ALBERT

Notes: Abstract The ultrastructural features of the ovary and oogenesis are described in the eastern oyster,Crassostrea virginica (Gmelin, 1791). The ovary is a diffuse organ consisting of highly branching acini in which oocytes develop. The acini are surrounded by a matrix of vesicular connective tissue (VCT tells) which serves a nutrient storage function. Each acinus is bathed by fluid within a surrounding connective tissue compartment, the hemocoel, which likely serves as a means of transporting nutrients to the oocytes. Oocytes begin growth while positioned near to the inner acinus wall. As differentiation proceeds and they enter the late stages of vitellogenesis, they become stalle-shaped and project into the acinus lumen. Follicle tells are closely associated with oocytes during the early and middle stages of vitellogenesis but they are largely confined to the basal, stalked region of late-stage oocytes. Vitellogenesis occurs through a process of autosynthesis, involving the combined activity of the Golgi complex and rough endoplasmic reticulum, and heterosynthesis in which extraovarian precursors are incorporated into oocytes via receptor-mediated endocytosis involving the basal surface of the oocytes. It is suggested that the follicle tells play some important role during oogenesis but probably are not the major source of yolk precursors. The VCT celas are probably the main source of nutrients for vitellogenesis.

Notes: Abstract The ultrastructural features of the testis and spermatogenesis have been described from the eastern oyster,Crassostrea virginica (Gmelin, 1791). The testis is a diffuse organ consisting of branching acini containing differentiating sperm in a variety of stages. Spermatogonia are located nearest the outer wall of the acinus, while spermatocytes and spermatids are positioned nearer the lumen. Mature spermatozoa are all confined to the central region of the acinus. The acinus is surrounded by an intermittent layer of myoepithelial cells and is bathed by a $uid-filled hemocoel. Vesicular connective tissue (VCT cells) fills the region between adjacent acini, and the cells contain glycogen granules and lipid droplets. Each acinus is divided radially into subcompartments that are partially separated by pleomorphic accessory cells which remain in close contact with sperm until late stages of development. Sperm are similar to those described in other oysters, except that five midpiece mitochondria were observed in some sperm rather than the usual four, and the acrosomal vesicle lacked the “whorled” substructure described in some other oyster sperm. We suggest that the neutral term “accessory cells” be applied to bivalve testicular somatic cells until more detailed studies are available to justify the use of “Sertoli cell” and other descriptive terms which have previously been adopted from other taxa only distantly related to bivalves.

Notes: Abstract A study has been made of the interaction between the thermosphere and the ionosphere at high latitudes, with particular regard to the value of the O+-O collision parameter. The European incoherent scatter radar (EISCAT) was used to make tristatic measurements of plasma parameters at F-region altitudes while simultaneous measurements of the neutral wind were made by a Fabry-Perot interferometer (FPI). The radar data were used to derive the meridional neutral winds in a way similar to that used by previous authors. The accuracy of this technique at high latitudes is reduced by the dynamic nature of the auroral ionosphere and the presence of significant vertical winds. The derived winds were compared with the meridional winds measured by the FPI. For each night, the value of the O+-O collision parameter which produced the best agreement between the two data sets was found. The precision of the collision frequency found in this way depends on the accuracy of the data. The statistical method was critically examined in an attempt to account for the variability in the data sets. This study revealed that systematic errors in the data, if unaccounted for by the analysis, have a tendency to increase the value of the derived collision frequency. Previous analyses did not weight each data set in order to account for the quality of the data; an improved method of analysis is suggested.

Notes: Abstract We present an analysis of the accuracy of the method introduced by Lockwood et al. (1994) for the determination of the magnetopause reconnection rate from the dispersion of precipitating ions in the ionospheric cusp region. Tests are made by applying the method to synthesised data. The simulated cusp ion precipitation data are produced by an analytic model of the evolution of newly-opened field lines, along which magnetosheath ions are firstly injected across the magnetopause and then dispersed as they propagate into the ionosphere. The rate at which these newly opened field lines are generated by reconnection can be varied. The derived reconnection rate estimates are then compared with the input variation to the model and the accuracy of the method assessed. Results are presented for steady-state reconnection, for continuous reconnection showing a sine-wave variation in rate and for reconnection which only occurs in square wave pulses. It is found that the method always yields the total flux reconnected (per unit length of the open-closed field-line boundary) to within an accuracy of better than 5%, but that pulses tend to be smoothed so that the peak reconnection rate within the pulse is underestimated and the pulse length is overestimated. This smoothing is reduced if the separation between energy channels of the instrument is reduced; however this also acts to increase the experimental uncertainty in the estimates, an effect which can be countered by improving the time resolution of the observations. The limited time resolution of the data is shown to set a minimum reconnection rate below which the method gives spurious short-period oscillations about the true value. Various examples of reconnection rate variations derived from cusp observations are discussed in the light of this analysis.

Notes: Abstract Early in 1996, the latest of the European inco-herent-scatter (EISCAT) radars came into operation on the Svalbard islands. The EISCAT Svalbard Radar (ESR) has been built in order to study the ionosphere in the northern polar cap and in particular, the dayside cusp. Conditions in the upper atmosphere in the cusp region are complex, with magnetosheath plasma cascading freely into the atmosphere along open magnetic field lines as a result of magnetic reconnection at the dayside magnetopause. A model has been developed to predict the effects of pulsed reconnection and the subsequent cusp precipitation in the ionosphere. Using this model we have successfully recreated some of the major features seen in photometer and satellite data within the cusp. In this paper, the work is extended to predict the signatures of pulsed reconnection in ESR data when the radar is pointed along the magnetic field. It is expected that enhancements in both electron concentration and electron temperature will be observed. Whether these enhancements are continuous in time or occur as a series of separate events is shown to depend critically on where the open/closed field-line boundary is with respect to the radar. This is shown to be particularly true when reconnection pulses are superposed on a steady background rate.

Notes: Abstract Superposed epoch studies have been carried out in order to determine the ionospheric response at mid-latitudes to southward turnings of the interplanetary magnetic field (IMF). This is compared with the geomagnetic response, as seen in the indices Kp, AE and Dst. The solar wind, IMF and geomagnetic data used were hourly averages from the years 1967–1989 and thus cover a full 22-year cycle in the solar magnetic field. These data were divided into subsets, determined by the magnitudes of the southward turnings and the concomitant increase in solar wind pressure. The superposed epoch studies were carried out using the time of the southward turning as time zero. The response of the mid-latitude ionosphere is studied by looking at the F-layer critical frequencies, foF2, from hourly soundings by the Slough ionosonde and their deviation from the monthly median values, δfoF2. For the southward turnings with a change in Bz of δBz 〉 11.5 nT accompanied by a solar wind dynamic pressure P exceeding 5 nPa, the F region critical frequency, foF2, shows a marked decrease, reaching a minimum value about 20 h after the southward turning. This recovers to pre-event values over the subsequent 24 h, on average. The Dst index shows the classic storm-time decrease to about −60 nT. Four days later, the index has still to fully recover and is at about −25 nT. Both the Kp and AE indices show rises before the southward turnings, when the IMF is strongly northward but the solar wind dynamic pressure is enhanced. The average AE index does register a clear isolated pulse (averaging 650 nT for 2 h, compared with a background peak level of near 450 nT at these times) showing enhanced energy deposition at high latitudes in substorms but, like Kp, remains somewhat enhanced for several days, even after the average IMF has returned to zero after 1 day. This AE background decays away over several days as the Dst index recovers, indicating that there is some contamination of the currents observed at the AE stations by the continuing enhanced equatorial ring current. For data averaged over all seasons, the critical frequencies are depressed at Slough by 1.3 MHz, which is close to the lower decile of the overall distribution of δfoFl values. Taking 30-day periods around summer and winter solstice, the largest depression is 1.6 and 1.2 MHz, respectively. This seasonal dependence is confirmed by a similar study for a Southern Hemisphere station, Argentine Island, giving peak depressions of 1.8 MHz and 0.5 MHz for summer and winter. For the subset of turnings where δBz 〉 11.5 nT and P ≤ 5 nPa, the response of the geomagnetic indices is similar but smaller, while the change in δfoF2 has all but disappeared. This confirms that the energy deposited at high latitudes, which leads to the geomagnetic and ionospheric disturbances following a southward turning of the IMF, increases with the energy density (dynamic pressure) of the solar wind flow. The magnitude of all responses are shown to depend on δBz. At Slough, the peak depression always occurs when Slough rotates into the noon sector. The largest ionospheric response is for southward turnings seen between 15–21 UT.

Notes: Abstract The open magnetosphere model of cusp ion injection, acceleration and precipitation is used to predict the dispersion characteristics for fully pulsed magnetic reconnection at a low-latitude magnetopause X-line. The resulting steps, as would be seen by a satellite moving meridionally and normal to the ionospheric projection of the X-line, are compared with those seen by satellites moving longitudinally, along the open/closed boundary. It is shown that two observed cases can be explained by similar magnetosheath and reconnection characteristics, and that the major differences between them are well explained by the different satellite paths through the events. Both cases were observed in association with poleward-moving transient events seen by ground-based radar, as also predicted by the theory. The results show that the reconnection is pulsed but strongly imply it cannot also be spatially patchy, in the sense of isolated X-lines which independently are intermittently active. Furthermore they show that the reconnection pulses responsible for the poleward-moving events and the cusp ion steps, must cover at least 3 h of magnetic local time, although propagation of the active reconnection region may mean that it does not extend this far at any one instant of time.

Notes: Abstract Possible configurations of the magnetic field in the outer magnetosphere during geomagnetic polarity reversals are investigated by considering the idealized problem of a magnetic multipole of order m and degree n located at the centre of a spherical cavity surrounded by a boundless perfect diamagnetic medium. In this illustrative idealization, the fixed spherical (magnetopause) boundary layer behaves as a perfectly conducting surface that shields the external diamagnetic medium from the compressed multipole magnetic field, which is therefore confined within the spherical cavity. For a general magnetic multipole of degree n, the non-radial components of magnetic induction just inside the magnetopause are increased by the factor 1 + [(n + 1)/n] relative to their corresponding values in the absence of the perfectly conducting spherical magnetopause. An exact equation is derived for the magnetic field lines of an individual zonal (m = 0), or axisymmetric, magnetic multipole of arbitrary degree n located at the centre of the magnetospheric cavity. For such a zonal magnetic multipole, there are always two neutral points and n – 1 neutral rings on the spherical magnetopause surface. The two neutral points are located at the poles of the spherical magnetopause. If n is even, one of the neutral rings is coincident with the equator; otherwise, the neutral rings are located symmetrically with respect to the equator. The actual existence of idealized higher-degree (n 〉 1) axisymmetric magnetospheres would necessarily imply multiple (n + 1) magnetospheric cusps and multiple (n) ring currents. Exact equations are also derived for the magnetic field lines of an individual non-axisymmetric magnetic multipole, confined by a perfectly conducting spherical magnetopause, in two special cases; namely, a symmetric sectorial multipole (m = n) and an antisymmetric sectorial multipole (m = n – 1). For both these non-axisymmetric magnetic multipoles, there exists on the spherical magnetopause surface a set of neutral points linked by a network of magnetic field lines. Novel magnetospheric processes are likely to arise from the existence of magnetic neutral lines that extend from the magnetopause to the surface of the Earth. Finally, magnetic field lines that are confined to, or perpendicular to, either special meridional planes or the equatorial plane, when the multipole is in free space, continue to be confined to, or perpendicular to, these same planes when the perfectly conducting magnetopause is present.