ALBERT

Beschreibung: Ongoing climate warming in the western Canadian Arctic is leading to thawing of permafrost soils and subsequent mobilization of its organic matter pool. Part of this mobilized terrestrial organic matter enters the aquatic system as dissolved organic matter (DOM) and is laterally transported from land to sea. Mobilized organic matter is an important source of nutrients for ecosystems, as it is available for microbial breakdown, and thus a source of greenhouse gases. We are beginning to understand spatial controls on the release of DOM as well as the quantities and fate of this material in large Arctic rivers. Yet, these processes remain systematically understudied in small, high-Arctic watersheds, despite the fact that these watersheds experience the strongest warming rates in comparison. Here, we sampled soil (active layer and permafrost) and water (porewater and stream water) from a small ice wedge polygon (IWP) catchment along the Yukon coast, Canada, during the summer of 2018. We assessed the organic carbon (OC) quantity (using dissolved (DOC) and particulate OC (POC) concentrations and soil OC content), quality (δ13C DOC, optical properties and source apportionment) and bioavailability (incubations; optical indices such as slope ratio, Sr; and humification index, HIX) along with stream water properties (temperature, T; pH; electrical conductivity, EC; and water isotopes). We classify and compare different landscape units and their soil horizons that differ in microtopography and hydrological connectivity, giving rise to differences in drainage capacity. Our results show that porewater DOC concentrations and yield reflect drainage patterns and waterlogged conditions in the watershed. DOC yield (in mg DOC g−1 soil OC) generally increases with depth but shows a large variability near the transition zone (around the permafrost table). Active-layer porewater DOC generally is more labile than permafrost DOC, due to various reasons (heterogeneity, presence of a paleo-active-layer and sampling strategies). Despite these differences, the very long transport times of porewater DOC indicate that substantial processing occurs in soils prior to release into streams. Within the stream, DOC strongly dominates over POC, illustrated by ratios around 50, yet storm events decrease that ratio to around 5. Source apportionment of stream DOC suggests a contribution of around 50 % from permafrost/deep-active-layer OC, which contrasts with patterns observed in large Arctic rivers (12 ± 8 %; Wild et al., 2019). Our 10 d monitoring period demonstrated temporal DOC patterns on multiple scales (i.e., diurnal patterns, storm events and longer-term trends), underlining the need for high-resolution long-term monitoring. First estimates of Black Creek annual DOC (8.2 ± 6.4 t DOC yr−1) and POC (0.21 ± 0.20 t yr−1) export allowed us to make a rough upscaling towards the entire Yukon Coastal Plain (34.51 ± 2.7 kt DOC yr−1 and 8.93 ± 8.5 kt POC yr−1). Rising Arctic temperatures, increases in runoff, soil organic matter (OM) leaching, permafrost thawing and primary production are likely to increase the net lateral OC flux. Consequently, altered lateral fluxes may have strong impacts on Arctic aquatic ecosystems and Arctic carbon cycling.

Beschreibung: Die Stammesgeschichte der Ordnung Ptenoglossa (Gastropoda, Caenogastropoda) wird anhand von 179 rezenten und fossilen Arten bis ins Paläozoikum zurückverfolgt. Ihre rezenten Überfamilien, die Cerithiopsoidea, Triphoroidea und Janthinoidea ernähren sich karnivor. Eine kollabral berippte Larvalschale stellt in den rezenten Überfamilien den ursprünglichen Zustand dar und ist die aussagekräftigste schalenmorphologische Symplesiomorphie. Dies erweist sich durch Radulauntersuchungen und den Fossilbericht. Cerithiopsoidea und Triphoroidea fressen Schwämme und bilden gemeinsam mit den mesozoischen Protorculidae ein Monophylum. Die Janthinoidea fressen Coelenteraten. Sie sind nahe mit den mesozoischen Zygopleuridae verwandt. Die Stammlinien der Schwamm- und Coelenteratenfresser waren schon in der Trias getrennt. Die Zygopleuroidea (Zygopleuridae, Pseudozygopleuridae und Protorculidae) sind folglich ein Parataxon. In allen drei Familien der Zygopleuroidea kommt die kollabral berippte Larvalschale vor und repräsentiert den ursprünglichen Zustand. Die paläozoischen Pseudozygopleuridae stellen wahrscheinlich die Schwestergruppe der rezenten und mesozoischen Ptenoglossa dar. Arten der Pseudozygopleuridae mit planktotropher und nicht planktotropher Frühontogenese können biometrisch getrennt werden. Die Abgrenzung nicht planktotropher Pseudozygopleuridae von den devonischen bis karbonischen Palaeozygopleuridae ist schwierig, weil letztere anhand eines nicht planktotrophen Protoconchs definiert wurden. Mit Hilfe von Protoconchvermessungen ist es jedoch meist möglich, beide Gruppen zu trennen. Die Außengruppe der Ptenoglossa sind die Cerithimorpha, die im Paläozoikum durch die Acanthonematidae und die Murchisoniidae repräsentiert werden. Die paläozoischen Acanthonematidae werden neu gefaßt und enthalten nun unter anderem die Gattungen Orthonema, Palaeostylus, Cerithioides und Knightella. Sie zeichnen sich durch einen heliciformen Protoconch aus, der sich grundsätzlich von dem der Pseudozygopleuridae (Ptenoglossa) unterscheidet. Mithin ist der Protoconch der Pseudozygopleuridae die wesentliche schalenmorphologische Apomorphie der Ptenoglossa. 30 Arten werden neu beschrieben (siehe Anhang A), davon 2 rezente, 8 aus dem Tertiär, 1 aus der Kreide, 13 aus der Trias, 1 aus dem Perm und 5 aus dem Karbon. Turritella hybrida MONSTER non DESHAYES erhält den neuen Namen Zygopleura hybridissima nom. nov. 5 neue Gattungen werden errichtet: Antiphora n. Gen. (Triphoroidea, Tertiär), Eorex n. gen. (Triphoroidea, Tertiär), Atorcula n. gen. (Protorculidae, Trias), Azyga n. gen. (Zygopleuridae, Trias) und Stiazyga n. gen. (Zygopleuridae, Trias). Ampezzopleurinae n. subfam. wird als Unterfamilie der Zygopleuridae WENZ errichtet. Nystiellinae CLENCH & TURNER erhält Familienstatus (Nystiellidae). Die Untergattung Cerithiopsis (Vatopsis) GRÜNDEL wird zur Gattung erhoben und von den Cerithiopsidae zu den Eumetulidae transferiert. Tembrockia GRÜNDEL wird von den Cerithiopsidae zu den Eumetulidae transferiert. Variseila DOCKERY wird von den Triforidae JOUSSEAUME zu den Eumetulidae GOLIKOV & STAROBOGATOV transferiert. Ampezzopleura BANDEL wird von den Protorculidae BANDEL zu den Zygopleuridae WENZ transferiert. Zygopleura tenuis (MÜNSTER) sensu Zardini wird als neue nominelle Art Ampezzopleura tenuis BANDEL betrachtet, für die ein Lectotyp hinterlegt wird. Teutonica SCHRÖDER wird von den Cerithiopsidae zu den Zygopleuridae transferiert. Orthonema MEEK & WORTHEN wird von den Turritellidae LOVÉN zu den Acanthonematidae WENZ zurücktransferiert. Palaeostylus MANSUY wird von den Procerithiidae COSSMANN zu den Acanthonematidae transferiert. Knightella LONGSTAFF wird von den Pseudozygopleuridae KNIGHT zu den Acanthonematidae transferiert. Cerithioides HAUGHTON wird von den Murchisoniidae zu den Acanthonematidae WENZ transferiert. Die systematische Stellung einiger weiterer Taxa wird kritisch hinterfragt. Etliche Arten werden anderen Gattungen zugeordnet (siehe Anhang A "comb. nov.").

Beschreibung: The phylogeny of the order Ptenoglossa (Gastropoda, Caenogastropoda) is traced back into the Paleozoic by studying 179 Recent and fossil species. The Recent superfamilies Cerithiopsoidea, Triphoroidea und Janthinoidea are carnivorous. A larval shell with collabral ribs represents the plesiomorphic state in the Recent superfamilies and it is the most informative symplesiomorphy that concerns to the shell morphology. Evidence for that is given by the radula morphology and the fossil record. Cerithiopsoidea and Triphoroidea feed on sponges and form a clade with the Mesozoic family Protorculidae. The Janthinoidea feed on coelenterates. They are closely related to the Mesozoic family Zygopleuridae. The stem lines of sponge eaters and coelenterate eaters have been separated from each other at least since the Triassic. Thus, the superfamily Zygopleuroidea (Zygopleuridae, Pseudozygopleuridae and Protorculidae) is a parataxon. In all three families of the Zygopleuroidea a larval shell with collabral ribs occurs and represents the plesiomorphic state. The Paleozoic Pseudozygopleuridae are presumably the sister-group of the Recent and the Mesozoic Ptenoglossa. Species of the Pseudozygopleuridae with planktotrophic and non-planktotrophic larval development can be separated from each other by measuring their protoconchs. The separation of non-planktotrophic Pseudozygopleuridae from Devonian and Carboniferous species of the Palaeozygopleuridae is difficult because Palaeozygopleuridae were defined by a non-planktotrophic protoconch, whereas Pseudozygopleuridae were defined by a larval shell of the planktotrophic type. But in most cases it is possible to identify the species by protoconch measurements. The outgroup of the Ptenoglossa are the Cerithimorpha which are represented in the Paleozoic by the families Acanthonematidae and Murchisoniidae. The Paleozoic Acanthonematidae are newly defined and contain genera like Orthonema, Palaeostylus, Cerithioides and Knightella. They have a heliciform protoconch which is fundamentally different from the protoconch of the pseudozygopleurids (Ptenoglossa). The protoconch of the Pseudozygopleuridae is the essential apomorphy concerning the shell of the Ptenoglossa. 30 species are described as new (see Anhang A), 2 of which are Recent, 8 are from the Tertiary, 1 from the Cretacous, 13 from the Triassic, 1 from the Permian and 5 from the Carboniferous. Turritella hybrida MÜNSTER non DESHAYES gets the new name Zygopleura hybridissima nom. nov. 5 new genera are erected: Antiphora n. gen. (Triphoroidea, Tertiary), Eorex n. gen. (Triphoroidea, Tertiary), Atorcula n. gen. (Protorculidae, Triassic), Azyga n. gen. (Zygopleuridae, Triassic) and Striazyga n. gen. (Zygopleuridae, Triassic). Ampezzopleurinae n. subfam. is erected as subfamily of the Zygopleuridae. Nystiellinae CLENCH & TURNER is raised on family level (Nystiellidae). The subgenus Cerithiopsis (Vatopsis) GRÜNDEL is raised on genus level and is transferred from Cerithiopsidae H. & A. ADAMS to Eumetulidae GOLIKOV & STAROBOGATOV. Tembrockia GRÜNDEL is transferred from Cerithiopsidae to Eumetulidae. Variseila DOCKERY is transferred from Triforidae JOUSSEAUME to Eumetulidae. Ampezzopleura BANDEL is transferred from Protorculidae BANDEL to Zygopleuridae WENZ. Zygopleura tenuis (MÜNSTER) sensu Zardini is deemed to be the new nominal species Ampezzopleura tenuis BANDEL (type species of Ampezzopleura) for which a lectotype is designated. Teutonica SCHRÖDER is transferred from Cerithiopsidae to Zygopleuridae. Orthonema MEEK & WORTHEN is retransferred from Turritellidae LOVÉN to Acanthonematidae WENZ. Palaeostylus MANSUY is transferred from Procerithiidae COSSMANN to Acanthonematidae. Knightella LONGSTAFF is transferred from Pseudozygopleuridae KNIGHT to Acanthonematidae. Cerithioides HAUGHTON is transferred from Murchisoniidae KOKEN to Acanthonematidae WENZ. The systematic position of several other taxa has been critically revised. The generic position of several species is changed (see Anhang A "comb. nov.").

Beschreibung: thesis

Beschreibung: DFG, SUB Göttingen

Beschreibung: Die Ordnungen Cerithiimorpha und Littorinimorpha stellen umfangreiche systematische Gruppen innerhalb der basalen Caenogastropoda mit einfachen konisch-orthostrophen Protoconchen dar. Diese Großgruppen lassen sich seit der späten Trias differenzieren und lassen seit der Kreidezeit eine zunehmende Diversität erkennen. Im folgenden werden ausgewählte Vertreter aus fünf Überfamilien und 20 Familien unter besonderer Berücksichtigung ihrer frühontogenetischen Schalen beschrieben und ihre Entwicklungsgeschichte wird seit der Kreide belegt. Die Überfamilie Cerithioidea stellt die Kerngruppe der Cerithiimorpha und war mit der Familie Procerithiidae seit dem Jura weltweit im Flachmarin verbreitet. Form und Skulptur der Larvalschale ermöglichte es, die Gattung Schroederium n.g. von den Gattungen Procerithium und Cryptaulax zu trennen. In der Oberkreide lassen sich die nahe verwandten Familien Cassiopidae, Potamididae, Melanopsidae und Scaliolidae über ihre Protoconchmorphologie sicher von den Procerithiidae differenzieren. Die Familien Batillariidae und Modulidae sind durch ihre charakteristische Embryonalschalenskulptur als Schwestergruppen ausgewiesen. Ihre nahe Verwandtschaft wird auch durch anatomische Daten gestützt. Innerhalb der seit dem Eozän in Protoconcherhaltung dokumentierten Familie Cerithiidae, Unterfamilie Cerithiinae, treten abhängig vom besiedelten Habitat zwei Protoconchtypen auf, die mit zu unterscheidenden Radulatypen korrelieren. Die Unterfamilie Bittiinae konnte über Details der Larvalschalenskulptur seit dem Eozän belegt und von den Cerithiinae differenziert werden. Die Adelphotaxa Planaxidae und Thiaridae mit dem gemeinsamen Merkmal einer Bruttasche im Kopffußbereich sind ebenfalls seit dem Eozän bekannt. Larvalschalen fossiler und rezenter Planaxinae werden vergleichend dargestellt. Larvalschalen der hinsichtlich der Morphologie des Teleoconches abweichenden Unterfamilie Fossarinae belegen die Nähe zur Nominatunterfamilie und ermöglichten eine sichere Abgrenzung von gehäusekonvergenten Vertretern der Littorinimorpha. Über die Formation der frühontogenetischen Schale, die eine charakteristische Embryogenese widerspiegelt, war es möglich, die Brackwasser und limnische Biotope besiedelnden Thiaridae bis in das mittlere Eozän zu belegen, was auch Anlaß zu neuen paläobiogeographischen Interpretationen gab. Pseudamauridae konnten über ihre Protoconche von der Oberkreide bis in das Eozän belegt und von gehäusekonvergenten Naticoidea (Neomesogastropoda) und Amphibolidae innerhalb der Archaeopulmonata (Heterostropha) abgegrenzt werden. Die Phylogenese der Überfamilien Vermetoidea und Turritelloidea wurde von der Unterkreide bis rezent dokumentiert, wobei konvergente Gruppen über ihre Protoconche differenziert wurden. Innerhalb der Littorinimorpha konnten charakteristische Merkmale der Embryonal- und Larvalschalen herausgearbeitet und zur Untergliederung der Littorinoidea und Rissooidea herangezogen werden. Pickworthiidae mit alloisostroph abgewinkelten Protoconchen konnten bis in das Danium belegt werden. Der direkte Vergleich mit Protoconchen der triassischen Protstyliferidae mit detaillierten Übereinstimmungen dokumentiert die unabhängige Evolution dieser Linie. Die konvergenten Vanikoridae ließen sich über ihre Protoconchmorphologie trennen, wobei auch signifikante Unterschiede zu den übrigen Littorinimorpha dargestellt werden. Micromphalina peyrerensis und Megalomphalus (M.) antwerpensis werden neu beschrieben und den Vanikoridae eingegliedert.

Beschreibung: The orders Cerithiimorpha and Littorinimorpha represent large systematic units within the stem of Caenogastropoda with simple conical-orthostrophic protoconchs. These groups are separated since the Upper Triassic showing increasing diversity since the Upper Cretaceous. Representatives of five superfamilies and 20 families are described with special reference to their early ontogenetic shells. The family Procerithiidae within the superfamily Cerithioidea has been distributed worldwide since Jurassic times as a faunal element in shallow marine environments. Regarding the shape and sculpture of its larval shell the procerithiid Schroederium n.g. could be differentiated from the genera Procerithium and Cryptaulax. Since the Upper Cretaceous brackish-water Cassiopidae, Potamididae and Melanopsidae can be differentiated from Procerithiidae by their protoconch-morphology. The families Batillariidae and Modulidae represent Adelphotaxa. This relation is proven by the uniting character of embryonic tuberculated sculpture and data concerning their anatomy. The Cerithiidae, subfamily Cerithiinae, are documented since the Eocene with preserved protoconchs. Two types of larval sculpture are present depending on the settled habitat, correlated with two different types of radulae. The subfamily Bittiinae is also documented and distinguished from the Cerithiinae by their protoconch-morphology since the Eocene. The Adelphotaxa Planaxidae and Thiaridae with the uniting character of a brood pouch within the head-foot are present since the Eocene. Larval shells of fossil and Recent Planaxinae are compared. Protoconchs of representatives of the subfamily Fossarinae with different teleoconch-morphology proved the close relation to the Planaxinae and also made a differentiation from convergent littorinimorphs feasible. The analysis of early ontogenetic shells of brackish water and limnic Thiaridae enabled to trace them back to the Middle Eocene and made new interpretations concerning their palaeobiogeography possible. Pseudamauridae could be documented from the Upper Cretaceous to the Middle Eocene with the aid of preserved protoconchs and could be discriminated from convergent Naticoidea and Archaeopulmonata. The phylogenetic history of Vermetoidea and Turritelloidea could be reconstructed since the Early Cretaceous and convergent groups could be differentiated. Within the Littorinimorpha characters regarding the early shell could be worked out to subdivide Littorinoidea and Rissooidea. Pickworthiidae with alloisostrophic protoconchs are documented since the Danian. Comparison with protoconchs of Triassic Prostyliferidae yielded detailed accordance and documents the independent history of this lineage. The convergent Vanikoridae could be discriminated by their protoconch-morphology and significant differences to the other Littorinimorpha are presented. Micromphalina peyrerensis and Megalomphalus (M.) antwerpensis are described as new species and are included within the Vanikoridae.

Beschreibung: thesis

Beschreibung: DFG, SUB Göttingen

Beschreibung: A fragmentary nerineoid gastropod collected from the Maasvlakte 2 beach may be identified as Ptygmatis cf. bruntrutana (Thurmann, 1832). Probably originating from Upper Jurassic strata in north-eastern France, it must have been transported here by a forerunner of the River Meuse (Maas).

Beschreibung: Ridsdalea philippinensis (Rubiaceae), a new species from the karst forest in El Nido (Palawan, Philippines), is described and illustrated. It is unique among Malesian Ridsdalea species in having a corolla tube that is distinctly inflated at the apex, a character state also displayed by R. sootepensis and R. thailandica from Laos and Thailand. Amongst others, R. philippinensis, however, differs from both of these species in having smaller flowers and anthers that do not emerge from the corolla tube. An updated key for Philippine Ridsdalea is also presented.

Beschreibung: In the present paper, the existence and location of the type series of the Japanese dancing mouse or waltzer, Mus wagneri variety rotans Droogleever Fortuyn, 1912, are established, and a lectotype is designated. Available type specimens are measured, and some morphological parameters, sex, and general condition of the specimens are recorded. A literature survey was conducted, and an attempt is made to clarify the position of M. wagneri variety rotans in the taxonomy of Mus. A genetic analysis suggests that the type series of the Japanese dancing mouse represent a crossbred, or derivation of a crossbred, between the original Japanese dancing mouse of Mus musculus molossinus Temminck 1844 origin and European fancy or laboratory mice of Mus musculus domesticus Schwarz & Schwarz, 1943 origin. Much of their genome was replaced and occupied by Mus musculus domesticus type genome, probably through extensive breeding with European mice.

Beschreibung: The genus Micronecta Kirkaldy, 1897 is the most species-rich genus in the family Micronectidae, containing more than 160 species. Micronecta is currently divided into 11 subgenera, five of which are monotypic. Moreover, the subgenus Micronecta is an empirical mixture group. The definitions of some subgenera were based on only a few aberrant morphological features, which are specializations with few phylogenetic significances. The relationship between these subgenera remains unclear. In this study, we newly sequenced mitochondrial genomes (mitogenomes) and nuclear rDNAs (nrDNAs) for 13 Micronecta species, representing seven subgenera, and those for ten other water bugs. Our phylogenetic analyses showed that the subgenus Lundbladella represents the sister group to all other studied subgenera of Micronecta. The subgenus Unguinecta was the sister group to the clade that contains Dichaetonecta and Sigmonecta. More importantly, the subgenus Micronecta represents a paraphyletic group, which further forms a monophyletic group together with the subgenera Basileonecta and Ctenonecta. This is for the first time that the phylogeny of the genus Micronecta was investigated based on molecular data and the paraphyly of the subgenus Micronecta was revealed. Such evidence suggested the necessity of the revision of the taxonomic system of the genus in the future, and may also serve as a reference for the delimitation of subgeneric characters.

Beschreibung: Climate change and land cover change often interactively affect plant species distributions. This study addresses the vulnerability of lowland and upland orchids to climate change and land cover change. Endemic orchids of New Guinea were grouped into four classes (lowland epiphyte, lowland terrestrial, upland epiphyte, upland terrestrial) based on their life form and elevation range. Forty occurrence records of endemic orchids were selected for each class, totaling 160 occurrence records. Ensemble modelling combining two machine learning algorithms was used to generate predictive current and future suitable areas for orchid classes. Model performance was evaluated using the AUC and TSS metrics. Suitable areas for both lowland and upland orchids (epiphyte and terrestrial) were predicted decrease in the future due to climate change and land cover change. The loss of suitable areas for upland terrestrial orchids was predicted to be most significant in the worst-case climate change scenario (SSP 5–8.5). Both lowland and upland orchids (epiphyte and terrestrial) tend to shift to higher elevation ranges from the present distributions. The predictive models have AUC values 〉0.90 and TSS value 〉0.80, indicating the models have excellent potential for predicting the impact of climate change and land cover change on orchid distributions.

Beschreibung: Snakebite envenoming is a priority Neglected Tropical Disease that causes an estimated 81,000–135,000 fatalities each year. The development of a new generation of safer, affordable, and accessible antivenom therapies is urgently needed. With this goal in mind, rigorous characterisation of the specific toxins in snake venom is key to generating novel therapies for snakebite. Monoclonal antibodies directed against venom toxins are emerging as potentially strong candidates in the development of new snakebite diagnostics and treatment. Venoms comprise many different toxins of which several are responsible for their pathological effects. Due to the large variability of venoms within and between species, formulations of combinations of human antibodies are proposed as the next generation antivenoms. Here a high-throughput screening method employing antibody-based ligand fishing of venom toxins in 384 filter-well plate format has been developed to determine the antibody target/s The approach uses Protein G beads for antibody capture followed by exposure to a full venom or purified toxins to bind their respective ligand toxin(s). This is followed by a washing/centrifugation step to remove non-binding toxins and an in-well tryptic digest. Finally, peptides from each well are analysed by nanoLC-MS/MS and subsequent Mascot database searching to identify the bound toxin/s for each antibody under investigation. The approach was successfully validated to rapidly screen antibodies sourced from hybridomas, derived from venom-immunised mice expressing either regular human antibodies or heavy-chain-only human antibodies (HCAbs).

Beschreibung: European marbled newts come in two species that have abutting ranges. The northern species, Triturus marmoratus, is found in France and the northern part of the Iberian Peninsula, whereas the southern species, T. pygmaeus, is found in the southwestern corner of the Iberian Peninsula. We study the intraspecific genetic differentiation of the group because morphological data show geographical variation and because the Iberian Peninsula is a recognized center of speciation and intraspecific genetic diversity for all kinds of organisms, amphibians included. We use target enrichment by sequence capture to generate c. 7 k nuclear DNA markers. We observe limited genetic exchange between the species, which confirms their distinctiveness. Both species show substantial genetic structuring that is only in part mirrored by morphological variation. Genetically differentiated groups are found in the south (T. marmoratus) and west (T. pygmaeus) of the species ranges. Our observations highlight the position of the Iberian Peninsula as a hotspot for genetic differentiation.