ALBERT

Note: Dissertation, Universität Potsdam, 2019 , Table of contents Abstract Zusammenfassung Abbreviations 1 Introduction 1.1 Scientific background 1.1.1 Permafrost in the Northern Hemisphere 1.1.2 The permafrost carbon climate feedback 1.1.3 Rapidly changing, deep permafrost environments 1.2 Aims of this dissertation 1.3 Investigated study areas 1.4 Basic method overview 1.4.1 Field work in the Arctic 1.4.2 Laboratory procedure 1.4.3 Analysis ofl andscape-scale carbon and nitrogen stocks 1.5 Thesis organization 1.6 Overview of publications 1.6.1 Publication#1 - Yedoma landscape publication 1.6.2 Publication#2 - Thermokarst lake sequence publication 1.6.3 Publication#3 - North Alaska Arctic river delta publication 1.6.4 Extended Abstract - Western Alaska river delta study 1.6.5 Appendices - Supplementary material and paper in preparation II Carbon and nitrogen pools in thermokarst-affected permafrost landscapes in Arctic Siberia 2.1 Abstract 2.2 Introduction 2.3 Material and methods 2.3.1 Study area 2.3.2 Field Work 2.3.3 Laboratory analysis 2.3.4 Landform classification and upscaling C and N pools 2.4 Results 2.4.1 Sedimentological results 2.4.2 Sampling site SOC and N stocks 2.4.3 Upscaling: Landscape SOC and N stocks 2.4.4 Radiocarbon dates 2.5 Discussion 2.5.1 Site specific soil organic C and N stock characteristics 2.5.2 Upscaling of C and N pools 2.5.3 Sediment and organic C accumulation rates 2.5.4 Characterizing soil organic carbon 2.5.5 The fate of organic carbon in thermokarst-affected yedoma in Siberia 2.6 Conclusions III Impacts of successive thermokarst lake stages on soil organic matter, Arctic Alaska 3.1 Abstract 3.2 Plain language summary 3.3 Introduction 3.4 Study site 3.5 Methods 3.5.1 Core collection 3.5.2 Biogeochemical analyses 3.5.3 Study area OC and N calculation 3.6 Results 3.6.1 Biogeochemistry 3.6.2 Sediment organic carbon and nitrogen stocks 3.6.3 Radiocarbon dates and carbon accumulation rates 3.6.4 Landscape C and N budget 3.7 Discussion 3.7.1 Impact of thermokarst lake dynamics on organic matter storage 3.7.2 High organic C and N stocks on the ACP 3.7.3 Landscape chronology 3.7.4 Organic matter accumulation 3.7.5 Future development 3.8 Conclusions IV Sedimentary and geochemical characteristics of two small permafrost-dominated Arctic river deltas in northern Alaska 4.1 Abstract 4.2 Introduction 4.3 Study area 4.4 Material and Methods 4.4.1 Soil organic carbon and soil nitrogen storage 4.4.2 Radiocarbon dating and organic carbon accumulation rates 4.4.3 Grain size distribution 4.4.4 Scaling carbon and nitrogen contents to landscape level 4.5 Results 4.5.1 Carbon and nitrogen contents 4.5.2 Radiocarbon dates and accumulation rates 4.5.3 Grain size distribution 4.5.4 Arctic river delta carbon and nitrogen storage 4.6. Discussion 4.6.1 Significance of carbon and nitrogen stocks in Arctic river deltas 4.6.2 SOC and SN distribution with depth 4.6.3 Sedimentary characteristics 4.6.3.1 Accumulation rates 4.6.3.2 Sediment distribution 4.6.4 Impacts of future changes 4.6.5 Significance of remotely sensed upscaling results 4.7 Conclusions V Soil carbon and nitrogen stocks in Arctic river deltas - New data for three Western Alaskan deltas 5.1 Abstract 5.2 Introduction 5.3 Study sites 5.4 Methods 5.5 Results and discussion 5.5 Conclusions VI Discussion 6.1 Interregional comparison 6.2 Changing thermokarst landscapes and their global impact 6.3 A growing C and N data base 6.4 Outlook - potential follow-up projects VII Synthesis VIII References Appendix A Synthesis of SOC and N inventories Appendix B Supplementary material to Chapter II Appendix C Supplementary material to Chapter III Appendix D Supplementary material to Chapter IV Appendix E Supplementary material to Chapter V Appendix F Arctic river delta data set - Version 1.0 Acknowledgements - Danksagung

Note: Dissertation, Universität Potsdam, 2014 , Table of contents I - Abstract II - Zusammenfassung Chapter 1 - Introduction 1.1. Introduction 1.1.1 Motivation 1.1.2 Organisation of thesis 1.1 Scientific background 1.2.1 Arctic and wetland bryophytes 1.2.2 Bryophyte remains as palaeo-environmental indicators 1.2.3 Regional setting 1.3 Objectives ofthe thesis 1.4 Overview of the manuscripts 1.5 Contribution of the authors Chapter 2 - Manuscript #1 Abstract 2.1 Introduction 2.2 Geographic setting 2.3 Materials and methods 2.3.1 Fieldwork 2.3.2 Radiocarbon dating 2.3.3 Geochemical, stable carbon isotope, and granulometric analyses 2.3.4 Analyses of moss remains and vascular plant macrofossils 2.3.5 Pollen analysis 2.3.6 Diatom analysis 2.3.7 Statistical analysis 2.4 Results 2.4.1 High-resolution spatial characteristics oft the investigated polygon and vegetation pattern 2.4.2 Geochronology and age-depth relationships 2.4.3 General properties of the sedimentary fill 2.4.4 Bioindicators 2.4.5 Characterization oftwo different types of polygon pond sediment 2.5. Discussion 2.5.1 Small-scale spatial structure of polygons 2.5.2 Age-depth relationships 2.5.3 Proxy value of the analysed parameters 2.5.4 The general polygon development 2.5.5 Polygon development as a function of external controls and internal adjustment mechanisms 2.6 Conclusions Chapter 3 - Manuscript #11 Abstract 3.1 Introduction 3.2 Material und methods 3.2.1 Regional setting 3.2.3 Field methods and environmental data collection 3.2.4 Data analysis 3.3 Results 3.3.1 Major characteristics of the investigated polygons 3.3.2 Vegetation cover and its relationships with micro-relief and vegetation type 3.3.3 Vegetation alpha-diversity and its relationship with micro-relief and vegetation type 3.3.4 Vegetation composition and its relationship with micro-relief and vegetation type 3.4 Discussion 3.4.1 Patterns of cover, alpha-diversity and compositional turnover of vascular plants and bryophytes along the rim-pond transect (local-scale) 3.4.2 Patterns of cover, alpha-diversity and compositional turnover of vascular plants and bryophytes along the regional-scale forest-tundra transect 3.4.3 Indicator potential ofvascular plant and bryophyte remains from polygonal peats for the reconstruction of local hydrological and regional vegetation changes 3.4.4. Implications of the performed vegetation transect studies for future Arctic warming 3.5 Acknowledgements 2.4.4 Bioindicators 2.4.5 Characterization of two different types of polygon pond sediment 2.5. Discussion 2.5.1 Small-scale spatial structure of polygons 2.5.2 Age-depth relationships 2.5.3 Proxy value of the analysed parameters 2.5.4 The general polygon development 2.5.5 Polygon development as a function of external controls and internal adjustment mechanisms 2.6 Conclusions Chapter 3 - Manuscript #II Abstract 3.1 Introduction 3.2 Material und methods 3.2.1 Regional setting 3.2.3 Field methods and environmental data collection 3.2.4 Data analysis 3.3 Results 3.3.1 Major characteristics of the investigated polygons 3.3.2 Vegetation cover and its relationships with micro-relief and vegetation type 3.3.3 Vegetation alpha-diversity and its relationship with micro-relief and vegetation type 3.3.4 Vegetation composition and its relationship with micro-relief and vegetation type 3.4 Discussion 3.4.1 Patterns of cover, alpha-diversity and compositional turnover of vascular plants and bryophytes along the rim-pond transect (local-scale) 3.4.2 Patterns of cover, alpha-diversity and compositional turnover of vascular plants and bryophytes along the regional-scale forest-tundra transect 3.4.3 Indicator potential of vascular plant and bryophyte remains from polygonal peats for the reconstruction of local hydrological and regional vegetation changes 3.4.4. Implications of the performed vegetation transect studies for future Arctic warming 3.5 Acknowledgements Chapter 4 - Manuscript #3 Abstract 4.1 Introduction 4.2 Material and methods 4.2.1 Sites 4.2.2 Sampling 4.2.3 Investigated moss species 4.2.4 Measurements 4.2.5 Statistical Tests 4.3 Results 4.4 Discussion Chapter 5 - Discussion 5.1 Bryophytes of polygonal landscapes in Siberia 5.1.1 Modern bryophytes in the Siberian Arctic 5.1.2 Biochemical and isotopic characteristics of mosses 5.1.3 Reliability and potential of fossil bryophyte remains as palaeoproxies 5.2 Dynamics of low-centred polygons during the late Holocene 5.3 Outlook Appendix I - Preliminary Report Motivation Material and methods Results and first interpretation Appendix II Additional tables and figures of manuscript #1 Appendix III Additional figures of manuscript #2 Appendix IV - Quantitative approach of Standard Moss Stem (SMS3) Bibliography Acknowledgements Eidesstattliche Erklärung

Note: Inhaltsverzeichnis Kurzfassung Abstract 1 EINFÜHRUNG 2 GRUNDLAGEN 2.1 Allgemeine Zirkulation 2.2 Barokline Instabiltät 2.3 Horizontale und vertikale Wellenausbreitung 2.4 Die Rolle der Arktis im Klimasystem 2.5 Einfluss des klimatischen Wandels in der Arktis auf die mittleren Breiten 2.6 Atmosphärisches Energiespektrum und Skalenwechselwirkung 3 DATEN UND METHODEN 3.1 Verwendete Reanalyse- und Modelldaten 3.1.1 ERA-Interim 3.1.2 AFES 3.1.3 Aufteilung der verwendeten Daten in Zeiträume mit hoher und niedriger Meereisbedeckung 3.2 Methoden 3.2.1 Instabilitätsanalyse für einen zonalgemittelten Grundzustand 3.2.2 Identifikation bevorzugter großskaliger Zirkulationsmuster 3.2.3 Energie- und Enstrophiespektren 3.2.4 Statistische Testverfahren 4 ERGEBNISSE 4.1 Instabilitätsanalyse für einen zonalgemittelten Grundzustand . 4.1.1 Klimatologie ERA-Interim und AFES 4.1.2 Sensititvitätsstudie 4.2 Identifikation bevorzugter großskaliger Zirkulationsmuster 4.2.1 September 4.2.2 Oktober 4.2.3 November 4.2.4 Dezember 4.2.5 Januar 4.2.6 Februar 4.2.7 Zugehörige synoptisch-skalige Aktivität und 2m-Temperatur 4.3 Kinetische Energiespektren und nichtlineare Wechselwirkungen 4.3.1 Kinetische Energiespektren 4.3.2 Enstrophiespektren 4.3.3 Nichtlineare Energiewechselwirkungen, Energieflüsse und Enstrophieflüsse 5 ZUSAMMENFASSUNG UND AUSBLICK Tabellenverzeichnis Abbildungsverzeichnis Variablen und Symbole Literaturverzeichnis Danksagung Anhang A.1 Hough-Funktionen und vertikale Strukturfunktionen A.2 Zugeordnete Legendre-Polynome und Kugelflächenfunktionen Eidesstattliche Erklärung

Note: Dissertation, Universität Potsdam, 2015 , Table of Contents Acknowledgements Abstract Zusammenfassung List of figures and tables List of Abbreviations 1. Introduction 1.1. Preface and thesis organization 1.2. Research motivation and relevance 1.3. Background knowledge 1.3.1. Terrigenous sediments 1.3.2. Hala Lake 1.3.3. The North Pacific 1.3.4. The Bering Sea 1.4. Aims and objectives 1.5. Methodological overview 1.5.1. Fieldwork 1.5.2. Age-depth modeling 1.5.3. Key proxies: grain size and clay minerals 1.5.4. Supplementary methodology: remote sensing, seismic sub-bottom profiling and geochemistry 1.6. Overview and status of the manuscripts 2 Manuscript 1 : Linkages between Quaternary climate change and sedimentary processes in Hala Lake, northern Tibetan Plateau, China Abstract 2.1. Introduction 2.2. Regional setting 2.3. Materials and methods 2.3.1. Remote sensing of the study area 2.3.2. Fieldwork 2.3.3. Radiocarbon dating of recovered sediment cores 2.3.4. Laboratory work 2.3.5. Statistical data treatment 2.4. Results and interpretation 2.4.1. Remote sensing on the spatial heterogeneity of lake ice and length of lake ice-free days 2.4.2. Seismic sub-bottom profiling 2.4.3. Age and sedimentary characteristics of the sediment core record 2.4.4. Grain-size modeling results 2.5. Discussion 2.5.1. Last Glacial Maximum (~24-17 cal. ka BP) 2.5.2. Time-equivalent of Heinrich Event 1 (~17-15.4 cal. ka BP) 2.5.3. Time-equivalent of Bolling-Allerod (~15.4-13 cal. ka BP) 2.5.4. Time-equivalent of Younger Dryas (~12.9-11.6 cal. ka BP) 2.5.5. Holocene (~11.6 cal. ka BP to present) 2.6. Conclusions Acknowledgments 3 Manuscript 2: Modern modes of provenance and dispersal of terrigenous sediments in the North Pacific and the Bering Sea: Implications and perspectives for palaeoenvironmental reconstructions Abstract 3.1. Introduction 3.2. Study area and regional setting 3.3. Material and methods 3.4. Results 3.4.1. Grain size distribution 3.4.2 Bulk mineralogy 3.4.3. Mineralogy of the clay fraction 3.5. Discussion 3.5.1. Sedimentary processes 3.5.2. Sediment provenance 3.5.3 Implications for palaeoenvironmental studies 3.6. Conclusions Acknowledgements 4 Manuscript 3: Provenance and dispersal of terrigenous sediments in the Bering Sea slope: Implications for late glacial land-ocean linkages Abstract 4.1. Introduction 4.2. Regional setting 4.3. Material and methods 4.4. Results and interpretation 4.4.1. Lithology and stratigraphy 4.4.2. Grain size distribution 4.4.3. Clay mineralogy 4.5. Discussion 4.5.1. Processes of terrigenous sediment supply 4.5.2. Detrital sediment sources 4.5.3. Detrital sediment supply and its relation to regionalpalaeoenvironmental changes 4.5.3.1. Time interval 32-15.7 ka BP: Background sedimentation at low sea level 4.5.3.2. Time interval 15.7-14.5 ka BP: Regional Meltwater Pulse 4.5.3.3. Time interval 14.5-12.9 ka BP: First biological bloom event 4.5.3.4. Time interval 12.9-6 ka BP: Cooling episode, rejuvenation of biological productivity and onset ofmodern conditions 4.5.4. Palaeoenvironmental implications 4.6. Conclusions Acknowledgements 5 Synthesis 5.1. The North Hemisphere synchronization of millennial climate oscillations during the last Glacial: teleconnections from Westerlies and thermohaline Circulation 5.2. The regional asynchronization of millennial climate oscillations during the last Glacial: discrepancy and "recording capacity" 5.3. Secondary connections between global climate transmissions: winter cyclone in the North Pacific 5.4. Future perspectives 6 References 7 Appendix Extended results: Core SO202-39-3 from the mid-latitude North Pacific 7.1. Material 7.2. Results 7.3. Oscillation of eolian sediment transport 7.4. Conclusions

Note: Dissertation, Universität Potsdam, 2021 , Table of Contents Acknowledgements Abstract Zusammenfassung List of figure List of tables List of abbreviation Chapter 1 1. Introduction 1.1 Research background 1.1.1 Response of mountain plant diversity to climate change 1.1.2 Response of Arctic vegetation composition and diversity to climate change 1.1.3 Understanding the critical mechanisms of community assembly are essential for sustaining ecosystem services 1.1.4 Pollen analysis as a traditional tool for representing palaeovegetation 1.1.5. Sedimentary ancient DNA (sedaDNA) is a useful tool for Quaternary ecology tracking 1.2 Study area 1.3 Aims and objectives 1.4 Structure of the thesis 1.4.1 Overview of the chapter 1.4.2 Author's contributions 1.4.3 Methods Chapter 2 2 Manuscript 1: Sedimentary ancient DNA reveals warming-induced alpine habitat loss threat to Tibetan Plateau plant diversity 2.1 Abstract 2.2 Introduction 2.3 Results and discussion 2.4 Methods 2.5 Acknowledgements · Chapter 3 3 Manuscript 2: Holocene vegetation and plant diversity changes in the north-eastern Siberian treeline region from pollen and sedimentary ancient DNA 3.1 Abstract 3.2 Introduction 3.3 Materials and methods 3.3.1 Study area 3.3.2 Lake sediment cores and subsampling 3.3.3 Dating 3.3.4 Pollen analysis 3.3.5 DNA extraction and amplification 3.3.6 Sequencing filtering and taxonomic assignment 3.3.7 Statistical analyses 3.4 Results 3.4.1 Chronology 3.4.2 SedaDNA and pollen assemblages 3.4.3 Gradient analysis and correlation analysis 3.5 Discussion 3 .5.1 Contributions of pollen and sedaDNA to vegetation reconstruction and taxon richness 3.5.2 Variation in Holocene vegetation composition in the Omoloy area, north-eastern Siberia 3.5.3 SedaDNA-based plant diversity changes within lake catchments of the Omoloy region 3.6 Conclusions 3.7 Acknowledgements Chapter 4 4 Manuscript 3: Vegetation reconstruction from Siberia and Tibetan Plateau using modern analogue technique - comparing sedimentary ancient DNA (sedaDNA) and pollen data 4.1 Abstract 4.2 Introduction 4.3 Materials and methods 4.3.1 Sites ofthe modern analogues 4.3.2 Sedimentary (ancient) DNA collection 4.3.3 Metabarcoding data processing and filtering 4.3.4 Pollen data collection 4.3.5 Numerical analysis 4.4 Results 4.4.1 Modern training-set, ROC curve analyses and AT results 4.4.2 Modern analogues for Lake Naleng and Omoloy lake II 4.4.3 Vegetation type reconstruction based on MAT 4.4.4 Projecting fossil samples in ordination space of modern assemblages 4.4.5 Comparing past and present intertaxa relationships 4.5 Discussion 4.5.1 Assessment of analogue quality using modem training-sets 4·5·2 Comparison of sed(a)DNA-based and pollen-based vegetation reconstruction for the Lake Naleng, Tibetan Plateau 4.5.3 Comparison of sedDNA based and pollen-based vegetation reconstruction for the Lake Omoloy, northern Siberia 4.6 Conclusions 4.7 Acknowledgements Chapter 5 5 Manuscript 4: Terrestrial-aquatic ecosystem links on the Tibetan Plateau inferred from sedaDNA shotgun sequencin 5.1 Abstract 5.2 Introduction 5.3 Results 5.4 Discussions 5.5 Methods 5.6 Acknowledgments Chapter 6 6 Synthesis 6.1 The ability of metabarcoding and metagenomic shotgun sequencing to reveal ecological community pattern 6.2 Driver of plant diversity change in high altitude and high latitudes 6.3 High-altitude and high-latitude vegetation type change 6.4 Past terrestrial and aquatic ecological change at ecosystem-scale 6.5 Conclusions and outlook Appendix 1 Appendix-1 Materials for Manuscript #1 1.1 Appendix discussion: Contamination in NTC6 2. Appendix-2 Materials for Manuscript #2 3. Appendix-3 Materials for Manuscript #3 4. Appendix-4 Materials for Manuscript #4 References Eidesstattliche Erklarung

Note: Dissertation, Universität Potsdam, 2021 , Contents List of Figures List of Tables I Preamble 1 Introduction 1.1.1 The Journey from Weather to Climate 1.1.2 The Climate Background 1.1.3 Pollen as Quantitative Indicators of Past Changes 1.2 Overview and Aims of Manuscripts 1.2.1 List of Manuscripts 1.2.2 Short Summaries of the Manuscripts 1.3 Author Contributions to the Manuscripts II Manuscripts 2 Comparing estimation of techniques for temporal Scaling 2.1 Introduction 2.2 Data and Methods 2.2.1 Scaling estimation methods 2.2.2 Evaluation of the estimators 2.2.3 Data 2.3 Results 2.3.1 Effect of Regular and Irregular Sampling 2.3.2 Effect of Time series length 2.3.3 Application to database 2.4 Discussion 2.5 Conclusions 3 Land temperature variability driven by oceans at millennial timescales 4 Variability of surface climate in simulations of past and future 4.1 Introduction 4.2 Data and Method 4.2.1 Model simulations 4.2.2 The Last Glacial Maximum experiment 4.2.3 The mid Holocene experiment (midHolocene) 4.2.4 The warming experiments 1pctCO2 and abrupt4xCO2 4.2.5 Preprocessing of model simulations 4.2.6 Comparisons across the ensemble 4.2.7 Diagnosing variability changes 4.2.8 Changes in precipitation extremes 4.2.9 Timescale-dependence of the variability changes 4.3 Results 4.3.1 Hydrological sensitivity across the ensemble 4.3.2 Changes in local interannual variability 4.3.3 Changes in modes of variability 4.3.4 Circulation patterns underlying extratropical precipitation extremes 4.3.5 Changes in. the spectrum of variability 4.4 Discussion 4.4.1 Changes in climate variability with global mean temperature 4.4.2 Temperature vs. precipitation scaling 4.4.3 Comparison to climate reconstructions and observations 4.4.4 Limitations 4.5 Conclusions 5 Holocene vegetation variability in the Northern Hemisphere 5.1 Introduction 5.2 Data and Methods 5.2.1 Pollen Database 5.2.2 Principal Component Analysis 5.2.3 Timescale-dependent Estimates of Variability 5.2.4 Biome Classification 5.3 Results 5.3.1 General Vegetation Variability Analysis 5.3.2 Comparison of Forested and Open Land Vegetations 5.3.3 Comparison of Broadleaf and Needleleaf Fore ts 5.3.4 Comparison of Temperate and Boreal Coniferous Forests 5.3.5 Comparison of Evergreen and Deciduous Boreal Forests 5.4 Discussion 5.5 Conclusion III Postamble 6 General discussion and conclusion 6.1 Overview 6.2 Timescale-Dependent Estimates of Variability 6.3 Climate and Vegetation Variabilities in the Holocene 6.4 Implications for the 21th Century 6.5 Outlook IV Appendix A Supplementary figures from "Comparing estimation techniques for temporal scaling in paleo-climate timeseries" A.1 Block Average Results A.2 First-Order Correction for the Effect of Interpolation A.3 Change in Bias and Standard Deviation B Methods and supplementary information from "Land temperature variability driven by oceans at millennial timescales" B.1 Methods B.1.1 Reconstructions B.1.2 Significance Testing B.1.3 Testing for Anthropogenic Impacts B.1.4 Instrumental Data B.1.5 Model Data B.1.6 Spectral Estimates B.1.7 Variance Ratios B.1.8 Sub-Decadal Variability Binning B.1.9 Correlation B.1.10 Moran's I B.2 Supplementary Information B.2.1 Tree Ring Data Analysis B.2.2 Energy-Balance Equations B.3 Extended Data Figures C Supplementary figures from "Variability of surface climate in simulations of past and future" D Supplementary figures from "Characterization of holocene vegetation variability in the Northern Hemisphere" Bibliography