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  • 1
    Electronic Resource
    Electronic Resource
    Oxford, UK : Blackwell Publishing Ltd
    Journal of fish biology 29 (1986), S. 0 
    ISSN: 1095-8649
    Source: Blackwell Publishing Journal Backfiles 1879-2005
    Topics: Biology
    Notes: Shoals composed of equal numbers of two size-classes of European minnows were observed undisturbed, feeding and after threat from a pike in a large arena tank.The time/frequency budget and analysed sequences of behaviour of the two size-classes were very similar. Irrespective of size, for standard behaviour measures, fish in the shoal behaved similarly under the same external influences, including predator threat.In contrast, however, the distribution of the two size-classes provided evidence of size segregation within the shoal. This was brought about by individual minnows making shoaling responses preferentially to their own size-class. After exposure to the predator, shoaling responses changed and differed between small and large minnows.The outcomes of contests at foraging patches were governed primarily by fish size and information asymmetry rather than by occupation of a feeding site.The experiment shows that asymmetrical pay-offs in foraging and in response to predator threat are the probable reasons for size-segregation behaviours. This conclusion supports the views of earlier workers that mechanical sorting by swimming speed is not an important factor in size segregation in shoals.
    Type of Medium: Electronic Resource
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  • 2
    Electronic Resource
    Electronic Resource
    Oxford, UK : Blackwell Publishing Ltd
    Journal of fish biology 29 (1986), S. 0 
    ISSN: 1095-8649
    Source: Blackwell Publishing Journal Backfiles 1879-2005
    Topics: Biology
    Notes: Adult dace, minnows and gudgeon (50 of each) were studied in a fluvarium. Each possible paired-and three-species combination was compared with single-species controls. Species position in the fluvarium and the behaviour of marked individuals were used to measure interactions. Individual behaviours were standardized for day-to-day changes in activity level. The presence of a second species caused significant increases in the frequency and total time of behaviours associated with positional change. Such behavioural changes may reduce competition by adjusting individual position within the shoal. Minnows moved higher in the water and downstream in the presence of dace. Gudgeon moved lower in the water and downstream in the presence of dace and of minnows. Dace made slight, but significant, movements downstream and higher in the water column in response to both minnows and gudgeon. These shifts were very distinct, and the resulting shoal structure mirrors the positions occupied in mixed-species shoals of these species in the wild.
    Type of Medium: Electronic Resource
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