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  • 1
    ISSN: 1365-2486
    Source: Blackwell Publishing Journal Backfiles 1879-2005
    Topics: Biology , Energy, Environment Protection, Nuclear Power Engineering , Geography
    Notes: Physiological and chemical responses of 17 birch (Betula pendula Roth) clones to 1.5–1.7 × ambient ozone were studied in an open-field experiment over two growing seasons. The saplings were studied for growth, foliar visible injuries, net photosynthesis, stomatal conductance, and chlorophyll, carotenoid, Rubisco, total soluble protein, macronutrient and phenolic concentrations in leaves. Elevated ozone resulted in growth enhancement, changes in shoot-to-root (s/r) ratio, visible foliar injuries, reduced stomatal conductance, lower late-season net photosynthesis, foliar nutrient imbalance, changes in phenolic composition, and reductions in pigment, Rubisco and soluble protein contents indicating accelerated leaf senescence. Majority of clones responded to ozone by changing C allocation towards roots, by stomatal closure (reduced ozone uptake), and by investment in low-cost foliar antioxidants to avoid and tolerate ozone stress. A third of clones, showing increased s/r ratio, relied on inducible efficient high-cost antioxidants, and enhanced leaf production to compensate ozone-caused decline in leaf-level net photosynthesis. However, the best ozone tolerance was found in two s/r ratio-unaffected clones showing a high constitutive amount of total phenolics, investment in low-cost antioxidants and N distribution to leaves, and lower stomatal conductance under ozone stress. The results highlight the importance of phenolic compounds in ozone defence mechanisms in the birch population. Depending on the genotype, ozone detoxification was improved by an increase in either efficient high-cost or less efficient low-cost antioxidative phenolics, with close connections to whole-plant physiology.
    Type of Medium: Electronic Resource
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  • 2
    Electronic Resource
    Electronic Resource
    Oxford, UK : Blackwell Science Ltd
    Plant, cell & environment 26 (2003), S. 0 
    ISSN: 1365-3040
    Source: Blackwell Publishing Journal Backfiles 1879-2005
    Topics: Biology
    Notes: A long-term free air ozone fumigation experiment was conducted to study changes in physiological ozone responses during tree ontogeny and exposure time in ozone sensitive and tolerant clones of European white birch (Betula pendula Roth), originated from south and central Finland. The trees were grown in soil in natural microclimatic conditions under ambient ozone (control) and 1.4–1.7 × ambient (elevated) ozone from May 1996 to October 2001, and were measured for stem and foliage growth, net photosynthesis, stomatal conductance, stomatal density, visible injuries, foliar starch content and bud formation. After 6 years of exposure, the magnitude of ozone-induced growth reductions in the sensitive clone was 12–48% (significant difference), levels similar or greater than those reported earlier for 2- and 3-year-old saplings undergoing shorter exposures. In the tolerant clone, growth of these larger trees was reduced by 1–38% (significant difference in stem volume), although the saplings had previously been unaffected. In both clones, ozone stress led to significantly reduced leaf-level net photosynthesis but significantly increased stomatal conductance rates during the late summer, resulting in a lower carbon gain for bud formation and the onset of visible foliar injuries. Increasing ozone sensitivity with duration of exposure was explained by a change in growth form (relatively reduced foliage mass), a lower photosynthesis to stomatal conductance ratio during the late summer, and deleterious carry-over effects arising from the reduced number of over-wintering buds.
    Type of Medium: Electronic Resource
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  • 3
    Electronic Resource
    Electronic Resource
    Oxford, UK : Blackwell Science Ltd
    Plant, cell & environment 22 (1999), S. 0 
    ISSN: 1365-3040
    Source: Blackwell Publishing Journal Backfiles 1879-2005
    Topics: Biology
    Notes: In the first experiment, saplings of ozone-sensitive and a more tolerant clone of Betula pendula Roth were exposed to ambient ozone (control treatment, accumulated exposure over a threshold 40 nmol mol−1 (AOT40) exposure of 1·0 μmol mol−1 h) and 1·5 × ambient ozone (elevated-ozone treatment, AOT40 of 17·3 μmol mol−1 h) over one growing season, 1996. After over-wintering, the dormant elevated-ozone saplings were transferred to the control blocks and assessed for short-term carry-over effects during the following growing season. In the second experiment, three sensitive, four intermediate and three tolerant clones were grown under ambient ozone (control treatment, AOT40 of 0·5–0·8 μmol mol−1 h per growing season) and 1·6–1·7 × ambient ozone (elevated-ozone treatment, AOT40 of 18·3–18·6 μmol mol−1 h per growing season) from May 1994 until May 1996, and were assessed for long-term carry-over effects during growing season 1997, after a 12–16 months recovery period. Deleterious short-term carry-over effects of ozone exposure included reduced contents of Rubisco, chlorophyll, carotenoids, starch and nutrients in leaves, lower stomatal conductance, and decreased new shoot growth and net assimilation rate, followed by a 7·5% (shoot dry weight (DW)), 15·2% (root DW) and 23·2% (foliage area) decreased biomass accumulation and yield over the long term, including a reduced root : shoot ratio. However, a slow recovery of relative growth rates during the following two seasons without elevated ozone was apparent. Several long-lasting structural, biochemical and stomatal acclimation, stress-defence and compensation reactions were observed in the ozone-tolerant clone, whereas in the sensitive clone allocation shifted from growth towards defensive phenolics such as chlorogenic acid. The results provide evidence of persistent deleterious effects of ozone which remain long after the ozone episode.
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  • 4
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    New York : Periodicals Archive Online (PAO)
    Explorations in economic history. 17:2 (1980:Apr.) 165 
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  • 5
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    Unknown
    New York : Periodicals Archive Online (PAO)
    Explorations in economic history. 11:1 (1973:Fall) 73 
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  • 6
    Electronic Resource
    Electronic Resource
    Springer
    Empirical economics 12 (1987), S. 97-106 
    ISSN: 1435-8921
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract Simulation experiments are used to compare OLS and several principal components (PCR) estimators of the classical linear regression model. According to mean square error and mean absolute error criteria OLS dominates PCR in these experiments.
    Type of Medium: Electronic Resource
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  • 7
    Electronic Resource
    Electronic Resource
    Springer
    Empirical economics 9 (1984), S. 139-150 
    ISSN: 1435-8921
    Source: Springer Online Journal Archives 1860-2000
    Topics: Economics
    Notes: Abstract This study provides a tentative explanation of the spatial distribution of economic activity in Canada, with a focus upon the role of inter-industry linkages. Location of economic activity for each of one hundred and nine three-digit S.I.C. manufacturing industries is explained by use of a “tobit” model incorporating backward and forward linkage variables. The two sets of linkage variables in each tobit equation were reduced by extraction of one principal component from each set, using the matrix of cosines of the variables. A set of control variables completes the set of explanatory variables. The overall explanatory power of our equations was remarkably high and the role of inter-industry linkages is unmistakable.
    Type of Medium: Electronic Resource
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  • 8
    Publication Date: 2015-09-25
    Description: Climate warming is having an impact on distribution, acclimation and defence capability of plants. We compared the emission rate and composition of volatile organic compounds (VOCs) from silver birch ( Betula pendula (Roth)) provenances along a latitudinal gradient in a common garden experiment over the years 2012 and 2013. Micropropagated silver birch saplings from three provenances were acquired along a gradient of 7° latitude and planted at central (Joensuu 62°N) and northern (Kolari 67°N) sites. We collected VOCs emitted by shoots and assessed levels of herbivore damage of three genotypes of each provenance on three occasions at the central site and four occasions at the northern site. In 2012, trees of all provenances growing at the central site had higher total VOC emission rates than the same provenances growing at the northern site; in 2013 the reverse was true, thus indicating a variable effect of latitude. Trees of the southern provenance had lower VOC emission rates than trees of the central and northern provenances during both sampling years. However, northward or southward translocation itself had no significant effect on the total VOC emission rates, and no clear effect on insect herbivore damage. When VOC blend composition was studied, trees of all provenances usually emitted more green leaf volatiles at the northern site and more sesquiterpenes at the central site. The monoterpene composition of emissions from trees of the central provenance was distinct from that of the other provenances. In summary, provenance translocation did not have a clear effect in the short-term on VOC emissions and herbivory was not usually intense at the lower latitude. Our data did not support the hypothesis that trees growing at lower latitudes would experience more intense herbivory, and therefore allocate resources to chemical defence in the form of inducible VOC emissions.
    Print ISSN: 0829-318X
    Electronic ISSN: 1758-4469
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
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  • 9
    Publication Date: 2012-09-14
    Description: Northern forests are currently experiencing increasing mean temperatures, especially during autumn and spring. Consequently, alterations in carbon sequestration, leaf biochemical quality and freezing tolerance (FT) are likely to occur. The interactive effects of elevated temperature and ozone (O 3 ), the most harmful phytotoxic air pollutant, on Norway spruce ( Picea abies (L.) Karst.) seedlings were studied by analysing phenology, metabolite concentrations in the needles, FT and gas exchange. Sampling was performed in September and May. The seedlings were exposed to a year-round elevated temperature (+1.3 °C), and to 1.4 x ambient O 3 concentration during the growing season in the field. Elevated temperature increased the concentrations of amino acids, organic acids of the citric acid cycle and some carbohydrates, and reduced the concentrations of phenolic compounds, some organic acids of the shikimic acid pathway, sucrose, cyclitols and steroids, depending on the timing of the sampling. Although growth onset occurred earlier at elevated temperature, the temperature of 50% lethality (LT 50 ) was similar in the treatments. Photosynthesis and the ratio of photosynthesis to dark respiration were reduced by elevated temperature. Elevated concentrations of O 3 reduced the total concentration of soluble sugars, and tended to reduce LT 50 of the needles in September. These results show that alterations in needle chemical quality can be expected at elevated temperatures, but the seedlings' sensitivity to autumn and spring frosts is not altered. Elevated O 3 has the potential to disturb cold hardening of Norway spruce seedlings in autumn, and to alter the water balance of the seedling through changes in stomatal conductance ( g s ), while elevated temperature is likely to reduce g s and consequently reduce the O 3 -flux inside the leaves.
    Print ISSN: 0829-318X
    Electronic ISSN: 1758-4469
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
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  • 10
    Publication Date: 2012-06-23
    Description: In the present experiment, the single and combined effects of elevated temperature and ozone (O 3 ) on four silver birch genotypes (gt12, gt14, gt15 and gt25) were studied in an open-air field exposure design. Above- and below-ground biomass accumulation, stem growth and soil respiration were measured in 2008. In addition, a 13 C-labelling experiment was conducted with gt15 trees. After the second exposure season, elevated temperature increased silver birch above- and below-ground growth and soil respiration rates. However, some of these variables showed that the temperature effect was modified by tree genotype and prevailing O 3 level. For instance, in gt14 soil respiration was increased in elevated temperature alone (T) and in elevated O 3 and elevated temperature in combination (O 3 + T) treatments, but in other genotypes O 3 either partly (gt12) or totally nullified (gt25) temperature effects on soil respiration, or acted synergistically with temperature (gt15). Before leaf abscission, all genotypes had the largest leaf biomass in T and O 3 + T treatments, whereas at the end of the season temperature effects on leaf biomass depended on the prevailing O 3 level. Temperature increase thus delayed and O 3 accelerated leaf senescence, and in combination treatment O 3 reduced the temperature effect. Photosynthetic : non-photosynthetic tissue ratios (P : nP ratios) showed that elevated temperature increased foliage biomass relative to woody mass, particularly in gt14 and gt12, whereas O 3 and O 3 + T decreased it most clearly in gt25. O 3 -caused stem growth reductions were clearest in the fastest-growing gt14 and gt25, whereas mycorrhizal root growth and sporocarp production increased under O 3 in all genotypes. A labelling experiment showed that temperature increased tree total biomass and hence 13 C fixation in the foliage and roots and also label return was highest under elevated temperature. Ozone seemed to change tree 13 C allocation, as it decreased foliar 13 C excess amount, simultaneously increasing 13 C excess obtained from the soil. The present results suggest that warming has potential to increase silver birch growth and hence carbon (C) accumulation in tree biomass, but the final magnitude of this C sink strength is partly counteracted by temperature-induced increase in soil respiration rates and simultaneous O 3 stress. Silver birch populations' response to climate change will also largely depend on their genotype composition.
    Print ISSN: 0829-318X
    Electronic ISSN: 1758-4469
    Topics: Agriculture, Forestry, Horticulture, Fishery, Domestic Science, Nutrition
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