ALBERT

Description: Accurate determination of stand establishment ages is important in developing growth and yield models and in studying stand dynamics of fire-origin stands. The study objective was to determine time to reach breast height for black spruce (Picea mariana (Mill.) BSP), jack pine (Pinus banksiana Lamb.), trembling aspen (Populus tremuloides Michx.), and white birch (Betula papyrifera Marsh.) from fire origin stands under different site conditions in northeastern Ontario. Stands were randomly selected from burns with known fire dates. In each stand, three to six dominant and codominant trees of a selected species were cored at breast height (1.3 m above the ground level) to determine time to reach breast height. Trembling aspen and white birch did not differ for time to reach breast height after fire, taking 6 or 7 years, jack pine took marginally longer (8 years), whereas black spruce took the longest (18 years). While time to reach breast height did not vary among site conditions as described by soil texture and moisture regime, it was positively related to time since fire. The results of this study indicate that stand establishment dates and total tree ages can be substantially underestimated if breast height age is used as the stand age, resulting in misinterpretations of growth and yield and forest succession.

Description: To evaluate the variation in trembling aspen (Populus tremuloides Michx.) productivity at a large geographic scale, we examined the relationships between site index and environmental factors from 142 even-aged, fully stocked stands located on a variety of sites across interior British Columbia. Site index was derived from stem analysis and the environmental measures included climate surrogates (latitude, longitude, and elevation), biogeoclimatic zone, slope aspect, actual soil moisture regime (SMR), and soil nutrient regime (SNR). The spatial gradients (latitude, longitude, and elevation), slopeaspect, SMR, and SNR affected aspen site index, but their relationships greatly varied with biogeoclimatic zone. At the provincial scale, these relationships were weaker than on the zonal scale. Among the models developed for predicting aspen site index, we recommend the zone-specific all-factor model for application, which explained 82% of the variation of site index and provided unbiased and precise predictions.

Description: To examine the relationships between trembling aspen (Populus tremuloides Michx.) productivity, environmental attributes, and foliar nutrients and to make accurate predictions of trembling aspen productivity, we sampled 60 naturally established, fire-originated, and even-aged trembling aspen stands in northern British Columbia. Trembling aspen site index significantly varied with latitude, elevation, aspect, slope position, edatopes, some forest floor and mineral soil physical and chemical properties, and concentrations of some foliar nutrients. To predict site index, we developed multiple linear regression models using climatic variables, topographic properties, edatopes, soil physical and chemical properties, or foliar nutrients as predictors. Model accountability for variation of site index differed in decreasing order from soil model, climatic model, forest floor model, foliar nutrient model, edatope model, topographic model, to mineral soil model. Examined by the test data set, all models were unbiased, but they had different levels of precision in prediction in decreasing order from edatope model, soil model, forest floor model, mineral soil model, foliar nutrient model, climatic model, to topographic model. The soil and foliar nutrients models may provide insight into ecosystem processes, but the models using climatic variables and topographic properties or edatopes as predictors are recommended for predicting trembling aspen site index.

Description: Canopy gaps play an important role in forest vegetation dynamics when fire return intervals are long. However, there is little known about the role of gaps in the development of forest stands that initially dominate following stand-replacing disturbance. We investigated gap disturbance during the breakup of trembling aspen (Populus tremuloides Michx.) stands at two scales: at the stand level we quantified gap fraction and gap size as stand development proceeds; at the gap level we determined causes of gap-maker mortality and evaluated resulting gap-maker structure and decay as stand development proceeds. We also evaluated the impact that gaps have on stand transition by quantifying the abundance and growth of juvenile trees in gaps of different sizes and ages. Ten stands between 60 and 120 years since fire in northeastern Ontario were sampled using line intersect transects. Gap fraction doubled (∼18%36%) and mean gap size was more than four times greater (∼45200 m2) over the time period. Standing dead gap makers in early states of decay were most frequent in young stands, whereas snapped gap makers in various states of decay were most frequent in old stands. Infection by fungal pathogens was the most frequent cause of mortality (56%) and was not related to time since fire. Balsam fir (Abies balsamea (L.) Mill.) was the most abundant juvenile recruit. However, transition probabilities for tree species were independent of gap type. These results indicate that gap creation is frequent during early stages of stand development in boreal forests; however, it is unlikely that successional trajectory is affected by their formation.

Description: Mixed-species stands are hypothesized to be more structurally diverse because of inherent differences in growth rates between tree species. We investigated three combinations of even-aged, single- and mixed-species coniferous stands in western Canada: (i) shade-tolerant western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western redcedar (Thuja plicata (Donn ex D. Don) Spach), (ii) shade-intolerant lodgepole pine (Pinus contorta Dougl. ex Loud.) and western larch (Larix occidentalis Nutt.), and (iii) shade-intolerant lodgepole pine and shade-tolerant black spruce (Picea mariana (Mill.) BSP). We quantified stand structural diversity based on tree diameter and height by using Shannon's index and the coefficient of variation. Regression analysis was applied to examine the relationships between structural diversity, species composition, and stand age. Mixed-species stands of western hemlock and western redcedar had similar structural diversity to their respective single-species stands. Pinelarch and pinespruce mixed-species stands had higher tree-size diversity than their respective single-species stands. Stand age was a significant factor for tree-size diversity, but its effect changed in different ways depending on the type of mixtures and index used to measure diversity.