ALBERT

Notes: In a study of the weed flora of dryland crops in the Córdoba region (southern Spain) in 1983 and 1984, a total of 256 species from 41 families was identified in the 201 fields surveyed. The families with the highest numbers of species were Compositae, Gramineae, Leguminosae and Cruciferae. Therophytes represented 79% of the biological spectrum, with hemicryptophytes and geophytes more especially in the spring-sown crops. At the early and late crop growth stages about 4% and 8%, respectively, of the fields had severe or very severe infestations. The seasonality of the crop had a marked influence on the weed flora, mainly due to the relationship of its phenological cycle with well defined climatic periods and to soil management practices. Cluster analysis clearly differentiated typically autumn-spring and spring-summer floras as well as a set of common species.

Notes: Studies were conducted in the field in 2 years comparing the phenology and growth of Orobanche crenata (Forsk) (crenate broomrape) in lentils (Lens esculenta L.) cv. Castellana, peas (Pisum sativum L.) cv. Orix, vetch (Vicia sativa L.) cv. comun and broadbean (Vicia faba L. cv. Alameda. First attachment of O. crenata to these crops took place 9–14 weeks after mid-November planting and earlier after later plantings, Differences in the first O. crenata attachment dates and in the duration of the underground growth period of O. crenata were much greater between years for any given crop than between crops in a given year: both attributes were apparently affected more by seasonal climatic conditions than by crop species, Furthermore, there was no consistent relationship found between crop growth stages and time after first attachment of the parante. This occurred in lentils and vetch while they were vegetative, in peas at late vegetative-early flowering stages, and at widely varying growth stages in broadbean, depending on planting dates and years. The maximum number of O. crenata plants successfully attached to each individual crop plant decreased in the order: peas 〉 broadbean 〉 lentil 〉 vetch, with 21, 14, 10 and 8 per plant, respectively, averaged over the two seasons. Similarly, plant parasites: host dry weight ratio were 1.0, 0.7, 0.3 and 0.2 For each of these crops, respectively.

Notes: The broomrape (Orobanche crenata Forsk) susceptibility to glyphosate applied on faba-bean (Vicia faba L.) as affected by the parasite growth stages at the time of application was studied under field conditions. Glyphosate treatments delayed O. crenata emergence. Single glyphosate application to faba-bean infected with O. crenata predominantly at the stage (a) (small nodule) and (b) (nodules with initial vestigial roots) resulted in a moderate to low control. When the stages (c) (shoot bud already visible) or (d) (shoot and vestigial roots well developed) were the predominant stages, an excellent control was achieved with a single glyphosate application at 60 g ha−1. Increased development stages, with the shoot emerged from the shoot bud, decreased its susceptibility to glyphosate. La lutte contre Orobanche crenata en culture de férerole avec le glyphosate; influence des doses d'herbicide et du stade de développement du parasite

Notes: [14C]-Imazethapyr was applied as a seed treatment and at plant pre-emergence and post-emergence to peas (Pisum sativum L.) parasitized by Orobanche crenata Forsk. Herbicide uptake increased with time regardless of the application method. Uptake reached about 98%, 89%, 81% and 94% of the total herbicide applied for the seed coating, seed soaking, pre-emergence and post-emergence treatments respectively. Herbicide translocation within the host plants consistently differed between O. crenata-infected and non-infected plants. High levels of 14C activity were accumulated by parasitic plants from the host. In non-infected pea plants, pods were stronger sinks for imazethapyr than the other parts of the plant, regardless of the application method. The herbicide distribution in the pea plant: O. crenata complex showed the same pattern regardless of the application methods. However, accumulation of radioactivity in the parasite was lower with pre-emergence and post-emergence application than with the seed treatments. In addition, radioactivity concentration in O. crenata plants was slightly higher when [14C]-imazethapyr was applied to pea seeds by coating than by soaking.

Notes: Field experiments were conducted from 1989 to 1992 to determine the effects of pre-emergence herbicides in sunflower (Helianthus armuus L.) on: (1) the control of Orobanche cemua Loefl. (broomrape) and (2) crop damage and crop yield. Herbicides tested belong to the imidazolinone, sulfonylurea and substituted amide families. Imazethapy r (20–40 g ha−1), imazapy r (12.5–25 gha−1) and chlorsulfuron (4–6 gha−1) controlled broomrape efficiently without crop injury. With good O. cernua control and good crop tolerance, sunflower seed yield from these treatments were generally similar to the non-infested checks and higher than the infestedchecks. Imazaquin (20–40 gha−1), triasulfuron(4gha−1), pdmisulfuron(3g ha−1), acetochlor (4–4 kg ha−1) and metazachlor (2 kg ha−1) were less effective. Imazamethabenz (200–600 g ha−1) and metolachlor (3–3 kg ha−1) were ineffective. Wetconditions aftercropsowing considerably decreased O. cemua control with pre-emergence herbicides probably caused by enhanced degrädation. Des herbicides de prelevee pour la lutte contre Vorobanche (Orobanche cemua Loefl.) dans le toumesol (Helianthus annuus L.) Des expérimentations au champ ont été conduites de 1989 á 1992 pour determiner les effets d'herbicides de pré1evée du toumesol sun (a) la destruction de l'orobanche (Orobanche cernua Loefl.) et (b) la phytotoxcité sur la culture. Les herbicides testés appartenaient aux imidazolinones, aux sulfonylurdes et aux amides substituées. L'imazethapyr (20 á 40 gha−1), l'imazapyr (12,5 á25gha−1)et le chlorsulfuron (4 á 6 gha−1) détruisaient efficacement l'orobanche sans occasionner de phytotoxidt6 sur la culture. Dans ces conditions, les rendements étaient généralement semblables à ceux des témoins non infestés et supérieurs à ceux des témoins infestés. L'imazaquin (20 à 40 g ha−1), le triasulfuron (4 gha−1), le primisulfuron (3 g ha−1), lacétolachlor (4,4 kg ha−1) et le métazachlor (2 kg ha−1) étaient moins efficaces. L'imazaméthabenz (200 á 600 g ha−1) et le métolachlor (3,3 kg ha−1) etaient inefficaces. Des conditions humides aprfes le semis diminuaient considérablement la destruction de O. cemua par les herbicides de prélevée, probablement à cause d'une dégradation plus élevée. Vorauflaufherbigide zur Bekdmpfung der Sommerwurz Orobanche cemua Loefl. in Sonnenblume (Helianthus annuus L.)Zwischen 1989 und 1992 wurden in Sonnenblume Feldversuche zur Wirkung von Vorauflaufherbiziden aus den Gruppen der Imidazolinone, Sulfonylharnstoffe und substitutierten Amide auf die Sommerwurz Orobanche cernua und auf Kulturpflanzenschaden sowie den Ertrag durchgefuhrt. Mit Imazethapyr (20 bis 40 g ha−1), Imazapyr (12,5 bis 25 g ha−1) und Chlorsulfuron (4 bis 6 g ha−1) lieβ sich die Sommerwurz wirksam bekampfen, ohne daβ Schaden an der Sonnenblume auftraten, und die Ertrage waren allgemein ahnlich oder hoher als bei der nichtparasitierten Kontrolle. Imazaquin (20 bis 40 ha−1), Triasulfuron (4 g ha−1).Primisulfuron (3 g ha −1). Acetochlor (4,4 kg ha −1 und Metazachlor (2 kg ha −1) waren weniger wirksam. Imazamethabenz (200 bis 600 g ha−1) und Metolachlor (3,3 kg ha−1) hatten keine Wirkung. Bei Niederschlagen nach der Saat der Sonnenblume war die Bekampfung der Sommerwurz mit Vorauflaufherbiziden vermutlich wegen verstärkten Abbaus erheblich schwächer.

Notes: Orobanche crenata seeds, collected in Syria, Egypt and Spain, were buried in the field in Syria (all three seed lots) and Spain (only Spanish seeds) and at regular intervals exhumed and tested for germination, to investigate whether the seeds exhibit an annual dormancy/non-dor- mancy cycle. When exposed directly to the synthetic germination stimulant GR24 for 7 days at 20°C, seeds only germinated in autumn after the first rains and to a limited extent in winter. When the seeds were conditioned for 11 days at 20°C prior to exposure to GR24, germination occurred during summer and autumn, but seeds were dormant in winter and early spring. The observed seasonal pattern in germinability, in relation to rainfall and soil temperature, was largely consistent with the results of an in vitro experiment by Van Hezewijk et al. (1993), investigating the effect of conditioning temperature and conditioning period on germination capacity and the development of secondary dormancy. Moisture and temperature can therefore be considered the major factors regulating induction and alleviation of dormancy in buried O. crenata seeds. There were no basic differences in response owing to site of collection of O. crenata seeds, nor to the location where they were buried. Variations saisonnières des exigences de germination de graines enfouies d'Orobanche crenata Forsk.Des graines d'Orobanche crenata récoltées en Syrie, en Égypte et en Espagne ont été enfouies au champ en Syrie (les 3 lots) et en Espagne (seules les graines d'Espagne) puis ont été exhumées a intervalles régulier pour que leur aptitude à la germination soil évaluée. Le but était de déterminer si les graines possédaient un cycle annuel dormance/non dormance. Quand elles étaient directement exposées au stimulant de germination synthétique GR24 pendant 7 jours à 20°C, les graines ne germaient qu'à l'automne après les premières pluies et peu en hiver. Quand les graines restaient pendant 11 jours à 20°C avant leur exposition au GR24, la germination seproduisait en été et à l'automne mais les graines restaient dormantes en hiver et au début du prin-temps. Les variations saisonnières d'aptitude à la germination, liées aux précipitations et à la temperature du sol, étaient en accord avec les résultats d'une expérience in vitro de Van Hezewijk et al. (1993) concernant l'effet de la température et de la durée pendant laquelle elle est appliquée, sur l'aptitude à la germination et le développement de la dormance secondaire. L'humidité du sol et sa température peuvent ainsi être considérées comme les principaux facteurs qui induisent et lèvent la dormance de graines de O. crenata enfouies. On n'observait pas de différences importantes dues au lieu de récolte ou à l'endroit oü elles étaient enfouies. Jahreszeitliche Änderungen der Keimung von vergrabenen Samen von Orobanche crenata Forsk.Proben von in Syrien, Ägypten und Spanien gesammelten Orobanche-crenata-Samen wurden in Syrien und Proben nur spanischer Herkunft in Spanien im Freiland im Boden ausgelegt und in regelmäßigen Zeitabständen ausgegraben und auf ihre Keimfähigkeit getestet, um zu untersuchen, ob die Samen einen jährlichen Dormanz-Zyklus haben. Beim direktem Auslegen in dem synthetischen Keimungsmittel GR24 öber 7 d bei 20°C keimten die Samen nur im Herbst nach den ersten Regenfällen und in beschränktem Umfang im Winter. Wenn die Samen för 11 d bei 20°C vor dem Auslegen in GR24 vorbehandelt worden waren, keimten sie im Sommer und Herbst, aber im Winter und fröhen Fröhjahr waren sie dormant. Das jahreszeitliche Verhalten der Keimfähigkeit in Abhängigkeit von Niederschlag und Bodentemperatur stimmte weitgehend mit den Ergebnissen eines In-vitro-Versuches von Van Hezewijk et al. (1993) öber die Wirkung einer Wärmevorbehandlung und Vorbehandlungszeit auf die Keimfähigkeit und die Ausprägung sekundärer Dormanz öberein. Bodenfeuchte und -temperatur können deshalb als die wichtigsten Faktoren för die Induktion und Aufhebung der Dormanz von Orobanche-crenata-Samen im Boden angesehen werden. Herkunft und Versuchsort hatten keinen erheblichen Einfluß auf die Ergebnisse.

Notes: The study was conducted on the weed flora of the main irrigated crops of the Guadalquivir Valley. The TWINSPAN (Two Way Indicator Species Analysis) method was applied for each crop separately to all the species present as well as to their degree of infestation. The results indicated that soil texture may markedly influence the flora, whereas the effects of pH and carbonate content were less important. The climatic differences found in the three areas studied were of little importance. The type of irrigation markedly influenced the flora, although it was always related to edaphic factors, especially texture and slope.In general, the weed flora found in the crops can be divided into two communities. One is characterized by species such as Ridolfia segetum, Picris echioides, Phalaris brachystachys, Phalaris paradoxa and Chrozophora tinctoria and the second by Capsella bursa-pastoris, Cyperus rotundus, Paspalum paspalodes, Digitaria sanguinalis, Sorghum halepense and Portulaca oleracea. The first community comprises weeds in the higher altitudes, further away from the riverbed, with a soil of clayey texture, a high slope and irrigated with sprinklers. The second community comprises the flora of flat areas, with sandy soils, located in the middle and low river terraces and furrow irrigated.

Notes: A floristic study of the weeds in irrigated crops in the Guadalquivir River Valley (southern Spain) revealed a total of 284 species in the 267 fields surveyed, of which about 100 can be considered highly harmful. The families with the highest numbers of species were Compositae, Gramineae, Leguminosae, Cruciferae and Umbelliferae. Although 77·8% of the species were therophytes, geophytes were important in spring-sown crops. Most species were of Mediterranean origin, especially in autumn-sown crops; Cosmopolitan and Neotropical species were more important in the spring-sown crops. At the early and late crop growth stages 7% and 10%, respectively, of the fields had severe or very severe infestations.The relationships between the main weed species and crops were studied by cluster analysis, using the corrected frequency of each species in each crop as the reference parameter. This allowed classification of the species into subgroups and hierarchical groups according to their agro-ecological preferences. A close relationship was found between a given crop, its cycle and the usual soil management practices and the phenology of certain species, reflecting the influence of these factors in restricting the germination period. The analysis differentiated the typically winter, summer and intermediate weed floras.

Notes: The influence of broomrape (Orobanche crenata Forsk) infection levels on commercial broad bean (Vicia faba L.) crops was investigated and an equation relating yield losses to O. crenata density was derived. O. crenata growth was weakly and negatively related to the final height or shoot number of the broad bean plant. O. crenata decreased seed yield mainly by reducing the number of pods. The number of seeds per pod was also decreased by O. crenata infection when competition occurred at late-crop growth stages when most of the pods had already developed. Bean unit seed weight was only weakly affected by infection. Correlation coefficients between O. crenata number and broad-bean seed yield per plant were significant at P=0·05 and greater than 0·70 in most locations. On average, about four O. crenata per broad-bean plant decreased seed yield by half. An estimation of the percentage of losses in any field can be calculated from the equation:%loss = 100 × 0·124 × OcN.where OcN is the average final number of emerged O. crenata per plant.

Notes: Studies on competition between Ridolfia segetum Maris, and sunflower (Helianthemum annuus L.) were conducted at eight locations in southern Spain in 1990 and 1991. in order to define competition models and to estimate from these economic thresholds as affected by crop inputs and potential yields. Competition losses in sunflower crops ranged from 19% to 56% of weed–free yields. There were slightly better correlations between percentage sunflower reduction and weed density than with weed dry weight, (−0.66 and −0.59, respectively). The weed competitive index, or sunflower crop dry weight reduction per unit dry weight of R. segetum, was 1.09. The percentage yield losses due to weed density (NPRt) were fitted to multiple linear, quadratic, exponential and hyperbolic models. The hyperbolic equation, %RSY=100 (1+1/b*NPRt)−1, where b=0.14 and is the R. segetum competitive ability index, had the lowest error sum of squares (SSE), and gave the best biological explanation for the competition response. Early emergence (before mid–March) made weeds about 1.5 times more competitive than late emergence. The economic threshold to offset the cost of a shallow post–emergence tillage, assuming 70% control efficiency, ranged from about 2.5 plants m −2 for low–yielding crops(1200kgha−1) to less than one plant m−2 for higher–yielding crops (2800 kg ha−1).