ALBERT

Description: A new species of the holopodid genus Cyathidium was found on rocks off Grande Comore in a depth of around 200 m. Based on external morphology of resting animals, the new species Cyathidium plantei sp. n is described, with emphasis on comparison to the only other extant species (C. foresti) as well as to the four extinct representatives of the genus. Concerning morphological characters, the new species is almost identical to the Cretaceous C. depressum. A cladistic analysis of the entire family, including the genus Holopus, shares a peculiar pattern of bending of the arms, which in principle is an apomorphic character of the family and in detail shows variations within the family. In addition, stratigraphic data are used for the determination of the evolutionary direction. This analysis reveals that the two recent species are closely related to each other, and to the fossil C. depressum. from which the entire family is probably derived.

Description: A protein unique to phosphorus stress observed in Dunaliella tertiolecta Butcher was studied in the context of phosphate‐limited cell physiology and is a potential diagnostic indicator of phosphate deficiency in this alga. Cells were grown over a range of limited, steady‐state growth rates and at maximum (replete) and zero (phosphate‐starved) growth rates. The stress protein, absent in nutrient‐replete cells, was produced under all steady‐state phosphate‐limited conditions and increased in abundance with increasing limitation (decreasing growth rate). Cellular carbon: phosphorus ratios and the maximum uptake rate of phosphate (Vm) increased with increasing limitation, whereas the ratio of chlorophyll a: carbon decreased. Alkaline phosphatase activity did not respond to limitation but was measurable in starved, stationary‐phase cells. Fv/Fm, a measure of photochemical efficiency, was a nonlinear, saturating function of p, as commonly observed under N limitation. The maximum Fv/Fm of 0.64 was measured in nutrient‐replete cells growing at μmax, and a value of zero was measured in stationary‐phase starved cells. When physiological parameters were compared, the P‐stress protein abundance and Fv/Fm were the most sensitive indicators of the level of deficiency. The stress protein was not produced under N‐ or Fe‐limited conditions. It is of high molecular weight (〉200) and is associated with internal cell membranes. The stress protein has several characteristics that make it a potential diagnostic indicator: it is 1) unique to phosphorus limitation (i.e. absent under all other conditions), 2) present under limiting as well as starved conditions, 3) sensitive to the level of limitation, and 4) observable without time‐course incubation of live samples.

Description: High-resolution reflection seismic data obtained around Gran Canaria allow a detailed and consistent correlation of seismic reflectors of the northern and southern Canary Basins with the lithology drilled by DSDP Leg 47A SSE of Gran Canaria, as well as with major phases of volcanic activity on Gran Canaria as mapped onshore. Two prominent reflectors were chosen as marker horizons and correlated with the drilled lithology. the results indicate that reflector R7 above the Miocene volcaniclastic debris flows V1-V3 reflects the shield-building phase of Gran Canaria. Reflector R3 is interpreted as corresponding with the Pliocene Roque Nublo formation. The top of the massive island flank of Gran Canaria, defined by seismically chaotic facies, extends 44 to 72 km off the coast of Gran Canaria. West of Gran Canaria the flank of Tenerife onlaps the steeper and older flank of Gran Canaria, which, in turn, is onlapping the older flank of Fuerteventura to the east in a similar way. Erosional channels, which can also be traced up to 50 km from the area between Gran Canaria and Fuerteventura into the deeper northern basin, have been identified in the bathymetry. The data presented provide new detailed information for modelling the submarine and subaerial evolution of the central Canary Islands of Gran Canaria and Tenerife, i.e. the timing of their shield-building phases and later stages of major volcanic activity, as reflected by the position of prominent seismic reflectors in the seismic stratigraphy.

Description: An initially resting ocean of stratification N is considered, subject to buoyancy loss at its surface of magnitude B0 over a circular region of radius r, at a latitude where the Coriolis parameter is f. Initially the buoyancy loss gives rise to upright convection as an ensemble of plumes penetrates the stratified ocean creating a vertically mixed layer. However, as deepening proceeds, horizontal density gradients at the edge of the forcing region support a geostrophic rim current, which develops growing meanders through baroclinic instability. Eventually finite-amplitude baroclinic eddies sweep stratified water into the convective region at the surface and transport convected water outward and away below, setting up a steady state in which lateral buoyancy flux offsets buoyancy loss at the surface. In this final state quasi-horizontal baroclinic eddy transfer dominates upright “plume” convection. By using “parcel theory” to consider the energy transformations taking place, it is shown that the depth, hfinal at which deepening by convective plumes is arrested by lateral buoyancy flux due to baroclinic eddies, and the time tfinal it takes to reach this depth, is given by both independent of rotation. Here γ and β are dimensionless constants that depend on the efficiency of baroclinic eddy transfer. A number of laboratory and numerical experiments are then inspected and carried out to seek confirmation of these parameter dependencies and obtain quantitative estimates of the constants. It is found that γ = 3.9 ± 0.9 and β = 12 ± 3. Finally, the implications of our study to the understanding of integral properties of deep and intermediate convection in the ocean are discussed.

Description: The incidence angles of the SSM/I radiometers on the DMSP satellites vary from satellite to satellite and exhibit variations of up to 1.5° during one orbit. The effects of these variations on the measured brightness temperatures are investigated on the basis of simulated and measured data for oceanic arm. A deviation of 1° from the nominal incidence angle of 53.0° causes brightness temperature changes of up to 2 K depending on surface and atmospheric conditions. Errors of retrieved geophysical parameters on the order of 5%–10% result when the incidence angle variation is not taken into account. This is a common property of most published statistical algorithms. For total precipitable water and cloud liquid water content the error increases with increasing parameter value. For wind speed the error is largest for low wind speed and decreases with increasing wind speed. Due to the slowly varying latitudinal dependence of the incidence angle, these errors do not cancel out when monthly means are computed. A correction method is developed on the basis of simulated data and tested successfully with measured data. Observed brightness temperature differences between DMSP F10 and F11 are reduced when using corrected data. If diurnal variations of geophysical parameters are investigated, the incidence angle correction is mandatory to obtain useful results, especially for DMSP F10.

Description: The compatibility of the Gent and McWilliams thickness mixing parameterization with perturbation thickness fluxes evaluated from eddy-resolving North Atlantic model results is investigated. After extensive spatial and temporal averaging, a linear correlation between the parameterized fluxes and those calculated directly from model fluctuations in the subtropics could be found. A direct estimate of a constant mixing parameter κ could be inferred in the order of 1.0 × 107 cm2 s−1.

Description: A simple point-vortex “heton” model is used to study localized ocean convection. In particular, the statistically steady state that is established when lateral buoyancy transfer, effected by baroclinic instability, offsets the localized surface buoyancy loss is investigated. Properties of the steady state, such as the statistically steady density anomaly of the convection region, are predicted using the hypothesis of a balance between baroclinic eddy transfer and the localized surface buoyancy loss. These predictions compare favorably with the values obtained through numerical integration of the heton model. The steady state of the heron model can be related to that in other convection scenarios considered in several recent studies by means of a generalized description of the localized convection. This leads to predictions of the equilibrium density anomalies in these scenarios, which concur with those obtained by other authors. Advantages of the heton model include its inviscid nature, emphasizing the independence of the fluxes affected by the baroclinic eddies from molecular processes, and its extreme economy, allowing a very large parameter space to be covered. This economy allows us to examine more complicated forcing scenarios: for example, forcing regions of varying shape. By increasing the ellipticity of the forcing region, the instability is modified by the shape and, as a result, no increase in lateral fluxes occurs despite the increased perimeter length. The parameterization of convective mixing by a redistribution of potential vorticity, implicit in the heton model, is corroborated; the heton model equilibrium state has analogous quantitative scaling behavior to that in models or laboratory experiments that resolve the vertical motions. The simplified dynamics of the heton model therefore allows the adiabatic advection resulting from baroclinic instability to be examined in isolation from vertical mixing and diffusive processes. These results demonstrate the importance of baroclinic instability in controlling the properties of a water mass generated by localized ocean convection. A complete parameterization of this process must therefore account for the fluxes induced by horizontal variations in surface buoyancy loss and affected by baroclinic instability.