ALBERT

Description: Statolith microstructure was studied in 162 specimens of Illex coindetii (mantle length ranging from 48 to 300 mm) captured on the shelves of Sierra Leone and Western Sahara between May and November 1987. Growth increments were revealed in all statoliths studied, but they were not well-resolved as in other congeneric species Illex illecebrosus and Illex argentinus . Age and growth rates were estimated assuming that growth increments within statoliths were produced. In both regions, differences in growth rates between sexes (females become larger than males) occurred first in length and then in weight. Geographical differences in sex-specific growth (Western Sahara squid grow faster and attain larger sizes than Sierra Leone squid) was apparent after 150 d in females and 120 d in males. Illex coindetii in Sierra Leone started maturing at smaller sizes but at approximately the same age as the Western Sahara squid. Small sized early maturing groups of I. coindetii with a life span of ~0.5 y occurred in both regions. However, large sized late maturing squid with a life span of ~1 y were encountered only in the Western Sahara. In waters of the west African shelf, I. coindetii spawn throughout the year.

Description: Stomach contents of Todarodes sagittatus caught by trawlers working from 100 to 800 m depth in the Balearic Sea (western Mediterranean) were studied. From the 348 stomachs examined (153 males and 195 females) 33.62% were empty (39.21% in males and 29.74% in females). The diet of the squid was composed of 58 different prey items belonging to four major groups: Osteichthya, Crustacea, Cephalopoda and Chondrichthya. Osteichthyes, crustaceans and cephalopods were the most common prey, with a frequency of occurrence value of 84.85, 48.92 and 29.87% respectively. A change in the diet as the squid grows was observed, since juveniles feed basically on fishes while adults prey more actively on crustaceans. Analysis of the diet by size-classes reflected an ontogenetic migration to deeper waters since, parallel to the increase of size, a raise in the percentage of prey species inhabiting deeper waters was detected. Cannibalism was quite frequent, since T. sagittatus was the second most common cephalopod prey. Females had higher fullness-weight index and lower emptiness index than males, which reflects their major energetic demand for egg production.

Description: The size of fish and squid prey of Loligo forbesi was investigated using otoliths, beaks and statoliths collected from stomach contents analysis of samples obtained from Scottish and Irish waters between 1990 and 1993. Loligo forbesi was found to consume a large range of prey sizes, but prey size was always less than the predator size. Season was shown to significantly influence the predator size-prey size relationship for sprat and sandeel prey, but this itself could be influenced by seasonal changes in the size of prey. Fish prey size increased with increased predator size up to a mantle length (ML) of 200 mm. Loligo forbesi of mantle length 〉200 mm consumed a range of prey sizes, with no clear increase in the size of prey. For most prey taxa the relationship between prey size and squid size was similar, the exceptions being dragonets and silvery pout. Cannibalism by L. forbesi was mostly limited to larger L. forbesi (〉150 mm ML) feeding on smaller (20–50 mm ML) conspecifics.

Description: The population structure of the European flying squid Todarodes sagittatus was studied using data of about 5000 squid caught in waters off the western Sahara between 1969 and 1997. The bulk of the population consisted of winter-spawned squid, which occurred as juveniles of 80–120 mm mantle length (ML) over the slope in spring, and as immature squid of 160–180 mm ML both on the shelf and slope in summer. In autumn, the squid attained 220–280 mm ML, matured, and shifted to the slope, where the spawning was supposed to occur in winter. Age and growth of T. sagittatus was studied using statolith ageing techniques. Assuming daily production of putative growth increments within statoliths, as well as sizes and proportions of immature and maturing females, the lifespan of the west Saharan populations of T. sagittatus lasts ∼1 y. Todarodes sagittatus is a fast growing squid at juvenile and immature ontogenetic phases. Early maturation (at ages 220–230 d in males and 250–260 d in females, respectively) and subsequent decrease of somatic growth rates caused rather small modal sizes of mature squid (250–300 mm ML) compared with those of their northern counterparts (350–420 mm ML). Both hatching dates and seasonality in occurrence of mature females shows that in waters off the western Sahara T. sagittatus spawns throughout the year with a well-pronounced winter peak.

Description: Size of ripe eggs and potential fecundity are described in the squid of the subfamilies Todaropsinae and Todarodinae (Oegopsida: Ommastrephidae)— Todaropsis eblanae from West Africa, Todarodes angolensis from Namibia, Todarodes sagittatus from north-west Africa and the Mediterranean Sea, Todarodes sp. from the south-east Pacific, Nototodarus hawaiiensis from the south-east Pacific and West Indian Ocean and Martialia hyadesi from the south-west Atlantic. Females of both subfamilies are characterized by a wide range of ripe egg size (0.7–2.4 mm) and low and medium values of potential fecundity (20,000–2,500,000).

Description: Records of unusual cephalopods, taken as by-catch in Irish and Scottish waters in the years 1985–1995, are presented. Of most interest are three specimens of giant squid (Architeuthis) that were caught in bottom trawls off the west of Ireland between April and June 1995, all were mature males of mantle length ∼1000 mm. Other records include a large mature female Histioteuthis bonnellii from the west of Ireland and three specimens of the gelatinous incirrate octopus, Haliphron atlanticus.

Description: Octopuses ( Eledone cirrhosa (Octopoda: Cephalopoda)) held in an aquarium were subjected to varying conditions of feeding and starvation to evaluate putative indices of feeding and growth. Specific growth rate (%d −1 ) was linearly related to feeding rate (% of the body mass d −1 ) in animals with a mean body mass of 250 g at 15°C. Maximum growth rates varied between 〉 2% d −1 (body weights 〈 300 g) to 〈 1% d −1 (body weights ≤ 900 g) but specific growth rates were not related to body weight. Growth rates became negative (weight loss) after one week without food. The digestive gland index (DGI) was significantly correlated with short and long-term feeding and specific growth rates, and with body weight. Muscle RNA concentration was linearly correlated with growth rate during the previous 1–3 weeks but not with feeding rate. RNA:protein ratios were not different between mid-arm and mantle sample sites but arm tip values were significantly higher. RNA:protein ratio was related to body weight only in feeding animals. It is concluded that DGI is an index of feeding rate and that RNA:protein ratio can be used as an index of recent (~ 4 weeks) growth rate.

Description: A specimen of Chaunoteuthis mollis (Cephalopoda: Onychoteuthidae), a mature mated female, from the Straits of Messina was examined. Measurements and indices were compared with those of Onychoteuthis banksii reported by previous authors. The differences found were attributed to the degeneration of the C. mollis specimen, due to sexual maturation. Two subocular (one under each eye) and two visceral photophores, similar to those of O. banksii , were present in the C. mollis specimen. The size of the posterior visceral photophore of C. mollis fit the regression line correlating posterior photophore diameter to mantle length in O. banksii. These results corroborate the hypothesis, put forward by other authors, that C. mollis represents the mature female of O. banksii.

Description: Three specimens of the giant squid Architeuthis were by-caught in demersal trawls to the west of Ireland, between April and June 1995. All three specimens were mature males, of mantle length 975–1084 mm. Although some intraspecific variation in fin, beak and gill were noticed, all three specimens were tentatively assigned to the species A. dux. The three specimens had food remains in their stomachs and food items identified included Micromesistius poutassou, Trachurus trachurus, Nephrops norvegicus and Eledone cirrhosa . Age estimates were made by counting putative daily growth increments in ground statoliths, and ranged from 294–122 d, giving percentage daily growth rates of 2·96–4·25% indicating a short life cycle and extremely rapid growth.

Description: Deep-water trawling for fish species on the shelf slope off Scotland results in a by-catch of rare cephalopod species. The cirrate octopus Opisthoteuthis grimaldii is normally present below 800 m; at least one other species of cirrate and five species of incirrate octopus are found; significant extensions of range and depth are recorded. An incidental catch of the ommastrephid squid, Todarodes sagittatus , included a fully mature specimen and is indicative of its probable breeding locations. The results highlight the many taxonomic and systematic uncertainties surrounding the rarer cephalopod forms, and emphasize the utility of the fishing by-catch.

Description: Two large (dorsal mantle length 42.5 and 47.5 cm), mated spent females of circum‐Antarctic bathypelagic cranchiid squid Galiteuthis glacialis were caught early in March 1992 at the surface of the ice hole in the western Weddell Sea over depths 1915‐1920 m by the team of the U.S.A.‐Russian Ice Station Weddell‐I. The structure of the reproductive system of adult females is described for the first time in detail. Both were gelatinous, devoid of tentacles, with empty or almost empty stomachs. The empty spermatangia (sperm reservoirs of spermatophores) 30‐35 mm in length were distributed in the mantle tissues parallel to the mantle surface and to each other in the dorso‐anterior part of the mantle: 13 in one female, parallel to the body axis, and 20 in the other, parallel (13) or perpendicular (7) to the body axis. In the latter case, they represented probably two mating events. The spermatangia lay nearer to the inner than the outer mantle side and opened by a round window on the inner side; the skin with chromatophores above them remained intact. The spermatozoa had one flagellum and rod‐like heads, length 5.0‐5.3 μm, width 1.2‐1.5 μm. The most characteristic features are: a very simple type of blood vessel branching making each micro‐gonad currant‐like, not grape‐like; a very compact disposition of oviducal, nidamental glands and gill, forming a united complex located on both sides of the mantle cavity; and an ovary connected by mesentery along all its length with the continuation of the stomach from the caecum to the end of the gastrogenital ligament. Only immature degenerating trophoplasmatic oocytes, length 0.9‐1.4, av. 1.0‐1.2 mm, were contained in ovaries; only one mature egg (length 3.3 mm, width 2.4‐2.5 mm) was found in each female. The absence of oocytes 〈0.9 mm and 1.5‐3.2 mm indicates that the maturation of oocytes proceeds rather synchronously, one large portion of eggs (some tens of thousands) matures in a short time while others degenerate. The residual fecundity is assessed to be approximately 20,000 eggs. It is hypothesized that mating occurs shortly before spawning and that mature males do not undergo gelatinous degeneration and do not lose tentacles. Spermatophores are placed on the inner side of the female's mantle with the aid of the male's tentacles and/or arms (less probably by the penis), but the exact mode of implantation is unclear. Spawning probably occurs at depths of adult habitat (approx. 500–2500m), may be multiportional but short; the exhausted female loses neutral buoyancy, rises to the surface and dies. Rising to the surface after spawning is a common feature of females of many meso‐ and bathypelagic squids undergoing gelatinous degeneration during maturation (Onychoteuthidae, Gonatidae, Histio‐teuthidae, Cranchiidae, etc.) and may explain the common occurrence of large deep‐water squids in the stomachs of seabirds, including those incapable of diving, and marine mammals.

Description: Growth in Loliolus noctiluca (Myopsida: Loliginidae) in Western Port, Victoria, Australia was studied from statolith growth increments. Tetracycline staining experiments verified previous work on tropical forms of this species that showed growth increments to be deposited daily. A logistic growth function described the relationship between length and increment number. There appear to be major differences in the form of growth, longevity and life history pattern between tropical and temperate forms of this species. These are probably attributable to differences in environmental conditions.

Description: Cephalopod prey of several cetacean species from Hawaiian waters were identified and quantified from the beaks in stomachs of stranded individuals. The different species of cetaceans all appear to target different species and sizes of cephalopods. Beaks from two sperm whales ( Physeter catodon ) included a total of 312 upper and 292 lower beaks (mandibles) of cephalopods. All of the cephalopods represented by lower beaks were oceanic squid belonging to 20 or more species in 14 families. The major constituents of the whale diet were Histioteuthis hoylei (45% by number and 10·9% by dry weight), Ommastrephes bartrami (7·6% by number, 30·6% by weight) and Architeuthis sp. (only 0·7% by number, but 26·5% by weight). In a Risso's dolphin ( Grampus griseus ) stomach, 1051 upper beaks and 1349 lower beaks were present. Eighteen or more genera of cephalopods in 15 families were present. The major constituents of the Grampus diet were Enoploteuthis spp. (36·1% by number and 30·1% by dry weight) and Abraliopsis spp. (23·6% by number, but only 4·9% by weight) and O. bartrami (1·3% and 32·1% respectively). The melon-headed whale ( Peponocephala electra ) eats fish as well as cephalopods which are represented by only six lower beaks belonging to five genera, Enoploteuthis, Teuthowenia, Abraliopsis, Abralia sp. and Bathyteuthis abyssicola . The beaked whale (unidentified) contained two lower beaks which were O. bartrami. Stenella attenuata also eats fish and the sample contained lower beaks, one of Enoploteuthis sp. and seven of an Abraliopsis sp.

Description: Observations on samples from Spanish trawlers between September and November of 1995 revealed the presence of mated females of Loligo gahi (Cephalopoda: Loliginidae) from 164–285 m depth, in the Western area of the Falkland Islands Conservation Zone. 93.8% of the mature females, and 31.0% of the maturing ones, were mated. Deposition of spermatophores always took place in the oral membrane between the connectives of arms IV. The relationships between sexual maturation and copulation have been analysed, and the hypothesis of mating acting as a ‘trigger’ of the final sexual maturation in Loligo gahi females is proposed and discussed.

Description: This paper gives morphometric variations and dorsal mantle length–total weight (DML-TW) relationships for Illex coindetii in the eastern Central Atlantic area. Positive allometry was observed in males and negative in females of the species. The most variable body measurements between males and females were width and perimeter of the head. In the study area, divergence of morphometric measurement starts at 95 mm. The point of divergence, however, varies with latitude; fluctuating from 104 mm in the north (Morocco and Sahara) to 76 mm in the central area (Mauritania and north of Senegal) and 73 mm in the south (Gulf of Guinea). The relationships between DML and TW showed that sexual dimorphism due to differential growth between males and females starts to occur at 56 mm ML. The starting point of sexual dimorphism (56 mm) varies according to the zone; the higher the latitude, the later it occurs. Thus, sexual dimorphism occurs at 49 mm in the south (Gulf of Guinea), at 54 mm in the central area (Mauritania and north of Senegal) and at 74 mm in the north (Morocco and Sahara). Females grow larger than males, but males were heavier at any given length. As latitude decreased, a slow down in the increase in weight-at-length was observed in both sexes.

Description: A total of 57 comparative hauls using a rectangular midwater trawl with a fishing mouth area of 50 m 2 (RMT 50) were carried out along the sides of an imaginary triangle south of Madeira in 1986. A total of 1258 cephalopods were caught, giving a mean of 22 per haul with a range from 0 to 67. The nets were used with a diver's light on the top bar which was either switched off or was operated with a 20, 70 or 150 W bulb, powered by a car battery. A significantly greater number of individuals per haul was caught with lights on than without lights, increasing from a mean of 13·5–25·1, a factor of 1·8. Similarly, the number of species caught was increased from a mean of 7 to 10·4, a factor of 1·5 and the volume of cephalopods was increased from a mean of 41·1–162·3ml, a factor of 3·9. Similar comparisons made for catches during day or night separately and on the three courses separately also showed marked increases with the lights. Samples show that increase in power of the lights increased the total number of cephalopod individuals caught. In the 12 species with more than ten individuals, in 33 of the 36 comparisons (of number of individuals, species and volumes) there is an increase with the light. The most influenced species was Taonius pavo which increased in numbers by a mean factor of 3·9 times with 20W, 4·0 times with 70W and 6·1 times with 150W when compared with the numbers caught with no light.

Description: Recent attention to members of the sepiolid squid genus Euprymna and symbiotic associations with luminescent bacteria ( Vibrio fischeri strains) has prompted a review of this poorly-resolved group of squids. Twelve nominal species have been placed in this genus of which the majority are ill-defined, known only from their original descriptions and separated on the basis of inadequate characters. As a first step in resolving this group, a temperate Australian species, the Southern dumpling squid, Euprymna tasmanica , is here redescribed in detail. As the genus Euprymna currently stands, most members are only distinguished on the number and position of enlarged suckers in mature males. No diagnostic characters are available to identify females. All nominal species placed in this genus are reviewed and a key to proposed valid species is presented. Six species are considered here to be valid: Euprymna berryi, E. hoylei, E. morsei, E. scolopes, E. tasmanica and an undescribed species treated here as Euprymna sp. 1. Euprymna similis is a synonym of E. morsei of Japan. Due to inadequate original descriptions, and lost or poor type material, two species are considered here to be nomen dubia ( E. schneehageni and E. pusilla ), while the taxonomic status of four additional species remain unresolved ( E. albatrossae, E. bursa, E. phenax and E. stenodactyla ).

Description: The statolith microstructure was studied in 142 females (mantle length, ML, ranging from 77–402 mm) and 119 males (72–328 mm ML) of Martialia hyadesi caught on the Patagonian and Falkland shelves and at the Antarctic Polar Frontal Zone between 1989–94. The statolith microstructure dark zone in this species, contains narrower and more numerous growth increments than the dark zones of other ommastrephid squids. Assuming daily production of putative growth increments within statoliths males live up to 12 months, and females live up to 13 months. It is likely that the life cycle lasts c. 1 yr, but immature squids with ages 〉330–340 d suggest that a part of M. hyadesi populations could have life span 〉1 yr. Growth in length was best described by the Gompertz function, whereas growth in weight was best described by the logistic function. M. hyadesi is characterized by slow juvenile growth (〈100 mm ML), fast growth of immature squids and a sharp decrease in growth rates during maturation. M. hyadesi mature later (at ages 〉270 d) than other temperate ommastrephids, but maturation is rather rapid (2–3 months). In the south-west Atlantic, M. hyadesi hatch throughout the year.

Description: The current status of Antarctic Odontocetes – sperm whales Physeter catodon , killer whales Orcinus orca , long-finned pilot whales Globicephala melaena , hourglass dolphins Lagenorhynchus cruciger and poorly known species of beaked whales (family Ziphiidae)–were studied in Anatarctic waters using data gathered in sighting surveys conducted from 1976/77 to 1987/88. Temporal variation in density demonstrated the different migration patterns by species, especially between sperm whale and killer whale. Spatial distributions during mid-summer demonstrated different peaks of occurrence for each species by latitude that suggest possible segregation between the species. Killer whales occur mainly in the very southernmost areas, sperm whales in the southern half of the study area, beaked whales (mostly southern bottlenose whales Hyperoodon planifrons ) ranged over a wide area, and long-finned pilot whales and hourglass dolphins were mainly in the northern regions of Antarctic waters. Several longitudinal peaks of occurrence and apparent distribution gaps were identified for sperm, beaked and killer whales. Abundance estimates for south of the Antarctic Convergence in January are based on line transect theory and were 28 100 animals (coefficient of variation CV 0.18) sperm whales, 599 300 (0.15) beaked whales (mostly southern bottlenose whales), 80 400 (0.15) killer whales, 200 000 (0.35) long-finned pilot whales, and 144 300 (0.17) hourglass dolphins. Based on this, biomass of these species were estimated as 0.77 (sperm whales), 2.70 (beaked whales), 0.32 (killer whales), 0.16 (long-finned pilot whales) and 0.01 (hourglass dolphins) million tonnes. Consumption of food (mostly squid) by the Odontocetes is estimated as 14.4 million tonnes with 67% of the total consumed by beaked whales. Indirect consumption of Antarctic krill through the predation of squid by beaked whales is estimated to be c . 24 million tonnes. This value is similar to the estimate of krill consumption by penguins in the Antarctic (33 million tonnes). Odontocetes, especially southern bottlenose whales, are suggested to have a much greater role in the Antarctic ecosystem than has previously been considered.

Description: Subantarctic Marion Island is one of the few localities where the congeneric albatrosses Phoebetria fusca and P. palpebrata breed sympatrically. Chicks of both species at Marion Island were induced to regurgitate their stomach contents after being fed. Liquid formed over half the diet by mass. Cephalopods occurred most frequently in both species' diets. In terms of mass, cephalopods formed the larger part of the diet of sooty albatrosses, whereas fish was more important to light-mantled sooty albatrosses. Crustaceans and birds were also recorded for both species. Squid of the families Onychoteuthidae, Histioteuthidae, Chiroteuthidae and Cranchiidae occurred most abundantly in both species. Most squid taken by both albatrosses were of species known to float after death, suggesting that scavenging plays an important role in the species' foraging behaviour. Light-mantled sooty albatrosses consumed more squid restricted to the south of the Antarctic Polar Front than did sooty albatrosses, supporting a trend to latitudinal segregation of the two species while foraging.

Description: Changes in the molecular structure of a highly ordered kaolinite, intercalated with urea and potassium acetate, have been studied using Raman microscopy. A new Raman band, attributed to the inner surface hydroxyl groups strongly hydrogen bound to the acetate, is observed at 3605 cm (super -1) for the potassium acetate intercalate with the consequential loss of intensity in the bands at 3652, 3670, 3684 and 3693 cm (super -1) . Remarkable changes in intensity of the Raman spectral bands of the low-frequency region of the kaolinite occurred upon intercalation. In particular, the 144 and 935 cm (super -1) bands increased by an order of magnitude and were found to be polarized. These spectroscopic changes provide evidence for the inner surface hydroxyl group-acetate bond being at an angle approaching 90 degrees to the 001 face. Decreases in intensity of the bands at 243, 271 and 336 cm (super -1) were observed. The urea intercalate shows additional Raman bands at 3387, 3408 and 3500 cm-1 which are attributed to N-H vibrations after formation of the urea-kaolinite complex. Changes in the spectra of the inserting molecules were also observed.

Description: Squid (Lolliguncula brevis) were exercised in a tunnel respirometer during a stepwise increase in water velocity in order to evaluate the anaerobic treshold, i.e. the critical swimming speed above which anaerobic metabolism contributes to energy production. The average anaerobic treshold was found at speeds of 1.5-2 mantle lenghts s-1. Above this velocity, α-glycerophosphate, succinate and levels fell and phospho-L-arginine was progressively depleted, while the levels of glucose 6-phosphate and inorganic phosphate rose. The finding of a simultaneous onset of anaerobic metabolism in the cytosol and the mitochondria indicates that a limited oxygen supply to the mitochondria elicits anaerobic energy production. This finding is opposite to the situation found in many other vertebrate and invertebrate species, in which energy covered by anaerobic energy production. This finding is opposite to the situation found in many other vertebrate and invertebrate species, in which energy requirements in exvess of aerobic energy production are covered by anaerobic metabolism, with mitochondira remaining aerobic. In L. brevis, swimming at higher speeds is associated with a small factorial increase in metabolic rate based on a high resting rate of oxygen cnsumption. Pressure recordings in the mantle cavity support this finding, indicating a high basal level of spontaneous activity at rest and a small rise in mean pressure at higher swimming velocity. Bursts of higher pressures from the jet support elevated swiming speeds and may explain the early transition to anaerobic energy production which occurs when pressure rises above 0.22-0.25kPa. The finding f mitochondrial hypoxia at a low critical speed in these squid is interpreted to be related to their life in shallow coastal and bay waters, which limits the necessity to maintain high swimming velocities. At increased swimming velocities, the animals oscilliate between periods of high and lo muscular activity. This behaviour is interpreted to reduce transport cost and to permit a longer-term net use of anaerobic resources when speed exceeds the critical value or when the squid dive into toxic waters. The simultaneous onset of anaerobic metabolism in the cytosol and the mitochondria emphasizes that squid generally make maximal use of available requirements are the highest among marine invertebrates.

Description: A mineralogical and 4OAr/39Ar study of 13 amphibole samples in the Kamila Amphibolite Belt and Kamila Shear Zone in northern Pakistan has found a correlation between the degree of greenschist facies alteration and quantity of excess 40Ar. Additionally, there is a north–south divide with amphibole samples from the northern region showing larger degrees of gree schist facies alteration, brittle deformation, and excess 40Ar incorporation compared to the predominantly plastically deformed, less altered, amphibole samples from the Kamila Shear Zone in the south. Acid leaching of two amphiboles from the Kamila Amphibolite Belt indicates that a large proportion of the excess 40Ar is correlated with later greenschist facies alteration hases, and can be easily removed by acid etching, thus revealing acceptable regional 40Ar/39Ar plateau ages.

Description: The results of an examination of one quarry within the gneisses of the Bartica Assemblage are described and the validity of these results throughout the whole of the Bartica Assemblage is discussed. The foliation and banding of the gneisses at Kereti Quarry are mainly the results of tectonism; concomitant metamorphism within the Amphibolite Facies was succeeded by a two-phase metasomatism involving successive additions of Na and K. These results appear valid throughout the Bartica Assemblage and it is further suggested that the main rock type distribution in the Bartica Assemblage is also controlled by tectonism.

Description: Textural differences in the occurrence of microcline define augen gneisses, subhedral porphyroblastic gneisses and euhedral porphyroblastic gneisses within the Bartica Assemblage, British Guiana. The introduction of microcline is metasomatic and the nature of the porphyroblasts appears to be a reflection of tectonic control during their development.

Description: In this paper we examine the steady, two-dimensional convective motion which occurs in a horizontal circular cylinder whose wall is non-uniformly heated. One observes several qualitatively different physical phenomena depending on the wall temperature distribution and the value of the Rayleigh number. The low-Rayleigh-number behaviour for the single convective cell heated from below is related to the classical Rayleigh stability problem. The critical Rayleigh number for the single circular cell is approximately five times the value for Rayleigh's multi-celluar configuration. The flow which exhibits a nearly parabolic velocity profile near the critical Rayleigh number, gradually changes to a rigidly-rotating-core behaviour as the Rayleigh number increases. The speed of core rotation is a function of the Prandtl number, whereas the boundary-layer thickness is primarily a function of the Rayleigh number. When the heating is from side to side, the solution shows that as the Rayleigh number increases the core motion is progressively arrested leaving a narrow circulating band of fluid adjacent to the wall. An oblique heating produces a hybrid phenomenon, a low-Rayleigh-number behaviour which is characteristic of the sideways heating case and a high-Rayleigh-number interior motion characteristic of the bottom heating case. To determine the core motion in the high-Rayleigh-number limit, Batchelor's work concerning the uniqueness of incompressible, exactly steady, closed streamline flows with small viscosity is extended to include flows with small thermal conductivity.

Description: The energy flux in a finite-depth gravity-wave spectrum resulting from weak non-linear couplings between the spectral components is evaluated by means of a perturbation method. The fifth-order analysis yields a fourth-order effect comparable in magnitude to the generating and dissipating processes in wind-generated seas. The energy flux favours equidistribution of energy and vanishes in the limiting case of a white, isotropic spectrum. The influence on the equilibrium structure of fully developed wave spectra and on other phenomena in random seas is discussed briefly.

Description: The genus Illex is likely to constitute a large portion of the annual world ommastrephid squid catches (Roper et al, 1984), even though specific official statistics are difficult to obtain. The broad-tail short-fin squid Illex coindetii is a widespread species ranging from the western to the eastern Atlantic (Roper et al., 1984) and east through the whole Mediterranean Sea (Mangold & Boletzky, 1987). Usually a by-catch of important fisheries, it is caught mainly by trawlers. Although its economic value is lower than that of other squid species (i.e. Loligo spp.), in the Sicilian Channel Illex coindetii may represent a valuable resource due to its abundance. In Italian waters, the available statistics (Cingolani et al., 1986) report that 2680 tonnes of ommastrephid squid were landed in 1982 (0.5% of the total landed catch). The main component of these was landed in Sicily (2183 tonnes), a consistently large part of which was no doubt Illex coindetii (Ragonese & Jereb, 1992). The catches came mainly from southern Sicilian waters (Sicilian Channel) where one of the major Mediterranean landing places is in Mazara del Vallo. Large trawlers (up to 200 gross tonnage) usually carry out long fishing trips (15–20 d), and Illex coindetii is caught mainly by those targeting Parapenaeus longirostris and Merluccius merluccius (Jereb & Ragonese, 1991).

Description: The natural feeding of the two most abundant ommastrephid squid (Cephalopoda: Ommastrephidae) in Galician waters was studied and compared. A sample of 334 stomach contents of Todaropsis eblanae (34–222 mm ML) and 267 stomach contents of Illex coindetii (50–379 mm ML) caught by commercial trawlers was examined. A total of 21 (T. eblanae) and 23 (I. coindetii) different prey items, belonging to three zoological groups (Teleostei, Crustacea and Cephalopoda), were taken by these cephalopods. However, 43% of the T. eblanae diet comprised only one fish species, Micromesistius poutassou. The diet of these squid species was significantly influenced by the geographical area (both species), size (T. eblanae) and maturation (I. coindetii). Feeding rate of both species decreased with size, but the percentage of stomachs with food remains increased in maturing and mature females. Weight of prey captured was dependent on available prey sizes and, in small individuals, maximum prey weight was very close to the squid weight. Both squid species are mainly neritic nekto-benthic predators, but I. coindetii seems to have a broader and more pelagic diet.

Description: This communication presents the first records of mated female Loligo gahi in Falkland Island waters. In October 1993 fully mature mated female L. gahi were identified in samples taken from the commercial fishery in waters east of Lively Island, East Falkland, at depths of 145–174 m. Spermatophores were found in both the mantle cavity and buccal sites of deposition. These records, combined with past records of spent females, suggest spawning periods in late October/early November and April/May. These concur with two of the three periods of spawning suggested from previous studies of juvenile and adult L. gahi.

Description: Age, growth and maturation of Loligo vulgaris were studied by examination of growth increments within statoliths of 294 specimens (mantle length, ML, ranging from 31 to 498 mm) caught on the west Saharan shelf between 1985 and 1988. Maximum age was 335 d (290 mm ML) for females, and 396 d (498 mm ML) for males. Growth rates varied considerably among individuals with a greater range in males. Sexual dimorphism in length was apparent after about 210 d. Males and females diverged considerably in weight, with males reaching a greater weight after about 180 d. Growth in length between 124 and 396 d was best described by a power function, while growth in weight was best described by the Gompertz function. Males started maturing at 180–210 d and mature males ranged in age between 250 and 396 d; while females started maturing at 240–270 d and mature females ranged between 285 and 335 d. Loligo vulgaris hatched throughout the year with two distinct peaks; in winter (December - early March) and summer (June-July). The life cycle of L. vulgaris populations on the west Saharan shelf lasts ~1 y, with large males (〉450 mm ML) living slightly longer.

Description: The reproductive strategies of male and female Loligo forbesi Steenstrup, 1856 were investigated from samples obtained from commercial catches and research cruises in Irish waters. In females maturity increased with size, but in males two modes in the size at maturity were identified, with approximately 40% mature at small size (180–200 mm mantle length), and the remainder mature at 〉250 mm mantle length. The difference in estimated age of the two modes of mature males was small, so size differences were probably due to different growth rates. Growth and maturation proceeded together in both sexes over much of the life-cycle. The effect of maturation on relative growth of somatic tissues was examined using analysis of covariance and multivariate regressions. In males there was a significant decline in total mass, and in mass of mantle, head and viscera, relative to mantle length with maturation. In females total mass was not significantly affected by maturation, but relative masses of head, mantle and viscera declined with maturation, indicating that energy was diverted from somatic growth to gonad production. Potential fecundity estimates were obtained by counting eggs and ova in the oviduct and ovary of mature females and were in the range 2500 to 10,500 (mean 5800). Fecundity was positively related to mantle length. The ovaries of mature females contained a range of egg sizes and developmental stages, indicating that spawning probably occurs intermittently.

Description: A total of 57 comparative hauls using a rectangular midwater trawl with a fishing mouth area of 50 m2 (RMT 50) were carried out along the sides of an imaginary triangle south of Madeira in 1986. A total of 1258 cephalopods were caught, giving a mean of 22 per haul with a range from 0 to 67. The nets were used with a diver's light on the top bar which was either switched off or was operated with a 20, 70 or 150 W bulb, powered by a car battery. A significantly greater number of individuals per haul was caught with lights on than without lights, increasing from a mean of 13·5–25·1, a factor of 1·8. Similarly, the number of species caught was increased from a mean of 7 to 10·4, a factor of 1·5 and the volume of cephalopods was increased from a mean of 41·1–162·3ml, a factor of 3·9. Similar comparisons made for catches during day or night separately and on the three courses separately also showed marked increases with the lights. Samples show that increase in power of the lights increased the total number of cephalopod individuals caught. In the 12 species with more than ten individuals, in 33 of the 36 comparisons (of number of individuals, species and volumes) there is an increase with the light. The most influenced species was Taonius pavo which increased in numbers by a mean factor of 3·9 times with 20W, 4·0 times with 70W and 6·1 times with 150W when compared with the numbers caught with no light.

Description: PEEM allows ‘real-time’ observations to be made of solid-state transformations and other high-temperature processes taking place during vacuum-heating up to c. 2000°C The solid state transformations of an amphibole-rich hornblendite specimen have been observed in the temperature range of 750–1000°C (± 50°C Between c. 970–990°C a rapid change in orientation contrast was observed, indicating a structural rearrangement from an oxyhornblende crystal lattice to a clinopyroxene structure. This phase retains the original amphibole shape and texture (including two 120°C intersecting cleavage traces), but possesses a clinopyroxene crystal structure. At higher temperatures this phase is seen to decompose, forming iron oxides and other fine-grained products. PEEM has provided useful information on both the nature and rates of transformation of natural amphiboles which has proved invaluable in our understanding of the mineralogically-controlled mechanisms of argon release during 40Ar/39Ar dating of amphibole samples.

Description: Density-dependent effects of Anguillicola crassus larval infections in the copepod intermediate host were examined experimentally. Three species of copepods (Cyclops vicinus, C. viridis and C. fuscus) were subjected to a range of doses of larval A. crassus within infection arenas. Prevalence, intensity and parasite dispersion (variance: mean abundance) values increase and then approach an asymptote as infection dose increases. Infection parameters differ between species of copepod. Increasing temperature has a negative effect on the establishment of the parasite population within the intermediate host. Parasite-induced host mortality increases with dose. These mechanisms have the potential to regulate populations of A. crassus larvae within the copepod population and hence the whole suprapopulation.

Description: The size distribution of benthic nematodes was investigated along different gradients of food availability in various regions of the north-eastern Atlantic: I, across the continental margin and II, with increasing distance from the continental rise. An overall trend for miniaturization with increasing distance from the food source was found. Moreover, our results indicate that seasonally varying food supply or a periodically pulsed input of organic matter to the sea floor affects nematode size spectra. The hypothesis is proposed that the life cycle of deep-sea nematode species and hence the size structure of their populations are related to seasonal energy availability. This dependence might result in one year life spans of deep-sea nematodes and probably other meiofauna.

Description: Arabian Sea sediments record changes in the upwelling system off Arabia, which is driven by the monsoon circulation system over the NW Indian Ocean. In accordance with climate models, and differing from other large upwelling areas of the tropical ocean, a 500,000-yr record of productivity at ODP Site 723 shows consistently stronger upwelling during interglaciations than during glaciations. Sea-surface temperatures (SSTs) reconstructed from the alkenone unsaturation index (U K′ 37) are high (up to 27°C) during interglaciations and low (22-24°C) during glaciations, indicating a glacial-interglacial temperature change of 〉3°C in spite of the dampening effect of enhanced or weakened upwelling. The increased productivity is attributed to stronger monsoon winds during interglacial times relative to glacial times, whereas the difference in SSTs must be unrelated to upwelling and to the summer monsoon intensity. The winter (NE) monsoon was more effective in cooling the Arabian Sea during glaciations then it is now.

Description: Metazoan meiofauna were studied in replicated multiple-corer samples obtained at a bathyal site (1320–1360 m depth) in the Porcupine Seabight (51°36′N 13°00′W) before (April) and after (July) the delivery to the seafloor of a phytodetrital pulse originating from the 1982 spring bloom. In all samples the metazoan meiofauna was dominated by nematodes; harpacticoid copepods and their nauplii were the second most abundant taxon. Population densities and biomass were very similar in both sample sets, the only significant differences being in the numbers of ostracods (higher in April) and nauplii (higher in July). Furthermore, vertical distribution patterns in the top 5 cm of sediment indicate that the meiofauna did not migrate towards the sediment surface following the phytodetrital pulse. The lack of a metazoan meiofaunal response contrasts with published evidence, based on the same samples, for a substantial increase in the foraminiferal abundance following the sedimentation event. Thus our results suggest that metazoans (as a whole) fail to exploit and utilize phytodetritus as rapidly as foraminifera. This probably reflects the energetic expense of egg production coupled with frequently slower rates of somatic growth among metazoans. In addition, foraminifera may outcompete metazoans for detrital food because they possess extremely efficient food-gathering organelles (granuloreticulate pseudopodia) and are able to raise their levels of metabolic activity very rapidly. However, metazoan responses at the species level, or over longer time periods (〉3 months), would not have been detected and so remain a possibility.

Description: The fully developed flow in a vertical cavity or duct subject to horizontal heating is considered. Solutions of the Boussinesq equations are obtained for rectangular and elliptic sections, in terms of Fourier series and polynomials, respectively. Both generalize the familiar odd-symmetric cubic profile of the plane cavity. Uniqueness is demonstrated under the restriction that the flow is independent of height. For cavities with rectangular sections, it is predicted and verified that the flow in the plane of spanwise symmetry is practically independent of the span if this exceeds 1.7 times the breadth.

Description: An experimental investigation on the initial instability of nonlinear deep-water wave trains including wind effects is reported. The experiment was conducted at the Ocean Engineering Laboratory wind-wave facility (50m long, 4.2m wide, 2.1m deep), with a fully computer-controlled mechanical wave generator to explore the parameter space: steepness; sideband frequency; wind speed. The estimated growth rates of the Benjamin-Feir instability from seeded wind-free experiments agreed well with the theoretical prediction derived from Krasitskii's four-wave reduced equation as computed here. Wind was added to the same wave system ; the growth rates of the sidebands were reduced for weak, and enhanced for strong wind forcing. Experiments with naturally selected sidebands, i.e. unseeded, were conducted as well; measurements showed that wind did not inhibit the growth of sidebands in the case of either two-dimensional or three-dimensional instabilities. A comparison of the results with earlier work suggests that there are two independent effects of wind: first, the alteration of the inviscid growth for a given modulational frequency as shown by comparison with the seeded experiments without wind; second, a change in the natural modulational frequency appearing in the presence of wind which is a function of the wave age, as observed in unseeded experiments. Both effects combined will determine whether the modulational instability is enhanced or suppressed; comparison of experimental results with theoretical predictions suggests that the effect of wind on the natural selection of the modulational frequency is the dominant effect. It was shown that for moderate to old waves, the net effect of wind on the modulational instability is small. For all the experiments except a few unseeded cases with weak breakers, the modulation was small and no breaking was observed within the tank.

Description: A coherent-vortex analysis is made of a computational solution for the free decay of homogeneous, Charney-isotropic geostrophic turbulence at large Reynolds number. The method of analysis is a vortex detection and measurement algorithm that we call a vortex census. The census demonstrates how, through non-conservative interactions among closely approaching vortices, the vortex population evolves towards fewer, larger, sparser, and more weakly deformed vortices. After emergence from random initial conditions and a further period of population adjustment, there is a period of approximately self-similar temporal evolution in the vortex statistics. This behaviour is consistent with a mean-vortex scaling theory based on the conservation of energy, vortex extremum, and vortex aspect ratio. This period terminates as the population approaches a late-time non-turbulent end-state vortex configuration. The end state develops out of merger and alignment interactions among like-sign vortices, and even during the scaling regime, local clusters of nearly aligned vortices are common.

Description: When two drops of radius R touch, surface tension drives an initially singular motion which joins them into a bigger drop with smaller surface area. This motion is always viscously dominated at early times. We focus on the early-time behaviour of the radius rm of the small bridge between the two drops. The flow is driven by a highly curved meniscus of length 2πrm and width Δ ≪ rm around the bridge, from which we conclude that the leading-order problem is asymptotically equivalent to its two-dimensional counterpart. For the case of inviscid surroundings, an exact two-dimensional solution (Hopper 1990) shows that Δ ∝ rm3 and rm ∼ (tγ/πη) ln [tγ/(ηR)]; and thus the same is true in three dimensions. We also study the case of coalescence with an external viscous fluid analytically and, for the case of equal viscosities, in detail numerically. A significantly different structure is found in which the outer-fluid forms a toroidal bubble of radius Δ ∝ rm3/2 at the meniscus and rm ∼ (tγ/4πη) ln [tγ/(ηR)]. This basic difference is due to the presence of the outer-fluid viscosity, however small. With lengths scaled by R a full description of the asymptotic flow for rm(t) ≪ 1 involves matching of lengthscales of order rm2 rm3/2, rm, 1 and probably rm7/4.

Description: In this paper we investigate the linear stability of detonations in which the underlying steady one-dimensional solutions are of the pathological type. Such detonations travel at a minimum speed, which is greater than the Chapman-Jouguet (CJ) speed, have an internal frozen sonic point at which the thermicity vanishes, and the unsupported wave is supersonic (i.e. weak) after the sonic point. Pathological detonations are possible when there are endothermic or dissipative effects present in the system. We consider a system with two consecutive irreversible reactions A→B→C, with an Arrhenius form of the reaction rates and the second reaction endothermic. We determine analytical asymptotic solutions valid near the sonic pathological point for both the one-dimensional steady equations and the equations for linearized perturbations. These are used as initial conditions for integrating the equations. We show that, apart from the existence of stable modes, the linear stability of the pathological detonation is qualitatively the same as for CJ detonations for both one- and two-dimensional disturbances. We also consider the stability of overdriven detonations for the system. We show that the frequency of oscillation for one-dimensional disturbances, and the cell size based on the wavenumber with the highest group velocity for two-dimensional disturbances, are both very sensitive to the detonation speed for overdriven detonations near the pathological speed. This dependence on the degree of overdrive is quite different from that obtained when the unsupported detonation is of the CJ type.

Description: This paper considers the role of long finite-amplitude Rossby waves in determining the evolution of flow along a rapidly rotating channel with an uneven floor. The Rossby waves travel on a potential vorticity interface in a channel with a cross-channel step change in depth, where step position varies slowly along the channel. A nonlinear wave equation is derived describing the evolution of the potential vorticity interface. To leading order this is the hydraulic equation derived by Haynes, Johnson & Hurst (1993). Dispersion appears at the next order. Various solution regimes are identified. As well as slowly varying hydraulic solutions, two further types of steady solutions appear: approach-controlled flows and twin supercritical leaps. Both these solutions are characterized by leaps between supercritical branches of the hydraulic function. It is shown how the position and size of these 'supercritical leaps' can be determined within the context of hydraulic theory. To fully resolve the internal structure of these leaps dispersive effects must be included and leaps are shown to correspond to kink soliton solutions of the steady unforced problem. It is also shown that increasing dispersion (decreasing topographic length scale) causes the loss of the subcritical solution branch in some subcritical flows. The only candidate for a steady solution in these regimes is then an approach-controlled flow. Integrations of initial value problems show that in general flows evolve towards the dispersive form of the solution predicted by hydraulic theory, at least near the topographic perturbation. However, in those subcritical flows where sufficiently large dispersion causes the subcritical branch to disappear, unsteady integrations evolve to approach-controlled flows even when the dispersion is sufficiently small that the subcritical branch still exists.

Description: The transient evolution of the bubble-size probability density functions resulting from the breakup of an air bubble injected into a fully developed turbulent water flow has been measured experimentally using phase Doppler particle sizing (PDPA) and image processing techniques. These measurements were used to determine the breakup frequency of the bubbles as a function of their size and of the critical diameter Dc defined as Dc = 1.26 (σ/ρ)3/5∈-2/5, where ∈ is the rate of dissipation per unit mass and per unit time of the underlying turbulence. A phenomenological model is proposed showing the existence of two distinct bubble size regimes. For bubbles of sizes comparable to Dc, the breakup frequency is shown to increase as (σ/ρ)-2/5 ∈3/5 √D/Dc-1, while for large bubbles whose sizes are greater than 1.63Dc, it decreases with the bubble size as ∈1/3 D-2/3. The model is shown to be in good agreement with measurements performed over a wide range of bubble sizes and turbulence intensities.

Description: Direct numerical simulations of turbulence resulting from Kelvin-Helmholtz instability in stratified shear flow are used to examine the geometry of the dissipation range in a variety of flow regimes. As the buoyancy and shear Reynolds numbers that quantify the degree of isotropy in the dissipation range increase, alignment statistics evolve from those characteristic of parallel shear flow to those found previously in studies of stationary, isotropic, homogeneous turbulence (e.g. Ashurst et al. 1987; She et al. 1991; Tsinober et al. 1992). The analysis yields a limiting value for the mean compression rate of scalar gradients that is expected to be characteristic of all turbulent flows at sufficiently high Reynolds number. My main focus is the value of the constant q that appears in both the Batchelor (1959) and Kraichnan (1968) theoretical forms for the passive scalar spectrum. Taking account of the effects of time-dependent strain, I propose a revised estimate of q, denoted qe, which appears to agree with spectral shapes derived from simulations and observations better than do previous theoretical estimates. The revised estimate is qe = 7.3±0.4, and is expected to be valid whenever the buoyancy Reynolds number exceeds O(102). The Kraichnan (1968) spectral form, in which effects of intermittency are accounted for, provides a better fit to the DNS results than does the Batchelor (1959) form.

Description: A methodology termed the 'filtered mass density function' (FMDF) is developed and implemented for large-eddy simulation (LES) of variable-density chemically reacting turbulent flows at low Mach numbers. This methodology is based on the extension of the 'filtered density function' (FDF) scheme recently proposed by Colucci et al (1998) for LES of constant-density reacting flows. The FMDF represents the joint probability density function of the subgrid-scale (SGS) scalar quantities and is obtained by solution of its modelled transport equation. In this equation, the effect of chemical reactions appears in a closed form and the influences of SGS mixing and convection are modelled. The stochastic differential equations (SDEs) which yield statistically equivalent results to those of the FMDF transport equation are derived and are solved via a Lagrangian Monte Carlo scheme. The consistency, convergence, and accuracy of the FMDF and the Monte Carlo solution of its equivalent SDEs are assessed. In non-reacting flows, it is shown that the filtered results via the FMDF agree well with those obtained by the 'conventional' LES in which the finite difference solution of the transport equations of these filtered quantities is obtained. The advantage of the FMDF is demonstrated in LES of reacting shear flows with non-premixed reactants. The FMDF results are appraised by comparisons with data generated by direct numerical simulation (DNS) and with experimental measurements. In the absence of a closure for the SGS scalar correlations, the results based on the conventional LES are significantly different from those obtained by DNS. The FMDF results show a closer agreement with DNS. These results also agree favourably with laboratory data of exothermic reacting turbulent shear flows, and portray several of the features observed experimentally.

Description: The objective of this study is to develop a method of controlling vortex shedding behind a bluff body using control theory. A suboptimal feedback control procedure for local sensing and local actuation is developed and applied to the flow behind a circular cylinder. The location of sensors for feedback is limited to the cylinder surface and the control input from actuators is the blowing and suction on the cylinder surface. Three different cost functionals to be minimized (J1 and J2) or maximized (J3) are investigated: J1 is proportional to the pressure drag of the cylinder, J2 is the square of the difference between the target pressure (inviscid flow pressure) and real flow pressure on the cylinder surface, and J3 is the square of the pressure gradient on the cylinder surface, respectively. Given the cost functionals, the flow variable to be measured by the sensors and the control input from the actuators are determined from the suboptimal feedback control procedure. Several cases for each cost functional have been numerically simulated at Re = 100 and 160 to investigate the performance of the control algorithm. For all actuations, vortex shedding becomes weak or disappears, and the mean drag and drag/lift fluctuations significantly decrease. For a given magnitude of the blowing/suction, reducing J2 provides the largest drag reduction among the three cost functionals.

Description: Using a combination of bifurcation theory for two-dimensional dynamical systems and numerical simulations, we systematically determine the possible flow topologies of the steady vortex breakdown in axisymmetric flow in a cylindrical container with rotating end-covers. For fixed values of the ratio of the angular velocities of the covers in the range from -0.02 to 0.05, bifurcations of recirculating bubbles under variation of the aspect ratio of the cylinder and the Reynolds number are found. Bifurcation curves are determined by a simple fitting procedure of the data from the simulations. For the much studied case of zero rotation ratio (one fixed cover) a complete bifurcation diagram is constructed. Very good agreement with experimental results is obtained, and hitherto unresolved details are determined in the parameter region where up to three bubbles exist. For non-zero rotation ratios the bifurcation diagrams are found to change dramatically and give rise to other types of bifurcations.

Description: The high Reynolds number flow through a circular pipe divided along a diameter by a semi-infinite splitter plate is considered. Matched asymptotic expansions are used to analyse the developing flow, which is decomposed into four regions: a boundary layer of Blasius type growing along the plate, an inviscid core, a viscous layer close to the curved wall and a nonlinear corner region. The core solution is found numerically, initially in the long-distance down-pipe limit and thereafter the full problem is solved using down-pipe Fourier transforms. The accuracy in the corners of the semicircular cross-section is improved by subtracting out the singularity in the velocity perturbation. The linear viscous wall layer is solved analytically in terms of a displacement function determined from the core. A plausible structure for the corner region and equations governing the motion therein are presented although no solution is attempted. The presence of the plate has little effect ahead of the bifurcation, but wall shear on the curved wall is found to increase from its undisturbed value downstream.

Description: The behaviour of the long-time self-diffusion tensor in concentrated colloidal dispersions is studied using dynamic simulation. The simulations are of a suspension of monodisperse Brownian hard spheres in simple shear flow as a function of the Péclet number, Pe, which measures the relative importance of shear and Brownian forces, and the volume fraction, φ. Here, Pe = γ̇a2/D0, where γ̇ is the shear rate, a the particle size and D0 = kT/6πηa is the Stokes-Einstein diffusivity of an isolated particle of size a with thermal energy kT in a solvent of viscosity η. Two simulations algorithms are used: Stokesian Dynamics for inclusion of the many-body hydrodynamic interactions, and Brownian Dynamics for suspensions without hydrodynamic interactions. A new procedure for obtaining high-quality diffusion data based on averaging the results of many short simulations is presented and utilized. At low shear rates, low Pe, Brownian diffusion due to a random walk process dominates and the characteristic scale for diffusion is the Stokes-Einstein diffusivity, D0. At zero Pe the diffusivity is found to be a decreasing function of φ. As Pe is slowly increased, 0(Pe) and 0(Pe3/2) corrections to the diffusivity due to the flow are clearly seen in the Brownian Dynamics system in agreement with the theoretical results of Morris & Brady (1996). At large shear rates, large Pe, both systems exhibit diffusivities that grow linearly with the shear rate by the non-Brownian mechanism of shear-induced diffusion. In contrast to the behaviour at low Pe, this shear-induced diffusion mode is an increasing function of φ. Long-time rotational self-diffusivities are of interest in the Stokesian Dynamics system and show similar behaviour to their translational analogues. An off-diagonal long-time self-diffusivity, Dxy, is reported for both systems. Results for both the translational and rotational Dxy. show a sign change from low Pe to high Pe due to different mechanisms in the two regimes. A physical explanation for the off-diagonal diffusivities is proposed.

Description: Based on energy principles, we propose a statistical model to describe the bubble size probability density function of the daughter bubbles resulting from the shattering of a mother bubble of size D0 immersed in a fully developed turbulent water flow. The model shows that the bubble size p.d.f. depends not only on D0, but also on the value of the dissipation rate of turbulent kinetic energy of the underlying turbulence of the water, ∈. The phenomenological model is simple, yet it predicts detailed experimental measurements of the transient bubble size p.d.f.s performed over a range of bubble sizes and dissipation rates ∈ in a very consistent manner. The agreement between the model and the experiments is particularly good for low and moderate bubble turbulent Weber numbers, Wet = ρ Δu2(D0)D0/σ where the assumption of the binary breakup is shown to be consistent with the experimental observations. At larger values of Wet, it was found that the most probable number of daughter bubbles increases and the assumption of tertiary breakup is shown to lead to a better fit of the experimental measurements.

Description: Using the continuum model of Pedley, Hill & Kessler (1988) for bioconvection in a suspension of swimming, gyrotactic micro-organisms, we investigate the existence and stability of a two-dimensional plume in tall, narrow chambers with stress-free sidewalls. The system is governed by the Navier-Stokes equations for an incompressible fluid coupled with a micro-organism conservation equation. These equations are solved numerically using a conservative finite-difference scheme. In sufficiently deep chambers, the plume is always unstable to both varicose and meandering modes. A linear stability analysis for an infinitely long plume predicts the growth rates of these instabilities, explains the mechanisms, and is in good agreement with the numerical results.

Description: A three-dimensional mathematical model based on the Brinkman extended Darcy equation has been used to study double-diffusive natural convection in a fluidsaturated porous cubic enclosure subject to opposing and horizontal gradients of temperature and concentration. The flow is driven by conditions of constant temperature and concentration imposed along the two vertical sidewalls of the cubic enclosure, while the remaining walls are impermeable and adiabatic. The numerical simulations presented here span a wide range of porous thermal Rayleigh number, buoyancy ratio and Lewis number to identify the different steady-state flow patterns and bifurcations. The effect of the governing parameters on the domain of existence of the three-dimensional flow patterns is studied for opposing flows (N 〈 0). Comprehensive Nusselt and Sherwood number data are presented as functions of the governing parameters. The present results indicate that the double-diffusive flow in enclosures with opposing buoyancy forces is strictly three-dimensional for a certain range of parameters. At high Lewis numbers multiple dipole vortices form in the transverse planes near the horizontal top and bottom surfaces, which the two-dimensional models fail to detect. The dipolar vortex structures obtained are similar to those created in laboratory experiments by the injection of fluid into a stratified medium.

Description: The influence of inertia and elasticity on the onset and stability of Taylor-vortex flow (TVF) is examined for an Oldroyd-B fluid. The Galerkin projection method is used to obtain the departure from Couette flow (CF). Only axisymmetric flow is examined. The solution is capable of capturing the dynamical behaviour observed experimentally for viscoelastic fluids in the inertio-elastic and purely elastic ranges. For flow with dominant inertia, the bifurcation picture is similar to that for a Newtonian fluid. However, transition from CF to TVF is oscillatory because of fluid elasticity. Steady TVF sets in, via supercritical bifurcation, as Re reaches a critical value, Rec. The critical Reynolds number decreases with fluid elasticity, and is strongly influenced by fluid retardation. As elasticity exceeds a critical level, a subcritical bifurcation emerges at Rec, similar to that predicted by the Landau-Ginzburg equation. It is found that slip along the axial direction tends to be generally destabilizing. The coherence of the formulation is established under steady and transient conditions through comparison with exact linear stability analysis, experimental measurements, and flow visualization. Good agreement is obtained between theory and the measurements of Muller et al. (1993) in the limit of purely elastic overstable TVF.

Description: We present simulation results of vortex-induced vibrations of an infinitely long flexible cylinder at Reynolds number Re = 1000, corresponding to a 'young' turbulent wake (i.e. exhibiting a small inertial subrange). The simulations are based on a new class of spectral methods suitable for unstructured and hybrid grids. To obtain different responses of the coupled flow-structure system we vary the structure's bending stiffness to model the behaviour of a vibrating inflexible (rigid) cylinder, a cable, and a beam. We have found that unlike the laminar flow previously studied, the amplitude of the cross-flow oscillation is about one diameter for the cable and the beam, close to experimental measurements, but is lower for the rigid cylinder. We have also found that for the latter case the flow response corresponds to parallel shedding, but for the beam and cable with free endpoints a mixed response consisting of oblique and parallel shedding is obtained, caused by the modulated travelling wave motion of the structure. This mixed shedding pattern which alternates periodically along the span can be directly related to periodic spatial variation of the lift force. In the case of structures with pinned endpoints a standing wave response is obtained for the cylinder; lace-like flow structures are observed similar to the ones seen in the laminar regime. Examination of the frequency spectra in the near wake shows that at Re = 1000 all cases follow a -5/3 law in the inertial range, which extends about half a decade in wavenumber. However, these spectra are different in all three cases both in low and high frequencies, with the exception of the beam and cable, for which the high-frequency portion is identical despite the differences in the displacement time history and the large-scale features of the corresponding flow.

Description: The incompressible boundary layer in the corner formed by two intersecting, semi-infinite planes is investigated, when the free-stream flow, aligned with the corner, is taken to be of the form U∞F(x), x representing the non-dimensional streamwise distance from the leading edge. In Dhanak & Duck (1997) similarity solutions for F(x) = xn were considered, and it was found that solutions exist for only a range of values of n, whilst for ∞ 〉 n 〉 -0.018, approximately, two solutions exist. In this paper, we extend the work of Dhanak & Duck to the case of non-90° corner angles and allow for streamwise development of solutions. In addition, the effect of transpiration at the walls of the corner is investigated. The governing equations are of boundary-layer type and as such are parabolic in nature. Crucially, although the leading-order pressure term is known a priori, the third-order pressure term is not, but this is nonetheless present in the leading-order governing equations, together with the transverse and crossflow viscous terms. Particular attention is paid to flows which develop spatially from similarity solutions. It turns out that two scenarios are possible. In some cases the problem may be treated in the usual parabolic sense, with standard numerical marching procedures being entirely appropriate. In other cases standard marching procedures lead to numerically inconsistent solutions. The source of this difficulty is linked to the existence of eigensolutions emanating from the leading edge (which are not present in flows appropriate to the first scenario), analogous to those found in the computation of some two-dimensional hypersonic boundary layers (Neiland 1970; Mikhailov et al. 1971; Brown & Stewartson 1975). In order to circumvent this difficulty, a different numerical solution strategy is adopted, based on a global Newton iteration procedure. A number of numerical solutions for the entire corner flow region are presented.

Description: A generalized model of flow in meandering subaqueous and subaerial channels is developed. The conservation equations of mass and momentum are depth/layer integrated, normalized, and represented as deviations from a straight base state. This allows the determination of integrable forms which can be solved at both linear and nonlinear levels. The effects of various flow and geometric parameters on the flow dynamics are studied. Although the model is not limited to any specific planform, this study focuses on sine-generated curves. In analysing the flow patterns, the turbidity current of the subaqueous case is simplified to a conservative density flow with water entrainment from above neglected. The subaqueous model thus formally corresponds to a subcritical or only mildly supercritical mud-rich turbidity current. By extension, however the analysis can be applied to a depositional or erosional current carrying sand that is changing only slowly in the streamwise direction. By bringing the subaqueous and subaerial cases into a common form, flow behaviour in the two environments can be compared under similar geometric and boundary conditions. A major difference between the two cases is the degree of superelevation of channel flow around bends, which is modest in the subaerial case but substantial in the subaqueous case. Another difference concerns Coriolis effects: some of the largest subaqueous meandering systems are so large that Coriolis effects can become important. The model is applied to meander bends on the youngest channel in the mid-fan region of the Amazon Fan and a mildly sinuous bend of the North-West Atlantic Mid-Ocean Channel. In the absence of specific data on the turbid flows that created the channel, the model can be used to make inferences about the flow, and in particular the range of values of flow velocity and sediment concentration that would allow the growth and downfan migration of meander bends.

Description: The collisional flow of a slightly inelastic granular material down a rough inclined plane is usually described by kinetic theories. We present an experimental study aimed at analysing the assumptions and the quantitative predictions of such theories. A two-dimensional channel coupled to a model granular material and image analysis allow detailed and complete measurement of the kinematics and structure of the flows. We determine the range of inclination and particle flux for which the flow is stationary and uniform. The characteristic profiles of solid fraction, mean velocity and granular temperature are systematically measured. Both the true collisional and the dilute kinetic regimes are examined. We show that a quasi-hydrodynamic description of these regimes seems relevant, and that the pressure and the viscosity terms are in good qualitative agreement with the prediction of the kinetic theory. The profiles are well described by the kinetic theory near the top of the flow, at low solid fraction. Conversely there are large discrepancies near the rough plane, where the material is structured in layers.

Description: A linear stability analysis is performed for the homogeneous state of a monodisperse gas-fluidized bed of spherical particles undergoing hydrodynamic interactions and solid-body collisions at small particle Reynolds number and finite Stokes number. A prerequisite for the stability analysis is the determination of the particle velocity variance which controls the particle-phase pressure. In the absence of an imposed shear, this velocity variance arises solely due to the hydrodynamic interactions among the particles. Since the uniform state of these suspensions is unstable over a wide range of values of particle volume fraction φ and Stokes number St, full dynamic simulations cannot be used in general to characterize the properties of the homogeneous state. Instead, we use an asymptotic analysis for large Stokes numbers together with numerical simulations of the hydrodynamic interactions among particles with specified velocities to determine the hydrodynamic sources and sinks of particle-phase energy. In this limit, the velocity distribution to leading order is Maxwellian and therefore standard kinetic theories for granular/hard-sphere molecular systems can be used to predict the particle-phase pressure and rheology of the bed once the velocity variance of the particles is determined. The analysis is then extended to moderately large Stokes numbers for which the anisotropy of the velocity distribution is considerable by using a kinetic theory which combines the theoretical analysis of Koch (1990) for dilute suspensions (φ ≪ 1) with numerical simulation results for non-dilute suspensions at large Stokes numbers. A linear stability analysis of the resulting equations of motion provides the first a priori predictions of the marginal stability limits for the homogeneous state of a gas-fluidized bed. Dynamical simulations following the detailed motions of the particles in small periodic unit cells confirm the theoretical predictions for the particle velocity variance. Simulations using larger unit cells exhibit an inhomogeneous structure consistent with the predicted instability of the homogeneous gas-solid suspension.

Description: A combined numerical/analytical study of the wide-vortex/wave interaction equations, describing boundary-layer instability, is presented. Depending on the obliqueness βof the wave input, different solution properties are obtained. For β= 1, oscillations in the wave amplitude lead to the evolution of a strongly three-dimensional mean flow, while for β= 2 the interaction is characterized by the development of a singularity in the wave pressure amplitude. This latter behaviour is modelled using an approximate form for the mean flow skin friction and the resulting amplitude equation is analysed using a combination of numerical and asymptotic techniques. A simple method is described for determining the singularity location for a given span wise wavenumber, and the asymptotic behaviour of the pressure amplitude as the singularity is approached is deduced.

Description: Using direct numerical simulations (DNS) and large-eddy simulations (LES) of velocity and passive scalar in isotropic turbulence (up to 5123 grid points), we examine directly and quantitatively the refined similarity hypotheses as applied to passive scalar fields (RSHP) with Prandtl number of order one. Unlike previous experimental investigations, exact energy and scalar dissipation rates were used and scaling exponents were quantified as a function of local Reynolds number. We first demonstrate that the forced DNS and LES scalar fields exhibit realistic inertial-range dynamics and that the statistical characteristics compare well with other numerical, theoretical and experimental studies. The Obukhov-Corrsin constant for the k-5/3 scalar variance spectrum obtained from the 5123 mesh is 0.87 %plusmn; 0.10. Various statistics indicated that the scalar field is more intermittent than the velocity field. The joint probabiltiy distribution of locally-averaged energy dissipation ∈r and scalar dissipation χr is close to log-normal with a correlation coefficient of 0.25 ± 0.01 between the logarithmic dissipations in the inertial subrange. The intermittency parameter for scalar dissipation is estimated to be in the range 0.43 ∼ 0.77, based on direct calculations of the variance of χr. The scaling exponents of the conditional scalar increment δ;rθ|χr,εr suggest a tendency to follow RSHP. Most significantly, the scaling exponent of δrθ|χr,εr over ∈r wass shown to be approximately -1/6 in the inertial subrange, confirming a dynamical aspect of RSHP. In agreement with recent experimental results (Zhu et al. 1995; Stolovitzky et a. 1995), the probability distributions of the random variable . βs = δrθ|χr,εr/χr 1/2εr-1/6r1/3 were found to be nearly Gaussian. However, contrary to the experimental results, we find that the moments of βs are almost identical to those for the velocity field found in Part 1 of this study (Wang et al. 1996) and are insensitive to Reynolds number, large-scale forcing, and subgrid modelling.subgrid modelling.

Description: This paper presents the results of experimental studies of a collapsing cylindrical cavity (the convergence of a liquid shell) in a rotating fluid as well as the formation and propagation of a jump (bore) at the interface. The basic parameters of the liquid shell dynamics for a pulsed one-dimensional load are estimated using the equation of cylindrical cavity pulsation in an unbounded fluid. The theoretical model of a rotationally symmetric hydraulic jump moving along the free surface of a hollow vortex is constructed. The jump is simulated by a discontinuous solution of the equations in the long-wave approximation for tornado-like and hollow vortices. For comparison with the experimental data, basic theoretical results are obtained for flows in a hollow vortex with constant circulation and axial velocity varying along the radius of the rotating liquid shell.

Description: In this paper, an experimental study of laminar magnetohydrodynamic (MHD) buoyancy-driven flow in a cylindrical cell with axis horizontal is described. A steady uniform magnetic field is applied vertically to the mercury-filled cell, which is also subjected to a horizontal temperature gradient. The main features of this internal MHD thermogravitational flow are made experimentally evident from temperature and electric potential measurements. Whatever the level of convection, raising the Hartmann number Ha to a value of the order of 10 is sufficient to stabilize an initially turbulent flow. At much higher values of the Hartmann number (Ha ∼ 100) the MHD effects cause a change of regime from boundary-layer driven to core driven. In this latter regime an inviscid inertialess MHD core flow is bounded by a Hartmann layer on the horizontal cylindrical wall and viscous layers on the endwalls. Since the Hartmann layer is found to stay electrically inactive along the cell, the relevant asymptotic (Ha ≫ 1) laws for velocity and heat transfer are found from the balance between the curl of buoyancy and Lorentz forces in the core, together with the condition that the flow of electric current between core and Hartmann layer is negligible. A modified Rayleigh number RaG/Ha2, which is a measure of the ratio of thermal convection to diffusion when there is a balance between buoyancy and Lorentz forces, is the determining parameter for the flow.

Description: The island of Minorca, Spain, has become a patchy mosaic landscape as a result of centuries of human-induced fragmentation. The dynamics of the landscape elements and the spatial pattern of Minorca were analysed to test whether this human-modelled landscape was at a stationary state, and, if it was the case, to see whether the system could be characterized by a particular spatial pattern. Landsat TM satellite images were processed to derive land cover classification and vegetation index maps corresponding to the years 1984 and 1992, the best compromise between null cloudiness and maximum time span. The classification was used to test the stationary state by estimating the transition matrix of land cover. The Normalized Difference Vegetation Index (NDVI) was used to analyse the spatial pattern of change at the patch level. The spatial analysis of NDVI was based on four indices: fractal dimension, number and size distribution of patches, and spatial Kappa index. Those pattern descriptors were compared with values obtained from stochastic landscape simulations. Temporal analysis showed that land cover proportions remained approximately constant over the 8-year period, although interchange amongst patches existed. This suggested that the landscape of Minorca was close to a stationary state. The study of NDVI changes revealed that the spatial structure was different from that of neutral models and presented scale invariance properties. Results from land cover transitions and from NDVI patterning suggest that the landscape of Minorca is a critical system, in the framework of the self-organized criticality hypothesis, i.e. the mosaic of patches would self-organize at the frontier of a dynamic equilibrium constituted by the balance between disturbances and successional processes.

Description: There are numerous reasons why land users do not always adopt innovations even though all evidence may suggest that they should. The purpose of this study was to review the literature in the field of extension and technology transfer drawing upon the rural sector primarily in Australia. Australia is now recognized as a world leader in adopting new technology and ideas, particularly those relating to the management of the physical environment but there has been little assessment of the circumstances and constraints surrounding this adoption. The reasons for non-adoption range from complexity of the technology, land user's beliefs and opinions towards the innovation, attitude to risk and conflicting information. For different land-use enterprises and innovations, different constraints will apply and it is argued that these are in three broad categories, characterized by: the land user and the adoption process; the characteristics of, and issues associated with the developers of, the innovation; and the role of extension agents and the transfer process. It is apparent from the changing nature of technology transfer that commercial organizations have assumed a leading role in this field and land users are being recognized as key stakeholders in both the adoption and technology development processes. The study, however, reveals a shortage of data on the effectiveness of corporations and other commercial organizations in the technology transfer and adoption processes.

Description: The Sierra de Tamaulipas is a biogeographically isolated mountain system in Northern Mexico, where habitat fragmentation by land-management practices is a possible threat to wildlife conservation. As a case example, we used GIS analyses to evaluate how human activities influence the landscape structure of jaguar (Panthera onca) habitat in the region. The study: (1) ranked potential habitat based on associations between environmental attributes (topography, streams and vegetation) and the frequency distribution of jaguar sighting records; (2) classified current land cover from a 1990 Landsat-TM image and mapped the landscape structure of high potential habitat; and (3) compared the degree to which mature natural vegetation is fragmented by different types of owners. Jaguar sites showed significant associations with tropical deciduous and oak forests, and low, west or south-east slopes, between 400 and 900 m. About 52% of the high potential habitat was mapped as mature natural vegetation, which was distributed as two large patches (28% of the land area) and many small forest patches (98% at 〈 80 ha). The number and size-class distribution of high-potential habitat patches varied little amongst four ownership types, but the dispersed distribution of more subsistence and commercial-based owners across the landscape suggests the need for collaborative participation in a conservation plan. From our study the need to scale up from managing individual land parcels is substantiated and areas that promote regional contiguity of jaguar habitat in the Sierra de Tamaulipas are identified.

Description: Having been deeply involved with the Convention on Biological Diversity (CBD) since it was first conceived at the World Parks Congress in Bali in 1982 (de Klemm 1984), I wish to respond to Guruswamy's (1999) critique of it.

Description: Wetland forest communities are seriously reduced and degraded throughout most of Europe, but significant areas still survive in the east and north-east. Data from 91 forest structures that support biodiversity are analysed in 369 relevés of Latvian wetland forests. These structures include variations in topography, stand composition, water movement and dead wood features occurring in semi-natural conditions undisturbed by management. There is clear structural differentiation between coniferous and deciduous wetlands. The study supports the widely held views on European forest quality: low volume of high-diameter (〉50 cm) snags and logs, too few overmature trees, and the threat to pattern heterogeneity caused by traditional forestry and silviculture. Twenty areas (totalling c. 3800 ha) are identified as reserves that should be strictly protected in Latvia in order to maintain representative and intact samples of wet forest belonging to Eurosiberian alder swamps (Alnetea glutinosae), broad-leaved forests (Querco-Fagetea) and boreal forest (Vaccinio-Piceetea) communities. Based on habitat features and site ecology, the concept of sustainable forestry must be incorporated widely in utilization of European wetland forests in order to prevent further loss of biodiversity.

Description: The environmental service provided by the great biodiversity of Amazonian forests is one of several factors leading to the conclusion that much greater efforts are warranted to reduce the destruction of these forests. Risks to biodiversity in Amazonian forests include deforestation, logging, fires, fragmentation, depletion of fauna, invasion by exotic species, and climate change. Financial values assigned to biodiversity depend strongly on the purposes of valuation. Utilitarian benefits include the values of presently-marketed and presently-unexploited forest products, and the monetary value of environmental benefits. Non-monetary values of Amazonian forests are also essential components of decision-making on conservation. Measures of ‘willingness to pay’ and ‘willingness to accept’ can be useful as indicators of potential financial flows, but should not be confused with the true values of the forests to society. Valuation for the purpose of setting penalties for destruction of biodiversity is an important legal question in Brazil and must take into consideration additional factors.Conservation of biodiversity in Brazil includes creation of various types of protected areas. The status of these areas varies greatly, with practice frequently deviating from official requirements. Creating reserves that include human occupants has a variety of pros and cons. Although the effect of humans is not always benign, much larger areas can be brought under protection regimes if human occupants are included. Additional considerations apply to buffer zones around protected areas. The choice and design of reserves depends on the financial costs and biodiversity benefits of different strategies. In Brazil, rapid creation of lightly-protected ‘paper parks’ has been a means of keeping ahead of the advance of barriers to establishment of new conservation units, but emphasis must eventually shift to better protection of existing reserves. Indigenous peoples have the best record of maintaining forest, but negotiation with these peoples is essential in order to ensure maintenance of the large areas of forest they inhabit. The benefits of environmental services provided by the forest must accrue to those who maintain these forests. Development of mechanisms to capture the value of these services will be a key factor affecting the long-term prospects of Amazonian forests. However, many effective measures to discourage deforestation could be taken immediately through government action, including levying and collecting taxes that discourage land speculation, changing land tenure establishment procedures so as not to reward deforestation, revoking remaining incentives, restricting road building and improvement, strengthening requirements for environmental impact statements (RIMAs) for proposed development projects, and creating employment alternatives.

Description: The Confidential Awards Committee voted Carolyn Crook and Roger Alex Clapp the joint recipients of the Volume 25 ‘Best Paper’ Prize for their paper ‘Is market-oriented forest conservation a contradiction in terms?’ (Crook & Clapp 1998). Their prize was a cheque for SFr 1000, the proceeds being shared between the co-authors.

Description: Rakiura Maori (a tribe of indigenous people in New Zealand) continue a centuries-old customary use of Sooty Shearwater (Puffinus griseus, titi, muttonbird) chicks from islands adjacent to Rakiura (Stewart Island). Some muttonbirders pluck chicks by hand, while others have recently changed to a plucking machine. We compared traditional and modern processing methods to see if new technology stands to increase the efficiency, size and cost effectiveness of harvest. On average, chicks were plucked 6 seconds quicker with a machine, which could potentially increase the catch by up to 4%. Innovation by using wax rather than water to remove down left after plucking saved muttonbirders 29–97 minutes per day, potentially allowing up to a 15% increase in the number of chicks harvested. Both wax and plucking machines increased costs, which led to a modest financial gain from using wax, but a net loss from using a plucking machine. Modern technologies have been introduced mainly for convenience and to ease labour in this customary use of wildlife. New technology may erode traditional skills, but does not necessarily pose a risk to the sustainability of a resource. Financial investment in harvest technologies might provide an incentive to increase harvest levels, but could equally provide an incentive to manage for sustainable use. Preservation lobbies are not justified in presuming that new technologies will always threaten wildlife traditionally used by indigenous people.

Description: Despite the increasing call for cooperation amongst the states parties to the 1992 Convention on Biological Diversity (UNTS 3069; see especially its Articles 5, 17, 18 and 20), the management of protected areas adjacent to political borders is rarely thought of as a common necessity for neighbouring countries. Setting up a zone dedicated to conservation is already a formidable challenge in the face of strongly competing interests for land. Bringing this about with two or more sovereign states sometimes seems a ‘gratuitous layer of complexity that spells almost certain failure’ (Westing 1998b). Nevertheless, transboundary protected areas are becoming increasingly common (IUCN, unpublished 1998). In fact, no less than 136 such protected area complexes involving over 406 individual protected areas and 112 different international boundaries (Zbicz & Green 1997) have been identified around the world. If proposed areas are also included, this figure can be pushed up to 200 complexes (Brunner 1998).

Description: Norman Myers recently challenged scientists to consider the social responsibilities that derive from their expertise and knowledge (Myers 1999). He raised important questions regarding how pro-active scientists can or should be in leading public debate about the state of the environment. This article is a response to that call, and hopefully will serve to stimulate further debate on the topic.

Description: The degradation of trail resources associated with expanding recreation and tourism visitation is a growing management problem in protected areas worldwide. In order to make judicious trail and visitor management decisions, protected area managers need objective and timely information on trail resource conditions. This paper introduces a trail survey method that efficiently characterizes the location and lineal extent of common trail problems. The method was applied to a large sample of trails within Great Smoky Mountains National Park, a high-use protected area in the USA. The Trail Problem-Assessment Method (TPAM) employs a continuous search for multiple indicators of pre-defined tread problems, yielding census data documenting the location, occurrence and extent of each problem. The present application employed 23 different indicators in three categories to gather inventory, resource condition, and design and maintenance data of each surveyed trail. Seventy-two backcountry hiking trails (528 km), or 35% of the Park's total trail length, were surveyed. Soil erosion and wet soil were found to be the two most common impacts on a lineal extent basis. Trails with serious tread problems were well distributed throughout the Park, although trails with wet muddy treads tended to be concentrated in areas where horse use was high. The effectiveness of maintenance features installed to divert water from trail treads was also evaluated. Water bars were found to be more effective than drainage dips. The TPAM was able to provide Park managers with objective and quantitative information for use in trail planning, management and maintenance decisions, and is applicable to other protected areas elsewhere with different environmental and impact characteristics.

Description: Field experiments were conducted at the farm of Qinghai Academy of Animal and Veterinary Science, Xining, China during 1996/97 to determine the effects of level of food intake on the urinary excretion of purine derivatives (PD), creatinine and nitrogen in yaks (Bos grunniens). Two experiments were carried out with three female yaks (initial body weight 173–187 kg, age 5 years). For Expt 1 a 3×3 Latin square experimental design was used with three levels of oat hay (nitrogen 13·5 g/kg dry matter (DM)) intake treatments, i.e. 0·3, 0·6 and 0·9 of voluntary intake (1·3–3·5 kg DM/d). Each treatment lasted for 17 days and the samples were collected during the last 7 days of each period. For Expt 2 the animals were fed the same oat hay as in Expt 1 for 3 weeks at a level equivalent to the estimated energy maintenance requirement (M) (1·5–2·2 kg DM/d). The intake was then reduced to 0·6 M on day 1, 0·3 M on day 2 and zero from day 3 until day 10. The animals were re-fed in the reverse order for 3 days. Of the PD, only allantoin and uric acid were present in the urine. The proportions of allantoin and uric acid were 0·86 and 0·14 respectively for both experiments. There was no response of creatinine and nitrogen excretions to feed intake. The rates of PD excretion per kg digestible organic matter (DOM) or digestible dry matter (DDM) were 13·5 and 13·6 mmol respectively. As expected, urinary PD excretion increased significantly (P