ALBERT

Description: Dissolution rates of small forsterite spheres in superheated melts of basalt, andesite and rhyolite composition have been measured at 1300°C, atmospheric pressure. The rate is constant (83 µm hr−1) in the basalt, regardless of run duration. In the andesite the initial dissolution rate is 200µm hr−1, followed by a decrease to a constant value of 16µmhr−1 in 2–3 hours. Dissolution rate in the rhyolite decreases from an initial value of 1.7 to 〈0.1 µmhr−1 over 280 hours and never reaches a constant rate. Once the rate of dissolution has become constant, the film of contaminated melt that forms in melt about a crystal does not thicken with time, indicating attainment of a steady-state condition. Steady state is attributed to natural convection arising from the difference in density between the film of contaminated melt surrounding a crystal and that beyond. The density difference is approximately 2% of the density of the rock melt.

Description: The deep eastern Arctic basin between the Lomonosov Ridge and the Eurasian continental margin differs from other ocean basins in the very slow spreading of its floor and unusual depositional environment under perennial sea-ice cover. The recent expedition ARK IV/3 of RV Polar stern for the first time made geoscientific investigations from the northern margin of the Barents Sea north to the Nansen-Gakkel Ridge. Much deeper than most other mid-ocean ridges, this ridge is poorly-surveyed, but has a central valley which in places is deeper than 5.5 km, 1–1.5 km below the basin floors on either side. Heat flow in the central part of the valley is very rapid; both basement rocks and overlying sediments showed unexpectedly the influence of intense and long-term hydrothermal activity. The sediments on the northern and southern flanks of the ridge are slightly calcareous pelagic mud layers alternating with carbonate-free horizons, where up to 40% of the sedimentary section is soft mud clasts. Similar mud aggregates were observed on the surface of the multi-year sea ice, appearing to represent a special type of sediment transport by sea ice in the Transpolar Drift. In contrast to the western Arctic, Fram Strait and the Norwegian-Greenland Sea, gravel is rarely found in sediment cores. Recovered cores indicate that icebergs and sea ice carrying coarse sediment seldom rafted detritus to the study area during the last approximately 300,000 years.

Description: This article describes the natural history of a large colony of emperor penguins Aptenodytes for steri , its size, dispersal pattern of chicks, and associations with other bird and mammal species. A mid-season count of 19,364 chicks indicated that about 20–25,000 breeding pairs had been present in June and July. The colony was fragmented into several sub-groups which showed different mean sizes of chicks and survival to fledging. Other species observed included leopard seals Hydrurga leptonyx , the only major predators, which preyed heavily on both adults and fledging chicks. Fledgelings left the colony over a period of about 10 days; departure was an active process in which the chicks walked to the ice edge and dispersed in groups, swimming consistently southward. At this time they were still in about 60% down and weighed about 10 kg, having lost some 30% of the heaviest mass achieved during parental feeding.

Description: Within the Western Ross Sea, there are six emperor penguin colonies of widely different size that occur exclusively on sea ice. In 1990 a survey of all six sites, two by close overflights and four from the ground, showed that the breeding habitats were highly variable. The most important physical characteristics of these habitats appear to be stable fast ice, nearby open water, access to fresh snow, and shelter from the wind.

Description: Light absorbed by the photoreceptor membranes of the retina is transformed by a series of biochemical steps into changes in receptor potential. The visual pigment, rhodopsin, and associated enzymes which relay the visual signal belong to a widespread family of receptorenzyme complexes which transmit sensory, hormonal and neurotransmitter signals into cells. In the invertebrate visual system, rhodopsin activates enzymes which increase cytoplasmic calcium. The highly specialised and structurally ordered photoreceptors of the squid retina provide a favourable system for studying the mechanisms of these enzyme pathways.

Description: Arm autotomy was induced in a living specimen of Metacrinus rotundus (Echinodermata: Crinoidea). An arm was autotomized at a ligamentary articulation known as a cryptosyzygy, following incision by scissors distal to the break point. Although sessile stalked crinoids cannot entirely escape from a predatory attack by arm autotomy and they do not have an active defense, arm autotomy at cryptosyzygies reduces damage and arm loss by effective distribution, and by minimizing trauma and facilitating subsequent regeneration. The paradigmatic distribution of cryptosyzygies in which arm loss is set at a minimum, compared with the actual distribution, shows that these two patterns are similar and that actual specimens successfully reduce arm loss by the effective distribution of cryptosyzygies. The crinoid branching pattern also affects arm loss, and two different paradigms are discussed: anti-predatory and harvesting. Arm branching patterns of various isocrinids have tended toward the anti-predatory configuration from the Jurassic to the Recent, suggesting that the isocrinids have coped with increased predation. Shallow-water comatulids generally adopt the anti-predatory paradigm in their branching pattern, whereas many deep-water, stalked crinoids adopt a harvesting paradigm, reflecting that shallow-water comatulids receive more predatory attacks than do deep-water crinoids.

Description: The functional morphology of the buccal mass of 23 species of cephalopod (Octopoda, 4 species; Teuthoidea, 17; Sepioidea, 2) was investigated by gross dissection, histology and observations on fresh preparations. Cephalopod beaks lack a joint or articulation point. The jaws slide and rotate around an area rather than a fixed point. During closing the superior mandibular muscle (SMM) provides the force of a bite and the largest movement vector, whilst the inferior mandibular muscle (IMM) acts to retract the upper beak, causing shearing action. Dorsal portions of the lateral mandibular muscles (LMM) flex the upper beak walls outwards, probably to accomodate the backwards sweep of the radula and buccal palps during closing. To open the beaks, the ventral portions to the lateral mandibular muscles pull the rear lateral walls of the two beaks towards each other, moving the lower beak back relative to the upper. The buccal mass weighs more in decapods (0.65-4.34% of body weight) than octopods (0.49-0.77%). The weight difference is mainly accounted for by the size of the superior mandibular muscle. Beak shape and muscle volume are related. Increasing the size of the upper beak hood and lateral wall area results in larger SMM and LMM respectively Increasing hood size in the lower beak increases IMM size, and altering the angle by which the wings meet the lateral wall changes the volume of the SMM and LMM. To accomodate the decapod pointed upper rostrum, the lateral walls of the lower beak have shortened in length, whilst increasing in breadth and surface relief to maintain the area available for muscle insertion. In species with a lateral wall ridge or fold (e.g. Onychoteuthis) this may mark the insertion point of the LMM.

Description: The influence of photoperiodicity on hatching of Loligo forbesi and Loligo vulgaris embryos was investigated under different experimental light-dark (LD) conditions. The transition from light to dark stimulated hatching and functions as a "Zeitgeber" or synchronizer. Independent of the timing and duration of the dark period most embryos hatched soon after termination of the light period. Embyros which had developed in constant light, showed no hatching rhythm at all. If these embryos were exposed to a dark shock most embryos hatched soon after the onset of darkness. A twilight shock, in which the light was reduced by 50% (i.e. 50 µE s -1m-²), could not simulate hatching. Embryos which were kept from stage X on in an artifically controlled LD cycle, preferentially hatch in a period which coincides with the period at which darkness usually occured when placed in constant illumination from stage XX onwards.

Description: Maturation in the onychoteuthid squid Moroteuthis ingens was found to be irreversible, with death following shortly after sexual maturation and spawning. Both males and females were found with spent gonads. The ovary reaches very large sizes in mature females and probably prevends feeding by constricting the caecum. There was also a marked difference in the tissue with an inelastic, gelantinous appearence. Histological examination of the mantle wall revealed that the tissue breakdown was due to a drastic histolysis of muscle tissue and, to a lesser extent, collagen fibres. Mature males also showed some tissue breakdown and loss of muscle fibres but this was not as dramatic as in the females. These features are considered in relation to processes contributing to terminal maturation in M. ingens.

Description: A correlation of the Campanian–Maastrichtian boundary is attempted using foraminiferal and nannoplankton data from two areas: the eastern North Atlantic and northwestern Germany. The Boreal benthic and Tethyan planktonic foraminiferal zonation schemes are applied to Site 548A, where both foraminiferal groups occur frequently. A direct comparison of both biozonations reveals that the base of the Maastrichtian, according to planktonic foraminifers, has to be placed in the Upper Campanian of the Boreal benthic foraminiferal biozonation, which concurs with the nannoplankton results. The Tethyan Middle and Upper Maastrichtian are probably equivalent to the Upper Maastrichtian in the Boreal sense. The bases of the Maastrichtian substages are thus diachronous between the Boreal and Tethyan realms. Palaeotemperatures (which were estimated using the oxygen isotopic composition of the Goban Spur chalks) indicate, in combination with palaeowind directions, that the faunal and floral distribution pattern recorded is the result of a stable, warm water outflow from the northwest European epicontinental seas through the Channel area to the Celtic Shelf sea and Goban Spur. This mechanism appears to have been a dominant separating factor of the Boreal and Tethyan bioprovinces on the western European Shelf.

Description: The groundmass of andesitic dykes at Sezaki, southwest Japan, has trachytic texture and contains microscopic shear zones. The shear zones comprise a conjugate pair formed by flattening of the solidifying dyke rock, probably caused by the magma pressure of the still molten part of the dyke. This pressure shortened the solidifying rock perpendicular to the dyke margins and caused it to extrude parallel to the magma flow direction. The groundmass shears indicate that locally the magma flowed 60° upward in the dykes. It is concluded that while groundmass shears are a useful indicator of flow direction in dykes, phenocryst alignment in dykes is strongly influenced by magma-pressure flattening and thus may be a poor indicator of flow direction.

Description: The diet of King Penguins Aptenodytes patagonica at Possession Island was studied in February 1989 by analysis of 20 stomach contents collected from adult birds, just before they fed their single chicks. The mean mass of the samples was 1.84 kg, equivalent to 15% of the unladen adult body-mass. Fish accounted for 99.8% and squids for 0.2% by mass. The main prey were mesopelagic myctophid fish which live in dense shoals and perform a daily vertical migration. Subadults/adults of Electrona carlsbergi and juveniles and subadults/adults of Krefftichthys anderssoni represented 73.7 and 13.4% of the diet by mass, respectively. No difference was found in the diet of male and female King Penguins. Comparison of individual samples suggests that these birds catch a large quantity of fish from only a limited number of shoals. The mean caloric content of the food was 7 kJ/g wet mass. The total energy requirement of each chick during its initial period of growth was estimated to amount to 328,000 kJ, equivalent to 55 kg of food. The rate at which energy was delivered to the chick was calculated to be 50 W during this period.

Description: Outer membrane (OM), cytoplasmic membrane (CM) and intracytoplasmic membranes (ICM) from the halophilic phototrophic purple sulphur bacterium Ectothiorhodospira mobilis 9903 were purified and characterized. The three membrane fractions were significantly different in regard to protein profiles on SDS-PAGE, and to the composition of amino acids, fatty acids and lipids. The presence of lipoproteins, the occurrence of lyso-phosphatidyl-ethanolamine and an increased content of saturated and short-chain fatty acids are characteristic properties of the OM. CM and ICM fractions are different on the basis of buoyant density, of protein profiles and amino acid composition, and due to the presence of succinate dehydrogenase activity in CM. In addition, CM and ICM showed significant differences in pigment content and absorption spectra.

Description: Three specimens of the rare giant squid (genus Architeuthis, 18 nominal species) are described from the waters around southern Africa. Consideration of their internal and external morphology, including beaks, radula, statocyst and statolith give no reason to suppose that the three specimens do not pertain to the same species. A standardized data collection form and a summary of existing morphometric data are presented for the first time and the morphology and systematics of the genus are discussed. The value of various systematic characters is examined and certain guidelines are proposed, with an appeal for a more scientific approach to Architeuthis studies. Features of the internal anatomy, including the female reproductive tract, are described and illustrated and the beak dimensions are used to evaluate existing regression formulae for beak and body parameters. The statocyst and statolith are described and illustrated in full, the statolith for the first time, and are compared with those of other squids. The value of recent data from trawl-caught specimens, as well as indirect evidence, is cited to replace speculation in discussions on distribution and habitat. Our specimens suggest that the waters off the south-western coast of Africa may be one of the natural habitats of Architeurhis, during both feeding and spawning periods of the life cycle.

Description: Benthoctopus piscatorum appears to be a multiple spawner. In the ovary of one specimen about 50 eggs were found at various stages of development.

Description: In Loligo forbesi Steenstrup, the female reproductive system consists of the ovary and accessory reproductive organs which include the oviducal gland, the nidamental gland, the accessory nidamental gland and seminal receptacle. Histological studies were made on the accessory reproductive organs of female L. forbesi. The various changes observed during maturation are described and the functional significance discussed. The secretions produced by the oviducal gland and nidamental gland apparently form the egg coats. The seminal receptacle serves to store spermatozoa after mating. The function of the accessory nidamental gland is unknown.

Description: Whether the study of external morphology can increase our knowledge of shallow-water octopuses and their biology is tested by multivariate morphometric analyses. The extent of morphological variation among 20 species from the Atlantic and eastern tropical Pacific Oceans is explored, and four hypotheses are addressed: (1) that octopuses show secondary sexual dimorphism; (2) that remote, ostensibly conspecific populations are morphologically distinct; (3) that morphometric characters contribute to resource partitioning among sympatric species; and (4) that morphology is predictably associated with habitat. Results of principal components analysis show most species to be very similar in shape. Arm length contributes most shape variation; other characters contribute little. Principal components and size-free discriminant analyses refute each of the hypotheses considered. The morphological similarity of isolated Atlantic populations may be a symplesiomorphy, but information on the planktonic phase of the octopus lire cycle supports the possibility of dispersal across the Atlantic Ocean. Size, not directly treated here, may significantly affect some aspects of octopus biology. Increased female size may evolve due to selection for increased fecundity and intraspecific niche partitioning. Size may also indicate interaction within and among species. Characters of shape, deemed essential to species descriptions, do not appear to be able to identify most specimens, nor to increase our understanding of octopus biology.

Description: The stomach contents of 1522 adult seals (Arctocephalus pusillus pusillus) and 673 pups, which were collected around the West and South Coasts of Southern Africa, were examined and cephalopod prey identified. About 20 cephalopod species (some identifications were uncertain) were found in the stomachs, of which only six were significant: Loligo vulgaris reynaudii. Sepia spp., Octopus sp., Todaropsis eblanae, Todarodes angolensis and Ocythoe tuberculata. This list suggests that A. pusillus feeds exclusively on the continental shelf, frequently on or near the bottom. Among adults, cephalopod prey varied in importance around the coast. It was most important on the South Coast of South Africa where it comprised 35.0% of the weight of all prey, of which L. v. veynaudii constituted about 88%. On the West Coast cephalopods comprised about 26.6% by weight of all prey, and Octopus sp. predominated, whereas in Namibian waters cephalopods were least important, constituting only 3.4% by weight of all prey, with Todarodes angolensis and Octopus sp. being most significant. Among pups (aged 8–10 months) in the Lüderitz (Namibia) area, cephalopods were about 16.9% of prey by weight, of which Ocythoe tuberculata, small Sepia spp. and Octopoda were most important.

Description: The structure of the reproductive systems of mature males and females of the nektonic, oceanic squid Thysanoteuthis rhombus are described. The main peculiarities of the female system are relatively low capacity oviducts, set in a tight spiral, and hypertrophically developed oviducal glands with a very large second section. The male reproductive system is characterized by a long, narrow Needham's sac containing 10–15 large spermatophores 80–100 mm in length. The mesentery supporting the gonad, and protruding into it dorsally, is a characteristic feature in both sexes. The hectocotylus structure differs markedly from that in other squids and resembles that of sepiids. The reproductive system of T. rhombus possesses primitive features (pattern of gonad attachment and hectocotylus) but mostly secondary characters (small oviducts, very large oviducal glands and ovary). The complex morpho-ecological adaptations of T. rhombus are reflected in the distinctive features of the reproductive system.

Description: Seasonal variation in the diet of common seals (Phoca vitulina) in the Moray Firth, north-east Scotland, was determined from analysis of faecal samples collected at haul-out sites during each month of 1988. Data on diet of common seals in 1987 are also presented. Limitations of the methods available for quantification of diet are discussed. Although some of the observed variation in diet from month to month may reflect changes in the sampling regime, a clear seasonal pattern was apparent, with clupeids predominating in the winter and sandeels in the summer. The trends observed are consistent with opportunistic feeding on the most abundant Prey.

Description: The cephalopod fauna collected during six surveys carried out in the bathyal basin of the north-western Mediterranean is discussed. Samples were taken at depths mainly between 1000 and 2000 m. Ten species were identified. Bathypolypus sponsalis and Neorossia caroli were the commonest species. Small individuals of both these species occurred at greater depths than did larger individuals, suggesting up-slope ontogenetic migration. The depth ranges recorded for all species collected are discussed and compared to the results of previous studies found in the literature.

Description: The upper bathyal sea-pen Kophobelemnon stelliferum extends to depths of about 1600 m in the Porcupine Seabight, to the south-west of Ireland, but is rare below about 1150 m. Photographic data suggest that the species attains numerical abundances of more than 2 m−2and a wet weight biomass of at least 4 g m−2. The highest densities, however, do not necessarily correspond to the highest biomass values since there is a clear depth-related change in population structure. The largest sea-pens are restricted to the deeper parts of the bathymetric range of the species. There is also a marked change in the growth form at a total colony length of about 250 mm, with larger colonies having relatively more polyps than smaller ones. The sexes are separate in Kophobelemnon stelliferum and the sex ratio of colonies is about 1:1. The maximum oocyte diameter is about 800 μm, but there is no evidence of seasonal reproduction by this pennatulid in the Porcupine Seabight.

Description: The free-running locomotor activity rhythms of freshly-captured swimming crabs Liocarcinus holsatus and L. depurator have been studied in constant conditions in the laboratory. L. holsatus captured in the intertidal zone of sandy beaches showed strong circatidal activity rhythms with maximum activity at high tide. L. holsatus captured in the sublittoral zone had a clear tendency to show circadian activity rhythms with highest activity during day-time hours. L. depurator occurred only sublittorally and showed circadian activity patterns with highest activity during the night. Exposure to hydrostatic pressure cycles of tidal amplitude and periodicity, entrained strong circatidal activity rhythms in previously arhythmic L. holsatus. This activity pattern also showed a marked circadian component. Exposure to the same regime entrained a circadian rhythm, but not a circatidal rhythm in L. depurator. In the sublittoral zone L. depurator is active mainly during the night, whereas L. holsatus, is active mainly during the day. This may constitute a behavioural mechanism for minimizing competitive interactions between these two sympatric crabs

Description: Serological methods for prey identification have been applied to detection of residues ofsandeel (Ammodytidae) protein in faeces of common seals (Phoca vitulina) and grey seals(Halichoerus grypus) from the Moray Firth, north-east Scotland. Antisera raised to muscleprotein from Ammodytes marinus were evaluated by testing their reactions with proteinextracts made from a range of North Sea fish species and protein residues in in vitro digestates,seal digestive tracts and seal faeces. It was concluded that, using fused rocketimmuno-electrophoresis, linkage of precipitin peaks from unknown samples with peaksfrom standard sandeel extract was a reliable indicator of the presence of sandeel in theunknown sample. Seasonal variation in the incidence of sandeels in common seal diet in theMoray Firth was examined by identifying otoliths, bones, and proteins, and all threemethods indicated that sandeels occurred in the majority of samples tested in the summer,but were less important during the winter. Proteins were detected in fewer samples thanotoliths, particularly in February and March. Possible reasons for this difference arediscussed. Serological identification of sandeel proteins is potentially applicable to dietarystudies on all marine predators.

Description: Age and growth were estimated in the European squid, Loligo vulgaris, by examining growth increments in the statoliths of 203 specimens collected from off the French Mediterranean coast. Length and increment data were analyzed assuming that the increments were formed daily. The relationships between age and length showed that: growth rate varied considerably among individuals; growth was double exponential; the squids grew on average to 240 mm ML at 240 d from hatching, with a maximum of 350 mm at 240 d; the life span is probably about one year.

Description: Some of the limits to the use of serology to identify prey species in the digestive tracts of cephalopods have been evaluated. Cuttlefish, Sepia officinalis, were given meals of krill slurry (Euphausia superba). Protein extracts of contents from four regions of the digestive tract, stomach, caecum, digestive gland and intestine, were tested for prey antigenicity. Digestion times (loss of antigenicity) ranged from 1 to 8 h depending on sampling site. Stomach and caecum emptied rapidly, but meal antigenicity persisted longer in the digestive gland. The Sepia experiments provide a basis for interpretation of results from natural predation by cephalopods).

Description: The stomachsof 23 striped dolphins (Stenella coeruleoalba Meyen, 1833, Cetacea), stranded along the Ligurian coast (western Mediterranean Sea), contained 32 species of cephalopods, crustaceans and fishes, totalling an estimated 2,723 prey specimens representing about 36 kg in weight. Cephalopods and bony fishes were equally important in the diet (50%). Todarodes sagittatus (34.5%) and Micromesistius poutassou (25.9%) were found to be the most important food species. Other species belonging to six cephalopod families, three crustacean families and nine bony fish families, contributed to the diet with variable numbers, weights, and occurrences, demonstrating the opportunistic character of striped dolphin feeding.

Description: Thirty-four adult individuals of Loligo forbesi (males and females with dorsal mantle lengths from 27–77 cm) were observed in captivity in a 3 m diameter closed sea-water system on Faial Island, Azores. Squids were caught by jigging and were fed with horse mackerel (Trachurus picturatus) and chub mackerel (Scomber japonicus), either alive or dead. The maximum survival was 73 days. Feeding behaviour was dependent upon both the size of prey and its state of preservation;e.g. the squid would eat the head of small fish (〈 about 15 cm), reject the head of medium-sized fish (about 15–35 cm) and would eat only the dorsal part of a big fish (〉 about 35 cm) or a fish poorly preserved. Seventeen chromatic, 9 postural and 6 movement components of body patterns were observed and described. Conspecific interactions considered to be aggression and dominance were observed among males; no such interactions occurred when one male and two females were kept together. Body patterns in relation to relaxation, stress, shock, feeding, locomotion and aggression are also described.

Description: The presence of Stoloteuthis leucoptera in the Mediterranean is recorded on the basis of three specimens, including an adult male, caught by IKMT and by commercial otter-trawl in the Ligurian Sea. The hypothesis of a recent immigration is discussed. The list of Mediterranean cephalopods (Mangold Wirz, 1963; Torchio, 1968; Bello, 1986; Mangold & Boletzsky, 1987) includes the Sepiolidae of the subfamily Heteroteuthinae, whose members are supposed to be pelagic throughout their life cycle. Mangold Wirz (1963) recognizes in the Mediterranean fauna the unique species Heteroteuthis dispar, the other authors include H. atlantis Voss, which Voss himself (1955) reported at Messina. To this group may now be added Stoloteuthis leucoptera (Verrill, 1878) a species until now recorded in limited Atlantic areas. Verrill (1881) wrote “This species is an exceedingly beautiful one, when living, owing to the elegance and brilliancy of its colours and the gracefulness of its movements. In swimming it moves its fins in a manner analogous to the motion of the wings of a butterfly.”

Description: Samples of two loliginid squids Alloteuthis africana and A. subulata were collected from the continental shelf off the west Sahara in August-September 1987. Statoliths were taken from 124 specimens and processed using statolith ageing techniques. Statoliths of both species were very similar in shape. In the ground statolith, growth increments were examined and grouped into four growth zones distinguished mainly by the width of the increments. Age of adult mature males of both species did not exceed eight months, that of females six months. Alloteuthis africana grew faster than A. subulata in weight and, particularly, in length. At age 180 d the mantle of A. africana was twice as long and the body weight 1·2–1·5 times as large. Both species matured over a wide range of sizes and ages (from 120 to 180 d). The life span of A. africana and A. subulata hatching between January and May on the west Saharan shelf is about six months, much shorter than that of A. subulata in its northern temperate range.

Description: Samples of female lllex argentinus were taken from the catch of a Japanese squid jigging vessel on the Patagonian Shelf during March 1986. Morphometrics of the somatic and reproductive organ systems and the histological structure of the mantle in relation to maturation were examined. The data suggest that growth and maturation occur simultaneously during most of the time that lllex argentinus females are on the feeding grounds. In a squid of a ‘standard’ mantle length the whole body mass increases relative to mantle length during maturation and growth of the reproductive organs. This is accompanied by a small but significant decrease in the relative mass of the mantle, head and viscera whilst the mass of the digestive gland remains constant. Although mantle mass of a ‘standard’ female squid decreases relative to mantle length with maturity this is not associated with degeneration of the mantle muscles. Energy and nutrient resources for maturation are apparently derived from the squid's food, not from reserves, and during the course of maturation there is an increasing shift of emphasis from somatic growth to production of gonad and accessory reproductive organs.

Description: The timing of spawning and recruitment in the squid Loligo forbesi in Scottish waters is described on the basis of data from three sources: monthly samples of squid caught by commercial trawls (1986–1988), egg masses found by fishermen (1987–1991), and statistical data on animals caught by research trawls (1978–1987). Spawning females were present in samples from December to June, with peak spawning occurring in March. Most records of egg masses were from these months, but eggs were also found in August and September. These results suggest that there is an extended spawning season. Small squid (≤100 mm dorsal mantle length) were rarely present in commercial samples, but were recorded in research samples almost all year round. Thus there appears to be more or less continuous recruitment into the catchable population. The results of the present study are consistent with published data from other parts of the geographic range in that there is a regular seasonal peak in spawning, and spawning adults disappear from the population in summer. Further interpretation of the life-cycle of this species is not justified on the basis of current knowledge, and more information is needed on migrations, geographical variation, and lifespan in Loligo forbesi.

Description: Cephalopod remains from the stomachs of a Risso's dolphin (Grampus griseus Cuvier, 1812, Cetacea) entangled in a fishing net off the Ligurian coast (central Mediterranean Sea) include squids Ancistroteuthis lichtensteini, Histioteuthis bonnellii, H. reversa and Todarodes sagittatus and the sepiolid Heteroteuthis dispar. All these cephalopods live in oceanic water including water over the steep continental slope where Risso's dolphin is frequently sighted. Histioteuthis reversa contributed 78% of the cephalopods by number, 81% of the wet weight and 73% of the dry weight and calorific value. The total calorific value of the cephalopods represented by lower beaks was 17,300 kj.

Description: The stomach contents of 235 specimens of the squid Sthenoteuthis oualaniensis (4·3–36·5 cm mantle length, ML) were examined. A detailed list of 60 species of prey, comprising young and adult squid, is given together with their frequency of occurrence and proportional contribution. The size and number of each food item was investigated. Three ontogenetic size-groups of S. oualaniensis were distinguished: I, fry and young (4–10 cm ML), micronektonic epipelagic plankton-eaters; II, transient critical size group (10–15 cm ML), converting from feeding on planktonic crustaceans and fish larvae to myctophid fishes; III, medium-sized (adult) nyctoepipelagic nektonic predators (15–36·5 cm ML), feeding primarily on myctophids and secondarily on squid. Myctophids (genera Symbolophorus, Myctophum and Hygophum) were the most abundant prey in the diet of adult S. oualaniensis from different parts of its distribution.

Description: Samples of male Illex argentinus were taken from the catch of a Japanese squid jigging vessel on the Patagonian Shelf during March 1986 and an analysis was carried out on the morphometrics of the somatic and reproductive organ systems in relation to maturation. The data show that growth and maturation occurred simultaneously during most of the time that Illex argentinus males were on the feeding grounds over the southern Patagonian Shelf. In a squid of a ‘standard’ mantle length the whole body mass increased relative to mantle length during maturation and this could be attributed to the increase in mass of the reproductive and accessory reproductive organs. During maturation the mantle and digestive gland mass showed no significant change relative to mantle length. The mass of the head increased and the mass of the viscera decreased relative to mantle length. In male Illex argentinus, as in the female, the energy and nutrient resources for maturation are derived from the squid's food and during the course of maturation there is an increasing shift of emphasis from somatic growth to production of gonad and accessory reproductive organs. The proportional investment of body mass in reproductive and accessory reproductive organs predicted for a fully mature male Illex argentinus was less than half that of the female.

Description: During a study based on catches taken in the northern North Sea by selected Scottish fishing boats during 1985–1992, large numbers of the normally rare short-fin squid, Todaropsis eblanae (Cephalopoda: Ommastrephidae), were recorded in 1987 and 1990. Our findings, supported by data obtained from plankton/young fish surveys in 1988 and 1989, suggest that in northern waters Todaropsis eblanae generally mates and spawns during late summer and early autumn (June-November). Successful hatching events appear to occur during October-March, producing juvenile (stage I) squid in the early part of the year (January-June). Estimations of maximum male reproductive output and female fecundity were up to 130 spermatophores and ~28,000 eggs per individual, respectively.

Description: Statoliths of Loligo gahi were sampled in the fishery region 45–47°S on the Patagonian shelf during September 1989. Peculiarities of the growth zones in the ground statoliths of adults are described. Maximum age of large maturing and mature females (130–160 mm of mantle length, ML) was estimated to be 325–345 d, that of large mature males (250–290 mm ML) ranged from 360 to 396 d. The squid Loligo gahi d'Orbigny, 1835, occurs in temperate shelf and upper slope waters of the Atlantic and Pacific coasts of South America and is caught commercially by the international fleet in the southern part of the Patagonian shelf within the Falkland Islands Interim Conservation Zone (FICZ) (Roper et al., 1984; Csirke, 1987). Occasionally, dense shoals of L. gahi appear in the fishery region 45–47°S off the Exclusive Economic Zone of Argentina (EEZA) and have been caught in significant numbers by trawlers at depths of 120–150 m in September-October (Chesheva, 1990). Loligo gahi is a medium sized loliginid; in Falkland waters males attain 350 mm ML, females 210 mm ML (Hatfield, 1991), while in the fishery region 45–47°S maximum size is 260 mm and 160 mm, respectively (Chesheva, 1990). Patterson (1988) revealed two Falkland spawning stocks of L. gahi of unclear status, spring-spawners and autumn-spawners (austral seasons) and pointed out that the life span of squid of each stock lasted ~1 y. Recently Hatfield (1991) used statoliths to elucidate Patterson's (1988) estimations of age and growth of Falkland stocks of L. gahi and confirmed the 1-y duration of L. gahi's life span.

Description: The presence of several typically northern hemisphere foraminiferal species in the upper Miocene and Recent deposits of the Río de la Plata, Paraná river and adjacent areas, and their absence along the Brazilian coast strongly suggests that in the late Miocene the Caribbean and the Southern Atlantic were linked by a marine connection. This connection was first envisioned by H. von Ihering, who called it the ‘Arm of the Tethys’. It was represented by a very broad strait which ran meridionally across the South American continent. In the Pliocene it shrank considerably and its salinity was lowered by Andean river runoff. At the end of the Pliocene - beginning of the Pleistocene it gave rise to the Paraná river, a series of brackish water lakes and lagoons, and the Río de la Plata. □Foraminifera, arm of the Tethys, South America.

Description: Organic-rich shales sampled from the aples Beach section of the Monterey Formation. Califomia. contain authigenic phosphorite phases which apparently formed within bacterial mat laminations. Light and scanning electron microscope observations show that the fossilized remains ofthese mats are similar to communities of filamentous sulfur-oxidizing bacteria found today at slope-depth, sediment, oxic anoxic interfaces. The crystal size and habits ofthe phosphatic minerals resemble contemporaf) prec1pitates in contact with pore water solutions. Mineralogical and chemical analyses ofthe most phosphatic layers show that some are a nearly pure carbonate fluorapatite (or francolite). These la)ers are lightly colored and are sandwiched between black siliceous muds with 〈0.2% CaCOJ . and organic carbon contents that excecd 25%. In othcr phosphatic intcrvals of thc Naplcs Beach section. ca leite is a major diluent oflight and dark layers. The C:P ratio of a single modern bacterial mat sample shows that these benthic communities are more enriched in phosphorus than planktonic organic matter. Thus, the generally believed premise that authigenic phosphorite formation results from organic matter diagenesis in near-surface anoxic sediments is hcre modified by the contention that such transformations are more highly favored in sediments supporting massive microbial communities.

Description: The data presented provides new information on the distribution of Antarctic squids and on the summer diet of the emperor penguins. The diet of 58 adult emperor penguins (Aptenodytes forsteri) on the fast ice of the Drescher Inlet, Vestkapp Ice Shelf (72°52′S, 19°25′W) in the eastern Weddell Sea was investigated. Prey consisted principally of squid, fish, krill, amphipods and isopods. Squids were identified by the lower beaks and allometric equations were used to estimate the squid biomass represented. Beaks occurred in 93% of the stomach samples. Each sample contained a mean of 27 beaks (range 1–206). Ninety-two percent of the squids could be identified by the lower beaks and belonged to four families (Onychoteuthidae, Psychroteuthidae, Neoteuthidae and Gonatidae). The most abundant squid was Psychroteuthis glacialis which occurred in 52 samples with lower rostral lengths (LRL) ranging from 1.4–7.2 mm. Forty-five samples contained Alluroteuthis antarcticus (LRL range 1.8–5.8 mm), 17 Kondakovia longimana (LRL range 4–12.1 mm), and four Gonatus antarcticus (LRL range 4.1–6.1 mm). In terms of biomass K. longimana was the most important species taken by the penguins comprising 50% of total estimated squid wet mass (245348 g) in 1990 and 48% in 1992 (154873 g). However, if only fresh beaks were considered for estimations of squid consumption, i.e. beaks that have been accumulated for not longer than 5–6 days in the stomachs, squid diet was of minor importance. Then total squid wet mass accounted for only 4809 g in 1990 and 5445 g in 1992 which implies that one penguin took c.30 g squid d−1 with P. glacialis and A. antarcticus being the most important by mass. The prey composition suggests that emperor penguins take squid at the steep slope regions of the eastern Weddell Sea.

Description: To study the impact of the gradient of primary production between the southern, central, and northern Red Sea, benthic metabolism and standing stocks were investigated in the axial trough between 17°N and 27°N. Data on sediment chloroplastic pigments, macrofauna and meiofauna abundance, particulate adenylate, protein and carbohydrate biomass, as well as electron transport activity of the sediment community give evidence for an enhanced benthic standing stock and activity in the southern Red Sea south of 18°N; this is related to the increased primary productivity in the area south 16°N. Despite a large primary production only small benthic standing stocks were found in all areas investigated; they are much smaller than in other parts of the world's deep oceans with comparable primary production. The low benthic biomass is caused by two factors: a large proportion of particulate organic carbon is remineralized in the water column, while only a relatively small amount sediments to the sea floor; nearly all sedimented particulate organic matter is respired by the benthic community of the deep Red Sea and only a minute proportion is used for the production of benthic biomass.

Description: Growth of the shallow-water gorgonian Lophogorgia ceratophyta was investigated in an infralittoral station located in La Spezia Gulf, Ligurian Sea. Mean annual height growth rate was estimated to be 2.57 cm · a-1. The fractal dimension of the colonies was found to gradually evolve in complexity, exhibiting a simpler branching pattern in younger specimens. The maintenance of a low, invariable ramification complexity as an optimal choice in managing relationships between water and the colony's living tissues is also discussed.

Description: The Stilbonemutinue (marine free-living nematodes) arc remarkable for cctosymbiotic bacteria, which cover the greatest part of their body in a highly ordered and species specific pattern. Using SEM we describe the main types of symbiotic cover and give evidence for the role of the bacteria in the nutrition of their host on the basis of stable carbon isotope ratios. In experimental systems the worms migrated repeatedly across a sulfide gradient during 12 h when sulfide concentrations were low, but stayed above the sulfide maximum at high concentrations. The migration across the chemocline exposes the symbionts alternately to reduced sulfur compounds and oxygen; this constitutes an alternative strategy to the ventilation/circulation systems in symbiotic macrofauna from sulfidic habitats

Description: We investigate the weakly nonlinear spatial evolution of helical disturbances of an axisymmetrical jet which are the analogue of three-dimensional disturbances, such as a single oblique wave (the wave vector is directed at an angle to the main flow velocity) in plane-parallel flows. It is shown that when a supercriticality is large enough, the perturbation amplitude A grows in the streamwise direction (along z) explosively: A ~ (z0—z)-5/2, though more slowly than in the case of essentially three-dimensional disturbances in the form of a pair of oblique waves (A ~ (z0 — z)-3; Goldstein & Choi 1989). The nonlinearity needed for such a growth, is due equally to the cylindricity of shear layer and to the spatial character of the evolution (in the temporal problem the ‘evolution’ contribution is absent). At a smaller supercriticality, the evolution equation has a non-local (integral in z) nonlinearity, unusual for the regime of a viscous critical layer. Scenarios of disturbance development for different levels of supercriticality are studied, with proper account taken of viscous broadening of the flow. © 1994, Cambridge University Press. All rights reserved.

Description: In an earlier paper (Davey 1978) the first author investigated the linear stability of flow in a straight pipe whose cross-section was an ellipse, of small ellipticity e, by regarding the flow as a perturbation of Poiseuille flow in a circular pipe. That paper contained some serious errors which we correct herein. We show analytically that for the most important mode n = 1, for which the circular problem has a double eigenvalue c0 as the ‘swirl’ can be in either direction, the ellipticity splits the double eigenvalue into two separate eigenvalues c0±e2c12, to leading order, when the cross-sectional area of the pipe is kept fixed. The imaginary part of c12 is non-zero and so the ellipticity always makes the flow less stable. This specific problem is generic to a much wider class of fluid dynamical problems which are made less stable when the symmetry group of the dynamical system is reduced from S1 to Z2. In the Appendix, P. G. Drazin describes simply the qualitative structure of this problem, and other problems with the same symmetries, without technical detail. © 1994, Cambridge University Press. All rights reserved.

Description: This paper describes a combined computational and experimental study of the flow between contra-rotating disks for — 1 ≤Γ≤0 and Reϕ= 105, where is the ratio of the speed of the slower disk to that of the faster one and Reϕ is the rotational Reynolds number of the faster disk. For Γ = 0, the rotor-stator case, laminar and turbulent computations and experimental measurements show that laminar Batchelor-type flow occurs: there is radial outflow in a boundary layer on the rotating disk, inflow on the stationary disk and a rotating core of fluid between. For Γ= — 1, the laminar computations produce Batchelor-type flow: there is radial outflow on both disks and inflow in a free shear layer in the mid-plane, on either side of which is a rotating core of fluid. The turbulent computations and the velocity measurements for Γ = — 1 show Stewartson-type flow: radial outflow occurs in laminar boundary layers on the disks and inflow occurs in a non-rotating turbulent core between the boundary layers. For intermediate values of Γ, transition from Batchelor-type flow to Stewartson-type flow is associated with a two-cell structure, the two-cells being separated by a streamline that stagnates on the slower disk; Batchelor-type flow occurs radially outward of the stagnation point and Stewartson-type flow radially inward. The turbulent computations are mainly in good agreement with the measured velocities for Γ = 0 and Γ= — 1, where either Batchelor-type flow or Stewartson-type flow occurs; there is less good agreement at intermediate values of Γ, particularly for F = —0.4 where the double transition of Batchelor-type flow to Stewartson-type flow and laminar to turbulent flow occurs in the two-cell structure. © 1994, Cambridge University Press. All rights reserved.

Description: Based on linear potential theory, the general three-dimensional problem of steady supersonic flow inside quasi-cylindrical ducts is formulated as an initial-boundary-value problem for the wave equation, whose general solution arises as an infinite double series of the Fourier-Bessel type. For a broad class of solutions including the general axisymmetric case, it is shown that the presence of a discontinuity in wall slope leads to a periodic singularity pattern associated with non-uniform convergence of the corresponding series solutions, which thus are unsuitable for direct numerical computation. This practical difficulty is overcome by extending a classical analytical method, viz. Kummer's series transformation. A variety of elementary flow fields is presented, whose complex cellular structure can be qualitatively explained by asymptotic laws governing the propagation of small perturbations on characteristic surfaces. © 1994, Cambridge University Press. All rights reserved.

Description: This paper determines the ray structure of a spinning acoustic mode propagating inside a semi-infinite circular cylindrical duct, and thereby determines the ray structure of the field radiated from the end of the duct. Inside the duct, but outside of a caustic cylindrical surface, the rays are piecewise linear helices; on striking the rim of the end-face of the duct, these rays produce ‘Keller cones’ of diffracted rays. The cones determine the structure of the radiated field: for example, no rays penetrate two cone-shaped far-field quiet zones centred on the duct axis; two rays pass through each point in a forward loud zone; and one ray passes through each point in a rearward loud zone. The two rays through each point in the forward loud zone interfere to produce an oscillatory directivity pattern. One quarter of the rays on each cone point back inside the duct and produce the reflected field. Thus the rim of the end-face of the duct acts as a ‘ring source’, in which the radiated and reflected fields have their origin. Every propagating duct mode determines a polar angle and an azimuthal angle; these are taken as parameters specifying the mode and are used to calculate the positions and angles of all the rays. The mathematical method on which the paper is based is Debye's approximation for the Bessel function which appears in the expression for the duct modes; the approximation shows also that the duct contains a region of smooth helical rays on which the field consists of inhomogeneous waves: this region is the inner cylinder, lying inside the annulus of piecewise linear helical rays. The results of the paper are very promising for the application of Keller's geometrical theory of diffraction to detailed calculations of the sound radiated from aeroengine ducts. An alternative description of the field, using Cargill's meridional rays, is summarized.

Description: The effects of thermal conditions on the patterns of two-dimensional Boussinesq convection are studied by numerical integration. The adopted thermal conditions are (i) the heat fluxes through both upper and lower boundaries are fixed, (ii) the same as (i) but with internal cooling, (iii) the temperature on the lower boundary and the heat flux through the upper boundary are fixed, (iv) the same as (iii) but with internal cooling, and (v) the temperatures on both upper and lower boundaries are fixed. The numerical integrations are performed with Ra = 104 and Pr = 1 over the region whose horizontal and vertical lengths are 8 and 1, respectively. The results confirm that convective cells with the larger horizontal sizes tend to form under the conditions where the temperature is not fixed on any boundaries. Regardless of the existence of internal cooling, one pair of cells spreading all over the region forms in the equilibrium states. On the other hand, three pairs of cells form and remain when the temperature on at least one boundary is fixed. The formation of single pairs of cells appearing under the fixed heat flux conditions shows different features with and without internal cooling. The difference emerges as the appearance of a phase change, whose existence can be suggested by the weak nonlinear equation derived by Chapman & Proctor (1980). © 1994, Cambridge University Press. All rights reserved.

Description: Gravity wave radiation by vortical flows in the /-plane shallow-water equations is investigated by direct nonlinear numerical simulation. The flows considered are initially parallel flows, consisting of a single strip in which the potential vorticity differs from the background value. The flows are unstable to the barotropic instability mechanism, and roll up into a train of vortices. During the subsequent evolution of the vortex train, gravity waves are radiated. In the limit of small Froude number, the gravity wave radiation is compared with that predicted by an appropriately modified version of the Lighthill theory of aerodynamic sound generation. It is found that the gravity wave field agrees well with that predicted by the theory, provided typical lengthscales of vortical motions are well within one deformation radius. It is found that the nutation time for vortices in the train increases rapidly with increasing Froude number in cases where the potential vorticity in the vortices is of the same sign as the background value, whereas the nutation time is almost independent of Froude number in cases where the potential vorticity in the vortices is zero or of opposite sign to the background. Consequently, in the former cases, the unsteadiness of the flow decreases with increasing Froude number, so the effect of the inertial cut-off frequency is increased, leading to an optimal Froude number for gravity wave radiation, above which the intensity of the radiated waves decreases as the Froude number is further increased. It is proposed that the existence of a finite range of interaction within the vortices, for flows with positive vortex potential vorticity, may account for the strong dependence of nutation time on Froude number in those cases. The interaction scale within the vortices becomes infinite in the limit of zero vortex potential vorticity, and so the arguments do not apply in those cases. © 1994, Cambridge University Press. All rights reserved.

Description: Oscillations of a tank at a near-resonant frequency have been shown to produce a response which changes from a ‘hard-spring’ to a ‘soft-spring’ response as the depth passes through a critical value. This paper investigates the transition region and it is shown, using a symbolic manipulator, that in fact the large-amplitude response is that of a soft spring on either side of this critical depth. © 1994, Cambridge University Press. All rights reserved.

Description: Premixed H2/02/N2 flames propagating in two-dimensional turbulence have been studied using direct numerical simulations (DNS: simulations in which all fluid and thermochemical scales are fully resolved). Simulations include realistic chemical kinetics and molecular transport over a range of equivalence ratios ©〉 {& — 0.35, 0.5, 0.7, 1.0, 1.3). The validity of the flamelet assumption for premixed turbulent flames is checked by comparing DNS data and results obtained for steady strained premixed flames with the same chemistry (flamelet ‘library’). This comparison shows that flamelet libraries overestimate the influence of stretch on flame structure. Results are also compared with earlier zero-chemistry (flame sheet) and one-step chemistry simulations. Consistent with the simpler models, the turbulent flame with realistic chemistry aligns preferentially with extensive strain rates in the tangent plane and flame curvature probability density functions are close to symmetric with near-zero means. For very lean flames it is also found that the local flame structure correlates with curvature as predicted by DNS based on simple chemistry. However, for richer flames, by contrast to simple-chemistry results with non-unity Lewis numbers (ratio of thermal to species diffusivity), local flame structure does not correlate with curvature but rather with tangential strain rate. Turbulent straining results in substantial thinning of the flame relative to the steady unstrained laminar case. Heat-release and H202contours remain thin and connected (‘flamelet-like’) while species including H-atom and OH are more diffuse. Peak OH concentration occurs well behind the peak heat-release zone when the flame temperature is high (of the order of 2800 K). For cooler and leaner flames (about 1600 K and for an equivalence ratio below 0.5) the OH radical is concentrated near the reaction zone and the maximum OH level provides an estimate of the local flamelet speed as assumed by Becker et al. (1990). © 1994, Cambridge University Press. All rights reserved.

Description: The modification of the mean and fluctuating characteristics of a flat-plate boundary layer subjected to nearly isotropic free stream turbulence (FST) is studied experimentally using hot-wire anemometry. The study is focussed on the region upstream of the transition onset, where the fluctuations inside the boundary layer are dominated by elongated flow structures which grow downstream both in amplitude and length. Their downstream development and scaling are investigated and the results are compared with those obtained by previous authors. This allows some conclusions about the parameters which are relevant for the modelling of the transition process. The mechanisms underlying the transition process and the relative importance of the Tollmien–Schlichting wave instability in this flow are treated in an accompanying paper (part 2 of the present report). © 1994, Cambridge University Press. All rights reserved.

Description: Linear birefringence measurements of dilute and semi-dilute polyisobutylene solutions following flow through a disordered fixed fibre bed of 2.47 % solids volume fraction provide both transient and steady measurements of chain deformation. Our results indicate that the flexible polyisobutylene polymers undergo a large conformation change, stretching in the direction of the average flow. This occurs even though the average flow in the bed is a plug flow which would not cause any polymer stretch by itself. The polymer stretch or conformation change increases with the number of chain interactions with bed fibres and ultimately reaches a steady-state value that can be correlated with the pore-size Deborah number (i.e. a characteristic polymer relaxation time divided by a characteristic flow time in the bed pore). Large changes in the polymer conformation are noted for values of the Deborah number, De 〉 5. In addition, the time to steady state scales with the characteristic flow time within a pore over a large range of Deborah numbers. The pressure drop across the fibre bed was also measured simultaneously with the birefringence measurement and was found to be directly proportional to the birefringence throughout the range of De investigated. Thus, we show empirically, for the first time, that chain elongation, which produces normal stress anisotropy within the fluid, is directly responsible for the increased flow resistance. These findings are then analysed in the light of recent theories for the response of polymer molecules in fixed bed flow fields (Shaqfeh & Koch 1992). It is shown that our results are consistent with the interpretation that these flows are stochastic strong flows, which create an apparent ‘coil-stretch’ transition. After extending the theory of Shaqfeh & Koch to account for the specifics in the experiments, including the bed geometry and statistics as well as the polydispersity of the polymer solutions, it is shown that the theory can predict most of the experimental results both qualitatively and quantitatively. © 1994, Cambridge University Press. All rights reserved.

Description: The existence of an organized streamwise vortical structure, which is superimposed on the well known coherent spanwise vorticity in nominally two-dimensional free shear layers, has been studied extensively. In the presence of stratification, however, buoyancy forces contribute to an additional mechanism for the generation of streamwise vorticity. As the spanwise vorticity layer rolls up and pulls high-density fluid above low-density fluid, a local instability results. The purpose of the current investigation is to force the three-dimensional instability in the stratified shear layer. In this manner, we experimentally observe the effect of buoyancy on the streamwise vortex tube evolution, the evolution of the buoyancy-induced instability, and the interaction between these two vortical structures. A simple numerical model is proposed which captures the relevant physics of the flow evolution. It is found that, depending on the location, streamwise vortices resulting from vortex stretching may be weakened or enhanced by the stratification. Buoyancy-induced vortex structures are shown to form where the unstable part of the interface is tilted by the streamwise vortex tubes. These vortices strengthen initially, then weaken downstream, the timescale for this process depending upon the degree of stratification. For initial Richardson numbers larger than about 0.03, the baroclinically weakened vortex tubes eventually disappear as the flow evolves downstream and the baroclinically generated vortices dominate the three-dimensional flow structure. © 1994, Cambridge University Press. All rights reserved.

Description: A flagellated, bottom-heavy micro-organism's swimming direction in a shear flow is determined from a balance between the gravitational and viscous torques (gyrotaxis). Hitherto, the cell has been assumed to be a spheroid and the flagella have been neglected. Here we use resistive-force theory to calculate both the magnitude and the direction of a biflagellate cell's swimming velocity and angular velocity relative to the fluid when there is an arbitrary linear flow far from the cell. We present an idealized model for the flagellar beat but, in calculating the velocity of the fluid relative to an element of a flagellum, the presence of the cell body is not neglected. Results are given for the case of a spherical cell body whose flagella beat in a vertical plane, when the ambient linear flow is in the same vertical plane. Results show that resistive-force theory can be used for organisms where the cell body has significant effect on the flow past the flagella and that the viscous torque on the flagella is a significant term in the torque balance equations. A model is presented for the calculation of a cell's velocity and angular velocity in a shear flow which is valid up to high magnitudes of rate of strain or vorticity. The main application of the results will be to modify a recent continuum model for suspensions of gyrotactic micro-organisms (Pedley & Kessler 1990). © 1994, Cambridge University Press. All rights reserved.

Description: The natural occurrence of Tollmien-Schlichting (TS) waves has so far only been observed in boundary layers subjected to moderate levels of free stream turbulence (Tu 〈 1%), owing to the difficulty in detecting small-amplitude waves in highly perturbed boundary layers. By introducing controlled oscillations with a vibrating ribbon, it is possible to study small-amplitude waves using phase-selective filtering techniques. In the present work, the effect of TS-waves on the transition is studied at Tu — 1.5%. It is demonstrated that TS-waves can exist and develop in a similar way as in an undisturbed boundary layer. It is also found that TS-waves with quite small amplitudes are involved in nonlinear interactions which lead to a regeneration of TS-waves in the whole unstable frequency band. This results in a significant increase in the number of turbulent spots, which promote the onset of turbulence. © 1994, Cambridge University Press. All rights reserved.

Description: We study the average mobilities and long-time self-diffusion coefficients of a suspension of bimodally distributed spherical particles. Stokesian dynamics is used to calculate the particle trajectories for a monolayer of bimodal-sized spheres. Hydrodynamic forces only are considered and they are calculated using the inverse of the grand mobility matrix for far-held many-body interactions and lubrication formulae for near-field effects. We determine both the detailed microstructure (e.g. the pair-connectedness function and cluster formation) and the macroscopic properties (e.g. viscosity and self-diffusion coefficients). The flow of an ‘ infinite’ suspension is simulated by considering 25, 49, 64 and 100 particles to be one ‘cell’ of a periodic array. Effects of both the size ratio and the relative fractions of the different-sized particles are examined. For the microstructures, the pair-connectedness function shows that the particles form clusters in simple shearing flow due to lubrication forces. The nearly symmetric angular structures imply the absence of normal stress differences for a suspension with purely hydrodynamic interactions between spheres. For average mobilities at infinite Peclet number, Ds0, our simulation results suggest that the reduction of as concentration increases is directly linked to the influence of particle size distribution on the average cluster size. For long-time self-diffusion coefficients, we found good agreement between simulation and experiment (Leighton & Acrovos 1987 a; Phan and Leighton 1993) for monodispersed suspensions. For bimodal suspensions, the magnitude of and the time to reach the asymptotic diffusive behaviour depend on the cluster size formed in the system, or the viscosity of the suspension. We also consider the effect of the initial configuration by letting the spheres be both organized (size segregated) and randomly placed. We find that it takes a longer time for a suspension with an initially organized structure to achieve steady state than one with a random structure. © 1994, Cambridge University Press. All rights reserved.

Description: It is shown that realizability of second-moment turbulence closure models can be established by finding a Langevin equation for which they are exact. All closure models currently in use can be derived formally from the type of Langevin equation described herein. Under certain circumstances a coefficient in that formalism becomes imaginary. The regime in which models are realizable is, at least, that for which the coefficient is real. The present method does not imply unrealizable solutions when the coefficient is imaginary, but it does guarantee realizability when the coefficient is real; hence, this method provides sufficient, but not necessary, conditions for realizability. Illustrative computations of homogeneous shear flow are presented. It is explained how models can be modified to guarantee realizability in extreme non-equilibrium situations without altering their behaviour in the near-equilibrium regime for which they were formulated. © 1994, Cambridge University Press. All rights reserved.

Description: The flow in a breaking-wave crest is represented by a complex velocity potential on a Riemann surface, satisfying the Bernoulli condition on two free boundaries. The flow is assumed to be stationary in the reference frame which moves with the wave crest, and at large distances approximates Stokes corner flow in the main part of the fluid and a parabolic descending flow in the jet. The interaction of the jet with the rest of the fluid is neglected. The solution is obtained by means of a conformal transformation from a bounded, teardrop-shaped domain, using a Faber polynomial expansion. The Bernoulli condition is applied at a number of discrete points on the boundaries, and the resulting nonlinear equations for the expansion coefficients are solved iteratively. The resulting surface form is similar to that obtained by laboratory experiments and time-dependent numerical simulations of waves up to the point of breaking, with a stagnation point at the top of the crest, an overturning loop with major axis « 8g~? ¥ 5, and a maximum acceleration of « 5.4 g, where g is the gravitational acceleration and ¥ is the flux in the jet. © 1994, Cambridge University Press. All rights reserved.

Description: The equilibrium dynamics in a homogeneous forced-dissipative f-plane shallow-water system is investigated through numerical simulations. In addition to classical two-dimensional turbulence, inertio-gravity waves also exist in this system. The dynamics is examined by decomposing the full flow field into a dynamically balanced potential-vortical component and a residual ‘free’ component. Here the potential-vortical component is defined as part of the flow that satisfies the gradient-wind balance equation and that contains all the linear potential vorticity of the system. The residual component is found to behave very nearly as linear inertio-gravity waves. The forcing employed is a mass and momentum source balanced so that only the large-scale potential-vortical component modes are directly excited. The dissipation is provided by a linear relaxation applied to the large scales and by an eighth-order linear hyperdiffusion. The statistical properties of the potential-vortical component in the fully developed flow were found to be very similar to those of classical two-dimensional turbulence. In particular, the energy spectrum of the potential-vortical component at scales smaller than the forcing is close to the ~ k~3 expected for a purely two-dimensional system. Detailed analysis shows that the downscale enstrophy cascade into any wavenumber is dominated by very elongated triads involving interactions with large scales. Although not directly forced, a substantial amount of energy is found in the inertio-gravity modes and interactions among inertio-gravity modes are principally responsible for transferring energy to the small scales. The contribution of the inertio-gravity modes to the flow leads to a shallow tail at the high-wavenumber end of the total energy spectrum. For parameters roughly appropriate for the midlatitude atmosphere (notably Rossby number ~ 0.5), the break between the roughly — k~3 regime and this shallower regime occurs at scales of a few hundred km. This is similar to the observed mesoscale regime in the atmosphere. The nonlinear interactions among the inertio-gravity modes are extremely broadband in spectral space. The implications of this result for the subgrid-scale closure in the shallow-water model are discussed. © 1994, Cambridge University Press. All rights reserved.

Description: The stability of an axisymmetric vortex with a single radial discontinuity in potential vorticity is investigated in rotating shallow water. It is shown analytically that the vortex is always unstable, using the WKBJ method for instabilities with large azimuthal mode number. The analysis reveals that the instability is of mixed type, involving the interaction of a Rossby wave on the boundary of the vortex and a gravity wave beyond the sonic radius. Numerically, it is demonstrated that the growth rate of the instability is generally small, except when the potential vorticity in the vortex is the opposite sign to the background value, in which case it is shown that inertial instability is likely to be stronger than the present instability. © 1994, Cambridge University Press. All rights reserved.

Description: The one-dimensional dissolution that occurs when a binary melt is placed above or below a solid of a different composition is examined both theoretically and experimentally. In the case considered, the dissolution is driven by vigorous compositional convection that results from a Rayleigh-Bénard instability of the compositional boundary layer in the vicinity of the dissolving solid. A scaling analysis is used to derive theoretical expressions for both the dissolving velocity and the interfacial fluid concentration. Laboratory experiments are also described in which ice is dissolved when it is overlain or underlain by aqueous solutions. The measured dissolving velocities are consistent with the theoretical expressions, and yield estimates of the critical Rayleigh number for boundary-layer instability. The results of this study are then applied to predict the rate at which dissolution will occur when undersaturated mixed magmas are generated during the periodic replenishment of large basaltic magma chambers in the Earth's crust. © 1994, Cambridge University Press. All rights reserved.

Description: The linear stability of boundary-layer flow over compliant or flexible surfaces has been studied by Carpenter & Garrad (1985), Yeo (1988) and others on the assumption of local flow parallelism. This assumption is valid at large Reynolds numbers. Nonparallel effects due to growth of the boundary layer gain in significance and importance as one gets to lower Reynolds number. This is especially so for a compliant surface, which can sustain a variety of wall-related instabilities in addition to the Tollmien-Schlichting instabilities (TSI) that are found over rigid surfaces. The present paper investigates the influence of boundary-layer non-parallelism on the TSI and wall-related travelling-wave flutter (TWF) on compliant layers. Corrections to the growth rate of locally parallel theory for boundary-layer non-parallelism are obtained through a multiple-scale analysis. The results indicate that flow non-parallelism has an overall destabilizing influence on the TSI and TWF. Flow non-parallelism is also found to have a very strong destabilizing effect on the branch of TWF that stretches to low Reynolds number. The results obtained have important implications for the design and use of compliant layers at low Reynolds numbers. © 1994, Cambridge University Press. All rights reserved.

Description: Building on results from two-dimensional magnetohydrodynamic (MHD) turbulence (Shebalin, Matthaeus & Montgomery 1983), the development of anisotropic states from initially isotropic ones is investigated numerically for fully three-dimensional incompressible MHD turbulence. It is found that when an external d.c. magnetic field (B0) is imposed on viscous and resistive MHD systems, excitations are preferentially transferred to modes with wavevectors perpendicular to B0. The anisotropy increases with increasing mechanical and magnetic Reynolds numbers, and also with increasing wavenumber. The tendency of B0to inhibit development of turbulence is also examined. © 1994, Cambridge University Press. All rights reserved.

Description: The collision efficiency in a dilute suspension of sedimenting drops is considered, with allowance for particle Brownian motion and van der Waals attractive force. The drops are assumed to be of the same density, but they differ in size. Drop deformation and fluid inertia are neglected. Owing to small particle volume fraction, the analysis is restricted to binary interactions and includes the solution of the full quasi-steady Fokker-Planck equation for the pair-distribution function. Unlike previous studies on drop or solid particle collisions, a numerical solution is presented for arbitrary Peclet numbers, Pe, thus covering the whole range of particle size in typical hydrosols. Our technique is mainly based on an analytical continuation into the plane of complex Peclet number and a special conformal mapping, to represent the solution as a convergent power series for all real Peclet numbers. This efficient algorithm is shown to apply to a variety of convection-diffusion problems. The pair-distribution function is expanded into Legendre polynomials, and a finite-difference scheme with respect to particle separation is used. Two-drop mobility functions for hydrodynamic interactions are provided from exact bispherical coordinate solutions and near-field asymptotics. The collision efficiency is calculated for wide ranges of the size ratio, the drop-to-medium viscosity ratio, and the Peclet number, both with and without interdroplet forces. Solid spheres are considered as a limiting case; attractive van der Waals forces are required for non-zero collision rates in this case. For Pe P 1, the correction to the asymptotic limit Pe- 〉 oo is 0(Pe~1/2). For Pe 1, the first two terms in an asymptotic expansion for the collision efficiency are C/Pe + C2, where the constant C is determined from the Brownian solution in the limit Pe~〉 0. The numerical results are in excellent agreement with these limits. For intermediate Pe, the numerical results show that Brownian motion is important for Pe 〈 0(1O2). For Pe = 10, the trajectory analysis for Peoo may underestimate the collision rate by a factor of two. A simpler, approximate solution based on neglecting the transversal diffusion is also considered and compared to the exact solution. The agreement is within 2–3 % for all conditions investigated. The effect of van der Waals attractions on the collision efficiency is studied for a wide range of droplet sizes. Except for very high drop-to-medium viscosity ratios, the effect is relatively small, especially when electromagnetic retardation is accounted for. © 1994, Cambridge University Press. All rights reserved.

Description: The statistical relationship between a passive scalar and its dissipation is important for both a basic understanding of turbulence small-scale properties and for various aspects of turbulent combustion modelling. This problem is studied in two different flows through spectral analysis as well as probability density functions using temperature as a passive scatar. Particular attention is paid to the experimental determination of the three squared derivatives involved in the temperature dissipation. As a first step, it is found that basic properties such as the correlation coefficient between temperature and its dissipation are strongly related to the asymmetry of the scalar fluctuations, so that the usually assumed statistical independence between these variables is not justified. These trends are the same for the two flows investigated here, a boundary layer and a jet. This connection appears to be related to fluctuations of small amplitude for both quantities which are associated with relatively low frequencies lying between the integral scale and the Taylor microscale. In regions where the temperature skewness factor is nearly zero, the correlation coefficient is also very small, and several tests show that the assumption of independence is then fully justified. Thus, the main parameter influencing joint statistics of temperature and its dissipation is the asymmetric feature of temperature fluctuations, but the asymmetry of the longitudinal temperature derivative, which results from the flow boundary conditions and is usually felt through the presence of the so-called temperature ramps, is also involved. Even though the magnitude of the derivative skewness factor is almost uniformly distributed in both flows, the secondary effect becomes the dominant one in flow regions where the influence of the temperature asymmetry is relatively weak. © 1994, Cambridge University Press. All rights reserved.

Description: We consider Lagrangian stochastic modelling of the relative motion of two fluid particles in the inertial range of a turbulent flow. Eulerian analysis of such modelling corresponds to an equation for the Eulerian probability distribution of velocity-vector increments which introduces a hierarchy of constraints for making the model consistent with results from the theory of locally isotropic turbulence. A nonlinear Markov process is presented, which is able to satisfy exactly, in the statistical sense, incompressibility, the exact results on the third-order structure function, and the experimental second-order statistics. The corresponding equation for the Eulerian probability density of velocity-vector increments is solved numerically. Numerical results show non-Gaussian statistics of the one-dimensional Lagrangian probability distributions, and a complex shape of the three-dimensional Eulerian probability density function. The latter is then compared with existing experimental data. © 1994, Cambridge University Press. All rights reserved.

Description: By means of large-eddy simulation, homogeneous turbulence is simulated for neutrally and stably stratified shear flow at gradient-Richardson numbers between zero and one. We investigate the turbulent transport of three passive species which have uniform gradients in either the vertical, downstream or cross-stream direction. The results are compared with previous measurements in the laboratory and in the stable atmospheric boundary layer, as well as with results from direct numerical simulations. The computed and measured flow properties agree with each other generally within the scatter of the measurements. At strong stratification, the Froude number becomes the relevant flow-controlling parameter. Stable stratification suppresses vertical overturning and mixing when the inverse Froude number based on a turn-over timescale exceeds a critical value of about 3. The turbulent diffusivity tensor is strongly anisotropic and asymmetric. However, only the vertical and the cross-stream diagonal components are of practical importance in shear flows. The vertical diffusion coefficient is much smaller than the cross-stream one at strong stratification. This anisotropy is stronger than predicted by second-order closure models. Turbulence fluxes in downstream and cross-stream directions follow classical mixing-length models. © 1994, Cambridge University Press. All rights reserved.

Description: The melting of a solid in contact with a hot fluid is quantified for the case in which a difference between the densities of the fluid and of the melted solid is able to drive vigorous compositional convection. A scaling analysis is first used to obtain a theoretical expression for the melting rate that is valid for a certain range of Stefan numbers. This expression is then compared with melting velocities measured in laboratory experiments in which ice and wax are melted when they are overlain or underlain by hot aqueous solutions. The melting velocities are consistent with the theoretical expression, and are found to depend on the heats of solution that are released when the melted solids mix with the solutions. The experiments also indicate that, for vigorous convection to occur during the melting of a floor, the unstable compositional buoyancy needs to be at least twice the stabilizing thermal buoyancy. An important geological situation in which melting occurs is when large volumes of basaltic magma are intruded into the Earth's continental crust. The theoretical and experimental results are used and extended to examine quantitatively the melting of the floor and walls of the magma chamber, and of crustal blocks that fall into the chamber. © 1994, Cambridge University Press. All rights reserved.

Description: We have conducted further numerical experiments on two-dimensional fully compressible convection in an imposed vertical magnetic field and interpreted the results by reference to appropriate low-order models. Here we focus on streaming instabilities, involving horizontal shear flows, which form an important mechanism for the breakdown of steady convection in relatively weak fields for boxes of sufficiently small aspect ratio. While these shearing modes can arise even in the absence of a field, they will typically lead only to travelling and modulated waves unless there is a field to provide a restoring force. For magnetoconvection a new and dramatic form of pulsating wave appears after a complex sequence of secondary bifurcations. © 1994, Cambridge University Press. All rights reserved.

Description: The dispersion of passive scalars by the steady viscous flow through a twisted pipe is both a simple example of chaotic advection and an elaboration of Taylor's classic shear dispersion problem. In this article we study the statistics of the axial dispersion of both diffusive and perfect (non-diffusive) tracer in this system. For diffusive tracer chaotic advection assists molecular diffusion in transverse mixing and so diminishes the axial dispersion below that of integrable advection. As in many other studies of shear dispersion the axial distribution ultimately becomes Gaussian as. Thus there is a diffusive regime, but with an effective diffusivity that is enhanced above molecular values. In contrast to the classic case, the effective diffusivity is not necessarily inversely proportional to the molecular diffusivity. For instance, in the irregular regime the effective diffusivity is proportional to the logarithm of the molecular diffusivity. For perfect tracer chaotic advection does not result in a diffusive process, even in the irregular regime in which streamlines wander throughout the cross-section of the pipe. Instead the variance of the axial position is proportional to t In t so that the measured diffusion coefflcent diverges like ln t. This faster than linear growth of variance is attributed to the trapping of tracer for long times near the solid boundary, where the no-slip condition ensures that the fluid moves slowly. Analogous logarithmic effects associated with the no-slip condition are well known in the context of porous media. A simple argument, based on Lagrangian statistics and a local analysis of the trajectories near the pipe wall, is used to calculate the constants of proportionality before the logarithmic terms in both the large- and infinite-Peclet-number limits. © 1994, Cambridge University Press. All rights reserved.

Description: When a rotating fluid over sloping topography is heated from below and/or cooled from above, horizontal temperature gradients develop which drive convection cells aligned with isobaths. We refer to these cells as topographic Hadley cells. Laboratory experiments reveal that sinking occurs in small cyclonic vortices situated in relatively shallow regions. This is balanced by slower upwelling in adjacent deeper regions. The cross-isobath motions which connect the upwelling and downwelling are accelerated by Coriolis forces, resulting in strong jets which follow isobathic contours. For anticlockwise rotation, the surface jets keep the shallows to their left when looking in the direction of flow, which is opposite to both Kelvin and Rossby wave propagation. The width of the jets scales with the Rossby deformation radius and if this is much less than the width of the slope region then a number of parallel jets form. Motions on the deeper side of the jets where the flow is accelerating are adequately described by linear inviscid theory. However, the strong shears generated by this acceleration lead to baroclinic instability. The resulting cross-stream momentum fluxes broaden and flatten the velocity profile, allowing the flow on the shallow side of the jet to decelerate smoothly before sinking. Topographic Hadley cells are dynamically similar to terrestrial atmospheric Hadley cells and may also be relevant to the zonal jet motions observed on Jupiter and Saturn. It is also suggested that in coastal seas they may represent an important mode of heat (or salt) transfer where surface cooling (or evaporation) drives convection. © 1994, Cambridge University Press. All rights reserved.

Description: Low-density flow of molecular hydrogen from a small nozzle is studied using numerical and experimental techniques. The conditions in the nozzle indicate that nonequilibrium effects will significantly influence the flow. Therefore, the numerical analysis is undertaken using a Monte Carlo approach. The experimental studies employ spontaneous Raman scattering. Comparisons of the measured data and computed results are made for total number density, rotational temperature, and for the number density of the first rotational level. The numerical results are found to be quite sensitive to the rotational relaxation rate, and a strong degree of thermal nonequilibrium is observed at the exit plane of the nozzle. Comparisons between experiment and analysis permit estimation of the rotational relaxation rate for hydrogen. Investigations are also conducted for expansion of the supersonic jet into a finite back pressure. The interaction of the plume with the chamber background gas is found to form shock waves in both the simulations and experiments. This phenomenon is investigated further by increasing the background pressure. Direct comparison of the simulation results and experimental measurements is very favourable. © 1994, Cambridge University Press. All rights reserved.

Description: Turning international agreement into national law has never been rapid or simple. Each country has its own parliamentary procedures, its own legal system and its own cultural interpretation of what the words in the international agreement actually mean. Last but not least, the importance attached to implementing Treaty law is clearly different between countries, as the amazingly patchy implementation of existing Treaty Recommendations shows.

Description: Values of attenuation coefficient, Kd(λ), for five visible wavelengths are reported for 14 sites around South Georgia and the Bransfield Strait, in the Southern Ocean. The mean chlorophyll-plus-phaeopigment concentrations in the upper 30 m of the water column ranged from 0.32–6.633 mg m−3 with one particularly high mean value of 31 mg m−3. Partition of attenuation between chlorophyll (Kc) and other factors in the water column (Ko) indicated that the spectral character of Ko in the Bransfield Strait was consistent with absorption by non chlorophyll-like pigments (Gelbstoff). Values were significantly different between the two areas. Values of the specific attenuation coefficient due to pigment concentration (Kc) were small compared to temperate values. These data support other studies in suggesting that the application of remotely sensed ocean colour data to global biogeochemical surveys requires the development of regional algorithms.

Description: SUMMARYThe ability of 12 Scottish Blackface ewes to discriminate and concentrate grazing activity in nitrogenfertilized heather patches was determined. The nitrogen content of a stand of heather (Calluna vulgaris) was increased and the fibre content decreased by adding ammonium nitrate fertilizer (7·5 g N/m2) 3 months prior to grazing, in Scotland during August 1991, to selected areas within a patchwork array. The sheep were offered either a high protein (300 g soyabean meal) or low protein (285 g ground barley) supplement daily before being allowed to graze within the heather mosaic during November 1991. The proportion of time spent grazing in fertilized and control patches (patch selection) and the proportion of daylight hours spent grazing (proportion of time spent grazing) was estimated for individual sheep for 6 days. Rumen ammonia concentrations were determined at the end of the grazing period. All animals spent approximately twice as much time grazing in fertilized as in control patches. About half of the daylight hours were spent grazing. The nature of the supplement did not influence patch selection, the proportion of time spent grazing or rumen ammonia concentrations. It was concluded that sheep are able to discern differences in the chemical composition of heather on a fine scale and that their foraging behaviour is not influenced by the type of supplement offered.

Description: SUMMARYThe effects of infection with T. vivax in mid- or late pregnancy on food intake and utilization, liveweight changes, abortion rate and lamb growth rate were investigated in West African Dwarf ewes at lbadan, Nigeria in 1990. Rate of liveweight gain by ewes infected during mid-pregnancy (IMH) was16 g/day compared with 33 and 37 g/day for the uninfected ewes offered medium (CM) or high (CH) plane diets. Although digestibility coefficients were not affected, intake of digestible organic matter was higher in CH ewes than in IMH and CM ewes. Nitrogen retention at mid-pregnancy on a metabolic size basis was higher in CH ewes than in CM and IMH ewes.Lamb birth weight and survival rate were lower in infected ewes. Ewes infected in mid-pregnancy (IMH) and in late pregnancy (ILH) had mean birth weights of 1·4 and 1·0 kg compared with CM and CH ewes, which had mean birth weights of 1·9 and 2·0 kg respectively. Observed survival rates were 63, 15, 75 and 80% for lambs nursed by IMH, ILH, CM and CH ewes respectively. During the first 6 weeks postpartum, lamb growth rate in all groups did not differ. However, during weeks 7–12 postpartum, lambs nursed by IMH ewes had significantly lower growth rates. Weaning weight was also lower in lambs from IMH (5·0 kg) dams than in lambs from CM and CH dams (7·1 kg). Infection during late pregnancy was more severe and all infected ewes lost weight due to reduced feed intake and fever. T. vivax infection in sheep is responsible for reproductive wastage, abortion, poor lamb growth and ewe mortality.

Description: SUMMARYThree rates of phosphate (0, 25 and 60 kg/ha P2O5) were applied for 7 years to phosphate-deficient grassland at Tel Hadya, north Syria. Liveweight, milk yield, wool production and supplementary feeding of Awassi sheep were monitored for six seasons (1985/86 to 1990/91). The experiment was grazed at low (0·8 sheep/ha per year) and high (1·7 sheep/ha per year) stocking rates from the second to the fourth season, while in the fifth to seventh seasons stocking rates were increased to 1·1 and 2·3 sheep/ha per year, respectively. The experimental site was typical of communally owned, degraded grasslands within the cereal zone of west Asia, where cropping is not possible because of shallow, stony soils and steep slopes.The results showed that annual applications of phosphate, even at 25 kg/ha, improved pasture and sheep productivity. Liveweights were higher on fertilized plots in five out of six years, significantly so in the last three. Milk production was also higher on phosphate-treated plots, and the need for supplementary feeding was reduced, especially in the last three years, when rainfall was below average. The results suggest that stocking rates can be significantly increased by annual applications of small amounts of superphospate, and that doing so is profitable.Use of the results depends on the presence of native legumes, the level of soil phosphate, and the ability of farmers to control grazing of these communally owned grasslands. A strategy to fulfil these criteria is suggested.

Description: SUMMARYThe progenies of six crosses, made in 1987, between the best available hooded barley line and six selected awned barley lines were used to study the effect of the hooded gene on the performance of barley under rainfed conditions in Cyprus. A single dominant gene was found to control the hooded character. Awned genotypes on average outyielded hooded genotypes by 17%, but the difference between the two genotypes was smaller in the central plain than in the coastal area. The effect of the hooded gene on yield varied also with the combination of parents, as the superiority of the awned genotypes varied with cross from 13 to 30%. Hooded lines giving a higher grain yield than their hooded parent were identified. On average the awned genotypes had higher 1000-grain weight (5·0 g), were slightly later in heading date (1·1 days), had slightly higher volume weight (1·3 kg/hl) than hooded genotypes, but had similar plant height. The 1000-grain weight of most hooded lines was significantly higher than that of their hooded parent.In spite of the negative effects of the hooded gene on grain yield, it was concluded that it is possible to increase the yield of hooded barley for hay production by continued crossing and selection (with special emphasis on 1000-grain weight), by selecting special combinations of parents through the Bulk-Pedigree method and by growing hooded barley in suitable areas, making use of the locationdependent performance of the hooded gene.

Description: SUMMARYFour experiments were conducted in the Yucatán, Mexico, in 1991 and 1992 to study feed intake in 24 h trials with chicks and pigs fed diets containing either raw or treated jack beans or conventional diets with added canavanine.Feed intake of chicks in the first hour was markedly reduced and feed intake of pigs was severely reduced when raw jack beans (300 g/kg for chicks, 250 g/kg for pigs) were included in their diets. Feed intake of pigs diminished to a greater extent than that of chicks when treated beans (300 g/kg) were included in the diet. Short-term feed intake of chicks was not affected when canavanine (3·5 g/kg) was added to a sorghum-soyabean diet. However, pigs showed a large drop in feed intake when canavanine was added to a maize-soya diet at only 0·8 g/kg. It was concluded that in both species the anti-nutritional factors in jack beans cause a rapid reduction in feed intake which precedes a depression in growth rate. The initial feed intake depression in pigs could be partly attributed to canavanine remaining in the jack beans after treatment. In the chicks' case, however, rejection of a diet containing treated jack bean was due to anti-nutritional factor(s) other than residual canavanine.