needed. Sixty per cent of the total herbage harvested in 1980 consisted of R. obtusifolius. These results indicate the damage that can be caused by this species and suggest that control measures should be taken before early summer.

Notes: A total of 104 cultivars of six agricultural grass species were compared under controlled conditions for their relative susceptibility to the common soil-borne fungus Fusarium culmorum (W.G.Sm.) Sacc. A number of the perennial ryegrass (Lolium perenne) cultivars showed a high degree of resistance, their emergence being little reduced under the stringent test conditions. The emergence of others was, however, severely affected. Italian ryegrass (L. multiflorum) cultivars generally had a high level of resistance, the outstanding exception being cv. Leda Daehnfeldt which proved very sensitive. None of the cultivars of cocksfoot (Dactylis glomerata), timothy (Phleum pratense), meadow fescue (Festuca pratensis) or tall fescue (F. arundinaceae) which were tested had more than a moderate level of resistance and most were highly susceptible.The vigour of ryegrass seedlings was, in most cases, closely correlated with percentage emergence but was less closely correlated in the more susceptible species. These often had a very low emergence but those seedlings which did establish were frequently of a size similar to that of healthy plants.Treatment of grass seed with a fungicide mixture of benomyl (15% a.i.) plus captan (60% a.i.) provided excellent protection against F. culmorum on even the most susceptible cultivars.Drazoxolon (40% a.i.) was, however, much less effective.

Notes: In field trials in 1979–81, perennial ryegrass cultivars Royal and Majestic (amenity) and Morenne (agricultural) produced maximum seed yields at levels of applied N ranging from 40 kg ha−1 to 160 kg ha−1. Available soil N levels were estimated at 55 kg ha−1; hence maximum seed yields were obtained at total nitrogen levels of 95–135 kg ha−1 in Royal, 95–215 kg ha−1 in Morenne, and 175–215 kg ha−1 in Majestic.Applied N at rates above 120 kg ha−1 either reduced or did not significantly increase seed yield, decreased seed numbers per unit area and decreased spikelets per tiller and seeds per spikelet.The use of a growth regulator increased seed yields by preventing lodging, but did so irrespective of nitrogen application rate. It is suggested that failure to increase seed yield at high N rates is a result not of poor pollination because of lodging, but seed abortion as a consequence of competition for assimilate supply by secondary vegetative tillers.

Notes: Swards were developed from plants of contrasting populations of Dactylis glomerata L. selected for high and low dry-matter (DM) production in monocultures and mixed cultures grown during 1970 and 1971. DM production of these mono-genotype swards was assessed during 1974. Those swards developed from the divergent selections from monocultures maintained a significant difference between the high and low selections, whereas those developed from selections out of mixed cultures showed no significant difference in yield. These results are consistent with earlier reports that there is no positive relationship between competitive ability in mixed cultures and vigour in pure stands. The implications of the physiological basis of these responses in terms of grass breeding are discussed.

Notes: Varieties of Italian, perennial and Italian × perennial ryegrasses, tall fescue and cocksfoot were used to determine the effects of soil moisture on grass growth. Weather conditions were monitored and herbage accumulation, leaf extension rate, leaf appearance rate and tillering were recorded under natural (control), covered and irrigated treatments.Water deficit reduced crop growth rate in the spring and drought was the major factor influencing crop growth rates in all varieties during the summer. The ryegrasses appeared most sensitive to drought, and particularly poor recovery growth was exhibited by the Italian ryegrass RvP and the hybrid ryegrass Snowdon.Leaf extension rate and leaf appearance rate were both reduced by increasing soil water deficit. Herbage accumulation was increased by irrigation when potential soil water deficits were greater than 100 mm. When water deficits were large, irrigation increased leaf extension more than leaf appearance or tiller number. Increasing moisture deficit had a greater effect upon tiller number than on leaf extension.

Notes: Records of grassland productivity were kept for 2 years on 152 dairy and 179 beef farms. Results were collated for six lowland zones, delineated primarily on the basis of average rainfall, and an upland zone. Contemporary and long-term average meteorological records were also collected.In most of the lowland zones stocking rates and use of fertilizer N were similar, but utilized metabolizable energy (UME) output from grass varied; it was 25% higher in the wet, cool zone of north-west England and east Wales than in the dry zone of eastern England. The ranking of zones for utilized output corresponded closely with the ranking for summer rainfall.On upland farms stocking rate was 25% lower and UME output 15% lower than on lowland farms, but this was achieved from little more than half the N input.The differences between zones were similar to those shown in other published farm data. They also showed a similar trend to that demonstrated in grass cutting experiments. This suggests that farmers were, on average, able to exploit the extra grass grown in wetter climates. The incidence of difficult topography and impeded drainage was similar in dry and wet zones, but did have a more serious effect in the wet zones. The much higher level of concentrate feeding in the driest zone may have reduced the utilization of grass.

Notes: Twelve varieties of cocksfoot, Dactylis glomerata L., were analysed for three mineral elements (Na, K, Ca) and their dry matter (DM) yields were measured in pot and field trials. Among these characteristics there were strong varietal differences in Na concentration, whereas in other cases the varietal differences were less significant. The concentrations of individual mineral elements were, in most cases, independent of yield and of one another.Parent plants and their clonal derivatives were highly correlated in Na and Ca concentration as well as in DM yield. Tiller size or position on the parental plant had no significant effects on the mineral concentrations or DM yields of clones derived from separated tillers.Within varieties continuous variation was observed for Na concentration and the range of variation increased with increasing mean Na concentration. When plants were grown under different experimental conditions there was a high repeatability in Na concentration within varieties over cuts and years. This result was taken as an indication of a strong genetic basis for Na concentration. The results obtained in this study show that it is possible to develop grass varieties of defined potential Na concentration.

Notes: Samples of 39 perennial ryegrass (Lolium perenne L.) and 24 Italian ryegrass (Lolium multiflorum Lam.) commercial seed-lots used at the Grassland Research Institute between 1974 and 1978 were examined for the presence of seed-borne Drechslera species. The five species of Drechslera isolated from the seeds were D. andersenii, D. siccans, D. nobleae, D. sorokiniana and an unidentified Drechslera sp.A high proportion of the seed-lots were infected but levels of infection were low except in the perennial ryegrass cv. S24. Infection was located more commonly inside the caryopses than externally on the lemmas and paleae, and about 12% of the seedlings growing from infected seeds were infected. The species recovered in the seedlings were the same as those observed on the seeds.There was no relationship between level of seed infection and level of damage observed in the field in the autumn of the first harvest year.

Notes: A high risk of pest damage to seedlings of white clover (Trifolium repens L.) and perennial ryegrass (Lolium perenne L.) was shown to exist after reseeding with a slot-seeder. Damage varied between years and sites; white clover appeared to be more affected than perennial ryegrass. Insecticides of feasible commercial use (chlorpyrifos granules and γ-HCH seed treatment) were found to prevent damage. There was no evidence of phytotoxicity from any of the pesticides studied at the dose rates used in the field.

Notes: The appearance of successive leaves on labelled tillers of S24 perennial ryegrass, and their subsequent death, were monitored for 1 year on cut plots on which four levels of N fertilizer were applied.A new leaf appeared about once every 15 d from the end of April until the beginning of September, and about once every 52 d from the end of October to the middle of February. Rate of appearance changed rapidly during September and October, and again from mid-February until the end of April. The response of leaf appearance rate to temperature changed early in the year; the effect of overwintering is discussed.Leaf lifespan was 50–60 d in summer and 120–140 d in autumn and early winter, and was influenced by conditions either during expansion of the leaf or shortly afterwards.The results are discussed in relation to the construction of a model to describe changes in the herbage available to animals grazing a grass sward.

Notes: A mathematical model which simulates the potential production and utilization of herbage on perennial ryegrass and grass-white clover swards in the context of an 18-month beef enterprise is described. Using the model the potential costs and benefits of replacing grass by grass-clover mixtures have been determined. Potentially grass-clover swards receiving no fertilizer N are shown to have annual DM yields equivalent to ryegrass swards receiving at least 300 kg N ha −1. Apart from the benefits of reduced fertilizer N usage there appear to be indirect benefits related to a lower concentrate requirement per head. In consequence, from an economic viewpoint grass-clover swards potentially compare very favourably with heavily fertilized grass swards, even if account is taken of the opportunity costs of lower stocking rates associated with grass-clover systems. However, this conclusion awaits experimental confirmation.

Notes: Earlier studies have concluded that for a dairy enterprise the most profitable conservation strategy, where cutting and grazing areas are separate, is generally associated with relatively low-digestibility silage. The aim of this study has been to investigate whether this conclusion remains true where cutting and grazing are integrated. A mathematical model of grass production and utilization by a dairy enterprise has been developed from information collected from a variety of experimental sources. It was found that cutting strategies aimed at making low-digestibility silage could not automatically be regarded as the most profitable. In good grass-growing conditions strategies resulting in low-digestibility silage did appear to offer the highest gross margins per unit area, but under average and poor growth conditions the greatest profits were associated with high-digestibility silage strategies. In particular, it was noted that the timing and frequency of conservation cuts influenced the efficiency with which grass could be utilized on the grazing areas and this fact rather than yield of conserved material influenced the comparative biological and economic efficiency of the different cutting strategies.

Notes: A study was conducted to determine the influence of herbage mass under continuous stocking management with sheep on rates of herbage growth (G), senescence (S) and net production of green herbage (NP) in a mixed-species sward of Lolium perenne. Poa annua and Trifolium repens. Plots were maintained in as nearly constant herbage mass as possible at 500, 700, 1000 and 1700 kg organic matter (OM) ha−1 by continuous but variable stocking with sheep for the period May to July inclusive. Estimates of G, S and NP were obtained for each species from repeated measurements over two 2-week periods on individually identified grass tillers or clover stolons. Estimates of species population densities enabled estimates of G. S and NP per unit area to be made.Rates of G were higher and of S were lower for Lolium tillers and Trifolium stolons than for Poa tillers. The laminae of Poa tillers occupied inferior positions in the canopy of the swards maintained at higher herbage masses and were less accessible to the grazing animals, so that Poa tissue was less well utilized.The total population densities of tillers and stolons declined at herbage masses both above and below 700 kg OM ha−1. The rate of G for the three species combined increased with herbage

Notes: The most likely reasons for seeding failures of sod-seeded lucerne (Medicago sativa L.) were determined from eight field experiments, where the effects of a range of pesticides on the establishment of and damage to lucerne were compared. The experiments were conducted during 1979–80 at four sites near Ithaca, N.Y., USA. Pesticides evaluated were: methyl bromide (MeBr), carbofuran, acylalanine (1979, 1980); fonofos, diazinon, methiocarb, metaldehyde and combinations of carbofuran with methiocarb or metaldehyde (1980). Seeded areas were pretreated with glyphosate for vegetation control.In 1979 seedling losses were decreased by carbofuran and methyl bromide whereas in 1980 seedling losses were decreased by methyl bromide at two sites and by methiocarb at all sites. Methiocarb was the only pesticide to reduce defoliation damage. The effect of combining methiocarb and metaldehyde each with carbofuran tended to be additive in terms of reducing seedling numbers and damage, and increasing yield. Treatment effects on lucerne yield were still apparent up to 12 months after seeding. Based on the consistent results of methiocarb, it was concluded that molluscs were probably the main factor limiting the establishment of sod-seeded lucerne but an arthropod component also appeared to be involved.

Notes: Book Reviews in this article: Silage in Milk Production By A. J. Corrall, H. D. St C. Neal and J. M. Wilkinson Sward Measurement Handbook Edited by J. Hodgson, R. D. Baker, Alison Davies, A. S. Laidlaw and J. D. Leaver Managing Agricultural Systems By G. E. Dalton

Notes: Faeces of cattle, pronghorn and rabbit were collected from the Central Plains Experimental Range, Colorado, 6, 18 and 30 months after being excreted. The faecal material was placed in moistened potting soil and seedling emergence observed. Bouteloua gracilis (blue grama grass) and Sporobolus cryptandrus (sand drop seed), emerged in abundance from cattle dung. Emergence of B. gracilis decreased with increasing period of exposure before collection, but emergence of S. cryptandrus increased with exposure. Forbs dominated the seedling populations in pronghorn dung, and few seedling populations of any species emerged from rabbit dung. The results are discussed in relation to the effect of large herbivores on the establishment of species from seed in rangeland.

Notes: The effect of temperature on the nutritive value of Macroptilium atropurpureum cv. Siratro and line 10–105 was determined in a controlled environment at day/night temperatures of 24/18, 30/24 and 36/30°C, and a 13-h daylength. Estimated dry matter digestibility, cell wall and lignin concentrations of leaf, stem and stubble were measured in two 5-week regrowth cuts.Siratro and line 10–105 were similar in digestibility and responded similarly to temperature with whole tops, stem and stubble decreasing by an average of 0.0023, 0.0026 and 0.0056 units of digestibility for each 1°C rise in growth temperature. These changes reflect the increase in cell wall and lignin concentrations with higher temperature.The digestibility of young, just-expanded leaves was increased by increased growth temperature. This effect may be explained by the faster rate of leaf expansion at high temperature which reduced cell wall and lignin concentrations at this stage of leaf development. However, for the bulk leaf fraction digestibility was unaffected by temperature. Leafiness declined at high temperatures.

Notes: The statistical properties and applicability of the Dry-Weight-Rank (DWR) method of pasture sampling are examined. Formulae for the expected value and variance of the DWR estimates are presented.The extent of bias in DWR is examined using several mathematical models. DWR is shown to break down when estimating multinomial proportions; however it provides good estimates for a subset of models simulated from a class of compound probability distributions.The success of DWR in practice does not rest on a theoretical basis. It appears to be a remarkable empirical discovery that DWR estimates are virtually unbiased on most pasture types. Several cases are considered where difficulties may be encountered in using DWR. Provided the theoretical reservations of this paper are borne in mind, DWR will continue to be a useful practical tool, especially in the absence of non-destructive observational alternatives.

Notes: Three groups of cows were continuously stocked on pasture and offered supplementary concentrates according to the herbage height measured weekly with a grass disc. Concentrates were offered when the mean herbage height fell below 9 cm for treatment A, 7 cm for treatment B and 5 cm for treatment C at a rate of 1 kg d-1 for each 0·2-cm decline below these threshold levels. If the herbage height declined by more than 1·2 cm (8 kg concentrates d-1) hay was offered in addition ad libitum. The stocking rates for all three treatments were 5·2, 3·2 and 3·2 cows ha-1 for three successive 8-week periods. For treatments A, B and C the mean concentrate DM intakes were 3·2, 1·2 and 1·2 kg d-1 (treatment A also received 0·2 kg hay DM d-1), and the mean daily milk yields were 17·2, 16·2 and 15·2 kg respectively. The increasing level of supplementary feeding from treatment C to A also resulted in an increase in liveweight and body condition score change, and a reduction in milk fat.

Notes: The effects of sheep grazing on the development of and production from heather swards and on intake and diet selection by grazing Scottish Blackface wether sheep were studied in an experiment in which sheep numbers were adjusted to remove 0, 40% or 80% by weight of current season's shoots in summer and in autumn for a period of 5 years. All combinations of season and level of grazing were provided. The plots were divided after 2 years and grazing was continued for a further 3 years on one subplot but discontinued on the other.Sward productivity during the 5 years of grazing was unaffected at the 40% level of shoot removal but at the 80% level was reduced by up to 66%.After a season's uninterrupted growth at the end of the experiment heather cover and herbage mass were lower and current season's shoots as a proportion of total mass were higher on those treatments which received the greatest severity of grazing. Weight of current season's shoots was unaffected on treatments which had received the 40% level of shoot removal but was reduced by 40% and 50% on treatments which had received the 80% level in summer and autumn respectively. On the rested subplots sward recovery was such that no treatment effects remained after 3 years except with respect to 80% shoot removal in autumn where herbage mass was reduced compared with other treatments.Intakes of digestible organic matter per sheep were higher at the 80% than the 40% level of grazing in the fifth year of the experiment. This was ascribed to the ingestion of new shoot growth from the twig bases on the 80% level of removal treatments. Intake and digestibility were higher in the summer on those treatments which had received the 80% level of removal in previous autumns. Over the 5 years of the experiment there was a small decline in intake and digestibility values, with the decline being greatest in the summer and at low levels of grazing.Floristic changes differed on the rested and grazed areas. On the rested subplots in the fourth year after grazing stopped, heather height was lower and cover by grasses, sedges and herbs was greater on treatments which previously had received the highest levels of utilization. On bare areas the rate of recovery of heather was rapid where shoots as well as seedlings contributed to recovery growth. On the grazed subplots, with the exception of the low-growing ruderal Rumex acetosella, cover due to grasses, sedges and herbs remained low as species other than heather were selectively grazed.Relationships among stocking rates, grazing behaviour and heather utilization are discussed and guidelines for the recognition of overgrazing are outlined.

Notes: In a study of growth rates and developmental morphology of simulated swards of two contrasting white clover varieties, Katrina and Kent, it was found that both varieties continued to produce dry matter throughout the winter at Aberystwyth. The gross crop growth rate in the coldest periods was about 7 kg ha-1 d-1. The rates of formation and loss of new leaves were approximately equal so that there was no net increase in weight until the latter part of March. This initial net weight increase in both varieties was observed in the same harvest interval, the most important factor being the difference in weight between the new leaf plus petiole unit and the weight of the unit which it replaced. The weight of laminae and petioles throughout the experiment was somewhat higher in Katrina which has come to be regarded as the ‘earlier’ variety.The prospects for improving spring production in white clover by variety improvement are discussed.

Notes: A technique for estimating the rate of disappearance of material from samples of dead herbage protected by simple nylon or wire mesh covers is described. Using this technique on a perennial ryegrass sward in southern England during August-November, relative rates of disappearance close to 0·204 g g-1 d-1 (DM) were obtained, but the variability was high. When herbage killed with paraquat was used, the rate of disappearance was higher and the variability lower. There were only small differences in the temperatures recorded in a normal sward and within samples of dead herbage under mesh covers.

Notes: Electrical capacitance and root diameter techniques for estimating root dry weight were evaluated in one test with red clover plants grown on slant-boards in plant growth chambers for 37–77 d, and in six tests with alfalfa grown in the field for 55–165 d. Root dry weights of red clover were estimated satisfactorily with regression equations from both diameter (R2=90%) and capacitance (R2=76%) measurements. Root dry weights of alfalfa were estimated satisfactorily throughout the entire season from diameter values (R2 from 51 to 92%). Alfalfa root weight was significantly (P 〈0·25) related to capacitance values at the beginning and at the end of the season but not at mid-season. Electrical capacitance measurements were similar for intact root systems and roots severed 4 cm below the crown in separate experiments with red clover and with alfalfa. The root diameter technique provided an accurate, rapid and inexpensive method of estimating root size throughout the first season of growth. The root capacitance method provided satisfactory estimates of roots at certain times during the summer. Neither method would be useful if a large portion of the root was naturally severed.

Notes: Italian ryegrass was slot-seeded into permanent grass in two field experiments at Oxford. In the first experiment August and September proved to be the most reliable months for slot-seeding, with April as a possible alternative. A row-spacing experiment showed that the optimum distance between rows was about 30 cm; seed rate in the row had a lesser influence on increasing herbage growth. In both experiments slot-seeding did not result in any substantial change in earliness of growth or alter total herbage harvested but it did even out the seasonal distribution of growth, particularly by increasing it in the following July.

Notes: An apparatus is described which enables excised pieces of plant material lo be weighed al frequent intervals as they dry in controlled conditions of temperature, humidity and air-speed. The effect of removing the epidermis on the drying rate of red clover (Trifolium pratense) leaflets, leaf petioles and stems is examined. Initially this treatment caused a very large increase in the drying rate of leaflets. Although the effect declined as water content fell, leaflets from which the epidermis had been removed still dried more rapidly than the controls at a water content equivalent to 50% of the dry weight. Removing the epidermis had a greater effect on the drying rate of leaf petioles and stems at low water contents than it did on the drying rate of leaflets.Slow drying at low water contents is a major factor responsible for the undesirably long periods for which hay may lie in the field. Slow drying occurs even though swath microclimate becomes more favourable for drying as water content falls. The results presented here suggest that treatments which reduce cuticular resistance have the potential to reduce field drying time.

Notes: A detailed study of the spring growth curves of two forage grasses, tall fescue and cocksfoot, over three successive years showed a large variability of growth over the vegetative phase. This variability was evident at levels of fertilizer N which were considered to be non-limiting (60 kg N ha-1 in autumn plus 120 kg N ha-1 in February).At this level of N there was a relationship between yield in the vegetative phase and accumulated temperatures from the last cut in the autumn which was described by a regression common to the three years. The slope of this regression represents the potential growth of a variety. At a lower level of N a separate regression was needed in each year to relate growth to accumulated temperature.Variation in spring growth was not related to accumulated net radiation. Growth in the reproductive phase was related to accumulated temperature and accumulated net radiation from the 10-cm ear stage.

Notes: In subtropical latitudes temperate crops can be grown during the cool months but the growing season is restricted by the termination and beginning of hot weather. Postponing sowing date in south Florida from October to November to January resulted in 150, 130 and 110-d growing seasons respectively. Dry matter (DM) yields of turnip, swede, rape and kale were lowered by each later sowing date and shorter growing season, and crude protein (CP) concentration was the same as for, or was increased by late sowing. In vitro organic matter digestibility (IVOMD) of turnip, swede and rape leaves was not affected by sowing date, but roots of turnip and swede from the earlier sowings and longer growing seasons were more digestible. Yield of the four species depended on harvest management as well as date of sowing and length of growing season. Total yield of turnip and swede were unaffected by harvest management but multiple cutting resulted in greater leaf yields and smaller root yields, whereas stockpiling resulted in smaller leaf yields and larger root yields. Kale produced more DM when harvested once at the end of the winter growing season but rape produced the greatest DM yield when sown in October and harvested

Notes: Plants of white clover (Trifolium repens L.) cv. Blanca and red clover (T. pratense L.) cv. Hungaropoly were grown singly in controlled environments. The effects of eight treatments on the two species were examined: 16- and 8-h photo-periods × 20/15 and 15/10°C day/night temperatures ×Rhizobium-free plants receiving nitrate N and inoculated plants receiving no combined N. Twice weekly measurements of the main axis leaf size, petiole length, rate of leaf production and the time period between the appearance of a leaf and its final size were carried out for 8 weeks after sowing.Increasing the day/night temperature from 15/10 to 20/15°C, or doubling the daylength from 8 to 16 h (doubling the daylength increased temperature by 1·2°C) accelerated the rate of leaf production and expansion and increased petiole length and the final area of leaves. Most aspects of main axis leaf growth were reduced in the inoculated plants dependent on their own N fixation compared with the nodule-free plants receiving abundant nitrate N. The results suggest that the temperature above which white clover exhibits appreciable leaf growth in the field could be relatively high compared with grasses. Further research is required to define these differences and relate them to seasonal changes in the environment.

Notes: Preharvest application of 1 and 2 litres ha-1 IPA glyphosate (glyphosate) to a seed crop of S24 perennial ryegrass with moisture concentrations of around 400 and 340 g kg-1 (40 and 34% moisture content w/w) failed to significantly lower seed and straw moisture concentration at harvest and consequently did not affect combine rate. Addition of extra surfactants and ammonium sulphate to glyphosate did not significantly increase the rate of desiccation.The quality of harvested seed was impaired at both rates and application times. Germination was significantly lowered through the production of abnormal seedlings. The germination of seed harvested in the previous year from glyphosatetreated plots decreased with storage. Seed vigour, germination rate and field emergence were also significantly decreased as a result of glyphosate application. The effect of glyphosate on seed quality precludes its preharvest use as a desiccant in the ryegrass seed crop.Harvesting difficulties due to the production of secondary vegetative tillers may be overcome by swathing rather than direct combining.

Notes: The fertility of forty-two heifers offered either red clover silage or grass silage prior to and during the period of insemination was compared. Pregnancy rate to first service was significantly higher (P 〈 0·25) on red clover silage at 76% compared with 43% on grass silage. Pregnancy rates from a mating period covering three oestrus cycles were similar on the two silage diets. The ratio of services to pregnancies was lower on red clover silage (1·2) than on grass silage (2·2). In a subsidiary trial with twenty-three heifers, pregnancy rate to first service on red clover silage was 78%.There was no evidence to indicate that herd fertility is depressed by red clover silage.

Notes: An estimate of the level of pest damage on enclosed upland permanent pasture in England and Wales was made by reducing invertebrate populations with insecticides at thirteen well-distributed sites and measuring the effect on herbage growth over a 3-year period. Potential pests were fewer in number than on lowland pastures and there was a wide range in herbage yield between sites. Insecticide treatment significantly increased annual yields at only three of the sites, and significantly decreased yield at two of the sites. Insect damage to grassland appears to be much less important in upland than in lowland areas: this may be due to the relative scarcity of frit-fly in these areas, but this could change if its preferred host, ryegrass, were encouraged by intensification of sward management in the uplands. At all but one of the sites insecticide treatment led to considerable soil compaction, probably as a consequence of suppressing earthworm populations.

Notes: Eight samples of perennial ryegrass-white clover herbage with in vivo dry matter digestibility (DMD) ranging from 0·279 to 0·264 were used to evaluate various cost-saving modifications to the two-stage pepsin–cellulase digestibility technique. The effect of sample size, cellulase quality, cellulase/sample ratio, digestion time and washing of residue following digestion were investigated. The loss of dry matter (DM) in the assay was correlated with in vivo DMD and each variation of the method was evaluated by comparing the s.d. between replicates and r.s.d. of the regression, as well as the convenience of the method for large-scale monitoring of digestibility of mixed ryegrass-clover herbage.It was found that the amount of cellulase used could be reduced by a factor of 25, compared with recently published methods, without increasing s.d. or r.s.d. appreciably. In addition stirring during digestion and washing of the residue could be omitted without any deleterious effects. Increasing digestion time did not reduce s.d. or r.s.d. and the low-grade cellulase proved to be slightly more economical.Increasing the sample size from 0·25 to 0·20 g improved the s.d. and r.s.d. but the residues from the larger samples were generally slower to filter, which made the assay unsuitable for routine use. Substantial reduction of digestion volume and use of a thermostatically controlled water bath instead of an incubator led to a considerable increase in efficiency and throughput of samples. Stirring the samples during digestion was found to be unnecessary, thus allowing more flexibility in the laboratory routine, for example using the weekend for digestion. Using the recommended method modification the repeatability between replicates and r.s.d. of the calibration regression was 0·204 and 0·215 respectively for samples ranging in DMD from 0·279 to 0·264.

Notes: Six varieties of white clover, each grown with perennial ryegrass, four intervals between cuts and two levels of applied nitrogen in all combinations, were compared in a field experiment during the first 27 months after sowing.Increasing the interval between harvests from 3 or 4 to 8–12 weeks increased the yield of white clover and generally did not reduce the proportion of clover in total herbage. Increasing the interval between harvests reduced the number of grass tillers but increased grass yield and the size of grass leaves and increased grass height more than clover height; it also increased the proportion of petiole relative to leaflet in the clover. Differences between varieties in response to interval between harvests were small but supported the view that medium large-leaved varieties can with advantage be defoliated rather less frequently than small-leaved ones. The adverse effect of applied N on clover appeared almost equally great with all four intervals between harvests and further research on this topic is suggested. Applied N increased grass height more than clover height and increased the number of grass tillers, the size of grass leaves and grass yield. The medium large-leaved varieties seemed more tolerant of applied N than the smaller varieties.

Notes: This second paper concerned with effects of prolonged spring defoliations on two early perennial ryegrass varieties (Cropper and RvP Hay Pasture), an intermediate perennial ryegrass (Talbot) and an early cocksfoot (Roskilde) examines the effects of different closing dates on ear emergence, weekly DOMD (in vitro) and conservation yields. The grasses were mown fortnightly to simulate sheep grazing from January until three closing dates, 19 April, 3 May or 17 May, in 1978 and 1979. A set of plots was left undefoliated. All plots were sampled weekly from 2 or 3 weeks after 17 May until the end of June or early July.Prolonged mowing resulted in a small delaying effect on 50% ear emergence of the ryegrasses. The maximum delay was 3 d with the latest closing date. Under this treatment, ear emergence of the cocksfoot was delayed by an average of 12 d but by only 0–4 d under the earlier closing dates. Delay in date of closing caused a significant delay in the time to reach a given DOMD but the effect was largely confined to the latest closing date and was greatest for the cocksfoot. The time when 670 g kg−1 DOMD was reached was delayed in the ryegrasses by no more than 5 d, except for a 12-d delay in the intermediate ryegrass in 1978 following the late closing date. Under this treatment the delay for the cocksfoot was 8 d in both years. The early ryegrasses produced stemmy regrowths.

Notes: For the 2 years 1977 and 1978 Masham ewes grazed a perennial ryegrass (Lolium perenne cv. S24) pasture from the beginning of April until the end of September each year. Twelve ewes were allocated to each of the three stocking rate treatments, 12 (L), 16 (M) and 20 (H) ewes per ha. Each treatment was rotationally grazed around six paddocks with a forward creep for the lambs. Conservation cuts were taken from each treatment in late May and were generally sufficient to cover a 3-month winter feeding period for treatments L and M but were always insufficient for treatment H. Lambing took place at pasture during the second half of May. No concentrates were fed to the ewes during late pregnancy or lactation but lamb birth weights (4–8 kg) and lamb growth rates over the first 4 weeks (240 g d−l) were satisfactory. Organic matter intakes (OMI)of grass by the pregnant ewes(1816,1844 g OMI ewe−1) were not affected by the stocking rate. Intake of grass by the lactating ewes was affected by year and by stocking rate. The overall growth rate of lambs was higher on treatment L than either M or H but decreased on all treatments during the latter half of the grazing season, resulting in 43% of L and only 3% of M and H lambs being fit for slaughter by the end of September. The main features of this time of lambing are low costs and the production of predominantly store lambs.

Notes: A trial was carried out to investigate whether or not sheep grazing at stocking rates of 22 (low) and 44 (high) ha−1 select red clover from a red clover-perennial ryegrass sward, The distribution of botanical and morphological components was assessed in consecutive 5-cm layers through the canopy inside and outside exclosure cages moved every 2 weeks for 6 weeks. The difference between inside and outside cages for a given layer was an indication of the amount of each component consumed from that position in the sward, and the corresponding difference for clover content was considered an indication of selection.Evidence of selection was observed over the third and fourth weeks of grazing, especially in the high stocking rate treatment. Laminae of red clover were selected in preference to other plant parts and were grazed throughout the depth of the canopy in contrast to grass. The basal layer at the high stocking rate during weeks 5 and 6 was not grazed suggesting that the crown of red clover may be protected from severe overgrazing by accumulation of dead matter.It was concluded that the low herbage density of red clover-grass swards allowed the occurrence of selection which was most pronounced when the sward was being grazed down relatively quickly.

Notes: An attempt was made to determine monetary values for grass which might be used by researchers to assess the likely economic implications of their findings. It is shown that the value of extra grass production will vary considerably depending on the use made of it. At 1982 price levels, estimated values for grass in the field ranged from 1–4 to 10–6 pence kg−1 DM, with a modal value around 3–5 pence kg−1 DM. At the same time, the seasonal distribution of the extra grass production is important since the value of extra grass at different points in the growing season may vary markedly. The way in which these values for grass may be used is illustrated, using as an example weed control during sward establishment. Finally, the sensitivity of investment returns to changes in the value placed on grass is discussed. Extreme caution is necessary in drawing general conclusions about benefits and costs from an evaluation based on a specific value for grass.

Notes: In three separate feeding experiments using a total of twenty-six individually-housed Ayrshire cows, three wilted silages made from Blanca white clover were offered ad libitum with either different supplements or different proportions of grass silage. The clover silages contained 680 g white clover kg−1 on a DM basis, and had a mean DM concentration of 263 g kg−1 with 231 g CP kg−1 DM and 91 g ammonia-N kg−1 N. The pH values averaged 4·16 and the DOMD concentrations 611 g kg−1. In experiment 1 the daily intake of clover silage given alone was 15.2 kg DM per cow, i.e. 30·1 g kg−1 live weight, and decreased by 0·76 kg DM kg−1 barley DM and by 0·66 kg DM kg−1barley plus soybean meal DM when these feeds were offered as supplements. Milk yield and fat concentration were higher on the supplement treatments than on the clover silage-only treatment. In experiments 2 and 3 the intakes of silage and total DM increased as the weight of clover in the diet increased from 0 to 700 g kg−1 with parallel increases in milk yield. The effects on milk composition were small and generally non-significant. Although white clover silages with excellent fermentations were made, it is concluded that the main role of white clover in a silage system will be in mixed swards with grass to reduce the input of fertilizer N and to increase the voluntary intake of silage.