ALBERT

Description: Recent collection efforts in the upper Campanian (∼76-73.5 Ma) Fruitland and Kirtland formations of northwestern New Mexico have significantly increased the taxonomic diversity of lizards in this historically poorly understood squamate assemblage. New lizard specimens from the “Hunter Wash Local Fauna” of the upper Fruitland and lower Kirtland formations include: (1) new specimens referable to Chamopsiidae; (2) new material belonging to Scincomorpha, (3) new material belonging to Anguidae; and (4) the first reported predatory lizard (Platynota) material from the Campanian of New Mexico. The increase in lizard diversity in the “Hunter Wash Local Fauna” expands our understanding of Late Cretaceous squamate taxonomy, distribution, and diversity in the Western Interior of North America (Laramidia). Collectively, the described specimens represent family-level diversity similar to that seen in other Campanian foreland basin deposits of the Western Interior, such as the mid-paleolatitude Kaiparowits Formation of southern Utah, the higher paleolatitude Dinosaur Park Formation of southern Alberta, and the lower paleolatitude Aguja Formation of southwestern Texas. The lizards of the “Hunter Wash Local Fauna” represent crucial mid-paleolatitude data from a coastal plain depositional setting in Laramidia—allowing for comparisons to more well-studied assemblages at different latitudes and in different depositional settings.

Description: Tyrannosaurids are hypothesized to be gregarious, possibly parasocial carnivores engaging in cooperative hunting and extended parental care. A tyrannosaurid (cf. Teratophoneus curriei) bonebed in the late Campanian age Kaiparowits Formation of southern Utah, nicknamed the Rainbows and Unicorns Quarry (RUQ), provides the first opportunity to investigate possible tyrannosaurid gregariousness in a taxon unique to southern Laramidia. Analyses of the site’s sedimentology, fauna, flora, stable isotopes, rare earth elements (REE), charcoal content and taphonomy suggest a complex history starting with the deaths and transport of tyrannosaurids into a peri-fluvial, low-energy lacustrine setting. Isotopic and REE analyses of the fossil material yields a relatively homogeneous signature indicating the assemblage was derived from the same source and represents a fauna living in a single ecospace. Subsequent drying of the lake and fluctuating water tables simultaneously overprinted the bones with pedogenic carbonate and structurally weakened them through wet-dry cycling. Abundant charcoal recovered from the primary bone layer indicate a low temperature fire played a role in the site history, possibly triggering an avulsion that exhumed and reburied skeletal material on the margin of a new channel with minimal transport. Possible causes of mortality and concentration of the tyrannosaurids include cyanobacterial toxicosis, fire, and flooding, the latter being the preferred hypothesis. Comparisons of the RUQ site with other North American tyrannosaur bonebeds (Dry Island-Alberta; Daspletosaurus horneri-Montana) suggest all formed through similar processes. Combined with ichnological evidence, these tyrannosaur mass-burial sites could be part of an emerging pattern throughout Laramidia reflecting innate tyrannosaurid behavior such as habitual gregariousness.