ALBERT

Description: SummaryA winter barley variety, from the USSR, 13031, lacking vernalization requirement but sensitive to short days and more frost resistant than varieties now commercially grown in Britain, was hybridized with Shimabara, a Japanese variety which requires vernalization but is less sensitive to short days and more susceptible to frost than 13031. The resulting segregating population was then selected under short days and non-vernalizing conditions and in artificial freezing tests, in order to isolate recombinant lines.Five lines derived in this way, together with the parents and the standard winter barley variety, Maris Otter, were included in a detailed developmental study made in a glasshouse under cool, short days.Four of the lines had very similar developmental characters, namely few leaves and a high rate but short duration of spikelet initiation. Developmentally these selections resembled the parent Shimabara more closely than 13031 but the rate of spikelet initiation was faster than that of either parent and the duration of spikelet initiation was shorter. The fifth selection also resembled Shimabara more closely than 13031 but had more leaves and a lower rate and longer duration of spikelet initiation than the other selections. This selection was found to have a strong vernalization requirement.One selection closely resembled 13031 in its frost resistance but had low short-day sensitivity and no detectable vernalization requirement. The complementary characters of the two parents were therefore recombined in this line.Vernalization had little effect on the development of any of the genotypes under cool, short days in a glasshouse.

Description: SummaryYields of two spring barley varieties, Golden Promise and Maris Mink, were greater at the Scottish Plant Breeding Station (SPBS) than at the Plant Breeding Institute (PBI) in both 1976 and 1977. In both seasons more ears per plant were produced and the number of grains per ear was greater at the SPBS. Weight per grain was greater at the Scottish Plant Breeding Station in both 1976 and 1977, but the distribution of grain size along the ears differed between seasons. Grain at the tip of the ears of plants grown at SPBS in 1976 was lighter than the corresponding grain on the Cambridge-grown plants. In 1977 the SPBS-grown plants had heavier grains at all the main shoot spikelet positions. At the SPBS the relative difference between main shoot and tillers 1 and 2 was smaller than at the PBI.These results are taken to indicate that the combination of climatic factors in Scotland affects plant development in such a way that intraplant competition is reduced and more spikelet primordia survive to form grains and more tillers produce fertile ears.

Description: SummaryAn analysis of development of Golden Promise and Maris Mink spring barley grown in England and in Scotland was made in 1976 and 1977. In both seasons the differences between varieties at a site were small. Rates of leaf emergence and primordium production and the number of primordia formed and the number which died were almost the same at both sites in 1976. In 1977 both leaf emergence and ear initiation started later in Scotland and the rates of leaf emergence and spikelet initiation were slower. Fewer spikelet primordia were formed in Scotland than in England in 1977 but a higher proportion survived to produce potentially fertile florets. Tillers emerged at the same leaf stage of the main shoot and the frequency of tillering differed at some tiller positions between England and Scotland, but the number of tillers produced was similar at both sites.These results show that in Scotland the effect of longer photoperiod which enhances the rate of leaf emergence and primordium production compensates for the lower temperature which slows down these processes. The lower temperature in Scotland particularly in 1977 during the period of spikelet primordium death increased the proportion of spikelets that survived.

Description: SummaryA number of commercial varieties and advanced breeding lines of wheat and of barley were each sown successively in the field from early in September until late in February in 3 or 2 years. Shoot apex development was monitored throughout the growing season and the dates at which the double ridge stage and the terminal spikelet stage were attained are reported. There was considerable variation in the date at which these stages occurred, associated both with variety and date of sowing. When sown early, spring wheat varieties and the winter wheat, Fenman, developed more rapidly than the other winter wheats, but the difference disappeared in sowings made in the middle of October or later. Spring barley developed more quickly than winter barley and the difference persisted until sowings made in mid-November.Length of the longest leaf sheath, number of emerged leaves on the main shoot and the time when stem elongation began (‘ear at 1 cm’), plant characters used to assess the stage for various agronomic treatments, were measured in parallel with apex development. The relationships between number of emerged leaves and the length of the longest leaf sheath and stage of development were found to vary with sowing date. The stage ‘ear at 1 cm’ provided a good guide to shoot apex development. Stem length and number of elongated internodes varied with date of sowing. Some of the variation in number of emerged leaves at a given stage and in the final number of elongated internodes was found to be correlated with total number of leaves on the main shoot. The form of analysis used indicated that sowing date may have important effects, via its effect on the number of leaves on the main shoot, on the duration of ear growth in wheat and barley and on the duration of ear formation in barley.

Description: SUMMARYLeaf emergence, apex development stage, internode length and Zadoks principal growth stage 3 were measured over 3 years at several sites.Internode elongation and Zadoks score were strongly related to the number of emerged culm leaves. The final length of the most basal internode was very variable and contributed to variation in the relationship between Zadoks stage 30 and the number of emerged culm leaves. Variation in the length of the basal internode was related to the final number of culm leaves. Most plants had six culm leaves but the number of leaves was affected by sowing date. In an experiment where sowings were made from September to March, stem elongation and Zadoks stage 30 started at a later stage of apex development in later sowings.Recognition and prediction of culm elongation and number of emerged culm leaves is important for the application of growth regulator and fungicides. Combined with functions to predict the rate of leaf emergence and final number of leaves, the relationships described in this paper may enable Zadoks principal growth stage 3 and number of emerged culm leaves to be predicted.

Description: SUMMARYEvapotranspiration from spring barley plots grown at a range of densities was estimated using the neutron probe technique. Early in the season rainfall led to percolation and kept the soil at field capacity, but from the beginning of June significant differences were found in the cumulative evapotranspiration. As the crop canopy was complete by this time in all densities, it seems probable that evapotranspiration at this time was dependent on rooting density. Although the total water use at the end of the season was similar at all densities, the more rapid use earlier in the season at higher densities may have led to a reduction in growth at the grain filling stage.

Description: SUMMARYThe growth and development of the spring wheat varieties Kolibri, Sirius and TB 435 ranging in height from relatively tall to semi-dwarf were compared when sown at 50, 200 and 800 plants m−2, with and without a spray application of CCC at the six-leaf stage. Growth analysis measurements were made throughout the season. Yield and yield components were determined at final harvest.A number of varieties x density and variety x CCC interactions were found which support the hypothesis that genotypic variation and environmentally induced differences in GA metabolism affect the agronomic performance of the crop.

Description: SUMMARYData from experiments with spring wheat, variety Kolibri, were used to examine size gradients within the mature ear in relation to ear development. The greatest number of grains and florets per spikelet, and the heaviest grains, occurred in the lower-mid part of the ear. Within the spikelets in this region grain 2 〉 grain 1 〉 grain 3.It is proposed that the timing and period of ear development and the number of spikelets can be described quantitatively in terms of (1) the rate of leaf initiation, (2) the number of leaves, (3) the rate of spikelet initiation and (4) the total number of primordia. Both leaf and spikelet initiation proceeded at more or less constant rates, but spikelet initiation was about three times as fast as leaf initiation. Differentiation of the spikelet primordia started at about the time the terminal spikelet was initiated, occurring first in the spikelets in lower-mid part of the ear. The rate of floret initiation was lower than that of spikelet initiation, but did not differ between spikelets. A maximum of nine florets per spikelet was formed and then, in all spikelets, some of the last-formed primordia died, leaving two to four potentially fertile florets.Morphologically, primordia in the lower-mid part of the ear were always the most advanced. The more rapid development of the terminal primordia tended to synchronize events such as meiosis and anthesis, thus shortening the development period of the later formed florets. Changes in meristem size were such that both length and diameter were the greatest when the primordia of the lower-mid part of the ear were initiated.The statistical model to describe primordium initiation and ear size is compared with other models describing primordium initiation rates. The analysis of ear development in quantitative terms is discussed in relation to analyses of ear development by morphological stages. A correlation between grain size and time of floret initiation is demonstrated and the significance of the size changes in the meristem is discussed.