ALBERT

Note: Dissertation, Universität Potsdam, 2024 , Contents 1 Introduction 1.1 The Arctic sea-ice cover 1.1.1 Sea ice in the coupled Arctic climate system 1.1.2 Recent changes of the Arctic sea ice 1.2 The atmosphere as driver of sea-ice variability 1.2.1 Large-scale circulation patterns 1.2.2 Role of cyclones 1.3 Thesis structure and research questions 2 Theory and methods 2.1 Synoptic cyclones 2.1.1 Related fundamentals of atmospheric dynamics 2.1.2 Cyclone activity in the Arctic 2.2 Cyclone tracking and cyclone occurrence mask 2.3 Dynamic and thermodynamic sea-ice variability related to cyclones 3 New insights into cyclone impacts on sea ice in the Atlantic sector of the Arctic Ocean in winter 3.1 Abstract 3.2 Introduction 3.3 Data and methods 3.3.1 Database and cyclone identification 3.3.2 Quantification of cyclone impacts on SIC 3.4 Cyclone impacts on SIC 3.4.1 Effects of different time scales and regions 3.4.2 Effects of SIC conditions and cyclone depth 3.4.3 Spatial variability of SIC response to cyclones 3.4.4 Relation to near-surface wind and surface energy budget 3.5 Signature of ’New Arctic’ conditions 3.6 Conclusions 3.7 Supplementary material 4 Impact of three intense winter cyclones on the sea ice cover in the Barents Sea: A case study with a coupled regional climate model 4.1 Abstract 4.2 Introduction 4.3 Data and methods 4.3.1 HIRHAM–NAOSIM simulation 4.3.2 Supplementary evaluation data 4.3.3 Dynamic and thermodynamic contributions to sea-ice changes 4.4 Results 4.4.1 Cyclone cases 4.4.2 Cyclone impacts on SEB 4.4.3 Cyclone impacts on sea-ice concentration (SIC) 4.4.4 Cyclone impacts on sea-ice thickness (SIT) 4.4.5 Context to other cyclone cases during the MOSAiC winter 4.5 Discussion and conclusions 4.6 Supplementary material 5 Cyclone impacts on sea ice concentration in the Atlantic Arctic Ocean: Annual cycle and recent changes 5.1 Abstract 5.2 Introduction 5.3 Data and methods 5.4 Changes in cyclones and traversed sea ice 5.5 Cyclone impacts on SIC 5.5.1 Annual cycle in the old Arctic 5.5.2 Changes in the new Arctic 5.5.3 Regional changes in autumn 5.6 Conclusions 5.7 Supplementary material 6 Conclusions and Outlook 6.1 What is the statistical impact of cyclone passages on sea-ice concentration (SIC) in the Atlantic Arctic Ocean? 6.2 What are the individual contributions of dynamic and thermodynamic processes to sea-ice changes related to cyclones? 6.3 Do the SIC impacts of cyclones change in a warming Arctic and what are the related mechanisms? 6.4 Ways forward Appendix: Cyclones modulate the control of the North Atlantic Oscillation on transports into the Barents Sea Bibliography

Note: Dissertation, Universität Potsdam, 2024 , Table of Contents Summary Deutsche Zusammenfassung 1 Introduction 1.1 Arctic ecosystems under global warming 1.2 The plant-associated microbiome 1.3 Drivers of soil development 1.4 Ancient DNA to unravel past ecosystems 1.4.1 Lake sediments as archives of past community changes 1.4.2 Metabarcoding for targeting specific communities 1.4.3 Shotgun sequencing for broader overview 1.5 Thesis objective 1.6 Thesis outline and author contributions 2 Manuscript I 2.1 Abstract 2.2 Introduction 2.3 Materials and Methods 2.3.1 Primer design and evaluation In silico analyses Evaluation of lake sediment core DNA for analyses of fungal paleoecology 2.4 Results Primer design and evaluation Evaluation of lake sediment core DNA for fungal paleoecology 2.4.1 Taxonomic resolution across the cores 2.4.2 Comprehensiveness: Rarefaction and accumulation curves 2.4.3 Amplicon length and GC content to assess bias through degradation 2.4.4 General taxonomic composition of fungi in Siberian lake sediment cores Diversity of fungal paleocommunities from lake CH12 2.5 Discussion 2.5.1 Preservation biases and potential contamination 2.5.2 Characteristics of the optimized sedaDNA ITS1 metabarcoding assay 2.5.3 Potential of lake sediment fungal DNA for paleoecology 2.6 Author contributions 2.7 Acknowledgements 2.8 Conflict of interest 2.9 References 3 Manuscript II 3.1 Abstract 3.2 Introduction 3.3 Geographic setting and study sites 3.4 Materials and Methods 3.4.1 Sampling 3.4.2 DNA extraction and amplification 3.4.3 Bioinformatic analysis 3.4.4 Assessment of negative controls and contamination 3.4.5 Statistical analysis and visualization 3.5 Results 3.5.1 Fungi: sedaDNA sequencing results and overall patterns of alpha diversity and taxonomic composition 3.5.2 Vegetation: sedaDNA sequencing results and overall patterns of alpha diversity and taxonomic composition 3.5.3 Site-specific plant-fungus covariation 3.5.3.1 Fungus and plant covariation in arctic Siberia from MIS3 to the Holocene 3.5.3.2 Quantitative relationships between fungi and plant richness and composition 3.6 Discussion 3.6.1 Fungus and plant diversity along a spatiotemporal gradient in Siberia 3.6.2 Changes in ecosystem functioning over a spatiotemporal gradient 3.6.3 Implications of our results for ecosystem functioning and future research avenues 3.7 Conclusions Funding Availability of data and material Author contribution Declaration of competing interest Acknowledgements 3.8 References 4 Manuscript III 4.1 Abstract 4.2 Introduction 4.3 Results and Discussion 4.3.1 Compositional changes of plants, fungi, and bacteria in ancient metagenomic datasets 4.3.2 Long-term soil development: a trajectory or environmentally driven processes? 4.3.3 Bioweathering supported by lichens and mycorrhiza 4.3.4 Turnover in carbon, nitrogen, and sulphur cycling 4.3.5 Tracing podzolization 4.4 Implications and conclusions 4.5 Material and methods 4.5.1 Geographical setting and study site 4.5.2 X-ray fluorescence scanning of the sediment core 4.5.3 Core sub-sampling 4.5.4 DNA extraction 4.5.5 Single stranded DNA library build 4.5.6 Bioinformatic pipeline for the analysis of the sequencing results 4.5.7 Data analysis 4.5.8 Analysis of the ancient patterns 4.5.9 Statistical analysis of the dataset Acknowledgements 4.6 References Declarations 5 Discussion and synthesis 5.1 Long-term rhizosphere establishment in tundra and taiga areas 5.1.1 SedaDNA as a proxy for soil microbiome 5.1.1.1 Fungal DNA metabarcoding 5.1.1.2 Targeting soil communities with shotgun sequencing 5.1.1.3 Comparison between metabarcoding and shotgun sequencing for the soil microbiome 5.1.2 Fungi-vegetation interaction changes over time 5.1.3 Soil development on a temporal gradient 5.2 Conclusion and future perspectives 6 References 7 Appendix 7.1 Appendix to manuscript I 7.2 Appendix to manuscript II 7.3 Appendix to manuscript III 7.4 Manuscript IV 7.4.1 Abstract 7.4.2 Introduction 7.4.3 Geographical setting and study sites 7.4.4 Material & Methods 7.4.4.1 Sub-sampling of the sediment cores 7.4.4.2 DNA extraction 7.4.4.3 Single stranded DNA library built 7.4.4.4 Bioinformatic pipeline for the analysis of the sequencing data 7.4.4.5 Data analysis 7.4.4.6 Statistical analysis of the datasets 7.4.5 Results 7.4.5.1 Compositional changes of representative plant taxa alongside dynamics in fungal ecologies and bacterial element cycling in ancient metagenomic datasets 7.4.5.2 Impact of abiotic and biotic drivers on soil establishment across geographical locations 7.4.5.3 Relative positive correlations of functional soil taxa with plants across the locations 7.4.5.4 Assessment of the plant taxon-specific microbiome across the locations 7.4.6 Discussion 7.4.6.1 Site-specific soil development 7.4.6.2 Differences in the bedrock 7.4.6.3 Correlation between the lake biota 7.4.6.3.1 General Trends in positively correlated rhizosphere taxa 7.4.6.3.2 Plant taxa specific microbiome 7.4.7 Implications and future directions 7.4.8 References 7.4.9 Supplement to manuscript IV Acknowledgements Eidesstattliche Erklärung Damage pattern analysis – Auflagen Doktorarbeit Summary Main References

Note: Contents Contributors Part I. Alaska's Past and Present Environment 1. The Conceptual Basis of LTER Studies in the Alaskan Boreal Forest / F. Stuart Chapin III, john Yarie, Keith Van Cleve, and Leslie A. Viereck 2. Regional Overview of Interior Alaska / James E. Beget, David Stone, and David L Verbyla 3. State Factor Control of Soil Formation in Interior Alaska / Chien-Lu Ping, Richard D. Boone, Marcus H. Clark, Edmond C. Packee, and David K. Swanson 4. Climate and Permafrost Dynamics of the Alaskan Boreal Forest / Larry D. Hinzman, Leslie A. Viereck, Phyllis C. Adams, Vladimir E. Romanovsky, and Kenji Yoshikawa 5. Holocene Development of the Alaskan Boreal Forest / Andrea H. Lloyd, Mary E. Edwards, Bruce P. Finney, Jason A. Lynch, Valerie Barber, and Nancy H. Bigelow Part II. Forest Dynamics 6. Floristic Diversity and Vegetation Distribution in the Alaskan Boreal Forest / F. Stuart Chapin III, Teresa Hollingsworth, David F. Murray, Leslie A. Viereck, and Marilyn D. Walker 7. Successional Processes in the Alaskan Boreal Forest / F. Stuart Chapin III, Leslie A. Viereck, Phyllis C. Adams, Keith Van Cleve, Christopher L. Fastie, Robert A. Ott, Daniel Mann, and Jill F. Johnstone 8. Mammalian Herbivore Population Dynamics in the Alaskan Boreal Forest / Eric Rexstad and Knut Kielland 9. Dynamics of Phytophagous Insects and Their Pathogens in Alaskan Boreal Forests / Richard A. Werner, Kenneth F. Raffa, and Barbara L. Illman 10. Running Waters of the Alaskan Boreal Forest / Mark W. Oswood, Nicholas F. Hughes, and Alexander M. Milner Part III. Ecosystem Dynamics 11. Controls over Forest Production in Interior Alaska / John Yarie and Keith Van Cleve 12. The Role of Fine Roots in the Functioning of Alaskan Boreal Forests / Roger W. Ruess, Ronald L. Hendrick, Jason C. Vogel, and Bjartmar Sveinbjornsson 13. Mammalian Herbivory, Ecosystem Engineering, and Ecological Cascades in Alaskan Boreal Forests / Knut Kielland, John P. Bryant, and Roger W. Ruess 14. Microbial Processes in the Alaskan Boreal Forest / Joshua P. Schimel and F. Stuart Chapin III 15. Patterns of Biogeochemistry in Alaskan Boreal Forests / David W. Valentine, Knut Kielland, F. Stuart Chapin III, A. David McCuire, and Keith Van Cleve Part IV. Changing Regional Processes 16. Watershed Hydrology and Chemistry in the Alaskan Boreal Forest: The Central Role of Permafrost / Larry D. Hinzman, W. Robert Bolton, Kevin C. Petrone, Jeremy B. Jones, and Phyllis C. Adams 17. Fire Trends in the Alaskan Boreal Forest / Eric S. Kasischke, T. Scott Rupp, and David L. Verbyla 18. Timber Harvest in Interior Alaska / Tricia L. Wurtz, Robert A. Ott, and John C. Maisch 19. Climate Feedbacks in the Alaskan Boreal Forest / A. David McCuire and F. Stuart Chapin III 20. Communication of Alaskan Boreal Science with Broader Communities / Elena B. Sparrow, Janice C. Dawe, and F. Stuart Chapin III 21. Summary and Synthesis: Past and Future Changes in the Alaskan Boreal Forest / F. Stuart Chapin III, A. David McCuire, Roger W. Ruess, Marilyn W. Walker, Richard D. Boone, Mary E. Edwards, Bruce P. Finney, Larry D. Hinzman, Jeremy B. Jones, Clenn P. Juday, Eric S. Kasischke, Knut Kielland, Andrea H. Lloyd, Mark W. Oswood, Chien-Lu Ping, Eric Rexstad, Vladimir E. Romanovsky, Joshua P. Schimel, Elena B. Sparrow, Bjartmar Sveinbjornsson, David W. Valentine, Keith Van Cleve, David L. Verbyla, Leslie A. Viereck, Richard A. Werner, Tricia L. Wurtz, and John Yarie Index

Note: Dissertation, Universität Potsdam, 2024 , Content Preface Acknowledgements Summary Zusammenfassung Abbreviations 1. Introduction 1.1. Peatlands 1.1.1. Peatland development and peat bog succession 1.1.2. Characteristic peatlands of the northern hemisphere 1.1.3. Anthropogenic threats of northern peatlands 1.1.4. Peat bog restoration 1.2. Peatland bryophytes 1.2.1. Brown mosses 1.2.2. Sphagnum mosses 1.3. Moss microbiota 1.3.1. Moss-associated bacteria 1.3.2. Moss-associated archaea 1.3.3. Endophytic prokaryotic communities 1.4. Biotic and abiotic influences on moss-associated microorganisms 1.5. Objectives 1.6. Study sites 1.6.1. High Arctic peatlands of Svalbard (SV) 1.6.2. Polygonal Tundra of Samoylov (SA) 1.6.3. Palsa Bogs of Neiden (NEI) 1.6.4. Kettle Bog Peatlands of Mueritz National Park (MUE) 2. Material and Methods 2.1. Sampling scheme overview 2.2. Sampling of pore water 2.3. Sampling of moss plantlets 2.4. Analysis of pore water chemistry 2.5. Cell wall analysis 2.5.1. Cation exchange capacity (CEC) 2.5.2. Holocellulose (HC) 2.5.3. Lignin and Lignin-like polymers (LLP) 2.5.4. Bulk moss litter analysis 2.6. Moss surface sterilisation and separation of putative epiphytic and endophytic microbial communities 2.7. DNA extraction and sequencing 2.8. Sequence analyses and bioinformatics 2.9. Statistical analyses 2.10. Potential methane production and oxidation assays 2.10.1. Surface sterilisation prior to activity tests 2.10.2. Methane production 2.10.3. Methane oxidation 3. Results 3.1. Peatland bulk and pore water characteristics 3.2. Diversity and structure of natural peatland microbial communities 3.3. Environmental drivers of moss-associated microbial communities 3.4. Microbial taxa associated with brown mosses and Sphagnum mosses 3.4.1. Moss-associated bacteria 3.4.2. Moss-associated archaea 3.4.3. Bacterial and archaeal core communities 3.4.4. Acetobacteraceae as dominant taxon of the bacterial core community 3.5. Sphagnum bacteriomes of disturbed, rewetted and pristine temperate kettle bog 3.6. Potential moss-associated methane production and methane oxidation rates 3.6.1. Moss-associated methane production 3.6.2. Moss-associated methane oxidation 4. Discussion 4.1. Environmental influences on moss-associated bacterial communities 4.2. Moss-associated archaeal communities and their environmental drivers 4.3. Distinct patterns of endophytic bacteria 4.4. The core microbiota and their possible role for peatland succession 4.5. The potential role of Acetobacteraceae for Sphagnum host mosses and bog ecosystems 4.6. Moss-associated microbial communities of the methane cycle and their potential metabolic activity 4.7. Diversity and structure of Sphagnum bacteriomes from pristine, disturbed and rewetted kettle bogs 5. Conclusion 6. Critical remarks and outlook 6.1. Critical remarks 6.2. Outlook Bibliography Supplementary

Note: Оглавление Роль В.Г. Зольникова в изучении мерзлотных почв Зольников Василий Георгиевич (к столетию со дня рождения) / Л.Г. Еловская Роль В.Г. Зольникова в изучении почвенного покрова Якутии / Д.Д. Саввинов Роль Центрального музея почвоведения им. В.В. Докучаева в исследовании мерзлотных почв России / Б.Ф. Апарин Региональные проблемы изучения мерзлотных почв Микростроение органопрофиля мерзлотных почв криолитозоны Забайкалья / В.М. Корсунов, В.М. Чиркова Криогенные почвы юга Восточной Сибири и вопросы их картографирования / В.А. Кузьмин Микроструктура почвенного покрова северной части О.Большевик (архипелаг Северная Земля) / Б.Ф. Апарин, В.Б. Апарин О количественной оценке разнообразия почв / А.И. Куликов, Е.Д. Канаева, М.А. Куликов К вопросу о мерзлотных почвах Юго-Восточного Алтая / Е.Н. Смоленцева Формы серы в мерзлотных почвах притундровых лесов Енисейского Севера / Т.В. Пономарева Влияние растительного покрова и микрорельефа на азотный фонд почв бугорковой тундры Сибири / К. Биази, В. Ванек, О. Русалимова, К. Кайзер, Х. Мейер, П. Барсуков, А. Рихтер Состав гумуса мерзлотных почв мелкодолинных ландшафтов Лено-Амгинского междуречья / М.В. Оконешникова, Р.В. Десяткин Особенности микробных комплексов островных почв дельты р. Селенга как показатели мерзлотности, обусловленной влиянием оз. Байкал / Э.О. Макушкин, Н.Д. Сорокин, В.М. Корсунов, Е.Ю. Шахматова, Е.Н. Афанасова Проблемы эволюции мерзлотных почв Центральной Якутии / А.П. Чевычелов, В.П. Скрыбыкина, А.А. Перк Особенности распространения доминирующих типов мерзлотных почв бассейна р. Биллях / Я.Р. Герасимов, Г.Н. Саввинов Биохимические и геохимические особенности почв урбанизированных территорий (на примере г. Мирный, Западная Якутия) / М.В. Щелчкова, Я.Б.Легостаева Экология мерзлотных почв Метанообразование в мерзлых почвах / А. Брушков, М. Фукуда Методические проблемы почвенно-геохимических исследований в экологии / Б.С. Ягнышев Криотурбации мерзлотных почв Севера при глобальном изменении климата (разнообразие, экология) / П.П. Гаврильев, Р.Н. Иванова Динамика гидротермических поясов / А.И. Дмитриев Влияние температурного режима почвы на радиальный прирост стволов лиственницы в Центральной Якутии / А.Н. Николаев, П.П. Федоров Сезонная динамика теплофизических свойств аласных почв / В.С. Макаров Изменение пространственной структуры растительного покрова аласа как показатель динамики почвенных условий / М.Х. Николаева, А.Р. Десяткин Агрофизические свойства мерзлотных палевых осолоделых почв Центральной Якутии / А.П. Пестерев Связь радиального прироста лиственницы Каяндера с температурным режимом почв / П.П. Федоров, А.Н. Николаев Естественные радионуклиды и радиоцезий в почвах тундровой и таежной зон Якутии / П.И. Собакин, А.П. Чевычелов Средняя годовая температура почвогрунтов на подошве деятельного слоя Западной Якутии и ее зависимость от средней годовой температуры воздуха / И.С. Васильев Баланс органогенного углерода в мерзлотных лесных экосистемах / Т.Х. Максимов, Б.И. Иванов, А.Й. Долман, Е.Й. Муре, А.П. Максимов, А.В. Кононов, Т. Ота, М. Хейманп Исследования динамики промерзания мерзлотных таежных почв около г. Якутска / П.Я. Константинов Охрана и рациональное использование мерзлотных почв Мерзлотно-экологическое районирование агроземель на примере Мегино-Кангаласского улуса / П.В. Ефремов Классификация геокриологических и почвенных процессов по степени опасности и безопасности для сельскохозяйственного освоения / П.П. Гаврильев Влияние погодных условий на солевой режим и продуктивность аласных дерново-луговых почв / А.Р. Десяткин, М.Х. Николаева Ритмика обводнения аласных почв Лено-Амгинского междуречья / Н.П. Босиков Урожайность зерновых в зависимости от варианта обработки и водно-физических свойств мерзлотной палевой осолоделой почвы / С.И. Баишев Почвенный покров и его деградация в окрестностях города Мирный / П.П. Данилов, Г.Н. Саввинов Особенности почвенного покрова территории месторождения "Таборное" / В.Г. Тарабукина, В.С. Макаров, В.С. Боескоров , English translation of Table of contents The role of V.G. Zolnikov in the study of permafrost soils Zolnikov Vasily Georgievich (on the occasion of the centenary of his birth) / L.G. Elovskaya The role of V.G. Zolnikov in the study of the soil cover of Yakutia / D.D. Savvinov The role of the Central Museum of Soil Science named after. V.V. Dokuchaev in the study of permafrost soils in Russia / B.F. Aparin Regional problems of studying permafrost soils Microstructure of the organoprofile of permafrost soils in the permafrost zone of Transbaikalia / V.M. Korsunov, V.M. Chirkova Cryogenic soils of the south of Eastern Siberia and issues of their mapping / V.A. Kuzmin Microstructure of the soil cover in the northern part of O. Bolshevik (Severnaya Zemlya archipelago) / B.F. Aparin, V.B. Aparin On the quantitative assessment of soil diversity / A.I. Kulikov, E.D. Kanaeva, M.A. Kulikov On the issue of frozen soils in South-Eastern Altai / E.N. Smolentseva Forms of sulfur in frozen soils of tundra forests of the Yenisei North / T.V. Ponomareva The influence of vegetation cover and microrelief on the nitrogen pool of soils in the hummock tundra of Siberia / K. Biasi, V. Vanek, O. Rusalimova, K. Kaiser, H. Meyer, P. Barsukov, A. Richter Composition of humus in permafrost soils in shallow valley landscapes of the Lena-Amga interfluve / M.V. Okoneshnikova, R.V. Desyatkin Features of microbial complexes of island soils of the river delta Selenga as indicators of permafrost caused by the influence of the lake. Baikal / E.O. Makushkin, N.D. Sorokin, V.M. Korsunov, E.Yu. Shakhmatova, E.N. Afanasova Problems of evolution of permafrost soils in Central Yakutia / A.P. Chevychelov, V.P. Skrybykina, A.A. Perk Features of the distribution of the dominant types of permafrost soils in the river basin. Billyakh / Ya.R. Gerasimov, G.N. Savvinov Biochemical and geochemical features of soils in urbanized areas (on the example of the city of Mirny, Western Yakutia) / M.V. Shchelchkova, Ya.B.Legostaeva Ecology of frozen soils Methane formation in frozen soils / A. Brushkov, M. Fukuda Methodological problems of soil-geochemical research in ecology / B.S. Yagnyshev Cryoturbation of permafrost soils of the North under global climate change (diversity, ecology) / P.P. Gavrilyev, R.N. Ivanova Dynamics of hydrothermal belts / A.I. Dmitriev The influence of soil temperature on the radial growth of larch trunks in Central Yakutia / A.N. Nikolaev, P.P. Fedorov Seasonal dynamics of thermophysical properties of alass soils / V.S. Makarov Changes in the spatial structure of the alas plant cover as an indicator of the dynamics of soil conditions / M.Kh. Nikolaeva, A.R. Desyatkin Agrophysical properties of frozen pale yellow solodized soils of Central Yakutia / A.P. Pesterev Relationship between the radial growth of Cajander larch and the temperature regime of soils / P.P. Fedorov, A.N. Nikolaev Natural radionuclides and radiocesium in soils of the tundra and taiga zones of Yakutia / P.I. Sobakin, A.P. Chevychelov Average annual soil temperature at the base of the active layer of Western Yakutia and its dependence on the average annual air temperature / I.S. Vasiliev Organogenic carbon balance in permafrost forest ecosystems / T.Kh. Maksimov, B.I. Ivanov, A.Y. Dolman, E.Y. Mure, A.P. Maksimov, A.V. Kononov, T. Ota, M. Heymanp Study of the dynamics of freezing of frozen taiga soils near the city of Yakutsk / P.Ya. Konstantinov Protection and rational use of permafrost soils Permafrost-ecological zoning of agricultural lands using the example of the Megino-Kangalassky ulus / P.V. Efremov Classification of geocryological and soil processes according to the degree of danger and safety for agricultural development / P.P. Gavriliev The influence of weather conditions on the salt regime and productivity of alas sod-meadow soils / A.R. Desyatkin, M.Kh. Nikolaev Rhythm of watering of alass soils of the Lena-Amga interfluve / N.P. Bosikov Grain yield depending on the treatment option and water-physical properties of frozen pale-yellow soil / S.I. Baishev Soil cover and its degradation in the vicinity of the city of Mirny / P.P. Danilov, G.N. Savvinov Features of the soil cover of the territory of the Tabornoe deposit / V.G. Tarabukina, V.S. Makarov, V.S. Fighters , In kyrillischer Schrift