ALBERT

Description: Males of Eledone cirrhosa grow to a size little over 600 g and normally have well-developed, and presumably active, reproductive organs from about 200 g upwards. Total weight of the genital bag is well correlated with total body weight (r= 0·906). Growth of the testis precedes that of the spermatophoric sac, and the size of neither of these reproductive components is predictable from body weight. The sizes of these organs and the estimated number and length of stored spermatophores are given for 100 g intervals of total body weight. No evidence was obtained for a seasonal trend in male maturity.

Description: A survey of the ecology of the octopus Eledone cirrhosa in Scottish waters is compiled from structured interviews with fishermen, records of occurrence in traps (for lobster and crab), and a research vessel survey. This species is widespread and common throughout the inshore waters covered by fishing activity (shoreline- 140 m) on bottom types ranging through rock, stones, sand and mud. It is caught in all months of the year but is especially common inshore in the summer (July-September) and further offshore on trawling grounds in October-December. The octopus is a normal and regular predator of large Crustacea (Hotnarus, Nephrops, Cancer) caught in commercial traps but gut contents yield little identifiable dietary remains.

Description: Markedly different cooling histories for the hanging- and footwall of theVari detachment on Syros and Tinos islands, Greece, are revealed by zircon and apatite fission-track data. The Vari/Akrotiri unit in the hangingwall cooled slowly at rates of 5–15 ◦CMyr−1 since Late Cretaceous times. Samples from the Cycladic blueschist unit in the footwall of the detachment on Tinos Island have a mean zircon fission-track age of 10.0±1.0 Ma, which together with a published mean apatite fission-track age of 9.4±0.5 Ma indicates rapid cooling at rates of at least ∼60 ◦CMyr−1. We derive a minimum slip rate of ∼6.5 kmMyr−1 and a displacement of 〉∼20 km and propose that the development of the detachment in the thermally softened magmatic arc aided fast displacement. Intra-arc extension accomplished the final ∼6–9 km of exhumation of the Cycladic blueschists from ∼60 km depth. The fast-slipping intra-arc detachments did not cause much exhumation, but were important for regionalscale extension and the formation of the Aegean Sea.

Description: Serological methods for prey identification have been applied to detection of residues ofsandeel (Ammodytidae) protein in faeces of common seals (Phoca vitulina) and grey seals(Halichoerus grypus) from the Moray Firth, north-east Scotland. Antisera raised to muscleprotein from Ammodytes marinus were evaluated by testing their reactions with proteinextracts made from a range of North Sea fish species and protein residues in in vitro digestates,seal digestive tracts and seal faeces. It was concluded that, using fused rocketimmuno-electrophoresis, linkage of precipitin peaks from unknown samples with peaksfrom standard sandeel extract was a reliable indicator of the presence of sandeel in theunknown sample. Seasonal variation in the incidence of sandeels in common seal diet in theMoray Firth was examined by identifying otoliths, bones, and proteins, and all threemethods indicated that sandeels occurred in the majority of samples tested in the summer,but were less important during the winter. Proteins were detected in fewer samples thanotoliths, particularly in February and March. Possible reasons for this difference arediscussed. Serological identification of sandeel proteins is potentially applicable to dietarystudies on all marine predators.

Description: Age and growth were estimated in the European squid, Loligo vulgaris, by examining growth increments in the statoliths of 203 specimens collected from off the French Mediterranean coast. Length and increment data were analyzed assuming that the increments were formed daily. The relationships between age and length showed that: growth rate varied considerably among individuals; growth was double exponential; the squids grew on average to 240 mm ML at 240 d from hatching, with a maximum of 350 mm at 240 d; the life span is probably about one year.

Description: Some of the limits to the use of serology to identify prey species in the digestive tracts of cephalopods have been evaluated. Cuttlefish, Sepia officinalis, were given meals of krill slurry (Euphausia superba). Protein extracts of contents from four regions of the digestive tract, stomach, caecum, digestive gland and intestine, were tested for prey antigenicity. Digestion times (loss of antigenicity) ranged from 1 to 8 h depending on sampling site. Stomach and caecum emptied rapidly, but meal antigenicity persisted longer in the digestive gland. The Sepia experiments provide a basis for interpretation of results from natural predation by cephalopods).

Description: Spirula spirula has stimulated considerable interest since it was first discovered. It is a member of one of the two genera of sepioids to frequent oceanic water (the other being Heteroteuthis); it has a unique spiral shell which acts as a buoyancy mechanism and can withstand considerable pressure (Denton, Gilpin-Brown & Howarth, 1967); and, until the capture by the Danish Oceanographical expeditions it was considered very rare, only 12 specimens having been captured. The Dana expeditions caught 193 individuals from 1909 to 1931 and these were described by Kerr (1931) and Bruun (1943,1955). Most of these were caught in the waters around the Canary Islands of the North Atlantic. Bruun (1943) arranged the specimens according to month and size and claimed that two size groups could be distinguished. The specimens were taken over a wide geographical area, in several years and during the months of February (1 specimen), March (40), April (3), May (8), June (1), August (1) and October (23). His conclusion concerning growth depends entirely upon his decision to split the March sample into two year-groups; those above 1.9 cm in ventral mantle length he put in a separate year-class to those below 1.9 cm in ventral mantle length. This division was arbitrary and, one suspects, based on a belief that a one-year life-span was likely. Clearly the growth of Spirula requires further study based on a larger collection and the present paper is an attempt to fulfil this need.

Description: During 1950, the Common Octopus (Octopus vulgaris Lamarck) was to be found along the south coast of England in greater numbers than at any time since Garstang (1900) reported on the ‘plague’ on the coasts of Devon and Cornwall in 1899–1900. In earlier papers (Rees, 1950, 1952) the distribution of the octopus in our northern waters was reviewed, and it was demonstrated that this species is an immigrant which breeds on our south coast only rarely. It reaches these coasts by being brought there as a planktonic larva by the water circulation in the English Channel and by migrations of the adult. The most important factor in controlling the movements of the adult, however, might be expected to be the water temperature in the English Channel—where the species is at the northern limit of its breeding range and might therefore be extremely sensitive to slight changes in temperature.

Description: The stomachsof 23 striped dolphins (Stenella coeruleoalba Meyen, 1833, Cetacea), stranded along the Ligurian coast (western Mediterranean Sea), contained 32 species of cephalopods, crustaceans and fishes, totalling an estimated 2,723 prey specimens representing about 36 kg in weight. Cephalopods and bony fishes were equally important in the diet (50%). Todarodes sagittatus (34.5%) and Micromesistius poutassou (25.9%) were found to be the most important food species. Other species belonging to six cephalopod families, three crustacean families and nine bony fish families, contributed to the diet with variable numbers, weights, and occurrences, demonstrating the opportunistic character of striped dolphin feeding.

Description: A new species of terebellid polychaete, Lanice arakani sp. nov., is described from two specimens collected in deep water at seamounts of the west Pacific by the submersible `JAGO', and comparisons are made with the established species of the genus. Special reference is given to the morphology of the worm's sediment tube and in situ observations.

Description: Arctic sea ice area (SIA) during late summer and early fall decreased substantially over the last four decades, and its decline accelerated beginning in the early 2000s. Statistical analyses of observations show that enhanced poleward moisture transport from the North Pacific to the Arctic Ocean contributed to the accelerated SIA decrease during the most recent period. As a consequence, specific humidity in the Arctic Pacific sector significantly increased along with an increase of downward longwave radiation beginning in 2002, which led to a significant acceleration in the decline of SIA in the Arctic Pacific sector. The resulting sea ice loss led to increased evaporation in the Arctic Ocean, resulting in a further increase of the specific humidity in mid-to-late fall, thus acting as a positive feedback to the sea ice loss. The overall set of processes is also found in a long control simulation of a coupled climate model.

Description: In the present study, the influence of some major tropical modes of variability on northern hemisphere regional blocking frequency variability during boreal winter is investigated. Reanalysis data and an ensemble experiment with the ECMWF model using relaxation towards the ERA-Interim reanalysis data inside the tropics are used. The tropical modes under investigation are El Niño Southern Oscillation (ENSO), the Madden-Julian Oscillation (MJO) and the upper tropospheric equatorial zonal-mean zonal wind . An early (late) MJO phase refers to the part of the MJO cycle when enhanced (suppressed) precipitation occurs over the western Indian Ocean and suppressed (enhanced) precipitation occurs over the Maritime Continent and the western tropical Pacific. Over the North Pacific sector, it is found that enhanced (suppressed) high latitude blocking occurs in association with El Niño (La Niña) events, late (early) MJO phases and westerly (easterly) . Over central to southern Europe and the east Atlantic, it is found that late MJO phases, as well as a suppressed MJO are leading to enhanced blocking frequency. Furthermore, early (late) MJO phases are followed by blocking anomalies over the western North Atlantic region, similar to those associated with a positive (negative) North Atlantic Oscillation. Over northern Europe, the easterly (westerly) phase of is associated with enhanced (suppressed) blocking. These results are largely confirmed by both the reanalysis and the model experiment.

Description: In a recent paper I stated that the hypothesis of a continuously cooling earth had “consistently failed to lead to any adequate explanations of fissure eruptions and other volcanic and tension phenomena, mountain-building processes and their distribution in time and space, and the alternation of marine transgressions and recessions”. It seems to be desirable that I should further justify this sweeping integration of objections, for Dr. Harold Jeffreys, who has actively explored the consequences of this theory during recent years, takes exception to my discouraging view of the position, and claims that the alleged failures include the principal successes of the theory. The evidence on which my summary indictment was based will be reviewed in this paper; and attention will be directed to certain other phenomena—such as the growth of geosynclines—of which the significance has not yet been generally realized. As a preliminary it will be convenient to discuss the gradually converging evidence bearing on the thickness and substructure of the continents.

Description: The Eddy Kinetic Energy (EKE) associated with the Subtropical Countercurrent (STCC) in the western subtropical South Pacific is known to exhibit substantial seasonal and decadal variability. Using an eddy-permitting ocean general circulation model, which is able to reproduce the observed, salient features of the seasonal cycles of shear, stratification, baroclinic production and the associated EKE, we investigate the decadal changes of EKE. We show that the STCC region exhibits, uniquely among the subtropical gyres of the world’s oceans, significant, atmospherically forced, decadal EKE variability. The decadal variations are driven by changing vertical shear between the STCC in the upper 300 m and the South Equatorial Current below, predominantly caused by variations in STCC strength associated with a changing meridional density gradient. In the 1970s, an increased meridional density gradient results in EKE twice as large as in later decades in the model. Utilizing sensitivity experiments, decadal variations in the wind field are shown to be the essential driver. Local wind stress curl anomalies associated with the Interdecadal Pacific Oscillation (IPO) lead to up- and downwelling of the thermocline, inducing strengthening or weakening of the STCC and the associated EKE. Additionally, remote wind stress curl anomalies in the eastern subtropical South Pacific, which are not related to the IPO, generate density anomalies that propagate westward as Rossby waves and can account for up to 30–40 % of the density anomalies in the investigated region.

Description: New light is shed on Worthington's concept of the North Atlantic circulation, postulating the existence of two anticyclonic gyres. This concept, which seems to have been laid to rest in the last decade, has now been reinforced by the results of a simple linear Sverdrup circulation model yielding a band of westward transport all across the North Atlantic at about the Azores latitude. This narrow band is called the Azores Countercurrent (AzCC) and matches the position of westward flow required by Worthington's “northern gyre.” An anomaly in the meridional change of the wind-stress curl in the eastern North Atlantic has been identified as the driving mechanism. A comparison with observations shows that the AzCC is verified in many analyses of historical datasets and synoptic surveys. A lack of the AzCC in other analyses is probably due to missing meridional sections, strong smoothing, and the superimposed Ekman flow close to the sea surface directed to the southeast. The AzCC has not been verified in low-resolution general circulation models applying simplified wind-stress fields and large friction coefficients, but there is evidence for its existence in recent high-resolution models driven by realistic wind stresses. Based on these findings, a new pattern for the wind-driven upper ocean circulation of the midlatitude North Atlantic is presented.

Description: The importance of feeding pattern is well documented in fish (Jenkins & Green, 1977; Simenstad & Cailliet, 1986) but there are not many reported studies in cephalopods. Feeding patterns, as defined by Jenkins & Green (1977) have been studied, to our knowledge, only in Todarodes pacificus (Okiyama, 1965), Loligo pealei (Vovk, 1972), Loligo opalescens (Karpov & Cailliet, 1978), Illex illecebrosus (Amaratunga et ah, 1979; Amaratunga, 1980) and Nototodarus gouldi (O'Sullivan & Cullen, 1983). Boyle (1983) dealt with aspects of feeding in several cephalopod species but not specifically with feeding pattern. Aspects of feeding in Sepia officinalis have been reviewed by Nixon (1987). The present work describes the daily feeding pattern in Sepia officinalis from data collected in the field.

Description: Thirty-four adult individuals of Loligo forbesi (males and females with dorsal mantle lengths from 27–77 cm) were observed in captivity in a 3 m diameter closed sea-water system on Faial Island, Azores. Squids were caught by jigging and were fed with horse mackerel (Trachurus picturatus) and chub mackerel (Scomber japonicus), either alive or dead. The maximum survival was 73 days. Feeding behaviour was dependent upon both the size of prey and its state of preservation;e.g. the squid would eat the head of small fish (〈 about 15 cm), reject the head of medium-sized fish (about 15–35 cm) and would eat only the dorsal part of a big fish (〉 about 35 cm) or a fish poorly preserved. Seventeen chromatic, 9 postural and 6 movement components of body patterns were observed and described. Conspecific interactions considered to be aggression and dominance were observed among males; no such interactions occurred when one male and two females were kept together. Body patterns in relation to relaxation, stress, shock, feeding, locomotion and aggression are also described.

Description: The Indian Ocean has sustained robust surface warming in recent decades, but the role of multi-decadal variability remains unclear. Using ocean model hindcasts, characteristics of low-frequency Indian Ocean temperature variations are explored. Simulated upper-ocean temperature changes across the Indian Ocean in the hindcast are consistent with those recorded in observational products and ocean reanalyses. Indian Ocean temperatures exhibit strong warming trends since the 1950s limited to the surface and south of 30°S, while extensive subsurface cooling occurs over much of the tropical Indian Ocean. Previous work focused on diagnosing causes of these long-term trends in the Indian Ocean over the second half of the 20th Century. Instead, the temporal evolution of Indian Ocean subsurface heat content is shown here to reveal distinct multi-decadal variations associated with the Pacific Decadal Oscillation and the long-term trends are thus interpreted to result from aliasing of the low-frequency variability. Transmission of the multi-decadal signal occurs via an oceanic pathway through the Indonesian Throughflow and is manifest across the Indian Ocean centered along 12°S as westward propagating Rossby waves modulating thermocline and subsurface heat content variations. Resulting low-frequency changes in the eastern Indian Ocean thermocline depth are associated with decadal variations in the frequency of Indian Ocean Dipole (IOD) events, with positive IOD events unusually common in the 1960s and 1990s with a relatively shallow thermocline. In contrast, the deeper thermocline depth in the 1970s and 1980s is associated with frequent negative IOD and rare positive IOD events. Changes in Pacific wind forcing in recent decades and associated rapid increases in Indian Ocean subsurface heat content can thus affect the basin’s leading mode of variability, with implications for regional climate and vulnerable societies in surrounding countries.

Description: The presence of Stoloteuthis leucoptera in the Mediterranean is recorded on the basis of three specimens, including an adult male, caught by IKMT and by commercial otter-trawl in the Ligurian Sea. The hypothesis of a recent immigration is discussed. The list of Mediterranean cephalopods (Mangold Wirz, 1963; Torchio, 1968; Bello, 1986; Mangold & Boletzsky, 1987) includes the Sepiolidae of the subfamily Heteroteuthinae, whose members are supposed to be pelagic throughout their life cycle. Mangold Wirz (1963) recognizes in the Mediterranean fauna the unique species Heteroteuthis dispar, the other authors include H. atlantis Voss, which Voss himself (1955) reported at Messina. To this group may now be added Stoloteuthis leucoptera (Verrill, 1878) a species until now recorded in limited Atlantic areas. Verrill (1881) wrote “This species is an exceedingly beautiful one, when living, owing to the elegance and brilliancy of its colours and the gracefulness of its movements. In swimming it moves its fins in a manner analogous to the motion of the wings of a butterfly.”

Description: We present chemical data on magmatically heterogeneous pyroclastic deposits of late Quaternary age erupted from zoned magma systems underlying Tenerife (Canary Islands), Sao Miguel and Faial (Azores), and Vesuvius. The most fractionated magmas present at each centre are respectively Na-rich phonolite, trachyte, and K-rich phonolite. Within any one deposit, chemical variation is either accompanied by changes in the phenocryst assemblage (petrographic zonation) or is largely manifested in trace element abundances, unaccompanied by any petrographic change (occult zonation). Zoning is analogous to that in calc-alkaline systems where the most fractionated products are high-silica rhyolites. When a range of magma types are considered, a correlation emerges between roofward depletion of trace elements (especially REE) in the zoned system and compatability of those same trace elements in the accessory phenocryst phases present. Thus, allanite- or chevkinite-bearing rhyolitic systems are light-REE depleted roofwards, the sphene-bearing Tenerife system is middle-REE depleted roofwards, the melanite-bearing Vesuvius system is heavy-REE depleted roofwards, while the Azores systems, which lack these phases, display roofward REE enrichment. Therefore, the behaviour of trace elements may in each case be explained by fractionation of observed phenocryst assemblages. The resemblance between features of zoned magma systems and published work on the dynamic consequences of cooling saturated aqueous solutions prompts us to suggest that sidewall crystallization and consequent boundary-layer uprise to form a capping layer at top of the system may be a plausible mechanism for the generation of both petrographic and occult zonation. Reverse zoning occurs among the first-erupted tephra of some deposits, demonstrating that the most highly differentiated magma available is not always the first to be tapped during an eruption from a zoned system.

Description: Changes in the relative size of the ovary, oviducal glands and eggs are described for Eledone cirrhosa captured from the North Sea off Aberdeen over a 3 year period (N = 488). The analysis is based only on freshly caught animals, excluding those held in aquarium conditions (〉 5 days). Ovary enlargement and egg size estimates are used as indices of sexual maturity. Between 0–15% and 18–95% of total body weight is contributed by the ovary. Maximum egg length in the ovary ranges up to 7 mm. On these criteria, sexual maturation typically occurs at body sizes between 400–1000 g although some animals of 1000–1200 g are found showing no evidence of ovary enlargement. The majority of the monthly sample is always immature but maturation can apparently occur at almost any time of the year. Increase in mean ovary index and mean values for egg size are strongly seasonal and indicate a peak incidence of sexual maturity over 2–3 months in the July-September period. Spawning is presumed to follow within 1 month. Estimates of the fecundity of the females, based on the egg sample from the ovary, range from 2·2 × 103 to 55 × 10 3 eggs with a mean of 11 × 10 3 and a mode of 7·5 × 10 3 eggs.

Description: Female Eledone cirrhosa held in aquarium conditions for periods of time of five daysand over show relatively enlarged ovary sizes. Values for ovary index considerably exceed thoseof freshly caught animals and the incidence of the final stage of maturity, in which eggs pack the oviducts, is greater. A comparison of maturity indices for fresh and aquarium males was inconclusive. The range of factors associated with aquarium conditions is briefly reviewed and it is concluded that studies of cephalopod reproductive maturation must distinguish fresh and aquarium animals. Introduction External factors effective in inducing sexual maturation in cephalopods have been suggested several times. The influence of the absence of light has been implicated since the experiments of Wells & Wells (1959) showed that blinded Octopus vulgar is matured precociously. An effect of short day length in stimulating the optic glands of Sepia has been found by Defretin & Richard (1967) and Richard (1967) but this is not clearly the case for Octopus (Buckley, 1977). Octopuses kept in aquarium conditions for lengthy periods are said to have larger relative gonad sizes than those fresh from the sea (Wells & Wells, 1975). One of the factors associated with aquarium conditions is often a degree of starvation, and this circumstance alone is held to be a factor in inducing precocious sexual maturation in Eledone (Mangold & Boucher-Rodoni, 1973). In the course of recent studies on the growth and reproduction of Eledone cirrhosa from the North Sea (Boyle & Knobloch, 1982,1983,1984) animals which had remained in aquarium conditions for 5 days or over were separated from the analysis.

Description: Santorini volcanic field has had 12 major (1–10 km3 or more of magma), and numerous minor, explosive eruptions over the last ~ 200 ka. Deposits from these eruptions (Thera Pyroclastic Formation) are well exposed in caldera-wall successions up to 200 m thick. Each of the major eruptions began with a pumice-fall phase, and most culminated with emplacement of pyroclastic flows. Pyroclastic flows of at least six eruptions deposited proximal lag deposits exposed widely in the caldera wall. The lag deposits include coarse-grained lithic breccias (andesitic to rhyodacitic eruptions) and spatter agglomerates (andesitic eruptions only). Facies associations between lithic breccia, spatter agglomerate, and ignimbrite from the same eruption can be very complex. For some eruptions, lag deposits provide the only evidence for pyroclastic flows, because most of the ignimbrite is buried on the lower flanks of Santorini or under the sea. At least eight eruptions tapped compositionally heterogeneous magma chambers, producing deposits with a range of zoning patterns and compositional gaps. Three eruptions display a silicic–silicic + mafic–silicic zoning not previously reported. Four eruptions vented large volumes of dacitic or rhyodacitic pumice, and may account for 90% or more of all silicic magma discharged from Santorini. The Thera Pyroclastic Formation and coeval lavas record two major mafic-to-silicic cycles of Santorini volcanism. Each cycle commenced with explosive eruptions of andesite or dacite, accompanied by construction of composite shields and stratocones, and culminated in a pair of major dacitic or rhyodacitic eruptions. Sequences of scoria and ash deposits occur between most of the twelve major members and record repeated stratocone or shield construction following a large explosive eruption. Volcanism at Santorini has focussed on a deep NE–SW basement fracture, which has acted as a pathway for magma ascent. At least four major explosive eruptions began at a vent complex on this fracture. Composite volcanoes constructed north of the fracture were dissected by at least three caldera-collapse events associated with the pyroclastic eruptions. Southern Santorini consists of pryoclastic ejecta draped over a pre-volcanic island and a ridge of early- to mid-Pleistocene volcanics. The southern half of the present-day caldera basin is a long-lived, essentially non-volcanic, depression, defined by topographic highs to the south and east, but deepened by subsidence associated with the main northern caldera complex, and is probably not a separate caldera.

Description: Samples of two loliginid squids Alloteuthis africana and A. subulata were collected from the continental shelf off the west Sahara in August-September 1987. Statoliths were taken from 124 specimens and processed using statolith ageing techniques. Statoliths of both species were very similar in shape. In the ground statolith, growth increments were examined and grouped into four growth zones distinguished mainly by the width of the increments. Age of adult mature males of both species did not exceed eight months, that of females six months. Alloteuthis africana grew faster than A. subulata in weight and, particularly, in length. At age 180 d the mantle of A. africana was twice as long and the body weight 1·2–1·5 times as large. Both species matured over a wide range of sizes and ages (from 120 to 180 d). The life span of A. africana and A. subulata hatching between January and May on the west Saharan shelf is about six months, much shorter than that of A. subulata in its northern temperate range.

Description: A new species of a monogenean Isancistrum subulatae (Gyrodactylidae) has been discovered on the arms and tentacles of the cephalopod mollusc Alloteuthis subulata at Plymouth and I. loliginis, on the gills of the same host, has been re-discovered for the first time since its original description in 1912. I. subulatae, like other gyrodactylids, is viviparous, and has been shown by experiments to transfer to new hosts by contagion. In nature such transfers probably take place during copulation of the hosts and since the parasite may occur in numbers of several thousands per host, it may thereby constitute a venereal disease.

Description: The Argentine margin contains important sedimentological, paleontological and chemical records of regional and local tectonic evolution, sea level, climate evolution and ocean circulation since the opening of the South Atlantic in the Late Jurassic–Early Cretaceous as well as the present-day results of post-depositional chemical and biological alteration. Despite its important location, which underlies the exchange of southern- and northern-sourced water masses, the Argentine margin has not been investigated in detail using scientific drilling techniques, perhaps because the margin has the reputation of being erosional. However, a number of papers published since 2009 have reported new high-resolution and/or multichannel seismic surveys, often combined with multi-beam bathymetric data, which show the common occurrence of layered sediments and prominent sediment drifts on the Argentine and adjacent Uruguayan margins. There has also been significant progress in studying the climatic records in surficial and near-surface sediments recovered in sediment cores from the Argentine margin. Encouraged by these recent results, our 3.5-day IODP (International Ocean Discovery Program) workshop in Buenos Aires (8–11 September 2015) focused on opportunities for scientific drilling on the Atlantic margin of Argentina, which lies beneath a key portion of the global ocean conveyor belt of thermohaline circulation. Significant opportunities exist to study the tectonic evolution, paleoceanography and stratigraphy, sedimentology, and biosphere and geochemistry of this margin.

Description: The North Atlantic Oscillation (NAO) and the Arctic Oscillation (AO) describe the dominant part of the variability in the Northern Hemisphere extratropical troposphere. Due to the strong connection of these patterns with surface climate, recent years have shown an increased interest and an increasing skill in forecasting them. However, it is unclear what the intrinsic limits of short-term predictability for the NAO and AO patterns are. This study compares the variability and predictability of both patterns, using a range of data and index computation methods for the daily NAO/AO indices. Small deviations from Gaussianity are found and characteristic decorrelation time scales of around one week. In the analysis of the Lyapunov spectrum it is found that predictability is not significantly different between the AO and NAO or between reanalysis products. Differences exist however between the indices based on EOF analysis, which exhibit predictability time scales around 12 - 16 days, and the station-based indices, exhibiting a longer predictability of 18 - 20 days. Both of these time scales indicate predictability beyond that currently obtained in ensemble prediction models for short-term predictability. Additional longer-term predictability for these patterns may be gained through local feedbacks and remote forcing mechanisms for particular atmospheric conditions.

Description: The 13th European Polar Low Workshop was organized by the European Polar Low Working Group (www.uni-trier.de/index.php?id=20308)and gathered scientists from nine countries focusing on polar mesocyclones in both hemispheres and other mesoscale weather phenomena such as katabatic winds, tip jets, boundary layer fronts, cold air outbreaks, and weather extremes in polar regions. Topics included experimental, climatological, theoretical, modeling, and remote sensing studies. The aim was to bring together scientists and forecasters to present their latest work and recent findings on these topics and to encourage discussions on improving forecasting and understanding of these phenomena.

Description: The regional climate model COSMOin Climate Limited-AreaMode (COSMO-CLM or CCLM) is used with a high resolution of 15km for the entire Arctic for all winters 2002/03–2014/15. The simulations show a high spatial and temporal variability of the recent 2-m air temperature increase in the Arctic. The maximum warming occurs north of Novaya Zemlya in the Kara Sea and Barents Sea between March 2003 and 2012 and is responsible for up to a 208C increase. Land-based observations confirm the increase but do not cover the maximum regions that are located over the ocean and sea ice.Also, the 30-km version of theArctic SystemReanalysis (ASR) is used to verify the CCLM for the overlapping time period 2002/03–2011/12. The differences between CCLM and ASR 2-m air temperatures vary slightly within 18C for the ocean and sea ice area. Thus,ASR captures the extreme warming as well. The monthly 2-m air temperatures of observations and ERA-Interim data show a large variability for the winters 1979–2016. Nevertheless, the air temperature rise since the beginning of the twenty-first century is up to 8 times higher than in the decades before. The sea ice decrease is identified as the likely reason for the warming. The vertical temperature profiles show that the warming has a maximum near the surface, but a 0.58Cyr21 increase is found up to 2 km. CCLM, ASR, and also the coarser resolved ERA-Interim data show that February and March are the months with the highest 2-m air temperature increases, averaged over the ocean and sea ice area north of 708N; for CCLM the warming amounts to an average of almost 58C for 2002/03–2011/12.

Description: A sigma-coordinate, primitive equation ocean circulation model is used to explore the problem of the remnant generation of trapped waves about a tall, circular, isolated seamount by an incident oscillatory barotropic current. The numerical solutions are used to extend prior studies into the fully nonlinear regime, and in particular to quantify and interpret the occurrence of residual circulation. Specific attention is also devoted to the dependence of the resonance and rectification mechanisms on stratification, forcing frequency, and choice of subgrid-scale viscous closure. Resonantly generated trapped waves of significant amplitude are found to occur broadly in parameter space; a precise match between the frequency of the imposed incident current and the frequency of the trapped free wave is not necessary to produce substantial excitation of the trapped wave. The maximum amplification factors produced in these numerical solutions, O(100) times the strength of the incident current, are consistent with previous studies. In the presence of nonlinear advection, strong residual currents are produced. The time-mean circulation about the seamount is dominated by a strong bottom-intensified, anticyclonic circulation closely trapped to the seamount. Maximum local time-mean current amplitudes are found to be as large as 37% of the magnitude of the propagating waves. In addition to the strong anticyclonic residual flow, there is a weaker secondary circulation in the vertical-radial plane characterized by downwelling over the top of the seamount at all depths. Maximum vertical downwelling rates of several tens of meters per day occur at the summit of the seamount. The vertical mass flux implied by this systematic downwelling is balanced by a slow radial flux of mass directed outward along the flanks of the seamount. Time-mean budgets for the radial and azimuthal components of momentum show that horizontal eddy fluxes of momentum are responsible for transporting net radial and azimuthal momentum from the far field to the upper flanks of the seamount. There, Coriolis and pressure gradient forces provide the dominant balances in the radial direction. However, the Coriolis force and viscous effects provide the primary balance for the azimuthal component.

Description: North Atlantic air-sea heat and freshwater flux data from several sources are used to estimate the conversion rate of water from one density to another throughout the range of sea surface density. This cross-isopycnal mass flux varies greatly over the ocean, with a maximum of 32.2 × 106 m3 s−1 at σ = 26.1 kg m−3 (toward greater densities) and a minimum of −7.6 × 106 m3 s−1 (toward lesser densities) at σ = 23.0 kg m−3. The air-sea fluxes force water to accumulate in three density bands: one at the lowest sea surface densities generated by heating; one centered near the density of subtropical mode water; and one spanning subpolar mode water densities. The transfer of water to the highest and lowest densities is balanced by mixing, which returns water to the middle density range, and also by boundary sources or sinks. Integrating the cross-isopycnal flux over all densities gives an annual average sinking of about 9 × 1O6 m3 s−1, which presumably escapes across the equator and must be balanced by a similar inflow. Comparison with estimates from tracer studies suggests that the renewal of tracer characteristics at a given density may occur without the existence of an annual average mass source at that density, because along- and cross-isopycnal mixing can renew a tracer without supplying mass.

Description: The formation of secondary ice in clouds, i.e. ice particles that are created at temperatures above the limit for homogeneous freezing without the direct involvement of a heterogeneous ice nucleus is one of the longest standing puzzles in cloud physics. Here we present comprehensive laboratory investigations on the formation of small ice particles upon the freezing of drizzle-sized cloud droplets levitated in an electrodynamic balance. Four different categories of secondary ice formation (bubble bursting, jetting, cracking, breakup) could be detected and their respective frequencies of occurrence as a function of temperature and droplet size are given. We find that bubble bursting occurs more often than droplet splitting. While we do not observe the shattering of droplets into many large fragments, we find that the average number of small secondary ice particles released during freezing is strongly droplet-size dependent and may well exceed unity for droplets larger than 300 μm in diameter. This leaves droplet fragmentation an important secondary ice process effective at temperatures around -10 °C in clouds where large drizzle droplets are present.

Description: Well-known problems trouble coupled general circulation models of the eastern Atlantic and Pacific Ocean basins. Model climates are significantly more symmetric about the equator than is observed. Model sea surface temperatures are biased warm south and southeast of the equator, and the atmosphere is too rainy within a band south of the equator. Near-coastal eastern equatorial SSTs are too warm, producing a zonal SST gradient in the Atlantic opposite in sign to that observed. The U.S. Climate Variability and Predictability Program (CLIVAR) Eastern Tropical Ocean Synthesis Working Group (WG) has pursued an updated assessment of coupled model SST biases, focusing on the surface energy balance components, on regional error sources from clouds, deep convection, winds, and ocean eddies; on the sensitivity to model resolution; and on remote impacts. Motivated by the assessment, the WG makes the following recommendations: 1) encourage identification of the specific parameterizations contributing to the biases in individual models, as these can be model dependent; 2) restrict multimodel intercomparisons to specific processes; 3) encourage development of high-resolution coupled models with a concurrent emphasis on parameterization development of finer-scale ocean and atmosphere features, including low clouds; 4) encourage further availability of all surface flux components from buoys, for longer continuous time periods, in persistently cloudy regions; and 5) focus on the eastern basin coastal oceanic upwelling regions, where further opportunities for observational–modeling synergism exist.

Description: Samples of female lllex argentinus were taken from the catch of a Japanese squid jigging vessel on the Patagonian Shelf during March 1986. Morphometrics of the somatic and reproductive organ systems and the histological structure of the mantle in relation to maturation were examined. The data suggest that growth and maturation occur simultaneously during most of the time that lllex argentinus females are on the feeding grounds. In a squid of a ‘standard’ mantle length the whole body mass increases relative to mantle length during maturation and growth of the reproductive organs. This is accompanied by a small but significant decrease in the relative mass of the mantle, head and viscera whilst the mass of the digestive gland remains constant. Although mantle mass of a ‘standard’ female squid decreases relative to mantle length with maturity this is not associated with degeneration of the mantle muscles. Energy and nutrient resources for maturation are apparently derived from the squid's food, not from reserves, and during the course of maturation there is an increasing shift of emphasis from somatic growth to production of gonad and accessory reproductive organs.

Description: The timing of spawning and recruitment in the squid Loligo forbesi in Scottish waters is described on the basis of data from three sources: monthly samples of squid caught by commercial trawls (1986–1988), egg masses found by fishermen (1987–1991), and statistical data on animals caught by research trawls (1978–1987). Spawning females were present in samples from December to June, with peak spawning occurring in March. Most records of egg masses were from these months, but eggs were also found in August and September. These results suggest that there is an extended spawning season. Small squid (≤100 mm dorsal mantle length) were rarely present in commercial samples, but were recorded in research samples almost all year round. Thus there appears to be more or less continuous recruitment into the catchable population. The results of the present study are consistent with published data from other parts of the geographic range in that there is a regular seasonal peak in spawning, and spawning adults disappear from the population in summer. Further interpretation of the life-cycle of this species is not justified on the basis of current knowledge, and more information is needed on migrations, geographical variation, and lifespan in Loligo forbesi.

Description: The distinguishing features of the common squid of British waters, Loligo forbesi, are summarized, and contrasted with those of L. vulgaris. The life-cycle and growth of L. forbesi are described, based on samples from trawl catches off Plymouth. This species seems to be an annual - young squid first appear in the trawl in late May, when their length is about 10 or 11 cm. Subsequent growth is rapid, and the males reach 30 cm and the females 25 cm by November. Spawning takes place mainly in December-January, but may continue into the spring. Neither sex survives beyond a single spawning season. Hatching of the spawn probably takes 30–40 days, and if the young squid taken in the trawl in late May hatched in the early part of the same year, a growth rate of about 25 mm/month would be required. Known growth rates for other species of Loligo are about 20 mm/month, so that indicated for L. forbesi does not seem to be impossibly high. The life-cycle is summarized in Fig. 8. There is also a summer spawning population, which grows to a rather smaller size at maturity, and which also seems to be annual. During the summer L. forbesi ranges throughout the English Channel and southern North Sea, particularly in inshore areas. In October the squid migrate farther offshore and tend to occupy the western part of the Channel. Values for total weight of squid/2 h trawl are given, on a monthly basis, for 1966–9. The largest quantities are usually taken in October and November, the highest single figure being 30.54 kg/2 h trawl, in November 1967.

Description: Cephalopod remains from the stomachs of a Risso's dolphin (Grampus griseus Cuvier, 1812, Cetacea) entangled in a fishing net off the Ligurian coast (central Mediterranean Sea) include squids Ancistroteuthis lichtensteini, Histioteuthis bonnellii, H. reversa and Todarodes sagittatus and the sepiolid Heteroteuthis dispar. All these cephalopods live in oceanic water including water over the steep continental slope where Risso's dolphin is frequently sighted. Histioteuthis reversa contributed 78% of the cephalopods by number, 81% of the wet weight and 73% of the dry weight and calorific value. The total calorific value of the cephalopods represented by lower beaks was 17,300 kj.

Description: The stomach contents of 235 specimens of the squid Sthenoteuthis oualaniensis (4·3–36·5 cm mantle length, ML) were examined. A detailed list of 60 species of prey, comprising young and adult squid, is given together with their frequency of occurrence and proportional contribution. The size and number of each food item was investigated. Three ontogenetic size-groups of S. oualaniensis were distinguished: I, fry and young (4–10 cm ML), micronektonic epipelagic plankton-eaters; II, transient critical size group (10–15 cm ML), converting from feeding on planktonic crustaceans and fish larvae to myctophid fishes; III, medium-sized (adult) nyctoepipelagic nektonic predators (15–36·5 cm ML), feeding primarily on myctophids and secondarily on squid. Myctophids (genera Symbolophorus, Myctophum and Hygophum) were the most abundant prey in the diet of adult S. oualaniensis from different parts of its distribution.

Description: In Britain Octopus vulgaris occurs on the Channel coast and only very rarely on other coasts. In Brittany and the Channel Islands it frequently makes its lair at low water, but on the English side of the Channel it does not come so close inshore except in abnormal years of high sea temperatures. The discovery of Octopus larvae of various sizes, from newly hatched to 6·0 mm. (mantle length), in plankton hauls taken to the north of the Channel Islands, proves that the species has a much longer planktonic life than hitherto supposed. The water circulation in the English Channel, as indicated by drift bottles, is admirably suited to the dispersal of larvae to our shores from breeding centres on the coasts of Brittany and the Channel Islands.

Description: The data presented provides new information on the distribution of Antarctic squids and on the summer diet of the emperor penguins. The diet of 58 adult emperor penguins (Aptenodytes forsteri) on the fast ice of the Drescher Inlet, Vestkapp Ice Shelf (72°52′S, 19°25′W) in the eastern Weddell Sea was investigated. Prey consisted principally of squid, fish, krill, amphipods and isopods. Squids were identified by the lower beaks and allometric equations were used to estimate the squid biomass represented. Beaks occurred in 93% of the stomach samples. Each sample contained a mean of 27 beaks (range 1–206). Ninety-two percent of the squids could be identified by the lower beaks and belonged to four families (Onychoteuthidae, Psychroteuthidae, Neoteuthidae and Gonatidae). The most abundant squid was Psychroteuthis glacialis which occurred in 52 samples with lower rostral lengths (LRL) ranging from 1.4–7.2 mm. Forty-five samples contained Alluroteuthis antarcticus (LRL range 1.8–5.8 mm), 17 Kondakovia longimana (LRL range 4–12.1 mm), and four Gonatus antarcticus (LRL range 4.1–6.1 mm). In terms of biomass K. longimana was the most important species taken by the penguins comprising 50% of total estimated squid wet mass (245348 g) in 1990 and 48% in 1992 (154873 g). However, if only fresh beaks were considered for estimations of squid consumption, i.e. beaks that have been accumulated for not longer than 5–6 days in the stomachs, squid diet was of minor importance. Then total squid wet mass accounted for only 4809 g in 1990 and 5445 g in 1992 which implies that one penguin took c.30 g squid d−1 with P. glacialis and A. antarcticus being the most important by mass. The prey composition suggests that emperor penguins take squid at the steep slope regions of the eastern Weddell Sea.

Description: Twenty-four out of 240 fishes caught by bottom lines at 366–3333 m had something in their stomachs. Stomach contents included parts of cephalopods, fish, cetaceans and bottom-living invertebrates, thin rubber sheet and terrestrial mammal bones. The material provides evidence that four species of cephalopod are at least partially demersal and suggests a means by which the tapeworm Phyllobothrium could pass from its secondary to its primary host. During the five biological cruises of R.R.S. ‘Discovery’ between 1967 and 1971 a total of 31 bottom lines with 1483 hooks were fished in depths of water between 366 and 3333 m. The stomachs of the 240 fish caught were examined and 216 (90%) proved to be empty. The high incidence of empty stomachs is thought to be due to frequent loss of food during the ascent from great depths and accounts for our poor knowledge of the feeding habits of demersal fish living at depths exceeding 400 m. The present collection of food from 25 stomachs (24 from ‘Discovery’ collections and one from a fish caught by Mr G. R. Forster from R. V. ‘Sarsia’) of fish belonging to 11 species (Table 1) probably gives little indication of the usual diet of the fish concerned, but its nature prompts some useful speculation and the rarity of such observations justifies placing them on record (Bigelow & Schroeder, 1948; Marshall, 1954). All the fish were caught on lines which lay on the bottom for several hours and it is our firm belief that they were hooked while on or very near the bottom.

Description: The genus Illex is likely to constitute a large portion of the annual world ommastrephid squid catches (Roper et al, 1984), even though specific official statistics are difficult to obtain. The broad-tail short-fin squid Illex coindetii is a widespread species ranging from the western to the eastern Atlantic (Roper et al., 1984) and east through the whole Mediterranean Sea (Mangold & Boletzky, 1987). Usually a by-catch of important fisheries, it is caught mainly by trawlers. Although its economic value is lower than that of other squid species (i.e. Loligo spp.), in the Sicilian Channel Illex coindetii may represent a valuable resource due to its abundance. In Italian waters, the available statistics (Cingolani et al., 1986) report that 2680 tonnes of ommastrephid squid were landed in 1982 (0.5% of the total landed catch). The main component of these was landed in Sicily (2183 tonnes), a consistently large part of which was no doubt Illex coindetii (Ragonese & Jereb, 1992). The catches came mainly from southern Sicilian waters (Sicilian Channel) where one of the major Mediterranean landing places is in Mazara del Vallo. Large trawlers (up to 200 gross tonnage) usually carry out long fishing trips (15–20 d), and Illex coindetii is caught mainly by those targeting Parapenaeus longirostris and Merluccius merluccius (Jereb & Ragonese, 1991).

Description: The natural feeding of the two most abundant ommastrephid squid (Cephalopoda: Ommastrephidae) in Galician waters was studied and compared. A sample of 334 stomach contents of Todaropsis eblanae (34–222 mm ML) and 267 stomach contents of Illex coindetii (50–379 mm ML) caught by commercial trawlers was examined. A total of 21 (T. eblanae) and 23 (I. coindetii) different prey items, belonging to three zoological groups (Teleostei, Crustacea and Cephalopoda), were taken by these cephalopods. However, 43% of the T. eblanae diet comprised only one fish species, Micromesistius poutassou. The diet of these squid species was significantly influenced by the geographical area (both species), size (T. eblanae) and maturation (I. coindetii). Feeding rate of both species decreased with size, but the percentage of stomachs with food remains increased in maturing and mature females. Weight of prey captured was dependent on available prey sizes and, in small individuals, maximum prey weight was very close to the squid weight. Both squid species are mainly neritic nekto-benthic predators, but I. coindetii seems to have a broader and more pelagic diet.

Description: The ratio of global mean surface air temperature change to cumulative CO2 emissions, referred to as transient climate response to cumulative CO2 emissions (TCRE), has been shown to be approximately constant on centennial time scales. The mechanisms behind this constancy are not well understood, but previous studies suggest that compensating effects of ocean heat and carbon fluxes, which are governed by the same ocean mixing processes, could be one cause for this approximate constancy. This hypothesis is investigated by forcing different versions of the University of Victoria Earth System Climate Model, which differ in the ocean mixing parameterization, with an idealized scenario of 1% annually increasing atmospheric CO2 until quadrupling of the preindustrial CO2 concentration and constant concentration thereafter. The relationship between surface air warming and cumulative emissions remains close to linear, but the TCRE varies between model versions, spanning the range of 1.2°–2.1°C EgC−1 at the time of CO2 doubling. For all model versions, the TCRE is not constant over time while atmospheric CO2 concentrations increase. It is constant after atmospheric CO2 stabilizes at 1120 ppm, because of compensating changes in temperature sensitivity (temperature change per unit radiative forcing) and cumulative airborne fraction. The TCRE remains approximately constant over time even if temperature sensitivity, determined by ocean heat flux, and cumulative airborne fraction, determined by ocean carbon flux, are taken from different model versions with different ocean mixing settings. This can partially be explained with temperature sensitivity and cumulative airborne fraction following similar trajectories, which suggests ocean heat and carbon fluxes scale approximately linearly with changes in vertical mixing.

Description: The Weddell Sea polynya is a large opening in the open-ocean sea ice cover associated with intense deep convection in the ocean. A necessary condition to form and maintain a polynya is the presence of a strong subsurface heat reservoir. This study investigates the processes that control the stratification and hence the buildup of the subsurface heat reservoir in the Weddell Sea. To do so, a climate model run for 200 years under preindustrial forcing with two eddying resolutions in the ocean (0.25° CM2.5 and 0.10° CM2.6) is investigated. Over the course of the simulation, CM2.6 develops two polynyas in the Weddell Sea, while CM2.5 exhibits quasi-continuous deep convection but no polynyas, exemplifying that deep convection is not a sufficient condition for a polynya to occur. CM2.5 features a weaker subsurface heat reservoir than CM2.6 owing to weak stratification associated with episodes of gravitational instability and enhanced vertical mixing of heat, resulting in an erosion of the reservoir. In contrast, in CM2.6, the water column is more stably stratified, allowing the subsurface heat reservoir to build up. The enhanced stratification in CM2.6 arises from its refined horizontal grid spacing and resolution of topography, which allows, in particular, a better representation of the restratifying effect by transient mesoscale eddies and of the overflows of dense waters along the continental slope.

Description: The current generation of marine biogeochemical modules in Earth system models (ESMs) considers mainly the effect of marine biota on the carbon cycle. We propose to also implement other biologically driven mechanisms in ESMs so that more climate-relevant feedbacks are captured. We classify these mechanisms in three categories according to their functional role in the Earth system: (1) "biogeochemical pumps", which affect the carbon cycling; (2) "biological gas and particle shuttles", which affect the atmospheric composition; and (3) "biogeophysical mechanisms", which affect the thermal, optical, and mechanical properties of the ocean. To resolve mechanisms from all three classes, we find it sufficient to include five functional groups: bulk phyto- and zooplankton, calcifiers, and coastal gas and surface mat producers. We strongly suggest to account for a larger mechanism diversity in ESMs in the future to improve the quality of climate projections.

Description: Introduction. Today, most countries have accepted a 2 degrees Celsius temperature increase above preindustrial levels as the maximum tolerable limit for global warming. An exceedance probability of below 20 percent for this limit implies an emission budget of less than 250 GtC from 2000 until 2049, however, extrapolating from current global CO2 emissions, this budget will only last until 2024. This sobering math should wake us up to the reality that all options, including climate engineering, need to be considered to address climate change. Climate engineering options can be classified broadly into two categories: solar radiation management and carbon dioxide removal measures. Solar radiation management schemes seek to decrease the incoming solar radiation or to increase the reflection of incoming solar radiation. These approaches can generate fast climate responses, but do not immediately address the cause of the problem. Carbon dioxide removal measures seek to decrease atmospheric carbon concentrations by enhancing or substituting natural carbon sinks. The terrestrial carbon sink can be enhanced by means of forestation; the oceanic sink may, in some regions, be enhanced by means of fertilization, for example by artificially enhanced upwelling of macronutrients or by purposeful addition of the micronutrient iron; the mineral carbon sink can be enhanced by means of chemically accelerated weathering. Some analysts have expressed doubts about the potential of mitigating climate change by sink enhancement, because of concerns about whether carbon can be stored permanently. Nevertheless, terrestrial vegetation sinks have entered the Kyoto Protocol (KP) as offsets for anthropogenic greenhouse gas emissions, but ocean sinks have not.

Description: Serological methods for prey identification have been applied to the gut contents of a field sample of 100 Eledone cirrhosa from the Moray Firth and 7 from the Sound of Jura. Protein extracts from the crop were electrophoresed (Laurell rockets) into antisera raised against potential crustacean prey species, Cancer pagurus, Carcinus maenas, Crangon crangon, Liocarcinus spp. and Nephrops norvegicus. The strengths of the resulting reactions were evaluated on the basis of peak height, staining density and the number of homocentric peaks of immune precipitates. Sixty-two of the 100 Moray Firth animals gave a positive reaction to one or more of the antisera and the most critical assessment of the results ranked the incidence of the prey as Liocarcinus (15) 〉 Nephrops (10) 〉 Cancer (8) 〉 Crangon (2) 〉 Carcinus (0), a total of 35 reactions from 28 animals. Absence of a reaction in the 38 animals from which appreciable sample volumes were also obtained indicates that alternative prey had been consumed. Of the 7 Sound of Jura animals, 4 gave a positive reaction to Nephrops alone. A total of only 24 animals gave any indication of diet by visual recognition of remains. The value and difficulties of the methodology are discussed.

Description: The growth of the octopus Eledone cirrhosa has been studied in a population from the North Sea off Aberdeen. Data are presented for the growth of individuals isolated in aquarium conditions; the growth of size classes in thefieldpopulation; and preliminary information on the growth relationships of gonad, somatic, cardiac and brain components of the body. At 15 °C Eledone cirrhosa is capable of growing from 10 to 1000 g in 270 days. From octopuses which feed readily in captivity, weight specific growth rates of up to about 3–5 % day-1 for animals of 100 g body weight are recorded, falling to a maximum of about 1–5 % day-1 at body sizes above 500 g. Females stop growing when sexually mature, but in the sample captured they were consistently larger than males, a feature which may account for the 7:1 bias towards the incidence of females. On a wet-weight basis, the mean food incorporation into growth is 37 % of the food ingested, which is 49% of the gross weight of crabs killed. Field data for 1978/79 suggest that animals recruited to the population at the beginning of the year grew steadily until December, overwintered without growing, then grew rapidly for several months in the subsequent year before disappearing from the samples. The estimated average age of those animals and by implication, the life span, is 20 months.

Description: Samples of male Illex argentinus were taken from the catch of a Japanese squid jigging vessel on the Patagonian Shelf during March 1986 and an analysis was carried out on the morphometrics of the somatic and reproductive organ systems in relation to maturation. The data show that growth and maturation occurred simultaneously during most of the time that Illex argentinus males were on the feeding grounds over the southern Patagonian Shelf. In a squid of a ‘standard’ mantle length the whole body mass increased relative to mantle length during maturation and this could be attributed to the increase in mass of the reproductive and accessory reproductive organs. During maturation the mantle and digestive gland mass showed no significant change relative to mantle length. The mass of the head increased and the mass of the viscera decreased relative to mantle length. In male Illex argentinus, as in the female, the energy and nutrient resources for maturation are derived from the squid's food and during the course of maturation there is an increasing shift of emphasis from somatic growth to production of gonad and accessory reproductive organs. The proportional investment of body mass in reproductive and accessory reproductive organs predicted for a fully mature male Illex argentinus was less than half that of the female.

Description: The Cycladic blueschist belt, Greece, is mostly submerged below sea level and regional correlations are difficult to establish. Marbles are widespread within the belt and locally used as marker horizons to subdivide monotonous schist sequences. However, owing to the lack of distinctive petrographic characteristics, the marbles have not been used for island-to-island correlations. This study aims to investigate the potential of Sr-, C- and O-isotope compositions of marbles as a tool for unravelling the litho- and/or tectonostratigraphic relationships across the Cycladic islands, and as a proxy for the time of sediment formation. For this purpose, we have studied metamorphic carbonate rocks from the islands of Tinos, Andros, Syros, Sifnos and Naxos. Identical 87Sr/86Sr values for certain marble horizons occurring on Tinos, Andros and Sifnos are interpreted to document coeval regional carbonate precipitation. The 87Sr/86Sr values of the apparently least altered samples intersect the seawater curve multiple times within the most likely time interval of original carbonate precipitation (〈 240 Ma; as indicated by previously published ion probe U–Pb zircon data) and thus an unequivocal age assignment is not possible. Very broad temporal correlations are possible, but more subtle distinctions are not feasible. On Andros, the overlapping Sr-isotope values of marbles representing the lowest and highest parts of the metamorphic succession are in accordance with a model suggesting isoclinal folding or thrusting of a single horizon, or very fast sedimentation. In contrast, distinct 87Sr/86Sr values for samples from Tinos, representing different levels of the metamorphic succession, suggest that these rocks represent a temporal succession and not the tectonic repetition of a single horizon. Based on Sr-,O- and C-isotope characteristics alone the time equivalence ofmarbles occurring on different islands could not be documented unambiguously. However, by using various combinations of these parameters, some occurrences can be discriminated from the overall sample population. The new data further accentuate the general potential of coupled Sr-, C- and O-isotope characteristics for identification of archaeological provenance and complement existing datasets for Aegean marbles.

Description: During a study based on catches taken in the northern North Sea by selected Scottish fishing boats during 1985–1992, large numbers of the normally rare short-fin squid, Todaropsis eblanae (Cephalopoda: Ommastrephidae), were recorded in 1987 and 1990. Our findings, supported by data obtained from plankton/young fish surveys in 1988 and 1989, suggest that in northern waters Todaropsis eblanae generally mates and spawns during late summer and early autumn (June-November). Successful hatching events appear to occur during October-March, producing juvenile (stage I) squid in the early part of the year (January-June). Estimations of maximum male reproductive output and female fecundity were up to 130 spermatophores and ~28,000 eggs per individual, respectively.

Description: Statoliths of Loligo gahi were sampled in the fishery region 45–47°S on the Patagonian shelf during September 1989. Peculiarities of the growth zones in the ground statoliths of adults are described. Maximum age of large maturing and mature females (130–160 mm of mantle length, ML) was estimated to be 325–345 d, that of large mature males (250–290 mm ML) ranged from 360 to 396 d. The squid Loligo gahi d'Orbigny, 1835, occurs in temperate shelf and upper slope waters of the Atlantic and Pacific coasts of South America and is caught commercially by the international fleet in the southern part of the Patagonian shelf within the Falkland Islands Interim Conservation Zone (FICZ) (Roper et al., 1984; Csirke, 1987). Occasionally, dense shoals of L. gahi appear in the fishery region 45–47°S off the Exclusive Economic Zone of Argentina (EEZA) and have been caught in significant numbers by trawlers at depths of 120–150 m in September-October (Chesheva, 1990). Loligo gahi is a medium sized loliginid; in Falkland waters males attain 350 mm ML, females 210 mm ML (Hatfield, 1991), while in the fishery region 45–47°S maximum size is 260 mm and 160 mm, respectively (Chesheva, 1990). Patterson (1988) revealed two Falkland spawning stocks of L. gahi of unclear status, spring-spawners and autumn-spawners (austral seasons) and pointed out that the life span of squid of each stock lasted ~1 y. Recently Hatfield (1991) used statoliths to elucidate Patterson's (1988) estimations of age and growth of Falkland stocks of L. gahi and confirmed the 1-y duration of L. gahi's life span.

Description: In this paper we examine the steady, two-dimensional convective motion which occurs in a horizontal circular cylinder whose wall is non-uniformly heated. One observes several qualitatively different physical phenomena depending on the wall temperature distribution and the value of the Rayleigh number. The low-Rayleigh-number behaviour for the single convective cell heated from below is related to the classical Rayleigh stability problem. The critical Rayleigh number for the single circular cell is approximately five times the value for Rayleigh's multi-celluar configuration. The flow which exhibits a nearly parabolic velocity profile near the critical Rayleigh number, gradually changes to a rigidly-rotating-core behaviour as the Rayleigh number increases. The speed of core rotation is a function of the Prandtl number, whereas the boundary-layer thickness is primarily a function of the Rayleigh number. When the heating is from side to side, the solution shows that as the Rayleigh number increases the core motion is progressively arrested leaving a narrow circulating band of fluid adjacent to the wall. An oblique heating produces a hybrid phenomenon, a low-Rayleigh-number behaviour which is characteristic of the sideways heating case and a high-Rayleigh-number interior motion characteristic of the bottom heating case. To determine the core motion in the high-Rayleigh-number limit, Batchelor's work concerning the uniqueness of incompressible, exactly steady, closed streamline flows with small viscosity is extended to include flows with small thermal conductivity.

Description: The ommastrephid squids are large active animals occurring in most of the world's oceans. Luminous organs or bioluminescence have been observed only in members of the subfamily Sthenoteuthinae, containing the genera Ornithoteuthis, Symplectoteuthis (= Eucleoteuthis), Hyaloteuthis, Ommastrephes and Dosidicus. The light organs of Ommastrephes pteropus are small sub-spherical bodies randomly distributed over the ventral surface of the mantle, head, arms and tentacles (Roper, 1963) and are aggregated dorsally to form a large luminous patch (Clarke, 1965). Relatively little is known about the organs, capabilities and biochemistry of luminescence in cephalopods (Harvey, 1952; Herring, in Press), and the size of the light organ and availability of O. pteropus provide an unusual opportunity for such studies. Although among the molluscs the luminescent systems of the gastropod Latia and the bivalve Pholas have been partially characterized (Shimomura & Johnson, 1968; Henry, Isambert & Michelson, 1970, 1973) the only cephalopod system which has been investigated to date is that of the enoploteuthid Watasenia scintillans (Goto et al., 1974; Inoue et al., 1975). This investigation examines the anatomy and biochemistry of the dorsal light organ of O. pteropus, which differs markedly in these respects from the brachial organs of Watasenia.

Description: Seasonal variability in pathways of warm water masses toward the Kangerdlugssuaq Fjord-Glacier system (KF/KG), southeast Greenland, is investigated by backtracking Lagrangian particles seeded at the fjord mouth in a high-resolution regional ocean model simulation in the ice-free and the ice-covered seasons. The waters at KF are a mixture of Atlantic-origin water advected from the Irminger Basin (FF for Faxaflói), the deep waters from the Denmark Strait and the waters from the Arctic Ocean, both represented by the Kögur section (KO). Below 200m depth, the warm water is a mixture of FF and KO water masses, and is warmer in winter than in summer. We find that seasonal differences in pathways double the fraction of FF particles in winter, causing the seasonal warming and salinification. Seasonal temperature variations at the upstream sections (FF and KO) have a negligible impact on temperature variations near the fjord. Successful monitoring of heat flux to the fjord therefore needs to take place close to the fjord, and cannot be inferred from upstream conditions.

Description: Squids (teuthoids) fall into two distinct groups according to their density in sea water. Squids of one group are considerably denser than sea water and must swim to stop sinking; squids in the other group are nearly neutrally buoyant. Analyses show that in almost all the neutrally buoyant squids large amounts of ammonium are present. This ammonium is not uniformly distributed throughout the body but is mostly confined to special tissues where its concentration can approach half molar. The locations of such tissues differ according to the species and developmental stage of the squid. It is clear that the ammonium-rich solution are almost isosmotic with sea water but of lower density and they are present in sufficient volume to provide the main buoyancy mechanism of these squids. A variety of evidence is given which suggests that squids in no less than 12 of the 26 families achieve near-neutral buoyancy in this way and that 14 families contain squids appreciably denser than sea water [at least one family contains both types of squid]. Some of the ammonium-rich squids are extremely abundant in the oceans.

Description: This communication presents the first records of mated female Loligo gahi in Falkland Island waters. In October 1993 fully mature mated female L. gahi were identified in samples taken from the commercial fishery in waters east of Lively Island, East Falkland, at depths of 145–174 m. Spermatophores were found in both the mantle cavity and buccal sites of deposition. These records, combined with past records of spent females, suggest spawning periods in late October/early November and April/May. These concur with two of the three periods of spawning suggested from previous studies of juvenile and adult L. gahi.

Description: Age, growth and maturation of Loligo vulgaris were studied by examination of growth increments within statoliths of 294 specimens (mantle length, ML, ranging from 31 to 498 mm) caught on the west Saharan shelf between 1985 and 1988. Maximum age was 335 d (290 mm ML) for females, and 396 d (498 mm ML) for males. Growth rates varied considerably among individuals with a greater range in males. Sexual dimorphism in length was apparent after about 210 d. Males and females diverged considerably in weight, with males reaching a greater weight after about 180 d. Growth in length between 124 and 396 d was best described by a power function, while growth in weight was best described by the Gompertz function. Males started maturing at 180–210 d and mature males ranged in age between 250 and 396 d; while females started maturing at 240–270 d and mature females ranged between 285 and 335 d. Loligo vulgaris hatched throughout the year with two distinct peaks; in winter (December - early March) and summer (June-July). The life cycle of L. vulgaris populations on the west Saharan shelf lasts ~1 y, with large males (〉450 mm ML) living slightly longer.

Description: The reproductive strategies of male and female Loligo forbesi Steenstrup, 1856 were investigated from samples obtained from commercial catches and research cruises in Irish waters. In females maturity increased with size, but in males two modes in the size at maturity were identified, with approximately 40% mature at small size (180–200 mm mantle length), and the remainder mature at 〉250 mm mantle length. The difference in estimated age of the two modes of mature males was small, so size differences were probably due to different growth rates. Growth and maturation proceeded together in both sexes over much of the life-cycle. The effect of maturation on relative growth of somatic tissues was examined using analysis of covariance and multivariate regressions. In males there was a significant decline in total mass, and in mass of mantle, head and viscera, relative to mantle length with maturation. In females total mass was not significantly affected by maturation, but relative masses of head, mantle and viscera declined with maturation, indicating that energy was diverted from somatic growth to gonad production. Potential fecundity estimates were obtained by counting eggs and ova in the oviduct and ovary of mature females and were in the range 2500 to 10,500 (mean 5800). Fecundity was positively related to mantle length. The ovaries of mature females contained a range of egg sizes and developmental stages, indicating that spawning probably occurs intermittently.

Description: The vertical distribution of the hydromedusa Aequorea forskalea was investigated using observations from the research submersible 'Jago' collected during 36 dives off the west coast of southern Africa during November 1997 and April 1999. The mean population depth of Aequorea? forskalea deepened with increasing sea surface temperature. We suggest that this behaviour enables individuals to avoid offshore advection, to minimize spatial overlap with other large medusae and to maintain their position over the middle of the shelf.

Description: The effect of medium dissolved-oxygen tension on the molar growth yield, respiration and cytochrome content of Beneckea natriegens in chemostat culture (D 0·37 hr-1) was examined. The molar growth yield (Y), the specific rate of oxygen (qo2) and glucose consumption, and the specific rate of carbon dioxide evolution were independent of the dissolved-oxygen tension above a critical value (〈 2 mmHg). However, the potential respiration rate increased with reduction in the dissolved-oxygen tension at values of the dissolved-oxygen tension well above the critical value. Changes in the cytochrome content occurred at dissolved-oxygen tensions well above the critical value. An increase in cytochrome c relative to cytochrome b was observed as the dissolved-oxygen tension was decreased. Reduction of the dissolved-oxygen tension to less than 1 mmHg caused a switch to fermentative metabolism shown by the apparent rise in Y o2 and decrease in the molar growth yield from glucose. At this point the potential respiration rate (q o2) increased to its highest value, while the cytochrome pattern reverted to that observed at dissolved-oxygen tensions above 96 mmHg. There appeared to be no correlation between cytochrome content, potential q o2, in situ q o2, and cyanide sensitivity of the organism at various dissolved-oxygen tensions.

Description: In Ammassalik, in southeast Greenland, downslope winds can reach hurricane intensity and represent a hazard for the local population and environment. They advect cold air down the ice sheet and over the Irminger Sea, where they drive large ocean–atmosphere heat fluxes over an important ocean convection region. Earlier studies have found them to be associated with a strong katabatic acceleration over the steep coastal slopes, flow convergence inside the valley of Ammassalik, and—in one instance—mountain wave breaking. Yet, for the general occurrence of strong downslope wind events, the importance of mesoscale processes is largely unknown. Here, two wind events—one weak and one strong—are simulated with the atmospheric Weather Research and Forecasting (WRF) Model with different model and topography resolutions, ranging from 1.67 to 60 km. For both events, but especially for the strong one, it is found that lower resolutions underestimate the wind speed because they misrepresent the steepness of the topography and do not account for the underlying wave dynamics. If a 5-km model instead of a 60-km model resolution in Ammassalik is used, the flow associated with the strong wind event is faster by up to 20 m s−1. The effects extend far downstream over the Irminger Sea, resulting in a diverging spatial distribution and temporal evolution of the heat fluxes. Local differences in the heat fluxes amount to 20%, with potential implications for ocean convection.

Description: Predicting tropical cyclone (TC) activity becomes more important every year while the understanding of what factors impact them continues to be complicated. El Niño–Southern Oscillation (ENSO) is one of the primary factors impacting the activities in both the Pacific and the Atlantic, but an extensive examination of the fluctuation in this system has yet to be studied in its entirety. This article analyzes the ENSO impacts on the Atlantic tropical cyclone activity during the assessed warm and cold years to show the dominant centennial-scale variation impact. This study looks to plausibly link this variation to the Southern Ocean centennial variability, which is rarely mentioned in any factors affecting the Atlantic tropical cyclone activity. This centennial variability could be used to enhance future work related to predicting tropical cyclones.

Description: Antarctic Bottom Water flows into the western North Atlantic across the equator, shifting from the western side to the eastern side of the trough between the American continents and the Mid-Atlantic Ridge as it continues north. This is puzzling because such large-scale motion is thought to be controlled by dynamics that disallows an eastern boundary current. Previous explanations for the transposition involve a (necessarily small-scale) density current that changes sides because of the change in sign of rotation across the equator, or a topographic effect that changes the sign of the effective mean vorticity gradient and thus requires an eastern boundary current. Here an alternative explanation for the overall structure of bottom flow is given. A source of mass to a thin bottom layer is assumed to upwell uniformly across its interface into a less dense layer at rest. A simple formula for the magnitude of the upwelling and thickness of the layer is derived that depends on the source strength to the bottom layer. For a strong enough source, the bottom layer thickness is zero along a grounding curve that separates the bottom water from the western boundary and confines it to the east. A band of recirculating interior flow occurs, supplied by an isolated northern and western boundary current. Similar structures appear to exist in the Antarctic Bottom Water of the western North Atlantic.

Description: Results of a three-dimensional primitive equation model are presented simulating turbulent mesoscale motions in the seasonal thermocline on an f plane. The model is based on a hybrid vertical coordinate scheme and conserves isopycnic potential vorticity. Mesoscale turbulence is modeled in terms of an unstable potential vorticity front. The model integration starts from a purely zonal, 60-km-wide geostrophically balanced jet, on which is superimposed a small initial perturbation. The most unstable mode exhibits a wavelength of 85 km and is driven by a mixed type of instability. Characteristic dynamical ingredients of the wave are enhanced cyclonic and anticyclonic relative vorticity in the troughs and the ridges, respectively, due to the curvature of the flow. Vertical motion of up to 10 m d−1 occurring downstream of the ridges (downwelling) and downstream of the troughs (upwelling) is driven by geostrophic advection of relative vorticity. The contrast of static stability across the front is changing during amplification of the instability: in troughs the stability is decreasing whereas in ridges it is increasing. The density field exhibits local anomalies of the isopycnals' depths (bumps) due to the ageostrophic cross-jet advection of potential vorticity streamers wound up in cyclones and anticyclones. Locally, the potential vorticity gradients are enhanced, creating a multiple front structure. The model results support observations and findings of earlier atmospheric and oceanic models. It is emphasized that mesoscale turbulent structures may have a profound influence on primary productivity, mixed-layer, and internal wave dynamics.

Description: Downward wave coupling (DWC) is an important process that characterizes the dynamical coupling between the stratosphere and troposphere via planetary wave reflection. A recent modeling study indicated that natural forcing factors, including sea-surface temperature variability and quasi-biennial oscillation, influence DWC and the associated surface impact in the Northern Hemisphere (NH). In light of this, we further investigate how DWC in the NH is affected by anthropogenic forcings, using a fully coupled chemistry-climate model CESM1 (WACCM). The results indicate that the occurrence of DWC is significantly suppressed in the future, starting later in the seasonal cycle, with more events concentrated in late winter (February-March). The future decrease in DWC events is associated with enhanced wave absorption in the stratosphere due to increased greenhouse gases. The enhanced wave absorption is manifest as more absorbing types of stratospheric sudden warmings, with more events concentrated in early winter. This early winter condition leads to a delay in the development of the upper stratospheric reflecting surface, resulting in a shift in the seasonal cycle of DWC towards late winter. The tropospheric responses to DWC events in the future exhibit different spatial patterns compared to those of the past. In the North Atlantic sector, DWC-induced circulation changes are characterized by a poleward shift and an eastward extension of the tropospheric jet, while in the North Pacific sector, the circulation changes are characterized by a weakening of the tropospheric jet. These responses are consistent with a change in the pattern of DWC-induced synoptic-scale eddy-mean flow interaction.

Description: Decadal variabilities in Indian Ocean subsurface ocean heat content (OHC; 50–300 m) since the 1950s are examined using ocean reanalyses. This study elaborates on how Pacific variability modulates the Indian Ocean on decadal time scales through both oceanic and atmospheric pathways. High correlations between OHC and thermocline depth variations across the entire Indian Ocean Basin suggest that OHC variability is primarily driven by thermocline fluctuations. The spatial pattern of the leading mode of decadal Indian Ocean OHC variability closely matches the regression pattern of OHC on the interdecadal Pacific oscillation (IPO), emphasizing the role of the Pacific Ocean in determining Indian Ocean OHC decadal variability. Further analyses identify different mechanisms by which the Pacific influences the eastern and western Indian Ocean. IPO-related anomalies from the Pacific propagate mainly through oceanic pathways in the Maritime Continent to impact the eastern Indian Ocean. By contrast, in the western Indian Ocean, the IPO induces wind-driven Ekman pumping in the central Indian Ocean via the atmospheric bridge, which in turn modifies conditions in the southwestern Indian Ocean via westward-propagating Rossby waves. To confirm this, a linear Rossby wave model is forced with wind stresses and eastern boundary conditions based on reanalyses. This linear model skillfully reproduces observed sea surface height anomalies and highlights both the oceanic connection in the eastern Indian Ocean and the role of wind-driven Ekman pumping in the west. These findings are also reproduced by OGCM hindcast experiments forced by interannual atmospheric boundary conditions applied only over the Pacific and Indian Oceans, respectively.

Description: The decadal-scale variability in the tropical Pacific has been analyzed herein by means of observations and numerical model simulations. The two leading modes of the sea surface temperature (SST) variability in the central western Pacific are a decadal mode with a period of about 10 yr and the ENSO mode with a dominant period of about 4 yr. The SST anomaly pattern of the decadal mode is ENSO like. The decadal mode, however, explains most variance in the western equatorial Pacific and off the equator. A simulation with an ocean general circulation model (OGCM) forced by reanalysis data is used to explore the origin of the decadal mode. It is found that the variability of the shallow subtropical–tropical overturning cells is an important factor in driving the decadal mode. This is supported by results from a multicentury integration with a coupled ocean–atmosphere general circulation model (CGCM) that realistically simulates tropical Pacific decadal variability. Finally, the sensitivity of the shallow subtropical–tropical overturning cells to greenhouse warming is discussed by analyzing the results of a scenario integration with the same CGCM.

Description: Oceanic ecosystems altered by interdecadal climate variability may provide a feedback to the physical climate by phytoplankton affecting heat fluxes into the upper ocean and dimethylsulfide fluxes into the atmosphere

Description: The circulation of the northeastern Atlantic Ocean at intermediate depths is characterized by watermass transformation processes that involve Iceland–Scotland Overflow Water (ISOW) from the northeast, Labrador Sea Water (LSW) from the west, and Mediterranean Water from the south. Field observations were carried out with 89 eddy-resolving floats (RAFOS and MARVOR types). The data coverage achieved is remarkably high and enables a comprehensive study of the eastern basins between Iceland and the Azores. The trajectories show typical pathways of the water masses involved and the role that the complex bottom topography plays in defining them. The ISOW paths tend to lean against the slopes of the Reykjanes Ridge and Rockall Plateau. Westward escapes through multiple gaps in the ridge are possible, superimposed on a sustained southward flow in the eastern basin along the Mid-Atlantic Ridge. LSW pathways leading to the eastern basins are subject to high variability in flow direction and eddy activity. In addition to a selection of characteristic trajectories, maps of the horizontal distributions of Lagrangian eddy kinetic energy and integral time scales are presented. These reveal distinct areas of intensified mixing in the Iceland Basin, as well as the sharp contrast between the subpolar and subtropical dynamics. A self-contained eddy detection scheme is applied to obtain statistics on individual eddy properties and their abundance. It is suggested that much of the intensified mixing can be related to cyclonic activity, particularly in the subpolar region.

Description: The current system east of the Grand Banks was intensely observed by World Ocean Circulation Experiment (WOCE) array ACM-6 during 1993–95 with eight moorings, reaching about 500 km out from the shelf edge and covering the water column from about 400-m depth to the bottom. More recently, a reduced array by the Institut für Meerskunde (IfM) at Kiel, Germany, of four moorings was deployed during 1999–2001, focusing on the deep-water flow near the western continental slope. Both sets of moored time series, each about 22 months long, are combined here for a mean current boundary section, and both arrays are analyzed for the variability of currents and transports. A mean hydrographic section is derived from seven ship surveys and is used for geostrophic upper-layer extrapolation and isopycnal subdivision of the mean transports into deep-water classes. The offshore part of the combined section is dominated by the deep-reaching North Atlantic Current (NAC) with currents still at 10 cm s−1 near the bottom and a total northward transport of about 140 Sv (Sv ≡ 106 m3 s−1), with the details depending on the method of surface extrapolation used. The mean flow along the western boundary was southward with the section-mean North Atlantic Deep Water outflow determined to be 12 Sv below the σθ = 27.74 kg m−3 isopycnal. However, east of the deep western boundary current (DWBC), the deep NAC carries a transport of 51 Sv northward below σθ = 27.74 kg m−3, resulting in a large net northward flow in the western part of the basin. From watermass signatures it is concluded that the deep NAC is not a direct recirculation of DWBC water masses. Transport time series for the DWBC variability are derived for both arrays. The variance is concentrated in the period range from 2 weeks to 2 months, but there are also variations at interannual and longer periods, with much of the DWBC variability being related to fluctuations and meandering of the NAC. A significant annual cycle is not recognizable in the combined current and transport time series of both arrays. The moored array results are compared with other evidence on deep outflow and recirculation, including recent models of different types and complexity.

Description: This study investigates and accounts for the influence of various ice cloud parameters on the retrieval of the surface solar radiation budget (SSRB) from reflected flux at the top of the atmosphere (TOA). The optical properties of ice clouds depend on ice crystal shape, size distribution, water content, and the vertical profiles of geometric and microphysical structure. As a result, the relationship between the SSRB and TOA-reflected flux for an ice cloud atmosphere is more complex and differs from that for water cloud and cloudless atmospheres. The sensitivities of the relationship between the SSRB and TOA-reflected flux are examined with respect to various ice cloud parameters. Uncertainties in the retrieval of the SSRB due to inadequate knowledge of various ice cloud parameters are evaluated thoroughly. The uncertainty study is concerned with both pure ice clouds and multiphase clouds (ice cloud above water cloud). According to the magnitudes of errors in the SSRB retrieval caused by different input variables, parameterized correction terms were introduced. If the input variables are known accurately, errors in the retrieval of the SSRB under a wide range of ice cloud conditions are expected to diminish substantially, to less than 10 W m−2 for 91% of the simulated ice cloud cases. In comparison, the same accuracy may be attained for only 19% of the retrievals for the same ice cloud cases using the retrieval algorithm designed for non-ice-cloud conditions.

Description: The Vasco—Cirene field experiment, in January—February 2007, targeted the Seychelles—Chagos thermocline ridge (SCTR) region, with the main purpose of investigating Madden—Julian Oscillation (MJO)-related SST events. The Validation of the Aeroclipper System under Convective Occurrences (Vasco) experiment (Duvel et al. 2009) and Cirene cruise were designed to provide complementary views of air—sea interaction in the SCTR region. While meteorological balloons were deployed from the Seychelles as a part of Vasco, the Research Vessel (R/V) Suroît was cruising the SCTR region as a part of Cirene. more: The Vasco—Cirene program explores how strong air—sea interactions promoted by the shallow thermocline and high sea surface temperature in the Seychelles—Chagos thermocline ridge results in marked variability at synoptic, intraseasonal, and interannual time scales. The Cirene oceanographic cruise collected oceanic, atmospheric, and air—sea flux observations in this region in January—February 2007. The contemporaneous Vasco field experiment complemented these measurements with balloon deployments from the Seychelles. Cirene also contributed to the development of the Indian Ocean observing system via deployment of a mooring and 12 Argo profilers. Unusual conditions prevailed in the Indian Ocean during January and February 2007, following the Indian Ocean dipole climate anomaly of late 2006. Cirene measurements show that the Seychelles—Chagos thermocline ridge had higher-than-usual heat content with subsurface anomalies up to 7°C. The ocean surface was warmer and fresher than average, and unusual eastward currents prevailed down to 800 m. These anomalous conditions had a major impact on tuna fishing in early 2007. Our dataset also sampled the genesis and maturation of Tropical Cyclone Dora, including high surface temperatures and a strong diurnal cycle before the cyclone, followed by a 1.5°C cooling over 10 days. Balloonborne instruments sampled the surface and boundary layer dynamics of Dora. We observed small-scale structures like dry-air layers in the atmosphere and diurnal warm layers in the near-surface ocean. The Cirene data will quantify the impact of these finescale features on the upper-ocean heat budget and atmospheric deep convection.

Description: The mechanisms involved in setting the annual cycle of the Florida Current transport are revisited using an adjoint model approach. Adjoint sensitivities of the Florida Current transport to wind stress reproduce a realistic seasonal cycle with an amplitude of ~1.2 Sv (1 Sv ≡ 106 m3 s−1). The annual cycle is predominantly determined by wind stress forcing and related coastal upwelling (downwelling) north of the Florida Strait along the shelf off the North American coast. Fast barotropic waves propagate these anomalies southward and reach the Florida Strait within a month, causing an amplitude of ~1 Sv. Long baroclinic planetary Rossby waves originating from the interior are responsible for an amplitude of ~0.8 Sv but have a different phase. The sensitivities corresponding to the first baroclinic mode propagate westward and are highly influenced by topography. Considerable sensitivities are only found west of the Mid-Atlantic Ridge, with maximum values at the western shelf edge. The second baroclinic mode also has an impact on the Florida Current variability, but only when a mean flow is present. A second-mode wave train propagates southwestward from the ocean bottom on the western side of the Mid-Atlantic Ridge between ~36° and 46°N and at Flemish Cap, where the mean flow interacts with topography, to the surface. Other processes such as baroclinic waves along the shelf and local forcing within the Florida Strait are of minor importance.

Description: The residual effect of surface gravity waves on mean flows in the upper ocean is investigated using thickness weighted mean (TWM) theory applied in a vertically Lagrangian and horizontally Eulerian coordinate system. Depth-dependent equations for the conservation of volume, momentum, and energy are derived. These equations allow for (i) finite amplitude fluid motions, (ii) the horizontal divergence of currents and (iii) a concise treatment of both the kinematic and viscous boundary conditions at the sea surface. Under the assumptions of steady and monochromatic waves and a uniform turbulent viscosity, the TWM momentum equations are used to illustrate the pressure- and viscosity-induced momentum fluxes through the surface that are implicit in previous studies of the wave-induced modification of the classical Ekman spiral problem. The TWM approach clarifies, in particular, the surface momentum flux associated with the so-called virtual wave stress of Longuet-Higgins. Overall the TWM framework can be regarded as an alternative to the three-dimensional Lagrangian mean framework of Pierson. Moreover the TWM framework can be used to include the residual effect of surface waves in large-scale circulation models. In specific models that carry the TWM velocity appropriate for advecting tracers as their velocity variable, the turbulent viscosity term should be modified so that the viscosity acts only on the Eulerian mean velocity.

Description: Downward wave coupling occurs when an upward propagating planetary wave from the troposphere decelerates the flow in the upper stratosphere, and forms a downward reflecting surface that redirects waves back to the troposphere. To test this mechanism and potential factors influencing the downward wave coupling, three 145-year sensitivity simulations with NCAR’s Community Earth System Model (CESM-WACCM), a state-of-the-art high-top chemistry-climate model, are analyzed. The results show that the QBO and SST variability significantly impact downward wave coupling. Without the QBO, the occurrence of downward wave coupling is significantly suppressed. In contrast, stronger and more persistent downward wave coupling occurs when SST variability is excluded. The above influence on the occurrence of downward wave coupling is mostly due to a direct influence of the QBO and SST variability on stratospheric planetary wave source and propagation. The strengths of the tropospheric circulation and surface responses to a given downward wave coupling event, however, behave differently. The surface anomaly is significantly weaker (stronger) in the experiment with fixed SSTs (without QBO), even though the statistical signal of downward coupling is strongest (weakest) in this experiment. This apparent mismatch is explained by the differences in the strength of the synoptic-scale eddy-mean flow feedback and the possible contribution of SST anomalies in the North Atlantic during DWC event. The weaker synoptic-scale eddy-mean flow feedback, and the absence of the positive NAO-related SST-tripole pattern in the fixed SST experiment are consistent with a weaker tropospheric response in this experiment. The results highlight the importance of synoptic-scale eddies in setting the tropospheric response to downward wave coupling.

Description: A method to derive salinity data from RAFOS float temperature and pressure measurements is described. It is based on evaluating the float's in situ density from its mechanical properties and in situ pressure and temperature data. The salinity of the surrounding water may then be determined, assuming that the float has reached equilibrium with its environment. This method, in comparison with the possible use of floatborne salinity cells, has the advantage of being both cost and energy neutral and highly stable in the long term. The effect on the estimated salinity of various parameters used in the determination of the float's in situ density is discussed. Results of seven RAFOS Boats deployed in the Brazil Basin are compared with corresponding CTD data to estimate the magnitude of these errors. At present, an accuracy of 0.3 psu is achieved. The accuracy may be improved to 0.02 psu by referring the float's calculated density to a reference density established by a CTD cast at the time of launch. Results from five floats deployed in the heterogeneous water masses of the Iberian Basin are compared with the corresponding CM casts to demonstrate the variability and interpretation of p-T-S float datasets from different areas.

Description: Data from a surface mooring located in the Sargasso Sea at 34°N, 70°W between May 1982 and May 1984 were compared with satellite data to investigate large diurnal sea surface temperature changes. Mooring and satellite measurements are in excellent agreement for those days on which no clouds covered the site at the time of the satellite pass. During the summer half-year at this site, there is a 20% charm of diurnal warming of more than 0.5°C, with values of up to 3.5°C observed in the two-year period. Diurnal warming observed at the mooring has been simulated well by a one-dimensional model driven by local beat and momentum fluxes. Under the conditions of very light wind and strong insolation that produce the Largest surface warming, the surface mixed-layer depth reduces to the convection depth, and wind-mixing becomes unimportant. The thermal response is then limited to depths between 1 and 2 m, making it likely that such events have been underreported in routine ship observations. In all cases observed, the spatial extent of warming events as determined by satellite data are well correlated with the corresponding atmospheric pressure patterns. Conditions giving rise to the largest diurnal warming events are often associated with a westward-extending ridge of the Bermuda high. In the region studied, 57°–75°W and 29°–43°N, diurnal warming of more than 1°C was found on occasion to cover areas in excess of 300 000 km2, with warming of more than 2°C coveting areas in excess of 130 000 km2.

Description: In this paper we use the historical hydrographic data base for the South Atlantic Ocean to investigate (i) the hydrographic boundary between the subtropical gyre and the Antarctic Circumpolar Current (ACC), the Sub-tropical Front (STF), and (ii) the southern current band of the gyre, which is called the South Atlantic Current (SAC). The STF begins in the west in the Brazil-Falkland (Malvinas) confluence zone, but at locations at and west of 45°W this front is often coincident with the Brazil Current front. East of 45°W the STF appears to be a distinct feature to at least the region south of Africa, whereupon it continues into the Indian Ocean. The associated current band of increased zonal speed is the SAC, which, except for one instance, is found at or north of the surface STF until Indian Ocean water from the Agulhas retroflection is reached. A reversal of baroclinicity in the STF is observed south of a highly saline Agulhas ring, causing the SAC to separate from the STF and turn north into the Benguela Current. Zonal flow south of the STF is generally weak and serves to separate the South Atlantic and circumpolar currents. In the Argentine Basin, the SAC has a typical volume transport of 30 Sv (1 Sv = 106m3s−1) in the upper 1000 m relative to a deep potential density surface (σ4 = 45.87 kg m−3), and can be as high as 37 Sv. It is thus comparable to, or stronger than, the Brazil Current. In the Cape Basin, the transport of the SAC is reduced to about 15 SY before it turns north to feed the Benguela Current. In late 1983 this flow was joined by about 8 Sv of water from the Agulhas Current.

Description: Historical data from the region between the Greenwich meridian and the African continental shelf are used to compute the offshore geostrophic transport of the Benguela Current. At 32°S, the Benguela Current is located near the African coast, transporting about 21 Sv (1 Sv = 106 m3 s−1) of surface water toward the north relative to a potential density surface lying between the upper branch of Circumpolar Deep Water and the North Atlantic Deep Watar. Two warm core eddies of probable Agulhas Current origin an observed west of the Benguela Current at 32°S. Near 30°S, the Benguela Current turns toward the northwest and begins to separate from the eastern boundary. It carries about 18 Sv of surface water across 28°S. The current then turns mainly toward the west to flow over a relatively deep segment of the Walvis Ridge south of the Valdivia Bank. A surface current with northward surface of about 10 cm s−1 flows along the western side of the Valdivia Bank, while another northward surface current flows at about 20 cm s−1 some 300 km west of the bank. About 3 Sv of surface now do not leave the Cape Basin south of the Vaidivia Bank, but instead drift northward as a wide. sluggish flow out of the northern end of the Cape Basin. Because of the more southerly seaward extensions of most of the Benguela Current, there are no deep-reaching interactions observed between this current and the cyclonic gyre in the Angola Basin east of the Greenwich meridian. Beneath the surface layer, about 4–5 Sv of Antarctic Intermediate Water are carried northward across 32° and 28°S by the Benguela Current, essentially all of which turns westward to cross the Greenwich meridian south of 24°S.

Description: An XBT interface is described for use with Commodore and other 6502 based microprocessors. This interface takes the form of a single circuit board mounted inside the microcomputer and is completely software controlled. The application of this digital XBT system to the real-time computation of density and dynamic height, using historical or recent temperature-salinity relationships, is also described. Comparison between XBT and CTD measured temperatures from the Northeast Atlantic yield a mean temperature difference of −0.08°C and an rms temperature difference of 0.33°C for the upper 800 m. Examples of dynamic topography maps and a temperature section computed using this technique are also presented and comparison between objectively analyzed XBT and CTD dynamic topographies demonstrates the reliability of the method for mapping the baroclinic flow.

Description: The upper bathyal sea-pen Kophobelemnon stelliferum extends to depths of about 1600 m in the Porcupine Seabight, to the south-west of Ireland, but is rare below about 1150 m. Photographic data suggest that the species attains numerical abundances of more than 2 m−2and a wet weight biomass of at least 4 g m−2. The highest densities, however, do not necessarily correspond to the highest biomass values since there is a clear depth-related change in population structure. The largest sea-pens are restricted to the deeper parts of the bathymetric range of the species. There is also a marked change in the growth form at a total colony length of about 250 mm, with larger colonies having relatively more polyps than smaller ones. The sexes are separate in Kophobelemnon stelliferum and the sex ratio of colonies is about 1:1. The maximum oocyte diameter is about 800 μm, but there is no evidence of seasonal reproduction by this pennatulid in the Porcupine Seabight.

Description: A correlation of the Campanian–Maastrichtian boundary is attempted using foraminiferal and nannoplankton data from two areas: the eastern North Atlantic and northwestern Germany. The Boreal benthic and Tethyan planktonic foraminiferal zonation schemes are applied to Site 548A, where both foraminiferal groups occur frequently. A direct comparison of both biozonations reveals that the base of the Maastrichtian, according to planktonic foraminifers, has to be placed in the Upper Campanian of the Boreal benthic foraminiferal biozonation, which concurs with the nannoplankton results. The Tethyan Middle and Upper Maastrichtian are probably equivalent to the Upper Maastrichtian in the Boreal sense. The bases of the Maastrichtian substages are thus diachronous between the Boreal and Tethyan realms. Palaeotemperatures (which were estimated using the oxygen isotopic composition of the Goban Spur chalks) indicate, in combination with palaeowind directions, that the faunal and floral distribution pattern recorded is the result of a stable, warm water outflow from the northwest European epicontinental seas through the Channel area to the Celtic Shelf sea and Goban Spur. This mechanism appears to have been a dominant separating factor of the Boreal and Tethyan bioprovinces on the western European Shelf.

Description: In this study a scenario is developed of two adjacent Mediterranean Water eddies (meddies) as they were observed merging and drifting through the Iberian Basin. Observations are based on four RAFOS floats (at 850–1050 dbar), two hydrographic surveys (centered roughly at 38°N, 24°W), and trajectories of surface drifters (drogued at 100 m). In April 1991, the meddy A was identified and labeled by surface drifters. During the revisit one month later two meddies were encountered, B1 and B2, in the vicinity of the former meddy A. The coalescence of B1 (subsequently identified as A, one month older) and B2 is inferred from a simple kinematic model describing the observed movement of the RAFOS floats for up to three months after the second CTD survey. The deduced vorticity front, radius ∼15 km, within B1 was of insufficient strength to keep the core waters of B1 isolated and prevent the absorption of B1 by B2. The resulting meddy (B1 + B2) showed a clear near-surface dynamical signal. Its deep root (1800 m) could explain the expulsion from the meddy of the remaining RAFOS float and surface drifter at the time of the meddy's collision with the Josephine Seamount. For the first time, a set of Lagrangian and hydrographic observations give direct evidence that neighboring meddies can merge as predicted by theoretical considerations.

Description: The total transport of Antarctic Bottom Water across the Rio Grande Rise, including the western boundary, the Vema Channel, and the Hunter Channel is estimated from hydrographic measurements across these pathways. The contribution of the Vema Channel is greatest at 3.9 × 106 m3 s−1, which is very close to earlier estimates. The western boundary current contribution is 2.0 × 106 m3 s−1 and that of the Hunter Channel 0.7 × 106 m3 s−1. The lower values outside the Vema Channel are offset by the important source of mass they form to the lower density classes of bottom water. About 40% of the flow is concentrated in the highest density class representing the source of Weddell Sea Deep Water to the Brazil Basin. The flow structure is characterized by horizontal and vertical recirculation.

Description: The existence of energetic anticyclonic mid-depth vortices of Mediterranean Water (meddies) questions the validity of a conventional advective–diffusive balance in the eastern Atlantic subtropical gyre. A mesoscale experiment in the Azores–Madeira region reveals a link of these meddies to large-scale subsurface meanders. For the first time it is shown that meddies may have strong surface vorticity, indicative of a generation process involving the Azores Current—a deep reaching near-surface jet.

Description: In this paper, the historical hydrographic database for the south Indian Ocean is used to investigate (i) the hydrographic boundary between the subtropical gyre and the Antarctic Circumpolar Current (ACC), the subtropical front (STF), and especially (ii) the southern current band of the gyre. A current band of increased zonal speeds in the upper 1000 m is found just north of the STF in the west near South Africa and at the surface STF in the open Indian Ocean until the waters off the coast of Australia are reached. As neither any other investigation of this current nor a name for it are known, the flow has been called the South Indian Ocean Current (SIOC). This name is anologous to the same current band in the South Atlantic Ocean, the South Atlantic Current. The STF is located in the entire south Indian Ocean near 40-degrees-S. The associated current band of increased zonal speeds is the SIOC, which is found at or north of the STF. East of 100-degrees-E the SIOC separates from the STF and continues to the northeast. The zonal flow south of the STF is normally weak and serves to separate the South Indian Ocean and Circumpolar currents. Near Africa the SIOC has a typical volume transport of 60 Sv (1 Sv = 10(6) m3 s-1) in the upper 1000 m relative to deep potential density surfaces of sigma(4) = 45.87 kg m-3 (2800-3500 m) or sigma(2) = 36.94 kg m-3 (1500-2500 m). Near western Australia the SIOC is reduced to about 10 Sv as it turns to the northeast.

Description: Many aspects of barnacle body form are known to be developmentally plastic. Perhaps the most striking examples of such plasticity occur in their feeding legs and unusually long penises, the sizes and shapes of which can change dramatically and adaptively with changes in conspecific density and local water flow conditions. However, whether variation in overall appendage form is mirrored by structural responses in cuticle and muscle is not known. In order to determine how structural variation underlies phenotypic plasticity in barnacle appendages, we examined barnacles occurring at low and high population densities from one wave-protected and one wave-exposed site. We used histological sectioning and fluorescence microscopy of feeding legs and penises to compare cuticle thickness, muscle thickness, and muscle organization, and artificial penis inflation to compare penis extensibility. We observed striking differences in cuticle thickness, muscle thickness, and muscle organization between sites that differed in water velocity, but we found no clear differences associated with variation in conspecific density. Penis extensibility also did not differ consistently between sites. These results are consistent with an adaptive explanation for much of the remarkable and complex variation in barnacle feeding leg and penis morphology among sites that differ in water velocity.

Description: There is evidence that the strengthened stratospheric westerlies arising from the Antarctic ozone hole–induced cooling cause a polar mesospheric warming and a subsequent cooling in the lower thermosphere. While previous studies focus on the role of nonresolved (gravity) wave drag filtering, here the role of resolved (planetary) wave drag and radiative forcing on the Antarctic mesosphere and lower thermosphere (MLT) is explored in detail. Using simulations with NCAR’s Community Earth System Model, version 1 (Whole Atmosphere Community Climate Model) [CESM1(WACCM)], it is found that in late spring and early summer the anomalous polar mesospheric warming induced by easterly nonresolved wave drag is dampened by anomalous dynamical cooling induced by westerly resolved wave drag. This resolved wave drag is attributed to planetary-scale wave (k = 1–3) activity, which is generated in situ as a result of increased instability of the summer mesospheric easterly jet induced by the ozone hole. On the other hand, the anomalous cooling in the polar lower thermosphere induced by westerly nonresolved wave drag is enhanced by anomalous dynamical cooling due to westerly resolved wave drag. In addition, radiative effects from increased greenhouse gases during the ozone hole period contribute partially to the cooling in the polar lower thermosphere. The polar MLT temperature response to the Antarctic ozone hole is, through thermal wind balance, accompanied by the downward migration of anomalous zonal-mean wind from the lower thermosphere to the stratopause. The results highlight that a proper accounting of both dynamical and radiative effects is required in order to correctly attribute the causes of the polar MLT response to the Antarctic ozone hole.

Description: The asymmetry of the continental margin around southern Africa can be related to Mesozoic sediment thicknesses, which were in turn controlled by the local structural setting. On the west coast, the Orange Basin sediments were built out as a thick wedge over the margin of the continent by discharge from the Orange River, whereas on the Agulhas Bank, sedimentation was confined to continental areas. Off the east coast the extremely narrow margin of the continent did not form an effective trap for sediments, which were readily carried beyond it. Cainozoic sediments are thin, and modify the Mesozoic sediment pile only locally on the outer shelf and slope.

Description: The Southern Hemisphere Subtropical Front (STF) is a narrow zone of transition between upper-level subtropical waters to the north and subantarctic waters to the south. It is found near 40 degrees S across the South Atlantic and South Indian Oceans and is associated with an eastward geostrophic current band, The current band in each basin is found at or just north of the surface front except near the eastern boundaries where most of the subtropical waters turn north into the eastern limbs of the subtropical gyres. The bands associated with the STF are thus distinct features separated from the strong zonal flows of the Antarctic Circumpolar Current farther south. The authors have referred to the current bands in the two respective oceans as the South Atlantic Current and the South Indian Ocean Current. In this paper the authors use the historical database from the South Pacific Ocean to investigate the geostrophic flow associated with the STF there. The STF extends across the southern Tasman Sea from south of Tasmania to southern New Zealand, and a weak eastward flow appears to be associated with it. The transport amounts to only about 3 Sv (1Sv = 10(6) m(3) s(-1)), little of which passes south of New Zealand. Mixing within the eddy-rich Tasman Sea may account for this weakness, while also setting up another more significant front in the northern Tasman Sea, the Tasman Front. It branches off from the East Australian Current toward the north of New Zealand, along which moves a flow of about 14 Sv. After passing north of New Zealand, a portion of this current flows east to contribute to a current band near 30 degrees S, while another portion turns south as the East Auckland Current and meets with subantarctic waters near Chatham Rise (44 degrees S), thus reestablishing the STF. An enhanced eastward current band is associated with the front there, one that extends across the remainder of the South Pacific and is referred to as the South Pacific Current. In comparison with its counterparts in the other basins, which typically begin by carrying 30 Sv (Atlantic) to 60 Sv (Indian) in the upper 1000 m in their western portions before weakening to 10-15 Sv in the east, the South Pacific Current is weak. Near Chatham Rise, it starts with a transport of approximately 5 Sv, and it remains near this strength as it shifts gradually north across the basin toward South America. The current appears to split into two smaller bands in the region of 115 degrees-85 degrees W, while near 28 degrees 5, 83 degrees W it begins to turn more strongly north and becomes shallower and weaker. Potential vorticity distributions indicate that this current acts as an impediment toward the northward spreading of Antarctic Intermediate Water, But why the South Pacific Current east of New Zealand should be so much weaker than its counterparts in the other basins is not particularly clear. It may be due to the presence of New Zealand and other topographic barriers to deep now east of Australia, to the axis of the subtropical gyre in the South Pacific shifting more rapidly southward with depth than those elsewhere, thus causing greater reductions in the underlying zonal velocities, and to strong poleward eddy heat and salt fluxes in the other two basins leading to smaller cross-STF gradients in the Pacific.

Description: The planktonic foraminifer Globigerinoides sacculifer (Brady) was cultured under two different light intensities and in continuous darkness. High light intensity (HLI = 4oo-soo einsteins/m2/s) resulted in a longer lifespan, a greater number of chambers formed, and a larger final shell size compared with individuals cultured under low light intensity (LLI = 20-50 einsteins/m2/s) or in continuous darkness. Shell growth rates were unaffected by increasing light intensity, but gametogenesis was delayed. Continuous darkness induced a rapid onset of gametogenesis in organisms with shell lengths larger than 250 m. Feeding frequency had a greater effect on growth and reproduction than light intensity under conditions of LLI and HLI, but continuous darkness had an overriding effect on growth and reproduction owing to the rapid onset of gametogenesis which terminated the life of the mother cell. Our previous data indicated that the longevity of G. sacculifer was dependent on feeding frequency, and that G. sacculifer cultured under LLI had a lifespan of approximately 2-4 weeks. Present results suggest that the lifespan can vary from a minimum of 8 days for organisms fed daily in continuous darkness to a maximum of 54 days for organisms fed once every 7 days and maintained in HLI. It is concluded that individual G. sacculifer attain a shell size greater than 6oo ,urn only if they maintain their position in the euphotic zone. Prolonged existence below the euphotic zone would result in premature death or gametogenesis following stunted shell growth.

Description: EGU2010-13373 The frequency of volcanic activity varies on a wide rangeof spatial and temporal scales, from 〈1 yr. periodicities in single volcanic systems to periodicities of 106 yrs. in global volcanism. The causes of these periodicities are poorly understood although the long-term global variations are likely linked to plate-tectonic processes. Here we present evidence for temporal changes in eruption frequencies at an intermediate time scale (104 yrs.) using the Pleistocene to recent records of widespread tephras of sub-Plinian to Plinian, and occasionally co-ignimbrite origin, along the Pacific Ring of Fire, which accounts for about half of the global length of 44,000 km of active subduction. Eruptions at arc volcanoes tend to be highly explosive and the well-preserved tephra records from the ocean floor can be assumed to be representative of how eruption frequencies varied with time. Volcanic activity along the Pacific Ring of Fire evolved through alternating phases of high and low frequency; although there is modulation by local and regional geologic conditions, these variations have a statistically significant periodicity of 43 ka that overlaps with the temporal variation in the obliquity of the Earth’s rotation axis, an orbital parameter that also exerts a strong control on global climate changes. This may suggest that the frequency of volcanic activity is controlled by effects of global climate changes. However, the strongest physical effects of climate change occur at 100 ka periods which are not seen in the volcanic record. We therefore propose that the frequency of volcanic activity is directly influenced by minute changes in the tidal forces induced by the varying obliquity resulting in long-period gravitational disturbances acting on the upper mantle.