ALBERT

Note: Enthaltener Beitrag: Russian-German Cooperation SYSTEM LAPTEV SEA: The Expedition COAST I / edited by Paul Overduin , Enthaltener Beitrag: The Expedition Lena 2005 / edited by Lutz Schirrmeister, Dirk Wagner, Mikhail N. Grigoriev and Dimitry Yu. Bolshiyanov , Enthaltener Beitrag: Russian-German cooperation Yakutsk - Potsdam: the expedition CENTRAL YAKUTIA 2005 / edited by Bernhard Diekmann, Sebastian Wetterich and Frank Kienast , Contents for "Russian-German Cooperation SYSTEM LAPTEV SEA: The Expedition COAST I" The Expedition COAST I 1. Background and Objectives 2. Logistics and Itinerary 3. Field Methods and Sample Recovery 3.1 Coring 3.2 Pore water analyses 3.3 Temperature profiles 4. Sample lists , Contents for "The Expedition Lena 2005" The Expedition Lena 2005 1. Introduction 2. Expedition itinerary and general logistics 3. Microbiological processes, trace gas fluxes and hydrobiology in permafrost ecosystems of the Lena Delta 3.1 Introduction 3.2 Dynamic of methane oxidising communities in permafrost soils 3.2.1 Introduction 3.2.2 Sampling procedure and field parameters 3.2.3 Pore water methane concentration 3.2.4 Sample processing and analyses 3.3 Microbial studies on nitrification from permafrost environments 3.3.1 Introduction 3.3.2 Field experiments: Impact of polygonal soil parameter on nitrification 3.4 Closed chamber measurements of carbon exchange between Arctic tundra and the atmosphere 3.5 Micrometeorological measurements of energy, water, and carbon exchange between Arctic tundra and the atmosphere 3.6 Energy and water budget of permafrost soils – long time meteorology and soil survey station on Samoylov Island 3.7 Isotopic Studies on the 13C-fractionation during CH4-production in polygonal and thermokarst lakes of the Lena Delta 3.7.1 Introduction and methods 3.7.2 Preliminary results and further plans 3.8 Hydrobiological investigations on Samoylov Island 3.8.1 Objectives 3.8.2 Research tasks 3.8.3 Material and methods 3.8.4 Preliminary results 3.9 References 4. Studies of periglacial landscape dynamics and surface characteristics studies in the western Lena Delta 4.1. Scientific background and objectives 4.2. Geological and geographical characteristics 4.3. Studies of oriented lakes and thermokarst depressions 4.3.1 Background 4.3.2 Study area 4.3.3 Topographical and geomorphological settings 4.3.3.1 Depressions 1, 2 and 3 4.3.3.2 Depression 4 4.3.3.3 Depression 5 4.3.4 Bathymetrical surveys 4.3.5 Field sampling 4.4. Characteristics and spectral properties of periglacial landforms 4.4.1 Introduction 4.4.2 Methods 4.4.3 First results 4.5. Studies of permafrost sequences for paleo-environmental reconstruction 4.5.1 The “Arga-Sands” on Turakh Island 4.5.1.1 Exposure Tur-1 4.5.1.2 Core Tur-2 4.5.1.3 Exposure T021 4.5.2 Sand sequences of Ebe Basyn Sise Island 4.5.2.1 Exposure Ebe-4 4.5.2.2 Exposure Ebe-2 4.5.2.3 Exposure Ebe-3 4.5.2.4 Exposure Ebe-5 4.5.3 Sand and Ice Complex sequences of Khardang Island 4.5.3.1 The sand deposits in the exposure Kha-1 4.5.3.2 The sequence Kha-2 4.5.3.3 Exposure Kha-3: large ice wedge and surrounding sediments 4.6 Subsurficial and bathymetrical Ground Penetrating Radar (GPR) Investigations 4.6.1 Subsurface mapping of the Arga sands stratigraphical unit 4.6.1.1 GPR survey configuration 4.6.1.2 Transects at exposure Ebe-4 4.6.1.3 Transects at exposure/borehole Tur-1/Tur-2 4.6.2 Arynskaya Channel bathymetry 4.7 Measuring of local weather and soil conditions by soil probe and weather station 4.8 Palaeontological collection of the “Mammoth” fauna from the museum of the Lena Delta Reserve 4.9 References 4.10 Appendices chapter 4 Appendix 4-1: Field spectrometry – description of measuring points and profiles (see chapter 4.4) Appendix 4-2: List of sediment samples (see chapter 4.5) Appendix 4-3: Modern soil profiles and surface samples Appendix 4-4: List of ground ice and surface water samples Appendix 4-5. Bone collection of the expedition LENA 2005 Appendix 4-6: Bone collection of Lena Delta Reserve Tiksi (see chapter 4.8) 5. Holocene ice wedges of the 1st Lena terrace 5.1 Introduction 5.2 Outcrops 5.2.1 Outcrop 1 5.2.2 Outcrop 2 5.2.3 Geocryolithology on Samoylov Island: General impressions 5.2.4 Outcrop 3 5.2.5 Outcrop 4 5.2.6 Outcrop 5 5.2.7 Outcrop 6 5.2.8 Outcrop 7 5.2.9 Outcrop 8 5.2.10 Outcrop 9 5.2.11 Outcrop 10 5.2.12 Pingo at Olenyekskaya Channel 5.2.13 Summary 5.3 Studies on recent cryogenesis on Samoylov Island 5.4 References 5.5 Appendices chapter 5 Appendix 5-1: Ice sample list Appendix 5-2: List of sediment samples and ice content measurement Appendix 5-3: List of water samples 6. Report of the hydrological work in the Lena River Delta in August 2005 6.1 Introduction 6.2 Methods 6.3 Preliminary results 6.4 Conclusion , Contents for "Russian-German cooperation Yakutsk - Potsdam: the expedition CENTRAL YAKUTIA 2005" Central Yakutia 2005 1. Expedition ‘Verkhoyansk 2005’ • Limnogeological studies at Lake Billyakh, Verkhoyansk Mountains, Yakutia 1.1 Introduction 1.2 Regional Setting of Lake Billyakh 1.3 Itinerary 1.4 Methods 1.4.1 Bathymetric measurements 1.4.2 Water sampling and measurements 1.4.3 Sediment coring 1.5 Results 1.5.1 Bathymetry 1.5.2 Water profiles 1.5.3 Sediment cores 1.6 Outlook 1.7 References 2 Limnological studies in Central and North-east Yakutia in summer 2005 2.1 Introduction 2.2 Study sites and lake types 2.3 Material and methods 2.4 Preliminary results 2.5 Outlook 2.6 References 2.7 Appendices Appendix 2-1: General characteristics and geographical position of the studied lakes in Central and North-east Yakutia Appendix 2-2: Some properties of the studied lakes in Central and North-east Yakutia, obtained during the fieldwork (unfilled table cells imply no data or information) Appendix 2-3: Sample list for further analyses on sediments, hydro-chemistry, water isotopes and aquatic organisms Appendix 2-4: Occurrence of zoobenthos organisms in the sampled Central-Yakutian lakes in July 2005 3. Vegetation studies in extremely continental regions of Yakutia 3.1 Introduction 3.2 Study areas and investigated vegetation types 3.3 Material and methods 3.4 Preliminary results 3.5 Appendix 3-1: Metadata of studied vegetation records

Note: Potsdam, Univ., Habil.-Schr., 2007

Note: Enthaltener Beitrag: Russian-German cooperation System Laptev Sea : the expedition Lena-Anabar 2003 / edited by Lutz Schirrmeister, Mikhail N. Grigoriev, Lars Kutzbach, Dirk Wagner and Dmitry Yu. Bolshiyanov , Enthaltener Beitrag: The Russian-German TRANSDRIFT IX Expedition of RV "Ivan Kireyev" 2003 / edited by Carolyn Wegner, Jens Hölemann and Vladimir Churun , Contents for "Russian-German cooperation System Laptev Sea : the expedition Lena-Anabar 2003" 1 Introduction 2 Expedition itinerary and general logistics 3 Ecological studies on permafrost soils and landscapes of the central Lena Delta6 3.1 Aims and study area 3.2 Energy and water budget of permafrost soils – long time soil survey station on Samoylov Island 3.3 Micrometeorological measurements of energy, water, and carbon exchange between Arctic tundra and the atmosphere 3.3.1 Introduction 3.3.2 Experimental set-up 3.3.3 The observation period 2003 3.3.3.1 Wind characteristics 3.3.3.2 Meteorological conditions 3.3.3.3 Turbulent fluxes 3.4 Microbial process studies on methane fluxes from permafrost environments 3.4.1 Introduction 3.4.2 Methane emission and microbial methane production 3.4.2.1 Methods and field experiments 3.4.2.2 Preliminary results 3.4.3 Process studies on methane oxidation 3.4.3.1 Introduction and objectives 3.4.3.2 Methods and field experiments 3.4.3.3 Preliminary results 3.4.4 Further investigations 3.5 Studies on recent cryogenesis 3.6 Seasonal progression of active-layer thickness dependent on microrelief 3.6.1 Introduction 3.6.2 Methods 3.6.3 First results 3.7 Air photography and surface classification of Samoylov Island 3.8 Hydrobiological investigations in the Lena Delta in summer 2003 3.8.1 Objectives 3.8.2 Research tasks 3.8.3 Material and methods 3.8.4 Preliminary results 3.9 Appendices Appendix 3-1 Collected variables determined by direct measurements within the micrometeorological campaign Samoylov, 2003 Appendix 3-2 Variables derived from calculations within the micrometeorological campaign Samoylov, 2003 Appendix 3-3 Constants required in calculations Appendix 3-4 List of samples for methane emission and microbial methane production studies Appendix 3-5 List of soil samples for methane oxidation studies 3.10 References 4 Periglacial studies around Cape Mamontov Klyk 4.1 Introduction 4.2 Geological and geographical background 4.3 Geomorphologic route along the Urasalakh River 4.3.1 Structure of Quaternary deposits along the Urasalakh River valley 4.3.2 Geomorphologic structure of the valley 4.3.3 Studies of lakes 4.4 Multi-sensor optical remote sensing of periglacial tundra landscapes 4.4.1 Research aim 4.4.2 Satellite data 4.4.3 General geomorphology in the Lena-Anabar interfluve 4.4.4 Geomorphology in the investigation area near Cape Mamontov Klyk 4.4.5 Field data 4.4.6 Tachymetric survey of periglacial surface features 4.4.7 Characterisation in situ surface properties with a soil probe at a typical elevated Edoma plain 4.5 Methane-related studies on recent tundra soils 4.5.1 Introduction and objectives 4.5.2 Methods 4.5.3 First results 4.6 The coastal section of Cape Mamontov Klyk 4.6.1 General profile 4.6.2 Cryolithological studies of permafrost deposits 4.6.2.1 The first composite profile 4.6.2.2 The second composite profile 4.6.2.3 Additional sampled subprofiles 4.6.3 Ice wedges of Cape Mamontov Klyk 4.6.3.1 Introduction 4.6.3.2 The ice wedges in the lower sands (Unit A) and in the sand-peat-complex (Unit B) 4.6.3.3 Ice wedges of the Late Pleistocene Ice Complex (Unit C) and of Holocene deposits (Unit D) 4.6.3.4 Ground ice of unknown origin 4.6.3.5 Ice wedge section west of the Nuchcha Dzhiele River mouth 4.6.3.6 General interpretation of the sampled profile 4.6.4 Geomicrobiological studies 4.6.4.1Introduction and objectives 4.6.4.2 Methods 4.6.4.3 First results: Methane content of permafrost samples 4.6.5 Paleontological studies 4.7 Studies of coastal dynamics and sub sea permafrost 4.7.1 Preliminary results of sub-sea permafrost drilling in the near-shore zone (spring 2003) 4.7.1.1 Introduction and background 4.7.1.2 Methods and preliminary results 4.7.1.3 Further investigations 4.7.2 Measurements of the coast relief in the area of Cape Mamontov Klyk and ice and sediment sampling 4.7.2.1 Introduction 4.7.2.2 Methods 4.7.2.3 Preliminary results 4.7.2.4 Further investigations 4.7.3 Shore face profiles in the area of Cape Mamontov Klyk: echo sounding, seawater and sea bottom deposits sampling 4.7.3.1 Introduction 4.7.3.2 Methods 4.7.3.3 Preliminary results 4.7.3.4 Further investigations 4.8 References 4.9 Appendices Appendix 4-1. Surface parameters for the studied geolocated sites around Cape Mamontov Klyk Appendix 4-2. Active layer data of the geo-located sites Appendix 4-3. List of soil samples (active layer); collected in the coastal lowland Appendix 4-4. List of permafrost sediment and paleosol samples for microbiological, molecular biological and biochemical analyses Appendix 4-5. List of sediment samples Appendix 4-6. List of ice and water samples Appendix 4-7. Collection of bone samples , Content for "The Russian-German TRANSDRIFT IX Expedition of RV "Ivan Kireyev" 2003" 1. Process Studies on Permafrost Dynamics in the Laptev Sea – An Introduction 2. The TRANSDRIFT IX Expedition: Process studies on submarine permafrost dynamics in the Laptev Sea 3. Motivation: Deployment of two seafloor observatories 4. Recent stability factors of submarine permafrost 4.1 Working program 4.2 Preliminary results 5. Hydrochemical structure of the water column 5.1 Working program 6. Diversity of nitrifying bacteria in submarine permafrost 6.1 Working program 6.2 Preliminary Results 7. Appendix: Station list of the TRANSDRIFT IX (IK03) expedition 8. References

Note: Zugl.: Hamburg, Univ., FB Geowiss., Diss., 1998 , Inhaltsverzeichnis: I Zusammenfassung. - II Summary. - III Abbildungsverzeichnis. - IV Tabellenverzeichnis. - V Abkürzungsverzeichnis. - VI Glossar. - 1 Einleitung und Fragestellung. - 2 Grundlagen. - 2.1 Methan in der Natur. - 2.2 Methan als klimarelevantes Spurengas. - 2.3 Der Methankreislauf. - 2.4 Methanogene Bakterien. - 2.5 Anaerober Abbau organischer Substanz. - 2.6 Isotopieeffekte beim mikrobiellen anaeroben Abbau organischer Substanz. - 2.7 Bedeutung terrestrischer Ökosysteme für die Methanfreisetzung. - 3 Charakterisierung der Untersuchungsgebiete. - 3.1 Asseler Sand (Assel, Norddeutschland). - 3.1.1 Lage und naturräumliche Gliederung. - 3.1.2 Landschaftsgenese. - 3.1.3 Klima. - 3.1.4 Hydrologie. - 3.1.5 Vegetation und Nutzung. - 3.1.6 Böden des Untersuchungsgebietes. - 3.2 Tai-Hu-Tiefland (Suzhou, Ostchina). - 3.2.1 Lage. - 3.2.2 Landschaftsgenese. - 3.2.3 Klima. - 3.2.4 Vegetation und Nutzung. - 3.2.5 Böden des Untersuchungsgebietes. - 4 Material und Methoden. - 4.1 Bodenprofilaufnahme und Probenentnahme. - 4.2 Behandlung und Aufarbeitung des Probenmaterials. - 4.3 Bodenphysikalische Analysen. - 4.3.1 Bestimmung der Porengrößenverteilung. - 4.3.2 Bestimmung der Korngrößenverteilung. - 4.4 Bodenchemische Analysen. - 4.4.1 Kohlenstoff- und Stickstoff-Analytik. - 4.4.2 δ13C-Bestimmung. - 4.4.3 pH-Wert- und Redoxpotential-Messung. - 4.5 Bestimmung von Methanbildungsraten. - 4.5.1 Aktuelle Methanbildungsaktivität. - 4.5.2 Potentielle Methanbildungsaktivität. - 4.5.3 Temperaturoptimum der Methanbildung. - 4.5.4 Methanbildungsaktivität in Gegenwart von Sauerstoff. - 4.5.5 Methanbildungsaktivität in Abhängigkeit vom Wassergehalt. - 4.5.6 Gasanalytik (CH4, O2). - 4.6 Mikrobiologische Untersuchungen. - 4.6.1 Nährmedien. - 4.6.2 Zellzahlbestimmung. - 4.6.2.1 MPN-Methode. - 4.6.2.2 Koch'sches Plattengußverfahren. - 4.6.3 Isolierung von methanogenen Bakterien. - 4.6.3.1 Anreicherungskultur. - 4.6.3.2 Agar-Verdünnungsreihe. - 4.6.3.3 Reinheitstest. - 4.6.3.4 Kulturführung. - 4.6.4 Charakterisierung der Isolate. - 4.6.4.1 Transmissionselektronenmikroskopie. - 4.6.4.2 Substratspektrum. - 4.6.4.3 Temperaturoptimum. - 4.6.4.4 Salz- und pH-Optimum. - 4.6.4.5 16S-rRNA-Sequenzierung. - 4.7 Fraktionierung der organischen Bodensubstanz. - 4.7.1 DOC-Extraktion. - 4.7.2 Dichtefraktionierung. - 4.8 Statistische Analysen. - 5 Ergebnisse. - 5.1 Bodenmikrobiologische Standortcharakterisierung. - 5.1.1 Marschboden (Asseler Sand). - 5.1.2 Reisboden (Suzhou). - 5.2 Temperaturcharakteristika der methanogenen Flora. - 5.2.1 Temperaturoptimum. - 5.2.2 Zellzahlen methanogener Bakterien in Abhängigkeit von Temperatur und Substrat. - 5.2.3 Einfluß verschiedener Substrate auf die Methanbildung. - 5.3 Isolierung und Charakterisierung von methanogenen Bakterien. - 5.3.1 Grundlagen. - 5.3.2 Morphologische Merkmale. - 5.3.3 Physiologische Merkmale. - 5.3.4 Phylogenetische Auswertung der 16S-rRNA-Sequenzen methanogener Bakterien. - 5.4 Methanbildung in Gegenwart von Sauerstoff. - 5.4.1 Hemmung der Methanoxidation durch Acetylen. - 5.4.2 Einfluß der Begleitflora. - 5.4.3 Einfluß der Bodenart. - 5.4.4 Methanbildung bei definierten Wassergehalten. - 5.5 Charakterisierung der organischen Bodensubstanz in Hinblick auf die CH4-Bildung. - 5.5.1 Marschboden (Asseler Sand). - 5.5.1.1 Kohlenstoff- und Stickstoff-Parameter. - 5.5.1.2 Isotopenanalytische Untersuchung der organischen Bodensubstanz. - 5.5.1.3 Charakterisierung der organischen Substanz mittels Dichtefraktionierung. - 5.5.2 Reisboden (Suzhou). - 5.5.2.1 Kohlenstoff- und Stickstoff-Parameter. - 5.5.2.2 Isotopenanalytische Untersuchungen der organischen Bodensubstanz. - 5.5.2.3 Charakterisierung der organischen Substanz mittels Dichtefraktionierung. - 5.5.2.4 Methanbildungspotentiale der Dichtefraktionen. - 6 Diskussion. - 6.1 Einfluß der Temperatur auf die Methanbildung. - 6.2 Einfluß von Sauerstoff auf die Aktivität methanogener Bakterien. - 6.3 Bedeutung der organischen Bodensubstanz für die Methanogenese. - 6.4 Wechselwirkungen der untersuchten Faktoren in hydromorphen Böden und ihre Bedeutung in Hinblick auf globale Klimaveränderungen. - 7 Literaturverzeichnis. - 8 Anhang. - Danksagung.

Note: Table of content Abstract Zusammenfassung 1. Introduction 1.1. Motivation 1.2. Scientific Background 1.2.1. Permafrost in arctic environments 1.2.2. Carbon storage and emission in arctic environments 1.2.3. Methane cycling in arctic environments 1.3. Study Sites 1.3.1. Lena-Delta, Siberia 1.3.2. El’gygytgyn Crater Lake, Chukotka 1.4. Objectives and approach 1.5. Thesis organization 1.6. Summary of the included manuscripts and contribution of the co-authors 1.6.1. Response of methanogenic archaea to Late Pleistocene and Holocene climate changes in the Siberian Arctic 1.6.2. Response of microbial communities to landscape and climatic changes in a terrestrial permafrost sequence of the El’gygytgyn crater, Far East Russian Arctic 1.6.3. Glacial-interglacial microbial community dynamics in Middle Pleistocene sediments in the Lake El’gygytgyn, Far East Russian Arctic 2. Response of methanogenic archaea to Late Pleistocene and Holocene climate changes in the Siberian Arctic 2.1. Abstract 2.2. Introduction 2.3. Materials and Methods 2.3.1. Study site 2.3.2. Permafrost drilling and sample preparation 2.3.3. Sediment properties 2.3.4. Potential methane production rates 2.3.5. Lipid biomarker analysis 2.3.6. Detection of archaeol and isoprenoid GDGTs 2.3.7. Detection of PLFAs and PLELs 2.3.8. DNA extraction and polymerase chain reaction (PCR) amplification 2.3.9. Phylogenetic analysis 2.4. Results and Discussion 2.4.1. Methane profile of the Kurungnakh permafrost sequence 2.4.2. Signals of living microbial communities in the Kurungnakh permafrost sequence 2.4.3. Reconstruction of past microbial communities in the Kurungnakh permafrost sequence 2.4.4. Climate impact on the distribution of microbial communities in the Kurungnakh permafrost sequence 2.4.5. Climatic impact on the composition of methanogenic communities in the Kurungnakh permafrost sequence 2.5. Conclusion 2.6. Acknowledgement 3. Response of microbial communities to landscape and climatic changes in a terrestrial permafrost sequence of the El’gygytgyn crater, Far East Russian Arctic 3.1. Abstract 3.2. Introduction 3.3. Materials and Methods 3.3.1.Study site 3.3.2. Drilling and sample material 3.3.3. Sediment properties 3.3.4. Lipid biomarker analysis 3.3.5. Detection of glycerol dialkyl glycerol tetraethers (GDGTs) and archaeol 3.3.6. Detection of phospholipid fatty acids (PLFA) 3.3.7. Deoxyribonucleic acid (DNA) extraction and amplification 3.3.8. Quantitative PCR analysis of archaeal and bacterial small sub unit (SSU) rRNA genes 3.3.9. Phylogenetic analysis 3.4. Results 3.4.1. TOC-contents 3.4.2. Distribution of glycerol dialkyl glycerol tetraethers (GDGTs) and archaeol 3.4.3. Distribution of phospholipid fatty acids (PLFA) 3.4.4. Composition of archaeol and isoprenoid GDGTs 3.4.5. Quantification of bacterial and archaeal genes 3.4.6. Analysis of methanogenic community fingerprints 3.5. Discussion 3.5.1. Microbial communities in subaquatic deposits 3.5.2. Microbial communities in subaerial deposits 3.5.3. Microbial succession in the Holocene sequence of Lake El’gygytgyn permafrost 3.6.Conclusion 3.7. Acknowledgements 4. Glacial-interglacial microbial community dynamics in Middle Pleistocene sediments in the Lake El’gygytgyn, Far East Russian Arctic 4.1. Abstract 4.2. Introduction 4.3. Materials and Methods 4.3.1. Study site 4.3.2. Drilling and sample preparation 4.3.3. Sediment properties 4.3.4. Lipid biomarker analyses 4.3.5. Deoxyribonucleic acid (DNA) extraction and quantitative polymerase chain reaction (qPCR) 4.3.6. PCR amplification of methanogenic SSU rRNA genes 4.4. Results 4.4.1. Sedimentary TOC and biogenic silica concentration 4.4.2. Quantification of bacterial and archaeal genes 4.4.3. Quantification and composition of lipid biomarkers 4.4.4. Potential methane production 4.4.5. Methanogenic community composition 4.5. Discussion 4.6. Acknowledgements 5. Synthesis 5.1. The reaction of microbial communities to past climatic change in the Arctic 5.2.The response of microbial communities to carbon composition and availability 5.3. Implications from this study for future research 6. Data collection 6.1. Manuscript I: Response of methanogenic archaea to Late Pleistocene and Holocene climate changes in the Siberian Arctic 6.1.1. Sediment properties 6.1.2. Isoprenoid glycerol dialkyl glycerol tetraethers and archaeol 6.1.3. Branched glycerol dialkyl glycerol tetraethers 6.1.4. Phospholipid ester and ether lipids (summary) 6.2. Manuscript II: Response of microbial communities to landscape and climatic changes in a terrestrial permafrost sequence of the El’gygytgyn crater, Far East Russian Arctic 6.2.1. Sediment properties and gene quantifications 6.2.2. Phospholipid fatty acids composition 6.2.3. Isoprenoid glycerol dialkyl glycerol tetraethers and archaeol 6.2.4. Branched glycerol dialkyl glycerol tetraethers 6.3. Manuscript III: Glacial-interglacial microbial community dynamics in Middle Pleistocene sediments in the Lake El’gygytgyn, Far East Russian Arctic 6.3.1. Sediment properties and gene quantifications 6.3.2. Isoprenoid glycerol dialkyl glycerol tetraethers and archaeol 6.3.3. Branched glycerol dialkylglycerol tetraethers 7. References 8. Final thoughts and acknowledgements 9. Curriculum vitae 10.Erklärung

Note: Dissertation, Universität Potsdam, 2020 , Contents Preface Acknowledgements Contents Summary Zusammenfassung List of abbreviations Chapter 1. Introduction 1.1 Motivation 1.2 Carbon storage in Arctic permafrost environments and the permafrost carbon feedback (PCF) 1.3 Methane cycling microorganisms 1.4 The microbial ecology of permafrost 1.5 Plasmids and their potential role in stress tolerance 1.6 Objectives Chapter 2. Study sites 2.1 Regional settings 2.2 Kurungnakh and Samoylov Island 2.3 Bol'shoy Lyakhovsky Island 2.4 Herschel Island Chapter 3. Manuscripts 3.1 Overview of manuscripts, including contribution of co-authors. 3.2 Manuscript I Methanogenic response to long-term permafrost thaw is determined by paleoenvironment 3.3 Manuscript II Methane production in thawing permafrost is constrained by methanogenic population size and carbon density 3.4 Manuscript III Metaplasmidome-encoded functional potential of permafrost active layer soils Chapter 4. Synthesis 4.1 Introduction 4.2 Constraints behind methane production from thawing permafrost 4.3 The methanogenic community response to long-term permafrost thaw 4.4 The adaptive potential of the permafrost micro biota to cope with stress factors during global warming 4.5 Conclusion Chapter 5. Future research directions and perspectives Chapter 6. References Chapter 7. Appendix 7.1 Supporting information for manuscript I 7.2 Supporting information for manuscript II 7.3 Supporting information for manuscript III 7.4 ESR collaboration, manuscript IV

Note: Dissertation, Universität Potsdam, 2015 , Table of contents Preface Table of contents Summary Zusammenfassung 1. Introduction 1.1. Environmental conditions on past and present Mars 1.2. Detection of methane on Mars 1.3. Methanogenic archaea 1.4. Description of study sites 1.5. Aims and approaches 1.6. Overview of the publications 2. Publication I: Methanosarcina soligelidi sp. nov., a desiccationandfreeze-thaw-resistant methanogenic archaeon from a Siberianpermafrost-affected soil 3. Publication II: Methanobacterium movilense sp. nov.,ahydrogenotrophic, secondary-alcohol-utilizing methanogen fromthe anoxic sediment of a subsurface lake 4. Publication III: Influence of Martian Regolith Analogs on the activityand growth of methanogenic archaea,with special regard to long-term desiccation 5. Publication IV: Laser spectroscopic real time measurements ofmethanogenic activity under simulated Martian subsurface conditions 6. Synthesis and Conclusion 6.1. Synthesis 6.2. Conclusion and future perspectives 7. References 8. Acknowledgments