ALBERT

Description: Graptoloid evolutionary dynamics show a marked contrast from the Ordovician to the Silurian. Subdued extinction and origination rates during the Ordovician give way, during the late Katian, to rates that were highly volatile and of higher mean value through the Silurian, reflecting the significantly shorter lifespan of Silurian species. These patterns are revealed in high-resolution rate curves derived from the CONOP (constrained optimization) scaled and calibrated global composite sequence of 2094 graptoloid species. The end-Ordovician mass depletion was driven primarily by an elevated extinction rate which lasted for c . 1.2 Ma with two main spikes during the Hirnantian. The early Silurian recovery, although initiated by a peak in origination rate, was maintained by a complex interplay of origination and extinction rates, with both rates rising and falling sharply. The global 13 C curve echoes the graptoloid evolutionary rates pattern; the prominent and well-known positive isotope excursions during the Late Ordovician and Silurian lie on or close to times of sharp decline in graptoloid species richness, commonly associated with extinction rate spikes. The graptoloid and isotope data point to a relatively steady marine environment in the Ordovician with mainly background extinction rates, changing during the Katian to a more volatile climatic regime that prevailed through the Silurian, with several sharp extinction episodes triggered by environmental crises. The correlation of graptoloid species diversity with isotopic ratios was positive in the Ordovician and negative in the Silurian, suggesting different causal linkages. Throughout the history of the graptoloid clade all major depletions in species richness except for one were caused by elevated extinction rate rather than decreased origination rate.

Description: Fourteen early Ordovician (Tremadoc-Llandeilo) graptolite sequences from around the world are precisely (infrazonally) correlated, based on the stratigraphic ranges of 130 species and species groups. The composite standard sequence (CSS) of graptolites has been determined from the six best regional sequences by a nonparametric graphic correlation. Two data sets were selected: one comprised first appearance events of 103 taxa, the other, first and last appearance events of 45 taxa. The results of the two runs accord well and reveal respectively 66 and 73 successive bioevents in early Ordovician time. Event spacing averages 0.7–0.8 Ma and enables fine subdivision, correlation and homotaxial testing for diachroneity. The strong correlation between each of the six regional sequences and the CSS indicates the high level of accordance among graptolite successions around the world.